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Following results of recent studies, Huperzia myriophyllifolia (Huperziaceae), Lycopodium subinundatum (Lycopodiaceae), and Polystichum pseudodeltodon, P. pseudostenophyllum, P. subapiciflorum, and P. subobliquum (Dryopteridaceae) are recognized/reinstated. A new combination, Pseudolycopodiella subinundata, is made based on L. subinundatum. Spinulum lioui (Lycopodiaceae) is reported as a new record for most part of China and other Asian countries.

4.10. Phlegmariurus recurvifolius (Rolleri) B.Øllg. Collections studied from the Guianas: 2 (GU: 2) Guyana: Samwarakna-tipu (Holi-tipu), Kamarang R., Wenamu trail, 1100 m, 10 Nov 1951, Maguire & Fanshawe 32519 (NY); Pakaraima Mts., Mount Aymatoi (sandstone), 1150 m, 15 Oct 1981, Maas et al. 5683 (AAU, NY, U)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 119, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.1. Phlegmariurus acerosus (Sw.) B.Øllg. Collections studied from the Guianas: 1 (FG: 1) French Guiana: Mont Itoupé - Sommet Tabulaire - 15, versant ouest, layon D, 780 m, 1 Apr 2010, Tostain et al. 5973 (AAU, CAY, P, US). Note: The small population recently found (2010) in Central French Guiana, at 780 m elevation, appears very isolated, the closest populations of the species being in Western Venezuela and in the Lesser Antilles. It could be expected in high mountains of Guyana and Suriname., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 111-112, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.8b. Phlegmariurus linifolius (L.) B.Øllg. var. jenmanii (Underw. & F.E.Lloyd) B.Øllg. Collections studied from the Guianas: 36 (GU: 11; SU: 14; FG: 11) Guyana: R. Padauiri, s.d., R. H. Schomburgk I 1017 (K, P, US); Kaieteur Savanna, 1897, Jenman s.n. (NY); Moruca R., Demerara, s.d., Jenman s.n. (BONN-Nessel 388, E, NY); Kaieteur Savanna, Potaro R., s.d., Jenman 1475 (K); Berbice-Rupununi, Cattle trail, Demerara, Sep 1919, Abraham 237 (BONN, BRG not found, K, NY); s.loc., s.d. (prob. 1937), Sandwith 1193 p.p. (K); Soesdyke, Jan 1974, Cooper 60 (NY, U); Mr. A. Thompson’s farm, S of Timehri, 13 Oct 1979, Maas & Westra 3505 (U); Cuyuni-Mazaruni, Pakaraima Mts, Kurupung Landing, 0.5-1 km SW of Kurupung R., 75 m, 15 Jun 1992, Hoffman et al. 2028 (US); Cuyuni-Mazaruni Region, Pakaraima Mts, Kurupung R., Imatta Creek, near Kurupung Landing, 80 m, 12 Aug 1992, Hoffman et al. 2399 (CAY, NY, U, US); Potaro-Siparuni Region, Upper Potaro R., 0–10 km S of camp, NE of toe slope of Mt. Wokomung, 620 m, 20 Jul 2003, Clarke et al. 11017 (US). Suriname: Distr. Para, May 1844, Hostmann 1199 A (B, P, S); s.loc., s.d. (prob. ca. 1850), Kegel 4816 (P); s.loc., s.d. (prob. 