Search

The distribution of some Scarabaeoidea of the Iranian fauna is specified. Holochelus (Miltotrogus) brenskei (Reitter, 1888), Holochelus (Miltotrogus) rusticus (Faldermann, 1835), Hoplia (s. str.) caucasica Kolenati, 1846, and Ochodaeus seleuciensis Petrovitz, 1963, are recorded from Iran for the first time. Two new species of Rhizotrogini belonging to the genera Lasiexis Semenov & Medvedev, 1936, and Holochelus Reitter, 1889, are described from Iran, illustrated and compared to their related species. Lasiexis glabrithoracica Keith, 2003 is synonymised with Lasiexis persicola Petrovitz, 1980., La répartition de quelques espèces de Scarabaeoidea de la faune iranienne est précisée. Holochelus (Miltotrogus) brenskei (Reitter, 1888), Holochelus (Miltotrogus) rusticus (Faldermann, 1835), Hoplia (s. str.) caucasica Kolenati, 1846, et Ochodaeus seleuciensis Petrovitz, 1963, sont cités d’Iran pour la première fois. Deux nouvelles espèces de Rhizotrogini appartenant aux genres Lasiexis Semenov & Medvedev, 1936, et Holochelus Reitter, 1889, sont décrites d’Iran, illustrées et comparées aux espèces proches. Lasiexis glabrithoracica Keith, 2003 est mis en synonymie avec Lasiexis persicola Petrovitz, 1980.

The distribution of some Scarabaeoidea of the Iranian fauna is specified. Holochelus (Miltotrogus) brenskei (Reitter, 1888), Holochelus (Miltotrogus) rusticus (Faldermann, 1835), Hoplia (s. str.) caucasica Kolenati, 1846, and Ochodaeus seleuciensis Petrovitz, 1963, are recorded from Iran for the first time. Two new species of Rhizotrogini belonging to the genera Lasiexis Semenov & Medvedev, 1936, and Holochelus Reitter, 1889, are described from Iran, illustrated and compared to their related species. Lasiexis glabrithoracica Keith, 2003 is synonymised with Lasiexis persicola Petrovitz, 1980., La répartition de quelques espèces de Scarabaeoidea de la faune iranienne est précisée. Holochelus (Miltotrogus) brenskei (Reitter, 1888), Holochelus (Miltotrogus) rusticus (Faldermann, 1835), Hoplia (s. str.) caucasica Kolenati, 1846, et Ochodaeus seleuciensis Petrovitz, 1963, sont cités d’Iran pour la première fois. Deux nouvelles espèces de Rhizotrogini appartenant aux genres Lasiexis Semenov & Medvedev, 1936, et Holochelus Reitter, 1889, sont décrites d’Iran, illustrées et comparées aux espèces proches. Lasiexis glabrithoracica Keith, 2003 est mis en synonymie avec Lasiexis persicola Petrovitz, 1980.

A new species of the genus Amphimallon Latreille is described from Greece (Sifnos Island), illustrated and compared with its most related species., Une nouvelle espèce du genre Amphimallon Latreille est décrite de Grèce (Ile de Sifnos), illustrée et comparée aux espèces proches.

International audience

Haplidia pilicollis temperei Baraud, 1988 Haplidia pilicollis temperei Baraud, 1988: 571; Montreuil, 2005: 131 (catalogue); Keith & Lo Cascio, 2005: 142 (catalogue). Aplidia petrovitzi: Chikatunov & Pavl��ček, 1997: 54 (catalogue). Material examined. LEBANON: 1 male, Caza Bcharre, 900 m, VI.2005, A. Kairouz leg. (RSCV). Remark. This specimen was collected together with Haplidia joannis. Distribution. This taxon was described from Lebanon (Saida) and is currently recorded also from Syria (Bezděk 2016). Distribution in Lebanon: see Fig. 4., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 543, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672","Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131.","Keith, D. & Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142.","Chikatunov, V. & Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65.","Bezdek, A. (2016) Rhizotrogini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, Boston, pp. 249 - 280."]}

