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Mantelliana speculum sp. nov. Fig. 4A–C 2022 Mantelliana sp. —Araripe et al., p. 7, fig. 5 (9a–b). Etymology: Speculum (lat.) due to the symmetry between the front and back ends that are reflected like a mirror. Holotype: Adult shell, Nº. DGEO-CTG-UFPE-1514 (Fig. 4A), Illustrated material from the Santo Antônio section 0.3 m. Paratypes: Adult shell, Nº. DGEO-CTG-UFPE-1515 (Fig. 4B) and Nº. DGEO-CTG-UFPE-1516 (Fig. 4C), Illustrated material from the Santo Antônio section 3.1 and 2.65 m respectively. Type locality: Brazil, State of Pernambuco, Municipality of Exu, Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type-horizon: Santo Antônio section, Araripe Basin. Calcarenite (F-Sl) corresponding to the Romualdo Formation, depth 0.30 m, dated as upper Aptian. Dimension. Nº. DGEO-CTG-UFPE-1514 (Fig. 4B); L= (LV) 0.870 / (RV) 0.976 mm, H= 0.623 and W= 0.443 mm. Diagnosis: Carapace large, triangular, inflated and perfectly symmetrical between both ends. Description: Carapace large, triangular outline in lateral view. Ventral margin straight. Dorsal margin with a well-marked median angularity that coincides with the greatest height. Perfectly rounded margin. Maximum length in VM. Greatest height in the median region coinciding with the location of the cardinal angle. LV larger than VR with characteristic overlaps in the ventral, posterodorsal and anterodorsal regions. In dorsal view, greatest width in the median region. Smooth surface. Observations. Mantelliana speculum sp. nov. was compared with the female specimen of Mantelliana wealdensis (Jones, 1888) described and illustrated by Anderson, (1985). However, the well-marked angle in the dorsal region described by Anderson (1985) is not centralized, thus not dividing the valves into two practically identical parts as in the new species. Occurrence and associated fauna. It was recorded in the Santo Antonio section, at the following depths (from base to top): 0.3; 2.25; 3.1; 3.55; 3.65; 3.85; 4.05; 5.25; 5.35; 5.55 and 5.65 m. Mantelliana speculum sp. nov. occurs associated with all species of ostracods. Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on pages 543-544, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092

Damonella grandiensis Tomé et al., 2014 Fig. 3P–R 1992 Mantelliana (?) sp.—Colin et al., p. 368, plate 3, fig. 1. 1999 Ostracode 207—Neumann, p. 93, figs. 2–4. 2002 Ostracode 207— Coimbra et al., p 691, plate IV, fig. 31. 2006 Candona sp —Ramos et al., p. 344, fig. 4. 2010 Damonella sp. — Tomé & Lima Filho, p. 63, plate. 2 (a–c). 2013 Candona ? sp.—Do Carmo et al., p. 95 and 103, fig. 5, nº.12–14. 2014 Damonella grandiensis —Tomé et al., p. 163, fig.10 A–D. Non 2014 Damonella grandiensis —Tome et al., p. 163, fig. 10 (D–F) and 167, fig. 10 (G–M). Non 2015 Damonella aff. grandiensis —Trabelsi et al., p. 10, fig. 5D, 1–3. 2017 Damonella grandiensis —Souza et al., p. 9, fig. 1, G–I. Non 2021 Damonella grandiensis —Araripe et al., p. 6, fig. 5 (M–P). 2022 Damonella grandiensis —Araripe et al., p. 7, fig. 5 (10a–b). Illustrated material. Adult shell, Nº. DGEO-CTG-UFPE-1513. Illustrated material from the Santo Antonio section (2.65 m). Dimension. Nº. DGEO-CTG-UFPE-1513 (Fig. 3 P–R); L= (LV) 0.576 / (RV) 0.611 mm, H= 0.325 mm, and W= 0.245 mm. Comparison. Material compared to D. grandiensis (Nº DGEO-CTG-UFPE-1157 and Nº-DGEO-CTG-UFPE- 1158) described and deposited by Tomé et al. (2014) in the Ivan de Medeiros Tinoco Microfossils Collection. The comparison was satisfactory, with the exception of the specimen Nº. DGEO-CTG-UFPE-1157, since they presented identical outlines and posterodorsal and ventral overlapping. Occurrence and associated fauna. It was recorded in the Santo Antonio section, in all depths and associated with all species. It is the only species that occurs associated with the nannofossils Thoracosphaera sp. and Biscutum sp. (Araripe at al. 2022). Observations. Tomé et al. (2014) emphasize the sexual dimorphism of this species and describe the male form, however in the present study these forms are described as distinct species, mainly due to the different outlines, since both specimens are smooth. Stratigraphic distribution. In Africa, it has been commonly recorded in the d’Hama-Koussou Basin, Lower Cretaceous (Aptian) (Colin et al. 1992). In Brazil, it has been recorded in the Araripe Basin, Crato Formation (Neumann 1999), Aptian; Grajaú Basin, Codó Formation, lower Aptian (Ramos et al. 2006); Cedro Basin, Crato Formation, upper Aptian (Tomé 2007; Tome & Lima Filho 2010); Potiguar Basin, Alagamar Formation, Aptian (Do Carmo et al. 2013). In the Sergipe / Alagoas Basin, Krommelbein & Weber (1971) observed the occurrence of this species and called it Ostracode 207 (Nº SMF Xe-22592), which is stored in slide PIA-3-AL (obtained from a depth of 900 to 930 m) in Frankfurt (Tomé et al. 2014). Additionally found in the Jatobá Basin, Serra Negra Formation, in the strata correlated to the Crato Formation, upper Aptian (Tomé et al. 2014), and this study in the Araripe Basin, Romualdo Formation, upper Aptian and lower Albian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on pages 542-543, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Tome, M. E. T. R., Lima Filho, M. F. & Neumann, V. H. M. L. (2014) Taxonomic studies of non-marine ostracods in the Lower Cretaceous (Aptian-lower Albian) of post-rift sequence from Jatoba and Araripe basins (Northeast Brazil): Stratigraphic implications. Cretaceous Research, 48, 153 - 176. https: // doi. org / 10.1016 / j. cretres. 2013.12.007","Colin, J. P., Brunet, M., Congleton, J. D., Dejax, J., Flynn, L. J., Hell, J. & Jacobs, L. (1992) Ostracodes lacustre des bassins d'age Cretace Inferieur du Nord Cameroun: Hama-Koussou, Koum et Babouri-figuil. Revue de Paleobiologie, 11 (2), 357 - 372.","Ramos, M. I. F., Rossetti, D. D. F. & Paz, J. S. (2006) Caracterizacao e significado paleoambiental da fauna de ostracodes da Formacao Codo (Neoaptiano), leste da Bacia de Grajau, MA, Brasil. Revista Brasileira de Paleontologia, 9, 339 - 348. https: // doi. org / 10.4072 / rbp. 2006.3.09","Tome, M. E. T. R. & Lima Filho, M. F. (2010) Ostracodes da Bacia de Cedro, Andar Alagoas, Estado de Pernambuco, NE, Brasil: Implicacoes paleoambientais e bioestratigraficas. Estudos Geologicos, 21, 56 - 70.","Do Carmo, D. A., Coimbra, J. C., Whatley, R. C., Antonietto, L. S. & Citon, R. T. P. (2013) Taxonomy of limnic Ostracoda (Crustacea) from the Alagamar Formation, middle-upper Aptian, Potiguar Basin, Northeastern Brazil. Journal of Paleontology, 87, 91 - 104. https: // doi. org / 10.1666 / 11 - 108 R. 1","Krommelbein, K. & Weber, R. (1971) Ostracoden des \" Nordost-Brasilianischen Wealden \". Geologisches Jahrbuch, 115, 1 - 93."]}

