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The morphological ontogeny of Oribatella reticulata Berlese, 1916 is described and illustrated. The nymphs of this species are apopheredermous, i.e. they carry the exuvial scalps of previous instar...

Oribatella similesuperbula Weigmann, 2001 (Figs. 1���15) Morphology of adult Adults studied here (Figs. 1, 2, 3a, 4, 5a) similar to those investigated by Weigmann (2001), but see Remarks. Mean length (and range) of females���374.2��8.5 (361���391, N= 30) and males���350.3��6.6 (range 343���361, N= 10); mean width of females���245.9��7.7 (235���259) and males���231.7��3.2 (229���235). Subcapitular setae h, m and a short and smooth (Fig. 2). Chelicera chelate, seta cha longer than chb, both barbed (Fig. 3b). Palp relatively short, palpal setae smooth, except for slightly barbed l��� on tibia (Fig. 3c). Formula of palp setae [trochanter to tarsus (+ solenidion ��)]: 0-2-1-3-9(1). Most epimeral setae short and smooth, except for longer and barbed 3b, 3c, 4a and 4b and clearly longer, thicker and barbed 4c (Fig. 2). Most leg setae barbed, l��� on genua and tibiae I and II, and l��� on genu and tibia IV longer and thicker than other setae (Fig. 6). Formulae of leg setae [trochanter to tarsus (+ solenidia)]: I���1-5-3(1)-4(2)-20(2); II���1-5-3(1)-4(1)-15(2); III���2-2-1(1)-3(1)-15; IV���1-2-2-3(1)-12. Legs heterotridactylous. Remarks: Females and males of O. similesuperbula investigated here are slightly longer than those investigated by Weigmann (2001) ���length of females 340���380, and males 320���340. In our adults, the lamellar complex is located more anterior than depicted by Weigmann (2001), and rostrum is not visible in dorsal aspect (versus visible). Other characters of adults are similar as in Weigmann (2001). Diagnosis of juveniles Juveniles light-brown and nymphs apopheredermous, i.e. they carry exuvial scalps of previous instars away from dorsal integument, using modified setal pair da. Prodorsal and most gastronotal setae of juveniles long and barbed, except for medium sized h 2 and minute h 3 in larva, and medium sized c 1, p 2 and p 3 in nymphs. Larva with 12 pairs of gastronotal setae, and central setae thicker than other setae, nymphs with 15 pairs, and long d -series. In all juveniles, humeral organ present. In nymphs, seta d on femora I and II, l��� on genu and tibia I and genu II, and l��� on genu III clearly thicker than other leg setae. Description of juvenile stages Larva oval in dorsal aspect (Figs. 5b���d, 7) and light brown. Prodorsum subtriangular, all prodorsal setae long and barbed, in longest, ro shortest (Table 1). Mutual distance between setal pairs le slightly longer than between pair ro, and between pair in about two times longer than between pair ro. Seta le inserted midway between pairs ro and in. Bothridium rounded, bothridial seta narrowly clavate and barbed. Gastronotum of larva with 12 pairs of setae, including minute h 3 inserted laterally to medial part of anal valves (Figs. 7, 8a, 9a). Most gastronotal setae long and thick, with clear barbs, except for medium sized h 2 and minute h 3 (Table 1), central setae thicker than lateral and posterior setae (Figs. 5b���d, 7, 9a). Anal valves (segment P) glabrous. Cupule ih lateral to anterior part of anal valves, cupules ia, im, and ip not observed, gland opening lateroventral to seta lp, humeral organ anterior to seta c 3 (Figs. 8a, 9a). Most leg setae barbed, l��� on genu and tibia I and genu II, and l��� on genu III thicker than other leg setae (Fig. 10). Seta ft��� on tarsus I and ft on tarsi II and III slightly thicker than other tarsal setae. Nymphs more stocky than larva, with setiform bothridial seta. Gastronotum of protonymph with 15 pairs of setae because setae of p -series appearing (Fig. 8b), and remaining in deutonymph and tritonymph (Figs. 9b, 11���13a, 13d, 14). In all nymphs, most gastronotal setae long and barbed, except for medium sized and slightly barbed c 1, p 2 and p 3, p 1 shorter than other posterior setae. Setal pair da long, barbed, inserted close to each other, with apical end adapted for bearing exuvial scalps of previous instars. Mutual distance between setal pair c 1 about two times longer than between setal pair da, mutual distance between setal pair dm and dp about 5.5, and 5 times longer than between setal pair da, respectively. Setae c 3, lm, lp, h -series and p 1 in marginal position on gastronotum. Protonymph, deutonymph and tritonymph carry exuvial scalps of previous instars (Fig. 9b), using long setal pair da. In protonymph, one pair of setae appearing on genital valves, and two pairs added in deutonymph and tritonymph each (Figs. 8b, 11); all short and smooth. In deutonymph, one pair of aggenital setae and three pairs of adanal setae appearing, and present in tritonymph; all short and smooth. Anal valves of protonymph (segment AD) and deutonymph (segment AN) glabrous, but in tritonymph two pairs of setae present, all short and smooth. In all nymphs, humeral organ placed as in larva. In deutonymph and tritonymph, cupule iad lateral to anterior part of anal valves, cupules ia, im, ip, ih and ips not observed, opisthonotal gland opening as in larva (Figs. 9b, 11). In deutonymph and tritonymph, leg setae d on femora I and II, l��� on genua I and II and tibia I, and l��� on genu III clearly thicker than other setae (Figs. 13a, 13b, 14, 15). Summary of ontogenetic transformations In all instars of O. similesuperbula, the prodorsal setae are long, and le is as thick as ro and in, but in the adult le is clearly thicker than other setae. In all juveniles, seta ex is long, whereas in the adult it is short. The opening of bothridium is rounded in all instars, but in the adult it gains small scales. In the larva, the bothridial seta is narrowly clavate, in the nymphs it is setiform, whereas in the adult it is fusiform. The larva has 12 pairs of gastronotal setae, including minute h 3, the nymphs have 15 pairs (p -series appear), whereas the notogaster of adult loses setae c 2, c 3 and of d -series, such that 10 pairs of setae remain. The formula of gastronotal setae is 12-15-15-15-10 (from larva to adult), and formulae of epimeral setae are 3-1-2 (larva, including scaliform 1c), 3-1-3-1 (protonymph), 3-1-3- 2 (deutonymph) and 3-1-3-3 (tritonymph and adult). Formula of genital setae is 1-3-5-6 (protonymph to adult), aggenital setae is 1-1-1 (deutonymph to adult), and formula of segments PS���AN is 03333-0333-022. The ontogeny of leg setae and solenidia of O. similesuperbula is given in Table 2. Distribution, ecology, and biology Oribatella similesuperbula has Central European distribution (Sub��as (2004, update 2021). It was recorded from Germany (Weigmann 2006) and Romania (Călugăr & Ivan 2013). Weigmann (2001, 2006) noted the preference of O. similesuperbula to moss and decaying wood. In Romania this species was first recorded from a forest plantation, with dominant Robinia pseudoacacia L. (Călugăr & Ivan 2013), but was more abundant in bryophyte covering calcareous cliffs, which suggests its preference to such a habitat. In this study, O. similesuperbula was distinctly more abundant in Horia (42 individuals per 500 cm 3) than in D��mbovicioara Gorges (2 individuals per 500 cm 3). In both habitats, the adults dominated the juveniles, which constituted 49% and 20% of sample populations, respectively. In the samples in Horia, the stage structure of this species was the following: 25 larvae, 45 protonymphs, 33 deutonymphs, one tritonymph and 107 adults. Among 40 investigated adults, the sex ratio (females: males) of O. similesuperbula was 1:0.25, and 16% of females were gravid. The gravid females carried 1���2 relatively large eggs, each 152 x 104, which constitutes about 41% of the total body length of females. In all habitats, females of O. similesuperbula were significantly longer and wider than males. Note: structures are indicated where they are first added and are present through the rest of ontogeny; pairs of setae in parentheses, dash indicates no additions., Published as part of Seniczak, Stanis��aw, Ivan, Otilia, Marquardt, Tomasz & Seniczak, Anna, 2021, Morphological ontogeny of Oribatella similesuperbula (Acari: Oribatida: Oribatellidae), pp. 90-110 in Zootaxa 5086 (1) on pages 91-105, DOI: 10.11646/zootaxa.5086.1.8, http://zenodo.org/record/5814523, {"references":["Weigmann, G. (2001) Contribution to the taxonomy of European Poronota I. Oribatella and Anachipteria. Spixiana, 24 (3), 235 - 240.","Subias, L. S. (2004 [update 2021]) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (Numero Extraordinario), 3 - 305. [15 ª actualizacion, 527 pp.] https: // doi. org / 10.3989 / graellsia. 2004. v 60. iextra. 218","Calugar, A. & Ivan, O. (2013) Diversity and distribution of the edaphic mites (Acari: Gamasina, Oribatida) in some forest plantations from the Central Moldavian Plateau. Scientific papers, Iasi, Agronomy Series, 56 (2), 97 - 103."]}