International audience

[Haplidia baudii Kraatz, 1882] Haplidia baudii Kraatz, 1882: 38; Reitter, 1902: 187 (key to species); Baraud, 1988: 567 (new data, key to species); Montreuil, 2005: 131 (catalogue); Keith & Lo Cascio, 2005: 141 (catalogue). Aplidia baudii: Chikatunov & Pavl��ček, 1997: 53. Remarks. Chikatunov & Pavl��ček (1997) recorded Aplidia baudii from Lebanon, after El-Hariri (1971), and reminded old records from Syria. This species was described from Cyprus and Baraud (1988) recorded it from Rhodos. Bezděk (2016) did not give credit to the records of this species from the continent. Miessen (2016) and Sehnal (in press), consider the synonymy between Haplidia baudii (Cyprus, patria falsa) and Haplidia hirticollis Burmeister, 1855, from southern Italy. This species does not belong to the Lebanese fauna., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 544, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202","Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen fauna und angrenzenden landern, enthaltend die gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn, 40, 93 - 303. [1901]","Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672","Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131.","Keith, D. & Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142.","Chikatunov, V. & Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65.","El-Hariri, G. (1971) A list of recorded insect fauna of Syria. Vol. 2. University of Aleppo, Faculty of Agriculture, Aleppo, 306 pp.","Bezdek, A. (2016) Rhizotrogini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, Boston, pp. 249 - 280.","Miessen, G. (2016) Essais sur le genre Aplidia Hope 1837 a Chypre et commentaires sur quelques especes limitrophes (Coleoptera, Scarabaeoidea, Melolonthidae, Rhizotrogini). Entomologie Faunistique-Faunistic Entomology, 69, 57 - 73.","Burmeister, H. C. C. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin, x + 569 pp."]}

Haplidia joannis Baraud, 1988 Haplidia joannis Baraud, 1988: 571; Keith & Lo Cascio, 2005: 141 (catalogue). Aplidia joannis: Chikatunov & Pavl��ček, 1997: 54 (catalogue). Material Examined. LEBANON: 1 specimen, Zeghrine, V.2001, C. Maalouf leg. (AUB); 1 specimen, Zebdine, VII.2000 (ULB); 1 specimen, Jbeil, Qartaba, VII.2001, C. Atallah leg. (ULB); 1 specimen, Kersrouane, Ghineh, 3.VII.2008, R. Chedid leg. (ULB); 3 males, 2 females, Caza Bcharre, Barhalioun, Chira, 700-800 m, VI.2009, A. Kairouz leg. (CMU); 2 males, Caza Bcharre, 900 m, VI.2005, A. Kairouz leg. (RSCV). SYRIA: 2 males, Buq���ata, 01.VIII.2002, V. Krachenko & V. Chikatunov leg. (CGS); 1 male, 1 female, Golan Heights, Odem forest, 22.VII.2009, E. Orbach leg. (CGS). Distribution. This species was described from Lebanon (Achkout, Jounieh) (Fig. 4) and Syria (Damascus). Remarks. See under H. petrovitzi., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 541, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672","Keith, D. & Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142.","Chikatunov, V. & Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65."]}

