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Understanding the richness and diversification processes in the Mediterranean basin requires both knowledge of the current environmental complexity and paleogeographic and paleoclimate events and information from studies that introduce the temporal dimension. The Carthamus–Carduncellus complex (Cardueae, Compositae) constitutes a good case study to investigate the biogeographic history of this region because it evolved throughout the basin. We performed molecular dating, ancestral area estimation, and diversification analyses based on previous phylogenetic studies of a nearly complete taxon sampling of the complex. The main aims were to determine the role of tectonic and climatic events in the disjunction of the complex and the expansion route of the two main lineages, Carduncellus s.l. and Carthamus. Our results suggest that the main lineages in the complex originated during the Miocene. Later, all main paleogeographic and paleoclimatic events during the Neogene and Pleistocene in the Mediterranean basin had an important imprint on the evolutionary history of the complex. The Messinian Salinity Crisis facilitated the dispersion of the genus Carduncellus from North Africa to the Iberian Peninsula and the split of the genera Phonus and Femeniasia from the Carduncellus lineage. The onset of the Mediterranean climate in the Pliocene together with some orogenic processes could be the main causes of the diversification of the genus Carduncellus. In contrast, Pleistocene glaciations played a key role in the species diversification of Carthamus. In addition, we emphasize the problems derived from secondary dating and the existing differences between two previous dating analyses of the tribe Cardueae. [ABSTRACT FROM AUTHOR]

In this work, the old Lungro mining site was studied by applying two classification methods (Brilha and IELIG Method), considering the site-specific scientific, educational, touristic, and degradation aspects, to promote it as a geosite. The results were compared with those from other worldwide areas, and a potential re-evaluation in the tourism sector was suggested, considering historical data and geological issues. The methods used evaluate the Lungro site an average score from a scientific perspective, and from high to very high for the tourist-educational aspects. On the contrary, lower scores were found regarding the degradation of the site due to natural phenomena, led to a lower total score. The obtained results are comparable with other similar restored mining areas, located in various parts of the world which today represent important geotourists sites recognized internationally. Based on these results, the safest, most cost-effective, and straightforward way to preserve and restore the site is by creating a “Geotourism Route.“. This choice would increase awareness of the area, providing the general public, including the curious and “non-experts”, with an understanding of various geological processes and the extensive history of the mine. A campaign to promote and preserve the Lungro site as a geoheritage will be encouraged, with significant implications for tourism at the local, national, and international levels. [ABSTRACT FROM AUTHOR]

Giant evaporite deposits formed during the Messinian Salinity Crisis (MSC) in the Mediterranean Sea during the upper Miocene. The primary cause is a restricted Atlantic-Mediterranean connection, but the detailed hydroclimate evolution of this event is still a matter of debate. Here we reconstruct the triple oxygen and hydrogen isotopic composition of paleo brines from structurally bonded water in Messinian gypsum from Cyprus. A two-stage evaporation model is constructed to best approximate the hydrological conditions at which Messinian gypsum formed in marginal basins of the Mediterranean Sea. Subsequently, hydroclimatic parameters like paleo relative humidity (RH) are modelled using an iterative curve fitting isotope model approach. The results of our limited dataset reveal a slightly lower RH during the third compared to the second stage of the MSC. This apparent drop in RH is consistent with previous observations from the literature. Besides absolute values of RH, our model allows reconstructing the triple oxygen and hydrogen isotopic composition of the open Mediterranean Sea. This provides an estimate for the proportions of continental water vs. Atlantic seawater flowing into the basin, serving as a proxy for the size of the strait of Gibraltar. The model implies a continental water fraction of around 75–81 % both for the second and third stage of the MSC. In addition, we determined the 87Sr/86Sr of our samples to: i) confirm their stratigraphy; and ii) estimate the relative proportion of continental water vs. Atlantic seawater entering the Mediterranean using Sr mass balance calculations. The combined oxygen and Sr isotope records indicate a persistent connection to the Atlantic supporting the hypothesis that the striking drop in 87Sr/86Sr at the beginning of MSC stage 3 is mainly related to an increased contribution of continental water with low 87Sr/86Sr and high Sr concentrations from the Paratethys. Our results show that the combination of triple oxygen, hydrogen, and Sr isotope data provides a powerful tool to disentangle paleo-hydroclimate even in such complicated hydrological settings as the Mediterranean Sea during the MSC. [ABSTRACT FROM AUTHOR]

Eucalathis, a member of the brachiopod superfamily Cancellothyridoidea, is common in Recent seas. The 15 known species have an almost cosmopolitan distribution, but no known occurrence in the Mediterranean Sea. Despite the rich Recent record, very little is known about the geological history of Eucalathis. In the Mediterranean, two fossil species are known from the Miocene of Italy. Two additional Eucalathis species from the Neogene of Italy are identified: E. davidi sp. nov. from NW Italy (Marmorito, Early Pliocene), and E. dorae sp. nov. from Sicily (Altavilla, Late Pliocene). It proves that Eucalathis did not disappear from the Mediterranean after the Messinian Salinity Crisis. Based on the four known species, Eucalathis (probably apart from the Messinian Crisis) was present in the Mediterranean from the Middle Miocene to the Late Pliocene. Its disappearance is possibly related to the Pleistocene cooling. Currently, three species of Eucalathis occur in the NE Atlantic, near the entrance to the Mediterranean. However, they are unable to enter because the distribution of all three species is restricted to deeper environments than the average 300 m deep Gibraltar sill. The circulation pattern of the Gibraltar area also prevents deeper water Atlantic species from entering the Mediterranean. [ABSTRACT FROM AUTHOR]

While the established paradigm of human evolution asserts that the lineages leading to the extant great apes and Homo arose in Africa, the large number of fossil discoveries from Europe in recent decades support arguments for a European origin of the Hominidae (all great apes) and plausibly, also a European common ancestor of the Homininae (African great apes, Australopithecus species, and the genus Homo). Meanwhile, a lack of consensus remains regarding the phylogenetic placement of australopithecine fossil species in Africa, with substantial evidence indicating that some of them may align more closely to extant African great apes than to Homo. Based on a novel interpretation of existing fossil, genetic, paleogeographic and paleoclimatic evidence, this paper aims to put forward a new hypothesis regarding the separate divergences of Gorilla, Pan, and Homo. We support existing arguments that the last common ancestor of African great apes and Homo may have lived in Europe in the late Miocene, and we put forward a new hypothesis as to where, when, and why the separate lineages may have started to diverge. Extreme conditions during the Vallesian Crisis (11.6-8.0 Ma) and the Messinian Salinity Crisis (6.0-5.3 Ma) may have forced separate branches of European hominids to migrate out of the Mediterranean region. We argue that the lineages leading to Gorilla and Pan independently migrated into Africa, while the lineage leading to Homo went in another direction. Thereafter, the Zanclean Megaflood (5.3 Ma)--which caused the Mediterranean to refill very quickly--may have cut off the migration route between Eurasia and Africa at the Sinai Peninsula, isolating a small population (the putative Homo lineage) on the Arabian Peninsula / Red Sea coast during a period of hyperaridity. The other group (Pan lineage) crossed into Africa, where it subsequently diversified into various species of Australopithecus.

