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This study examines parameters, causes, and spatio-temporal patterns of handaxe shape variation from Tabun cave in the Levant, and Khall Amayshan 4B and Khabb Musayyib in northern Arabia. These assemblages span the range of most pointy to most rounded handaxes found anywhere during the Acheulean. The AGMT3D program is used to conduct high resolution geometric morphometric analysis of handaxe form from 3D models. Shape variation is tested against blank type, allometry, and reduction intensity. None of these factors appears to be a strong influence, but there are significant assemblage-wise differences in form, suggesting the different shapes were intentionally produced. The analysis quantifies a pattern of high diversity in the assemblages from Tabun versus low diversity in the shorter occupations at the Arabian sites. We suggest possible explanations of emerging specificity in utilitarian functions, as well as the manifestation of social identities in artefacts at the end of the Acheulean. Introduction Materials and Methods Results Discussion

The early part of Marine Isotope Stage (MIS) 3 (ca. 60–50 ka) is a crucial period for studying human demography and behaviour in south-west Asia, and how these relate to climatic changes. However, the archaeological and palaeoenvironmental records for MIS 3 in critical areas such as the Arabian Peninsula remain poorly developed. Here, we present findings from the Al Marrat basin in the Nefud desert, which provides the first clear evidence for both increased humidity and human occupation of the interior of northern Arabia during early MIS 3. A Middle Palaeolithic assemblage, dated by optically stimulated luminescence to ca. 55 ka, was found stratified within a sequence of relict palustrine deposits indicative of shallow water body formation in the Al Marrat basin. Hominin presence in northern Arabia at this time coincides with the intensification and northward displacement of monsoon rainfall systems during a period of maximum insolation. These findings add to a growing corpus of palaeoenvironmental evidence, which indicates that the Arabian interior was neither arid nor unpopulated during early MIS 3, and that hydrodynamic responses to enhanced moisture availability facilitated demographic expansions into the Arabian interior.

Microlith production is a distinctive and significant stone tool technology. However, inter-regional comparative analyses of microlithic industries are rare, and have tended to homogenise these industries by focussing analytical attention on retouched tool typologies alone. This thesis provides the first demonstration and exploration of variability in two of the earliest microlithic industries in the world – the Howiesons Poort of southern Africa and the Late Palaeolithic of South Asia. Analysis of this variation has implications for the long-standing debates concerning modern human behaviour and dispersals. In order to assess variability in underlying technological processes and manufacturing trajectories, detailed attribute analyses were conducted on lithic assemblages. Metric and qualitative variables were recorded on cores, debitage and tools from three southern African Howiesons Poort sites (Rose Cottage Cave and Umhlatuzana, South Africa, and Ntloana Tsoana, Lesotho) and four South Asian Late Palaeolithic sites (Batadomba-lena and Kitulgala Beli-lena, Sri Lanka, and Patne and Jwalapuram 9, India). Analysis of the results reveals variability within sites, over time, and between sites and regions, demonstrating that microlith production is not a homogenous technology. Underlying technological processes are shown to differ more between regions than do retouched tool forms. It is argued that this pattern is more parsimoniously explained by independent innovation of microlithic technology situated within local lithic traditions, rather than by cultural diffusion. Additionally, the exploration of variability in microlithic assemblages highlights the benefits of using a methodological approach to the modern human behaviour debate which focusses on technological variability rather than the presence of particular tool types. It is this behavioural and technological variability that is key to understanding our species.

The Jwalapuram Locality 9 rockshelter in southern India dates back to 35 000 years ago and it is emerging as one of the key sites for documenting human activity and behaviour in South Asia. The excavated assemblage includes a proliferation of lithic artefacts, beads, worked bone and fragments of a human cranium. The industry is microlithic in character, establishing Jwalapuram 9 as one of the oldest and most important sites of its kind in South Asia.

BACKGROUND The relationship between frequency and severity of pain symptoms and site, stage and morphological characteristics of endometriotic lesions was analysed in a multicentre cross-sectional observational study. METHODS A total of 469 women (median age 31 years, range 18-45) who met the following criteria were consecutively observed in the participating centres during the study period: age 18-45 years, first laparoscopic or laparotomic diagnosis of endometriosis, pain symptoms lasting > or = 6 months, pain as the main or only complaint of the condition, absence of pelvic anomalies and no previous pelvic surgery. Dysmenorrhoea and non-menstrual pain were evaluated using a multidimensional verbal rating scale. The women were requested to grade the severity of dysmenorrhoea, non-menstrual pelvic pain and deep dyspareunia using a 10-point linear analogue scale. RESULTS Dysmenorrhoea was present in 77% of subjects with ovarian endometriosis, 88% of those with endometriosis of the peritoneum, 92% of subjects with endometriosis of both ovary and peritoneum and in all the subjects with endometriosis of rectovaginal septum. These differences were not statistically significant after Bonferroni's correction. No marked difference emerged between the severity of dysmenorrhoea and site of endometriosis, but women with ovarian endometriosis tended to have lower scores (not significant). No clear association emerged between frequency and severity of non-menstrual pain, dyspareunia and site of endometriosis and the presence and severity of dysmenorrhoea, non-menstrual pain and dyspareunia. Dyspareunia was more frequently reported in women with only atypical endometriosis (56.8%) versus 47.7% in women with typical endometriosis, but with borderline significance (P = 0.05). Dyspareunia occurred in 68.2% of patients with both typical and atypical lesions. CONCLUSIONS The results of this study find no clear-cut association between stage, site or morphological characteristics of pelvic endometriosis and pain.