Oribatella calcarata (C.L. Koch, 1835) Oribatella calcarata Bernini 1977, p. 476, figs VIII, IX. Oribatella calcarata Weigmann 2006, p. 363, fig. 194a-e. Parco Naturale delle Alpi Marittime: AB single record. General distribution: Northern, southern Italy; Italian, Swiss, Austrian Alps, Piedmont (Torino), southern France; Holarctic. Habitat preferences: silvicolous; hygrophilous., Published as part of Schatz, Heinrich, Fortini, Lorenzo, Fusco, Tommaso, Casale, Francesca, Jacomini, Carlo & Giulio, Andrea Di, 2021, Oribatid mites (Acari, Oribatida) from " Parco Naturale delle Alpi Marittime " (Piedmont, Italy), pp. 501-540 in Zootaxa 5082 (6) on page 527, DOI: 10.11646/zootaxa.5082.6.1, http://zenodo.org/record/5797624, {"references":["Bernini, F. (1977) Notulae Oribatologicae XVII. Nuovi dati sul genere Oribatella in Italia (Acarida, Oribatei). Redia, 60, 457 - 505.","Weigmann, G. (2006) Hornmilben (Oribatida). Die Tierwelt Deutschlands. Teil 76. Goecke & Evers, Keltern, 520 pp."]}

Oribatella quadricornuta (Michael, 1880) Oribatella quadricornuta Bernini 1975, p. 445, figs 7-11. Oribatella quadricornuta Weigmann 2006, p. 364, fig. 194f-h. Parco Naturale delle Alpi Marittime: PAL sparse. General distribution: Northern, southern Italy; Italian, Swiss, Austrian Alps, Piedmont (Torino); Holarctic. Habitat preferences: silvicolous, praticolous; xerophilous., Published as part of Schatz, Heinrich, Fortini, Lorenzo, Fusco, Tommaso, Casale, Francesca, Jacomini, Carlo & Giulio, Andrea Di, 2021, Oribatid mites (Acari, Oribatida) from " Parco Naturale delle Alpi Marittime " (Piedmont, Italy), pp. 501-540 in Zootaxa 5082 (6) on page 527, DOI: 10.11646/zootaxa.5082.6.1, http://zenodo.org/record/5797624, {"references":["Bernini, F. (1975) Notulae Oribatologicae XIII. La famiglia Oribatellidae (Acarida, Oribatei) nell'Arcipelago Toscano. (Studi sulla Riserva Naturale dell'Isola di Montecristo, X). Lavori della Societa Italiana di Biogeografia, Nuova Serie, 5, 429 - 504. https: // doi. org / 10.21426 / B 65110013","Weigmann, G. (2006) Hornmilben (Oribatida). Die Tierwelt Deutschlands. Teil 76. Goecke & Evers, Keltern, 520 pp."]}

Oribatella hungarica Balogh, 1943 Oribatella hungarica Balogh 1943, p. 95, 147, tab. XVII, fig. 8. Oribatella hungarica Bernini 1978, p. 506, fig. Ia-i, Tab. I-III. Parco Naturale delle Alpi Marittime: AND-I sparse. General distribution: Northern, southern Italy; Italian, Austrian Alps, Piedmont (Torino); Central, South, Southeast Europe; Mediterranean. Habitat preferences: silvicolous; xerophilous? Remark: Body length 320 ��m., Published as part of Schatz, Heinrich, Fortini, Lorenzo, Fusco, Tommaso, Casale, Francesca, Jacomini, Carlo & Giulio, Andrea Di, 2021, Oribatid mites (Acari, Oribatida) from " Parco Naturale delle Alpi Marittime " (Piedmont, Italy), pp. 501-540 in Zootaxa 5082 (6) on page 527, DOI: 10.11646/zootaxa.5082.6.1, http://zenodo.org/record/5797624, {"references":["Balogh, J. (1943) Magyarorszag Pancelosatkai (Conspectus Oribateorum Hungariae). Matematikai es termeszettudomanyi kozlemenyek, 39 (5), 1 - 202.","Bernini, F. (1978) Notulae Oribatologicae XX. Il Genere Oribatella in Italia (Acarida, Oribatida). Redia, 6, 503 - 538."]}

Oribatella euthricha Berlese, 1908 Oribatella eutricha (sic!) Bernini 1977, p. 486, figs X, XI, Tab. VIII, VIII. Parco Naturale delle Alpi Marittime: FAG, PAL very numerous. General distribution: Northern, southern Italy; Italian, Swiss, Austrian Alps; Central, South, Southeast Europe, Caucasus; Mediterranean. Habitat preferences: silvicolous., Published as part of Schatz, Heinrich, Fortini, Lorenzo, Fusco, Tommaso, Casale, Francesca, Jacomini, Carlo & Giulio, Andrea Di, 2021, Oribatid mites (Acari, Oribatida) from " Parco Naturale delle Alpi Marittime " (Piedmont, Italy), pp. 501-540 in Zootaxa 5082 (6) on page 527, DOI: 10.11646/zootaxa.5082.6.1, http://zenodo.org/record/5797624, {"references":["Bernini, F. (1977) Notulae Oribatologicae XVII. Nuovi dati sul genere Oribatella in Italia (Acarida, Oribatei). Redia, 60, 457 - 505."]}

