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Maculauger sudchinensis Malcolm, Terryn & Fedosov newspecies (Fig. 13D, E) Type material: Holotype: MNHN IM-2013-61887, 1 lv, 19.9 mm; N Macclesfield Bank, S China Sea, 16 ° 07 'N, 114 ° 23 'E, 161 m (ZhongSha 2015 Stn DW4144). Paratypes: N Macclesfield Bank, S China Sea, 16 ° 08 'N, 114 ° 19 'E, 180–226 m (ZhongSha 2015 Stn CP4145). Paratype 1: MNHN IM-2013-61902, 1 lv, 20.2 mm. Paratype 2: MNHN IM-2013-61895, 1 lv, 20.7 mm. Other examined material: N Macclesfield Bank, S China Sea, 16 ° 07 'N, 114 ° 19 'E, 218–281 m (ZhongSha 2015 Stn CP4148), 2 dd. Zoobank registration: urn:lsid:zoobank.org:act: BB5F5C93-C9FE- 44F0-B1EC-C00AA31EFF23 Diagnostic nucleotide positions: Table 11. Description: Shell to 21 mm; narrow apical angle. Protoconch paucispiral, 1.5 translucent whorls. Teleoconch of holotype with 14 whorls; outline of whorls slightly convex. Subsutural band formed of deep pits. Axial ribs narrow with wide interspaces; ribs c. 18–20 on penultimate whorl, straight on subsutural band, becoming oblique at subsutural groove, then curved on abapical part of whorls. Axial ribs intersected by 2–3 evenly spaced spiral grooves. Aperture long, with straight columella. Ground colour chalkywhite, sometimes with vague brown marks on subsutural band. Distribution: Only known from N Macclesfield Bank. Habitat: Depths of 160– 220 m. Etymology: Named after the South China Sea. Remarks: Maculauger campbelli has a similar protoconch and slender overall shape with slightly convex whorls, but in comparison with M. sudchinensis n. sp. has a more continuous subsutural groove, less pronounced sculpture, with 5–7 spiral grooves and a distinctive pattern of straw colour with random square spots. Maculauger sudchinensis n. sp. is similar to Terebra helichrysum (Fig. 13F), which however has a much shorter curved columella, a multispiral protoconch of 3.5 whorls, shorter whorls and a mottled orange colour pattern. Terebra levantina (Fig. 13G) has similar sculpture, but its whorls are much shorter, with 4–6 spiral grooves and it has a mottled colour pattern.

Genus Maculauger new genus (Fig. 7E, F) Type species: Terebra pseudopertusa Bratcher & Cernohorsky, 1985. Zoobank registration: urn:lsid:zoobank.org:act: 3565DB17-E4EF- 478B-8757-3A682BB8F6CF Definition: Includes all species included in clade E5A of Modica et al. (2019) and those that show a combination of conchological, anatomical and distribution characteristics closely comparable with Maculauger pseudopertusus or any genetically proven member of the clade. Diagnosis: Diagnostic nucleotide combinations in Table 10. Shell: Small to medium-sized (< 60 mm); typically, orange-brown with irregular maculations on subsutural band. Whorl outline flattened or gently convex. Subsutural band lightly ribbed or flattened, separated from lower portion of whorl by narrow groove or row of punctations. Axial and spiral sculpture of about equal strength, producing cancellate sculpture below subsutural band. Anatomy: Proboscis, radula, venom gland and accessory proboscis structure absent; salivary glands present or absent. Distribution: Indo-Pacific; subtidal and bathyal depths to about 600 m. Remarks: The new genus Maculauger comprises several species, formerly placed in Hastulopsis, that can be differentiated from similar species in Myurella by the equal prominence of spiral and axial spiral elements, producing a fine cancellate sculpture. Hastulopsis species have very weak sculpture and are found in the Indian Ocean or Japan. Some species of Maculauger, such as M. campbelli, are conchologically very close to Profunditerebra orientalis —a case where only anatomical and molecular characters can ensure correct identification (see Remarks on Profunditerebra). Etymology: The name combines ‘macula’ (Latin: spot, blotch), referring to the commonly maculated subsutural band in the species of the genus, with ‘auger’, the English epithet for Terebridae (‘auger shells’). Gender masculine. Included species: Maculauger alveolatus (Hinds, 1844) 1 n. comb.; M. campbelli (R. D. Burch, 1965) 1 n. comb.; M. castigatus (A. H. Cooke, 1885) 1 n. comb.; M. cinctella (Deshayes, 1859) 3 n. comb.; M. minipulcher (Bozzetti, 2008) 1 n. comb.; M. pseudopertusus (Bratcher & Cernohorsky, 1985) 1 n. comb.; M. sudchiniensis n. sp. Malcolm, Terryn & Fedosov 1 n. sp.