1849-1855), Wullschlägel 489 (P), 1212 (P); s.loc., s.d., Miquel s.n. (BR); Upper Suriname R., 1 Oct 1908, Tresling 398 (U); Boven-Nickerie, 24 Feb 1915, Gonggrijp 1062 (U); Kaboeri, Corantijn R., 27 Jun 1916, Gonggrijp 2110 (U); Piai Kreek, 13 Oct 1920, Pulle 577 (U); Brownsberg, 9 Oct 1923, Zaandam 6200 (U, UC, US); Brownsberg, 21 Apr 1925, Zaandam 6862 (U); Para –rivier, 2–4 km S of Hout-water, 28 Jan 1962, Kramer & Hekking 2749 (U); Para, Powaka, 13 Jan 1986, Elias 16903 (BBS); Mts. Bakhuis, concession BMS: zone 2, Nord, 330 m, 12 Dec 2006, Bordenave et al. 8623 (BBS, CAY). French Guiana: Oyapock, 1834, Leprieur s.n. (P); s.loc., 1836, Leprieur s.n. (P); s.loc., 1838, Leprieur s.n. (P); riv. Mana, crique Tamanoir, 20 Aug 1962, Hallé 614 (P, U, US); saut Dame Jeanne, bassin de la Mana, 29 Aug 1962, Hallé 724 (CAY, P, U, US, VEN); sauts Koueki, bassin de l’Yaloupi, 28 Apr 1970, Oldeman-B 3099 (CAY, P); Ilet Matain - saut Grand Ako, bassin de l’Oyapock, 19 Jul 1980, Prévost & Grenand 866 (CAY, MPU); montagne des Nouragues, bassin de l’Approuague, 1 Sep 1987, Feuillet & Riéra 4475 (CAY); montagne des Nouragues, bassin de l’Approuague, 1 Jun 1989, Sarthou 541 (CAY); station des Nouragues, bassin de l’Approuague, 380 m, 16 Nov 1989, Cremers 10797 (CAY); crique Grande Waki nord, bassin de la Waki, 140 m, 18 Aug 2012, Girault & Pelletier 817 (CAY); début de la piste de la montagne Plomb, Kourou (H. Galliffet, 28 Oct 2019, website photos!)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 118, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

5.4a. Pseudolycopodiella meridionalis (Underw. & F.E.Lloyd) Holub var. meridionalis Collections studied from the Guianas: 106 (GU: 45; SU: 32; FG: 29) Guyana: s.loc., s.d., M. R. Schomburgk 1159 (P); Rivulets between Suddie and Huribisei, 1897, Jenman s.n. (NY); between Suddie and Huribisei, Essequibo, s.d., Jenman s.n. (NY); Corentyne R., s.d., Jenman 374 (K); Kaieteur Savanna, s.d., Jenman 1474 (K); s.loc., s.d., Jenman 2230 (US); Kako Creek, s.d. (prob. 1864–1872), Appun 1027 (K); Potaro Road, s.d. (prob. 1909–1912), Stockdale 8858 (K); Penal Settlement, 3–9 Dec 1919, Hitchcock 17153 1/2 (NY, S, US); Mt. Roraima, on rocks at side of the Kukenam R., Paulo, 1220 m, 4 Nov 1927, Tate 185 (NY); Mt. Roraima, Philipp swamp, 1570 m, 11 Nov 1927, Tate 351 (NY); Mazaruni Station, 14 Aug 1937, Sandwith 1081 (K, U); Parabaru Savanna, s.d. (prob. 1938), Smith A. C. 3012 (K); between Rupununi and Kuyuwini R., Parabaru Savanna, 16 Feb 1938, Smith A. C. 3052 (F, K, MO, NY, P, S, U, US); Kaieteur Plateau, 30 Apr 1944, Maguire & Fanshawe 23111 (BR, F, K, NY, P, S, U, UC, US, VEN); North Kopinang Savanna, on Waipa trail, 850 m, 4 Sep 1961, Maguire 