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Premise Methods Results Conclusions Ceanothus (Rhamnaceae) is a large genus of shrubs that dominate California chaparral and are resilient to fires. Persistence is ensured by resprouting and/or seedling recruitment from dormant seed banks. Some species do both and others, the obligate seeders, are entirely dependent on seedling recruitment. The distribution of these two modes within the genus is poorly documented.We used all available publications that document species responses to fire and filled most gaps in the literature based on extensive field studies of more than 60 recent wildfires in California.The genus is divided into two subgenera, Ceanothus and Cerastes. Ceanothus is widely considered to comprise mostly resprouting species and Cerastes to consist of only obligate seeders. The subgenus Ceanothus includes resprouting species throughout their range from the eastern United States and Midwest to western United States. Within the California Floristic Province (CFP), a few species are unique in producing massive lignotubers that develop from repeated fires; however, within the CFP, the majority of species in this subgenus do not resprout and are obligate seeders. Two have disjunct subspecies that are facultative seeders or obligate seeders.Previously, speciation in this genus was contended to have occurred in the late Miocene within the CFP. The syndrome of obligate seeding is most strongly represented in this region, and we hypothesize that evolution of this syndrome was a response to increased predictability of fire driven by the Mediterranean climate and the long interval between fires. [ABSTRACT FROM AUTHOR]

This study was conducted on the distribution of manganese content in the soils of the city of Mitrovica and its surroundings in Kosovo. A total of 156 topsoil samples were taken in the entire study region (sampling grid of 1.4 × 1.4 km2). The average Mn content of the entire study area is 1600 mg/kg (range 170–17000 mg/kg), which is 4.2 times higher than the EU average and 3.3 times higher than the global average. The spatial distribution maps show that the higher Mn contents occur mainly in soils from Miocene latite as well as quartz-latite and from Triassic volcanic-sedimentary formations in the eastern and north-eastern part of the study area. The lower values were found in the Flysch and Cretaceous flysch outcrops. It was found that the increased Mn contents in soils from this area are mostly of lithogenic origin. The values of the enrichment factor (EF) show that the soils of the entire study area are very highly enriched with Mn (20 < EF < 40). The values of the geo-accumulation index (1< I-geo ≤ 2), show that the soils of the entire study area are moderately contaminated with Mn. [ABSTRACT FROM AUTHOR]

ABSTRACT This response addresses the comments made by Saraswati et al. on our recent publication in Geological Journal. In our study, we used a multi‐analytical screening approach, including visual, XRD and trace element analysis, to evaluate diagenetic alteration of carbonate shell samples. Contrary to the claims made by Saraswati et al. our findings are based on well‐preserved samples with minimal alteration. The Sr‐isotope data from the Khari Nadi Formation (23.07–18.09 Ma) are largely consistent with existing biostratigraphic constraints, but suggest a possible extension of the upper age limit into the Burdigalian. Our Sr‐isotope ages for the Chhasra Formation (15.11–12.29 Ma) are younger than those suggested previously, based on larger benthic foraminifers, but largely corroborate recent nannoplankton data. As a result of the new age data from Kutch, we suggest that the sedimentation rates, derived from multiple samples, increased significantly from the Oligocene to Miocene. We assert that ‘established’ ages are also subject to refinements as the rock units become amenable to new dating techniques. We welcome any criticism that is constructive, and remain open to any further data that could refine or challenge our interpretations. [ABSTRACT FROM AUTHOR]

ABSTRACT Characterising the spatial and temporal distribution of the S‐N‐trending rift in southern Tibet is crucial for elucidating the dynamics of E‐W extension within the Tibetan Plateau since the Miocene. The Kung Co–Tangra Yumco rift, located in the central part of the rift system, was initiated at the peak of rifting development. While the initiation of rifting has been ascertained through low‐temperature thermochronology, direct timing constraints remain absent for the Gangga Graben in the southernmost region. Utilising the quartz electron spin resonance (ESR) dating technique, we ascertained the onset age of the eastern boundary fault of the Gangga Graben to be 12.00 ± 1.80 Ma, with an accelerated activity phase at 9.17 ± 1.19 Ma, and an activity age for the western boundary fault at 2.22 ± 0.24 Ma. U‐series dating conducted on the western boundary disclosed a time span of 20–13 ka for hot spring fissure activity. Seismic evidence, inclusive of those from the Gongdapu Horst, indicates persistent activity of the Gangga Graben since the Pleistocene. Comparative analysis of age data from the Kung Co–Tangra Yumco rift suggests that it commenced almost synchronously along its strike at ~14.5–12 Ma. Considering the rifts trending from west to east in southern Tibet and the genetic mechanism of leucogranite, we propose an E‐W extension mechanism in southern Tibet. At ~26–14 Ma, the lithosphere experienced weakened delamination and asthenosphere upwelling, leading to a series of magmatic activities and onset of E‐W extension in southern Tibet. At ~17–7 Ma, as the basal shearing of the underthrusting Indian Plate increased, it marked the peak phase of rifting in southern Tibet. [ABSTRACT FROM AUTHOR]

As a passive rift basin in the Cenozoic, the formation and evolution process of the Weihe Basin has not been well explained. Here we estimate the stretching factor and basal heat flow for each rift period along four geological profiles using a 2D multi-stage finite extension model. The modelling is based on a reconstruction of pre-rift crustal and lithospheric thicknesses, and is calibrated with the current heat flow. The Xi'an Depression had the biggest stretching factor while the Pucheng Uplift had the smallest throughout the Cenozoic period, which saw a range of stretching factors from 1.02 to 2.06. The basal heat flow in the Weihe Basin exhibits a corresponding change. At the end of the Oligocene, Miocene, and Pliocene periods, the basal heat flow varied between 58.5 ~ 64.0 mW·m−2, 59.2 ~ 67.0 mW·m−2, and 59.6 ~ 78.7 mW·m−2, respectively. Increasing stretching factor and basal heat flow indicate that the Weihe Basin is currently in a state of sustained extension. The formation and development of the Weihe Basin are analysed in the context of larger tectonic changes, and it is considered that the Weihe Basin is controlled by pre-existing faults, and formed in a regional tectonic context of sliding and pulling apart first, and tensioning and lifting up later. Since the late Miocene (7.3 Ma), anomalously rapid subsidence has occurred, induced by a new phase of rifting and depth-related lithospheric extension. The thermal genesis of the geothermal system in the Weihe Basin is also thought to be closely related to the latest phase of extension. [ABSTRACT FROM AUTHOR]

Myotragus balearicus is the terminal species of a Caprinae lineage that inhabited the Balearic Islands since the Messinian. Myotragus balearicus had a relatively small brain, reported to be half the size of a contemporary bovid of similar size. This study presents a reassessment of its brain size and anatomy. We demonstrate that the brain of M. balearicus is only 17% smaller when compared to Late Miocene taxa. Since the ancestors of M. balearicus were isolated on the Balearics in the Late Miocene, we deduce that the small brain size of M. balearicus is a feature largely retained from its continental ancestor and only partially the result of geographic isolation. The reduction in brain size can be attributed to reduction of the occipital region. We hypothesise that neuroanatomical changes during the evolutionary process of dwarfism were likely influenced by the absence of ecologically relevant predators. [ABSTRACT FROM AUTHOR]

