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Bivalves of the genus Gigantopecten Rovereto, 1899 (Bivalvia: Pectinida) are common in the Middle Miocene (Badenian) deposits of the Central Paratethys (e.g., Schultz, 2001 ; Mandic, 2004 ; Studencka, 2019 and references therein). Large-sized species Gigantopecten (originally: Pecten) nodosiformis (Pusch, 1837) is particularly well researched, as it can be clearly recognized in the sediment, due to its large shells with nodes on top of radial ribs. It is recorded in the Miocene deposits of the Atlantic and Mediterranean provinces at least since the Early Miocene (e.g., Studencka, 2019 and references therein), and it is widespread in the Middle Miocene (Badenian) deposits of the Central Paratethys (e.g., Studencka, 2019 and references therein ; Mandic, 2004). At the last (8th) NCSEE Workshop, this scallop was chosen for the workshop logo (Studencka, 2019). These facultatively mobile scallops are also recorded from several Middle Miocene localities in Northern Croatia, e.g., on the Medvednica Mt., Zrinska Gora Mt., Ivanščica Mt., Moslavačka Gora Mt. and Psunj Mt., and part of these specimens are today housed in the Croatian Natural History Museum (CNHM) in Zagreb. Findings of G. nodosiformis in the Badenian deposits of Northern Croatia are so far best described in detail in Kochansky (1944), where the author recorded more than 20 specimens at several localities on the Medvednica Mt., near Zagreb. These specimens are also stored in the CNHM. During the 2020 we conducted the field work with the biology student E. Gujić in the area of Klenovnik, Hrvatsko Zagorje, searching for the possible topic of his master thesis. From the two outcrops of the Miocene sedimentary rocks, we collected several new specimens of this species. Pectinid shells were collected from the biocalcarenites with corraline algae. We plan to conduct a further research, which will give more insight into the paleoceology and paleogeography of this area in the Northern Croatia. Klenovnik Miocene calcarenites were deposited during the beginning of the Middle Miocene (Badenian) at the southwestern margin of the Central Paratethys. These findings will be very helpful in the correlation with the neighbouring Badenian localities and research on possible migration corridors during the Badenian in the Central Paratethys. As described in Studencka (2019 and references therein), the oldest occurrences of this species in the Paratethys area are recorded from the Karpatian deposits of southwestern Moravia, Austria and Hungary. In Austria it was also found in the earliest Badenian (Mandic, 2004). G. nodosiformis occurs in Polish and Ukrainian part of the Carpathian Foredeep Basin in the Lower Badenian deposits, with no records from the Upper Badenian deposits, while in Austria and Slovakia this species is recorded also in the Upper Badenian. Findings in Romania range from the Lower to the Upper Badenian. Mikuž (2009) lists the previous findings of these large scallops in Slovenia and describes a rare finding of a complete specimen of G. nodosiformis with both valves from Dobruška Vas in Dolenjska, associated with the Middle Miocene (Badenian) "Lithothamnion limestones". The comparison among the localities in the vicinity of the area of the assumed „Trans-Tethyan“ corridor gives a further insight into the possible open marine corridors and migration routes during the Badenian, similar to those proposed by Kováč et al. (2017).

Adamussium colbecki is one of the most common bivalve mollusks found in the coastal waters of the Antarctic continent. Its widespread distribution, easiness of collection and sensitivity to slight alterations of water temperature and presence of pollutants make this cold-adapted scallop a potentially interesting sentinel species for the biomonitoring of the impact of anthropic activities on the Antarctic benthic communities. However, while the availability of genetic and molecular data would represent a resource of the utmost importance to enable the use of this species as a model for environmental studies, this data is presently nearly non-existing. Here we report a high quality de novo assembled and annotated transcriptome for A. colbecki, discussing the long-debated phylogenetic position of this species within the order Pectinida through a Bayesian phylogenomics approach, based on the concatenated multiple sequence alignment of 978 universally conserved orthologous genes.

Dijkstra, Henk H., Beu, Alan G. (2018): Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae). Records of the Australian Museum (Rec. Aust. Mus.) 70 (2): 113-330, DOI: 10.3853/j.2201-4349.70.2018.1670, URL: http://dx.doi.org/10.3853/j.2201-4349.70.2018.1670

In order to enhance the understanding of Pacific Colombia’s deep-water marine fauna, a benthic research cruise (2012 TUM Offshore 6 and 7) was conducted off the coast of the Department of Nariño, in southern Colombia. Biological, oceanographic and sediment samples from the continental shelf and slope were collected at depths between 350 and 941 m. A new species of Malletia obtained on that cruise is described and compared with other species from the eastern Pacific. Sixteen species of bivalve mollusks (belonging to 12 families and 15 genera) were identified. Five of them were the first records for Pacific Colombia (Jupiterialobula, Limatulasaturna, Lucinomaheroica, Cuspidariapanamensis, and Dallicordiaalaskana). Four of them had geographic distributions that now extend to Tumaco at the southern end of Nariño.

Complicachlamys Iredale, 1939 Complicachlamys Iredale, 1939: 362. Type species (by original designation): Complicachlamys wardiana Iredale, 1939; Recent, QLD, Australia. Diagnosis. Elongate, medium-sized Pedini with 7–9 primary radial plicae and numerous secondary radial costae; valves flattened, subequilateral; interstitial commarginal sculpture present, but no shagreen microsculpture; auricles very unequal, anterior large, posterior small; hinge teeth small, with obsolete resilial teeth and prominent dorsal teeth, intermediate teeth lacking; byssal notch deep, ctenolium well-developed. Distribution. Recent. Tropical southwestern Pacific, living in the littoral zone, bysally attached to rocks or other hard substrates. Discussion. Iredale (1939: 362) introduced Complicachlamys and included several “representative” species, viz. Complicachlamys fulvicostata A. Adams & Reeve, 1850 [sic], Complicachlamys luculenta Reeve, 1853 [sic] = Complicachlamys dringi Reeve, 1853 [sic], and Complicachlamys crouchi Smith, 1892 [sic] [all the authors and years should have been placed between parentheses]. All these species have shagreen microsculpture and should be placed in Semipallium [Jousseaume] Lamy, 1928. Complicachlamys wardiana does not have this peculiar microsculpture, although Iredale (1939: 362) mentioned sculpture in the interstices of the secondary radial costae “producing a honeycomb effect”, which is somewhat similar to the shagreen sculpture of Semipallium species. Complicachlamys wardiana has interstitial commarginal sculpture of curved lamellae, producing hollow sections on most specimens. Other characters (outline, primary plicae and secondary costae, highly unequal auricles, deep byssal notch, well-developed ctenolium) are almost indistinguishable from those of Semipallium, and it is possible that C. wardiana is simply a species of Semipallium that has lost shagreen sculpture. This would be resolved best by comparison of DNA sequences. Hertlein (1969: N366) treated Complicachlamys as a junior synonym of Semipallium, and was followed by Vaught (1989: 119). Wagner (1989b: 111) considered Complicachlamys to be a junior synonym of Chlamys Röding, 1798., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 227, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Smith, E. A. 1892. Descriptions of new species of shells from Mauritius and California. Annals and Magazine of Natural History, Series 6, 9: 255 - 256. https: // doi. org / 10.1080 / 00222939208677314","Lamy, E. 1928. Les peignes de la Mer Rouge (d'apres les materieux recueillis par le Dr Jousseaume). Bulletin du Museum national d'Histoire naturelle 34: 166 - 172.","Vaught, K. C. 1989. A Classification of the Living Mollusca, ed. R. T. Abbott and K. J. Boss. Melbourne, Florida: American Malacologists Inc.","Wagner, H. P. 1989 b. Taxonomy and nomenclature of the genus Complicachlamys Iredale, 1939, and its species (Bivalvia, Pectinidae). Basteria 53: 111 - 116."]}

Mimachlamydini Waller, 1993 Diagnosis. Pedinae with chlamydoid, aequipectinoid or pectinoid shape, simple primary radial sculpture of solid plicae with (in some taxa) secondary and tertiary flanking costellae; microsculpture of simple, smooth, oval pits in pre-radial stage, antimarginal or divaricated (herringbonelike) interstitial microsculpture on remainder of shell; true shagreen microsculpture never present, although some taxa have “pseudo-shagreen” sculpture in which cells are not united across dorsal surface; internal rib carinae near ventral margin of most taxa; hinge with prominent resilial and dorsal teeth; deep byssal notch and prominent functional ctenolium in late ontogeny of almost all taxa., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 274, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Waller, T. R. 1993. The evolution of \" Chlamys \" (Mollusca: Bivalvia: Pectinidae) in the tropical western Atlantic and eastern Pacific. American Malacological Bulletin 10: 195 - 249."]}

Mimachlamys funebris (Reeve, 1853) Figs 86D,G, 88E,G, 92 Pecten funebris Reeve, 1853: sp. 85, pl. 22, fig. 85 (not a junior homonym of Chlamys funebris Lacordaire, 1848, Insecta; ICZN, 1999: 60, Article 57.8); E.A. Smith, 1884: 116; Küster & Kobelt, 1888: 171, pl. 47, fig. 6. Chlamys funebris (Reeve).–Wells & Bryce, 1988: 160, fig. 588; Wells, Slack-Smith & Bryce, 2000: 42. Chlamys (Mimachlamys) funebris (Reeve).– Rombouts, 1991: 28, pl. 11, figs 2–2a; Lamprell & Whitehead, 1992: [22], pl. 9, fig. 54. Mimachlamys funebris (Reeve).–Beu & Darragh, 2001: 181, fig. 47E; Slack-Smith & Bryce, 2004: 237; Raines & Poppe, 2006: 270, 273, upper figs; pl. 217, figs 1–5; Huber, 2010: 210. Type data. Type material not traced in NHMUK. Type locality: Bathurst Island, Western Australia. The illustrated articulated specimen, from WA, Broome, Entrance Point (WAM S55490; Figs 86D, G, 88E, G) is here designated the neotype of Pecten funebris Reeve, 1853. Additional material examined. —AUSTRALIA: WESTERN AUSTRALIA: Cardabia, S of Point Cloates, 23°06'S 113°48'E, dead (1 v, C.375604); Point Cloates, 22°43'S 113°40'E, dead (1 v, C.375605); Exmouth Gulf, 17 ml S of Exmouth townside, 22°13'S 114°06'E, dead (1 v, C.093621; 1 v, C.375607); North West Cape, Lighthouse Beach, 21°49'S 114°11'E, dead (2 v, C.100740); Turtle Beach, W side of North West Cape, 21°48'S 114°10'E, dead (4 v, C.375606); S Muiron Island, off SW coast, 21°41.55'S 114°18.17'E, dead, 19–20 m (1 v, WAM S12834); S Muiron Island, off NE coast, 21°40.46'S 114°20.91'E, dead, 3–5 m (1 v, WAM S12785; 1 v, WAM S12836); S Muiron Island, off North West Point, 21°39.45'S 114°20.22'E, dead, 11–13 m (1 v, WAM S12835); Onslow Beach, 21°38'S 115°07'E, dead (1 v,C.119519); Onslow area, 21°38'S 115°07'E, dead (2 v, C.375608); Onslow, W side of Peak Island, 21°36'S 114°30'E, alive, 18 m (4 pr + 2 v, WAM); Bundegi Reef, 21°51'S 114°10'E, dead (1 v, WAM); Exmouth, Bundegi Reef, 21°15'E 114°11'E, alive (3 pr, C.344047); Exmouth, Bundegi Reef, 21°51'S 114°11'E, alive, 0–3 m (1 pr, C.344046); 40 ml S of Dampier, 21°0'S 116°06'E, dead (1 v, C.375609); Passage Island, Sholl Island, NE side, 20°57'S 113°53'E, dead, intertidal (1 pr, WAM); between Cape Dupuy and Cape Malouet, Barrow Island, 20°40'-42'S 115°25'E, dead (1 v, WAM); Montebello Islands, Buttercup Island, 20°29.16'S 115°32.04'E, dead, intertidal (1 pr, WAM); Montebello Islands, Stephenson Channel near S end of Hermite Island, 20°28'21"S 115°32'27"E, dead, 4 m (2 v, WAM S12838);Montebello Islands, E side of Trimouille Island,dead (3 v, WAM);Montebello Islands,S of Gannet Island to E of Delta Island,stn MB#12, 20°27'S 115°33.59'E, dead, 4–6 m (1 v, WAM S12974); Montebello Islands, 20°26'S 115°32'E, alive (1 pr, C.049666;1 pr,C.132070);Montebello Islands, near rock islands,SW end of Bluebell Island, 20°23'30"S 115°31'04"E,dead, 0–9 m (1 v, WAM S12839);Montebello Islands, 20°21'16"S 115°30'53"E, inside reef of North-West Island, dead, 3–4 m (1 v, WAM S12837); Dampier Archipelago, Rosemary Island, 20°29'S 116°35'E,alive (2 pr, WAM);DampierArchipelago,NE corner of Delambre Island,approx. 20°26'S 117°10'E, alive (1 pr, WAM); Dampier Archipelago, N end of Flying Foam Passage, 20°28'S 116°50'E, alive (1 pr, WAM); Port Hedland, 20°18'S 118°35'E,alive (1 pr, WAM); Port Hedland, 20°18'S 118°35'E, alive (1 pr, C.097536; 1 pr,C.119518);Lagrange, 18°37'S 121°43'E, alive (1 pr, WAM);Lagrange Bay, 18°38'S 121°42'E, alive (2 pr,C.103817); c. 230 ml W of Roebuck Bay, 18°30'S 118°03'E, dead, 238 m (1 v, C.165163);Broome, 17°58'S 122°14'E,alive (3 pr, WAM);Broome, Entrance Point, 18°01'S 122°12'E,alive, intertidal (4 pr, WAM);Broome, Entrance Point, 18°01'S 122°12'E,dead (3 v,C.119521; 2 pr, C.121369); Broome, Roebuck Bay, 18°0'S 122°15'E, alive (1 pr, C.097537; 1 pr, C.132071; 3 pr,C.344037); Gantheaume Point, W of Broome, 17°59'S 122°11'E, dead (1 pr, C.057197);Black Ledge,Roebuck Bay, S of Broome, 17°59'S 122°17'E, dead (4 pr,C.303776);Broome, 17°58'S 122°14'E, alive 0–16 m (1 v, C.075235; 3 v, C.375613; 1 pr,C.375612);Broome, Cable Beach, 17°56'S 122°12'E,dead (5 v,C.375610);Rowley Shoals, Imperieuse Reef, 17°31'S 118°56'E, dead (2 v, WAM);Kimberley,Beagle Bay, 16°56'S 122°32'E, dead (many v, C.119520; 10 v, C.119522); Kimberley, Beagle Bay, 16°52'S 122°32'E, dead, 9 m (1 pr, WAM); King Sound, 16°50'S 123°30'E, alive (2 pr, C.097538); Kimberley, Sunday Island, Derby, 16°25'S 123°11'E, dead (1 pr + 1 v, WAM); Kimberley, Sunday Island, S side, 16°26'S 123°09'E, dead, 23 m (1 v, WAM); Cape Leveque, 16°24'S 122°55'E,alive (2 v, C.160997; 1 pr, C.061584; 1 v, C.375614; 1 pr, C.095818; 1 v, C.375615). Description. Shell up to c. 70 mm high, most specimens smaller, to c. 50 mm; dorsoventrally elongate, weakly inflated, almost equally convex, equivalve, almost equilateral, anterior auricles much larger and longer than posterior ones, umbonal angle c. 85–90°; most specimens brownish, a few orange, red or yellow; mottled in juvenile stage, more uniform in adults. Both valves sculptured with c. 20–25 evenly spaced, squamose radial plicae; a narrow secondary riblet present on each side of each plica, commencing on central part of disc of mature shells. Anterior auricle of left valve with c. 10 squamose radial riblets, weaker and fewer (c. 6) on posterior auricle; anterior auricle of right valve with 4–5 prominent radial riblets, much weaker on posterior. Dorsal margin somewhat declined on posterior auricle. Byssal notch deep, byssal fasciole rather broad. Functional ctenolium well-developed, with 4–6 teeth. Interior radially furrowed, internal rib carinae prominent in narrow zone around ventral margin. Resilial and dorsal teeth prominent. Dimensions. Illustrated specimen: WA, Broome, Entrance Point (WAM S55490): H 48.4, L 42.4 mm. Habitat. Living in the upper littoral zone, byssally attached to undersides of rock slabs on hard reef at minus tide levels only. Usually slabs are in shallow pools or in almost dried out gutters (pers. comm. H. Morrison, 2000). Distribution. Mainly along the shores of northern Western Australia, from Shark Bay to the Kimberley. Present specimens alive in the intertidal zone to 18 m. Remarks. This species easily could be confused in the juvenile stage with Mimachlamys punctata (Gmelin, 1791), a common species from the tropical Southwest Pacific, also occurring in the tropical region of Australia, although more offshore than M. funebris in Western Australia. The two species can be distinguished by size, sculpture and colour (Table 7)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 282, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Lacordaire, J. T. 1848. Monographie des coleopteres subpentameres de la famille des phytophages. Tome second. Memoires de la Societe Royale des Sciences de Liege 5: i - vi, 1 - 890.","Smith, E. A. 1884. Mollusca. In Report on the Zoological Collections Made in the Indo-Pacific Ocean During the Voyage of the H. M. S. \" Alert \" 1881 - 2, pp. 34 - 116, 487 - 508. London: Trustees of the British Museum.","Rombouts, A. 1991. Guidebook to Pecten Shells. Recent Pectinidae and Propeamussiidae of the World [edited and revised by H. E. Coomans, H. H. Dijkstra, R. G. Moolenbeek and P. L. van Pel]. Oegstgeest: Universal Book Services."]}

