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Panaspis cabindae (Bocage, 1866) Material (two specimens): One adult specimen (MHNCUP/REP 266, formerly UP-MHNFCP-017381), collected from Lucálla [=Lucala] [-9.27268º, 15.25206º, 708 m a.s.l.] Kwanza Norte Province, in 1903; one adult specimen (MHNCUP/REP 267, formerly UP-MHNFCP-017380; Fig. S37), collected from Catete [-9.11667º, 13.70000º, 21 m a.s.l.] Bengo Province, in September 1903. Comments: Panaspis cabindae is a species with a distribution extending from southwestern Democratic Republic of the Congo to southwestern Angola (Marques et al. 2018; Ceríaco et al. 2020b). The presented specimens were cited as “ Ablepharus cabindae, Boc ” by Ferreira (1904) and are the only published records for the species from “Lucala” and “Catete” localities. Recently, Ceríaco et al. (2016a, 2018b, 2020b) and Medina et al. (2016) published new records for the species in Angola, including the first accounts for Bié, Luanda, Uíge, Malanje, Lunda Norte and Huíla provinces. This species is one of the most widespread in the country, despite the lack of published records (Ceríaco et al. 2020b)., Published as part of Santos, Bruna S., Marques, Mariana P., Bauer, Aaron M. & Ceríaco, Luis M. P., 2021, Herpetological results of Francisco Newton's Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection, pp. 1-80 in Zootaxa 5028 (1) on page 23, DOI: 10.11646/zootaxa.5028.1.1, http://zenodo.org/record/5453967, {"references":["Bocage, J. V. B. (1866) Reptiles nouveaux ou peu connus recueillis dans les possessions portugaises de l'Afrique occidentale, qui se trouvent au Museum de Lisbonne. Jornal de Sciencias Mathematicas, Physicas e Naturaes, I, 57 - 78.","Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501.","Ceriaco, L. M. P., Heinicke, M. P., Parker, K. L., Marques, M. P. & Bauer, A. M. (2020 b) A review of the African snake-eyed skinks (Scincidae: Panaspis) from Angola, with the description of a new species. Zootaxa, 4747 (1), 077 - 112. https: // doi. org / 10.11646 / zootaxa. 4747.1.3","Ferreira, J. B. (1904) Reptis e amphibios de Angola da regiao ao norte do Quanza (Colleccao Newton 1903). Jornal de Sciencias Mathematicas Physicas e Naturaes, Segunda Serie, 7 (25), 111 - 117.","Ceriaco, L. M. P., De Sa, S. A. C., Bandeira, S., Valerio, H., Stanley, E. L., Kuhn, A. L., Marques, M. P., Vindum, J. V., Blackburn, D. C. & Bauer, A. M. (2016 a) Herpetological survey of the Iona National Park and Namibe National Park, Angola, with a synoptic list of the amphibians and reptiles of Namibe Province, southwestern Angola. Proceedings of the California Academy of Sciences, Series 4, 63 (2), 15 - 61.","Ceriaco, L. M. P., Marques, M. P., Bandeira, S., Blackburn, D. C. & Bauer, A. M. (2018 b) Herpetological survey of Cangandala National Park with a synoptic list of the amphibians and reptiles of Malanje Province, central Angola. Herpetological Review, 49 (3), 408 - 431.","Medina, M. F., Bauer, A. M., Branch, W. R., Schmitz, A., Conradie, W., Nagy, Z. T., Hibbitts, T. J., Ernst, R., Portik, D. M., Nielsen, S. V., Colston, T. J., Kusamba, C., Behangana, M., Rodel, M. - O. & Greenbaum, E. (2016) Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution, 100, 409 - 423. https: // doi. org / 10.1016 / j. ympev. 2016.04.026"]}