New post-cranial remains from the early Late Miocene have been discovered in the lowermost part of the Irrawaddy Formation, Tebingan area, central Myanmar. Three genera and one indeterminate taxon were identified: Rhinoceros sp. Rhinoceros cf. R. sondaicus, Dicerorhinus sp., Brachypotherium perimense, and Rhinocerotidae indet. The evolutionary history of the Rhinocerotidae is still poorly known in Southeast Asia. Few Rhinoceros species, Rhinoceros sp. ‘B’. fatehjangense, and B. perimense, has already been identified in the Tebingan area by dental remains. The present discovery of the post-cranial remains of Rhinoceros cf. R. sondaicus and Dicerorhinus sp. that complements previous studies based on dental remains. The early Late Miocene Tebingan record is the oldest fossil record of Dicerorhinus in Southeast Asia, indicating a continental origin of the genus. Furthermore, the possible presence of R. sondaicus in the Tebingan area suggests that it may have appeared in Southeast Asia as early as the early Late Miocene. [ABSTRACT FROM AUTHOR]

Dolichandrone spathacea is a mangrove associate with high medicinal and ecological values. However, due to the dual-pressure of climate change and human activities, D. spathacea has become endangered in China. Moreover, misidentification between D. spathacea and its terrestrial relative D. cauda-felina poses further challenges to field protection and proper medicinal usage of D. spathacea. Thus, to address these problems, we sequenced and assembled mitochondrial (mt) and chloroplast (cp) genomes for both D. spathacea and D. cauda-felina. Comparative analysis revealed apparently different size and scaffold number between the two mt genomes, but a high similarity between the cp genomes. Eight regions with high sequence divergence were identified between the two cp genomes, which might be used for developing candidate DNA markers for distinguishing the two species. The splitting between D. spathacea and D. cauda-felina was inferred to occur at ~6.8 - 7.7 million years ago (Mya), which may be driven by the environment fluctuations in late Miocene. In the cp genome, 12 genes related to the expression of photosynthesis-associated proteins were detected with signatures of positive selection, which may contribute to the origin and evolutionary adaptation of Dolichandrone mangrove species. These new findings do not only enrich organelle genomic resources of Dolichandrone species, but also provide important genetic clues for improving the conservation and proper usage of endangered mangrove associate D. spathacea. [ABSTRACT FROM AUTHOR]

A new fossil epifoliar ascomycete (Ascomycota), Meliolinites miocenicus nov. sp. (fossil Meliolaceae), occurs in situ on a compressed monocot leaf from middle Siwalik (Late Miocene, ca. 12–8 Ma) sediments of Himachal Pradesh, Western Himalaya. The fossil consists of a well-preserved mycelium of superficial, brown to dark brown, septate, thick-walled, branched hyphodiate hyphae; a sub-globose, dark brown putative ascoma, and an oblong to broadly cylindrical, five-celled, four-septate, mature germinating ascospore. The new fossil differs from earlier reported melioloid fossils primarily in the morphology of appressoria. Meliolinites miocenicus nov. sp. on its host is evidence of the existence of a biotrophic relationship at the time of deposition. Qualitative climate data using plant megafossils recovered from the same fossil locality indicate that M. miocenicus nov. sp. and its host thrived in a warm and humid tropical environment. [ABSTRACT FROM AUTHOR]

Prosantorhinus is a genus of small extinct teleoceratine rhinoceroses with shortened limb bones, widely distributed in Europe. However, the Asian evolution of the teleoceratine Prosantorhinus has remained unclear because of a scarcity of fossil records of the genus. Here, we report the first record of Prosantorhinus in East Asia from the Middle Miocene of Tongxin, Ningxia. The new specimen is characterized by a concave dorsal skull profile and elevated nasals; short and stout nasal bones with drooping margins on both sides; the thickened, enlarged, and roughened nasal extremity supporting a small horn; the semi-molarized upper premolars with a lingual bridge between the protocone and hypocone; the metaloph constriction present on P2–4; the protocone equal to the hypocone on P2; the crista present on P3; and the cement on the cheek teeth developed. With all the morphological evidence considered, we establish a new species, Prosantorhinus yei sp. nov.. A phylogenetic analysis based on 282 morphological characters scored for 36 taxa reveals that Prosantorhinus yei sp. nov. is a relatively derived taxon in the genus. We hypothesize that Prosantorhinus yei sp. nov. lived in relatively moist environments. [ABSTRACT FROM AUTHOR]

We here present a comprehensive integrative taxonomic review of the genus Hylomys , using molecular (mitochondrial genomes and up to five nuclear loci) and morphological data from museum specimens across its distribution, resulting in the description of two new species and the elevation of three subspecies to specific status. This revision significantly increases the known diversity of Hylomys from two to seven extant species, challenging the traditional view of species-level diversity within gymnures. We discuss the implications of the taxonomic findings for conservation, particularly in relation to the restricted distribution ranges of several species that may be threatened by habitat loss and/or climate change. Our research emphasizes the importance of scientific collections and underscores the potential of museum genomics and additional field sampling to identify new species and improve our understanding of species diversity in poorly studied regions. Speciation events within Hylomys occurred during the Late Miocene and Early Pliocene, possibly driven by shifting climate conditions such as the strengthening of the Indian monsoon and the expansion of seasonally dry conditions. This study supports northern Sumatra and the southern Annamites as centres of localized endemicity and suggests the need for additional small mammal surveys across Sumatra's Barisan Range. [ABSTRACT FROM AUTHOR]

The Neotropical region is the most diverse on the planet, largely owing to its mosaic of tropical rainforests. Multiple tectonic and climatic processes have been hypothesized to contribute to generating this diversity, including Andean orogeny, the closure of the Isthmus of Panama, the GAARlandia land bridge and historical connections among currently isolated forests. Micrathena spiders are diverse and widespread in the region, and thus a complete phylogeny of this genus allows the testing of hypotheses at multiple scales. We estimated a complete, dated phylogeny using morphological data for 117 Micrathena species and molecular data of up to five genes for a subset of 79 species. Employing event‐based approaches and biogeographic stochastic mapping while considering phylogenetic uncertainty, we estimated ancestral distributions, the timing and direction of dispersal events and diversification rates among areas. The phylogeny is generally robust, with uncertainty in the position of some of the species lacking sequences. Micrathena started diversifying around 25 Ma. Andean cloud forests show the highest in‐situ speciation, while the Amazon is the major dispersal source for adjacent areas. The Dry Diagonal generated few species and is a sink of diversity. Species exchange between Central and South America involved approximately 23 dispersal events and started ~20 Ma, which is consistent with a Miocene age for the Isthmus of Panama closure. We inferred four dispersal events from Central America to the Antilles in the last 20 Myr, indicating the spiders did not reach the islands through the GAARlandia land bridge. We identified important species exchange routes among the Amazon, Andean cloud forests and Atlantic forests during the Plio‐Pleistocene. Sampling all species of the genus was fundamental to the conclusions above, especially in identifying the Andean forests as the area that generated the majority of species. This highlights the importance of complete taxonomic sampling in biogeographic studies. [ABSTRACT FROM AUTHOR]

The Amal oil field is located in the southern part of the Gulf of Suez and produces hydrocarbons mainly from middle Miocene reservoirs. It is marked by uncertainties in the southern part of the Gulf of Suez province because of the structural complexity, lateral facies change, different lithologies, and diverse reservoir quality. Therefore, in this approach, different reservoir properties of the Hammam Faraun Member of the Belayim Formation, the Markha Member of the Kareem Formation, and the Upper Rudies Member are obtained and evaluated. The main reservoirs are composed of sand and shale intercalations. The wireline logging data (gamma ray, density, neutron, sonic, and resistivity logs) of four wells were used mainly for the petrophysical analyses of hydrocarbon reservoirs. The three most important parameters in petrophysical evaluation: shale content, porosity, and fluid saturation are calculated for the main reservoirs to construct the litho-saturation model for each well. Each of these parameters is mapped to study their areal variation across the study area and estimate the appropriate locations for the new development wells. Fluid saturations are estimated based on the Indonesian formula due to shale presence. The study exhibited that Hammam Faraun, Markha, and Upper Rudies Formations have porosity of 0.15-0.17, 0.15-0.23, 0.12-0.27; shale volume of 0.3-0.36, 0.12-0.51, 0.19-0.37; and water saturation of 0.34-0.6, 0.33-0.51, 0.48-0.6, respectively. The results show an increasing trend toward the central part in porosity, sandy facies, and oil saturation. Based on the petrophysical evaluation and the mapping, the central part of the field is very promising for development and production activities. [ABSTRACT FROM AUTHOR]

