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We calculate the hydrodynamic force on a small spherical, unsteady squirmer moving with a time-dependent velocity in a fluid at rest, taking into account convective and unsteady fluid inertia effects in perturbation theory. Our results generalise those of Lovalenti and Brady (1993) from passive to active spherical particles. We find that convective inertia changes the history-contribution to the hydrodynamic force, as it does for passive particles. We determine how the hydrodynamic force depends on the swimming gait of the unsteady squirmer. Since swimming breaks the spherical symmetry of the problem, the force is not completely determined by the outer solution of the asymptotic matching problem, as it is for passive spheres. There are additional contributions due to the inhomogeneous solution of the inner problem. We also compute the disturbance flow, illustrating convective and unsteady effects when the particle experiences a sudden start followed by a sudden stop., Comment: Revised version, 16 pages, 2 figures

It is well known that the reversibility of Stokes flow makes it difficult for small microorganisms to swim. Inertial effects break this reversibility, allowing new mechanisms of propulsion and feeding. Therefore it is important to understand the effects of unsteady and fluid inertia on the dynamics of microorganisms in flow. In this work, we show how to translate known inertial effects for non-motile organisms to motile ones, from passive to active particles. The method relies on a principle used earlier by Legendre and Magnaudet (1997) to deduce inertial corrections to the lift force on a bubble from the inertial drag on a solid sphere, using the fact that small inertial effects are determined by the far field of the disturbance flow. The method allows for example to compute the inertial effect of unsteady fluid accelerations on motile organisms, and the inertial forces such organisms experience in steady shear flow. We explain why the method fails to describe the effect of convective fluid inertia., Comment: 10 pages, revised version

We compute the drag force on a sphere settling slowly in a quiescent, linearly stratified fluid. Stratification can significantly enhance the drag experienced by the settling particle. The magnitude of this effect depends on whether fluid-density transport around the settling particle is due to diffusion, to advection by the disturbance flow caused by the particle, or due to both. It therefore matters how efficiently the fluid disturbance is convected away from the particle by fluid-inertial terms. When these terms dominate, the Oseen drag force must be recovered. We compute by perturbation theory how the Oseen drag is modified by diffusion and stratification. Our results are in good agreement with recent direct-numerical simulation studies of the problem at small Reynolds numbers and large (but not too large) Froude numbers., Comment: 10 pages, 1 figure