Parvamussium slacksmithae sp. nov. Figs 10G–I, 11 Holotype (lv) AM C.157685, Western Australia, 72 n. ml NNW of Dampier, 19°28.9'– 19°29.0'S 116°29.4'– 116°29.0'E, dead, 110 m, coll. B. W. Jenkins, 26.X.1983 (FRV Soela stn 26-18), (H 5.1 mm, L 6.0 mm). Paratypes (64, lv + rv): 50 AM C.209769, 14 ZMA Moll.409002; paratypes from the same locality. Largest paratype (lv): H 5.9 mm, L 6.5 mm. Additional material examined. — AUSTRALIA: WESTERN AUSTRALIA: 85 n. ml NNW of Port Hedland, 19°00.4'– 19°03'S 118°01'E, dead, 116–120 m, sand & gravel, FRV Soela stn 29-26, coll. B.W. Jenkins, 29.X.1983 (1 v, C.157713); 94 n. ml NNE of Port Hedland, 18°48'S 119°0'E,dead, 92–94 m, FRV Soela stn 23-06, coll. B. W. Jenkins, 23.X.1983 (10 v, C.157658). QUEENSLAND: SE of Swain Reefs, 22°20'09"– 22°26'16"S 153°17'05"E, dead, 187 m, HMAS Kimbla stn 7, coll. P. H. Colman, VI.2008 (5 v, C.462541). Description. Shell small, up to c. 6 mm high, fragile, translucent, left and right valves equally convex, almost circular, inequivalve, equilateral, auricles almost equal in size, umbonal angle c. 90°. Left valve with small milky white maculations, right valve with milky white dot umbonally. Both valves smooth and glossy, with traces of commarginal growth lines on left valve, some specimens with weak, closely spaced commarginal lamellae and rudimentary radial sculpture on anterior and posterior ends of disc. Anterior auricle of left valve smooth, some specimens with weak commarginal sculpture near periphery. Anterior auricle of right valve with one radial riblet near pseudo-fasciole, posterior auricle smooth. Dorsal margin straight. Byssal notch shallow. Interior surface with rudimentary short riblets, two anteriorly and two posteriorly. Discussion. Parvamussium slacksmithae sp. nov. is somewhat similar to P. torresi (E. A. Smith, 1885) from northeastern Australia, but differs in size (P. slacksmithae is c. 6 mm high, P. torresi is larger, up to c. 9 mm high), in external sculpture (right valve of P. slacksmithae is smooth, right valve of P. torresi has weak commarginal lamellae) and internal sculpture (P. slacksmithae has a few rudimentary riblets, P. torresi has 10 prominent, well-developed radial riblets). Parvamussium vesiculatum Dijkstra, 1995 from New Caledonia also has rudimentary internal riblets, somewhat similar to the present species, but differs strongly externally (P. slacksmithae is almost smooth, P. vesiculatum is prominently sculptured) (Dijkstra, 1995b: 39). Habitat. The present specimens are so far only dead-taken on the continental shelf on soft sediment. Distribution. Northwestern Australia near Port Hedland from 18°48'– 19°29'S, and 116°29'– 119°0'E, also from Queensland, taken dead at 92– 187 m. Remarks. It is possible that the present species was collected alive at the type locality. Specimens are very fresh and soft parts are often lacking after sorting the material, as the valves become separated easily due to the weak external ligament. Specimens from Queensland have more prominent commarginal sculpture near the disc flank, but other characters are similar to the type material. Etymology. Named after Mrs Shirley Slack-Smith, formerly curator of the Mollusca Section of the WAM, who generously supported this project and assisted in many ways., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on pages 140-141, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Smith, E. A. 1885. Report on the Lamellibranchiata collected by H. M. S. Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of H. M. S. Challenger During the Years 1873 - 76 […] Zoology 13 (35): 1 - 341."]}

Key to species of Cryptopecten from Australia 1 Shell c. 30 mm high, thin, almost circular, weakly inflated, rv more convex than lv, weakly prosocline, auricles unequal, valves with 17–20 squamose, bi-vesicular radial plicae, byssal fasciole narrow........................................................................................................... C. bullatus —— Shell with 18–22 vesicular-sculptured radial plicae.................................................................. 2 2 Shell c. 20 mm high, solid, circular to subcircular, moderately to strongly inflated, rv more convex than lv, auricles highly unequal, valves with 18–22 evenly spaced, vesicular-sculptured radial plicae, tripartite when worn, byssal fasciole wide.................................................. C. nux

Juxtamusium Iredale, 1939 Juxtamusium Iredale, 1939: 368. Type species (by original designation): Juxtamusium oblectatum Iredale, 1939 (= Pecten (Chlamys) coudeini Bavay, 1903); Recent, 0.5 miles W of North Direction Isle, QLD, in 20 fathoms [37 m]. Diagnosis. Small, thin-shelled, subcircular to circular Decatopectinini with depressed, unevenly to evenly spaced, low radial costae; narrowly spaced, delicate commarginal microsculpture throughout ontogeny; resilifer oblique, strongly erect on one side, strong internal hinge ligament, hinge plate very flat, lacking crura or denticles; ctenolium on both valves (see Waller, 1984: 211), byssal notch shallow. Distribution. Early Miocene–Recent [Juxtamusium pseudojamviniensis (Eames & Cox, 1956: 41), early Miocene, Zanzibar; Waller (2006a) agreed that this belongs in Juxtamusium]. Tropical Indo-West Pacific, living in the littoral to sublittoral zones amongst coral rubble on soft sediment. Discussion. Hertlein (1969: N357) treated Juxtamusium as a subgenus of Chlamys Röding, 1798 in the Chlamys group. Waller (1986: 40) raised Juxtamusium to a genus in Decatopectinini., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 197, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Waller, T. R. 1984. The ctenolium of scallop shells: functional morphology and evolution of a key family-level character in the Pectinacea (Mollusca: Bivalvia). Malacologia 25: 203 - 219.","Waller, T. R. 2006 a. New phylogenies of the Pectinidae (Mollusca: Bivalvia): reconciling morphological and molecular approaches. In Scallops: Biology, Ecology and Aquaculture. Second Edition, ed. S. E. Shumway and G. J. Parsons, pp. 1 - 44. Amsterdam: Elsevier. https: // doi. org / 10.1016 / S 0167 - 9309 (06) 80028 - 1","Waller, T. R. 1986. A new genus and species of scallop (Bivalvia: Pectinidae) from off Somalia, and the definition of a new tribe Decatopectinini. The Nautilus 100: 39 - 46. https: // doi. org / 10.5962 / bhl. part. 26491"]}

Scaeochlamys Iredale, 1929 Scaeochlamys Iredale, 1929: 162. Type species (by original designation): Pecten lividus Lamarck, 1819; Recent, southwestern, southern and southeastern Australia. Diagnosis. Pedini attached by byssus, prosocline or prosogyrate in form, right valve flattened (distorted in most specimens), left valve convex; macrosculpture of unevenly spaced, strongly scaly, primary radial plicae and secondary interstitial riblets at least in early ontogeny; prominent interstitial shagreen microsculpture on anterior auricle of left valve and on disc at least in early ontogeny; byssal notch deep, ctenolium well-developed. Hinge with prominent resilial and dorsal teeth. Interior weakly plicate. Distribution. Pleistocene [e.g., Japan (Masuda, 1962), Red Sea (Cox, 1929)]–Recent. (Sub)tropical Indo-West Pacific to temperate waters of the NW and SW Pacific, living in the intertidal to sublittoral zones, byssally attached to rocks, gravel or other hard substrates. Discussion. Hertlein (1969: N355) treated Scaeochlamys as a junior synonym of Chlamys Röding, 1798, placed in the suprageneric Chlamys group. Vaught (1989: 118) followed Hertlein. Waller (1993: 203) recognized Scaeochlamys as a genus in Chlamydini (i.e., Pedini)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 255, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1929. Mollusca from the continental shelf of eastern Australia. No. 2. Records of theAustralian Museum 17 (4): 157 - 189. https: // doi. org / 10.3853 / j. 0067 - 1975.17.1929.759","Lamarck, J. B. P. A. de M. de. 1819. Histoire Naturelle des Animaux sans Vertebres […]. Vol. 6 (1). Paris: Lamarck.","Masuda, K. 1962. Tertiary Pectinidae of Japan. Science Reports of the TohokuUniversity, Series 2, Geology 33: 117 - 238.","Vaught, K. C. 1989. A Classification of the Living Mollusca, ed. R. T. Abbott and K. J. Boss. Melbourne, Florida: American Malacologists Inc.","Waller, T. R. 1993. The evolution of \" Chlamys \" (Mollusca: Bivalvia: Pectinidae) in the tropical western Atlantic and eastern Pacific. American Malacological Bulletin 10: 195 - 249."]}