Scientific ocean drilling cores recovered years ago (legacy cores), especially as recovered by rotary drilling, commonly show incomplete recovery and core disturbance. We present a novel method to date such cores by presenting the first high-precision U-Pb zircon ages targeting the duration of the Miocene Climate Optimum (MCO; ca. 17-14 Ma) from volcanic ashes at Ocean Drilling Program Site 1000 (on the Nicaragua Rise in the Caribbean Sea). We place these ages within a newly developed framework to address incomplete core recovery and use them to calibrate a high-resolution bulk carbonate δ13C and δ18O record. Our Site 1000 ages show that volcanism of the Columbia River Basalt Group (CRBG) large igneous province was coincident with the interval of greatest sustained MCO warmth at this site. However, if the CRBG were the primary driver of the MCO, our chronology may allow for outgassing preceding volcanism as a major source of CO2. We thus document a promising new way to obtain highly resolved, accurate, and precise numerical age models for legacy deep-sea sediment cores that does not depend on correlation to other records. [ABSTRACT FROM AUTHOR]

Premise: Molecular studies based on chloroplast markers have questioned the monophyly of the fern genus Pecluma (Polypodioideae, Polypodiaceae), which has several species of Polypodium nested within it. We explored the delimitation of Pecluma and its biogeographic pattern by evaluating the phylogenetic position of four Polypodium species not sequenced thus far and integrating the first fossil evidence of Pecluma. Methods: Using herbarium material, we applied a genome‐skimming approach to obtain a phylogenetic hypothesis of Polypodioideae; assessed the combination of character states observed in the fossil from Miocene Dominican amber using a previously published phylogeny of Polypodioideae based on four plastid markers as framework; calculated divergence times; and conducted an ancestral area estimation. Results: Within Polypodioideae, Pecluma was recovered as sister to Phlebodium. Three of the newly sequenced species—Polypodium otites, P. pinnatissimum, and P. ursipes—were recovered with maximum support within the Pecluma clade, whereas P. christensenii remained within Polypodium. The closest combination of character states of the fossil was found within Pecluma. Our biogeographic analyses suggest an Eocene origin of the genus in South America, with several subsequent Oligocene and Miocene colonization events to Mexico–Central America and to the West Indies. Conclusions: Although the circumscription of Pecluma is still challenging, our results elucidate the origin and age of the genus. The newly described fossil, Pecluma hispaniolae sp. nov., supports the hypothesis that the epiphytic communities of the Greater Antilles exhibit a constant generic composition since the Miocene. We propose new combinations (Pecluma otites, Pecluma pinnatissima, and Pecluma ursipes) to accommodate three species previously classified in Polypodium. [ABSTRACT FROM AUTHOR]

The palynological analysis of the Paleogene and Neogene sequences of the southern part of West Siberia (borehole 2, Novosibirsk oblast) showed that the conifer bisaccate pollen in the spore–pollen spectra exhibits a series of morphological features allowing its comparison with the modern pollen of Cathaya argyrophylla Chun et Kuang. These palynomorphs belong to relict vegetation, which currently occurs in the wet subtropical climate of South China, and are important climate indicators. The palynomorphs are ascribed to the ancestral forms of the species Cathaya as Abietineaepollenites sellowiiformis (Zaklinskaja) Doweld 2018. The pollen is described, and its distribution along the section is shown. The maximum amount of the A. sellowiiformis pollen (up to 23.7%) is confined to the upper part of the Oligocene and the Lower Miocene and its last appearance is recognized in the Middle–Upper Miocene. These data expand our knowledge on the geographic range of the Cathaya ancestral form and its role in plant communities of Central Asia in the Cenozoic. [ABSTRACT FROM AUTHOR]

The genus Notiosorex is the only group of shrews in North America with adaptations to arid or semiarid zones. The genus was described with a single variable species, Notiosorex crawfordi , from which 5 new species have since been distinguished. To date, the phylogenetic relationships of Notiosorex species have only been partially analyzed and it is possible that there are still distinct species included within the catch-all of N. crawfordi. Here, we use geometric morphometrics on cranial and mandibular characters of the described extant species, 3 fossil species, and a distinctive population of N. crawfordi as a proxy for an integrated assessment of phylogenetic relationships of all Notiosorex species. Our results indicate that the population from the Altiplano Mexicano is more similar to, yet distinct from, N. villai than it is to N. crawfordi —we describe it as a new species. We also used the resulting tree to reconstruct phylogeographic history within the genus, which suggests that Notiosorex sp. nov. N. villai , N. evotis , and N. tataticuli all diverged allopatrically as populations from the ancestral area that is currently occupied by N. crawfordi (and in the deeper past by the fossil species N. harrisi), and pushed south into the Gulf Coast, the Altiplano, and Baja California within the last 5 million years following the end of the Miocene. [ABSTRACT FROM AUTHOR]

Ceratophryid frogs inhabit the lowlands of South America and exhibit greater diversity in the semiarid Gran Chaco. The presence of Ceratophryidae dates back to the late Miocene on the eastern slopes of the Andes in Argentina. Herein, we describe the remains of Ceratophrys from late Miocene of Palo Pintado Formation (Salta, Argentina). Fragmentary bones were recovered from the base of the unit in beds dated between 8 and 7.7 million years ago. The specimen IBIGEO-P141 exhibits the fusion of dentary and mentomeckelian bones forming a fang, a sharp-edged lower jaw, and sculptured bones as Ceratophryidae. The granular pattern of sculpturation, differences between pars facialis and pars dentalis of maxillary, and the shape of lower jaw, support its assignment to the genus Ceratophrys. Combined bioclimatic data and geographic distribution of living Ceratophrys indicate a strong influence of precipitation variables in the distribution of these frogs with adaptations to survive long dry periods. These facts suggest a seasonal rainfall with a well-defined dry season during the deposition of lower levels of Palo Pintado Formation. The palaeoclimatic conditions resemble those occurring in the actual semiarid Chaco, and reveal that the historical geographic distribution of Ceratophryidae was retracted eastward due to the Andean uplift. [ABSTRACT FROM AUTHOR]

This study focuses on the post-Miocene neotectonic evolution of the slope-perched sedimentary basins around the Eastern Korean Continental Margin and which category these basins could be classified into. The Hupo Basin, the largest neotectonic sedimentary basin on the Eastern Korean Continental Margin, comprises three key geological components: 1) Neotectonic Sequence, which has divergently filled the Hupo Basin since the Early Pliocene, 2) Hupo Fault, a N-S striking, high-angle thrust (or reverse) fault that bounds the Hupo Basin to the east, and 3) Hupo Bank, a N-S elongated submarine wave-cut platform that sits on top of the uplifted eastern Hupo Fault block. On the basis of stratigraphic and structural restoration, we conclude that the Hupo Basin was formed by the compressional uplift of the Hupo Bank during the incipient subduction of the Ulleung Basin crust after the Miocene. Given the morphostructural features of the Hupo Basin and its host incipient subduction system, we categorize it as a sort of piggyback basins. The basin-forming process shown in this study is commonly found in numerical models that simulate modern cases of subduction initiation around the world. [ABSTRACT FROM AUTHOR]