Oribatella hungarica Balogh, 1943 Morphology of adult Adults studied here (Figs. 1, 2, 3a, 4 a���c, 5) similar to those investigated by Balogh (1943) and Bernini (1978), but see Remark below. Mean length (and range) of females 311.3��8.87 (295���337, N=100) and males 294.4��6.38 (range 277���307, N=50); mean width of females 201.5��5.94 (187���217) and males 189.2��6.11 (175���205). Subcapitular se- tae h and m longer than a, all barbed (Fig. 2). Chelicera chelate, seta cha longer than chb, both barbed (Fig. 3b). Palp relatively short, palpal setae sup and inf longer than other setae, sup, inf and l��� on tibia barbed, other setae smooth (Fig. 3c). Formula of palp setae [trochanter to tarsus (+ solenidion ��)]: 0-2-1-3-9(1). Epimeral setae 1a, 1c, 2a and 3a short (Fig. 2) and smooth, seta 1b slightly longer, setae 3b, 3c, 4a and 4b clearly longer and barbed, and seta 4c longest and thickened; all barbed. Most leg setae barbed, seta l��� on genu I and II and tibia I and II, and l��� on genu III and genu and tibia IV thickened (Fig. 5, Table 2). Formulae of leg setae [trochanter to tarsus (+ solenidia), Table 3)]: I���1-5-3(1)-4(2)-20(2); II���1-5-3(1)-4(1)-15(2); III���2-3-1(1)-3(1)-15; IV���1-2-2-3(1)-12. Legs bidactylous. Remark. The adults of O. hungarica investigated here are smaller than those investigated by Bernini (1978; length of females 330���390, males 290���390; width of females 255���260, males 195���255), but other characters are similar as in Balogh (1943) and Bernini (1978). Note: structures are indicated where they are first added and are present through the rest of ontogeny; pairs of setae in parenthe- ses, dash indicates no additions. 1 In some individuals present on one leg, if not it appears in the next stage. Diagnosis of juveniles Juveniles light-brown and nymphs apopheredermous, i.e. they carry exuvial scalps of previous instars away from dorsal integument, using modified setal pair da. Larva with 12 pairs of gastronotal setae, most with long barbs, except for c 3 with short barbs and minute h 3, nymphs with 13 pairs of these setae, with short barbs (dm and dp are lacking). In tritonymph, leg seta d on all femora and genu IV, l��� on femora I and II and pair l on genua and tibiae I and II, l��� on femur III and genua and tibiae III and IV, and ft on tarsi II���IV thicker than other setae on these segments. In all juveniles, humeral organ present. Description of juvenile stages Larva oval in dorsal aspect (Figs. 4d, 6) and light brown. Prodorsum subtriangular, all prodorsal setae long, with short barbs, in longest, ro shortest (Table 4). Mutual distance between setal pairs le about two times longer than between pair ro, and between pair in about three times longer than between pair ro. Seta le inserted midway between pairs ro and in. Bothridium rounded, bothridial seta narrowly clavate and barbed. Gastronotum of larva with 12 pairs of setae, including minute h 3 inserted laterally to medial part of anal valves (Figs. 6, 7a, 8a). Most gastronotal setae long, with long barbs, except for long c 3 and medium sized h 2 with short barbs, and minute h 3; length of setae decreasing from anterior to posterior (Figs. 6, 8a, Table 4). Anal valves (seg- ment P) glabrous. Cupules ia, im, ip and ih not observed, gland opening lateroventral to seta lp. Humeral organ anterior to seta c 3 present (Fig. 8a). Most leg setae barbed, l��� on genu and tibia I and genu II, and l��� on genu III longer than other leg setae (Fig. 9). Nymphs more stocky than larva, with setiform bothridial seta, but in protonymph slightly clavate, length and distribution of prodorsal setae as in larva. Gastronotum of protonymph with 13 pairs of setae because setae of p - series appearing (Fig. 7b), and remaining in deutonymph and tritonymph (Figs. 10a, 10b), but setae of dm and dp lost and remaining absent in other nymphs. In all nymphs, most gastronotal setae long (Table 4), except for medium sized p 2 and p 3; p 2 slightly longer than p 3. Setal pair da long, smooth, inserted close to each other and posterior to setal pair c 1, apical end of da adapted for bearing exuvial scalps of previous instars. Other gastronotal setae in marginal position on gastronotum and barbed, but setae p 2 and p 3 with shorter barbs than other setae. All nymphs carry exuvial scalps of previous instars (Fig. 8b), using long setal pair da. In protonymph, one pair of setae appearing on genital valves, and two pairs added in deutonymph and tritonymph each (Figs. 7b, 10a, 10b); all short and smooth. In deutonymph, one pair of aggenital setae and three pairs of adanal setae appearing, and present in tritonymph; all short and smooth. Anal valves of protonymph (segment AD) and deutonymph (segment AN) glabrous, but in tritonymph two pairs of anal setae present, all short and smooth. Cupules ia, im, ip, ih, ips and iad not observed, opisthonotal gland opening as in larva (Figs. 7b, 8b, 10a, 10b, 11). In tritonymph, leg setae d on all femora and genu IV, l��� on femora I and II and pair l on genua and tibiae I and II, l��� on femur III and genua and tibiae III and IV, and ft on tarsi II���IV thicker than other setae on these segments (Fig. 12, Table 2). Solenidia �� 1 and �� 2 on tibia I curved outwards. In all nymphs, humeral organ as in larva. Summary of ontogenetic transformations In all instars of O. hungarica, the prodorsal setae are long, and seta in is longest and ro is shortest, but in the juveniles le is clearly longer than ro, whereas in the adult these setae are of similar length, and le is clearly thicker than other setae. In all juveniles, seta ex is long, whereas in the adult it is short. The opening of bothridium is rounded in all instars, but in the adult it gains small scales. In the larva, the bothridial seta is narrowly clavate, whereas in other instars it is fusiform. The larva has 12 pairs of gastronotal setae, including minute h 3, the nymphs have 13 pairs (p - series appears, dm, dp are lost), whereas the notogaster of adult loses setae c 1, c 3 and da, such that 10 pairs of setae remain. The formula of gastronotal setae is 12-13-13-13-10 (from larva to adult), and formulae of epimeral setae are 3-1-2 (larva, including scaliform 1c), 3-1-3-1 (protonymph), 3-1-3-2 (deutonymph) and 3-1-3-3 (tritonymph and adult). Formula of genital setae is 1-3-5-6 (protonymph to adult), aggenital setae is 1-1-1 (deutonymph to adult), and formula of segments PS���AN is 03333-0333-022. The ontogeny of leg setae is shown in Table 3. Distribution, ecology and biology Oribatella hungarica has a Mediterranean distribution (Sub��as 2020) and is considered sylvicolous (Mahunka & Mahunka-Papp 2004, Ivan & Vasiliu 2006). This species was described from Hungary (Balogh 1943), and recorded from Italy (Bernini et al. 1995), southern Austria (Krisper et al. 2017), Hungary (Mahunka & Mahunka-Papp 2004), Bulgaria (Csisz��r & Jeleva 1962) and Romania (Ivan &Vasiliu 2000). Oribatella hungarica is frequent and rela- tively abundant in litter and moss of mesophilous forests in hilly regions (Ivan & Vasiliu 2000). This species tolerated the cement dust pollution in beech forests in Romanian Southern Subcarpathians (Ivan & Vasiliu 2012), and also chemicals used against defoliating insects in oak forests (Ivan 2009), but is sensitive to the forest soil pollution with heavy metals and fluorine (Ivan & Vasiliu 2009). 1 plant cover as in Table 1; a,b,cdifferent letters in superscript mean significantly different between plots at ��= 0.05; *significantly different between females and males. In this study, O. hungarica was present in the upper soil horizon of different oak forests (Table 1), with the highest density in Roşcani (26.8 individuals per 500 cm 3), and lowest in Heltiu (7.8 individuals per 500 cm 3). In all soils, the adults highly dominated the juveniles, which constituted 1���11% of sample populations or were absent. In the soil in Ampoiţa, all instars of O. hungarica were present, and the stage structure of this species was the following: one larva, one protonymph, two deutonymphs, three tritonymphs and 41 adults. In these soils, females of O. hungarica were more abundant than males, the sex ratio (females: males) varied (1:0.43���1:0.58), and 15���58% of females were gravid (Table 5). The gravid females carried 1���2 relatively large eggs, each 163���169 x 102���108, which constituted about 53% of the total body length of females. In most oak forests, the differences of body length and width of females and males of O. hungarica were indistinct. Only in Heltiu, females were significantly longer than in Ampoiţa and Roşcani, and only in Roşcani males were significantly shorter than in other forests. In Ampoiţa, females were significantly wider than in Copălău and Heltiu, and in Urecheşti males were significantly wider than in Roşcani, Copălău and Ampoiţa (Table 5). In all for- ests, females of O. hungarica were significantly longer and wider than males., Published as part of Seniczak, Stanis��aw, Ivan, Otilia, Marquardt, Tomasz & Seniczak, Anna, 2020, Morphological ontogeny of Oribatella hungarica (Acari: Oribatida: Oribatellidae), pp. 137-159 in Zootaxa 4857 (1) on pages 139-150, DOI: 10.11646/zootaxa.4857.1.8, http://zenodo.org/record/4396627, {"references":["Balogh, J. (1943) Magyarorszag Pancelosatkai (Conspectus Oribateorum Hungariae). Matematikai es termeszettudomanyi kozlemenyek, 39 (5), 1 - 202.","Bernini, F. (1978) Notulae Oribatologicae XX. Il Genere Oribatella in Italia (Acarida, Oribatida). Redia, 61, 503 - 538.","Mahunka, S. & Mahunka-Papp, L. (2004) A Catalogue of the Hungarian oribatid mites (Acari: Oribatida). In: Csuzdi, Cs. & Mahunka, S. (Eds.), Pedozoologica Hungarica. No. 2. Hungarian Natural History Museum and Systematic Zoology Research Group of the Hungarian Academy of Sciences, Budapest, pp. 1 - 363.","Ivan, O. & Vasiliu, N. (2006) Diversity and distribution of the oribatid mites (Acari, Oribatida) in the forest ecosystem from the middle section of the Prut riverside. Scientific Annals of the Danube Delta Institute, 12, 55 - 64.","Bernini, F., Castagnoli, M. & Nannelli, R. (1995) Arachnida, Acari. In: Minelli, A., Rufo, S. & La Posta, S. (Eds.), Checklist delle specie della fauna italiana. Vol. 24. Calderini, Bologna, pp. 1 - 131.","Krisper, G., Schatz, H. & Schuster, R. (2017) Oribatida (Arachnida: Acari). In: Schuster, R. (Ed.), Checklists of the Austrian Fauna. No. 9. Biosystematics and Ecology Series No. 33. Austrian Academy of Sciences Press, Vienna, pp. 25 - 90. https: // doi. org / 10.2307 / j. ctvdjrr 69.5","Csiszar, J. & Jeleva, M. (1962) Oribatid mites (Acari) from Bulgarian soils. Acta Zoologica Academiae Scientiarum Hungaricae, 8 (3 - 4), 273 - 301.","Ivan, O. & Vasiliu, M. (2000) Oribatid fauna (Acari, Oribatida) in Romanian Quercus forests. Anuarul Complexului Muzeal Bucovina, Suceava, 15, 67 - 116.","Ivan, O. & Vasiliu, N. A. (2012 / 2010) Influence of the pollution with cement dust on the oribatid communities (Acari, Oribatida) in forest soils from Campulung Muscel zone (Romania). Scientific papers, Agronomy Series, University of Agricultural Sciences and Veterinary Medicine Iasi, 53 (3), 186 - 192.","Ivan, O., Vasiliu, N. A. (2009) Oribatid mites (Acari, Oribatida) - bioindicators of forest soils pollution with heavy metals and fluorine. Annals of Forest Research, 52, 11 - 18."]}