Key to species of Glorichlamys from Australia 1 Shell c. 20 mm high, circular to oblong, flattened to weakly inflated, 9–13 radial costae, 1–3 intercalated radial riblets, interstitial commarginal lamellae, byssal notch deep, byssal fasciole broad........................................................................................ G. elegantissima —— Shell with quadripartite radial costae........................................................................................ 2 2 Shell c. 25 mm high, subcircular to oblong, valves weakly inflated, radial costae quadripartite, byssal notch shallow, byssal fasciole narrow.................................................................................................... G. quadrilirata, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 192, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Key to genera of Propeamussiidae occurring in Australia 1 Shell with auricles equal in size, lacking byssal notch in adult, internal riblets commencing early, and external sculpture weak.......................... Propeamussium —— Shell with unequal auricles, and with byssal notch and internal riblets......................................................................................................................................... 2 2 Shell prominently commarginally sculptured on rv, internally with riblets commencing after early growth stage.............................................. Parvamussium —— Shell of most specimens lacking internal riblets....................................................................... 3 3 Shell prominently radially and/or commarginally sculptured externally, with vesicles on many specimens; internal riblets lacking from most specimens, rudimentary on a few.............................................. Cyclopecten —— Shell lacking commarginal sculpture on rv............................................................................... 4 4 Shell with raised anterior auricle and broad hinge dentition................................... Similipecten —— Shell commarginally undulated................................................................................................. 5 5 Shell flattened, commarginally sculptured on both valves..................................... Catillopecten, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 121, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Ylistrum Mynhardt & Alejandrino, 2014 Ylistrum Mynhardt & Alejandrino in Mynhardt et al., 2014: 407. Type species (by original designation): Pecten balloti Bernardi, 1861; Pliocene to living, Southwest Pacific. Diagnosis. Amusiini with (sub)circular shape, up to c. 140 mm high, very weakly inflated, both valves slightly convex; umbonal angle c. 140°; exterior surface of disc and auricles smooth, interior with c. 42–54 narrow, prominent radial riblets arranged irregularly or in vaguely discernable pairs; auricles equal in length, intermediate hinge teeth absent, byssal notch and ctenolium weak in early ontogeny, absent later. Distribution. Middle Miocene–Recent. The earliest species referable to Ylistrum seems to be Amusium morganense Beu & Darragh (2001: 175, figs 64A, F, 65A–E), from Morgan Limestone (Balcombian Australian local stage, middle Miocene), River Murray cliffs near Blanchetown, South Australia, and Nullarbor Limestone (also Balcombian), Nullarbor Plain, southern Western Australia. Ylistrum morganense is smaller and has larger auricles and much more closely paired internal rib carinae than Y. balloti. Tropical and subtropical Indo-West Pacific, living littorally. Discussion.As discussed under Annachlamys and Amusium, we retain Ylistrum in Amusiini at present, awaiting a more final molecular phylogeny of Pectinidae., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 207, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Mynhardt, G., A. Alejandrino, L. Puslednik, J. Corrales, and J. M. Serb. 2014. Shell shape convergence masks biological diversity in gliding scallops: description of Ylistrum n. gen. (Pectinidae) from the Indo-Pacific Ocean. Journal of Molluscan Studies 80: 400 - 411. https: // doi. org / 10.1093 / mollus / eyu 038"]}

Bractechlamys Iredale, 1939 Bractechlamys Iredale, 1939: 366. Type species (by original designation): Bractechlamys evecta Iredale, 1939 (= Pecten vexillum Reeve, 1853). Recent, 0.5 ml SE of Lizard Island, QLD, 19 fathoms [35 m]; widespread in tropical western Pacific. Diagnosis. Decatopectinini with 8–15 primary radial plicae, some species with nodules on plicae; with secondary radial riblets; shape oblong to subcircular; with prominent intermediate teeth, a relatively deep byssal notch, prominent costae on auricles, and an obsolete ctenolium. Distribution. Recent. Tropical Indo-West Pacific and eastern Atlantic, living in the littoral to sublittoral zones amongst coral rubble on soft sediment. Discussion. Grau (1959: 120) and subsequently Hertlein (1969: N366) treated Bractechlamys, together with Comptopallium Iredale, 1939 and Complicachlamys Iredale, 1939, as junior synonyms of Semipallium Jousseaume in Lamy, 1928 in the Decatopecten group. Waller (1972a: 245) pointed out that Semipallium has reticulate (“shagreen”) microsculpture, lacking on Bractechlamys and its relatives, and belongs in tribe Pedini. At present Bractechlamys is treated as a valid genus, placed in Decatopectinini by Waller (1986: 40)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 181, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Lamy, E. 1928. Les peignes de la Mer Rouge (d'apres les materieux recueillis par le Dr Jousseaume). Bulletin du Museum national d'Histoire naturelle 34: 166 - 172.","Waller, T. R. 1972 a. The Pectinidae of Eniwetok Atoll, Marshall Islands. The Veliger 14: 221 - 264.","Waller, T. R. 1986. A new genus and species of scallop (Bivalvia: Pectinidae) from off Somalia, and the definition of a new tribe Decatopectinini. The Nautilus 100: 39 - 46. https: // doi. org / 10.5962 / bhl. part. 26491"]}

Minnivola Iredale, 1939 Minnivola Iredale, 1939: 363. Type species (by original designation): Minnivola isomeres Iredale, 1939 (= Ostrea pyxidata Born, 1778); Recent, Queensland. Diagnosis. Moderate-sized Mimachlamydini with a somewhat concave left valve and convex right valve, mimicking the appearance of Pecten; semi-circular, with auricles slightly unequal in size, byssal notch moderately deep, functional ctenolium present, radial macrosculpture present, shallowly pitted microsculpture on pre-radial area of left valve, reticulate “pseudo-shagreen” microsculpture on left valve at least in early ontogeny, weak commarginal microsculpture on right valve, and on left valve in late ontogeny. Internal rib carinae moderately prominent around ventral margin. Hinge teeth weak. Distribution. Miocene–Recent (Hayami, 1989: 16). Indo-West Pacific, living in the littoral zone on soft sediment. Discussion. Hertlein (1969: N371) placed Minnivola in the Pecten (Patinopecten) subgroup, merely based on the reticulate microsculpture of the left valve. Other morphological characters are similar to those of Pecten, although it is now clear that this is another confusing convergence resulting from similarity in life habit rather than from close phylogentic relationship. Patinopecten is currently placed in Pedinae, tribe Fortipectinini (see Kafanov, 1986; Waller, 1991, 2006a). The molecular phylogeny by Mahidol et al. (2007: fig. 3) placed Minnivola in the same clade as several Mimachlamys species. T. R. Waller (USNM, pers. comm. 1 July 2017) has advised us that both Minnivola and Volachlamys species have a shallowly pitted pre-radial area on the left valve, as in Mimachlamys. They also share a unique form of “pseudoshagreen” microsculpture, in which the individual cells are not closed over the outer surface to form a secondary shell surface, as occurs in true shagreen microsculpture in Pedini. In both Minnivola and Volachlamys, the internal rib carinae are weaker than in Mimachlamys species, suggesting that they occupy a primitive position in Mimachlamydini. A position within Mimachlamydini was also suggested for Volachlamys by the molecular phylogeny of Matsumoto (2003: fig. 2) and is clearly implied by the molecular phylogeny of Feng et al. (2011: figs 1–3). T. R. Waller has therefore concluded that both Minnivola and Volachlamys likely belong in Mimachlamydini, and we follow his opinon here. We have not rearranged the plates to follow this conclusion., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 294, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Kafanov, A. I. 1986. Comparison of the geographical and stratigraphical ranges of Fortipectininae and Patinopectininae (Bivalvia: Pectinidae). Monograph of the Mizunami Fossil Museum 6: 23 - 40.","Waller, T. R. 1991. Evolutionary relationships among commercial scallops (Mollusca: Bivalvia: Pectinidae). In Scallops: Biology, Ecology and Aquaculture, ed. S. E. Shumway, pp. 1 - 73. Amsterdam: Elsevier.","Waller, T. R. 2006 a. New phylogenies of the Pectinidae (Mollusca: Bivalvia): reconciling morphological and molecular approaches. In Scallops: Biology, Ecology and Aquaculture. Second Edition, ed. S. E. Shumway and G. J. Parsons, pp. 1 - 44. Amsterdam: Elsevier. https: // doi. org / 10.1016 / S 0167 - 9309 (06) 80028 - 1","Mahidol, C., U. Na-Nakorn, S. Sukmanomon, W. Yoosuk, N. Taniguchi and T. T. T. Nguyen. 2007. Phylogenetic relationships among nine scallop species (Bivalvia: Pectinidae) inferred from nucleotide sequences of one mitochondrial and three nuclear gene regions. Journal of Shellfish Research 26: 25 - 32. https: // doi. org / 10.2983 / 0730 - 8000 (2007) 26 [25: PRANSS] 2.0. CO; 2","Matsumoto, M. 2003. Phylogenetic analysis of the subclass Pteriomorphia (Bivalvia) from mtDNA COI squences. Molecular Phylogenetics and Evolution 27: 429 - 440. https: // doi. org / 10.1016 / S 1055 - 7903 (03) 00013 - 7"]}

Pedum Bruguière, 1792 Pedum Bruguière, 1792: pl. 178, figs 1–4 (ICZN, 1999: 17, Article 12.2.7; P. Bouchet, MNHN, pers. comm.). Type species (by subsequent monotypy, Lamarck, 1799: 88): Ostrea spondyloidea Lamarck, 1799 (ICZN, 1999: 73, Article 69.3) (= Ostrea spondyloidea Gmelin, 1791); Recent, tropical Indo-West Pacific. Diagnosis. Coral-embedded, dorso-ventrally elongate Pedini with a flattened left valve and weakly convex right valve; microsculpture of numerous scabrous radial riblets and closely arranged commarginal lamellae; anterior auricle strongly curved in late ontogeny, posterior auricle almost absent; hinge teeth lacking, resilial insertions high and narrow, ligament migrating strongly in postero-ventral direction; byssal fasciole broad, byssal gape very wide; ctenolium present early in ontogeny, absent late in ontogeny. Distribution. Pleistocene and Recent; Pleistocene of Massaua, Eritrea (Selli, 1973). Tropical Indo-West Pacific, living in the littoral zone embedded in massive hard corals. Discussion. Hertlein (1969: N364) placed Pedum in the suprageneric Hinnites group, Vaught (1989: 119) in Hinnitini. Habe (1977: 92) established a new subfamily Peduminae [sic, emended to Pedinae] of Pectinidae. Waller (1993: 200, 203) placed Pedum in Chlamydini and reduced Pedinae to a synonym, but we here adopt Pedinae and Pedini as the valid senior synonyms., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 253, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Lamarck, J. B. P. A. de M. de. 1799. Prodome d'une nouvelle classification des coquilles. Memoires de la Societe d'Histoire Naturelle de Paris 1: 63 - 91.","Selli, R. 1973. Molluschi quaternari di Massaua e di Gibuti. Accademia Nazionale dei Lincei, Missione Geologica dell' AGIP nella Dancalia Meridionale e Sugli Altipiani Ararini, 1936 - 1938, 4, 2. Documentazione Paleontologica, pp. 153 - 444.","Vaught, K. C. 1989. A Classification of the Living Mollusca, ed. R. T. Abbott and K. J. Boss. Melbourne, Florida: American Malacologists Inc.","Waller, T. R. 1993. The evolution of \" Chlamys \" (Mollusca: Bivalvia: Pectinidae) in the tropical western Atlantic and eastern Pacific. American Malacological Bulletin 10: 195 - 249."]}