The results of regional studies of Early Miocene deposits in Western Ciscaucasia, based on the seismic stratigraphic analysis, are presented. The spatial pattern of sediment accumulation is analyzed and the paleogeographic conditions during the Late Miocene regressive stages in Western Ciscaucasia are refined. Erosional incisions of several levels, developed during the base-level fall in the course of major regressions in the studied time interval, were identified. Based on the spatial correlation of paleoincisions using a chosen series of intersecting seismic profiles, the buried Paleo-Don and Paleo-Donets river valleys were reconstructed at the Sarmatian‒Meotian boundaries and within the Meotian‒upper Pontian interval. [ABSTRACT FROM AUTHOR]

The fossil record of the genus Carcharodon (Lamniformes: Lamnidae) dates to the late Oligocene and has a near global distribution. Today the genus is represented by a single species, the white shark Carcharodon carcharias. However, multiple extinct species are recognised in the Cenozoic including Carcharodon hastalis, Carcharodon hubbelli and Carcharodon planus comb. nov. Here we present the first occurrences of Carcharodon hubbelli and Carcharodon planus from New Zealand/Aotearoa, from the South Island/Te Waipounamu. Carcharodon hubbelli is previously reported from late Miocene deposits in the Pacific basin of North and South America, Asia, and Australia. Whereas Carcharodon planus has been reported from the Miocene of North America and Asia. One tooth from Carcharodon hubbelli is reported from the presumed late Miocene of Motunau Beach, North Canterbury, while one upper early Miocene tooth recovered from the Kakahu River, South Canterbury and an earliest Miocene tooth from the Big River mouth, Tasman District are referred to Carcharodon planus. In addition to being the first reported occurrences of these two species in New Zealand, the teeth of Carcharodon planus represent some of the earliest occurrences of the taxon as well as its southernmost occurrence. [ABSTRACT FROM AUTHOR]

The Late Oligocene is a period of high penguin diversity, following major changes in the marine environment at the Eocene/Oligocene boundary and prior to the emergence of crown penguins in the Miocene. Historically, a large morphological gap existed between the most crownward Platydyptes among the Oligocene penguins from New Zealand and the Early Miocene stem penguins such as Palaeospheniscus from South America. Here we describe a new species that contributes to filling this gap. Pakudyptes hakataramea gen. et sp. nov. is the earliest tiny penguin, overlapping in size with the little penguin Eudyptula minor. Its distinctive combination of a well-developed proximal end of the humerus and an archaic elbow joint provides clues to the evolution of penguin wings. Phylogenetic analysis indicates that penguin wings evolved rapidly from the Late Oligocene to the Early Miocene, together with the acquisition of morphofunctional and hydrodynamical characteristics that enable the excellent swimming ability of modern penguins. As an indicator of aquatic adaptation, bone microanatomy shows a comparable structure to that of Eudyptula. The appearance of the smallest body size and the evolution of modern wings may have led to the ecological diversity of modern penguins, which confirms the importance of Zealandia in penguin evolution. [ABSTRACT FROM AUTHOR]

The earliest Miocene (Aquitanian, 23–20 Ma) remains a critically under-sampled 'dark age' in cetacean evolution. This is especially true of baleen whales (mysticetes), Aquitanian specimens of which remain almost entirely unknown. Across the globe, the nature of the cetacean fossil record radically shifts at the Oligocene-Miocene boundary, with mysticetes and some archaic odontocete lineages suddenly disappearing despite the availability of cetacean-bearing rock units. New Zealand is the only place worldwide where this change is not readily apparent, with baleen whales apparently persisting into the earliest Miocene. Whether this is a genuine pattern has so far remained obscured by a lack of biostratigraphic resolution associated with the Oligo-Miocene boundary. Here, we report 23 new strontium (87Sr/86Sr) dates from Lentipecten shells associated with 16 mysticete and seven odontocete specimens, respectively. Of these, eight fall within the Early Miocene and seven – including five mysticetes – specifically within the Aquitanian. Our findings confirm the unique nature and global importance of the cetacean fossil record from New Zealand, and provide a foundation for investigations into the causes and effects of the Early Miocene cetacean 'dark age'. [ABSTRACT FROM AUTHOR]

Albatrosses are among the most intensely studied groups of living birds, yet their fossil record remains sparse. Despite modern albatrosses being more abundant and widespread in the Southern Hemisphere, the vast majority of fossil albatrosses identified to date come from Northern Hemisphere localities. Here, we describe Plotornis archaeonautes sp. nov., a new albatross species from the earliest Miocene that represents the earliest record of Procellariiformes in New Zealand and the earliest uncontroversial record of the clade Pan-Diomedeidae from the Southern Hemisphere. Phylogenetic analyses support the placement of Plotornis outside of the clade uniting all extant albatrosses. The new fossil reveals that stem lineage albatrosses were widespread by the onset of the Neogene. Although the humerus of Plotornis archaeonautes exhibits a short processus supracondylaris dorsalis, this early species may have possessed at least one of the unique ossifications associated with the patagial bracing system present in modern albatrosses. [ABSTRACT FROM AUTHOR]

Ctenium has about 20 species distributed in Africa, Arabia, and the Neotropics, with no species occurring on more than one continent. Its relationship with the morphologically similar Kampochloa is still unclear and has not been investigated. In this study, we performed phylogenomic analyses using 71 protein‐coding genes of 24 newly sequenced plastomes, of 13 species of Ctenium and 1 species of Kampochloa to understand their biogeography and the phylogenetic and evolutionary relationships between these two genera and within Ctenium. Our results support two major clades of Ctenium: the Neotropics and Old‐World clades. Kampochloa is shown to be a close relative of the genus Ctenium, and their morphological similarities support this. Divergence time estimation analyses show that Kampochloa and Ctenium diverged around 7.99 Ma, and might have originated in the late Miocene during the rapid expansion of C4 grasslands. With this evidence from morphology and phylogenomic data, we propose the inclusion of the previously unplaced Kampochloa into the subtribe Cteniinae. [ABSTRACT FROM AUTHOR]

Background: The superorder Forcipulatacea is a major clade of sea stars with approximately 400 extant species across three orders (Forcipulatida, Brisingida, Zorocallida). Over the past century, the systematics of Forcipulatacea have undergone multiple revisions by various authors, with some considering numerous families such as Asteriidae, Zoroasteridae, Pedicellasteridae, Stichasteridae, Heliasteridae, Labidiasteridae, and Neomorphasteridae, while others recognized only two families (i.e., Asteriidae and Zoroasteridae). Recent molecular analyses have shown the artificial nature of some of these groupings. Notably, four well-supported clades (Zorocallida, Brisingida, Stichasteridae, and Asteriidae) emerged from a synthesis of morphological and molecular evidence. The majority of extinct forcipulatacean species have been placed in modern families. However, many of these fossil species are in need of revision, especially those species placed within the Asteriidae, the largest of all forcipulatacean families. Methods: In light of recent advancements in forcipulatacean systematics, we comprehensively reassess six well-preserved Jurassic forcipulatacean taxa, including the earliest crown-group members from the Hettangian (∼201.4 Ma), and also describe two new Jurassic genera, Forbesasterias gen. nov. and Marbleaster gen. nov. We assembled the largest and most comprehensive phylogenetic matrix for this group, sampling 42 fossil and extant forcipulatacean species for 120 morphological characters. To infer phylogenetic relationships and construct an evolutionary timeline for the diversification of major clades, we conducted a Bayesian tip-dating analysis incorporating the fossilized birth-death process. A total of 13 fossil species were sampled in our analysis, including six taxonomically revaluated herein, two recently reappraised species from the Jurassic, and five additional species from the Cretaceous and Miocene. Results: Contrary to prior assumptions, our results indicate that none of the Jurassic taxa investigated belong to Asteriidae or any other modern families, and instead represent stem-forcipulatids. Furthermore, our phylogenetic results suggest that Asteriidae likely originated during the late Cretaceous. Our findings highlight a greater early diversity within the Forcipulatacea than previously presumed, challenging existing perceptions of the evolutionary history of this significant clade of marine invertebrates. [ABSTRACT FROM AUTHOR]