The late Miocene‐early Pliocene (7.4‐4.5 Ma) is a key interval in Earth's history where intense reorganization of atmospheric and ocean circulation occurred within a global cooling scenario. The Southern African monsoon (SAFM) potentially played an important role in climate systems variability during this interval. However, the dynamics of this important atmospheric system is poorly understood due to the scarcity of continuous records. Here, we present an exceptional continuous late Miocene to early Pliocene reconstruction of SAFM based on elemental geochemistry (Ca/Ti and Si/K ratios), stable isotope geochemistry (δ18O and δ13C recorded in the planktonic foraminifera Orbulina universa), and marine sediment grain size data from the International Ocean Discovery Program (IODP) Site U1476 located at the entrance of the Mozambique Channel. Spectral characteristics of the Si/K ratio (fluvial input) was used to identify the main orbital forcing controlling SAFM. Precession cycles governed precipitation from 7.4 to ∼6.9 Ma and during the early Pliocene. From ∼6.9 to ∼5.9 Ma, the precession and long eccentricity cycles drove the SAFM. The major Antarctic ice sheet expansion across this interval appear to influence the isotopic records of O. universa imprinting its long‐term variability signal as a response to the ocean and atmospheric reorganization. Precession cycles markedly weakened from 5.9 to 5.3 Ma, almost the same period when the Mediterranean Outflow Water ceased. These findings highlight important teleconnections among the SAFM, Mediterranean Sea, and other tropical regions. Key Points: From 7.4 to 4.5 Ma, the Southern African Monsoon (SAFM) was driven by precession and the long eccentricity cycles related to internal feedbacksWeak Northern African monsoon was coeval with intense SAFM during the acme of the Messinian Salinity CrisisThe SAFM intensity and Orbulina universa responded to changes in the Antarctic ice sheet during the latest Miocene [ABSTRACT FROM AUTHOR]

The evolution of marine gateways and sea straits exerts major control on bottom current depositional systems. A well‐known interval in geological history characterized by frequent changes in marine connectivity is the Messinian Salinity Crisis (5.97 to 5.33 Ma) when the Mediterranean allegedly experienced major (>1 km) sea‐level drawdown followed by a catastrophic marine replenishment at the base of the Zanclean. Controversy exists around the timing and mode of this event as unambiguous flood deposits have so far never been drilled or recognized in outcrops. In the Sicilian Caltanissetta Basin (Italy), the Messinian/Zanclean boundary is directly underlain by the Arenazzolo Formation. This 5 to 7 m thick sandy sedimentary interval may reveal a genetic link with the abrupt refilling of the Mediterranean, but at present a detailed study to understand its origin is lacking. In this work, the Arenazzolo Formation at Eraclea Minoa has been studied by a multi‐method approach, employing detailed facies description, grain‐size analyses, petrographic analyses and palaeocurrent analyses. Palaeogeographical reconstructions and facies associations show that the Arenazzolo Formation sands were deposited on the northern flank of the Gela thrust front by persistent bottom currents, flowing parallel to the regional slope physiography, during a transgression. It is hypothesized that these currents are associated with the active circulation of surface and intermediate water masses coeval with a terminal Messinian flood, when basin margins overtopped and a reconnection between western and eastern Mediterranean was created. The Arenazzolo Formation is a unique example of a contouritic deposit formed by bottom currents that establish during the reconnection of major isolated water bodies. [ABSTRACT FROM AUTHOR]

Tectonic, paleoenvironmental, and paleoclimatic unstable conditions preceding the onset of the Messinian Salinity Crisis (MSC) highly affected marine life. Changes in calcareous plankton association are overall registered in the Mediterranean. They consist of a general transition from abundant and well-diversified planktonic associations to strictly oligotypic assemblages that precede their total disappearance at the onset of evaporitic precipitation. In this work, an accurate quantitative analysis of calcareous plankton, both foraminifers and nannofossils, has been carried out in the Torrente Vaccarizzo Section of Sicily (southern Italy). The aim is to independently define a chronostratigraphic pattern of bioevents preceding the MSC in the absence of magnetostratigraphic or radiometric constraints. The fluctuating abundance of the genus Orbulina fits well with the 100 ky Eccentricity maxima, and it is successfully applied to build an astronomically calibrated age-model for the section. On this basis, all the biohorizons have been recalibrated and discussed with regard to the previous literature. Abundant influxes of selected species demonstrated to be of local significance since they are highly affected by paleoenvironmental and paleoclimatic conditions. A chronological sequence of foraminifer and nannofossil events marks the onset of the MSC with a derived age of 5.957 My, which agrees well with previous findings from other Mediterranean sections. This methodology and the new biostratigraphic events may be useful for future studies on pre-evaporitic successions of the Mediterranean. [ABSTRACT FROM AUTHOR]

Twelve available two-way time high-resolution seismic reflection profiles located in the central part of the middle Rhône valley are interpreted. In addition, one of the profiles was reprocessed to determine the P-wave velocities of the main geological units and to convert this profile into a depth cross section. The Lower and Upper Cretaceous units are clearly identifiable on all the profiles, along with the Messinian Erosion Surface (MES) carved out during the Messinian Salinity Crisis (MSC) by the paleo-Rhône and its western tributaries, the Ardèche and Cèze paleo-canyons. The Plio-Quaternary fill of these paleo-canyons shows at least 4 main units with an overall transgression. The combination of geological data from geological maps, geological field surveys and borehole data made it possible to model the MES in 3D at the scale of the region, and to produce depth/elevation model. From a geological point of view, the interpretation of the seismic profiles enabled us to reconstruct the stages in the sub-aquatic filling of the Messinian-Pliocene aggradation of the paleo-river. Several Mass Transport Deposits (MTDs) were identified both during the drop and during the rise in the Mediterranean Sea level. From a geomorphological point of view, this study provides new insights in the route and longitudinal profile of paleo-rivers and, in particular, it deepens the profile of the Paleo-Rhône at the latitude of the Tricastin region (up to −700 m b.s.l.) and significantly modifies the course and depth of the Ardèche proposed in previous studies. The N-Ardèche river, known to develop a karstic system during the MSC, is connected to a deep canyon, most likely through a karstic pocket valley, as suggested by the very steep longitudinal profile of the MES. Finally, from a structural point of view, our interpretation of the seismic profiles shows a broad ENE-trending anticline structure associated with a normal fault which apparently did not affect the Mio-Pliocene fill. In the southern part of the area, near the Uchaux anticline, the imaged structures suggest the presence of a recent (syn- to post-Pliocene) fault propagation fold. In addition to all the new information on the geology, morphology and methods of excavation and filling of the Messinian paleo-canyon, the proposed topographic model of the paleo-canyon is crucial for modelling seismic movement in the context of a basin with a complex geometry and, in particular, for the numerical assessment of site effects in a context of low seismicity.