Cyclopecten cancellus Dijkstra, 1991 Figs 15H–I,K, 16 Cyclopecten cancellus Dijkstra, 1991: 21, figs 66–70; Dijkstra, 2001: 90, figs 33–35; Dijkstra & Moolenbeek, 2008: 18; Dijkstra & Maestrati, 2008: 97; Dijkstra & Maestrati, 2013b: 474. Type data. Holotype (lv) RMNH56560, 14 paratypes (v) RMNH56561–56566. Type locality: Indonesia, Flores Sea, off SW Salayer, 6°22.4'S 120°26.3'E, dead, 130–155 m (SNELLIUS-II stn 4.153). Additional material examined. — AUSTRALIA: QUEENSLAND: off Cairns, 16°36'12"– 16°35'54"S 146°15'24"– 146°14'18"E, dead, 110–201 m (1 v, C.165464). WESTERNAUSTRALIA: S of Cowaramup Bay, near mouth of Margaret River, 33°57'S 114°59'E, dead (2 v, C.165468); Kilcarnup, N side of Margaret River, 33°57'S 114°59'E, dead, beach (1 v, C.374666); Ellensbrook, 33°55'S 115°0'E, dead, beach (2 v, WAM S12952); Cowaramup, 33°52'S 115°0'E, dead (2 v [rudimentary internal riblets], WAM S12559; 1 v, WAM S12535); Rottnest Island, 32°0'S 115°30'E, dead, beach (9 v, WAM415.91); Rottnest Island, Bathurst Point, 32°0'S 115°30'E, dead, beach (1 v, WAM414.91); Rottnest Island, Ricey Beach, 32°0'S 115°30'E, dead, beach (3 v, WAM413.91; 1 v, C.165465); Cottesloe, Cable Station, 33°0'S 115°45'E, dead (1 v, WAM481.91); NW of Beagle Island, 29°43'S 114°17'E, dead, 274–283 m (1 v, C.165466). — NEW CALEDONIA: off New Caledonia, 20°16'S 169°51'E, alive, 85–100 m (2 pr, C.165234). Figure 16. Distribution of Cyclopecten cancellus Dijkstra (circles), C. horridus Dijkstra (star), C. powelli Dell (triangle) and C. reticulatus sp. nov. (squares). Description. Shell small, up to c. 3 mm high, convex, subcircular, opaque, whitish to cream, wider than high, inequivalve, almost equilateral, left valve slightly more convex than right, auricles almost equal in size, umbonal angle c. 100°. Left valve with coarse reticulate sculpture with scaly intersections, radial riblets (c. 10) more prominent than commarginal lamellae, both widely spaced, commencing at 0.5 mm shell height and increasing in prominence ventrally, with secondary interstitial radial riblets. Auricles with 1–2 radial riblets crossed by prominent commarginal lamellae. Right valve with closely spaced commarginal lamellae. Anterior auricle with 2–4 weak radial riblets, posterior smooth. Dorsal margin straight. Byssal notch moderately deep, byssal fasciole narrow. Habitat. Living in the sublittoral and bathyal zones amongst rubble on soft bottoms. Distribution. Indonesia, 130–375 m, dead; Vanuatu, 140–175 m, dead (Dijkstra, 1991, 2001); Fiji, 500–614 m, dead (Dijkstra & Maestrati, 2008). Now also from New Caledonia (85–100 m) and Australia (beach drift and 110–283 m, dead). Remarks. The present species is similar to the type specimens from Indonesia, although some specimens from Western Australia have some rudimentary internal riblets, which are lacking in typical specimens. Cyclopecten cancellus is a new record for Australia (empty shells only)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 149, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Parvamussium retiolum Dijkstra, 1995 Figs 8C,E,K, 9 Parvamussium retiolum Dijkstra, 1995b: 29, figs 39–42, 97; Dijkstra, 2001: 85; Dijkstra & Marshall, 2008: 10, figs 8A–G, 9; Dijkstra & Maestrati, 2008: 93; Spencer et al., 2009: 198; Dijkstra & Maestrati, 2013b: 473. Type data. Holotype (pr) MNHN Moll 21171. Paratypes (39pr+6v): 35 MNHN Moll 21172, 2 C.201713, 2 ZMA Moll. 395026, 2 NMNZ M.268537, 2 NSMT-Mo70541, 2 USNM890860. Type locality: Coral Sea, Chesterfield Is., 19°47'S 158°44'E, alive, 685–700 m (MUSORSTOM 5 stn CP 363). Additional material examined. — AUSTRALIA: QUEENSLAND: SE of Swain Reefs,22°20'09"– 22°26'16"S 153°17'05"E,dead, 187 m (1 v,C.462543); E of Lady Elliott Island, 24°0'S 153°06'E, dead, 476–531 m (4 v, C.165535). WESTERN AUSTRALIA: SW of Cape Naturaliste, 33°44.5'S 114°26.1'E, dead, 183–238 m (3 v, C.165545); W of Garden Island, 32°15'S 115°03'E, dead, 250–258 m (1 v, C.165550); W of Rottnest Island, 31°0'S 114°51'E, dead (1 v, C.165548); 80 n. ml NNE of Port Hedland, 19°03.6'– 19°04'S 119°03.4'– 119°05'E,dead, 82 m (3 v, C.157711); N of Port Hedland, 18°40'S 117°55'E, dead, 150 m (1 v, C.157679); c. 240 ml NE of Broome, 14°37'S 123°40'E,dead, 80 m (19 v, C.165494 [in part]); c. 140 ml N of Cape Leveque, 14°29'S 123°03'E,dead, 124 m (many v [atypical],C.165493). CORAL SEA:Elizabeth Reef, 29°53.82'S 159°01.65'E, alive, 420 m (1 pr, C.165246). —NEW CALEDONIA: S of Ile des Pins, 22°50'S 167°34'E, dead, 274 m (many v, C.165441; 4 v, C.165439); 4 ml S of Ile des Pins, 22°50'S 167°34'E, dead, 275 m (16 v, C.165443 [in part]; 1 v, C.165442 [in part]; 5 v, C.165444). Description. Shell up to c. 16 mm high, fragile, inequivalve, inequilateral, left valve slightly more convex than right, translucent white, auricles unequal, umbonal angle c. 95°. Inner surface with 10 riblets, plus 1 posterior auricular riblet and 4 rudimentery auricular riblets on left valve and 3 on right, ends of riblets somewhat nodose. Left valve sculptured with delicate commarginal lamellae crossing closely spaced radial riblets, somwhat coarser near umbonal area than elsewhere, more delicate near ventral margin.Auricles with fine radial riblets crossed near margin by fine commarginal lamellae. Right valve with regular commarginal lirae, closely spaced near umbonal area, more widely spaced near ventral margin. Microscopic interstitial radial scratches near margins. Auricles with commarginal lamellae, more prominent anteriorly. Strongly developed scales on anterodorsal margin. Byssal notch narrow. Habitat. Living on the continental shelf and in the bathyal zone, free amongst soft sediment. Distribution. Chesterfield Islands, Coral Sea, 620–700; New Caledonia, 720–800 m (Dijkstra, 1995b, 2001); New Zealand: many lots listed by Dijkstra & Marshall (2008: 10, fig. 9) from the Norfolk Ridge N of Norfolk Island (26°25.2'S 167°11.2'E, 714–756 m, 11 v, NMNZ M.171056) to South Ritchie Trough, off Mahia Peninsula, central E North Island (39°58.59'S 178°14.18'E, alive, 990 m, 2 pr, NIWA V466), in 316–1000 m (Dijkstra & Marshall, 2008: 11, fig. 9); Fiji, 310–699 m; Tonga, 391–510 m (dead) (Dijkstra & Maestrati, 2008). Now also Queensland and Western Australia. Maximum depth range of live-taken specimens is 310–1000 m. Present specimens in 82–531 m, alive in 420 m. Remarks. Parvamussium retiolum is closely similar to P. maorium, but differs in the sculpture of the left valve (P. maorium is smooth on the central part of the disc and radially sculptured laterally, P. retiolum commarginally and radially sculptured throughout). Other morphological characters are very similar. Parvamussium retiolum is a new record for Australia., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 138, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Spencer, H. G., B. A. Marshall, and R. C. Willan. 2009. Checklist of New Zealand living Mollusca. In New Zealand Inventory of Biodiversity. Volume one. Kingdom Animalia. Radiata, Lophotrochozoa, Deuterostomia, ed. D. P. Gordon, pp. 196 - 219. Christchurch: University of Canterbury Press."]}

Semipallium Lamy, 1928 Semipallium [Jousseaume] Lamy, 1928: 169. Type species (by original designation): Pecten tigris Lamarck, 1819 (= Ostrea flavicans Linnaeus, 1758). Recent, Indo-West Pacific. Belchlamys Iredale, 1929: 164. Type species (by original designation): Pecten aktinos Petterd, 1886. Pleistocene and Recent, southern Australia. Diagnosis. Byssate Pedini with a prosocline disc, evenly or unevenly spaced strong to weak primary radial plicae, delicate secondary radial riblets in late ontogeny of most species; shagreen microsculpture throughout ontogeny; auricles very unequal in size; inner surface plicate with edges carinate close to ventral margin; byssal notch deep, ctenolium well-developed. Hinge with moderately weak resilial and dorsal teeth., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 260, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Lamy, E. 1928. Les peignes de la Mer Rouge (d'apres les materieux recueillis par le Dr Jousseaume). Bulletin du Museum national d'Histoire naturelle 34: 166 - 172.","Lamarck, J. B. P. A. de M. de. 1819. Histoire Naturelle des Animaux sans Vertebres […]. Vol. 6 (1). Paris: Lamarck.","Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis […]. Tomus I, editio decima, reformata. Holmiae [Stockholm]: L. Salvii.","Iredale, T. 1929. Mollusca from the continental shelf of eastern Australia. No. 2. Records of theAustralian Museum 17 (4): 157 - 189. https: // doi. org / 10.3853 / j. 0067 - 1975.17.1929.759","Petterd, W. F. 1886. New species of Tasmanian marine shells. Papers and Proceedings of the Royal Society of Tasmania 1885: 320 - 321."]}

Gloripallium Iredale, 1939 Gloripallium Iredale, 1939: 357. Type species (by original designation): Ostrea pallium Linnaeus, 1758; Recent, Indonesia. Diagnosis. Medium-sized Decatopectinini with 7–14 radial plicae; subcircular to circular; with prominent commarginal lamellae on plicae; secondary sculpture of radial riblets (lacking from some specimens); noduliferous radial macrosculpture on auricles, commarginal microsculpture on auricles and on disc in early growth stage of some specimens; internal rib carinae prominent; hinge with prominent dorsal teeth and weak resilial teeth; byssal notch deep, ctenolium well-developed. Distribution. Miocene–Recent (Hayami, 1989: 15). Tropical Indo-West Pacific, living in the littoral to sublittoral zones amongst coral on soft sediment. Discussion. Hertlein (1969: N357) treated Gloripallium as a subjective junior synonym of Cryptopecten Dall, Bartsch & Rehder, 1938, a subgenus of Chlamys Röding, 1798, placed in the Chlamys group. Waller (1986: 40) recognized Gloripallium as a valid genus in Decatopectinini., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 194, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis […]. Tomus I, editio decima, reformata. Holmiae [Stockholm]: L. Salvii.","Waller, T. R. 1986. A new genus and species of scallop (Bivalvia: Pectinidae) from off Somalia, and the definition of a new tribe Decatopectinini. The Nautilus 100: 39 - 46. https: // doi. org / 10.5962 / bhl. part. 26491"]}

Key to species of Anguipecten from Australia 1 Shell c. 50 mm high, lv flat and rv weakly inflated, auricles short and almost equal, 12–14 primary radial plicae, secondary radial riblets on ribs and interstices, byssal notch shallow, byssal fasciol lacking, ctenolium weakly developed......................................................... A. picturatus —— Shell with 8 primary radial plicae............................................................................................. 2 2 Shell c. 37 mm high, lv flat and rv weakly inflated, auricles almost equal, 8 primary radial plicae, secondary radial riblets in late growth stage on lirae and interstices, byssal notch distinct, byssal fasciole weak, ctenolium rudimentary........................................................... A. simoneae —— Shell with 20–25 primary radial plicae...................................................................................... 3 3 Shell c. 70 mm high, valves weakly inflated, auricles short and almost equal, 20-25 obsolete primary radial plicae, byssal notch and byssal fasciole almost lacking in adult stage, ctenolium rudimentary.............................................................................................. A. superbus, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 177, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Key to species of Mirapecten from Australia 1 Shell c. 40 mm high, roundly triangular, thin, posteriorly oblique, lv almost flat, rv inflated, 4–5 distinct unevenly spaced radial plicae (obsolete on some specimens) with variable scales on lv, 7–9 (4 bifurcated) radial plicae with variable scales (or lacking scales) on rv................................................................................. M. mirificus —— Shell with nodulous radial plicae.............................................................................................. 2 2 Shell c. 35 mm high, subcircular, radially undulated, weakly inflated, almost equivalve and equilateral, valves with 8–9 nodulous radial plicae, secondary radial riblets in interspaces.................................................... M. moluccensis —— Shell with evenly spaced radial plicae...................................................................................... 3 3 Shell c. 40 mm high, subcircular, lv flattened, rv more inflated, almost equivalve and equilateral, valves with 7–9 evenly spaced radial plicae, obsolete or with widely spaced curved scales................................... M. rastellum, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 201, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Palliolinae Korobkov in Eberzin, 1960 Diagnosis. Pectinidae with an inner shell surface microstructure of irregularly foliated calcite, producing small reflective surfaces on the inner surface of well-preserved shells. Remarks. This subfamily was defined for the first time by Waller (2006a: 10, fig. 1.2) for genera related to Palliolum and Mesopeplum. Based on morphology and time ranges, he proposed that Palliolum was the stem genus of both Palliolini and Mesopeplini, and evolved from the largely Mesozoic genus Dhontichlamys during late Paleocene time., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 170, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Waller, T. R. 2006 a. New phylogenies of the Pectinidae (Mollusca: Bivalvia): reconciling morphological and molecular approaches. In Scallops: Biology, Ecology and Aquaculture. Second Edition, ed. S. E. Shumway and G. J. Parsons, pp. 1 - 44. Amsterdam: Elsevier. https: // doi. org / 10.1016 / S 0167 - 9309 (06) 80028 - 1"]}

Excellichlamys Iredale, 1939 Excellichlamys Iredale, 1939: 366. Type species (by original designation): Pecten spectabilis Reeve, 1853; Recent, unknown locality [(sub)tropical Indo-West Pacific]. Diagnosis. Small to medium-sized Decatopectinini with 9–12 radial plicae, elongate to subcircular, most species with large unequal auricles; with commarginal macro- and microsculpture, prominent noduliferous radial sculpture on auricles, short internal rib carinae, weak resilial hinge teeth (dorsal and intermediate teeth absent), byssal notch shallow, ctenolium obsolete to prominent. Distribution. Miocene–Recent (Hayami, 1989: 15). Tropical Indo-West Pacific, living in the littoral to sublittoral zones amongst coral or coral rubble on soft sediment. Discussion. Hertlein (1969: N366) treated Excellichlamys as a subgenus of Semipallium Lamy, 1928, placed in the Decatopecten group. However, Excellichlamys differs strongly in morphology from Semipallium, which has shagreen microsculpture and is placed in Pedini. Waller (1986: 40) raised Excellichlamys to a genus and placed it in Decatopectinini., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 190, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Lamy, E. 1928. Les peignes de la Mer Rouge (d'apres les materieux recueillis par le Dr Jousseaume). Bulletin du Museum national d'Histoire naturelle 34: 166 - 172.","Waller, T. R. 1986. A new genus and species of scallop (Bivalvia: Pectinidae) from off Somalia, and the definition of a new tribe Decatopectinini. The Nautilus 100: 39 - 46. https: // doi. org / 10.5962 / bhl. part. 26491"]}