Ferromanganese nodules are important marine storehouses for critical metals and windows for changing oceans. Although advanced in situ analytical techniques have been applied to visualize the elemental distribution in the nodule cross‐sections, their spatial distribution remains largely uncertain. This study addresses this gap by employing micro X‐ray fluorescence mapping of parallel nodule cross‐sections to delineate the spatial distributions of critical metals (Mn, Fe, Co, Ti, Ni, and Cu) in three Co‐rich ferromanganese nodules from the northwestern Pacific Ocean. The 10‐layer Os isotopic compositions of one nodule closely align with the well‐documented marine Os isotope evolution of seawater, providing a chronological framework and a maximum age of ∼36 Ma for these nodules. Three concentric chemostratigraphic layers, labeled L1, L2, and L3, were identified from the inside out, based on microscopic structures and the distributions of critical metals. The early growth stage was marked by Mn‐rich, Si‐rich, and high Mn/Fe ratios, suggesting a diagenetic‐driven process attributed to high paleoproductivity conditions because of the low latitude of the study area at that time. The subsequent growth stages are all hydrogenetic in origin to be rich in Fe, Co, and Ti with low Mn/Fe ratios. The apparent detritus present during the second growth stage of the nodules may correspond to the stronger bottom currents in the early Miocene. The final mineralization stage indicates a more stable environment with diminished bottom current activity, leading to the formation of a dense, laminar hydrogenetic layer. Plain Language Summary: The present study bridges the knowledge gap on the spatial distribution of critical metals through the innovative use of micro X‐ray fluorescence mapping on parallel sections of three nodules in the northwestern Pacific Ocean. Os isotope stratigraphy establishes a chronological framework and a maximum age of ∼36 Ma for these nodules. Distinct concentric chemostratigraphic layers, labeled L1, L2, and L3, have been identified in these nodules. The early growth stage of the nodules is characterized by Mn‐rich, Si‐rich and high Mn/Fe ratios, indicative of a diagenetic‐driven process under high paleoproductivity conditions due to the low historical latitude. The subsequent stages are all hydrogenetic in origin to be rich in Fe, Co, and Ti with low Mn/Fe ratios. The apparent detritus present during the second growth stage of the nodules may correspond to the stronger bottom currents in the early Miocene. The final mineralization stage suggested a period of environmental stability with reduced current activity, forming of a hydrogenetic layer enriched in Fe, Co, and Ti. This study not only enhances our understanding of the complex processes that govern the formation and evolution of ferromanganese nodules but also provides deeper insight into the historical shifts within our oceans. Key Points: Ferromanganese nodule was dated via Os isotope stratigraphy and has a maximum age of ∼36 MaMulti‐element integrated maps were produced to assist in obtaining clearer chemostratigraphy images of ferromanganese nodulesSpatial distribution of critical metals and chemostratigraphy in ferromanganese nodules were visualized via multiple parallel cross‐sections [ABSTRACT FROM AUTHOR]

Hitherto unreported whale fall fauna has been found from the Lower Miocene of Khari Nadi Formation of Kutch, India.This vertebrate-invertebrate assemblage comprises jumbled up bone fragments of different parts of the whale along with molluscs and echinoids. Some representatives in this assemblage are characteristic whale fall fauna and some are characteristic shallow marine fauna. The molluscs belong to the families Buccinidae, Provannidae, Naticidae, Turritellidae, Borsoniidae, Volutidae, Haminoeidae, Cypraeidae, Strombidae, Rostelleriidae, Cassidae, Osteridae, Veneridae, Astartidae and the echinoids to family Spatangidae and Schizasteridae. The present study reveals that the deposition of this entire assemblage was in a shallow marine environment and also sheds some light on the molluscan association related to a whale fall from the western Indian sea waters in the Lower Miocene. [ABSTRACT FROM AUTHOR]

I here describe lacertid and snake remains from the classic Middle Miocene (MN 6) locality Bonanza of the Devínska Kobyla Hill near Bratislava (Slovakia). The locality is famous for being a type locality of an early seal Devinophoca Koretsky & Holec, 2002. During the Middle Miocene, the area of Devínska Nová Ves was part of an archipelago in the western part of the Central Paratethys in the northern part of the Vienna basin. The fossils described here comprise an incomplete left maxilla of a lacertid lizard and an articulated portion of a vertebral column of a small colubrid snake with several ribs still attached. The maxilla is tentativelly allocated here to Lacerta Linnaeus, 1758 and is a rare instance of the occurrence of the clade Lacertidae Oppel, 1811 in this environment during the Middle Miocene. Moreover, it represents the oldest known occurence of this clade in Slovakia. Articulated snake specimens from the Cenozoic are rare and although only party articulated, the specimen from Bonanza is therefore exceptional. On the basis of vertebrae alone, determination of fossil colubrids is very difficult. The morphology of the Bonanza specimen is most similar to the extant Dolichophis Gistel, 1868, as also to other small fossil snakes assigned to 'Coluber' Linnaeus, 1758 when that genus was a catch-all grade taxon recognised from both Europe and North America, notably 'Coluber' dolnicensis Szyndlar, 1987 and 'Coluber' pouchetii (Rochebrune, 1880). Because the potentially closest living relative of the European Miocene 'Coluber' is Dolichophis, I assigned tentatively the material from Bonanza to cf. Dolichophis, rather than to the wastebasket taxon 'Coluber'. [ABSTRACT FROM AUTHOR]

The NE Tibet experienced complex and distinct basin developments and uplifts in different areas. However, the reasons for such distinct surface deformation and their relationship to deep crustal geodynamic processes are not well understood. Here, we obtain a crust model of NE Tibet by jointly inverting Rayleigh wave ellipticity and phase velocity. Our results reveal that deep crustal strength contrasts across NE Tibet play an important role in controlling basin development. Extrusion of the significantly weak Qilian crust is obstructed by rigid Alxa block, resulting in deep foreland basin with dramatic topographic step. In contrast, the relatively weak crust of Longzhong absorbs outward extrusion of NE Tibet within a wide transition zone, leading to small intermontane basins. Furthermore, the systematic thinning of basins from north to south around the western Ordos Block demonstrates the tectonic transformation from extension to compression due to expansion of NE Tibet since the late Miocene. Plain Language Summary: In this study, we obtain a high‐resolution model of NE Tibet by jointly inverting Rayleigh wave phase velocity and ellipticity (the radial‐to‐vertical amplitude ratio), which provides complementary constraints to the shallow structure. Our results show a clear correlation between velocity variations in the deep crust and basin structures at the surface. We infer that the extrusion of mechanically weak mid‐to‐lower crust of Qilian is obstructed by the strong Alxa block, leading to the steep topography and deep foreland Hexi Basin. To the east, in contrast, the outward extrusion of Songpan‐Ganzi is absorbed in a wide range by the relatively weak Longzhong region, developing gently sloping topography and small intermontane basins. Furthermore, the significant structural differences of rift basins from north to south around the western Ordos likely result from tectonic regime transformation induced by the continuous expansion of NE Tibet since the late Miocene. Our results improve the understanding of how deep crustal geodynamic processes influence surface uplift and basin developments in the expanding NE Tibet. Key Points: A high‐resolution 3‐D crustal model of NE Tibet is obtained by joint inversion of Rayleigh wave ellipticity and phase dispersionWe infer that deep crust extrusion and strength differences across plateau boundaries control distinct surface deformation in NE TibetThe systematic thinning of basins in west Ordos indicates tectonic regime transformation due to expansion of Tibet since Miocene [ABSTRACT FROM AUTHOR]