The Messinian Salt Giant in the Provençal Basin represents a good example to study salt tectonics: salt deposition occurred throughout the basin well after basin opening, with a tectonic context stable since ∼16 Ma, in a closed system. Also, the youth of salt tectonics has led to less mature structures and an evolutionary history that is easier to decipher than in older salt-bearing margins. We conducted an analysis of the chronology of salt deformation, from its deposition to the present-day, thanks to the basin-wide correlation of the Late Miocene and Pliocene-Pleistocene stratigraphic markers. The large seismic dataset provided detailed analysis of the causes and timing of salt deformation at a regional level. The salt tectonics started relatively early, during the Messinian Upper Unit (UU) deposition (phase 1) in the deepest part of the basin. From the Pliocene to the present-day, salt movement is divided into two more main phases (phases 2 and 3), the first of small intensity, occurred during the Pliocene and the second, more intense, during the Pleistocene. The geometric relationship between salt tectonics and crustal nature domains has revealed, regardless of the timing deformation phases, a more rapid and intense salt deformation above the proto-oceanic crust domain than in the continental or transitional crust domain. This observation, remaining unexplained, emphasizes the role of the influence of crustal nature, associated thermal regime and fluid circulation system on salt tectonics.

The Central Mallorca Depression (CMD) located in the Balearic Promontory (Western Mediterranean) contains a well‐preserved evaporitic sequence belonging to the Messinian Salinity Crisis (MSC) salt giant, densely covered by high‐ and low‐resolution seismic reflection data. It has been proposed recently that the MSC evaporitic sequence in the CMD could be a non‐deformed analogue of the key MSC area represented by the Caltanissetta Basin in Sicily. This presumed similarity makes the CMD an interesting system to better understand the MSC events. Physics‐based box models of the water mixing between sub‐basins, built on conservation of mass of water and salt, help constrain the hydrological conditions under which evaporites formed during the MSC. Those models have been widely used in the literature of the MSC in the past two decades. They have been mostly applied to the Mediterranean Sea as a whole focusing on the Mediterranean–Atlantic connection, or focusing on the influence of the Sicily Sill connecting the Western and Eastern Mediterranean Sea. In this study, we apply a downscaled version of such modelling technique to the CMD. First, we quantify the present‐day volumes of the MSC units. We then use a reconstructed pre‐MSC paleo‐bathymetry to model salinity changes as a function of flux exchanges between the CMD and the Mediterranean. We show that a persistent connection between the CMD and the Mediterranean brine near gypsum saturation can explain volume of Primary Lower Gypsum under a sea level similar to the present. For the halite, on the contrary, we show that the observed halite volume cannot be deposited from a connected CMD‐Mediterranean scenario, suggesting a drawdown of at least 850 m (sill depth) is necessary. Comparison between the deep basin halite volume and that of the CMD shows that it is possible to obtain the observed halite volume in both basins from a disconnected Mediterranean basin undergoing drawdown, although determining the average salinity of the Western Mediterranean basin at the onset of drawdown requires further investigation. [ABSTRACT FROM AUTHOR]

The biogeochemical cycles of carbon and silicon are tightly coupled in modern marine environments due to the pivotal role of planktonic diatoms. Once diatoms are sedimented, the fate of organic matter and biogenic silica is initially governed by bottom water oxygen levels and bacterial communities involved in remineralization processes. The early diagenesis of biosiliceous sediments may result in drastic changes in composition, sometimes hampering palaeoenvironmental reconstruction. Sedimentary successions deposited in the Mediterranean region during the Messinian salinity crisis (5.97–5.33 Ma) allow to explore the early diagenetic transformation of organic matter and biogenic silica in a restricted basin experiencing a severe palaeoceanographic turnover. Sedimentological and petrographic observations coupled to elemental, mineralogical, inorganic and organic geochemical analyses were carried out on biosiliceous (diatomaceous shales and diatom‐bearing mudstones) and associated clay‐rich and dolomite‐rich sediments (dolomitic mudstones) interbedded with primary gypsum layers in the Piedmont Basin (north‐west Italy). The state of preservation of biogenic silica was governed by different pathways of organic matter remineralization, depending on bottom and pore water redox conditions, which were controlled by the structure of the water column. Aerobic respiration of organic matter in a mixed water column and an oxygenated seafloor promoted biogenic silica preservation. In contrast, bottom water anoxia induced by permanent stratification of the water column favoured the concomitant formation of dolomite and authigenic clays. In particular, organic matter degradation through bacterial sulphate reduction increased pore water alkalinity, promoting the precipitation of dolomite. At the same time, the rise in pH promoted the dissolution of biogenic silica which, reacting with pore water cations, ultimately caused the formation of authigenic clays. This study suggests that the apparent annihilation of Mediterranean marine biota during the Messinian salinity crisis partially reflects an early diagenetic bias produced by interactions of the carbon, silicon and sulphur biogeochemical cycles in a restricted basin. [ABSTRACT FROM AUTHOR]

The current distribution patterns of many Mediterranean species are often a consequence of large and impactful past geoclimatic events, such as the Messinian Salinity Crisis (MSC) and the Quaternary glacial cycles. Cicadas are flying insects with poor dispersal ability, which have experienced intense local differentiation in the Mediterranean, where the genus Tettigettalna has surfaced as a biogeographic model. The genus includes 10 species with species‐specific calling songs but identical morphology. All Tettigettalna species are restricted to Southern Iberia, with the exception of T. estrellae (northwest Iberia), the widespread T. argentata (mainly Iberia, France and Italy), and T. afroamissa (Morocco). With an expanded genetic dataset involving nuclear (EF1α) and mitochondrial (5′ and 3′ COI and ATP) loci, we reconstructed the phylogeny of the genus and estimated divergence dates for Tettigettalna species under a Bayesian framework. Phylogeny with the new mitochondrial dataset was in agreement with previous studies, whereas the nuclear EF1α supported T. josei and T. afroamissa as monophyletic clades but lacked resolution to resolve the remaining taxa. Some sister taxa share mitochondrial haplotypes, hinting for incomplete lineage sorting. Estimates of divergence time settled T. josei as the earliest diverging lineage, likely as a pre‐ or early‐MSC event. As for the origin of T. afroamissa in Morocco, though time estimates could not entirely rule out post‐MSC dispersal, the most likely scenario points to isolation of African Tettigettalna after the reopening of the strait of Gibraltar. The Pleistocene glaciations that followed likely impacted on the diversification of the remaining species of the genus in southern Iberia refugia. [ABSTRACT FROM AUTHOR]

Field analogues illustrating the early stage of deformation of shortening structures in salt‐bearing orogenic fold‐and‐thrust belts are not yet well illustrated in literature. The relatively young Messinian salt of the Algerian basin could represent a good case example of such systems. The Algerian Basin is a deep‐water Miocene back‐arc basin including a layer of mobile Messinian evaporites up to 2 km thick. The Messinian salt was deposited in an already inverted basin, after its extensive stage. Its inversion initiated in the Tortonian, with a N‐NW shortening induced by the north‐westward motion of the African plate. In this study, we use new 2D multichannel seismic data to identify, classify and map salt structures throughout the central Algerian Basin and re‐assess its structural style. We interpret contractional salt tectonic structures, such as buckle folds, squeezed diapirs and related salt sheets as evidence of regional thick‐skinned shortening episodes. We conclude that horizontal displacement loading has driven salt deformation within the basin since its deposition. We also show spatial variations in the structural style of the central Algerian Basin, both along‐ and down‐dip. We demonstrate that the initial shortening‐related salt deformation in the late Messinian was focussed along the Algerian margin and later shifted outward toward the Balearic margin in the Plio‐Quaternary. The shifting of the deformation front is interpreted to be a result of the thickening and strengthening of the overburden. The second peak of deformation may have reactivated faults along the Emile‐Baudot escarpment with thick‐skinned deformation. We also observe a variation in the intensity of the salt deformation along the margin from SW to NE, which may be due to variable tectonic loading applied along the Algerian margin or the pre‐shortening distribution of salt. [ABSTRACT FROM AUTHOR]