Key to species of Pecten from Australia 1 Shell c. 150 mm high, (sub)circular, solid, lv flat or slightly concave, rv weakly to strongly inflated, radial macrosculpture prominent, simple, commarginal microsculpture closely spaced............................... P. fumatus —— Shell circular, thin, lv strongly concave, rv strongly inflated.................................................... 2 2 Shell c. 55 mm high, circular, thin, lv strongly concave, rv strongly inflated, radial macroculpture very weak, commarginal microsculpture weak.................................................................................................. P. dijkstrai, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 219, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Amusiini Thiele, 1934 [emend. Waller, 2006a] Diagnosis. Pectininae with weak radial macrosculpture or smooth, internal rib carinae present, disc microsculpture of closely spaced commarginal lamellae, hinge teeth welldeveloped, consisting of resilial and dorsal teeth, intermediate teeth weak or lacking, auricular crura prominent. Remarks. Waller (2006a: 22–27, fig. 1.3) separated the Amusium and Pecten groups into their own tribes for the first time because of phylogenetic separation. He considered that Amusium, Euvola Dall, 1898 and Leopecten Masuda, 1971 all evolved from Amussiopecten Sacco, 1897 (all included in Amusiini), whereas Annachlamys Iredale, 1939 and Pecten evolved from Gigantopecten Rovereto, 1899 (all included in Pectinini). There has been continuing confusion over the relative priority of Macrochlamys Sacco, 1897 and Gigantopecten, but following Waller & Bongrain (2006), the International Commission on Zoological Nomenclature (2008) accepted that Macrochlamys Sacco, 1897 is a junior homonym of Macrochlamys Benson, 1832, and adopted Gigantopecten as the valid name. The tribe name has usually been attributed to Habe (1977), but Waller (2011: 93) attributed it to Thiele (1934: 805). Waller (2007, 2011) further discussed the evolution of weakly sculptured to smooth scallops resembling Pecten and Amusium, pointing out the complex origins of the living taxa, and adding the new genus Zamorapecten Waller, 2011 in Amusiini., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 204, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Thiele, J. 1934. Handbuch der systematischen Weichtierkunde. Band 2. Jena: G. Fischer.","Waller, T. R. 2006 a. New phylogenies of the Pectinidae (Mollusca: Bivalvia): reconciling morphological and molecular approaches. In Scallops: Biology, Ecology and Aquaculture. Second Edition, ed. S. E. Shumway and G. J. Parsons, pp. 1 - 44. Amsterdam: Elsevier. https: // doi. org / 10.1016 / S 0167 - 9309 (06) 80028 - 1","Masuda, K. 1971. On some Patinopecten from North America. Transactions and Proceedings of the Palaeontological Society of Japan, new series, 83: 166 - 178.","Iredale, T. 1939. Mollusca. Part 1. British Museum (Natural History) Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5: 209 - 425.","Rovereto, G. 1899. Rectification de nomenclature. Revue critique de Paleozoologie 3: 90.","Waller, T. R. 2011. Neogene paleontology of the northern Dominican Republic. 24. Propeamussiidae and Pectinidae (Mollusca: Bivalvia: Pectinoidea) of the Cibao Valley. Bulletins of American Paleontology 381: 1 - 198.","Waller, T. R. 2007. The evolutionary and biogeographic origins of the endemic Pectinidae (Mollusca: Bivalvia) of the Galapagos Islands. Journal of Paleontology 81: 929 - 950. https: // doi. org / 10.1666 / pleo 05 - 145.1"]}

Key to species of Parvamussium from Australia 1 Shell small, c. 9 mm high, circular, almost opaque, whitish, auricles unequal, byssal notch small, lv with radial and commarginal sculpture, 3 rudimentary internal riblets............................................ P. araneum —— Shell small, subcircular, translucent white, auricles unequal, byssal notch shallow, lv with commarginal and interstitial radial scupture, 11–13 internal riblets................................................................................................. 2 2 Shell c. 7 mm high, whitish, left valve more convex than right valve, equivalve, almost equilateral, auricles dissimilar in shape, lv with commarginal lamellae and weak interstital radial riblets, rv with regularly spaced commarginal lirae, lv with 12–13 internal riblets, rv with 11 internal riblets........................................................... P. cancellorum —— Shell small, circular, hyaline, both valves smooth, 12–13 internal riblets......................................................................................................................................... 3 3 Shell c. 9 mm high, whitish, both valves weakly inflated, almost equally convex, inequivalve, equilateral, 12–13 narrow internal riblets........................................................................................................... P. lamprelli —— Shell small, oblong, hyaline, lv delicate sculptured, 10 internal riblets......................................................................................................................................... 4 4 Shell c. 14 mm high, slightly posteriorly oblique, whitish, lv delicately radially sculptured, central part almost smooth, commarginal lamellae near ventral margin, 10 or 11 internal riblets, most specimens with 10............................................................................... P. maorium —— Shell small, fragile, circular, hyaline, lv commarginally sculptured, 14 internal riblets.................................................................................................... 5 5 Shell c. 9 mm high, flattened, lv with widely spaced prominent commarginal lamellae, 14 internal riblets plus a few rudimentary ones................................................................................................. P. multiliratum —— Shell small, circular, hyaline or opaque, auricles almost equal, a few rudimentary internal riblets................................................................................................ 6 6 Shell c. 8 mm high, both valves almost smooth, lv flattened, 1–3 rudimentary internal riblets................................................................................ P. pauciliratum —— Shell small, oblong, posteriorly oblique, lv with delicate reticulate sculpture, 10 internal riblets...................................................................................... 7 7 Shell c. 16 mm high, opaque, whitish, lv with fine radial and commarginal sculpture, 10 prominent internal riblets with a few rudimentary............................................................................................................... P. retiolum —— Shell small, almost circular, lv flattened, delicately radially sculptured, 9–10 internal riblets................................................................................................ 8 8 Shell c. 10 mm high, lv stained and opaque, weakly radially and commarginally sculptured, rv almost smooth, whitish, 9–10 internal riblets, with a few rudimentary in some specimens..................................... P. scitulum —— Shell small, circular, hyaline, valves equally convex, smooth, auricles almost equal, internal riblets rudimentary................................................................... 9 9 Shell c. 6 mm high, valves smooth and glossy, with white maculations, internal riblets rudimentary, 2 anteriorly and 2 posteriorly........................................................................................................... P. slacksmithae —— Shell small, almost circular, lv sculptured with fine scaly radial riblets, 10 internal riblets......................................................................................................... 10 10 Shell c. 11 mm high, lv stained and opaque, sculptured with fine unevenly spaced scaly radial riblets, rv whitish, 10 internal riblets, with a few rudimentary ones in some specimens.................................... P. squalidulum —— Shell small, almost circular, opaque, lv heavily sculptured, 10 internal riblets.......................................................................................................................... 11 11 Shell c. 10 mm high, creamy with white spots, lv with unevenly set prominent radial sculpture and close-set commarginal lamellae, 8–10 internal riblets, 10 in most specimens................................................. P. thetidis —— Shell small, almost circular, lv smooth and glossy, 10 internal riblets....................................................................................................................................... 12 12 Shell c. 9 mm high, lv opaque and whitish, lv sculptured with only delicate commarginal lamellae near ventral margin, 10 internal riblets, with a few rudimentary in some specimens................................... P. torresi —— Shell small, almost circular, lv coarsely sculptured with vesicles, few rudimentary internal riblets.............................................................................................. 13 13 Shell c. 6 mm high, lv with commarginal lirae and radially arranged vesicles, few rudimentary internal riblets anteriorly and posteriorly.............................................................................................................. P. vesiculatus, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on pages 130-131, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Aequipectinini von Teppner, 1922 Diagnosis. Pedinae with most taxa of aequipectinoid shape, with a complex sculpture of radial costae or narrow plicae, evenly lamellose with a series of hollow sections or secondary interstitial commarginal lamellae; many taxa with commarginal ridges developing ventrally directed loops, convex in the dorsal direction and concave in the ventral direction, on flanks of radial plicae; microsculpture of prominent, deep, smooth pits in pre-radial stage; equilateral and with auricles becoming almost equal in late ontogeny; internal rib carinae prominent; hinge with enlarged resilial teeth., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 298, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Teppner, W. von. 1922. Lamellibranchiata tertiaria. Pars 15. \" Anisomyaria \" II. In Fossilium Catalogus. I: Animalia, ed. C. Diener, pp. 67 - 296. Berlin: W. Junk."]}

Key to species of Decatopecten from Australia 1 Shell c. 120 mm high, valves inequivalve and inequilateral, lv flat to weakly inflated, rv more convex, auricles subequal, 9–13 radial plicae, fine to coarse lamellae on plicae, secondary radial riblets in late growth stage, byssal notch shallow, byssal fasciole moderately wide, ctenolium weak.................................................................. D. radula —— Shell oblong, with 5–7 radial plicae.......................................................................................... 2 2 Shell c. 45 mm high, almost equally convex, equivalve and equilateral, auricles subequal, 5 radial plicae, rarely 6–7, secondary radial riblets on plicae and interstices in late growth stage, byssal notch, byssal faciole and ctenolium lacking in adult stage.............................................................................................................. D. strangei, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 185, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Veprichlamys Iredale, 1929 Veprichlamys Iredale, 1929: 164 (proposed as a subgenus of Mimachlamys). Type species (by original designation): Chlamys perillustris Iredale, 1925. Recent, off Victoria, Australia. Diagnosis. Small to medium-sized byssate Pedini of strongly prosocline shape, with narrow umbonal angle, auricles highly unequal; pre-radial antimarginal microsculpture; narrow, squamous primary radial costae, secondary costae intercalated near ventral margin (late in ontogeny on most specimens); interstitial antimarginal microsculpture present, with a few long, straight ridgelets in centre of central radial interspaces, shagreen present in a few species; shallow internal radial furrows present, rib carinae lacking; byssal notch deep, ctenolium well-developed. Hinge with moderately prominent resilial and dorsal teeth. Distribution. Miocene–Recent (Beu & Darragh, 2001: 115–118; Balcombian Australian Stage, middle Miocene). Southwestern Pacifc, southern Australia, New Zealand, and the Galapagos Islands, living in the sublittoral to bathyal zones. Discussion. Iredale (1929: 164) proposed Veprichlamys as a subgenus of Mimachlamys. Hertlein (1969: N355) treated Veprichlamys as a junior synonym of Chlamys, placed in the suprageneric Chlamys group. Waller (1993: 236; 2007: 942) considered Veprichlamys to be an extant genus of Chlamydini (i.e., Pedini), based on “its pre-radial microsculpture and lack of internal rib carinae”., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 271, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1929. Mollusca from the continental shelf of eastern Australia. No. 2. Records of theAustralian Museum 17 (4): 157 - 189. https: // doi. org / 10.3853 / j. 0067 - 1975.17.1929.759","Iredale, T. 1925. Mollusca from the continental shelf of eastern Australia. Records of the Australian Museum 14 (4): 243 - 270. https: // doi. org / 10.3853 / j. 0067 - 1975.14.1925.845","Waller, T. R. 1993. The evolution of \" Chlamys \" (Mollusca: Bivalvia: Pectinidae) in the tropical western Atlantic and eastern Pacific. American Malacological Bulletin 10: 195 - 249.","Waller, T. R. 2007. The evolutionary and biogeographic origins of the endemic Pectinidae (Mollusca: Bivalvia) of the Galapagos Islands. Journal of Paleontology 81: 929 - 950. https: // doi. org / 10.1666 / pleo 05 - 145.1"]}

Key to genera of Cyclochlamydidae occurring in Australia 1 Shell small, triangular prodissoconch on lv, smooth or radially and/or commarginally sculptured on lv, rv with hexagonal microstructure................................................................................................................. Cyclochlamys —— Shell small with a weakly inflated prodissoconch on lv........................................................... 2 2 Shell smooth or radially and/or commarginally sculptured on lv, many specimens posteriorly oblique, rv with hexagonal microstructure................................................................................................................... Chlamydella, Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 157, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017

Key to species of Serratovola from Australia 1 Shell c. 17 mm high, lv slightly concave, rv strongly convex, 19–20 broad angular radial plicae, interspaces narrow on lv, functional ctenolium weak........................................................................................... S. pallula —— Shell with flat lv, small angular plicae, equal interspaces......................................................... 2 2 Shell c. 35 mm high, lv flat, rv convex, 17–19 small angular radial plicae on lv, interspaces equal to or wider than plicae, ctenolium lacking.................................................................................................... S. rubicunda

Talochlamys Iredale, 1929 Talochlamys Iredale, 1929: 188. Type species (by original designation): Chlamys famigerator Iredale, 1925 (= Pecten pulleineanus Tate, 1887), Recent, southern and eastern Australia. Diagnosis. Small to medium-sized, byssate Pedini with unevenly spaced squamose primary radial costae or narrow plicae, and with secondary interstitial (mimachlamydoidlike) riblets in late ontogeny; microsculpture of weak antimarginal striae (also lacking in late ontogeny) and interstitial, widely spaced, prominent commarginal lamellae; shagreen microsculpture absent from post-Eocene species; internal rib carinae lacking; weak dorsal and resilial hinge teeth; byssal notch deep, ctenolium prominent. Distribution. Paleocene–Recent (Beu & Darragh, 2001; del Rio & Martinez, 2015: appendix 1). Eastern Atlantic, Indo-West Pacific, southern Australia and New Zealand, living in the littoral to bathyal zones. Discussion. Hertlein (1969: N355) treated Talochlamys as a junior synonym of Chlamys Röding, 1798, placed in the suprageneric Chlamys group. Waller (1993: 202) considered Talochlamys to be a valid genus of Chlamydini (i.e., Pedini). Beu (1995: 18) for the first time included several fossil and Recent species from New Zealand in Talochlamys (see also Beu & Darragh, 2001; Dijkstra & Marshall, 2008)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 268, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Iredale, T. 1929. Mollusca from the continental shelf of eastern Australia. No. 2. Records of theAustralian Museum 17 (4): 157 - 189. https: // doi. org / 10.3853 / j. 0067 - 1975.17.1929.759","Iredale, T. 1925. Mollusca from the continental shelf of eastern Australia. Records of the Australian Museum 14 (4): 243 - 270. https: // doi. org / 10.3853 / j. 0067 - 1975.14.1925.845","Waller, T. R. 1993. The evolution of \" Chlamys \" (Mollusca: Bivalvia: Pectinidae) in the tropical western Atlantic and eastern Pacific. American Malacological Bulletin 10: 195 - 249."]}