Haplidia tarsensis Kraatz, 1882 Figs. 3 A���O Haplidia tarsensis Kraatz, 1882: 37; Reitter, 1902: 185 (new data, key to species); Dalla Torre, 1912: 210 (catalogue); Baraud, 1988: 564 (new data, key to species); Montreuil, 2005: 131 (new data); Keith & Lo Cascio, 2005: 142 (catalogue). Material examined. LEBANON: 4 males, Bcharre, North Lebanon, Alt. 1300 m, JULY 2011, Andre Kairouz lgt. (NMPC, RSCV); 1 male, Tannourine El Faouka, Casa El Batroun 1420 m, Aout 2011, A. Kairouz Nord Liban (COM); 2 males, 1 female, Bahn��s, VI.2004, O. Montreuil leg. (COM); 1 male, Baskinta (Metn), 3.IV.2005, M. Moubarak leg. (ULB); 1 male, 2 females, Aarsal, 09.VII.1997 (NHM, CMU); 5 males, 1 female, North. Gov., Tannourine, 2 Km N Harissa, Cedars Nat. Res., 1750 m, 34��12���34���N, 35��55���45���E, 18.VI.2016, A. Kotan leg. (NHM, CMU); 1 male, Tannourine, 23.III.2001 N. Nemer leg. (HNHM); 11 males, 2 females, Jbeil, Laqlouq, 1300 m, 34��09���N 35��51���E, 02.VII.2009, A. Kairouz leg. (CGS); 5 males, 1 female, same data, 17.VII.2009 (CGS); 8 ex, Bcharre, 14.VI.1998, D. Baiocchi leg. (CGS); 1 male, Mtein, Matn, 33��53���N 35��44���E, V.2009 (CGS). SYRIA: 2 males, Damas (CGS). TURKEY: 1 male, 1 female, prov. Mersin, Taurus (city) 50 m, 36.917, 34.896, 29.4.2018 (CMU); 4 males, Gaziantep, 30km NW, 7.6.2001, Sn��žek leg. (RSCV); 5 males, 3 females, Iskanderun, 10. 6. 1999, R. Sehnal leg. (RSCV); 30 males, 15 females, Aslani, 35 km N of Erdemli, 6.-14.VI.1996, Martin Johanides (RSCV); 5 males, 5 females, Mersin, Kuzucubelen, 1.6.2001, M. ����rovec leg. (RSCV); 1 male, Taurus, Erdemli, 25.V.1995 R. Reitmaier lgt. (RSCV); 2 males, 2 females, Erdemli - Aslanli, 4.-8. VI. 1996. P. Viktora lgt. (CPV). Remarks. Study of 108 specimens referable to Haplidia tarsensis from the full span of its geographic distribution (Turkey, Lebanon, Syria) enabled us to critically evaluate the characters that have been used in the keys to species (Baraud 1988). Our observations show that characters such as setation and punctation of the head, pronotum and elytra, the shape of paramere apex, and the shape of sclerites in the endophallus are markedly variable, which could lead to difficulties in routine identification of specimens. Such situation can also cast some doubts on the current taxonomic treatment of such populations. However, in specimens of H. tarsensis collected at the same time (IV. 1996) in Erdemli, Turkey, all the said characters were found variable. Setation of the pronotum and humeral area of the elytra varied from none (due to wear) to short and recumbent (Figs. 3 A���E), punctation of the pronotum varied from fine and even to coarse and uneven (Figs. 3 A���E), and the shape of the parameres (Figs. 3 F���J) and of the sclerites in the endophallus (Figs. 3 K���O) were also considerably variable. A similar variability was also present in other populations from Turkey, Lebanon and Syria. We therefore regard all the examined populations as belonging to a single species, H. tarsensis. These findings suggest that the variability of conditions usually regarded as diagnostic should be scored across the entire genus, similarly to what has recently been done for the Cypriote fauna of Haplidia by Miessen (2016), resulting in the identification of two new synonymies. It would of course be ideal to combine the traditional morphological approach with molecular analyses, but we are not aware of any molecular study of Haplidia in progress at the present time. Distribution. Lebanon (Fig. 4), Turkey (Tarsus, type locality), Syria., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 543, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202","Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen fauna und angrenzenden landern, enthaltend die gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn, 40, 93 - 303. [1901]","Dalla Torre, K. W. von. (1912) Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. & Schenkling S. (Eds): Coleopterorum Catalogus. Vol. XX. W. Junk, Berlin, pp. 135 - 290.","Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672","Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131.","Keith, D. & Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142.","Miessen, G. (2016) Essais sur le genre Aplidia Hope 1837 a Chypre et commentaires sur quelques especes limitrophes (Coleoptera, Scarabaeoidea, Melolonthidae, Rhizotrogini). Entomologie Faunistique-Faunistic Entomology, 69, 57 - 73."]}

Haplidia chaifensis Kraatz, 1882 Haplidia chaifensis Kraatz, 1882: 38; Reitter, 1902: 187 (key to species); Dalla Torre, 1912: 210 (catalogue); Baraud, 1988: 577 (catalogue, key to species); Keith & Lo Cascio, 2005: 142 (catalogue). Aplidia chaifensis: Chikatunov & Pavl��ček, 1997: 53 (new data); Mienis, 2017: (new data). Material examined. LEBANON: 1 male, Ayta El Jabal, V.2002, R. Abadi leg. (AUB). Distribution. This species, described from Israel (Haifa), is recorded from Lebanon (Fig. 4). for the first time (first country record), on the base of a single male found in a place close to the border with Israel. This species is morphologically very close to H. villosicollis., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 544, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202","Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen fauna und angrenzenden landern, enthaltend die gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn, 40, 93 - 303. [1901]","Dalla Torre, K. W. von. (1912) Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. & Schenkling S. (Eds): Coleopterorum Catalogus. Vol. XX. W. Junk, Berlin, pp. 135 - 290.","Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672","Keith, D. & Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142.","Chikatunov, V. & Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65.","Mienis, H. K. (2017) Activity pattern of the Cockchafer Aplidia chaifensis in Kibbutz Netzer Sereni, Israel, in 2017. Natuurhistorische en Andere Notities, 14, 3 - 4."]}