Small mammal remains have been discovered in three new outcrops from the Afoud locality. This site is significant from a paleontological point of view because it has yielded fauna of European affinities (Castillomys , Occitanomys , Stephanomys, Eliomys, and Prolagus). The presence of two different species of Arvicanthis from the Afoud locality represents the oldest record in North Africa. The association of small mammals from the AF12-2 deposits provides an approximate view of the local environment and climate at the time of the formation of this locality. This faunal assemblage indicates a warm, temperate, semi-arid climate and an open vegetation cover of wooded savannah. Fossil assemblages documented in this paper expand our knowledge on the Late Miocene–Early Pliocene small mammal communities of the Aït Kandoula Basin. This study provides useful data for understanding the paleobiogeography of the Mediterranean region and faunal exchanges between North Africa and the Iberian Peninsula that took place before the Messinian Salinity Crisis. [ABSTRACT FROM AUTHOR]

The definition of pre-Messinian source rocks in the eastern Mediterranean is of paramount importance for hydrocarbon exploration because of the ability of salt to act as a high-quality seal rock. This research evaluates the organic geochemical features of the Upper Miocene (Tortonian—Messinian) sedimentary succession onshore Crete Island, Greece. The study employs original (Messinian, Agios Myron Fm) and published (Tortonian, Viannos Fm, Skinias Fm, Moulia Fm, and Messinian Ploutis section) results from organic geochemical analyses of mudstone samples. One hundred and one samples were examined using standard organic geochemistry methodology (Rock-Eval II and VI-TOC) to define the origin, type, and degree of organic matter maturity. The data indicate that the studied samples have poor to fair gas-prone source rock potential. These possible source rock units have not experienced great temperatures during burial, and, thus, their organic matter is thermally immature. The sub-salt (Tortonian—Messinian) source rock units are likely to be of higher thermal maturity in the western and eastern south Cretan trenches because of tectonic subsidence and a thicker sedimentary overburden. Several traps can grow in these regions, associated with normal faults, rotated blocks and unconformities (both below and above the unconformities). This research provides a basis for the further evaluation of the hydrocarbon potential in Crete Island. It is an area that shares geological similarities with the surrounding regions that contain proven reserves and is of crucial economic and strategic importance. [ABSTRACT FROM AUTHOR]

The Messinian Salinity Crisis (5.97-5.33Ma) may be one of the most significant periods of sea-level change in recent geologic history. During this period, evaporite deposition throughout the Mediterranean basin records a series of dramatic environmental changes as flow through the Strait of Gibraltar was restricted. In the first stage of evaporite deposition, cycles of gypsum appear in shallow basins on the margins of the Mediterranean. The complex environmental history giving rise to these cycles has been investigated for decades but remains controversial. Notably, whether the evaporites are connected to significant changes in Mediterranean sea level is an open question. In one proposed model, competition between tectonic uplift and erosion at the Strait of Gibraltar gives rise to selfsustaining sea-level oscillations—limit cycles—which trigger evaporite deposition. Here I show that limit cycles are not a robust result of the proposed model and discuss how any oscillations produced by this model depend on an unrealistic formulation of a key model equation. First, I simplify the model equations and test whether limit cycles are produced in 64 million unique combinations of model parameters, finding oscillations in only 0.2% of all simulations. Next, I examine the formulation of a critical model equation representing stream channel slope over the Strait of Gibraltar, concluding that a more realistic formulation would render sea-level limit cycles improbable, if not impossible, in the proposed model.

The Messinian Salinity Crisis (MSC) resulted from changes in the Atlantic-Mediterranean connectivity in the Alboran Basin, a region with a complex and debated geodynamic configuration. Since the MSC, this basin's topography and its record of the Messinian Erosional Surface have been subject to vertical motions due to sediment accumulation, tectonic deformation, isostasy, and latent effects of thermal cooling after extension and magmatic arc formation. The objective of this work is to restore these contributions to post-Messinian subsidence in order to quantify the original depth of formation of the MSC features. We do this by performing a pseudo-3D planform flexural isostatic reconstruction of the Messinian Erosion Surface mapped from an extensive set of seismic reflection data. We focus on identifying the most likely position of the gateway between Atlantic and Mediterranean, the effect of a drawdown on gateway topography and connectivity, and the depth of proposed erosional features related to the Messinian lowstand. The results indicate that the depth of the Alboran Basin by the end of the Messinian was about 500 m shallower than nowadays, but over 500 m deep on average, reaching depths of >1000 m in most subbasins, even when accounting for the possible ∼300 m isostatic rebound caused by water unloading in a largely desiccated Alboran Sea during the MSC. Although these results are compatible with volcanic cones locally emerging above sea level at the East Alboran Volcanic Arc during the Messinian, several lows remaining in the reconstruction suggest that that region is unlikely to have been the sill between Atlantic and Mediterranean at that time, unless the basin saw unconstrained dynamic topography contributions of over −500 m. Full desiccation of the Alboran Basin implies an uplift of up to 100 m at the Strait of Gibraltar, and uplift rates too high to be counteracted by erosion, suggesting that full disconnection and the main corresponding evaporative drawdown took place only once. The terraces and canyons in the West Alboran are restored to depths between 250 and 550 m (shallowest terrace) and 750–1500 m (deepest terrace), and cannot be clearly linked to a single, stable water level during the MSC, pointing to climate-controlled variations in the water level during the isolation phase. • The depth of the Alboran Basin during the MSC was similar to the modern day. • Erosional features formed from <500 to >1000 m depth. • Atlantic-Mediterranean barrier in the Gibraltar arc, not the Alboran Volcanic Arc. [ABSTRACT FROM AUTHOR]

Tectonic, paleoenvironmental, and paleoclimatic unstable conditions preceding the onset of the Messinian Salinity Crisis (MSC) highly affected marine life. Changes in calcareous plankton association are overall registered in the Mediterranean. They consist of a general transition from abundant and well-diversified planktonic associations to strictly oligotypic assemblages that precede their total disappearance at the onset of evaporitic precipitation. In this work, an accurate quantitative analysis of calcareous plankton, both foraminifers and nannofossils, has been carried out in the Torrente Vaccarizzo Section of Sicily (southern Italy). The aim is to independently define a chronostratigraphic pattern of bioevents preceding the MSC in the absence of magnetostratigraphic or radiometric constraints. The fluctuating abundance of the genus Orbulina fits well with the 100 ky Eccentricity maxima, and it is successfully applied to build an astronomically calibrated age-model for the section. On this basis, all the biohorizons have been recalibrated and discussed with regard to the previous literature. Abundant influxes of selected species demonstrated to be of local significance since they are highly affected by paleoenvironmental and paleoclimatic conditions. A chronological sequence of foraminifer and nannofossil events marks the onset of the MSC with a derived age of 5.957 My, which agrees well with previous findings from other Mediterranean sections. This methodology and the new biostratigraphic events may be useful for future studies on pre-evaporitic successions of the Mediterranean.