Oribatella palustris Hammer, 1962 (Figures 1���11), Published as part of Bayartogtokh, Badamdorj & Ermilov, Sergey G., 2019, Ontogeny of morphological traits in Oribatella palustris Hammer, 1962, with remarks on juveniles of Oribatellidae (Acari: Oribatida), pp. 85-103 in Zootaxa 4717 (1) on page 87, DOI: 10.11646/zootaxa.4717.1.8, http://zenodo.org/record/3602068, {"references":["Hammer, M. (1962) Investigations on the oribatid fauna of the Andes Mountains. III. Chile. Det Kongelige Danske Videnskabernes Selskab Biologiske Skrifter, 13 (2), 1 - 96."]}

In this work, the larva and tritonymph of Oribatella palustris, an Antarctic and Subantarctic species of the Neotropical region are investigated along with supplementary re-description of adults. Comparative characteristics of juveniles of Oribatellidae, especially larvae and tritonymphs, are given based on this study and available literature sources; keys for identification of larvae and tritonymphs of Oribatella are also presented. Previous record of Oribatella palustris from a tropical island of Mexico is questioned.

Oribatella pawnee Behan-Pelletier and Walter, 2012 Geographic Location: AB: ABMI-OG-1122-1 (53.1648407, -111.919342), along Township road 484 E of Bruce; ABMI-1125 (53.07548141, -111.074829), E of juncture Routes 619 and 881; ABMI-OG-1192-1 (52.66613007, -110.693031), north of Wainwright Dunes Ecological Reserve; ABMI-1224 (52.50389099, - 110.730148), W of juncture Highways 41 and 13; ABMI-1247 (52.51489258, -112.811302), SE of juncture Highways 53 and 56 near Buffalo Lake; ABMI-1556 (50.07164764, -112.112526), near Vauxhall; ABMI- 1558 NE (50.00374222, -111.578972), south of Ronalane (Behan-Pelletier & Walter 2012); ABMI Sites (Walter et al. 2014). Habitats: from upper soil organic layers on untreed site with shrubby vegetation, in dry grassland; rich fen with shrubs, sedge, and grass understory. Distribution: Nearctic., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 116, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of Western North America. Zootaxa, 3432 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3432.1.1","Walter, D. E., Latonas, S., Byers, K. & Lumley, L. M. (2014) Almanac of Alberta Oribatida Part I. Ver. 2.4. Royal Alberta Museum, Edmonton, AB. 542 pp. Available from: https: // www. royalalbertamuseum. ca / research / lifeSciences / invertebrateZoology / research. cfm (accessed 20 March 2019)"]}