Panaspis cabindae (Bocage, 1866) (Fig. 7–10) Chresonymy (includes only citations unambiguously pertaining to specimens referable to the species, or explicitly stating or depicting its geographic range). Ablepharus cabindae (Bocage 1866a: 64, 1895: 51; Peters 1877: 614; Boulenger 1887: 352; Ferreira 1904: 116; Parker 1936: 136) Ablepharus Cabindae (Bocage 1866b: 45, 1867: 224, 1887: 179, 1897: 196) Ablepharus aeneus (Boulenger 1887: 352) Riopa (Panaspis) cabindae (Smith 1937: 229) Panaspis cabindae (Greer 1974: 29; Perret 1975: 8; Schmitz et al. 2005: 23; Ceríaco et al. 2016a: 57; 2016b: 65; Medina et al. 2016: 411; Ceríaco et al. 2018a: 141; Ceríaco et al. 2018c: 423; Marques et al. 2018: 247; Branch et al. 2019a: 317) This species was described by Bocage (1866) based on three specimens collected by the Portuguese explorer José Alberto de Oliveira de Anchieta (1832–1897) in the Cabinda enclave, northwestern Angola. Cope (1868) described a new genus and species Panaspis aeneus. The description of this species was problematic as Cope was unable to provide a specific type locality (according to the original description Habitat. Probably Swan River, Australia; possibly from South-Western Africa.), and the holotype is currently lost. The usage of the nomen aeneus was short-lived, as the only other reference to aeneus as a valid species was that of Boulenger (1887), when referring to specimens from Angola. The two specimens labelled as aeneus by Boulenger (1887) were collected by the London-born Portuguese colonial officer Joaquim John Monteiro (1833–1878) in the Luanda region. These specimens are still extant in the collections of the BMNH under the accession number 1873.7.28.11 and are unambiguously conspecific with P. cabindae. In his scholarly revision of the herpetofauna of Angola, Bocage (1895) was the first to synonymize aeneus with cabindae, a decision then followed by all subsequent authors (e.g. Fuhn 1972; Perret 1975). In the same work Bocage (1895) provided the first illustration of the species, including detailed views of the head scalation (Fig. 7). Despite the unavailability of aeneus type material, the original description provides enough morphological data to confidently confirm that it is conspecific with cabindae, namely the presence of supranasals, pre-ablepharine eye and frontoparietals separated, diagnostic features of cabindae (see below). Due to this lack of ambiguity, a neotype designation for aeneus is not needed. The same holds true for cabindae itself, as the original type series was destroyed in the fire that engulfed the zoological collections of Museu Bocage, Lisbon, in 1978. The most recent data on the species was provided by Medina et al. (2016) who confirmed, through molecular data, that the species extended from southwestern Democratic Republic of the Congo to southwestern Angola; our sequence data here further fills in the Angolan distribution of the species. Other recent published records include those from Cangandala National Park, Malanje Province, Central Angola reported by Ceríaco et al. (2016b) and Ceríaco et al. (2018c). Marques et al. (2018) presented and mapped all available records of the species for Angola. Despite being a quite common species in Angola (LMPC pers. obs.), not much has been published about the distriution, natural history, ecology or conservation status of this species. Diagnosis. Panaspis cabindae can be distinguished from other members of the genus occurring inAngola and surroundingsregionbyhaving1)presenceofsupranasals;2)pre-ablepharineeye(asdefinedbyGreer1974);3)frontoparietalsseparated;dorsumcoppery-brown,withfourveryvaguelongitudinalstripesextendingapproximatelytomidbody, and a thin darker band starting on the temporals and extending to midbody; 5) absence of rows of light spots on the neck; 6) absence of a white ventrolateral stripe; 7) 23 to 26 midbody scales rows (Figs. 8–9). Specimens examined. ANGOLA: Cabinda Province: Chinchoxo [-5.1º, 12.1º, 45 m] (ZMB 9168, 9276); Zaire Province: Soyo [-6.134903º, 12.368935º, 10 m] (PEM 20256 –20260); Uíge Province: Kimpa Vita University Campus [-7.618614º, 15.065536º, 1122 m] (MTD 48612); Luanda Province: Luanda [-8.83333º, 13.26667º, 81 m] (NHM 1951.1.1.94), Cabiri [-8.91667º, 13.66667º, 30 m] ( AMB 10881, 