Extensive canyons, excavated into the margins of the Levant Basin during the Oligocene–Miocene, are interesting case-studies for canyon fills in carbonate settings. The carbonate Pattish Formation, developed along the margins of the pre-evaporitic Messinian Beer Sheva Canyon in Israel, was investigated using both onshore seismic imaging and field data. The canyon has three main seismic facies of fill (1) Subparallel reflections mimicking the canyon´s morphology; (2) chaotic reflections overlying the subparallel ones, and (3) sigmoidal reflections, locally with sharp edges at the canyon margins. The first seismic facies corresponds to the pelagic marls of the Bet Eshel Formation. The other two seismic facies are, respectively, equivalent to bioclastic calcarenite clinobeds with slumps and channels, and to coral–stromatolite reefs and reef slopes of the Pattish Formation observed at outcrop. There were three phases of canyon development: (1) slope incision and headward erosion due to tectonic uplift and eustatic sea-level fall during the Early Oligocene, and large slope failure during the latest Middle Miocene; (2) platform incision and connection with a fluvial system in the Late Miocene related to falling sea level and tectonic uplift; and (3) canyon filling first by pelagic marls at the centre of the canyon, followed by calcarenite clinobeds at the canyon flanks formed by gravity flows. Finally, carbonate production at the margins of the canyon resulted in reefs and associated slopes prograding towards the canyon axis. The late canyon filling phase ended with the deposition of evaporites during the Messinian Salinity Crisis. [ABSTRACT FROM AUTHOR]

The present study proposes a characterization of textural and mineralogical features of branching selenite gypsum lithofacies from the Monferrato area (NW Italy). This facies is considered to have appeared during the sequence of the primary lower gypsum of the Messinian Salinity Crisis starting from the sixth stratigraphical cycle, providing a useful tool for stratigraphic correlation throughout the Mediterranean. It is often associated with thick and continuous beds that are exploited by open-pit and underground quarries. We provide the results of a characterization of the non-gypsum minerals that represent approximately 10% in weight of the rock. Mineralogical data were collected with energy-dispersive X-ray spectroscopy (EDS) and X-ray powder diffraction (XRPD) techniques. In addition, a detailed characterization of clay families was performed through dedicated XRPD analyses. Results highlight the presence of detrital minerals (quartz, feldspars and clay minerals), calcite, dolomite and epsomite. Clay analysis registered the existence of five different families (illite, chlorite, smectite and two mixed layers). These mineralogical data suggest that the origin of this lithofacies is connected to both depositional processes and in situ precipitation processes under different conditions of saturation of the brine (respectively undersaturated and supersaturated in sulfates). The mineralogical and textural heterogeneity also represents an important element in controlling the mechanical response of the material and the consequent stability of quarry sites. [ABSTRACT FROM AUTHOR]

During the Messinian Salinity Crisis (MSC, 5.97–5.33 Ma), thick evaporites were deposited in the Mediterranean Sea associated with major margin erosion. This has been interpreted by most authors as resulting from water level drop by evaporation but its timing, amplitude and variations between subbasins are poorly constrained due to uncertainty in post‐Messinian vertical motions and lack of a clear time‐correlation between the marginal basin and offshore records. The Balearic Promontory and surrounding basins exemplify a range of responses to this event, from margin erosion to up to a kilometre thick Messinian units in the abyssal areas containing the majority of the MSC halite. The Balearic Promontory contains unique patches of halite with thickness up to 325 m at intermediate depths that provide valuable information on water level during the stage of halite deposition. We compile seismic markers potentially indicating ancient shorelines during the drawdown phase: the first is marked by the transition from the MES to UU based on seismic data. The second is the limit between the bottom erosion surface (BES) and abyssal halite deposits. We restore these shorelines to their original depth accounting for flexural isostasy and sediment compaction. The best‐fitting scenario involves a water level drop of ca. 1,100 ± 100 m for the Upper unit level and 1,500 ± 100 m for the BES level. According to our results, halite deposition began in the Central Mallorca Depression at 1,300–1,500 m depth, perched hundreds of metres above the deep basins, which were at 1,500–1,800 m (Valencia Basin) and >2,900 m (Algerian Basin). The hypothesis that erosion surfaces were formed subaerially during the drawdown phase is consistent with a model of halite deposition before/during the water level drop of at least 1,000 m, followed by the deposition of the Upper unit until the MSC is terminated by the reinstatement of normal marine conditions. [ABSTRACT FROM AUTHOR]

The Mediterranean Basin has experienced extensive change in geology and climate over the past six million years. Yet, the relative importance of key geological events for the distribution and genetic structure of the Mediterranean fauna remains poorly understood. Here, we use population genomic and phylogenomic analyses to establish the evolutionary history and genetic structure of common wall lizards (Podarcis muralis). This species is particularly informative because, in contrast to other Mediterranean lizards, it is widespread across the Iberian, Italian, and Balkan Peninsulas, and in extra-Mediterranean regions. We found strong support for six major lineages within P. muralis , which were largely discordant with the phylogenetic relationship of mitochondrial DNA. The most recent common ancestor of extant P. muralis was likely distributed in the Italian Peninsula, and experienced an "Out-of-Italy" expansion following the Messinian salinity crisis (∼5 Mya), resulting in the differentiation into the extant lineages on the Iberian, Italian, and Balkan Peninsulas. Introgression analysis revealed that both inter- and intraspecific gene flows have been pervasive throughout the evolutionary history of P. muralis. For example, the Southern Italy lineage has a hybrid origin, formed through admixture between the Central Italy lineage and an ancient lineage that was the sister to all other P. muralis. More recent genetic differentiation is associated with the onset of the Quaternary glaciations, which influenced population dynamics and genetic diversity of contemporary lineages. These results demonstrate the pervasive role of Mediterranean geology and climate for the evolutionary history and population genetic structure of extant species. [ABSTRACT FROM AUTHOR]

Bovidae from the Messinian localities of Italian Peninsula are poorly known and documented except in the Monticino Quarry record (Brisighella, central Italy), which represents, therefore, a key locality for the understanding of Bovidae diversity and biogeography during the Messinian in Italy. In the present paper, the material published in the 1989 along with new, unpublished bovid material from Monticino Quarry are revised in order to clarify the taxonomic attribution of the Monticino' bovids and to better understand their geographic and chronological distribution, as well as the Bovidae diversity in Italy during the Messinian. The updated and revised data on the Italian small, spiral-horned antelope from Monticino Quarry, and the additional specimens from Borro Strolla (Tuscany) and Verduno (Piedmont), suggests it has to be referred to the genus Oioceros, i.e., Oioceros occidentalis nov. comb. In addition, the presence of four other bovids at Monticino is attested here: Bovidae indet. (small-size), Bovinae indet. (medium-large size cf. Boselaphini), Bovinae indet. cf. Parabos, and Antilopinae indet. (medium size). The biochronological correlations and paleobiogeographic implications of these findings are discussed, suggesting a close affinity in faunal assemblage among the northern and central Italian latest Messinian localities and supporting the dispersal of bovids from East to West during the Messinian (before and during the Messinian Salinity Crisis). [ABSTRACT FROM AUTHOR]