Perissocytheridea oculusutilis sp. nov. Fig. 4G–K 2022 Perissocytheridea sp.1 —Araripe et al., p 7, fig. 5 (4a–b). Etymology: Oculus (lat.) and sutilis (lat.) in reference to the presence of a subtle ocular tubercle. Holotype: Adult carapace, Nº. DGEO-CTG-UFPE-1519 (Fig. 4G–K). Type locality: Brazil, State of Pernambuco, Municipality of Exu, Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type-horizon: Santo Antônio section, Araripe Basin. Calcarenite (F-Sl) corresponds to Romualdo Formation, depth 3.10 m, dated as upper Aptian interval. Dimension. Nº. DGEO-CTG-UFPE-1519 (Fig. 4G–K); L= (LV) 0.358 / (RV) 0.349 mm, H= 0.194 mm, and W= 0.174 mm. Diagnosis: Carapace very small, with a subtle ocular tubercle and a ventral projection with a triaxial cap in the posteroventral region. Description: Carapace very small. Sub-rectangular outline in the lateral view. Ventral margin straight on both valves. Dorsal margin straight and slightly arched, descending to the posterior end. Anterior margin sub-rounded, starting dorsally straight and rounding to ventral margin. Sub-rectangular posterior margin tapering down to the caudal process, which coincides with the greatest length. Greatest height at the anterior portion. The left valve slightly overlaps the right valve in the dorsal margin. In the dorsal view, the ends are sharp. Surface ornamented with primary reticulation and secondary punctuations inside of the reticles. Discrete winged processes in the posteroventral margins, with a smooth triaxial cap, which coincides with the greatest width. It has a very smooth lateral groove. Ocular tubercle evident. Observations. The main differences between the species described in the present work are the greatest evidence of both secondary ornamentation in punctuation and the ocular tubercle. The triaxial structure on the posteroventral margin, which in the dorsal view appears as two lateral “pins”. It is worth emphasizing the need for a study covering a larger population of the genus Perissocytheridea found in this study. Occurrence and associated fauna. It was recorded in the Santo Antônio section, at the following depths (from base to top): 0.3; 2.25; 2.65; 3.1; 4.05; 3.55 and 5.45 m. Perissocytheridea oculusutilis sp. nov. does not occur with Dicrorygma (Orthorygma ?) dimorpha. Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 545, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092

Pattersoncypris salitrensis (Krömmelbein & Weber, 1971) Fig. 3A–C 1971 Hourcqia angulata salitrensis —Krommelbein & Weber, p. 22 and 50, plate 6. 1972 Pattersoncypris angulata salitrensis —Bate, p. 389, text-fig. 11 (c, d, f). 2006 Harbinia salitrensis —Ramos et al., p. 344, fig. 4Q, non 4R, 4S and 4T. 2008 Harbinia salitrensis —Do Carmo et al., p. 795, fig.6, nº 8. 2012 Harbinia salitrensis —Antonietto et al., p. 662, fig.4, nº 1-10. 2012a Pattersoncypris salitrensis —Poropat and Colin, p.708, fig. 4, nº 3. 2014 Pattersoncypris salitrensis —Tomé et al., p.167, fig. 10 Q–S and 14A, C. 2021 Pattersoncypris salitrensis — Araripe et al., p. 6, fig.5 (A–C). 2022 Pattersoncypris salitrensis —Araripe et al., p. 7, Fig. 5 (13a–b) Illustrated material. Adult shell, Nº. DGEO-CTG-UFPE-1505 (Fig. 3A and B) and Nº. DGEO-CTG-UFPE-1506 (Fig. 3C). Illustrated material from the Santo Antônio section sample at 2.65 and 3.55 m respectively. Dimension. Nº. DGEO-CTG-UFPE-1505 (Fig. 3A and B); L= (LV) 1.045 / (RV) 1.373 mm, H= 0.69 mm, and W= 0.744 mm and Nº. DGEO-CTG-UFPE-1506 (Fig. 3C); L= (LV) 1.045 / (RV) 1.373 mm. Comparison. Material compared to Pattersoncypris salitrensis (Nº DGEO-CTG-UFPE-1971 and Nº DGEO- CTG-UFPE-1166), described and deposited by Tomé et al. (2014) in the Ivan de Medeiros Tinoco Microfossils Collection of the Paleontology Laboratory of the Federal University of Pernambuco. Occurrence and associated fauna. It was recorded in the Santo Antonio section, at the following depths (from base to top): 0.3; 1.8; 2.1; 2.25; 2.65; 3.4; 3.65; 3.85; 4.05 and 4.25 m. P. salitrensis is associated with all species of ostracods, except Dicrorygma (Orthorygma ?) dimorpha Weiss and Theriosynoecum cf. quadrinodosum Krömmelbein & Weber. Observations. The specimens recovered were compared to the illustrations of Pattersoncypris cretapa Krömmelbein & Weber, published in Do Carmo et al. (2008). Some similarities were seen, but a comparison between both specimens is necessary for a future synonymy of these species. The Santo Antônio site is one of the few sections that marks the boundary between the Aptian and lower Albian in the Romualdo Formation. It is worth mentioning that this species was not recorded in the lower Albian at this locality (Araripe et al. 2022). Stratigraphic distribution. In Brazil, distributed in the Araripe Basin, Santana Group (above the layers with fish occurrence), Aptian (Krommelbein & Weber 1971; Bate 1999; Poropat & Colin 2012a) and Romualdo Formation, lower Albian (Tomé et al. 2014; Araripe et al. 2021); Grajaú Basin, Codó Formation, upper Aptian (Ramos et al. 2006); Potiguar Basin, Alagamar Formation, Aptian (Do Carmo et al. 2008). In the present study, found in the Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on pages 538-539, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Krommelbein, K. & Weber, R. (1971) Ostracoden des \" Nordost-Brasilianischen Wealden \". Geologisches Jahrbuch, 115, 1 - 93.","Tome, M. E. T. R., Lima Filho, M. F. & Neumann, V. H. M. L. (2014) Taxonomic studies of non-marine ostracods in the Lower Cretaceous (Aptian-lower Albian) of post-rift sequence from Jatoba and Araripe basins (Northeast Brazil): Stratigraphic implications. Cretaceous Research, 48, 153 - 176. https: // doi. org / 10.1016 / j. cretres. 2013.12.007","Do Carmo, D. A., Whatley, R., Queiroz Neto, J. V. Q. & Coimbra, J. C. (2008) On the validity of two Lower Cretaceous non-marine ostracode genera: biostratigraphic and paleogeographic implications. Journal of Paleontology, 82, 790 - 799. https: // doi. org / 10.1666 / 07 - 008.1","Araripe, R., Pedrosa Lemos, F. A., do Prado, L. A. C., Tome, M. E. T. R., Oliveira, D. H. D., Pereira, P. A., Nascimento, L. R. S. L., Asakura, Y., Ng, C., Viviers, M. C. & Barreto, A. F. (2022) Upper Aptian - lower Albian of the southern-central Araripe Basin, Brazil: Microbiostratigraphic and paleoecological inferences. Journal of South American Earth Sciences, 116, 103814. https: // doi. org / 10.1016 / j. jsames. 2022.103814","Bate, R. H. (1999) Non-marine ostracod assemblages of the Pre-Salt rift basins of West Africa and their role in sequence stratigraphy. In: Cameron, N. R., Bate, R. H. & Clure, V. S. (Eds.), The Oil and Gas Habitats of the South Atlantic. Geological Society of London Special Publication, 153, pp. 283 - 292. https: // doi. org / 10.1144 / GSL. SP. 1999.153.01.17","Poropat, S. F. & Colin, J. P. (2012 a) Early Cretaceous ostracod biostratigraphy of eastern Brazil and western Africa: An overview. Gondwana Research, 22, 772 - 798. https: // doi. org / 10.1016 / j. gr. 2012.06.002","Araripe, R., Oliveira, D., Tome, M. E., Mello, R. M. & Barreto, A. M. F. (2021) Foraminifera and Ostracoda from the lower Cretaceous (Aptian - lower Albian) Romualdo formation, Araripe Basin, northeast Brazil: Paleoenvironmental inferences. Cretaceous Research, 122, 104766. https: // doi. org / 10.1016 / j. cretres. 2021.104766","Ramos, M. I. F., Rossetti, D. D. F. & Paz, J. S. (2006) Caracterizacao e significado paleoambiental da fauna de ostracodes da Formacao Codo (Neoaptiano), leste da Bacia de Grajau, MA, Brasil. Revista Brasileira de Paleontologia, 9, 339 - 348. https: // doi. org / 10.4072 / rbp. 2006.3.09"]}