Although the barred sand burrower, Limnichthys fasciatus, is widely distributed throughout the western Pacific, including Japan, Taiwan, and Australia, its morphology and genetics are poorly known. We discovered four cryptic species of Limnichthys from the western Pacific based on mtDNA cytochrome c oxidase subunit I (COI) and 16S ribosomal RNA (rRNA) sequences. Genetic distances showed remarkably large differences (12.7%–24.3% in COI and 7.9%–19.6% in 16S rRNA) between true L. fasciatus (type locality: southeastern Australia) and the others. A relaxed clock model with optimized selected substitution models showed that their deep divergence began in the middle Miocene epoch and subsequently diverged into the current cryptic species in the Plio‐Pleistocene. A eurythermal common ancestor may have evolved independently in each region due to geographical events and paleoclimatic fluctuations, which made it possible for L. fasciatus complex to be an anti‐equatorial species. Despite their deep genetic divergence, they showed marked phenotypic similarity, suggesting that they have experienced similar selective pressures related to their specific behavior. [ABSTRACT FROM AUTHOR]

The paper presents the lithologies used in the stone products of Neviodunum (modern Drnovo in Slovenia), a Roman municipium in south‐western Pannonia. For this purpose, 95 stone monuments were assessed. Petrographic and biostratigraphic analyses were carried out on 56 archaeological and 57 geological samples. Our research shows that, besides some rare exceptions, most stone products refer to three lithostratigraphic units: the Middle Miocene ‘Lithothamnium’ Limestone Member of the Laško Formation, the Upper Cretaceous Krško Formation and the Early Jurassic Krka Limestone Member of the Podbukovje Formation. [ABSTRACT FROM AUTHOR]

According to GNSS/INSAR measurements, the Konya Basin in Central Anatolia is undergoing rapid subsidence within an uplifting orogenic plateau. Further, geophysical studies reveal thickened crust under the basin and a fast seismic wave speed anomaly in the underlying mantle, in addition to a localised depression in calculated residual topography (down to 280 m) over the Konya Basin, based on gravity-topography considerations. Using scaled laboratory (analogue) experiments we show that the active formation of the Konya Basin may be accounted for by the descent of a mantle lithospheric drip causing local circular-shaped surface subsidence. We interpret that the Konya Basin is developing through a secondary drip pulse that is contemporaneous with broad plateau uplift caused by a larger-scale lithospheric drip since the Miocene. The research reveals that basin evolution and plateau uplift may be linked in a multistage process of lithospheric removal during episodic development of orogenic systems. Active subsidence of the Konya region, Turkiye, is attributed to descent of a lithospheric drip into the mantle, according to analogue models, and suggests that major tectonic events may be linked to multistage dynamic flow processes in the mantle. [ABSTRACT FROM AUTHOR]

The Burdigalian/Langhian (B/L) boundary has not yet been designated as a Global Stratotype Section and Point (GSSP), despite various proposed zonal schemes. In the Gulf of Suez region of Egypt, the Burdigalian-Langhian successions are notable for hosting significant hydrocarbon reservoirs within a tectonic rift setting. Therefore, biostatigraphy plays a crucial role in exploration endeavors in this area. The nannofossil biostratigraphy is investigated in two sections, Wadi Baba and Wadi Gharandel, of the lower-middle Miocene from west-central Sinai. Three biozones, NN3 (Sphenolithus belemnos) Zone, NN4 (Helicosphaera ampliaperta) Zone, and NN5 (Sphenolithus heteromorphus) Zone, are identified from the studied interval. The NN4 Zone could be divided into MNN4a/b and MNN4c. Important biovents are discussed, such as the S. heteromorphus paracme interval and the first occurrence and evolution of the Discoaster exilis and Discoaster variabilis groups. Based on the cluster analysis, the recorded taxa can be subdivided into four groups that reflect their palaeoclimatic preferences. The paleoecological interpretation of the studied Rudies Formation indicates prevailing cool and eutrophic nutrient conditions based on the dominance of taxa such as Coccolithus pelagicus, Reticulofenestra minuta, and Cyclicargolithus floridanus. The nannofossil taxa responses to sea level curve are interpreted. Fluctuations in taxa abundance and diversity indicate a slight rise in the sea level at the base of the Burdigalian followed by sudden drop in the sea level at the middle Burdigalian. High sea-level conditions prevailed again until the B/L boundary. During the Langhian period, many small-scale fluctuations in sea-level curve are detected. [ABSTRACT FROM AUTHOR]

The Zama discovery was identified off the coast of Tabasco, Mexico, in the Sureste Basin of the Gulf of Mexico and is hosted in a three‐way closure in the Upper Miocene. This study conducted a detailed detrital zircon (DZ) U–Pb and (U‐Th)/He provenance analyses on samples from sandstone reservoirs in the Zama‐3 and Zama‐2ST1 wells. A total of 22 sandstone samples (11 from each well) were collected for DZ U–Pb and (U‐Th)/He dating from different reservoir zones, testing the hypothesis that different zones were whether originally derived from varied sedimentary source terranes and associated transport pathways to the Zama mini‐basin depositional site. Additional objectives include determination of maximum depositional ages, reconstruction of paleofluvial systems, and exploring the temporal evolution of the drainage region and hinterland tectonics. The DZ U–Pb age spectra from both Zama wells have remarkably homogenous DZ signatures with very similar DZ age modes and modal percentages, displaying dominant Permian/Chiapas Batholith (ca. 35%–45%), Mesoproterozoic/Oaxaquian (ca. 20%–35%), Early Palaeozoic/Acatlán (ca. 8%–20%), and Cenozoic magmatic arc (ca. 15%–25%) age modes, as well as some subsidiary (<5%) early Proterozoic/Archean and Early Cretaceous DZ age components, linked to recycled lower Palaeozoic strata and the Guerrero Terrane and Alisitos arc, respectively. Despite differences in paleocurrent directions, deduced from image logs, there are no systematic differences in DZ spectra, indicating a consistent sediment provenance and no changes in source area. All Zama samples analysed in the study are characterized by abundant syn‐depositional Late Miocene DZ grains, clustering between 8.6 and 10.2 Ma, corroborating a Tortonian (Late Miocene) depositional age, and yield rapid sediment accumulation rates of ca. 200 m in <1.4 Ma (13 m/Myr). Doubled zircon U–Pb and (U‐Th)/He age pairs are indicative of recycling of early Mesozoic rift strata and Paleogene and younger Chiapas basement. These new DZ U–Pb and (U‐Th)/He data have a nearly invariant Tortonian sediment provenance that is similar to the modern Grijalva River flowing generally northward out of the Chiapas highlands. The paleo‐Grijalva drainage, providing sediment to the Late Miocene Zama mini‐basin, was likely drastically larger than the present catchment as it involved 10 Ma plutonic sources that were subsequently downfaulted along the Pacific coast in the latest Miocene. Importantly, DZ U–Pb age components are consistent with Oaxaquia, early Acatlán, and Guerrero/Alisitos signatures and point to sourcing from the Chortis block during its tectonic eastward translation. Such a scenario would allow for a substantially larger Miocene paleo‐drainage that would have encompassed both Chiapas and portions of Chortis. The Miocene tectonic translation of Chortis and erosion of a large and tectonically active hinterland would also reconcile the dramatically larger Middle to Late Miocene sediment supply, funnelling a large influx of sediment into the southern Gulf of Mexico. Rapid Late Miocene sediment accumulation in the salt‐defined Zama mini‐basin must have involved sustained sediment flux via the paleo‐Grijalva drainage and was likely facilitated and focused by continued salt deflation due to sediment loading, as described in Part II. Thus, our work provides new scientific insights into one of the largest hydrocarbon discoveries in Mexico in recent years. [ABSTRACT FROM AUTHOR]