Aim: In the Mediterranean biodiversity hotspot, freshwater fishes are key biogeographic indicators, while their habitats are vulnerable to ongoing anthropogenic pressures. The freshwater blenny is a widespread endemic taxon in this area. However, our understanding of the overall diversity of specific populations and their phylogenetic relationships is sparse. Thus, we aim to investigate the genetic diversity, infer relationships among sampled populations related to major palaeoenvironmental changes and suggest insights for future research and conservation targets. Location: Mediterranean Basin Methods: We studied 171 Salaria specimens from 51 rivers and lakes, and from 13 countries surrounding the Mediterranean Sea. For assessing the phylogenetic relationships among different locations, we used mtDNA d‐loop and intron S7 sequences. Furthermore, we inferred absolute divergence times and demographic changes using secondary calibrations and investigated the diversity within major lineages using haplotype networks and several geographical and genetic clustering methods. Results: We found eight well‐differentiated lineages, each of which being confined to a particular geographical region. The onset of the freshwater blenny radiation was dated around the Messinian salinity crisis. Further differentiation happened during the Plio‐ and Pleistocene with signatures of population expansion after the Last Glacial Maximum in some of the main lineages. Main conclusions: Due to their unusually widespread distribution, the freshwater blennies represent an excellent model for studying phylogeographic structure across the Mediterranean basin biodiversity hotspot. The extant diversity and distribution of the freshwater Salaria species mirrors palaeoenvironmental changes in the region, but there are still large gaps in knowledge, particularly in the Levant. Even though the main lineages described are statistically well supported, the phylogenetic relationship among several of them remains poorly resolved. Despite the fact that the most widespread species, S. fluviatilis, is not globally threatened, it harbours some distinct populations that are of conservation concern. [ABSTRACT FROM AUTHOR]

Recent studies on the genesis of sedimentary native sulfur deposits indicate diagenetic mid‐low temperature Bacterial Sulfate Reduction (BSR) as the main process, involving organic compounds (kerogen/hydrocarbons), bacterial colonies and gypsiferous rocks. In the peri‐Mediterranean area (Southern Spain, Sicily, Northern Apennines, Israel), the main sulfur accumulations are always associated with late Miocene sulfates and organic‐rich successions encompassing the Messinian salinity crisis (MSC). In particular, the Messinian successions of the Apennine‐Adriatic foreland basin system, due to a large amount of high‐resolution stratigraphic data, represent a perfect case study for understanding the diagenetic conditions controlling the development of the BSR process during sedimentary basin evolution. In this work, thermal models performed in three sub‐basins in a sector of the Northern Apennines comprised of the Sillaro and Marecchia rivers (Italy), calibrated by means of organic and inorganic geothermometers, indicate a general thermal immaturity of the studied successions attained as a result of a constant heat flow similar to the present day one (ca. 40 mW/m2) since Late Tortonian and lithostatic loads between 615 and 1,710 m depending on different sub‐basins. These results suggest that the MSC deposits experienced maximum temperatures between about 39°C and 65°C. Temperatures derived from thermal models have been used to constraint occurrence of the diagenetic BSR associated with evaporitic deposits providing thermal constraints in sulfur genesis as well as new useful thermal‐constraints for basin analysis studies. [ABSTRACT FROM AUTHOR]

Evaporites are typically described as impermeable seals that create some of the world's highest reservoir pressures beneath the salt seal. However, several laboratory studies demonstrate that evaporites can retain open pore spaces that hydraulically connect the sediments above and below them in sedimentary basins. During the Messinian Salinity Crisis (5.97–5.33 Ma), up to 2,400 m thickness of evaporites were rapidly deposited in the Western Mediterranean, which may have generated high pore fluid overpressure in the basin sediments. Here we use one‐dimensional numerical modelling to quantify the temporal evolution of overpressure at two distinct locations of the Western Mediterranean, the Liguro‐Provençal and Algero‐Balearic basins, from the Miocene to Present. We reconstruct the sedimentation history of the basin, considering disequilibrium compaction as an overpressure mechanism and constraining model parameters (such as permeability and porosity) using laboratory experiments and the literature. In the Liguro‐Provençal basin the highest overpressure of 11.2 MPa occurs within the halite during deposition of Pliocene to Quaternary sediment, while in the Algero‐Balearic basin at the base of the Emile Baudot Escarpment, the highest overpressure of 3.1 MPa also occurs within the halite but during stage 3 of the Messinian Salinity Crisis (5.55–5.33 Ma). In the Algero‐Balearic basin an overpressure of 3.1 MPa could have been sufficient to hydro fracture the sediments, which agrees with the development of fluid escape features observed on seismic reflection profiles. In general, our models with evaporite deposition rates above 20 m kyr−1 and permeabilities below 10–18 m2, suggest that high overpressure, approaching lithostatic, can be generated in salt basins. [ABSTRACT FROM AUTHOR]

We present a new 87Sr/86Sr curve for the time interval between 6.5 and 5.5 Ma of the Ain El Beida section (AEB, Atlantic side of Morocco). The location and the precessional‐scale resolution of our curve allow to investigate the possible relationships between global paleoclimatic and palaeoceanographic trends and the high‐amplitude hydrological changes of the Mediterranean during the Messinian salinity crisis (MSC). The values, obtained from planktic foraminifers, plot in the upper portion of the global ocean curve and show long‐ (at eccentricity scale) and short‐term (precessional scale) oscillations; the latter are more evident during the maximum of eccentricity for both the 100 and 400 ka components, between 6.00 and 5.85 Ma. Negative 87Sr/86Sr anomalies are observed at insolation minima in phase with δ18O maxima, marking the glacial stages from TG34 to TG26. It is unclear whether these oscillations originated from local (river runoff) or global (glacial/interglacial) climatic forcing. Conversely, the evaporites of the coeval Primary Lower Gypsum (5.97–5.62 Ma) accumulated in the Mediterranean at insolation minima show positive anomalies at glacial stages TG32, 30, 28, and 26. Such an opposite trend is possibly related to the greater sensitivity of the Mediterranean Sea to river runoff during insolation maxima compared to the global ocean, as recorded at the Moroccan Atlantic margin. These observations and the deviation toward lower values of the Mediterranean Sr isotope curve starting at around 6.5 Ma, support the hypothesis of a progressive restriction of the Atlantic connections since the early Messinian, leading to the MSC. Key Points: A new planktic foraminifers‐based high‐resolution 87Sr/86Sr reference curve for the Atlantic (Ain El Beida section, Morocco) is providedLong‐ (at eccentricity scale) and short‐term (precessional scale) oscillations characterize the 87Sr/86Sr curveNegative 87Sr/86Sr anomalies are observed at insolation minima in phase with marine isotopic glacial stages from TG34 to TG26 [ABSTRACT FROM AUTHOR]