Similipecten colmani sp. nov. Figs 19E–I, 20 Holotype (lv) AM C.165504, Arafura Sea, Northern Territory, c. 45 ml N of Croker Island, 10°17'S 132°38'E, dead, 65 m, leg. P. H. Colman, 9 Nov 1969, MV San Pedro Sound; (H 3.0 mm, L 3.4 mm). Paratypes c. 100 (v), AM C.209771 (lv & rv), ZMA Moll.409004 (3 lv and 3 rv), same locality data as holotype. Additional material examined. — AUSTRALIA: QUEENSLAND: N Gulf of Carpentaria, 11°07'S 139°32'E, dead, 59 m (4 v, C.165497); NE Gulf of Carpentaria, 11°56'S 140°25'E,dead, 67 m (1 v,C.165495); Central Gulf of Carpentaria, 14°29.5'S 138°14'E,dead, 60 m (1 v,C.165496). WESTERN AUSTRALIA: off North West Cape, 22°51'S 113°41.2'E, dead, 78 m (2 v, C.165512); c. 190 ml NW of Port Hedland, 19°32'S 115°49'E, dead, 183 m (2 v, C.165491 [in part]); N of Port Hedland, 18°40'S 117°55'E, dead, 150 m (3 v, C.157678 [in part]); c. 135 ml NW of Roebuck Bay, 17°34'S 120°22'E,dead, 188 m (4 v,C.165508); c. 100ml NW of Broome, 16°58'S 120°47'E, dead, 194 m, 5 v (5 v, C.165510; 4 v, C.165511); c. 240 ml NE of Broome, 14°37'S 123°40'E, dead, 80 m (1 v, C.165494 [in part]). NORTHERN TERRITORY: 100 km NE of Melville Island, 10°22.5'S 131°37'E, dead, 71 m (4 v, C.165506); 68 km NE of Croker Island, 10°39'S 133°05'E, dead, 62 m (4 v, C.165507); Arnhem Land, c. 60 ml NE of Goulburn Islands, 11°08'S 134°18.5'E,dead, 50 m (1 v,C.165501); Arnhem Land, 365 km N of Miinggimbi Island, 8°48'S 134°58'E, dead, 100 m (1 v, C.165502); c. 190 ml NW of Gove, 10°16'S 135°09'E, dead, 60 m (1 v, C.165505); Arnhem Land, c. 200 ml NW of Wessel Islands, 8°39'S 135°21'E,dead, 65 m (4 v, C.165499); Arnhem Land, N of Wessel Islands, 8°26'S 135°22'E, dead, 75 m, (1 v, C.165500); c. 250 ml NE of Croker Island, 9°11'S 135°43'E,dead, 70 m (1 v, C.165505); N of Arnhem Land, 10°50'S 137°10'E, dead, 59 m (1 v, C.165498). Description. Shell small, fragile, hyaline or opaque, triangularly subcircular, somewhat wider than high, rather compressed, inequilateral, auricles unequal in size and proportions; posterior ones longer, taller, weakly differentiated from disc, with weakly concave posterior margin, anterior ones with obvious, narrow byssal notch (right valve) or shallow byssal sinus (left valve); umbonal angle c. 90°. Stained milky white. Exterior of left valve disc sculptured with weak, closely spaced commarginal lirae towards ventral margin. Anterior auricle somewhat demarcated from shell disc, bearing c. 20 fine commarginal lamellae, prominent near disc flank. Antero-marginal area of anterior auricle somewhat curved, antero-dorsal area somewhat raised. Posterior auricle weakly demarcated from disc, bearing very weak, fine, close-set commarginal lirae; postero-dorsal margin straight. Additional description (paratypes, rv). Shell up to c. 4 mm high, hyaline or opaque, dull milky white without maculations, slightly more convex than left valve, sculptured with closely spaced commarginal lamellae or lirae, also very weak or even absent. Anterior auricle sharply demarcated from disc, with semicircular anterior end, beraing very closely spaced fine lirae, absent from some specimens. Posterior auricle weakly demarcated from disc, bearing same sculpture as disc. Byssal notch narrow. Figure 20. Distribution of Similipecten colmani sp. nov. (circles), Catillopecten murrayi (Smith) (star) and C. tasmani Dijkstra & Marshall (triangle). Discussion. A similar-looking species, Similipecten eous (Melvill in Melvill & Standen, 1907), from the Red Sea and Arabian Sea, differs from the present species in its very weak sculpture of commarginal growth lines or a smooth shell surface on the left valve in early growth stages. The present species has more prominent sculpture of commarginal lirae on the left valve. The exterior surface of the right valve of S. colmani has commarginal lamellae or lirae, whereas it is smooth in S. eous. Similipecten similis (Laskey, 1811) from the eastern Atlantic is larger (up to c. 6 mm in height), more brightly coloured on the left valve and most specimens have completely smooth discs on both valves. Habitat. Dead specimens only collected so far, in the sublittoral and bathyal zones on soft sediment (sand and clay). The presence of an obvious byssal notch indicates that Similipecten colmani sp. nov. lives byssaly attached to hard objects. Ockelmann (1958: 68–72) described the byssal attachment of Similipecten greenlandicus [under Propeamussium (Arctinula)]: “Several adult animals were found with a bundle of exceedingly fine byssus threads, and possibly the species, when undisturbed, normally lives attached to objects on the sea bottom”. Distribution. Similipecten colmani sp. nov. is distributed from off North West Cape (Western Australia) northeastwards to the Arafura Sea in the Gulf of Carpentaria (Queensland). Probably also from the Flores Sea and Savu Sea (Indonesia) (see Remarks). So far only empty specimens have been collected, in a bathymetric range of 50– 194 m. Remarks. It is most likely that the specimens with abraded sculpture from Indonesia, identified as Similipecten eous by Dijkstra (1991: 23) are also this species. Etymology. Named after Mr Phil H. Colman, senior technical officer in the Malacology Department of the Australian Museum, Sydney (now retired), who collected many pectinoids treated in this project and provided valuable ecological data from his field books., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 154, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Laskey, J. 1811. Account of north British Testacea. Memoirs of the Wernerian Natural History Society 1: 370 - 417.","Ockelmann, K. W. 1958. The zoology of East Greenland. Marine lamellibranchs. Meddelelser om GrOnland 122 (4): 1 - 256."]}

Laevichlamys Waller, 1993 Laevichlamys Waller, 1993: 204. Type species (by original designation): Pecten multisquamatus Dunker, 1864. Recent, tropical western Atlantic. Diagnosis. Non-cemented or embedded Pedini with weak radial costae or riblets (evenly or unevenly spaced), shagreen microsculpture secondarily absent from most specimens, intercalated antimarginal striae, commarginal ridges absent, byssal notch shallow to deep, ctenolium well-developed, hinge teeth weak. Distribution. Upper Miocene–Recent (Waller, 1993: 204). Western Atlantic and Indo-West Pacific, living in the subtidal to sublittoral zones. Discussion. Waller (1993: 204) created a new genus for the chlamydoid taxa with weak to almost smooth macrosculpture. Most of the representative extant species live in the Indo-West Pacific, the exception being the type species (see above)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 237, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Waller, T. R. 1993. The evolution of \" Chlamys \" (Mollusca: Bivalvia: Pectinidae) in the tropical western Atlantic and eastern Pacific. American Malacological Bulletin 10: 195 - 249."]}

Mesopeplini Waller, 2006 Diagnosis. Palliolinae with radial costae clustered into broad radial plicae, posterior auricle with a distinctive posterior margin (at least in primitive members of the tribe), prominent widely spaced commarginal ridges in costal interspaces, with internal rib carinae in most taxa, and with byssal notch reduced in depth. Remarks. Waller (2006a: 20–21) stated that the tribe Mesopeplini was established “to contain a phylogenetically closely knit group of five palliolinine genera and one subgenus … all of which are endemic to New Zealand and Australia …: Mesopeplum (Mesopeplum) Iredale, 1929, … Mesopeplum (Borehamia) Beu, 1978, … Sectipecten Marwick, 1928, … Phialopecten Marwick, 1928, … Kaparachlamys Boreham, 1965, … and Towaipecten Beu, 1995, … Collectively, this group displays a higher level of variation in ribbing patterns and shell shapes than almost any other group of Pectinidae. What ties these genera together are demonstrations of morphological transitions in carefully collected stratigraphic sequences.” Waller (2006a) established the tribe in part because of their demonstrated phylogeny and geographical restriction, but also because they are the only Palliolinae with obvious commarginal ridges throughout ontogeny (limited to the early pre-radial disc in other Palliolinae), as well as the only Palliolinae that develop internal rib carinae. Further study of the New Zealand fossils has demonstrated that almost all of them develop internal rib carinae, including Kaparachlamys (which distinguishes it from the morphologically similar genus Placopecten Verrill, 1897). These genera were discussed by Beu (1995) and Beu & Darragh (2001). Waller (2006a: 32) indicated that the relationship of Mesopeplum to the Palliolini needs to be re-evaluated, implying that he thought it possible that Mesopeplini represents a subfamily separate from Palliolinae. The molecular phylogeny by Sherratt et al. (2016: fig. 2) and Serb et al. (2017) placed Mesopeplum as almost the most basal of Pectinidae, separating Camptonectinae from Chlamydinae (i.e., Pedinae) plus Pectininae, and distinct from Palliolinae. This needs confirmation from further comparisons. Only a single species of this tribe remains in Australia at present, along with one other living species in New Zealand, although Mesopeplum has a fossil record from Oligocene time onwards in Australia and from late Eocene time onwards in New Zealand. A further Australian fossil genus is possibly represented by Mesopeplum (?) contrainflatum Beu & Darragh (2001: 163, fig. 58A–D; late Miocene, eastern Victoria).A radiation of genera and species from Mesopeplum in New Zealand late Miocene–Pliocene rocks (the genera listed above, citing Waller) was described by Beu (1995)., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 172, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Waller, T. R. 2006 a. New phylogenies of the Pectinidae (Mollusca: Bivalvia): reconciling morphological and molecular approaches. In Scallops: Biology, Ecology and Aquaculture. Second Edition, ed. S. E. Shumway and G. J. Parsons, pp. 1 - 44. Amsterdam: Elsevier. https: // doi. org / 10.1016 / S 0167 - 9309 (06) 80028 - 1","Iredale, T. 1929. Mollusca from the continental shelf of eastern Australia. No. 2. Records of theAustralian Museum 17 (4): 157 - 189. https: // doi. org / 10.3853 / j. 0067 - 1975.17.1929.759","Marwick, J. 1928. The Tertiary Mollusca of the Chatham Islands including a generic revision of the New Zealand Pectinidae. Transactions of the New Zealand Institute 58: 432 - 506.","Verrill, A. E. 1897. A study of the family Pectinidae, with a revision of the genera and subgenera. Transactions of the Connecticut Academy of Arts and Sciences 10: 41 - 96.","Sherratt, E., A. Alejandrino, A. C. Kraemer, J. M. Serb, and D. C. Adams. 2016. Trends in the sand: directional evolution in the shell shape of recessing scallops (Bivalvia: Pectinidae). Evolution 70: 2061 - 2073. https: // doi. org / 10.1111 / evo. 12995","Serb, J. M., E. Sherratt, A. Alejandrino and D. C. Adams. 2017. Phylogenetic convergence and multiple shell shape optima for gliding scallops (Bivalvia: Pectinidae). Journal of Evolutionary Biology 2017, 12 pp. https: // doi. org / 10.1111 / jeb. 13137"]}

Pectinidae Rafinesque, 1815 Pectinidae Rafinesque, 1815 [emend. Waller, 1978]; formerly attributed to Wilkes (1810), but that work is not consistently binominal (Waller & Stanley 2005: 38; Dijkstra & Marshall 2008: 2; Bouchet & Rocroi, 2010: 65). Diagnostic characters. Byssate, cemented, or free-living Pectinoidea with outer prismatic calcite layer on right valve limited to early dissoconch stage; some taxa with crossedlamellar aragonite inside pallial line; with an aragonitic myostracum; byssal notch with ctenolium, at least in early growth stages; resilium single, triangular., Published as part of Dijkstra, Henk H. & Beu, Alan G., 2018, Living Scallops of Australia and Adjacent Waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae), pp. 113-330 in Records of the Australian Museum (Rec. Aust. Mus.) (Rec. Aust. Mus.) 70 (2) on page 161, DOI: 10.3853/j.2201-4349.70.2018.1670, http://zenodo.org/record/5299017, {"references":["Rafinesque, C. S. 1815. Analyse de la Nature, ou Tableau de l'Univers et des Corps Organises. Palermo: Rafinesque.","Waller, T. R. 1978. Morphology, morphoclines and a new classification of the Pteriomorphia. Philosophical Transactions of the Royal Society of London, B 284: 345 - 365. https: // doi. org / 10.1098 / rstb. 1978.0072","Wilkes, J. 1810. Conchology. In Encyclopaedia Londinensis; or, Universal Dictionary of Arts, Sciences, and Literature, pp. 14 - 41. London: J. Adlard."]}

Mitotic chromosome preparations of the interspecific hybrids Chlamys farreri (Jones & Preston, 1904) × Patinopecten yessoensis (Jay, 1857), C. farreri × Argopecten irradians (Lamarck, 1819) and C. farreri × Mimachlamys nobilis (Reeve, 1852) were used to compare two different scallop genomes in a single slide. Although genomic in situ hybridization (GISH) using genomic DNA from each scallop species as probe painted mitotic chromosomes of the interspecific hybrids, the painting results were not uniform; instead it showed species-specific distribution patterns of fluorescent signals among the chromosomes. The most prominent GISH-bands were mainly located at centromeric or telomeric regions of scallop chromosomes. In order to illustrate the sequence constitution of the GISH-bands, the satellite Cf303 sequences of C. farreri and the vertebrate telomeric (TTAGGG)n sequences were used to map mitotic chromosomes of C. farreri by fluorescence in situ hybridization (FISH). The results indicated that the GISH-banding pattern presented by the chromosomes of C. farreri is mainly due to the distribution of the satellite Cf303 DNA, therefore suggesting that the GISH-banding patterns found in the other three scallops could also be the result of the chromosomal distribution of other species-specific satellite DNAs.