Genus Mongolianella Mandelstam, in Galeeva, 1955 Type species: Mongolianella palmosa Mandelstam, 1956 Observations. This genus is widely distributed in the basins of India (Khosla & Sahni 2000; Whatley & Bajpai, 2000a, b and c; Bajpai & Whatley 2001; Whatley et al. 2003a, b and c; Khosla & Nagori 2005; Khosla et al. 2009a and Khosla et al. 2011). Khosla et al. (2009a) transferred this genus to Stenocypris Sars, (1889). In the present study, the species found was treated as belonging to the genus Mongolianella, but we emphasize the need for a comparative study with the forms described by Khosla et al. (2011)., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 540, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Galeeva, L. I. (1955) Ostracoda from the Cretaceous deposits of the Peoples Republic of Mongolia: Moscow. Geologo-Razved Neft Institute, 1955, 94.","Mandelstam, M. I. (1956) Material on Palaeontology. New families and genera: Ostracoda. Trudy VSEGEI, New Series, Palaeontology, 12, 87 - 144 + 306 - 323.","Khosla, A. & Sahni, A. (2000) Late Cretaceous (Maastrichtian) ostracodes from the Lameta Formation, Jabalpur Cantonment area, Madhya Pradesh, India. Journal of the Palaeontological Society of India, 45, 57 - 78.","Whatley, R. & Bajpai, S. (2000 a) A new fauna of Late Cretaceous non-marine Ostracoda from the Deccan intertrappean beds of Lakshmipur, Kachchh (Kutch) District, Gujarat, Western India. Revista Espanola de Micropaleontologia, 32, 385 - 409.","Bajpai, S. & Whatley, R. C. (2001) Late Cretaceous non- marine ostracods from the Deccan Intertrappean beds, Kora (Western Kachchh, India). Revista EspanTola de Micropalaeontologiia, 33, 91 - 111.","Whatley, R., Bajpai, S. & Whittaker, J. E. (2003 a) Freshwater Ostracoda from the Upper Cretaceous intertrappean beds at Mamoni (Kota District), southeastern Rajasthan, India. Revista Espanola de Micropaleontologia, 35 (1), 75 - 86.","Khosla, S. C. & Nagori, M. L. (2005) A restudy of ostracode fauna from the Inter-trappean beds of Anjar, Kachchh District, Gujarat. Journal of the Geological Society of India, 66 (5), 573 - 580.","Khosla, S. C., Nagori, M. L., Jakhar, S. R. & Rathore, A. S. (2009 a) Mixed marine, brackish water and non-marine microfaunal association in the Inter-trappean beds (Early Palaeocene) of Jhilmili, Chhindwara District, Madhya Pradesh. Journal of the Geological Society of India, 73 (5), 724 - 732. https: // doi. org / 10.1007 / s 12594 - 009 - 0056 - 2","Khosla, S. C., Nagori, M. L., Jakhar, S. R. & Rathore, A. S. (2011) Early Danian lacustrine - brackish water Ostracoda from the Deccan Intertrappean beds near Jhilmili, Chhindwara District, Madhya Pradesh, India. Micropaleontology, 57 (3), 223 - 245. https: // doi. org / 10.47894 / mpal. 57.3.02"]}

Genus Perissocytheridea Stephenson, 1938 Type species: Perissocytheridea matsoni Stephenson, 1938 Remarks: Specimens attributed to the genus Perissocytheridea found in Cretaceous strata have been reported by several authors (Swain & Brown, 1964; Piovesan et al. 2014a, b; Piovesan et al. 2015; Trabelsi et al. 2021). Nogueira & Ramos (2016), showed the distribution of Perissocytheridea over geological time according to the global record and emphasized the need for studies that prove the relationship between species attributed to the Cretaceous and species found in the present. Specimens analyzed in the present study were attributed to this genus since the morphological characteristics related to the carapace are similar, that is, they have small and fragile carapaces, are subpiriform, with a straight but angled dorsal margin, have a slightly convex ventral margin, a broadly rounded anterior margin, a posterior margin presenting a posterodorsal caudal process, and a reticular surface (with pits and ridges), but are sometimes smooth. The ocular tubercle is small but visible., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 545, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Stephenson, M. B. (1938) Lower Eocene Ostracoda of the genus Cytheridea. Journal of Paleontology, 10 (8), 695 - 703.","Swain, F. M. & Brown, P. M. (1964) Cretaceous Ostracoda from wells in the southeastern United States. Bulletin North Carolina Department of Conservation and Development, 78, 1 - 55. https: // doi. org / 10.5962 / bhl. title. 63514","Piovesan, E. K., Cabral, M. C., Colin, J. - P., Fauth, G. & Bergue, C. T. (2014 a) Ostracodes from the Upper Cretaceous deposits of the Potiguar Basin, Northeastern Brazil: taxonomy, paleoecology and paleobiogeography, part 1: Turonian. Carnets de Geologie / Notebooks on Geology, 14, 211 - 252. https: // doi. org / 10.4267 / 2042 / 54003","Piovesan, K. E., Cabral, M. C., Boavida., E. A. & Fauth, G. (2015) Fossocytheride a Swain and Brown and Perissocytheridea Stephenson (Ostracoda): insights into paleosalinity gradients of Late Cretaceous deposits from Brazil and Portugal. Revista Brasileira de Paleontologia, 18 (1), 21 - 30. https: // doi. org / 10.4072 / rbp. 2015.1.02","Trabelsi, K., Sames, B., Nasri, A., Piovesan, E. K., Elferhi F., Skanji, A., Houla, Y., Soussi, M. & Wagreich, M. (2021) Ostracods as proxies for marginal marine to non-marine intervals in the mid-Cretaceous carbonate platform of the Central Tunisian Atlas (North Africa): Response to major short-term sea-level falls. Cretaceous Research, 117, 1 - 32. https: // doi. org / 10.1016 / j. cretres. 2020.104581","Nogueira, A. A. E. & Ramos, M. I. F. (2016) The genus Perissocytheridea Stephenson, 1938 (Crustacea: Ostracoda) and evidence of brackish water facies along the Oligo-Miocene, Pirabas Formation, eastern Amazonia, Brazil. Journal of South American Earth Sciences, 65, 101 - 121. https: // doi. org / 10.1016 / j. jsames. 2015.11.007"]}

Theriosynoecum cf. quadrinodosum Krömmelbein & Weber, 1971 Fig. 5P–R 1978a Bisulcocypris quadrinodosa —Silva: 1018–20; est. 1, figs. 9–12 2002 Theriosynoecum quadrinodosum — Coimbra et al., Fig. 4:5–6. 2020 Theriosynoecum quadrinodosum — Guzmán-González et al., Fig. 5 (b). 2022 Theriosynoecum quadrinodosum — Araripe et al., p 7, fig. 5 (11a–b). Illustrated material. Juvenile shell, Nº. DGEO-CTG-UFPE-1528. Illustrated material from the Santo Antônio section (3.1 m). Dimension. Nº. DGEO-CTG-UFPE-1528 (Fig. 5P–R); L= (LV) 0.603/ (RV) 0.688 mm, W= 0.365 mm and H= 0.352 mm. Comparison. Material compared to Theriosynoecum silvai (Silva, 1978a) (Nº DGEO-CTG-UFPE-1970 e DGEO-CTG-UFPE-1166), described and deposited by Tomé et al. (2014) in the Ivan de Medeiros Tinoco Microfossils Collection of the Paleontology Laboratory of the Federal University of Pernambuco. It was observed that the specimen found in the present study corresponds to a juvenile specimen and it has nodules in the posterior margin that differentiate it from the species deposited in the mentioned collection. Occurrence and associated fauna. It was recorded in the Santo Antonio section, just in the depths (from base to top): 0.3 m. T. cf. quadrinodosum is associated with the species of ostracods: P. angulata, P. micropapillosa, D. grandiensis, D. cf. minuta, Mantelliana speculum sp. nov. and Mongolianella aptianensis sp. nov. Observations. So far four species belonging to the genus Theriosynoecum Branson (1936) have been recorded in the Aptian/Albian: Theriosynoecum silvai (Silva, 1978a), Theriosynoecum munizi (Silva, 1978a), Theriosynoecum colini Do Carmo et al., 2013 and Theriosynoecum guzzoia Do Carmo et al., 2013. Regarding the first two, Tomé et al. (2014) placed T. munizi in synonymy as being a young form of T. silvai. The specimen observed in the present manuscript does not resemble the forms found for this interval. However, it corresponds to a young male of Theriosynoecum quadrinodosum, with characteristic nodules in the posterior margin. This species is recorded in interior basins in Northeast Brazil, of Jurassic age (Silva et al. 1978a; Coimbra et al. 2002; Guzmán-González et al. 2020), so if the range of the T. quadrinodosum is confirmed, it would be recorded as far back as the Aptian. What made the authors leave this species in confer is the fact that they found only a single, unique specimen among all the samples. Stratigraphic distribution. Brazil, Araripe Basin, Formação, Brejo Santo, Jurassic (Coimbra et al. 2002). Jatobá Basin, Aliança Formation, Jurassic (Guzmán-González et al. 2020). Araripe Basin, Romualdo Formation, upper Aptian.