Haplidia houryae Montreuil, 2005 Haplidia houryae Montreuil, 2005: 130. Material examined. LEBANON: 1 male, Mansourieh, IV.2004, E. Abi Khalil leg. (AUB); 1 male, Aramoun, 33��45���N 35��31���E, V.2009 (CGS); 1 male, Beiruth, AUB, 26.V.2009 (CGS); 1 male, Caza du Chouf, Deir el Kamar, 900 m, 27-30.V.1972, P. Brignoli leg. (CGS). Distribution. Species endemic to Lebanon (Fig. 4). It was described from Hermel and Bchamoun., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 539, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131."]}

[Haplidia endroedii Baraud, 1988] Haplidia endroedii Baraud, 1988: 576; Montreuil, 2005: 131 (misidentification); Keith & Lo Cascio, 2005: 142 (catalogue). Aplidia endroedii: Chikatunov & Pavl��ček, 1997: 53 (catalogue). Remarks. Haplidia endroedii was described from Jordan (Wadi Sir, type locality) and recorded from Lebanon by Montreuil (2005) on the base of a specimen from Bziza. The re-examination of this specimen invalidates its former identification: it is in fact a male of Haplidia karlaeschneiderae Sehnal, Montreuil & Uliana, new species. As no other data attests the presence of H. endroedii in Lebanon, this species does not seem to occur in Lebanon and should be removed from the list of Haplidia occurring in this country. Haplidia endroedii is recorded from Israel and Jordan., Published as part of Montreuil, Olivier, Uliana, Marco & Sehnal, Richard, 2020, The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini), pp. 536-548 in Zootaxa 4718 (4) on page 544, DOI: 10.11646/zootaxa.4718.4.7, http://zenodo.org/record/3602746, {"references":["Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672","Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131.","Keith, D. & Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142.","Chikatunov, V. & Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65."]}

Genus Trigonochilus Brenske, 1896 Trigonochilus Brenske, 1896: 33. Suralcis Machatschke, 1970: 158, junior objective synonym Type species: Trigonochilus coriaceus Brenske, 1896 , by monotypy. Machatschke (1970) introduced the name Suralcis to replace Trigonochilus Brenske, 1896, in reason of the homonymy of the later with Trigonochilus Agassiz, 1846 (Gyrinidae), an unjustified emendation of Trigonocheilus Dejean, 1833. However, Trigonocheilus Dejean is a nomen nudum (Bousquet & Bouchard 2013) as it does not include any valid species name: the only included name ��� rostratus De Hann��� was not published. Thus, Trigonochilus Agassiz, 1846 is also a nomen nudum as it doesn���t include any available species name (Gustafson & Miller 2013). For these reasons, replacement of Trigonochilus Brenske, 1896 is superfluous, and Suralcis Machatschke, 1970 is a synonym of Trigonochilus Brenske., Published as part of Montreuil, Olivier & Frolov, Andrey V., 2019, A review of the Malagasy species of the genus Trigonochilus Brenske (Coleoptera Scarabaeidae: Rutelinae: Adoretini) with description of a new species, pp. 596-600 in Zootaxa 4679 (3) on page 596, DOI: 10.11646/zootaxa.4679.3.12, http://zenodo.org/record/3471116, {"references":["Brenske, E. (1896) Trigonochilus n. g. Vertreter einer neuen Ruteliden-Gruppe aus der Verwandschaft der Anoplognathiden. Entomologische Nachrichten, 22, 33 - 37.","Machatschke, J. W. (1970) Rutelina (Col., Scarabaeidae): synonymische Bemerkungen. Entomologist's Monthly Magazine, 105, 157 - 158. [1969]","Agassiz, L. (1846) Nomenclatoris Zoologici. Index Universalis, continens Nomina Systematica Classum, Ordinum, Familiarum et Generum Animalium Omnium. Jent et Gassmann, Soloduri, 1135 pp.","Bousquet, Y. & Bouchard, P. (2013) The genera in the second catalogue (1833 - 1836) of Dejean's Coleoptera collection. ZooK- eys, 282, 1 - 219. https: // doi. org / 10.3897 / zookeys. 282.4401","Gustafson G. T. & Miller K. B. (2013) On the family- and genus-series nomina in Gyrinidae Latreille, 1810 (Coleoptera, Adephaga). Zootaxa, 3731 (1), 77 - 105. https: // doi. org / 10.11646 / zootaxa. 3731.1.3"]}