Oribatella maryae Behan-Pelletier and Walter, 2012 Geographic Location: BC: Vancouver Is., Trail Is. Ecological Reserve, 48°24N, 123°18'W; Cowichan Lake Research Station, 48°49'N, 124°07'W; Carmanah PP (Behan-Pelletier & Walter 2012). Habitats: lichen on rocks; bark and moss on trunk of Douglas fir. Distribution: Nearctic., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 115, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of Western North America. Zootaxa, 3432 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3432.1.1"]}

Oribatella brevicornuta Jacot, 1934 Geographic Location: QC: Parc National du Mont-Orford; Mont Joli; Cedarville, Descente 20 (Behan- Pelletier 2011); Abitibi (Déchêne & Buddle 2009, 2010); NS: Cape Breton Highlands NP, Mackenzie Mountain (Behan-Pelletier 2011). Habitats: beech litter, some Lycopodium; litter under Hericium on fallen dead beech; boreal mixedwood forest. Distribution: Holarctic, Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 114, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Dechene, A. D. & Buddle, C. M. (2009) Effects of experimental forest harvesting on oribatid mite biodiversity. Forest Ecology and Management, 258, 1331 - 1341. https: // doi. org / 10.1016 / j. foreco. 2009.06.033","Dechene, A. D. & Buddle, C. M. (2010) Decomposing logs increase oribatid mite assemblage diversity in mixedwood boreal forest. Biodiversity Conservation, 19, 237 - 256. https: // doi. org / 10.1007 / s 10531 - 009 - 9719 - y","Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56. https: // doi. org / 10.11646 / zootaxa. 2973.1.1"]}

Oribatella ewingi Behan-Pelletier and Walter, 2012 Geographic Location: BC: Vancouver Is., Smuggler's Cove PP; Manning PP; Osoyoos, Haynes Lease Reserve; Penticton, Madeline Lake (Behan-Pelletier & Walter 2012); AB: ABMI-996 (54.39379883, - 119.614182); ABMI-1231 (52.99143219, -117.452957) (Behan-Pelletier & Walter 2012); ABMI Sites (Walter et al. 2014). Habitats: Douglas fir and sassafras litter; Rhododendron litter; Heuchera litter; upper soil organic layers in mesic upland pine forest with Rhododendron and feathermoss. Distribution: Nearctic., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 114, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of Western North America. Zootaxa, 3432 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3432.1.1","Walter, D. E., Latonas, S., Byers, K. & Lumley, L. M. (2014) Almanac of Alberta Oribatida Part I. Ver. 2.4. Royal Alberta Museum, Edmonton, AB. 542 pp. Available from: https: // www. royalalbertamuseum. ca / research / lifeSciences / invertebrateZoology / research. cfm (accessed 20 March 2019)"]}

Oribatella banksi Behan-Pelletier and Walter, 2012 Geographic Location: BC: Vancouver Is., 48°18.8N, 123°36.07W (Behan-Pelletier & Walter 2012); AB: Wood Buffalo NP (Behan-Pelletier & Walter 2012); ABMI Sites (Walter et al. 2014). Habitats: mixed coniferous and deciduous forest; soil and litter under Pacific yew; mesic upland pine and aspen forests. Distribution: Nearctic., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 114, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of Western North America. Zootaxa, 3432 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3432.1.1","Walter, D. E., Latonas, S., Byers, K. & Lumley, L. M. (2014) Almanac of Alberta Oribatida Part I. Ver. 2.4. Royal Alberta Museum, Edmonton, AB. 542 pp. Available from: https: // www. royalalbertamuseum. ca / research / lifeSciences / invertebrateZoology / research. cfm (accessed 20 March 2019)"]}

Oribatella canadensis Behan-Pelletier and Eamer, 2010 Geographic Location: AK: Fairbanks (Behan-Pelletier & Walter 2012); YT: Ivvavik NP, British Mtns, Firth River floodplain, 68°54'N, 140°23'W (Behan-Pelletier & Walter 2012); BC: Cathedral PP (Behan- Pelletier & Walter 2012); AB: Writing-on-Stone PP; Waterton Lakes NP; Cypress Hills PP; ABMI-82H-5 (49.01377, -112.29149), south of Route 501 (Behan-Pelletier & Eamer 2010; Behan-Pelletier & Walter 2012); ABMI Sites (Walter et al. 2014); ON: near Chapleau (Rousseau et al. 2018). Habitats: under bark of fallen Balsam poplar; dry litter of Larix lyallii; Loblolly pine litter; under hawthorn; bracket fungi on poplar log; Jack pine forest. Distribution: Nearctic., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 114, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. & Eamer, B. (2010) The first sexually dimorphic species of Oribatella (Acari, Oribatida, Oribatellidae) and a review of sexual dimorphism in the Brachypylina. Zootaxa, 2332, 1 - 20. https: // doi. org / 10.11646 / zootaxa. 2332.1.1","Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of Western North America. Zootaxa, 3432 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3432.1.1","Walter, D. E. & Latonas, S. (2012) Almanac of Alberta Acari Part II. Ver. 2.1. Royal Alberta Museum, Edmonton, AB. 159 pp. Available from: https: // www. royalalbertamuseum. ca / research / lifeSciences / invertebrateZoology / research. cfm (accessed 20 March 2019)","Walter, D. E., Latonas, S., Byers, K. & Lumley, L. M. (2014) Almanac of Alberta Oribatida Part I. Ver. 2.4. Royal Alberta Museum, Edmonton, AB. 542 pp. Available from: https: // www. royalalbertamuseum. ca / research / lifeSciences / invertebrateZoology / research. cfm (accessed 20 March 2019)","Rousseau, L., Venier, L., Hazlett, P., Fleming, R., Morris, D. & Handa, I. T. (2018) Forest floor mesofauna communities respond to a gradient of biomass removal and soil disturbance in a boreal Jack pine (Pinus banksiana) stand of northeastern Ontario (Canada). Forest Ecology and Management, 407, 155 - 165. https: // doi. org / 10.1016 / j. foreco. 2017.08.054"]}

Oribatella abmi Behan-Pelletier and Walter, 2012 Geographic Location: AB: ABMI 1126 (53.05752182, -110.746841), 7 km east of Auburndale along Township Road 473; ABMI-OG-1160-1 (52.78925323, -110.302109), 11 km north-northeast of Edgerton along Township Road 442; ABMI-OG-1190-1 (52.71193, -111.23734), 2 km east of Route 881 along Township Road 433 northeast of Hardisty; ABMI-OG-1283-1 (52.13767624, -110.796417), 2 km NW Gooseberry Lake PP; ABMI-72L-3 (50.75932, -110.76165); ABMI-72L-4 (50.77691, -110.90216); Edmonton, Parkallen; ABMI Sites (Behan-Pelletier & Walter 2012; Walter et al. 2014). Habitats: upper organic soil layer in open field; untreed site with lichen, bryophytes, grasses and sedges; upper organic soil layer under liverworts and mosses in urban garden. Distribution: Canada., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 113, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of Western North America. Zootaxa, 3432 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3432.1.1","Walter, D. E., Latonas, S., Byers, K. & Lumley, L. M. (2014) Almanac of Alberta Oribatida Part I. Ver. 2.4. Royal Alberta Museum, Edmonton, AB. 542 pp. Available from: https: // www. royalalbertamuseum. ca / research / lifeSciences / invertebrateZoology / research. cfm (accessed 20 March 2019)"]}