Paracypris undulareventralis sp. nov. Fig. 4D–F 2022 Paracypris sp. —Araripe et al., p 7, fig. 5 (8a–b). Etymology: Undulare (lat.), suggested due to the significant concave undulation, and ventralis (lat.) where the undulation is present. Holotype: Adult shell, Nº. DGEO-CTG-UFPE-1517 (Fig. 4D). Illustrated material from the Santo Antônio section 2.65 m. Paratypes: Adult shell, Nº. DGEO-CTG-UFPE-1518 (Fig. 4E and F). Illustrated material from the Santo Antônio section 0.3 m. Type locality: Brazil, State of Pernambuco, Municipality of Exu, Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type-horizon: Santo Antônio section, Araripe Basin. Calcarenite (F-Sl) corresponds to Romualdo Formation, depth 0.3 m, dated as upper Aptian. Dimension. Nº. DGEO-CTG-UFPE-1517 (Fig. 4D); L= (LV) 0.568 / (RV) 0.635mm, H= 0.299mm, and W= 0.220mm. Diagnosis: Carapace smooth, with a pronounced concave sinuosity in the posteroventral region. Description: Carapace medium, sub-triangular in lateral view. VM with a concave sinuosity in the posteroventral region. Dorsal margin angled in the interior portion that coincides with the greatest height. Rounded anterior and posterior margins, the latter being lower. Greater length in the ventral region, just above the ventral sinuosity. Greater anterodorsal height coinciding with the location of the cardinal angle. LV larger than RV with evident overlaps on the central-ventral margin. In the dorsal view, greatest width in the posterior region. Smooth surface. Observations. Paracypris undulareventralis sp. nov. was compared with Candonopsis alagoensis Tomé et al. (2014) deposited in the Ivan de Medeiros Tinoco Microfossils Collection. The studied species has a very marked sinuosity in the posteroventral region while C. alagoensis does not. Occurrence and associated fauna. It was recorded in the Santo Antônio section, at the following depths (from base to top): 0.3; 2.25; 2.65; 3.55; 3.65; 5.25; 5.45 and 5.55 m. Paracypris undulareventralis sp. nov. does not occur with Dicrorygma (Orthorygma ?) dimorpha and Theriosynoecum cf. quadrinodosum . Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 544, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Tome, M. E. T. R., Lima Filho, M. F. & Neumann, V. H. M. L. (2014) Taxonomic studies of non-marine ostracods in the Lower Cretaceous (Aptian-lower Albian) of post-rift sequence from Jatoba and Araripe basins (Northeast Brazil): Stratigraphic implications. Cretaceous Research, 48, 153 - 176. https: // doi. org / 10.1016 / j. cretres. 2013.12.007"]}

Pattersoncypris angulata (Krömmelbein & Weber, 1971) Fig. 3D–F 1966 Paraschuleridea sp. 5 —Braun, p. 66–67, fig. 1, 2a–b. 1971 Hourcqia angulata angulata —Krömmelbein & Weber, p. 20, plate 6, fig. 23a–c. 1989 Hourcqia angulata angulata —Viana et al., p. 216, fig. 2a–c. 1990 Hourcqia angulata angulata —Silva Telles Jr. & Viana, plate 3, fig. 3. 1990 Hourcqia angulata angulata —Dépêche et al., plate 3, figs.17–18. 2002 Pattersoncypris angulata angulata — Coimbra et al., p. 691, fig. 4, nº 29. 2006 Harbinia angulata —Ramos et al., p. 344, fig. 4E–H. 2008 Harbinia angulata —Do Carmo et al., p. 795, fig. 6, nº 11. 2010 Pattersoncypris angulata — Tomé & Lima Filho, p. 62, plate. 1 (d–f). 2012a Kroemmelbeincypris angulata —Poropat & Colin, p. 708, fig. 4, nº 4 and 5. 2022 Pattersoncypris angulata —Araripe et al., p. 7, fig. 5 (14a–b). Illustrated material. Adult shell, Nº. DGEO-CTG-UFPE-1507. Illustrated material from the Santo Antônio section (2.65 m). Dimension. Nº. DGEO-CTG-UFPE-1507 (Fig. 3D–F); L= (LV) 0.917/ (RV) 1.197 mm, H= 0.571 mm, and W= 0.729 mm. Comparison. Material compared to Pattersoncypris angulata (Nº. DGEO-CTG-UFPE-1963), described and deposited by Tomé et al. (2014) in the Ivan de Medeiros Tinoco Microfossils Collection from Paleontology Laboratory of the Federal University of Pernambuco. The comparison was satisfactory, due to the posterodorsal angle being well marked, however, the species found in the collection is smooth with no surface ornamentation. Occurrence and associated fauna. It was recorded in the Santo Antonio section, at the following depths (from base to top): 1.8; 2.1; 2.25; 3.1; 3.4; 3.85; 5.25 and 5.35 m. P. salitrensis is associated with all species of ostracods, except Dicrorygma (Orthorygma ?) dimorpha. Observations. Among the forms belonging to the genus Pattersoncypris found in the interval corresponding to the Aptian-Albian, in the Romualdo Formation, Pattersoncypris angulata presents a well-marked angle at the posterodorsal end, which helps its identification. This species can be smooth or with a tessellated ornamentation on the surface. Stratigraphic distribution. Brazil, Araripe Basin, Santana Formation, Romualdo Member (Silva Telles Jr. & Viana, 1990; Poropat & Colin, 2012b), range dated as Aptian (Regali 1990; Coimbra et al. 2002). Potiguar Basin, Alagamar Formation, Aptian (Do Carmo et al. 2008). Cedro Basin, Crato Formation, range dated as Aptian (Tomé 2007; Tomé & Lima Filho 2010). Jatobá Basin, Serra Negra, in strata correlated to the Crato Formation, upper Aptian (Tomé et al. 2014). In the present study, found in the Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 539, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Krommelbein, K. & Weber, R. (1971) Ostracoden des \" Nordost-Brasilianischen Wealden \". Geologisches Jahrbuch, 115, 1 - 93.","Tome, M. E. T. R., Lima Filho, M. F. & Neumann, V. H. M. L. (2014) Taxonomic studies of non-marine ostracods in the Lower Cretaceous (Aptian-lower Albian) of post-rift sequence from Jatoba and Araripe basins (Northeast Brazil): Stratigraphic implications. Cretaceous Research, 48, 153 - 176. https: // doi. org / 10.1016 / j. cretres. 2013.12.007","Poropat, S. F. & Colin, J. P. (2012 b) Reassessment of the Early Cretaceous non-marine ostracod genera Hourcqia Krommelbein, 1965 and Pattersoncypris Bate, 1972 with the description of a new genus, Kroemmelbeincypris. Journal of Paleontology, 86, 699 - 719. https: // doi. org / 10.1666 / 11 - 140 R. 1","Regali, M. S. P. (1990) Biocronoestratigrafia e paleoambiente do Eocretaceo das bacias do Araripe (CE) e Rio do Peixe (PB), NE, Brasil. Simposio sobre a Bacia do Araripe e Bacias Interiores do Nordeste, 1, 163 - 171.","Coimbra, J. C., Arai, M. & Carreno, A. L. (2002) Biostratigraphy of Lower Cretaceous microfossils from the Araripe Basin, northeastern Brazil. Geobios, 35, 687 - 698. https: // doi. org / 10.1016 / S 0016 - 6995 (02) 00082 - 7","Do Carmo, D. A., Whatley, R., Queiroz Neto, J. V. Q. & Coimbra, J. C. (2008) On the validity of two Lower Cretaceous non-marine ostracode genera: biostratigraphic and paleogeographic implications. Journal of Paleontology, 82, 790 - 799. https: // doi. org / 10.1666 / 07 - 008.1","Tome, M. E. T. R. & Lima Filho, M. F. (2010) Ostracodes da Bacia de Cedro, Andar Alagoas, Estado de Pernambuco, NE, Brasil: Implicacoes paleoambientais e bioestratigraficas. Estudos Geologicos, 21, 56 - 70."]}