The Neogene and Quaternary hinterland basins of the Northern Apennine have been the subject of different tectonic interpretations. Several studies considered these basins as the result of polyphase normal faulting framed in a continuous crustal extensional regime since the middle Miocene. On the contrary, geophysical and geological studies provided evidence of the important role played by out‐of‐sequence thrusts and backthrusts in the evolution of these basins during a prolongated and intense period of shortening. Here we present an integrated analysis of 2D stacked seismic reflection profiles, stratigraphic and geophysical data from deep exploration wells, gravity data, and published geological and biostratigraphic data for the Valdera‐Volterra basin (central Tuscany, Italy). The results support a polyphase and composite evolution of the basin, subdivided into three main phases. During the late Tortonian–Zanclean, the growth of major thrust‐related anticlines controlled the evolution of the sedimentary basin. The growth of a syncline determined the creation of accommodation space for the sediments. This main compressional deformation occurred during the Messinian and ended during the Late Zanclean. NE migration of the depocentre during the Early Zanclean was identified, likely possibly due to a differential activity growth between the bordering anticlines. During the Piacenzian, an extensional phase has been recognised, superposed to the previous compressive phase. During the Latest Piacenzian–Early Pleistocene (?), a final compressional phase took place resulting in the positive inversion of the Piacenzian WSW dipping main border fault. [ABSTRACT FROM AUTHOR]

The provenance of most basin systems today is interpreted based on radiogenic ages or the geochemical composition of detrital minerals, which has all but replaced the use of whole-rock geochemical approaches that can effectively complement provenance information inferred from detrital approaches. Here, we further investigate previous provenance models developed using detrital zircon U-Pb geochronology by applying whole-rock major and trace element geochemistry of fine-grained clastic rocks from the late Oli-gocene–middle Miocene Tyonek Formation, late Miocene Beluga Formation, and Pliocene Sterling Formation in the Cook Inlet Basin, Alaska, USA. When taken alone, our new geochemical data suggest solely intermediate igneous sediment sources to the basin. When paired with existing detrital zircon U-Pb data, however, significant mixing of felsic and mafic sediment sources is evident, which indicates that thorough mixing of geochemically distinct source terranes can mask the input from individual sources in whole-rock geochemical studies. Furthermore, we demonstrate that both weathering and provenance influence the major element chemistry of sediment source terranes as well as the resultant basinal strata. Our conclusions indicate that the combination of whole-rock geochemistry with other detrital approaches provides a robust interpretation of sedimentary basin provenance. [ABSTRACT FROM AUTHOR]

Fossils can unveil a long-vanished combination of character states that inform inferences about the timing and patterns of diversification of modern fungi. By examining the well-preserved stacked chained vesicular conidiophores developed in clusters from the basal stroma, we describe a new taxon of fossil Zygosporiaceae with a combination of characters unknown among extant taxa on compressed serrated-margined dicot leaf (cf. Fagaceae) recovered from the Siwalik sediments (Late Miocene; ca. 12–8 Ma) of Himachal Pradesh, western Himalaya. Based upon conidiophore morphology, our Siwalik fungal remains, similar to Zygosporium Mont. (Zygosporiaceae: Xylariales: Sordariomycetes), are recognized as a new fossil species, Z. stromaticum Kundu & Khan, sp. nov. Zygosporium stromaticum is the only known fossil anamorphic fungus that occurs on plant cuticles and has a cluster of stacked chained vesicular conidiophores arising from a poorly preserved basal stroma formed by irregular, thick-walled cells. Its combination of morphological characteristics is unknown in extant fungal taxa, so Z. stromaticum likely represents a new anamorphic foliicolous fungus that may now be extinct. This unique evidence may be essential for the calibration of divergence time estimations of fungal lineages. [ABSTRACT FROM AUTHOR]

An association of eighteen coprolites (specimens 01–18) and one isolated coprolite (specimen 209,210) were found in a vertebrate fossil-rich paleosol at the Gratkorn site (south-eastern Austria; late Middle Miocene). The specimens consist mostly of calcium phosphate (apatite) and a matrix formed by microglobules. Coprolites 01–18 show cylindrical and spherical morphologies and are considerably smaller than the tube-shaped specimen 209,210, in which no inclusions were observed. In contrast, coprolites 01–18 contain numerous, highly altered bone fragments (sub-mm-sized long bones and several mm-sized trabecular bone remains) as well as hair imprints, plant detritus and palynomorphs. Based on composition, morphology, size, microstructure, and inclusions, and considering the body fossil record of this site, we assume the hyaenid Protictitherium and the barbourofelid Albanosmilus, as producers of coprolites 01–18 and 209,210, respectively. The preserved bone remains in specimens 01–18 suggest that Protictitherium fed on small vertebrates, but possibly also cracked bones of medium-sized animals. The hair imprints found were either from the hyaenid itself or its prey, while the plant material was probably ingested accidentally. The lack of inclusions in specimen 209,210 is related to the presumably hypercarnivorous diet of Albanosmilus, which was certainly the apex predator in this biome. [ABSTRACT FROM AUTHOR]

Two carnivoran mandibles were recovered from the Late Miocene Dhok Pathan Formation outcrops of Hasnot village of northern Pakistan. Thorough morphometric comparison of the specimens verifies an assignment to 'Martes' lydekkeri. The present specimens represent new records of this rare mustelid species and confirm the extension of its stratigraphic range from the Chinji Formation to the Dhok Pathan Formation. [ABSTRACT FROM AUTHOR]

Valgus cantalensis sp. nov., second fossil species of the scarabaeid subfamily Valginae, is described from the Upper Miocene Konservat-Laggerstätte of Sainte-Reine (France). The fossil is compared to the extant valgine genera and Valgus spp., showing it is a new species. But its exact affinities in the genus Valgus remain undetermined because of the lacking of a recent morphological phylogenetic analysis of the Valgus spp. [ABSTRACT FROM AUTHOR]

The terrestrial isopod genus Ligidium includes 58 species from Europe, Asia, and North America. In Eastern North America four species are recognized: L. floridanum and L. mucronatum, known just from their type localities in Florida and Louisiana respectively, L. blueridgensis, endemic to the southern Appalachians, and L. elrodii, widespread from Georgia to Ontario. The genus shows a marked morphological conservatism, and species are differentiated mostly by small morphological differences; it is not always easy to determine if such variability represents inter‐ or intraspecific variation. Here, we explore the diversity of Ligidium from the southern Appalachian Mountains, exploring the congruence of morphologically defined groups with multilocus phylogenetic reconstructions and molecular species delimitation methods. We have studied a total of 130 specimens from 37 localities, mostly from the southern Appalachians, and analysed mtDNA (Cox1) and nuclear (28S, NaK) sequences. Morphologically, we recognized eight morphotypes, most of them assignable to current concepts of L. elrodii and L. blueridgensis. Phylogenetic analyses supported the evolutionary independence of all morphotypes, and suggest the existence of 8–9 species, including limited cryptic diversity. Single‐locus delimitation analyses based on mtDNA data suggest the existence of a much higher number of species than the multilocus analyses. The estimated age of the ancestors of sampled lineages indicates a long presence of the genus in eastern North America and old speciation events through the Miocene. Our results indicate a higher diversity than previously thought among the Ligidium populations present in the southern Appalachian Mountains, with several species to be described. [ABSTRACT FROM AUTHOR]