Oribatella nortoni Behan-Pelletier, 2011 Geographic Location and Habitats: NB: Kouchibouguac NP, moss and litter at base of living and dead Red oak; moose dung in mixedwoods; mixed deciduous litter (Behan-Pelletier 2011). Distribution: Nearctic., Published as part of Behan-Pelletier, Valerie M. & Lindo, Zoë, 2019, Checklist of oribatid mites (Acari: Oribatida) of Canada and Alaska, pp. 1-180 in Zootaxa 4666 (1) on page 115, DOI: 10.11646/zootaxa.4666.1.1, http://zenodo.org/record/4000595, {"references":["Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56. https: // doi. org / 10.11646 / zootaxa. 2973.1.1"]}

Two new species of oribatid mites, Oribatella paraumaetluisorum sp. n. and Taiwanoppia nepalica sp. n., are described from Nepal. The genera Oribatella and Taiwanoppia are recorded from this country for the first time.

Oribatella (Oribatella) serrata Balogh & Mahunka, 1969 Oribatella serrata Balogh & Mahunka, 1969 Neotropical region. Brazil: States of Amazonas and Roraima. References: Ribeiro & Schubart (1989): 260, Adis et al. (2002): 11, Woas (2002): 287, Franklin et al. (2006): 1016., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 61, DOI: 10.5281/zenodo.437584, {"references":["Ribeiro, E. F. & Schubart, H. O. R. (1989) Oribatideos (Acari: Oribatida) colonizadores de folhas em decomposicao de tres sitios florestais da Amazonia Central. Boletim do Museu Paraense Emilio Goeldi. Serie Zoologia, 5 (2), 243 - 276.","Adis, J., Bonaldo, A. B., Brescovit, A. D., Bertani, R., Cokendolpher, J. C., Conde, B., Kury, A. B., Lourenco, W. R., Mahnert, V., Pinto-da-Rocha, R., Platnick, N. I., Reddell, J. R., Rheims, C. A., Rocha, L. S., Rowland, J. M., Weygoldt, P. & Woas, S. (2002) Arachnida at ' Reserva Ducke', Central Amazonia / Brazil. Amazoniana, 17 (1 - 2) 1 - 14.","Franklin, E., Santos, E. M. & Albuquerque, M. I. (2006) Diversity and distribution of oribatid mites (Acari: Oribatida) in a lowland rain forest in Peru and in several environments of the Brazilians states of Amazonas, Rondonia, Roraima and Para. Brazilian Journal of Biology, 66 (4), 999 - 1020."]}

Oribatella (Oribatella) serrula Pérez-Iñigo & Baggio, 1985 Oribatella serrula Pérez-Iñigo & Baggio, 1985 Reported only from Brazil. Brazil: States of Minas Gerais, Pará and São Paulo. References: Pérez-Iñigo & Baggio (1985): 193, Balogh & Balogh (1990): 126, Pérez-Iñigo & Baggio (1996): 70, Ferreira et al. (2012):14.

Oribatella nigra Kulijev, 1967 Distribution in Georgia. W: Ritsa Reserve, Musera, Beshumi, Kolkheti National Park, Kintrishi Reserve, Mtirala National Park, Sataplia Reserve, Itkhvisi, Sairme, Tsaghveri; E: Kojori, Norio, Lagodekhi Reserve (Mumladze et al. 2015; Murvanidze & Arabuli 2015; Murvanidze et al. 2015; Shtanchaeva & Sub��as 2010) Global distribution. Caucasus Ecology. Humid forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 62, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella ornata (Coggi, 1900) Distribution in Georgia. W: Mtirala National Park (Murvanidze et al. 2015) Global distribution. Palaearctic Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 62, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella tenuis Csiszar, 1961 Distribution in Georgia. W: Mtirala National Park, Sairme (Mumladze et al. 2015; Murvanidze et al. 2015) Global distribution. Mediterranean Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 62, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella foliata Krivolutsky, 1974 Distribution in Georgia. W: Ritsa Reserve, Sakeni, Bzyb (Shtanchaeva & Sub��as 2010) Global distribution. Palaearctic Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 61, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella superbula (Berlese, 1904) Syn.: Oribatella meridionalis Berlese, 1908 sensu Djaparidze 1974; Karppinen et al. 1987; Murvanidze & Darejanashvili 2000 Distribution in Georgia. E: Tbilisi, Algethy Reserve (Shtanchaeva & Subías 2010) Global distribution. Palaearctic Ecology. Forest soils Remark. In the Catalogue (Shtanchaeva & Subías 2010) the presence of O. meridionalis Berlese, 1908 is registered for Tbilisi. This species is known as a junior synonym of O. superbula (Weigmann 2001)., Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 62, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella berlesei (Michael, 1898) Distribution in Georgia. whole country (Murvanidze et al. 2013; Murvanidze 2014; Murvanidze & Mumladze 2014; Murvanidze & Arabuli 2015; Shtanchaeva & Subías 2010) Global distribution. Palaearctic Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 61, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella colchica Krivolutsky, 1974 Distribution in Georgia. W: Banguriani, Kintrishi Reserve, Mtirala National Park, Tskaltubo, Itkhvisi, Sairme, Kvabiskhevi Reserve; E: Tbilisi (Murvanidze & Mumladze 2014; Murvanidze & Arabuli 2015; Murvanidze et al. 2015; Shtanchaeva & Sub��as 2010) Global distribution. Mediterranean Ecology. Wet to humid forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 61, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella calcarata (C.L. Koch, 1835) Distribution in Georgia. (Shtanchaeva & Subías 2010) Global distribution. Holarctic Ecology. Forest soils and canopy, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 61, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Oribatella heterodentata Karppinen & Shtanchaeva, 1987 Distribution in Georgia. W: Sakeni (Shtanchaeva & Subías 2010) Global distribution. Caucasus Ecology. Forest soils

Oribatella bulanovae Kulijev, 1967 Distribution in Georgia. W: Musera, Batumi Botanical Garden (Shtanchaeva & Sub��as 2010) Global distribution. Europe Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 61, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

A new subgenus, Sacculoribatella subgen. n., of Oribatella and 3 new species the families Hermanniellidae (Hermanniella aliverdievae sp. n.), Oribatellidae (Oribatella caspica sp. n.), and Scheloribatidae (Pachygena makarovae sp. n.) are described from the Caucasus.