Mongolianella aptianensis sp. nov. Fig. 3J–O 2021 Damonella grandiensis — Araripe et al., p. 6, fig. 5 (M–P). 2022 Mongolianella sp. —Araripe et al., p. 7, fig. 5 (7a–b). Etymology: Aptian (lat.) derived from the range where this species was found. Holotype: Adult carapace, Nº. DGEO-CTG-UFPE-1511 (Fig. 3J–L). Paratype: Juvenile carapace, Nº. DGEO-CTG-UFPE-1512 (Fig. 3M–O). Type locality: Brazil, State of Pernambuco, Municipality of Exu, Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type-horizon: Santo Antônio section, Araripe Basin. Shale (F1) corresponding to Romualdo Formation, depth 3.00m, dated as upper Aptian. Dimension. Nº. DGEO-CTG-UFPE-1511 (Fig. 3J–L); L= (LV) 0.901 / (RV) 0.979mm, H= 0.397mm, and W= 0.307mm. Diagnosis: Large, smooth carapace with an outline that makes a sharp angle at the posteroventral margin. Description. Carapace slightly concave in the central-ventral margin. Dorsal margin angular in the antero and posterodorsal portions in adults, while in juveniles the anterior angular portion promotes a strong fall to the anterior margin. In juveniles the posterodorsal cardinal angle is lower. Anterior margin rounded in both stages, unlike the posterior margin in adults, which is more rectangular, and in juveniles it is triangular. Maximum length at the ventral region. Maximum height at anterodorsal region, coinciding with the location of the cardinal angle. LV larger than RV with evident overlap along the central ventral margin, in juvenile specimens there is overlap along the dorsal margin, making the cardinal angle more prominent. In dorsal view, the greatest width is at the posterior margin, but narrower in the juvenile form. Smooth outer surface. Observations. A very large and elongated species compared to the others of the genus. This species stands out for its size. First, it was considered that this species is of the marine genus Krithe Brady, Crosskey & Robertson, 1867, but this genus is typical of deep waters, and so this idea was discarded. Detailed comparisons were also made with the species of Damonella described and deposited by Tome et al. (2014) and Araripe et al. (2021) in the Ivan de Medeiros Tinoco Microfossils Collection. The studied form is very large when compared to Damonella grandiensis Tomé et al., 2014 and Damonella ultima Tomé et al., 2014. When compared, it presents the same characteristics of the species found by Araripe et al. (2021). It is worth noting that they are totally smooth specimens with a difficult level of identification, since they do not have ornamentation to use for comparisons. In the present study, we chose to call holotype Nº. DGEO-CTG-UFPE-1511 as an adult form and the paratype Nº. DGEO-CTG-UFPE-1512 as a young form, however further studies with a larger population would be necessary to recognise the male carapace and others ontogenetic stages. Occurrence and associated fauna. It was recorded in the Santo Antônio section, at the following depths (from base to top): 1.8; 2.1; 2.25; 3.1; 3.4; 3.45; 3.55; 3.65; 3.85 and 4.05 m. Mongolianella aptianensis sp. nov. does not occur in association with Dicrorygma (Orthorygma ?) dimorpha. Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian.

Perissocytheridea poruslinearis sp. nov. Fig. 4L–O 2022 Perissocytheridea sp.2 —Araripe et al., p 7, fig. 5 (2a–b). Etymology: Porus (lat.) and linearis (lat.) in reference to the sequence of linear pores that occurs along the dorsal margin. Holotype: Adult carapace, Nº. DGEO-CTG-UFPE-1520 (Fig. 4L–O). Type locality: Brazil, State of Pernambuco, Municipality of Exu, Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type horizon: Santo Antônio section, Araripe Basin. Calcarenite (F-Sl) corresponds to the Romualdo Formation, depth 0.3 m, dated as upper Aptian. Dimension. Nº. DGEO-CTG-UFPE-1520 (Fig. 4L–O); L= (LV) 0.383 / (RV) 0.320 mm, H= 0.153 mm, and W= 0.142 mm. Diagnosis: Carapace very small. Presence of pores aligned on the dorsal margin. Reticular ornamentations but more subtle when compared to other species of Perissocytheridea described in this study. Description: Carapace very small. Subrectangular outline in lateral view. Ventral margin rectilinear on both valves. Dorsal margin rectilinear with drop to posterior end. Anterior margin rounded. Posterior margin sub-triangular tapering to a subtle caudal process, not clear in lateral view but coincides with the greatest length. Greatest height at anterior margin. Left valve slightly overlaps right valve. In the dorsal view, tapering more to anterior to the posterior margin, sharp and short ends. Surface ornamented with smooth primary reticulations and pores spread over the surface, with emphasis on the sequence of three pores along the dorsal margin of each valve. Winged process discrete in the posteroventral margin. Ocular tubercle evident. Observations. Compared to the other two species of Perissocytheridea, the surface has a very slightly primary reticulated ornamentation and the presence of pores spread on the surface with the set of six sequenced pores on the dorsal margin. Occurrence and associated fauna. It was recorded in the Santo Antônio section, at the following depths (from base to top): 0.3; 2.25; 2.65; 3.1; 4.05; 3.55 and 5.45 m. Perissocytheridea poruslinearis sp. nov. does not occur with Dicrorygma (Orthorygma ?) dimorpha. Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 547, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092

Microceratina retangularis sp. nov. Fig. 5J–L 2022 Microceratina sp. —Araripe et al., p 7, fig. 5 (5a–b). Etymology: Retangularis (lat.) in reference to the carapace outline. Holotype: Adult carapace, Nº. DGEO-CTG-UFPE-1525 (Fig. 5J–L). Type locality: Brazil, State of Pernambuco, Municipality of Exu, Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type horizon: Santo Antônio section, Araripe Basin. Calcarenite (F-Sl) corresponds to the Romualdo Formation, depth 3.10 m, dated as upper Aptian. Dimension. Nº. DGEO-CTG-UFPE-1525 (Fig. 5J–L); L= (LV) 0.353 / (RV) 0.335 mm, H= 0.223 mm, and W= 0.191 mm. Diagnosis: Carapace small and rectangular. It has reticular ornamentation and a ventral posterolateral projection. Description: Carapace very small. Rectangular outline in the lateral view. Ventral margin rectilinear and parallel. Dorsal margin rectilinear, descends slightly towards the posterior. Anterior margin sub-rectangular. Posterior margin rounded, truncated at ventral margin by small winged process, greater mid-length. Greatest height along the anterior margin. Left valve slightly overlaps right valve in the posterodorsal margin. In the dorsal view, valves are narrow, with the posterior margin subtly wider, ends sharp and short. Surface ornamented with smooth primary reticulations. Winged process at the posteroventral margin. Observations. Small species, easy to recognize as it presents a rectangular shape. The main difference with the genus Perissocytheridea studied in the present work is the non-occurrence of a caudal process. Occurrence and associated fauna. It was recorded in the Santo Antonio section, at the following depths (from base to top): 2.65 and 3.4 m. Microceratina retangularis n. sp. does not occur with Alicenula cf. leguminella, Theriosynoecum cf. quadrinodosum and Dicrorygma (Orthorygma ?) dimorpha. Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 549, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092

Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo, Ng, Christiano (2022): Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species. Zootaxa 5159 (4): 535-557, DOI: https://doi.org/10.11646/zootaxa.5159.4.4