A deep-marine sedimentary succession of Whaingaroan to Altonian age (Early Oligocene–Early Miocene), cropping out near Weber, southern Hawke's Bay, records abrupt changes in depositional paleoenvironments and sedimentary lithofacies. Highly calcareous early Waitakian (latest Oligocene) Weber Formation (Mangatu Group/Waka Supergroup) is unconformably overlain by terrigenous-dominated late Waitakian–Otaian (Early Miocene) Coast Road or Whakataki formations (Tolaga Group/Māui Supergroup). Most notably, in some localities, lowermost parts of the Coast Road Formation (Mangapuku Mudstone Member) contain prominent matrix-supported extra-formational olistoliths of centimetre to decametre scale derived from the underlying Weber Formation. Deposition of these blocks is attributed to deep-water collapse of parts of rapidly growing, over-steepened, reverse-fault-controlled thrust ridge(s). The onset of Coast Road and Whakataki formation deposition and associated abrupt changes in sedimentary lithofacies correlate to prominent Early Miocene lithological, mineralogical, and paleoenvironmental changes elsewhere along the Hikurangi Margin. We attribute these bio- and lithostratigraphic changes as evidence for significant shortening and upper-plate reverse faulting marking the onset of subduction beneath eastern North Island during the mid-Waitakian (c. 23 Ma). Proposed lithostratigraphic revisions presented herein help clarify depositional events and correlative sedimentary packages within the Miocene Hikurangi margin. [ABSTRACT FROM AUTHOR]

With the high demand for fossil fuels, exploring the frontier areas for hydrocarbon reserves has become imperative. The recent discoveries in Gojalia, Sonamura, Baramura, and Sundalbari fields emphasize the need to explore additional anticlinal structures in Tripura for hydrocarbon exploration. Tulamura anticline (the study area) produced gas from Upper Bhuban, establishing hydrocarbon prospectivity in the northern part, but the southern part remains largely unexplored. An electro-log interpretation revealed the presence of sand facies deposited in a fining upward sequence, suggesting channel deposition. An integrated geophysical approach using seismic inversion and machine learning techniques was performed to delineate and characterize the litho-facies dispersal patterns in the Tulamura field. Spectral decomposition (12, 20 and 28 Hz) of stacked seismic data were RGB (red-green-blue) blended, revealing the southward striking channel geometry of the Bhuban Formation at a depth of 2220 m. The 3D P-impedance and Vp/Vs ratio volumes were estimated using the model-based pre-stack seismic inversion. Inversion results help discriminate among sand, shale and siltstone litho-facies. Petrophysical property (effective porosity) was predicted by combining the post-stack seismic attributes and well-log data using neural network modelling. The identified sand facies within the channel geometry exhibit relatively moderate to low P-impedance (9800–10600 m/s * gm/cm3), low Vp/Vs ratio (1.68–1.76), and moderately high effective porosity (8–13%) from surroundings, indicating favourable conditions for hydrocarbon accumulations. Shale between channels and major faults can create favourable stratigraphic entrapment, while an upward fining sequence suggests an intact top seal. This study advocates an integrated approach involving geophysical inversion and machine learning to identify optimal conditions for hydrocarbon accumulation within sand facies, supported by structural and stratigraphic entrapment. [ABSTRACT FROM AUTHOR]

The "Diahot Tooth" is an isolated postcanine tooth of a large herbivorous mammal, discovered in the Diahot region of northern New Caledonia in 1875. Most authors have identified it as an upper premolar of a rhinocerotid, but an alternative proposal is that it belongs to a diprotodontoid marsupial that has been named Zygomaturus diahotensis. Either possibility raises biogeographical difficulties, because New Caledonia has been isolated from other major landmasses for 80 million years, and neither rhinocerotids nor diprodotontoids appear to be good candidates for such a long-distance overwater dispersal event. Here, we present a novel interpretation of the affinities and origin of the Diahot Tooth, based on qualitative study of its preserved morphology and quantitative phylogenetic analyses that include both rhinocerotids and diprotodontoids. We show that the Diahot Tooth most closely resembles the first deciduous premolar of Western Eurasian Miocene teleoceratine rhinocerotid Brachypotherium brachypus, with the few discrepancies relating to traits that are known to be variable in B. brachypus. Our phylogenetic analyses also support this relationship. The preservation of the Diahot Tooth closely resembles that of B. brachypus teeth from the "Faluns Sea" of the Loire basin, and we propose that the New Caledonian specimen originated there and was taken to New Caledonia by a European colonist during the mid-19th century, where it was lost, rediscovered, and incorrectly assumed to be autochthonous. [ABSTRACT FROM AUTHOR]

Ascidians, or sea squirts, are sac‐like sessile tunicates commonly found in modern seas. Although the oldest ascidians, or at least ascidian‐like tunicates, are mid‐Cambrian in age, the quality of the ascidian fossil record is extraordinarily poor. It mostly consists of isolated finds or low‐diversity assemblages, often represented by mineral spicules. Our study reports the world's most diverse assemblage of ascidian spicules, originating from the Middle Miocene Bogutovo Selo near Ugljevik, Bosnia and Herzegovina. This material includes 31 spicule types, with four assignable to species (Lissoclinum cf. perforatum, Polysyncraton cf. bilobatum, Didemnum cf. vexillum and D. cf. leopardi) and 10 to genera, all belonging to aplousobranchiate ascidians. Comparing our findings with those of two other studies that use modern biological nomenclature, our assemblage shows a closer resemblance to the distant Eocene Australian assemblage than to the Miocene ascidians from Eastern Paratethys. This suggests the persistence of a once‐homogeneous and widely distributed Eocene fauna into the Miocene, while the Eastern Paratethyan basin developed an endemic fauna after the isolation of Eastern Paratethys from the open sea. Some taxa of our assemblage currently inhabit the Mediterranean Sea. The discovery of one species in distant South African waters suggests that tectonic and environmental changes during the Late Miocene, including the closure of the Gibraltar Strait and prior disconnection from the Indian Ocean, may have limited its survival to remote refugia. The study underscores the importance of including spicules in reconstructing ascidian faunas in the geological past, an area that has been surprisingly understudied. [ABSTRACT FROM AUTHOR]

Cheilolejeunea sicilicula S. Fish & Glenny, a new species of liverwort from lowland indigenous forest reserves in Westland and Western Nelson provinces in South Island, New Zealand, is described and illustrated. In New Zealand it is superficially similar to C. campbelliensis (Steph.) R. M. Schust., C. novaezelandiae R. M. Schust. and C. albovirens (Hook.f. & Taylor) E. A. Hodgs, which are all small-leaved species with leaves that range from 195 to 500 µm long. Cheilolejeunea sicilicula differs from these species in having a curved second tooth, and in having elongated cells along the antical free margin of the lobule. Molecular phylogenetic analyses resolve C. sicilicula as a member of the tropical section Cyrtolejeunea which has five extant species and a Miocene fossil species. This section was previously unknown in New Zealand. The addition of Cheilolejeunea sicilicula to the New Zealand flora brings the number of accepted New Zealand Cheilolejeunea species to 12, six of which are endemic. [ABSTRACT FROM AUTHOR]

Massive salt accumulations, or salt giants, have formed in highly restricted marine basins throughout geological history, but their impact on biodiversity has been only patchily studied. The salt giant in the Mediterranean Sea formed as a result of the restriction of its gateway to the Atlantic during the Messinian Salinity Crisis (MSC) 5.97 to 5.33 million years ago. Here, we quantify the biodiversity changes associated with the MSC based on a compilation of the Mediterranean fossil record. We conclude that 86 endemic species of the 2006 pre-MSC marine species survived the crisis, and that the present eastward-decreasing richness gradient in the Mediterranean was established after the MSC. [ABSTRACT FROM AUTHOR]