Recent archaeological investigations in Sri Lanka have reported evidence for the exploitation and settlement of tropical rainforests by Homo sapiens since c. 48,000 BP. Information on technological approaches used by human populations in rainforest habitats is restricted to two cave sites, Batadomba-lena and Fa-Hien Lena. Here, we provide detailed study of the lithic assemblages of Kitulgala Beli-lena, a recently excavated rockshelter preserving a sedimentary sequence from the Late Pleistocene to the Holocene. Our analysis indicates in situ lithic production and the recurrent use of the bipolar method for the production of microliths. Stone tool analyses demonstrate long-term technological stability from c. 45,000 to 8,000 years BP, a pattern documented in other rainforest locations. Foraging behaviour is characterised by the use of lithic bipolar by-products together with osseous projectile points for the consistent targeting of semi-arboreal/arboreal species, allowing for the widespread and recurrent settlement of the wet zone of Sri Lanka. Introduction Mobility and hunter-gatherer tool-kits Kitulgala Beli-lena rockshelter Materials and methods Results - Late Pleistocene (c. 45,000–31,000 cal BP) - Terminal Pleistocene (17,157–11,314 cal BP) - Holocene (10,577–8,029 cal BP) Summary Discussion

The Armenian highlands encompasses rugged and environmentally diverse landscapes and is characterized by a mosaic of distinct ecological niches and large temperature gradients. Strong seasonal fluctuations in resource availability along topographic gradients likely prompted Pleistocene hominin groups to adapt by adjusting their mobility strategies. However, the role that elevated landscapes played in hunter-gatherer settlement systems during the Late Pleistocene (Middle Palaeolithic [MP]) remains poorly understood. At 1640 m above sea level, the MP site of Kalavan 2 (Armenia) is ideally positioned for testing hypotheses involving elevation-dependent seasonal mobility and subsistence strategies. Renewed excavations at Kalavan 2 exposed three main occupation horizons and ten additional low densities lithic and faunal assemblages. The results provide a new chronological, stratigraphical, and paleoenvironmental framework for hominin behaviors between ca. 60 to 45 ka. The evidence presented suggests that the stratified occupations at Kalavan 2 locale were repeated ephemerally most likely related to hunting in a high-elevation within the mountainous steppe landscape. info:eu-repo/semantics/publishedVersion

Revealing the temporal and geographic pattern of human and animal dispersal and migration has been a major goal within anthropology, archaeology, and palaeontology. Here, I focus on the use of ancient DNA to delve beneath the migration palimpsests. More specifically, I firstly describe the use of ancient DNA derived from archaeological pig remains as a proxy to understand the initial dispersal of farmers into Europe, the back migration into Anatolia, and the spread of people into the Pacific. I then show how ancient DNA from foxes in the northern and southern hemisphere has been used to understand their dispersal onto islands in the absence of landbridges. These case studies highlight the value of DNA derived from long-dead organisms to detail the pattern and process of ancient dispersals.

Despite ecological, anthropological, and archaeological debate surrounding their desirability as habitats for human occupation, tropical rainforests have received relatively little attention in discussions of Homo sapiens' Pleistocene dispersal. Sri Lanka has yielded some of the earliest dated fossil and material culture evidence (c. 38-35,000 cal. years BP) for our species in a modern rainforest context beyond Africa. Nevertheless, assertions in Sri Lanka, and elsewhere, regarding early human rainforest reliance have been largely based on coarse or 'off-site' palaeoenvironmental records, and the overall role of these environments in human subsistence strategies has remained uncertain. This study applies stable carbon and oxygen isotope analysis to Sri Lankan human fossil, and associated faunal, tooth enamel dated to between 36-29,000 and 3,000 cal. years BP, in order to directly test human rainforest resource reliance, reconstruct a stable isotope ecology, and develop 'on-site' palaeoenvironmental records for Late Pleistocene-Holocene Sri Lankan rainforest foragers. Stable carbon and oxygen isotope analysis of modern Sri Lankan primates, and stable carbon, oxygen, and nitrogen isotope analysis of modern plant samples from the Polonnaruwa Nature Sanctuary, are also performed to investigate the ecology of Sri Lankan primates on which Late Pleistocene-Holocene forager subsistence strategies were focused. The results demonstrate that Homo sapiens relied on rainforest resources in Sri Lanka from c. 36-29,000 cal. years BP until the Iron Age c. 3 cal. years BP, even when open environments, and their corresponding resources, were available. This remains the case through periods of evident environmental change at the Last Glacial Maximum and even upon the arrival of agriculture in the island's tropical forests. The primate stable isotope data prove difficult to interpret as ecological niche separation in the absence of observation data. Nonetheless, humans were evidently able to not only use but also rapidly specialise in the exploitation of South Asia's rainforests.