Hybalus punicus Baraud, 1991 Hybalus punicus Baraud, 1991: 22. The species is only known from the type series collected in Fondouk-Djedid (Fig. 2)., Published as part of Frolov, Andrey, Akhmetova, Lilia & Montreuil, Olivier, 2019, A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species, pp. 290-294 in Zootaxa 4671 (2) on page 293, DOI: 10.11646/zootaxa.4671.2.10, http://zenodo.org/record/3442338, {"references":["Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28."]}

Frolov, Andrey, Akhmetova, Lilia, Montreuil, Olivier (2019): A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species. Zootaxa 4671 (2): 290-294, DOI: https://doi.org/10.11646/zootaxa.4671.2.10

[Hybalus bigibber Reitter, 1892] Hybalus bigibber Reitter, 1892e: 260; Normand 1936: 376; Baraud 1991: 14. This species was described from Sicily and recorded from Tunisia by Normand (1936) (Ghardimaou and Souk El Arba [Jendouba]), although the later author was unsure about the identification of the Hybalus specimens he studied. The record from Jendouba probably belongs to H. algiricus. We don���t know any specimen to confirm the presence of this species on Tunisian territory., Published as part of Frolov, Andrey, Akhmetova, Lilia & Montreuil, Olivier, 2019, A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species, pp. 290-294 in Zootaxa 4671 (2) on page 293, DOI: 10.11646/zootaxa.4671.2.10, http://zenodo.org/record/3442338, {"references":["Normand, H. (1936) Contribution au Catalogue des Coleopteres de la Tunisie. Fascicule 10. Bulletin de la Societe d'Histoire naturelle de l'Afrique du Nord, 27, 355 - 383.","Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28."]}

Hybalus normandi Baraud, 1980 Hybalus normandi Baraud, 1980: 60; 1991: 20. This species was described from Tamera (Baraud 1980) and later recorded also from Cap Bon (Baraud 1991) (Fig. 2)., Published as part of Frolov, Andrey, Akhmetova, Lilia & Montreuil, Olivier, 2019, A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species, pp. 290-294 in Zootaxa 4671 (2) on page 293, DOI: 10.11646/zootaxa.4671.2.10, http://zenodo.org/record/3442338, {"references":["Baraud, J. (1980) Les Hybalus Brulle de Tunisie. Nouvelle Revue d'Entomologie, 10, 59 - 62.","Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28."]}

[Hybalus parvicornis Lucas, 1855] Hybalus parvicornis Lucas, 1855: 556; Normand 1936: 376; Baraud 1991: 21. This species is known from a few localities in Algeria. It was recorded from Tunisia by Normand (1936) but he mentioned that the identification was doubtful. We don���t know any specimens to confirm the presence of this species on Tunisian territory., Published as part of Frolov, Andrey, Akhmetova, Lilia & Montreuil, Olivier, 2019, A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species, pp. 290-294 in Zootaxa 4671 (2) on page 293, DOI: 10.11646/zootaxa.4671.2.10, http://zenodo.org/record/3442338, {"references":["Normand, H. (1936) Contribution au Catalogue des Coleopteres de la Tunisie. Fascicule 10. Bulletin de la Societe d'Histoire naturelle de l'Afrique du Nord, 27, 355 - 383.","Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28."]}

Hybalus algiricus Petrovitz, 1968 Hybalus algiricus Petrovitz, 1968: 6; Baraud, 1991: 11. Material examined. Tunisia: Jendouba, El Feija National Park, N36��27��48�� E08 14��11��, 590 m, 3.III.2012, W. Ziegler leg., 1 male (ZIN), 2 males (CWZ); Sillana Province, N Kesra, 8.III.2012, N35��51��20�� E9��23��41��, 1000m, W. Ziegler leg., 1 male (CWZ); Beja, 11 km of Teboursouk Pass, N36��29��48�� E09��09��12��, 1.III.2012, Meybohm leg., 1 male (CWZ); 2 males MNHN La Kesra Tunisie De Vauloger. This species was described from Algeria (without more precise locality) and known from the single type specimen. It is recorded from Tunisia for the first time (Fig. 2)., Published as part of Frolov, Andrey, Akhmetova, Lilia & Montreuil, Olivier, 2019, A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species, pp. 290-294 in Zootaxa 4671 (2) on page 293, DOI: 10.11646/zootaxa.4671.2.10, http://zenodo.org/record/3442338, {"references":["Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28."]}