46090 (K, NY, U); Ité Swamp, Rupununi Northern Savanna, 12 mi N of Manari Creek, 150 m, 26 Sep 1963, Goodland 882 (NY, P, US); Imbaimadai, upper Mazaruni R., Aug 1967, Davis 321 (BRG not found, K); Soesdyke, Mar 1974, Cooper 55 (U); vicinity of the village of St-Cuthbert’s, ca. 50 km SSE of Georgetown, 8 Aug 1976, Mori et al. 8047 (K, NY); Pakaraima Mts., Mount Membaru, 400 m, 12 Nov 1979, Maas 4339 (K, U); Pakaraima Mts., Kako Amerindian village, Kako R., 13 Nov 1979, Maas 4401 (K, U); Pakaraima Mts., Mt. Aymatoi, 1150 m, 15 Oct 1981, Maas 5673 (K, U); Upper Mazaruni Distr., N slope of Mount Roraima, 2300 m, 16 Feb 1985, Renz et al. 14265a (U); Potaro-Siparuni, Kaieteur National Park 2, near airstrip, 395 m, 24 Jan 1987, Pipoly & Gharbarran 9816 (AAU, CAY, FDG not found, NY, US); Cuyuni-Mazaruni, Chinoweing Village, Ayanganna Plateau, 660 m, 20 Feb 1987, Pipoly et al. 10372 (AAU, NY); Upper Mazaruni R. Region, Karowtipu Mt., Eastern end of mountain, 1000 m, 21 Apr 1987, Boom & Gopaul 7591 (NY); Potaro-Siparuni, Kaieteur Falls National Park, along rim downs, 500 m, 13 Apr 1988, Hahn et al. 4530 (US); U. Takutu-U. Essequibo, Gunn’s Village, Essequibo R., 250 m, 2 Sep 1989, Jansen-Jacobs 1386 (CAY, NY, U); Cuyuni-Mazaruni, Savanna between Partang and Karowrieng rivers, 580 m, 20 Dec 1989, Gillespie & Smart 2796 (NY, U, US); Cuyuni-Mazaruni, Mt. Holitipu, 1240 m, 1 Jun 1990, McDowell & Gopaul 3043 (CAY, NY, U, US); Cuyuni-Mazaruni Region, Pakaraima Mts., 2–5 km N of Imbaimadai settlement, 500 m, 22 Oct 1992, Henkel 34 (U); Potaro-Siparuni, Pakaraima Mts., Upper Ireng R., 750 m, 3 Feb 1993, Henkel et al. 1158 B (NY); U. Takutu-U. Essequibo, Gunn’s Village, Essequibo R., 250 m, 18 Feb 1994, Henkel et al. 4563 (CAY, NY, U, US); Kaieteur National Park, between guest house and Manzies Landing, 300 m, 12 Jun 1994, Kelloff 1061 (US); Potaro-Siparuni Region, Kurupukari-Annai road, mi 28, Iwokrama National Rainforest Reserve, 60 m, 30 Sep 1996, Clarke 2660 (NY, U); Potaro-Siparuni, Mt. Wokomung, Little Ayanganna, upper slopes of highest point of Mt. Wokomung massif, 1525 m, 5 Jul 2003, Clarke 10568 (NY, US); Boy Scout Lookout and Johnson Lookout, 465 m, 30 Aug 2006, Wurdack et al. 4199 (NY); Cuyuni-Mazaruni, Imbaimadai, across from airstrip in savanna and along creekside, 515 m, 3 May 2009, Redden et al. 6406 (BRG not found, NY, US); Cuyuni-Mazaruni, Kato R. On savanna near old farm, 485 m, 6 May 2009, Wurdack al. 4854 (BRG not found, NY, US); Cuyuni-Mazaruni, Kato R., 490 m, 19 May 2009, Redden et al. 6679 (BRG not found, US); Cuyuni-Mazaruni, Kako R. vicinity, above Kako Village, near Camp 3, ca. 22 air km east of Tulameng summit, 495 m, 1 Jul 2010, Wurdack et al. 5205 (BRG not found, K, US); Cuyuni-Mazaruni, Imbaimadai – Mazaruni R., 515 m, 29 Jul 2010, Redden et al. 7113 B (US); Cuyuni-Mazaruni, Imbaimadai airstrip vicinity, west of airstrip, 505 m, 30 Jul 2010, Redden et al. 7154 (BRG not found, US); Cuyuni-Mazaruni, below lst escarpment (of four) of Kamakusa Mt., 735 m, 19 May 2012, Redden et al. 7246 (BRG not found, CAY, US). Suriname: In virgultis aridis, 1842, Hostmann & Kappler 1155–(4761) (P ); s.loc., 1843, Hostmann 1155 [‘ M. Hortmann’] (P); s.loc., s.d., Hostmann 1155 (P, U); s.loc., 1845, Hooker s.n. (P); s.loc., s.d. (prob. 1849–1855), Wullschägel 1701 (P); Zanderij I, Savanne, Oct 1909, Boldingh 3001 (U); Zanderij I, Sep 1914, Kuijper s.n. (U); Zanderij I, 22 Jul 1933, Lanjouw 148 (U); Zanderij I, 29 Jul 1933, Lanjouw 289 (BR, K, U); Zanderij II, 3 Jun 1944, Maguire & Stahel 23690 (NY); vicinity of Arawak village of Mata, Zanderij I, 18 Oct 1944, Maguire & Stahel 24981 p.p. (A, U); Zanderij I, 4 Sep 1948, Lanjouw & Lindeman 117 (U); road Moengo tapoe – Cr. Zwibelzwamp, 7 Oct 1948, Lanjouw & Lindeman 701 (NY, U); road Moengo tapoe – Cr. Zwibelzwamp, 8 Oct 1948, Lanjouw & Lindeman 748 (U); road Moengo tapoe – Cr. Zwibelzwamp, 21 Oct 1948, Lanjouw & Lindeman 953 (K, U, US); along rail-road, near km 98, 20 Apr 1949, Lanjouw & Lindeman 3400 (U, US); Zanderij, 22 Dec 1950, Florschütz & Florzchütz 800 (U); Zanderij, Pontjikreek, new road to Matta, 9 Dec 1955, Jonker-Verhoef & Jonker 148 (U, US); Zanderij: Troelie, Zanderij-Mata, 27 Nov 1960, Kramer & Hekking 2209 (BBS, NO, U); Zanderij, Colakreek, 27 Jan 1961, Kramer & Hekking 2779 (U, VEN); just E of Zanderij, 12 Feb 1961, Tryon & Kramer 5616 (COL, P, S, US); Distr. Saramacca, Kappel Savanna at S foot of Tafelberg, 300 m, 25 Feb 1961, Kramer & Hekking 3039 (U); Zanderij airstrip, Feb 1963, Schultz 9608 (BBS, U); Kappel-Savanna near the S foot of the Tafelberg, 8 Jun 1963, Wessels Boer 1520 (P, U); savanne between Zanderij and Troelie, 2 Jan 1964, Schultz 9966 (BBS, U); Distr. Brokopondo, 4 km E of village of Brownsweg, 22 Dec 1965, Donselaar 2836 (U); savanne W of Brinckheufel, between Amoedoejoekreek and Spikrihoedoekreek, 26 Aug 1967, Teunissen & Wildschut 14487 (U); wet savanna S of Bigi Poika, 16 Sep 1970, Gier 72 (U); Zanderij savanne near Troeli, Mar 1976, Jansma 15682 (BBS); Zanderij near Matta, 19 Sep 1976, Mori et al. 8324 (BBS, NY, U, US); intersection Zanderij/Avenavero hwy, and road to Matta, 9 km W of Zanderij, 25 m, 11 Jul 1979, Webster 24049 (BBS); Sipaliwini, around Rudi Kappel savanna, near airstrip, 325 m, 23 Aug 2013, Aguirre-Santoro & Michelangeli 1903 (NY). French Guiana: s.loc., s.d. (prob. 1781–1789), Richard s.n. (P); Montsinéry, Sept 1837, Leprieur 