Pattersoncypris micropapillosa Bate, 1972 Fig. 3G–I 1972 Pattersoncypris micropapillosa — Bate, p. 379–393, est. 66–71, figs. 1–12. 1990 Pattersoncypris micropapillosa —Arai & Coimbra, p. 239. plate. 1, fig. 2. 1990 Pattersoncypris micropapillosa —Dépêche et al., plate 1, fig. 9. 1991 Hourcqia angulata sinuata —Andreu-Boussut, plate 1, figs. 4–9. 2008 Harbinia micropapillosa —Do Carmo et al., p. 795, fig. 6, nº 6. 2000 Pattersoncypris micropapillosa —Smith, p. 70–83, plates 1–9, text-figs. 2–9. 2002 Pattersoncypris micropapillosa — Coimbra et al., fig. 3. 2010 Pattersoncypris micropapillosa — Tomé & Lima Filho, p. 62, plate 1 (a–c). 2012a Pattersoncypris micropapillosa —Poropat & Colin, p. 708, fig. 4, nº. 1–2. 2014 Pattersoncypris micropapillosa —Tomé et al., p. 167, fig. 10 J–P. 2017 Harbinia micropapillosa —Souza et al., p. 7, fig. 1, A–D. 2020 Pattersoncypris micropapillosa — Melo et al., fig. 5, nº 5–6. 2022 Pattersoncypris micropapillosa —Araripe et al., p 7, fig. 5 (15a–b). Illustrated material. Juvenile carapace (A-1), Nº. DGEO-CTG-UFPE-1508, depth 3.1 m; Nº. DGEO-CTG-UFPE- 1509, depth 4.05 m and Nº. DGEO-CTG-UFPE-1510, depth 3.1 m. Dimension. Nº. DGEO-CTG-UFPE-1508 (Fig. 3G); L= (LV) 1,045 / (RV) 1.173 mm, H= 0.69 mm, and W= 0.744 mm. Comparison. Material compared to P. micropapillosa (Nº. DGEO-CTG-UFPE-1164) described and deposited by Tome et al. (2014) in the Ivan de Medeiros Tinoco Microfossils Collection. Occurrence and associated fauna. It was recorded in the Santo Antônio section, at the following depths (from base to top): 1.8; 2.65; 3.1; 4.25; 5.25; 4.25; 5.35 and 5.55 m. P. micropapillosa is not associated with the species Alicenulla cf. leguminella, Dicrorygma (Orthorygma ?) dimorpha and the species of the genus Perissocytheridea. Observations. P. micropapillosa was not found in the same abundance as the other species of Pattersoncypris so it is noteworthy because this genus was described for the first time in this basin. Further studies are required to assess whether this species is a synonym of P. salitrensis or P. angulata. Stratigraphic distribution. Brazil, Araripe Basin, Santana Group, Romualdo Formation (Silva Telles Jr. & Viana 1990; Poropat & Colin 2012b), interval dated as Aptian (Regali 1990; Coimbra et al. 2002). Potiguar Basin, Alagamar Formation, Aptian (Do Carmo et al. 2008). Cedro Basin, Crato Formation, interval dated as Aptian (Tomé 2007; Tomé & Lima Filho 2010). Jatobá Basin, Serra Negra, in strata correlated to the Crato Formation, upper Aptian (Tomé et al. 2014). In the present manuscript, it was observed in the Araripe Basin, Romualdo Formation, upper Aptian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 540, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Bate, R. H. (1972) Phosphatized Ostracods with Appendages from the Lower Cretaceous of Brazil. Paleontology, 15, 379 - 393.","Tome, M. E. T. R., Lima Filho, M. F. & Neumann, V. H. M. L. (2014) Taxonomic studies of non-marine ostracods in the Lower Cretaceous (Aptian-lower Albian) of post-rift sequence from Jatoba and Araripe basins (Northeast Brazil): Stratigraphic implications. Cretaceous Research, 48, 153 - 176. https: // doi. org / 10.1016 / j. cretres. 2013.12.007","Poropat, S. F. & Colin, J. P. (2012 b) Reassessment of the Early Cretaceous non-marine ostracod genera Hourcqia Krommelbein, 1965 and Pattersoncypris Bate, 1972 with the description of a new genus, Kroemmelbeincypris. Journal of Paleontology, 86, 699 - 719. https: // doi. org / 10.1666 / 11 - 140 R. 1","Regali, M. S. P. (1990) Biocronoestratigrafia e paleoambiente do Eocretaceo das bacias do Araripe (CE) e Rio do Peixe (PB), NE, Brasil. Simposio sobre a Bacia do Araripe e Bacias Interiores do Nordeste, 1, 163 - 171.","Coimbra, J. C., Arai, M. & Carreno, A. L. (2002) Biostratigraphy of Lower Cretaceous microfossils from the Araripe Basin, northeastern Brazil. Geobios, 35, 687 - 698. https: // doi. org / 10.1016 / S 0016 - 6995 (02) 00082 - 7","Do Carmo, D. A., Whatley, R., Queiroz Neto, J. V. Q. & Coimbra, J. C. (2008) On the validity of two Lower Cretaceous non-marine ostracode genera: biostratigraphic and paleogeographic implications. Journal of Paleontology, 82, 790 - 799. https: // doi. org / 10.1666 / 07 - 008.1","Tome, M. E. T. R. & Lima Filho, M. F. (2010) Ostracodes da Bacia de Cedro, Andar Alagoas, Estado de Pernambuco, NE, Brasil: Implicacoes paleoambientais e bioestratigraficas. Estudos Geologicos, 21, 56 - 70."]}

Perissocytheridea florisdorsalis sp. nov. Fig. 5A–C 2022 Perissocytheridea sp.3 —Araripe et al., p 7, fig. 4 (3a–b). Etymology: Floris (lat.) due to the design of a flower in the ornamentation of the carapace and dorsalis (lat.) marks the region and view in which the said design is observed. Holotype: Adult C, Nº. DGEO-CTG-UFPE-1521 (Fig. 5A–C). Type locality: Brazil, State of Pernambuco, Municipality of Exu, Sítio Santo Antônio section. Coordinates 7° 30’ 06” S and 39° 32’ 36” W. Type horizon: Santo Antônio section, Araripe Basin. Shale (F1) corresponding to Romualdo Formation, depth 4.05 m, dated as upper Aptian. Dimension. Nº. DGEO-CTG-UFPE-1521 (Fig. 5A–C); L= (LV) 0.373 / (RV) 0.356 mm, H= 0.189 mm, and W= 0.171 mm. Diagnosis: Carapace very small, reticulated surface and a subtle caudal process at the dorsal margin. Additionally with a flower design in the dorsal view. Description: Carapace very small. Sub-rectangular outline in lateral view. Ventral margin rectilinear on both valves. Dorsal margin straight descends to the posterior end. Anterior margin sub-rounded, beginning dorsally rectilinear and rounding up to ventral margin. Posterior margin sub-rectangular tapering to the caudal processes, which coincides with greatest length. Greatest height at anterior margin. Left valve slightly overlaps right valve. In the dorsal view the caudal process is evident, giving the posterior end a sharp appearance. Anterior margin also presents a sharp appearance. Surface with primary reticulations. Winged process discrete in the postero-ventral margin. Presents a lateral sulcus, where a flower-like design is seen in the dorsal view. Discrete ocular tubercle blending with ornamentation. Observations. This species differs from others species to the genus described in this paper by the presence of a flower-shaped ornamentation in the dorsal view. However, a further detailed study with a larger population is needed to differentiate the male and others ontogenetic stages. Occurrence and associated fauna. It was recorded in the Santo Antônio section, at the following depths (from base to top): 0.3; 2.25; 2.65; 3.1; 4.05; 3.55 and 5.45 m. Perissocytheridea florisdorsalis sp. nov. does not occur with Dicrorygma (Orthorygma ?) dimorpha. Stratigraphic distribution. Brazil, Araripe Basin, Romualdo Formation, upper Aptian.

Dicrorygma (Orthorygma ?) dimorpha Weiss, 1996 Fig. 5G–I 1990 Ostracoda gen. et sp. indet,—Zihrul: 89, Taf. 6, Fig. 60. 1996 Dicrorygma (Dicrorygma ?) n. sp. a—Weiss: 119, Abb. 10 Fig. 7–8; Tar. 1, Fig. 10–15; Taf. 2, Fig. 8. 2022 Dicrorygma (Orthorygma?) dimorpha —Araripe et al., p 7, fig. 5 (6a–b). Illustrated material. Adult carapace, Nº. DGEO-CTG-UFPE-1524 (Fig. 5 G–I) Dimension. Nº. DGEO-CTG-UFPE-1524 (Fig. 5 G–I); L= (LV) 0.426 / (RV) 0.396 mm, H= 0.233 mm, and W= 0.213 mm. Comparison. It was not possible to compare with reference collection materials. Dicrorygma (Orthorygma ?) dimorpha was compared to illustrations in Michael Weiss (1996); both species are very small, with a smooth surface and subtriangular carapace. The posterior end is sharp, joining the small caudal process in lateral view. The dorsal view is very similar, but the caudal process is smaller in the specimen found. We also compared the species studied with the illustrations of the species Parataxodonta inornate Kaye 1965, but they are different in the dorsal view. Occurrence and associated fauna. It was recorded in the Santo Antônio section, exclusively at following depth (from base to top) of 5.65 m. Dicrorygma (Orthorygma ?) dimorpha occurs with D. grandiensis and Dicrorygma cf. minuta, which occurs throughout most of the section. Observations. Dicrorygma (Orthorygma ?) dimorpha and Dicrorygma cf. minuta comprise the first record of this genus in the Araripe Basin (Aptian). Weiss (1996), characterized the sexual dimorphism for this species, and the specimen found is similar to the male described by this author. It is worth pointing out that it is a very small species, and like the species belonging to the genus Perissocytheridea Stephenson, 1938, they are only recovered in the smallest sieves and in the sorting process in the finest granulometry, thus requiring greater attention and persistence to recover. Stratigraphic distribution. NW-German Kimmeridgian, Upper section (sample 5/7a), Lower Kimmeridge (ostracod-zone 10, Kimmeridgian) (Babinot et al. 2007). Brazil, Araripe Basin, Romualdo Formation, upper Aptian (this work)., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on pages 548-549, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Weiss, M. (1996) Neue Ostracoden aus dem Kimmeridgium NW-Deutschlands. Palaontologische Zeitschrift, 70 (3 / 4), 461 - 474. https: // doi. org / 10.1007 / BF 02988085","Kaye, P. (1965) Some new British Albian Ostracoda. Bulletin of the British Museum (Natural History), 11 (5), 215 - 253.","Stephenson, M. B. (1938) Lower Eocene Ostracoda of the genus Cytheridea. Journal of Paleontology, 10 (8), 695 - 703.","Babinot, J. F., Moullade, M. & Tronchetti, G. (2007) The upper Bedoulian and lower Gargasian Ostracoda of the Aptian strato- type: Taxonomy and biostratigraphic correlation. Carnets de Geologie, 5, 1 - 35. https: // doi. org / 10.4267 / 2042 / 8877"]}