10883), Kissama National Park [-9.18303°, 13.37063°, 136 m] (FLMNH 187235–241); Bengo Province: Catete [-9.11667º, 13.7º, 21 m] (MHN- CUP 017380), Riverine Forest [-9.11667º, 13.7º, 21 m] PEM R 21594; Benguela Province: Lobito Bay [-12.35º, 13.55º, 7 m] (AMNH R-48501), Hanha [-13.3º, 14.2º, 953 m] (AMNH R-40671, 48502, 48504, 40718); Malanje Province: Duque de Bragança / Kalandula [-9.1º, 15.95º, 1011 m] (ZMB 7775, CAS 263590), Cangandala National Park [-9.84606º, 16.72233º, 1111 m] (CAS 258403–407); Laúca [-9.762749º, 15.143798º, 750 m] (ANG 40-41, AMB 9190, 9209, 9214, 9230, 9250, 9251, 9254, 9255, 9256, 9257, 9258, 9259, 9260, 9261, 9262, 9676, 9677, 9678, 9685, 9686); Lunda Norte Province: Lagoa Carumbo [-7.74422º, 19.95467º, 789 m] (PEM R 19467); Kwanza-Norte Province: Lucalla [-9.4º, 15.03333º, 411 m] (MHNCUP 017381); N’Dallatando [-9.3º, 14.91667º, 782 m] (NHM 1909.10.29.103); Kwanza-Sul Province: Congulu [-10.86667º, 14.28333º, 639 m] (NHM 1936.8.1.622–624, MCZ R- 112222); Bié Province: Cassumbe [-11.08615º, 16.66619º, 1235 m] (AMB 11696, 11717, 11753, 11788); Chitau [-11.43333º, 17.15º, 1510 m] (FMNH 74295); Namibe Provice: Mamué Riparian área [-13.80147º, 13.12056º, 660 m] (CAS 10315, AMB 10316, CAS 263554, 10321, CAS 263550); Capangombe [-15.1º, 13.15º, 553 m] (MHNCUP 017379, NHMW 10236, CAS 264244). REPUBLIC OF THE CONGO: Pointe-Noire [-4.762602º, 11.866835º, 33 m] (MNHN 1966.554). DEMOCRATIC REPUBLIC OF THE CONGO: Bombo-Lumene Reserve Headquarters [-4.42218º, 16.04904º, 619 m] (UTEP Herp-21173); Malela [-5.982438º, 12.621458º, 1 m] (CAS 54820 –821; MCZ R-11286, 14011–14012). Distribution. The species occurs in the southwestern region of Republic of the Congo and the Democratic Republic of the Congo, extending to all parts of northern Angolan and descending south to central and southwestern Angola through the Angolan escarpment (Fig. 10). Habitat and Natural History notes. The species is found in a wide variety of habitats, from some more coastal, Euphorbia and baobab dominated savannahs, in northwestern Angola, to typical Miombo woodland habitats (Fig. 11) in the central plateau of Angola. The species appears to be absent from the more xeric and desertic areas of the southwest of Angola and northwestern Namibia, however it may possibly be found in more humid areas close to the escarpment. The species is active during the day in leaf-litter, but is rarely seen exposed., Published as part of Ceríaco, Luis M. P., Heinicke, Matthew P., Parker, Kelly L., Marques, Mariana P. & Bauer, Aaron M., 2020, A review of the African snake-eyed skinks (Scincidae: Panaspis) from Angola, with the description of a new species, pp. 77-112 in Zootaxa 4747 (1) on pages 90-93, DOI: 10.11646/zootaxa.4747.1.3, http://zenodo.org/record/3693431, {"references":["Bocage, J. V. B. (1866 a) Lista dos reptis das possessoes portuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 37 - 56.","Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls.","Peters, W. C. H. (1877) Ubersicht der Amphibien aus Chinchoxo (Westafrika), welche von der Afrikanischen Gesellschaft dem Berliner zoologischen Museum ubergeben sind. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1877, 311 - 621, 1 pl.","Boulenger, G. A. (1887) Catalogue of the Lizards in the British Musuem (Natural History). Vol. 3. Lacertid ӕ, Gerrhosaurid ӕ, Scincid ӕ, Anelytropsid ӕ, Dibamid ӕ, Chamaeleontid ӕ. Trustees of the British Museum, London, xii + 575 pp., 40 pls.","Ferreira, J. B. (1904) Reptis e amphibios de Angola da regiao ao norte do Quanza (Colleccao Newton- 1903). Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 7, 111 - 117.","Parker, H. W. (1936) Dr. Karl Jordan's expedition to South-West Africa and Angola: herpetological collection. Novitates Zoologicae, 40, 115 - 146.","Bocage, J. V. B. (1866 b) Reptiles nouveaux ou peu connus recueillis dans les possessions portugaises de l'Afrique occidentale, qui se trouvent au Museum de Lisbonne. Jornal de Sciences Mathematicas, Physicas e Naturaes, 1, 57 - 78.","Bocage, J. V. B. (1867) Segunda lista dos reptis das possessoes portuguezas d'Africa occidental que existem no Museu de Lisboa. Jornal de Sciences Mathematicas, Physicas e Naturaes, 1, 217 - 228.","Bocage, J. V. B. (1887) Melanges erpetologiques. I. Reptiles et batraciens du Congo. Jornal de Sciences Mathematicas, Physicas e Naturaes, 11, 177 - 192.","Bocage, J. V. B. (1897) Mammiferos, reptis e batrachios d'Africa de que existem exemplares typicos no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 187 - 206.","Smith, M. A. (1937) A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum, 39 (3), 213 - 234.","Greer, A. E. (1974) The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology, 31, 1 - 67. https: // doi. org / 10.1071 / ajzs 031","Perret, J. L. (1975) La differentiation dans le genre Panaspis Cope (Reptilia: Scincidae). Bulletin de la Societe Neuchateloise des Sciences Naturelles, 98, 5 - 16.","Schmitz, A., Ineich, I. & Chirio, L. (2005) Molecular review of the genus Panaspis sensu lato (Reptilia: Scincidae) in Cameroon, with special reference to the status of the proposed subgenera. Zootaxa, 863 (1), 1 - 28. https: // doi. org / 10.11646 / zootaxa. 863.1.1","Ceriaco, L. M. P., de Sa, S. A. C., Bandeira, S., Valerio, H., Stanley, E. L., Kuhn, A. L., Marques, M. P., Vindum, J. V., Blackburn, D. C. & Bauer, A. M. (2016 a) Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a synoptic list of the amphibians and reptiles of Namibe Province, southwestern Angola. Proceedings of the California Academy of Sciences, Series 4, 63, 15 - 61.","Ceriaco, L. M. P., Marques, M. P. & Bandeira, S. A. (2016 b) Anfibios e Repteis do Parque Nacional da Canganda. Instituto Nacional da Biodiversidade e Areas de Conservacao, Angola & Museu Nacional de Historia Natural e da Ciencia, Lisboa, 97 pp.","Medina, M. F., Bauer, A. M., Branch, W. R., Schmitz, A., Conradie, W., Nagy, Z. T., Hibbitts, T. J., Ernst, R., Portik, D. M., Nielsen, S. V., Colston, T. J., Kusamba, C., Behangana, M. & Rodel, M. - O. (2016) Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution, 100, 409 - 423. https: // doi. org / 10.1016 / j. ympev. 2016.04.026","Ceriaco, L. M. P., Branch, W. R. & Bauer, A. M. (2018 a) A new species of African snake-eyed skink (Scincidae: Panaspis) from central and northern Namibia. Zootaxa, 4527 (1), 140 - 150. https: // doi. org / 10.11646 / zootaxa. 4527.1.12","Ceriaco, L. M. P., Marques, M. P., Bandeira, S., Blackburn, D. C. & Bauer, A. M. (2018 c) Herpetological survey of Cangandala National Park with a synoptic list of the amphibians of Malanje Province, central Angola. Herpetological Review, 49 (3), 408 - 431.","Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (supplement 2), 1 - 501.","Branch, W. R., Pinto. P. V., Baptista, N. & Conradie, W. (2019 a) The Reptiles of Angola: History, Diversity, Endemism and Hotspots. In: Huntley, B, Russo, V., Lages & Ferrand, N. (Eds.) Biodiversity of Angola, Science & Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13","Jacobsen, N. H. G. & Broadley, D. G. (2000) A new species of Panaspis Cope (Reptilia: Scincidae) from southern Africa. African Journal of Herpetology, 49 (1), 61 - 71. https: // doi. org / 10.1080 / 21564574.2000.9650017","Cope, E. D. (1868) Observations on reptiles of the old world. Proceedings of the Academy of Natural Sciences of Philadelphia, 20 (1868), 316 - 323.","Fuhn, I. E. (1972) Revision du phylum forestier du genre Panaspis Cope (Reptilia, Scincidae, Lygosominae). Revue Roumaine de Biologie, serie de Zoologie, 17 (4), 257 - 272."]}