Parvamussium retiaculum Dijkstra, 1995 Parvamussium retiaculum Dijkstra, 1995: 28, figs 35-38. MATERIAL EXAMINED. — New Caledonia. Southern New Caledonia, 23°05’S, 167°45’E, 680-700 m, BIOCAL, stn DW51, holotype spm (MNHN-IM-2000-24266). Solomon Sea. MADEEP 2014, stn DW4323, 08°38’S, 151°46’E, 720 m, 1 spm. — Stn DW4326, 08°19’S, 149°45’E, 640-660 m, 1 spm (MNHN). Bismarck Sea. PAPUA NIUGINI 2012, stn CP4055, 03°03’S, 142°18’S, 370-374 m, 1 spm (MNHN). DISTRIBUTION. — See Dijkstra & Maestrati (2008: 93). Now also from the Solomon Sea and the Bismarck Sea, live in 370- 660 m., Published as part of Dijkstra, Henk H. & Maestrati, Philippe, 2017, New species and new records of littoral and bathyal living Pectinoidea (Bivalvia: Propeamussiidae, Cyclochlamydidae, Pectinidae) from the western and southwestern Pacific, pp. 473-485 in Zoosystema 39 (4) on page 477, DOI: 10.5252/z2017n4a3, http://zenodo.org/record/4578634, {"references":["DIJKSTRA H. H. 1995. - Bathyal Pectinoidea (Bivalvia: Propeamussiidae, Entoliidae, Pectinidae) from New Caledonia and adjacent areas, in BOUCHET P. (ed.), Resultats des Campagnes MUSORSTOM volume 14. Museum national d'Histoire naturelle, Paris, 654 p. (Memoires du Museum national d'Histoire naturelle; 167): 9 - 73.","DIJKSTRA H. H. & MAESTRATI P. 2008. - New species and new records of deep-water Pectinoidea (Bivalvia: Propeamussiidae, Entoliidae and Pectinidae) from the South Pacific, in HEROS V., COWIE R. H. & BOUCHET P. (eds), Tropical Deep-Sea Benthos volume 25. Museum national d'Histoire naturelle, Paris, 806 p. (Memoires du Museum national d'Histoire naturelle; 196): 77 - 113."]}

Twenty-five species of Pectinoidea (24 Propeamussiidae, 1 Cyclochlamydidae) are herein listed from the Mozambique Channel, northwestern and southern Madagascar, and northeastern South Africa. New species: Propeamussium rosadoi, Parvamussium catillus, Parvamussium kilburni, Parvamussium puillandrei, Parvamussium strongae, Cyclopecten cassiculus, Cyclopecten kantori, Cyclochlamys bacachorda. New synonym: Amussium sewelli Knudsen, 1967 = Propeamussium watsoni (E.A. Smith, 1885). New records for the Mozambique Channel and northwestern Madagascar: Propeamussium andamanicum, Propeamussium arabicum, Propeamussium caducum, Propeamussium jeffreysii, Propeamussium sibogai, Propeamussium watsoni, Parvamussium formosum, Parvamussium scitulum, Parvamussium torresi, Parvamussium vesiculatum, Cyclopecten kapalae, Similipecten eous. New records for southern Madagascar: Propeamussium jeffreysii, Propeamussium sibogai, Propeamussium watsoni, Parvamussium formosum, Parvamussium scitulum, Parvamussium thyrideum, Parvamussium vesiculatum, Parvamussium vidalense, Cyclopecten kapalae, Similipecten eous. New record for South Africa: Propeamussium jeffreysii, Parvamussium formosum, Parvamussium scitulum, Cyclopecten horridus, Similipecten eous.

Cyclopecten kantori sp. n. Fig. 7A–G Etymology:After Dr Yuri I. Kantor, leading researcher of the Department of Invertebrate Morphology at the A.N. Severtzov Institute of Ecology and Evolution, Russian Academy of Sciences in Moscow. Participant of several French expeditions. Description: Shell up to 11.9 mm in height, fragile, subcircular, inequivalve, almost equilateral, left valve more inflated than right valve (nearlY flat), auricles dissimilar in shape, anterior auricle larger than posterior one, umbonal angle 120°, internal ribs lacking, semi-translucent whitish. Prodissoconch 220 µm long. Left valve reticulately sculptured throughout shell disc by narrow radial riblets and overrunning commarginal lirae with delicate solide nodules on intersections. On central part of disc c. 10 radial riblets per mm, c. 7–8 commarginal lirae. Reticular sculpture commences in early growth stage, radial riblets enlarge gradually in ontogeny by intercostal riblets towards the ventral margin. Commarginal lirae more prominent than radial riblets, very closely spaced in early growth stage, gradually wider in later ontogeny. Auricular sculpture similar to that of shell disc with stronger solid nodules on the intersections. Right valve sculptured with commarginal lirae, closely spaced (c. 30 per mm) near umbonal area, coarser and wider on central part of disc (c. 8 per mm). Posterior auricle continuous with shell disc, sculptured with fine close-set commarginal lirae, anterior auricle and byssal fasciole seperated from shell disc by sharply incised groove, sculptured with prominent commarginal lirae and 8 crowded radial riblets with nodular intersections. Byssal notch moderately deep, byssal fasciole narrow. Dimensions: Holotype: Height 11.9 mm, width 11.1 mm, convexity 3.2 mm. Type material: Holotype, pr (MNHN IM-2007-38440). Type locality: NW MADAGASCAR: In front of Narendry Bay (14°30'S 47°27'E), - 274–325 m, live, campaign Miriky, stn CP3241, 06.vii.2009. Distribution and habitat: So far northwestern Madagascar, living bathyally on soft substrata at - 274– 325 m. Remarks: The morphologically closest congeneric species is the abyssally (- 2928– 2930 m) living similar sized Cyclopecten textus Dijkstra & Marshall, 2008, recorded from the Lord Howe Rise. This species could be distinguished from C. kantori by the following characters: by shell shape (C. textus is almost circular, C. kantori somewhat higher than wide), by shape of auricles (C. textus has smaller auricles (c. 2 mm) measured between end of hinge and disc, C. kantori somewhat broader (2.5 mm)), by commarginal sculpture of the left valve (C. textus has less commarginal sculpture in early ontogeny, P. kantori more close-set), and by reticulate sculpture in the central part of the disc of left valve (C. textus more squarely, C. kantori more elongate). A somewhat similar reticulated sculptured bathyally living species is Parvamussium retiolum Dijkstra, 1995, recorded from the southwestern Pacific. This species could be easily distinguished from C. kantori by its more oblique elongate shape (C. kantori sub-circular), and by its well-developed internal ribs (lacking in C. kantori)., Published as part of Dijkstra, H. H. & Maestrati, P., 2015, Pectinoidea (Bivalvia: Propeamussiidae and Cyclochlamydidae) from the southwestern Indian Ocean, pp. 585 in African Invertebrates 56 (3) on pages 618-619, DOI: 10.5733/afin.056.0307, http://zenodo.org/record/7914533

Parvamussium catillus sp. n. Fig. 2A–G Etymology: From the Latin catillus (m.) (small dish), after the smooth circular shell that resembles a small dish. Description: Shell up to 7.2 mm in height, fragile, semi-transparent, circular, inequivalve, left valve slightlY more inflated than right valve, equilateral, auricles almost equal in size, umbonal angle 110°–115°, colour whitish. Prodissoconch 100 µm long. Left valve smooth and dead throughout shell disc, umbonal part glossy and transparent. Anterior auricle with very weak and delicate widely spaced commarginal lamellae, more prominent near shell disc, posterior auricle almost smooth with traces of very closely spaced commarginal lirae laterally. Right valve with weak, regularly spaced, commarginal lirae (c. 8 per mm).Anterior auricle with weak and verY fine commarginal lirae and a noduliferous ridge along the byssal fasciole, posterior auricle continuous with disc, smooth with very close-set commarginal lirae laterally. Byssal notch moderately deep, byssal fasciole rather small, no ctenolium. Hinge line straight. Resilifer triangular. Internal ribs 13 commence in early growth stage and are exposed almost to the periphery, one interstitial rudimentary riblet antero-ventrally, and one riblet on both auricles. Dimensions (holotype): Height 7.2 mm, width 7.2 mm, convexity 2.5 mm. Type material: Holotype (pr) (MNHN IM-2007-41966), paratype (pr) (MNHN IM-2007-41964). Type locality: MOZAMBIQUE CHANNEL: Inhambane transect (23°35'S 36°06'E), - 1092–1195 m, live, campaign Mainbaza, stn CP3139, RV Vizconde de Eza, leg. P. Bouchet, J. Rosado & E. Strong, 11.iv.2009. Distribution and habitat: See type locality. Remarks: The closest resembling species is Parvamussium multiliratum Dijkstra, 1995, recorded from New Caledonia, Vanuatu, Fiji, Tonga, Wallis and Futuna (Dijkstra & Maestrati 2012: 392). Parvamussium catillus morphologically differs from P. multiliratum by a smooth shell disc of the left valve (P. multiliratum with prominent commarginal lamellae and weak radial sculpture postero-laterally), by having fewer interior ribs (P. multiliratum 14 and two rudimentary). Another somewhat resembling species is Propeamussium arabicum Dijkstra & Janssen, 2013, recorded from the Gulf of Aden and herein also from northwestern Madagascar and the Mozambique Channel. This species is also circular in shape, but larger sized (up to 14.9 mm in height, P. catillus up to 7.2 mm) and has a fine reticular sculpture on the left valve (P. catillus smooth). Moreover, fewer internal ribs (7 without rudimentary, P. catillus 13 and one rudimentary)., Published as part of Dijkstra, H. H. & Maestrati, P., 2015, Pectinoidea (Bivalvia: Propeamussiidae and Cyclochlamydidae) from the southwestern Indian Ocean, pp. 585 in African Invertebrates 56 (3) on pages 599-601, DOI: 10.5733/afin.056.0307, http://zenodo.org/record/7914533, {"references":["DIJKSTRA, H. H. & JANSSEN, R. 2013. Bathyal and abyssal Pectinoidea from the Red Sea and Gulf of Aden (Bivalvia: Propeamussiidae, Entoliidae, Pectinidae). Archiv fur Molluskenkunde 142: 181 - 214."]}