Genus Maculauger new genus (Fig. 7E, F) Type species: Terebra pseudopertusa Bratcher & Cernohorsky, 1985. Zoobank registration: urn:lsid:zoobank.org:act: 3565DB17-E4EF- 478B-8757-3A682BB8F6CF Definition: Includes all species included in clade E5A of Modica et al. (2019) and those that show a combination of conchological, anatomical and distribution characteristics closely comparable with Maculauger pseudopertusus or any genetically proven member of the clade. Diagnosis: Diagnostic nucleotide combinations in Table 10. Shell: Small to medium-sized (< 60 mm); typically, orange-brown with irregular maculations on subsutural band. Whorl outline flattened or gently convex. Subsutural band lightly ribbed or flattened, separated from lower portion of whorl by narrow groove or row of punctations. Axial and spiral sculpture of about equal strength, producing cancellate sculpture below subsutural band. Anatomy: Proboscis, radula, venom gland and accessory proboscis structure absent; salivary glands present or absent. Distribution: Indo-Pacific; subtidal and bathyal depths to about 600 m. Remarks: The new genus Maculauger comprises several species, formerly placed in Hastulopsis, that can be differentiated from similar species in Myurella by the equal prominence of spiral and axial spiral elements, producing a fine cancellate sculpture. Hastulopsis species have very weak sculpture and are found in the Indian Ocean or Japan. Some species of Maculauger, such as M. campbelli, are conchologically very close to Profunditerebra orientalis —a case where only anatomical and molecular characters can ensure correct identification (see Remarks on Profunditerebra). Etymology: The name combines ‘macula’ (Latin: spot, blotch), referring to the commonly maculated subsutural band in the species of the genus, with ‘auger’, the English epithet for Terebridae (‘auger shells’). Gender masculine. Included species: Maculauger alveolatus (Hinds, 1844) 1 n. comb.; M. campbelli (R. D. Burch, 1965) 1 n. comb.; M. castigatus (A. H. Cooke, 1885) 1 n. comb.; M. cinctella (Deshayes, 1859) 3 n. comb.; M. minipulcher (Bozzetti, 2008) 1 n. comb.; M. pseudopertusus (Bratcher & Cernohorsky, 1985) 1 n. comb.; M. sudchiniensis n. sp. Malcolm, Terryn & Fedosov 1 n. sp., Published as part of Fedosov, Alexander E, Malcolm, Gavin, Terryn, Yves, Gorson, Juliette, Modica, Maria Vittoria, Holford, Mandë & Puillandre, Nicolas, 2019, Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea), pp. 359-388 in Journal of Molluscan Studies The Malacological Society of London 85 (4) on page 20, DOI: 10.1093/mollus/eyz004, http://zenodo.org/record/4469844, {"references":["MODICA, M. V., GORSON, J., FEDOSOV, A. E., MALCOLM, G., TERRYN, Y., PUILLANDRE, N. & HOLFORD, M. 2019. Macroevolutionary analyses suggest environmental factors, not venom apparatus, play key role in Terebridae marine snail diversification. Systematic Biology. DOI: https: // doi. org / 10.1093 / sysbio / syz 059."]}