The paper summarizes the data on oribatid mites of the family Oribatellidae Jacot, 1925 from the Caucasus. The redescriptions of the genus Ferolocella Grabovsky, 1971 = Gendzella Kuliev, 1977 (Subias, 2004), the subgenus Multoribatella, the endemic species Ferolocella cribraria (Kuliev, 1977), and the new species Oribatella abdurachmanovi from mountain Daghestan are given. Two species are synonymized (Ferolocella cribraria Kuliev, 1977 = Oribatella sitnikovae Dzaparidze, 1989, syn. n. and Oribatella foliata Krivolutsky, 1974 = Oribatella brevipila Bernini, 1977, syn. n.). The species Oribatella bulanovae Kuliev, 1962; O. reticulata Berlese, 1916; O. asiatica Krivolutsky, 1974; O. foliata, O. (M.) colchica Krivolutsky, 1974; and O. (M.) nigra Kuliev, 1967 are redescribed. Original illustrations of O. heterodentata Karppinen et Shtanchaeva, 1987; O. krivolutskyi Karppinen et Shtanchaeva, 1987; O. sexdentata Berlese, 1916; O. exilicornis Berlese, 1910; O. cf. calcarata (C.L. Koch, 1836) and O. cf. foliata, demonstrating some differences from the nominotypical species, are given. The diagnoses, differential diagnoses, and keys to the Caucasian species of Joelia Oudemans, 1906 and Oribatella Banks, 1895 are presented. The distribution of Oribatellidae species in the territory of the Caucasus is shown.

Oribatella sp. Iran localities. Behshahr (Akrami et al. 2008). Rasht (Mortazavi 2009). Shiraz (Behmanesh & Akrami 2012 a).

Oribatella vicinus (Ryabinin, 1975) Fenestrobates vicinus Ryabinin, 1975 p. 540 Material examined. Holotype male, kindly loaned from the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia. Diagnosis. Total length of adult male about 576. Integument of prodorsum, notogaster, venter and mentum micropunctate. Integument of lenticulus and coxisternum with distinct striae, as in Oribatella jacoti Behan- Pelletier 2011. Rostrum medially rounded with pair of small lateral teeth on margin, as in O. brevicornuta Jacot, 1934 (see Fig. 2D, E, F in Behan-Pelletier 2011). Lamella (with cusp) about 162 long, cusp about 108 long, 62 wide at level of insertion of seta le. Lamellar cusps diverging anteromedially; separated posteromedially, leaving prodorsum visible through oval opening about 14 at widest. Medial dens of lamellar cusps about 52 – 55 µm, subequal to length lateral dens; medial dens smooth; lateral dens with about three small teeth on lateral edge. Long striae extending longitudinally along outer margin of lamella. Translamella without tooth, about 14 at greatest width, about 26 deep. Setae ro strongly barbed along length, acuminate, strongly directed medially. Setae le about 79 long, thick, heavily barbed, tapered. Setae in 172 long, thick (less so than le), heavily barbed, tapered; mutual distance of pair about 81 µm. Sensillus 141 long, subequal in width to seta le, except head slightly fusiform, tapered, stalk with small barbs distally; head directed slightly anteromedially, as in O. brevicornuta (see Fig. 3, insert in Behan-Pelletier 2011). Exobothridial seta not visible. Genal tooth broad, subtriangular, rounded distally, without dens ventrodistally, with longitudinal ridge ventrally. Tutorium including cusp about 168 long, rectangular, distal margin with eight to 10 dentes. Ventral depression between pedotectum I and the lateral body wall less defined than that in O. brevicornuta. Custodium about 58 long, extending to anterior margin of pedotectum II. Anterior margin of notogaster undulating, convex lateral of bothridium, with about 10 transverse ridges. Pteromorph with dens anteroventrally. Four pairs of porose areas present, Aa circular about 12 at widest. Ten pairs of long, barbed notogastral setae present; setae barbed, acuminate, almost flagellate 52 – 79 long (Fig. 4 of Ryabinin 1975). Setae h 1 directed posteriorly and parallel as in O. jacoti (Behan-Pelletier, 2011). Distance h 1 –h 1 and p 1 –p 1 subequal, about 23 – 26. Lenticulus subtriangular. Epimeral setation 3(?) – 1 – 3 – 3; barbed, tapered, length not established except 3b, about 72, 4a, 4b about 29 – 36, long, spinous. Setae 3b, 4a, 4b, strongly directed anteriorly (Fig. 5 of Ryabinin 1975). Genital setae 6 pairs, two seta on anterior of genital plate. Mentum with anterior tectum covering base of gena, reflexed anteriorly, similar in size and shape to that for Oribatella yukonensis Behan- Pelletier and Walter, 2012 (their Fig. 14G). Seta l” of tibia I about 53. Tarsi heterotridactylous. Remarks. The description of this species by Ryabinin (1975) is brief and lacks measurements. We redescribe this species based on the slide mounted holotype male. This is a dorsal mount which precludes evaluation of ventral and lateral characters. Ryabinin (1975) assigned this species to Fenestrobates on the basis of its similarity to O. rossicus. He stated “The new species is similar to F. rossicus from Primorye territory (Krivolutsky 1974)”. The species are similar but can be easily separated by the long, acuminate notogastral seta in O. vicinus vs. the short notogastral setae of O. rossicus; the very long epimeral seta 3b (96) in O. rossicus vs. shorter seta (72) in O. vicinus; setae h 1 directed posteriorly and parallel in O. vicinus vs. curved h 1 in O. rossicus. We were unable to verify whether O. rossicus had 11 pairs of notogastral setae vs. the 10 pairs found in O. vicinus., Published as part of Ermilov, Sergey G. & Behan-Pelletier, Valerie M., 2014, Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae, pp. 258-272 in Zootaxa 3827 (2) on pages 267-268, DOI: 10.11646/zootaxa.3827.2.7, http://zenodo.org/record/4920524, {"references":["Ryabinin, N. A. (1975) New and little known species of oribatid mites from the Khabarovsk Territory and the Amur Region. Zoologichesky Zhurnal, 54 (4), 533 - 542. [in Russian]","Jacot, A. P. (1934) A new four horned mossmite (Oribatoidea-Acarina). American Midland Naturalist, 15 (1), 706 - 712.","Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56.","Krivolutsky, D. A. (1974) New oribatid mites of the USSR. Zoologichesky Zhurnal, 53 (12), 1880 - 1885. [in Russian]"]}