Alicenula cf. leguminella (Forbes, in Lyell 1855) Fig. 5M–O 1855 Cypris leguminella —Forbes (in Lyell, 1855), p. 294, Fig. 334c. 1855 Darwinula leguminella —Jones, p. 346, Taf. 8, Figs. 3 –31. 1856 Darwinula leguminella —Jones, p. 147, Taf. 4, Fig. 4a–c. 1940 Darwinula leguminella —Martin, p. 317, Taf. 4, Figs. 58–61. 1954 Darwinula leguminella —Bartenstein & Burri, Taf. 26. 1957 Darwinula leguminella —Grekoff, p. 61, Plate 3, Figs. 56 and 57. 1961 Darwinula leguminella —Martin, p. 119, Taf. 14, Fig. 19. 1962 Darwinula leguminella —Kringler et al., p. 187–188, Taf. 25, Fig. 14. 1963 Darwinula leguminella —Oertli, p. 20, Taf. 6, Fig. 40. 1963 Darwinula leguminella —Christensen, p. 21–23, Taf. 2, Fig. 2 aec. 1966 Darwinula leguminella —Barker, p. 472, Taf. 7, Fig. 9. 1971 Darwinula leguminella —Anderson, p. 110, Taf. 18. 1975 Darwinula leguminella —Bielecka, p. 368, Taf. 14, Fig. 5. 1976 Darwinula leguminella —Brenner, p. 141, Taf. 10, Figs. 23–25. 1985 Darwinula leguminella —Colin & Oertli, p.160, Taf. 40, Fig.1. 1978b Darwinula martinsi —Silva, p. 1028–31, Plate 1, Figs. 1–2. 1990 Darwinula cf. martinsi —Silva Telles Jr. & Viana, Plate 2, Fig. 4. 1997 Darwinula cf. martinsi —Colin & Dépêche, Plate 2:18. 2002 Darwinula martinsi — Coimbra et al., Fig. 4:32. 2004 Darwinula martinsi —Do Carmo et al., p. 50–51, Fig. 9:13 2010 Darwinula cf. martinsi —Tomé & Lima Filho, p. 63, Plate 2, Fig. d–f. 2013 Darwinula martinsi — Do Carmo et al., p. 95e103, Fig. 5, N?.9. 2014 Alicenula leguminella (Forbes in Lyell 1855)—Tomé et al., p. 170, Fig.14 (T–Z). 2020 Alicenulla sp.—Melo et al., p 9, Fig. 5 (11–12). 2022 Alicenula leguminella —Araripe et al., p 7, fig. 5 (12a–b). Illustrated material. Juvenile shell, Nº. DGEO-CTG-UFPE-1526. Illustrated material from the Santo Antônio section (0.3 m). Dimension. Nº. DGEO-CTG-UFPE-1526 (Fig. 5M–N); L= (LV) 0.376 mm, and W= 0.172 mm and Nº. DGEO- CTG-UFPE-1527 (Fig. 5O); L= (RV) 0.424 mm and H= 0.143 mm. Comparison. Material compared to Alicenulla leguminella (Nº DGEO-CTG-UFPE-1174), described and deposited by Tomé et al. (2014) in the Ivan de Medeiros Tinoco Microfossils Collection of the Paleontology Laboratory of the Federal University of Pernambuco. It was observed that the specimen found in the present study was much smaller, possibly a juvenile form. The ventral covering of the right valve was not observed. Thus, the specimen studied was assigned the confer . Occurrence and associated fauna. It was recorded in the Santo Antonio section, at the following depths (from base to top): 0.3; 2.1 and 2.25 m. A. cf. leguminella is associated with all species of ostracods, except Dicrorygma (Orthorygma ?) dimorpha, Microceratina retangularis sp. nov. and T. cf. quadrinodosum. Observations. It is worth pointing out the association of this species with the planktonic foraminifera Pseudoguembelitria sp. and Liliputinella? sp., thus suggesting that such forms may occur in typically brackish-marine environments. Stratigraphic distribution. Brazil, Araripe Basin, Santana Formation, Crato Member, and base of Ipupi Member (Silva 1978b; Silva Telles Jr. & Viana 1990; Colin & Dépêche 1997); interval dated as Aptian (Regali 1990; Coimbra et al. 2002). Potiguar Basin, Alagamar Formation, Aptian (Do Carmo et al. 2013). Cedro Basin, in strata correlated to the Crato Formation, Aptian (Tomé 2007; Tomé & Lima Filho 2010). Jatobá Basin, Serra Negra, in deposits correlated to the Crato Formation, upper Aptian (Tomé et al. 2014). The rocks of the Puberck and Wealden, in England, contain records of this species from the Jurassic to the lower Aptian (Anderson 1985). In this work, Araripe Basin, Romualdo Formation, upper Aptian - lower Albian., Published as part of Tomé, Maria Emilia, Araripe, Rilda, Oliveira, David, Barreto, Alcina, Prado, Ludmila, Pedrosa, Flavia, Pereira, Priscilla, Nascimento, Luiz Ricardo & Ng, Christiano, 2022, Early Cretaceous Ostracoda (Crustacea) from south-centralAraripe Basin, Brazil with descriptions of seven new species, pp. 535-557 in Zootaxa 5159 (4) on page 550, DOI: 10.11646/zootaxa.5159.4.4, http://zenodo.org/record/6786092, {"references":["Tome, M. E. T. R., Lima Filho, M. F. & Neumann, V. H. M. L. (2014) Taxonomic studies of non-marine ostracods in the Lower Cretaceous (Aptian-lower Albian) of post-rift sequence from Jatoba and Araripe basins (Northeast Brazil): Stratigraphic implications. Cretaceous Research, 48, 153 - 176. https: // doi. org / 10.1016 / j. cretres. 2013.12.007","Silva, M. D. (1978 b) Ostracodes da Formacao Santana (Cretaceo Inferior), Grupo Araripe, Nordeste do Brasil. 3. Nova especie do genero Darwinula Brady & Robertson, 1885. Congresso Brasileiro de Geologia, 30, 1028 - 1031.","Colin, J. P. & Depeche, F. (1997) Faunes d'ostracodes lacustres des basins intra-cratoniques d'age albo-aptien en Afrique de l'Ouest (Cameroun, Tchad) et au Bresil: considerations d'ordre paleoecologique et paleobiogeographique. African Geoscience Review, 4 (3 / 4), 431 - 450.","Regali, M. S. P. (1990) Biocronoestratigrafia e paleoambiente do Eocretaceo das bacias do Araripe (CE) e Rio do Peixe (PB), NE, Brasil. Simposio sobre a Bacia do Araripe e Bacias Interiores do Nordeste, 1, 163 - 171.","Coimbra, J. C., Arai, M. & Carreno, A. L. (2002) Biostratigraphy of Lower Cretaceous microfossils from the Araripe Basin, northeastern Brazil. Geobios, 35, 687 - 698. https: // doi. org / 10.1016 / S 0016 - 6995 (02) 00082 - 7","Do Carmo, D. A., Coimbra, J. C., Whatley, R. C., Antonietto, L. S. & Citon, R. T. P. (2013) Taxonomy of limnic Ostracoda (Crustacea) from the Alagamar Formation, middle-upper Aptian, Potiguar Basin, Northeastern Brazil. Journal of Paleontology, 87, 91 - 104. https: // doi. org / 10.1666 / 11 - 108 R. 1","Tome, M. E. T. R. & Lima Filho, M. F. (2010) Ostracodes da Bacia de Cedro, Andar Alagoas, Estado de Pernambuco, NE, Brasil: Implicacoes paleoambientais e bioestratigraficas. Estudos Geologicos, 21, 56 - 70."]}