The genus Pygopleurus includes some of the most important beetle pollinators of the Eastern Mediterranean islands, showing a particular affinity to red-guild flowers. Focussing on the Aegean Archipelago, 75 individuals belonging to five Pygopleurus species were sampled and screened for two mtDNA loci (COI and 16S). The phylogenetic analysis implemented both by Bayesian and Maximum Likelihood approaches, showed a deep and robust structure for three species, mostly reflecting the palaeogeographic history of the islands. Interspecific divergence among them appears to have emerged at the beginning of the Messinian Salinity Crisis (5.99-5.33 Mya), with differentiation dating back to the eustatic sea level changes of the Pleistocene (2.56-0.0117 Mya). For P. foina, the most abundant species in the Aegean Archipelago, a finer sampling (121 individuals from 8 islands) and analysis of COI locus showed a distinct separation between northern and southern islands of East Aegean that disconnected non-simultaneously from the continent during the Pleistocene. A strong geographic structure was also evident at smaller taxonomic scales, with island populations exhibiting unique haplotypes. Thus, we concluded that the major palaeogeographic events of the region are reflected in the observed patterns of inter- and intraspecific divergence among and within Pygopleurus species in the Aegean Archipelago, something that is facilitated by this genus' low dispersal ability. [ABSTRACT FROM AUTHOR]

The bulk mineralogy and geochemistry of mudrock samples from a representative section of the Calcare di Base Formation outcropping in the Catanzaro Basin (Calabria) were studied using X-Ray Diffraction and X-Ray Fluorescence techniques. Phyllosilicates are the main mineralogical constituents followed by quartz, calcite, feldspar, hematite and gypsum in the lower part of the studied succession, whereas calcite is the main mineralogical component in the upper part. The geochemical composition reveals that SiO 2 , Fe 2 O 3 , Al 2 O 3 , CaO and MgO are the most abundant oxides on average. The chemical weathering indices, such as the CIA (Chemical Index of Alteration) and its modifications, show low-medium values and thus suggest weak-moderate source area(s) weathering conditions, under a mainly dry/arid paleoclimate environment. The low kaolinite amounts confirm a low-moderate degree of weathering processes. The studied samples originate from an environment in which non-steady-state weathering conditions prevailed and active tectonism allows erosional processes. This trend suggests a paleoclimate characterized by persistent dry (arid) conditions alternating with relatively wet (humid) conditions. The studied mudrocks are characterized by recycling effects as recorded in the Al–Zr–Ti plot. The provenance of the studied samples is mainly related to a felsic source such as the rocks composing the Sila Massif. The composition of the studied samples was compared to that of similar samples from the Calcare di Base formation of the Rossano Basin to reconstruct the paleoclimate scenario of the Mediterranean area during the Messinian Salinity Crisis. The analogies between the mudrocks investigated in this work and the argillaceous marls from the Calcare di Base formation of the Rossano Basin suggest that they formed in the same (i) non-steady-state weathering and (ii) dry/arid paleoclimatic conditions. All the evidences allow to hypothesize that semi-closed basins, even if distant and not necessarily synchronous from each other, may have developed in a comparable way due to similar local climate condition, geochemistry of the body water and river run-off. The geochemical and mineralogical proxies from the mudrocks of the Calcare di Base Formation may open a new scenario for the understanding of paleoclimatic condition during the Messinian Salinity Crisis. • Climatic and environmental changes of the Mediterranean basin in response to the Messinian Salinity Crisis (MSC). • Geochemistry and mineralogy of Calcare di Base (CdB) mudrocks as paleoenvironmental proxies of the first stage of the MSC. • Persistent dry conditions alternating with relatively wet phases during the deposition of the CdB. [ABSTRACT FROM AUTHOR]

Before the Messinian Salinity Crisis (MSC) left its imprint on the sediment record of the Mediterranean Sea in the form of evaporites, the basin had already undergone significant changes. At 7.17 Ma, a drop in δ13C values, as well as a basin-wide shift in the abundance of benthic foraminifers, already attest to a sudden change in the Mediterranean conditions. This event coincides with an increase in the amplitude of the insolation curve. It thus stands to question whether a change in the freshwater budget or a change in the connection between the Mediterranean Sea and the Atlantic was the driver for this event. Answering this question would not only help to understand the event itself, but might also help to decipher the early dynamics of the MSC. With a computational box model, we investigate the response of the Mediterranean Sea to a varying freshwater budget for a wide range of restriction. The results then let us define scenarios in which we analyse how a gradually changing restriction would express itself in the basin dynamics. We find that the change in the freshwater budget alone cannot explain the changes that are attributed with the 7.2 event, but coupled with an increase in restriction most differences can be accounted for. Our results also show that a gradual change in restriction can provoke a non-linear response in the behaviour of the basin, which can appear abrupt when happening on a short enough timescale. Such a change would also enhance the influence of said changes in the freshwater budget. This tells us that the processes that most likely triggered the Messinian Salinity Crisis started much earlier and incrementally increased the restriction of the Mediterranean Sea. • Model allows analysis of response to changes in restriction and freshwater budget. • increased variation expressed by proxies can be caused by an increased restriction. • Analysis links 7.2 Ma event to restriction and as such to MSC. [ABSTRACT FROM AUTHOR]

We carried out an integrated stratigraphic study aiming the identification of the onset of the Messinian salinity crisis (MSC) in the Peri-Adriatic Depression, a region forming the northern front of the Albanides fold-and-thrust belt. The onset was identified in the Kavaje area within an interval of euxinic shales representing a time-equivalent deposit of the Primary Lower Gypsum of stage 1 of the MSC, recognized elsewhere in the Mediterranean realm (CIESM, 2008). This interpretation is supported by the integration of magneto-stratigraphic (base of Gilbert chron) and bio-stratigraphic data (last influx of T. multiloba; last occurrence of foraminifera) calibrated with precessional cyclicity recognized in the pre-evaporitic sediments by rock magnetic analyses. Detailed sedimentological observations revealed that the overlying Gips-Sharrë Gypsum unit consists of chaotic and clastic evaporites floored by a regional-scale erosional surface. Their 87Sr/86Sr values (0.708937 to 0.708967) are typical of stages 1–2 of the MSC. Consequently, their assignment to the Resedimented Lower Gypsum unit is here proposed. The halite deposits, that have been intercepted by boreholes above the Gips-Sharrë Gypsum unit, have been also framed in the RLG. Our results confirm the applicability of the MSC 3-stage model (CIESM, 2008) also in the eastern side of the Adriatic Sea. • Identified the onset of the Messinian Salinity Crisis in the Peri-Adriatic Depression. • New chronostratigraphic framework for the MSC sedimentary record of Albania. • Rock-magnetic analysis as a powerful tool to reconstruct a climatic cyclicity. [ABSTRACT FROM AUTHOR]