Oribatella rossicus (Krivolutsky, 1974) Fenestrobates rossicus Krivolutsky, 1974 p. 1883 Material examined. Cotypes (male, female, on single slide) kindly loaned from the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia. Diagnosis. Total length of adult 630 (measurement from Krivolutsky 1974). Integument of prodorsum, notogaster, venter and mentum micropunctate. Integument of coxisternum with distinct striae. Rostrum medially rounded with pair of small lateral teeth on margin, as in Oribatella brevicornuta Jacot, 1934 (see Fig. 2D, E, F in Behan-Pelletier 2011). Lamella (with cusp) about 191 long, cusp about 128 long, 63 wide at level of insertion of seta le. Lamellar cusps diverging anteromedially; separated posteromedially, leaving prodorsum visible through oval opening × about 12 at widest. Medial dens of lamellar cusps about 61, subequal or slightly shorter than lateral dens (68 long); medial dens smooth; lateral dens with about 3–4 small teeth on lateral edge (varying on either side of type female specimen). Long striae extending longitudinally along outer margin of lamella. Translamella without tooth, about 12 at greatest width, about 22 deep. Seta ro about 132 long, strongly barbed along length, acuminate, directed anteromedially. Seta le about 110 long, thick, heavily barbed, tapered. Setae in 213 long, thick (less so than le), heavily barbed, tapered; mutual distance of pair about 104 µm. Sensillus 127 long, subequal in width to seta le, except head slightly fusiform, stalk with small barbs distally; head directed slightly anteromedially. Exobothridial setae not visible. Genal tooth broad, subtriangular, rounded distally, without dens ventrodistally, with longitudinal ridge ventrally. Tutorium including cusp about 235 long, rectangular, distal margin with about eight dens. Ventral depression between pedotectum I and the lateral body wall similar to that in O. brevicornuta. Custodium about 56 long, Anterior margin of notogaster undulating, convex lateral of bothridium, with about 10 transverse ridges. Pteromorph with dens anteroventrally. Four pairs of porose areas present, Aa oval about 34 at widest. Eleven pairs of long, barbed notogastral setae present (data from Krivolutsky 1974); setae barbed, tapered, 14–30 long. Distance h 1 –h 1 and p 1 –p 1 could not be established. Lenticulus subtriangular. Epimeral setation 3(?)–1–3–3; about 12 to 48, barbed, tapered, except 4a, 4b, 4c, 3b, 3c long, spinous; 3b about 96, reaching anteriorly 3c about 80, 4a, 4b about 77, 4c about 48. Setae 3b, 4a, 4b, 4c strongly directed anteriorly to level of indentation of pedotectum I, seta 3b in longitudinal alignment with long spinous seta l’ of tibia IV when legs folded. Genital setae six pairs, two seta on anterior of genital plate; genital and aggenital setae about 11 long. Mentum with anterior tectum covering base of gena, reflexed anteriorly, similar in size and shape to that for Oribatella yukonensis Behan-Pelletier and Walter, 2012 (their Fig. 14G). Seta l” of tibia I about 54, seta l’ of tibia IV about 54. Tarsi heterotridactylous. Remarks. The description of this species by Krivolutsky (1974) is brief and lacks measurements other than for body length. The expanded description above is based on the slide mounted type specimens. These are mounted dorsal and ventral and thus some characters could not be re-evaluated. For example, Krivolutsky (1974) noted “Trochanters III, IV with long, thin, finger-like structure, directed anteriorly”; we were unable to evaluate this further. Krivolutsky (1974) assigned this species to Fenestrobates on the basis of “Pore-like structure on coxes of legs II rounded, clearly visible in ventral view”. This is referring to the ventral depression between pedotectum I and the lateral body wall as seen through the coxisternal integument. However, we do not consider this depression to be as extensive as in the species of Fenestrobates, or to be flask-shaped. This species shares the long spinous setae 3b with species of Fenestrobates, but seta 4c is present in O. rossicus, whereas this seta is absent in species of Fenestrobates. Furthermore O. rossicus has about 10 ridges on the anterior margin of the notogaster, ridges that are present in all described species of Oribatella, but absent in Fenestrobates., Published as part of Ermilov, Sergey G. & Behan-Pelletier, Valerie M., 2014, Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae, pp. 258-272 in Zootaxa 3827 (2) on page 267, DOI: 10.11646/zootaxa.3827.2.7, http://zenodo.org/record/4920524, {"references":["Krivolutsky, D. A. (1974) New oribatid mites of the USSR. Zoologichesky Zhurnal, 53 (12), 1880 - 1885. [in Russian]","Jacot, A. P. (1934) A new four horned mossmite (Oribatoidea-Acarina). American Midland Naturalist, 15 (1), 706 - 712.","Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56.","Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of western North America. Zootaxa, 3432, 1 - 62."]}

Oribatella flagellata Behan-Pelletier, 2011: 13 — Holotype male (CNC, No. 23954), Trametes versicolor (L.) polypore fungi and lichen on hardwood, 6km W Troutdale, Hwy 603 at Fox Creek, Grayson Co., Virginia, USA., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 77, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56."]}

Oribatella oregonensis Behan-Pelletier & Walter, 2012: 31 — Holotype female (CNC), Oregon, Tillamook Co., Cliff Creek Trail, USA., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 78, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of western North America. Zootaxa, 3432, 1 - 62."]}

Oribatella sintranslamella Behan-Pelletier & Walter, 2012: 45 — Holotype female (CNC), from deep, damp, mixed maple-hemlock litter, rotten wood, Oregon, Deschutes Co., Proxy Falls, St. Hwy 242, 28 mi SW Sisters, ca. 3000’, USA.

Oribatella heatherae Behan-Pelletier & Walter, 2012: 19 — Holotype female (CNC), from moss and lichens among rocks on alpine scree slope, Alberta, Kananaskis Country, Fortress Mt., Canada., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 78, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of western North America. Zootaxa, 3432, 1 - 62."]}

Oribatella abmi Behan-Pelletier & Walter, 2012: 5 — Holotype female (CNC), from upper organic soil, layer in open field with grasses and sedges, Alberta, ABMI Site 1438 (51.10178757, -111.487175), 8 km east of Pollockville along Township Road 250, Canada.

Oribatella ewingi Behan-Pelletier & Walter, 2012: 16 — Holotype female (CNC), from redwood litter, California, Matea Co., 6 mi SE Halfmoon Bay, USA.

Oribatella maryae Behan-Pelletier & Walter, 2012: 27 — Holotype male (CNC), from litter of Lawson’s Cypress and Tan-oak, Oregon, Curry Co., Siskiyou National Forest, Shrader Old Growth Trail, 10 mi on Jerry’s Flat Road to Agness, USA (42°29'28''N 124°17'04''W)., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 78, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of western North America. Zootaxa, 3432, 1 - 62."]}

Oribatella banksi Behan-Pelletier & Walter, 2012: 11 — Holotype female (CNC), from Quercus durata, California, Tehama Co., 4mi SW Platina Beegum Creek, 2250’, USA.

Oribatella yukonensis Behan-Pelletier & Walter, 2012: 48 — Holotype female (CNC), from semi-stabilized dunes with grass, sedge, Polemonium, Stellaria, Yukon, Carcross Dunes, Canada., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 78, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of western North America. Zootaxa, 3432, 1 - 62."]}

Oribatella texana Behan-Pelletier, 2011: 47 — Holotype female (CNC, No. 23958), palmetto and oak litter, Guadeloupe State Park, Kendall Co., Texas, USA (29'52.443N 98'29.139W)., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 77, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56."]}

Oribatella pawnee Behan-Pelletier & Walter, 2012: 37 — Holotype male (CNC), Colorado, Weld Co., Central Plains Experimental Range, Rd. 37, USA., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 78, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. & Walter, D. E. (2012) Oribatella (Acari, Oribatida, Oribatellidae) of western North America. Zootaxa, 3432, 1 - 62."]}

Oribatella jacoti Behan-Pelletier, 2011: 16 — Holotype female (FM, FMHD #85–128), red and chestnut oak, sugar maple litter on ridge top, Mo. Campus, Hinkson Creek S Univ, Boone Co., Missouri, USA., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 77, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56."]}

Oribatella metzi Behan-Pelletier, 2011: 21 — Holotype female (CNC, No. 23956.), Duke Forest, Durham Co., North Carolina, USA., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 77, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56."]}

Oribatella nortoni Behan-Pelletier, 2011: 30 — Holotype female (CNC, No. 23957), Auburn in coastal plain, nr., Lee Co., Alabama, USA., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 77, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Behan-Pelletier, V. M. (2011) Oribatella (Acari, Oribatida, Oribatellidae) of eastern North America. Zootaxa, 2973, 1 - 56."]}