Orientadora: Profa. Dra. Cristina Silveira Vega Coorientadora: Dra. Maria da Saudade Araujo Santos Maranhão Tese (doutorado) - Universidade Federal do Paraná, Setor de Ciências da Terra, Programa de Pós-Graduação em Geologia. Defesa : Curitiba, 18/06/2019 Inclui referências Resumo: A sucessão sedimentar permiana da Bacia do Paraná inclui o Grupo Passa Dois, que engloba quatro formações, Irati, Serra Alta, Teresina e Rio do Rasto, e representa uma transição de ambientes deposicionais marinhos para continentais. Este intervalo é palco de várias discussões paleoambientais, porém microfácies haviam sido ainda pouco estudadas. Além disso, soma-se a escassez de trabalhos micropaleontológicos, a falta de calibração geocronológica e ausência de fósseisguia marinhos. Diante deste cenário, propôs-se um estudo de microfácies abrangente para a unidade, com análise de amostras amplamente distribuída e representativa da faixa de ocorrência do grupo, divididas em três grandes domínios, norte, central e sul. Foram analisadas lâminas petrográficas distribuídas entre as quatro unidades com ênfase nas camadas carbonáticas. Os dados foram confrontados com um quadro biocronoestratigráfico revisado e integrado com biozonas de bivalves, macroflora, palinologia, conchostráceos e vertebrados. As análises microfaciológicas resultaram no reconhecimento de 13 microfácies, quatro delas siliciclásticas, oito carbonáticas e uma silicosa. Microfácies siliciclásticas finas predominam no Domínio Sul; microfácies carbonáticas rudáceas no Domínio Central, com maior quantidade de aloquímicos e bioclastos; e maior influência de siliciclásticos no Domínio Norte da bacia, onde ocorrem apenas as formações Irati e Teresina. As microfácies carbonáticas ao sul são menos variadas que aquelas ao norte e, ainda no Domínio Sul, há uma escassez de microfósseis. No Domínio Central ocorrem os carbonatos com maior granulação, relacionados à maior agitação da água. Entre os microfósseis foram identificados ostracodes, espículas de espongiários, bivalves, oósporos carbonificados, microfósseis tubulares de parede microgranular, microbialitos, fragmentos de artrópodes, restos fosfáticos e fitoclastos. Bivalves, ostracodes e espículas de espongiários são os principais bioclastos presentes em rudstones e grainstones nas formações Irati e Teresina, e foram interpretados como alóctones a parautóctones. Apresentam-se ainda os resultados petrográficos dos estudos em esponjas do Grupo Passa Dois. A morfologia, forma e estrutura dos fósseis pode ser atribuída à Ordem Haplosclerida, como associado previamente, de ambientes marinhos como indicados pela associação com glauconita. As microfácies presentes em três furos de sondagem de Anhembi (SP) refletem processos sedimentares atuando em ambientes marinhos marginais sob influência de fluxos oscilatórios, tais como ondas de tempestade e marés, próximos a corpos de água doce e desembocaduras de rios, que trazem consigo elementos continentais. Neste cenário, microfósseis marinhos coexistem regionalmente com elementos não marinhos. Evidências da literatura foram somadas aos resultados e possibilitaram a proposta de um modelo de distribuição de microfácies para a Formação Teresina, plataformal raso em clima progressivamente mais seco, com shoals e mounds microbialíticos nos altos e depósitos pelíticos e cherts nos fundos estagnados e com baixa oxigenação. A assembleia de microfósseis encontrada, embora abundante, é pouco diversificada em comparação com uma assembleia marinha rasa normal, o que deve relacionar-se possivelmente à condições de alto estresse ambiental, ou condições paleobatimétricas inviáveis ao desenvolvimento de outros organismos estenohalinos, o que pode estar relacionado às grandes extinções do Permiano, em apoio às datações geocronológicas mais recentes que indicam idades detríticas mais jovens para os estratos do Grupo Passa Dois. Palavras-chave: Petrografia, Microfácies, Esponjas, Guadalupiano, Permiano. Abstract: Permian sedimentary sucession of Paraná Basin include the Passa Dois Group, which encompasses four formations: the Irati, Serra Alta, Teresina, and Rio do Rasto, and represents a transition from marine to continental depositional environments. This interval has been speculated to have formed under several different paleoenvironmental scenarios; where microfacies have rarely been studied. In addition, there is a scarcity of micropaleontological studies and geochronological calibration, besides an absence of marine guide fossils. In view of this, a comprehensive microfacies study for the unit was proposed, with a large and representative surface and subsurface sample analysis of the Passa Dois Group, divided into three large geographic domains: north, central, and south. Thin sections distributed among the four units were analyzed with an emphasis on carbonate layers. The data were interpreted within a revised biochronostratigraphic framework and integrated with bivalves, macroflora, palynology, conchostracans, and vertebrates biozones. Microfacies analysis resulted in the recognition of 13 microfacies, four siliciclastic, eight carbonate, and one siliceous. Fine siliciclastic microfacies, rich in organic matter, predominate in the Southern Domain; rudaceous carbonate microfacies in the Central Domain, with greater diversity of allochemicals and bioclasts; and the increase on siliciclastic sediments influx in the Northern Domain of the basin, where only the Irati and Teresina formations occur. The carbonate microfacies to the south are less varied than those to the north along with a shortage of microfossils. In the Central Domain, coarse-grained carbonates formed by the high hydrodynamic conditions of the sedimentary processes are present. Among the microfossils identified were ostracods, sponge spicules, bivalves, organic oospores, tubular microfossils, microbialites, fragments of arthropods, phosphatic remains and phytoclasts. Bivalves, ostracods, and sponge spicules are the main bioclasts on rudstones and grainstones from the Irati and Teresina formations, and have been interpreted as allochthonous to parautochthonous. This study also presents the results of petrological studies on sponges from Passa Dois Group. The morphology, shape and structure of the sponges remains could be attributed them to the Order Haplosclerida as associated previously, from marine environments as indicated by the associated glauconite grains. Microfacies from three boreholes of Anhembi (SP), reflect sedimentary processes of marginal marine environments on influence of oscilatorious fluxes, such as tides and storm waves, close to fresh water bodies and river deltas, that bring with them continental elements. In this context, marine microfossils regionally coexist with non-marine bioclasts. Evidences from literature were added to these results and allow to propose a model of microfacies distribution for the Teresina Formation, a shallow platform in a progressively drier climate, with microbialitic shoals and mounds on shallow settings and pelitic deposits and cherts on hypoxic waters. The microfossil assemblage, although abundant, is poorly diversified compared to a normal shallow marine assemblage, possibly related to conditions of high environmental stress, or incompatible paleo-bathymetric depths for the development of other stenohaline organisms. On the other hand, it may be related to the great Permian extinctions, in support of the more recent geochronological dates that indicate younger detrital ages for the strata of the Passa Dois Group. Keywords: Petrography, Microfacies, Sponges, Guadalupian, Permian.

Resumo: A Formação Ponta Grossa, de idade devoniana, aflora na borda leste da Bacia do Paraná ao redor do Arco de Ponta Grossa. O objetivo deste trabalho foi realizar um levantamento da faciologia, bioestratigrafia e do paleoambiente acerca da formação. Para tanto, foram feitos trabalhos de campo, análises petrográficas e micropaleontológicas. Foram estudados afloramentos ao longo dos municípios de Palmeira, Ponta Grossa, Tibagi, Ventania e Jaguariaíva (PR), com os quais se determinou espessura atual de ~110 m de litologias pelíticas e psamíticas finas, considerando-se falhamentos e diques de diabásios presentes na área. Quatro associações de fácies identificadas representam as rochas na região de afloramentos, e associam-se a pelo menos três sequências estratigráficas de 3ª ordem que representam a passagem de trato de sistema transgressivo iniciado com o afogamento da Formação Furnas; instalação de trato de sistema de mar alto com progradação de fácies; e retomada do trato de sistema transgressivo, interrompido por superfície de discordância regional que representa hiato temporal de regressão forçada e sobreposição por arenitos conglomeráticos e diamictitos do Grupo Itararé de trato de sistema de mar baixo, encerrando a chamada Supersequência Paraná, transgressiva, marinha, de águas frias. O paleoambiente inferido para o intervalo da Formação Ponta Grossa analisado varia de nerítico profundo anóxico a raso óxido, dominado por ondas de tempestade, entre o nível de base das ondas de tempo bom até abaixo do nível das ondas de tempestade. A caracterização petrográfica subsidiou a identificação e descrição com detalhe de elementos sedimentológicos e micropaleontológicos para a unidade. Entre os elementos micropaleontológicos, ressalta-se a descoberta inédita dos mais antigos foraminíferos bentônicos aglutinantes arenáceos e também de radiolários, no Brasil, o que abre caminho a novas fronteiras científicas. Escolecodontes, fitoclastos (Spongiophyton spp.), quitinozoários, Tasmanites spp., moldes e elementos indeterminados compõem os principais elementos reconhecidos em seções delgadas. Suas distribuições ambientais e estratigráficas são apontadas, com concentração de fitoclastos, escolecodontes e quitinozoários, nas porções mais proximais, enquanto radiolários e Tasmanites spp. concentram-se em folhelhos abaixo do nível das ondas de tempestade. Verificou-se ainda a aplicabilidade do arcabouço bioestratigráfico proposto na literatura para tentaculitoideos, encerrando assim a deposição das litologias aflorantes da Formação Ponta Grossa entre o Emsiano e o Frasniano.