Cyclopecten cassiculus sp. n. Fig. 6A–G Etymology: From the Latin cassiculus (m.) (small spider’s web), after the sculpture of left valve that resembles a spider’s web with fine raindrops on the intersections. Description: Shell up to 8.9 mm in height, fragile, lv opaque, rv semi-transparent, somewhat higher than wide, inequivalve, flattened and somewhat rough, left valve slightlY more convex than right valve, inequilateral, anterior auricle larger than posterior, umbonal angle 90°–95°, colour of left valve creamy, right valve whitish. Prodissoch 170 µm long. Left valve sculptured with very closely spaced commarginal lirae in early growth stage (c. 10 per mm), more prominent and widely spaced on central shell disc (c. 4–5 per mm), 8–10 fine radial lirae commence c. 0.8–1.0 mm below umbo, increasing to c. 15 strong primary lirae near periphery with interstitial secondary lirae, intersections with small nodules on primary radial lirae in early growth stage, more lamellar in adult stage, small nodules on secondary interstitial riblets, similar sculptured auricles almost continuous with shell disc. Right valve with very weak, regularly spaced, delicate commarginal lamellae (c. 10 per mm). Anterior auricle with 6–7 antimarginal noduliferous lirae, posterior auricle continuous with disc with verY fine and close-set commarginal lirae. Byssal notch moderately deep, byssal fasciole rather small, no ctenolium. Hinge line straight. Resilifer triangular. Internal ribs lacking. Dimensions (holotype): height 8.9 mm, width 8.0 mm, convexity 2.9 mm. Type material: Holotype, pr (MNHN IM-2007-41951), paratype, pr (MNHN IM-2007-41954).Type locality: MADAGASCAR: Near Mahajamba Bay (14°50'S 46°57'E), - 340–446 m, live, campaign Miriky, stn CP3248, RV Miriky, leg. P. Bouchet & Y. Kantor, 07.vii.2009. Paratype: MADAGASCAR: Between Majunga and Cap Saint-André (15°32'S 45°45'E), - 327–730 m, live, campaign Miriky, stn CP3265, RV Miriky, leg. P. Bouchet & Y. Kantor, 10.vii.2009. Distribution and habitat: NW Madagascar, living upper bathyally at - 327–340 m (minimum depth range). Remarks: The closest resembling species is Parvamussium cassium Dijkstra, 1991, recorded from deep water (- 450–600 m) off Indonesia. Cyclopecten cassiculus morphologically differs from P. cassium bY having prominent radial and fine commarginal lirae (in P. cassium weaker and more equal of prominence), in having secondary intercostal radial lirae in late ontogeny (P. cassium in earlier growth stage), in having fine interstitial radial lirae, up to 6 near ventral margin (one radial lira in P. cassium), and in having coarser nodules and lamellae on intersections of lirae (P. cassium very small nodules). Both species are lacking internal ribs, although P. cassium has a short auricular riblet (very weak in the present species). A somewhat resembling species is Parvamussium vidalense (Barnard, 1964), recorded from eastern South Africa and herein also from southern Madagascar. This species has a reticular sculpture on the left valve of more closelY spaced, finer radial and commarginal lirae with short spines or curved lamellae on the radial and commarginal intersections. Moreover, several rudimentary interior ribs are present (lacking in C. cassiculus)., Published as part of Dijkstra, H. H. & Maestrati, P., 2015, Pectinoidea (Bivalvia: Propeamussiidae and Cyclochlamydidae) from the southwestern Indian Ocean, pp. 585 in African Invertebrates 56 (3) on pages 616-617, DOI: 10.5733/afin.056.0307, http://zenodo.org/record/7914533, {"references":["BARNARD, K. H. 1964. Contributions to the knowledge of South African marine Mollusca. Part 5. Lamellibranchiata. Annals of the South African Museum 47: 361 - 593."]}

Family Cyclochlamydidae Dijkstra & Maestrati, 2012 Remarks: For diagnosis and remarks on this family see Dijkstra and Maestrati (2012: 393)., Published as part of Dijkstra, H. H. & Maestrati, P., 2015, Pectinoidea (Bivalvia: Propeamussiidae and Cyclochlamydidae) from the southwestern Indian Ocean, pp. 585 in African Invertebrates 56 (3) on page 622, DOI: 10.5733/afin.056.0307, http://zenodo.org/record/7914533

Genus Parvamussium Sacco, 1897 Parvamussium Sacco, 1897 a: 102 (Proposed as a subgenus of Amussium Herrmannsen, 1846, an unjustified emendation of Amusium Röding, 1798); no diagnosis given, but type species designated; Sacco 1897 b: 48 (diagnosis). Type species (OD): Pecten (Pleuronectes) duodecimlamellatus Bronn, 1832; Pliocene, northern Italy (Waller 2011: 25). Remarks: For synonymy, diagnosis, distribution and discussion see Dijkstra (1995: 25)., Published as part of Dijkstra, H. H. & Maestrati, P., 2015, Pectinoidea (Bivalvia: Propeamussiidae and Cyclochlamydidae) from the southwestern Indian Ocean, pp. 585 in African Invertebrates 56 (3) on page 598, DOI: 10.5733/afin.056.0307, http://zenodo.org/record/7914533, {"references":["SACCO, F. 1897 a. I molluschi dei terreni terziarii del Piemonte e della Liguria, 24. Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Universita di Torino 12 (298): 101 - 102.","BRONN, H. G. 1832. Ergebnisse meiner naturhistorisch-okonomischen Reisen. Zweyter Teil: Skizzen und Ausarbeitungen uber Italien nach einem zweiten Besuche im Jahre 1827. Heidelberg & Leipzig: Goos."]}

Propeamussium sibogai (Dautzenberg & Bavay, 1904) Fig. 1I, J Amussium sibogai Dautzenberg & BavaY, 1904: 207, figs 1–4; Dautzenberg & BavaY 1912: 31, pl. 28, figs 1–4. Type locality: Indonesia, Bali Sea (7°15'S 115°15'E), - 289 m, live, mud and broken shells, 14.iii.1899 (Siboga, stn 12). Propeamussium sibogai: Knudsen 1967: 272, pl. 1, figs 23–24; Dijkstra 1995: 23, figs 19–22; 2001: 81; 2013: 14, pl. 1, figs 4a–d, pl. 4, fig. 4; Dijkstra & Kastoro 1997: 253, figs 24–29.; Dijkstra & Marshall 1997: 79, pl. 3, figs 1–3.; 2008: 5, figs 1, 3H–J; Wang 2002: 151, pl. 5, fig. 7; Dijkstra & Maestrati 2008: 84; 2013: 470. Amussium cf. sibogai: Barnard 1969: 655, pl. 1, figs a–d. Description: Shell fragile, up to 65 mm in height (most specimens smaller up to 40– 50 mm), hyaline, almost circular, inequivalve, somewhat inequilateral, slightly oblique, left valve more inflated than right valve, laterallY gaping rather stronglY, auricles small and equal in size, umbonal angle 120°–125°. Left valve creamy brown, right valve creamy. Prodissoconch 220 µm high. Left valve covered with a few delicate commarginal growth lines; a few minute commarginal lamellae present near ventral margin of some specimens. Delicate radial lirae developed near posterior margin. Auricles smooth, somewhat raised on lateral margins. Right valve with widely spaced commarginal lirae, weaker at periphery, and microscopic interstitial granules. Marginal apron lacking from most specimens due to the verY thin laYer of prismatic calcite (HaYami 1988: fig. 4). Auricles smooth. Hinge line straight. Internal costae 7 in most specimens, deep brown on left valve, white and broader on right valve. Marginal costae with 4 or 5 small nodules, visible through lateral gape. Resilifer triangular, elongate. No byssal notch. Type material examined: Holotype (pr) ZMA Moll. 304001. Other material examined: NW MADAGASCAR: Between Nosy-Bé and Banc du Leven (12°38'S 48°14'E), - 420–436 m, live, campaign Miriky, stn CP3183, 26.vi.2009 (MNHN); Northwest region, between Nosy-Bé and Banc du Leven (12°40'S 48°12'E), - 492–524 m, live, campaign Miriky, stn CP3184, 26.vi.2009 (MNHN); Northwest region, between Nosy-Bé and Banc du Leven (12°30'S 48°18'E), - 346–376 m, dead, campaign Miriky, stn CP3189, 27.vi.2009 (MNHN); Northwest region, between Nosy-Bé and Banc du Leven (12°31'S 48°15'E), - 415–416 m, dead, campaign Miriky, stn CP3190, 27.vi.2009 (MNHN); Northwest region, between Nosy-Bé and Banc du Leven (12°44'S 48°12'E), - 442–491 m, live, campaign Miriky, stn CP3210, 29.vi.2009 (MNHN); Northwest region, between Nosy-Bé and Banc du Leven (12°33'S 47°56'E), - 391–438 m, live, campaign Miriky, stn DW3217, 30.vi.2009 (MNHN); Northwest region, between Nosy-Bé and Banc du Leven (12°46'S 48°11'E), - 430–488 m, live, campaign Miriky, stn CP3223, 02.vii.2009 (MNHN); Mahajamba Bay (14°50'S 46°57'E), - 340–446 m, live, campaign Miriky, stn CP3248, 07.vii.2009 (MNHN); Majunga (15°22'S 46°00'E), - 493–750 m, live, campaign Miriky, stn CP3250, 08.vii.2009 (MNHN); in front of Narendry Bay (14°29'S 47°26'E), - 409–425 m, live, campaign Miriky, stn CP3290, 14.vii.2009 (MNHN). S MADAGASCAR: Between Lokaro and Sainte Luce (24°53'S 47°28'E), - 184–203 m, dead, expedition Atimo Vatae, stn DW3515, 30.iv.2010 (MNHN). MOZAMBIQUE CHANNEL: Maputo transect (25°12'S 35°17'E), - 503–505 m, dead, campaign Mainbaza, stn CP3136, 10.iv.2009 (MNHN); Maputo transect (23°31'S 35°50'E), - 446–475 m, live, campaign Mainbaza, stn CP3142, 11.iv.2009 (MNHN). MOZAMBIQUE: E of Inhaca Island, - 457 m, live, don. Mrs K. Eastwood (NMSA 5294, 1 pr); Bazaruto Island, - 549 m, live, ex colln K. Eastwood (NMSA J5095). SOUTH AFRICA: Zululand: Off Matigulu River mouth (29°22.3'S 31°57.7'E), - 400 m, dead, mud, NMDP stn ZR12, dredged RV Meiring Naudé, 16.vi.1989 (NMSA E8820). KwaZulu­Natal: Off Durban (29°55.8'S 31°13.0'E), - 337–346 m, live, stn B3.1, trawled ORI, 20.ii.2006 (NMSA W8601); Off Durban, - 366 m, live, don. R. Kilburn, 01.ix.1969 (NMSA 4492). Distribution and habitat: Propeamussium sibogai is recorded throughout the tropical Indo-West Pacific from southern Japan (HaYami 2000), southwards to the Philippines and Indonesia (Dautzenberg & Bavay 1912; Knudsen 1967; Dijkstra & Kastoro 1997; Dijkstra 2013), eastwards to the Solomon Islands, Vanuatu Archipelago, Loyalty Islands, New Caledonia, Wallis and Futuna Islands, Fiji, Tonga, and Kermadec Islands (Dijkstra 1995; 2001; Dijkstra & Maestrati 2008; 2013; Dijkstra & Marshall 2008), and westwards only recorded from off Durban, South Africa (Knudsen 1967). Now also from the Mozambique Channel and northwestern and southern Madagascar (new record). Propeamussium sibogai is living on soft substrata of mud or muddy sand at a bathyal depth of - 90–750 m (Dijkstra 2013; present records).Present material live at - 340– 750 m. Remarks: All morphological characters of the present material are identical to the holotype from the Bali Sea, Indonesia. These characters are rather constant and can slightly vary in number of the internal ribs. Occasionally on one or both sides the lateral rib is divided into two separated ribs. The southwestern Indian Ocean morph does not have divided lateral ribs and thus only 6 internal ribs (typically 7). Propeamussium sibogai is the type species of Luteamussium Oyama, 1951: 82, which was treated by Hertlein (1969: N350) as a junior synonym of Propeamussium, subsequently followed in literature by most authors (see references). Descriptions of the soft parts, reproduction and diet are given by Knudsen (1967: 272)., Published as part of Dijkstra, H. H. & Maestrati, P., 2015, Pectinoidea (Bivalvia: Propeamussiidae and Cyclochlamydidae) from the southwestern Indian Ocean, pp. 585 in African Invertebrates 56 (3) on pages 594-595, DOI: 10.5733/afin.056.0307, http://zenodo.org/record/7914533, {"references":["DAUTZENBERG, P. & BAVAY, A. 1904. Description d'un Amussium drague par le \" Siboga \" dans la mer de Celebes. Journal de Conchyliologie 52: 207 - 211.","KNUDSEN, J. 1967. The deep-sea Bivalvia. In: The John Murray Expedition 1933 - 34. Scientific Reports 11 (3). London: British Museum (Natural History), pp. 237 - 343.","DIJKSTRA, H. H. & KASTORO, W. W. 1997. Mollusca Bivalvia: Pectinoidea (Propeamussiidae and Pectinidae) from eastern Indonesia. In: Crosnier, A. & Bouchet, P., eds, Resultats des Campagnes MUSORSTOM, 16. Memoires du Museum national d'Histoire naturelle, Zoologie 172: 245 - 285.","DIJKSTRA, H. H. & MARSHALL, B. A. 1997. Pectinoidea (Mollusca: Bivalvia: Propeamussiidae: Pectinidae) of Lord Howe Island, Norfolk Island and the Kermadec Islands. Molluscan Research 18: 73 - 114.","WANG, Z. 2002. Fauna Sinica. Invertebrata 31. Mollusca. Bivalvia. Pteriina. Beijing: Science Press. (in Chinese)","DIJKSTRA, H. H. & MAESTRATI, P. 2008. New species and new records of deep-water Pectinoidea (Bivalvia: Propeamussiidae, Entoliidae and Pectinidae) from the South Pacific. In: Heros, V., Cowie, R. H. & Bouchet, P., eds, Tropical Deep - Sea Benthos, 25. Memoires du Museum national d'Histoire naturelle 196: 77 - 113.","HAYAMI, I. 1988. Functional and taxonomic implications of internal ribs of Propeamussium. Transactions and Proceedings of the Palaeontological Society of Japan, New Series 150: 476 - 490.","DIJKSTRA, H. H. & JANSSEN, R. 2013. Bathyal and abyssal Pectinoidea from the Red Sea and Gulf of Aden (Bivalvia: Propeamussiidae, Entoliidae, Pectinidae). Archiv fur Molluskenkunde 142: 181 - 214.","OYAMA, K. 1951. Amusiinae in Japan. List of marine molluscan genera, subgenera and sections (Tertiary of Recent) established by Japanese authors (1925 - 1949), 4. In: Kuroda, T., ed., Illustrated catalogue of Japanese shells 1 (13): 85 - 86.","HERTLEIN, L. G. 1969. FamilY Pectinidae Rafinesque, 1815. In: Moore, R. C., ed., Treatise on invertebrate paleontology, Part N, Vol. 1, Mollusca 6, Bivalvia. Boulder, Colorado: The University of Kansas, Lawrence and The Geological Society of America, pp. N 348 - N 372."]}