Abstract The evaporites of the Realmonte salt mine (Sicily, Italy) are important archives recording the most extreme conditions of the Messinian Salinity Crisis (MSC). However, geochemical approach on these evaporitic sequences is scarce and little is known on the response of the biological community to drastically elevating salinity. In the present work, we investigated the depositional environments and the biological community of the shale–anhydrite–halite triplets and the K–Mg salt layer deposited during the peak of the MSC. Both hopanes and steranes are detected in the shale–anhydrite–halite triplets, suggesting the presence of eukaryotes and bacteria throughout their deposition. The K–Mg salt layer is composed of primary halites, diagenetic leonite, and primary and/or secondary kainite, which are interpreted to have precipitated from density-stratified water column with the halite-precipitating brine at the surface and the brine-precipitating K–Mg salts at the bottom. The presence of hopanes and a trace amount of steranes implicates that eukaryotes and bacteria were able to survive in the surface halite-precipitating brine even during the most extreme condition of the MSC. Graphical abstract

The Messinian Salinity Crisis (MSC) created a substantial negative impact on the Mediterranean marine fauna and flora because hypersaline conditions were established in many marginal basins. At the northern margin of the Tertiary Piedmont Basin, Northern Italy, the Messinian successions exposed at Banengo and Moncalvo recorded the transition into and through the MSC with well-preserved nanno-, micro- and macrofossils. Siliceous sponge spicules present in these pre-MSC sediments reflect the sponge population in this area with twenty different morphological types of spicule identified, twelve of them attributable to sponge genera of the modern Mediterranean sponge fauna. The most common genera were Stelletta, Geodia and Dercitus, with fewer records of Tethya, Chondrilla, Alectona and Samus. Only two types of spicule belong to species not currently present in the Mediterranean sponge fauna and are putatively attributed to the tropical species Stelletta purpurea and Diplastrella megastellata. The sponge assemblage thriving before the onset of the MSC provided new information on the sponge diversity and palaeoenvironment of this northern sector of the palaeo-Mediterranean basin over this crucial period. These data, integrated with information derived from the associated fossils (nannofossils, benthic and planktic foraminifers, and bryozoans), indicate that the considered sites were coastal localities, with sediments rich in an organic matter likely derived from phytoplankton blooms. At both sites, the presence of patchy rocky substrates on the seafloor is testified by a diversified group of sponges typical of hard substrata. These rocks were, very likely, carbonate as suggested by the presence of spicules of typical boring sponges. In the studied sites, common epiphytic foraminifers and bryozoan segments indicate the presence of Posidonia seagrass or algal meadows. [ABSTRACT FROM AUTHOR]

The Calcare di Base is a carbonate/evaporitic unit formed before the deposition of the massive halite bodies in the main depozones of the Mediterranean region, and has been at the centre of a fiery debate between different 'schools of thought' about its genesis, environmental conditions and timing of deposition. It crops out extensively in Southern Italy, across Neogene basins located on the Tyrrhenian and Ionian margins of northern Calabria, where it was studied through a detailed facies and stratigraphic analysis that revealed the presence of a continental/shallow‐marine to slope system, active during the onset of the Messinian Salinity Crisis. During the Calcare di Base deposition, an arid climate regime with intermittent humid phases occurred with a precession forced cyclicity, facing an overall generally cold climatic period. During arid periods, stressed environmental conditions permitted the establishment of widespread microbial‐mediated carbonate production, frequently associated with evaporitic (gypsum‐dominated) deposition, along a sabkha‐type environment with extensive salinas and hypersaline coastal lagoons grading seaward into a carbonate/evaporite shallow‐marine shelf. This is testified to by microbial boundstones and dissolution breccias commonly associated with nodular gypsum and arenites affected by desiccation, typically forming metre‐scale shallowing‐upward peritidal cycles. Seaward, gravity flow deposits are interpreted as re‐sedimented shelf limestones, defining a slope apron. Mixed carbonate/evaporite megabreccias and disorganized floatbreccias were deposited along the upper slope, whereas soft deformation with slides, slumps and channelized turbidites further developed along the lower slope to basin. During the humid climate episodes, that affected the platform system with a precession‐forced cyclicity, the carbonate production was reduced together with the evaporitic deposition, leading to the widespread deposition of marlstones and fine to coarse siliciclastic deposits. [ABSTRACT FROM AUTHOR]

The Mediterranean realm, comprising the Mediterranean and Macaronesian regions, has long been recognized as one of the world’s biodiversity hotspots, owing to its remarkable species richness and endemism. Several hypotheses on biotic and abiotic drivers of species diversification in the region have been often proposed but rarely tested in an explicit phylogenetic framework. Here, we investigate the impact of both species-intrinsic and -extrinsic factors on diversification in the species-rich, cosmopolitan Limonium, an angiosperm genus with center of diversity in the Mediterranean. First, we infer and time-calibrate the largest Limonium phylogeny to date. We then estimate ancestral ranges and diversification dynamics at both global and regional scales. At the global scale, we test whether the identified shifts in diversification rates are linked to specific geological and/or climatic events in the Mediterranean area and/or asexual reproduction (apomixis). Our results support a late Paleogene origin in the proto-Mediterranean area for Limonium, followed by extensive in situ diversification in the Mediterranean region during the late Miocene, Pliocene, and Pleistocene. We found significant increases of diversification rates in the “Mediterranean lineage” associated with the Messinian Salinity Crisis, onset of Mediterranean climate, Plio-Pleistocene sea-level fluctuations, and apomixis. Additionally, the Euro-Mediterranean area acted as the major source of species dispersals to the surrounding areas. At the regional scale, we infer the biogeographic origins of insular endemics in the oceanic archipelagos of Macaronesia, and test whether woodiness in the Canarian Nobiles clade is a derived trait linked to insular life and a biotic driver of diversification. We find that Limonium species diversity on the Canary Islands and Cape Verde archipelagos is the product of multiple colonization events followed by in situ diversification, and that woodiness of the Canarian endemics is indeed a derived trait but is not associated with a significant shift to higher diversification rates. Our study expands knowledge on how the interaction between abiotic and biotic drivers shape the uneven distribution of species diversity across taxonomic and geographical scales.

The southern part of the French Alps is studied for years in mapping and understanding of large-scale gravitational deformations. The identification and the knowledge of large-scale slope deformation (deep-seated gravitational slope deformation or DSGSD and deep-seated landslide or DSL) in the previous work of Jomard (2006) and Zerathe (2013) open a new vision of landslide processes with the reinterpretation of their dynamics and the characterization of the time scales involved. We identify DSGSD (10.109 m3) in the Var Valley associated with geological and geomorphological anomalies linked to the Messinian Salinity Crisis (MSC) and the alpine orogenesis. We use field observations, geological information (geological map, boreholes), and topographic analysis performed in a GIS environment in order to describe these anomalies. This old and partly eroded slide mass is associated with three typical DSGSD features: (1) a double-crested ridge, the Sinne Valley, (2) a large formation (2.7 × 108 m3) of slope deposit dated from the Messinian (Carros breccia), and (3) the kilometric deviation (1 to 2 km) of the Var River. We relate all these anomalies to the MSC (5.97 to 5.46 Ma) and the incision of deep canyons during this period related to this major eustatic variation (≈ 1300 m). The incision of the canyon triggered the collapse process of the DSGSD of the Sinne Valley and so destabilized the entire massif. At present, three DSLs resulting from the DSGSD deformation are still present in the area in a dormant state. Indeed, since the infilling of the Var Canyon during the Pliocene, the activity of the DSGSD has stopped. [ABSTRACT FROM AUTHOR]