159 (P, RB, US); s.loc., 1838, Leprieur 15 (P); s.loc., 1840, Leprieur s.n. (P); Charvein, 10 Nov 1913, Benoist 190 (P); bourg de Saint-Elie, bassin du Sinnamary, 4 Aug 1974, de Granville-B 5136 (CAY, P); montagne des Chevaux, bassin de la Comté, 23 Jan 1978, Raynal-Roques 19879 (CAY, P, U); montagne des Chevaux, bassin de la Comté, 18 Oct 1981, Billiet & Jadin 1044 (BR, CAY, U); route Nationale R. N. 2, pk 36, bassin de la Comté, 25 Oct 1983, Billiet & Jadin 1859 (BR, CAY); savane Lambert 1, région de Cayenne, 28 Mar 1986, Cremers 9419 (AAU, B, CAY, P, U, Z); route Nationale R. N. 2, pk 36, bassin de la Comté, 100 m, 22 Aug 1986, Sastre & Bell 8022 (CAY, P, US); savane de Corossony, région littorale, 5 m, 27 Dec 1986, Cremers 9557 (AAU, B, CAY, NY, P, U); piste de Saint-Elie, 50 m, 28 Mar 1988, Philippe 214 (CAY); savane Macoua, région littorale, 10 m, 26 May 1989, Hoff & Cremers 5575 (AAU, B, CAY, US); savane Aubanèle, région littorale, 20 m, 9 Jun 1989, Cremers & Hoff 10709 (AAU, CAY, GOET, P, Z); bourg de Saint-Elie, bassin du Sinnamary, 27 Nov 1989, Billiet & Jadin 4644 (BR, CAY); crique Favard, plaine et montagne de Kaw, 7 m, 8 Feb 1996, Jansen-Jacobs 5126 (CAY, U); piste de Rocoucoua - 1, région littorale, 10 m, 25 Jan 1997, Cremers 14574 (CAY, P); secteur de Saint-Elie - 1, environs du village, bassin du Sinnamary, 130 m, 14 Dec 1997, Boudrie 2864 (CAY); piste de Rocoucoua - 1, région littorale, 15 May 2001, Héquet 1266 (CAY); Placer Trésor - 2, plaine et montagne de Kaw, 380 m, 30 Jan 2003, Ek et al. 1447 (CAY, U); Petite Montagne Tortue, bassin de l’Approuague, 50 m, 26 Feb 2003, Christenhusz et al. 2535 (AAU, CAY, TUR); route nationale R. N. 2, entre les pk 87 et 88, à 16,5 km à l’WNW de Régina, 30 m, 14 Mar 2006, Boudrie et al. 4150 (CAY); savanes de Karouabo, région littorale, 10 m, 11 May 2007, de Granville & Tostain 17438 (CAY, US); montagne des Chevaux, carrière SCC, 30 m, 26 Jul 2007, Tostain 732 (CAY, NY, P); bords de la RN 2, vers le PK 86, et à 17 km à l’WNW de Régina (obs. M. Boudrie, 3 Nov 2007!); bord sud-ouest de la RN 2, à 7,8 km au SSE du carrefour du Galion, au niveau de la Montagne des Chevaux, commune de Roura (obs. M. Boudrie, 15 Aug 2008!); bord sud-ouest de la RN 2, à 11,7 km au SSE du carrefour du Galion, à 5 km environ au NW du pont de la Comté, commune de Roura (obs. M. Boudrie, 15 Aug 2008!); montagne des Chevaux, carrière SCC, 30 m, 31 Oct 2008, Tostain et al. 2271 (CAY, P); route nationale R. N. 2, pk 78, commune de Régina, 30 m, 14 Oct 2015, Petitbon 1629 (CAY), 1630 (CAY), 1631 (CAY); savane Matiti, près du lieu-dit Caussade, com. de Kourou (obs. M. Boudrie, 26 Jul 2016!); Anton, vers le PK 173,8 de la RN 1 (nord de la RN 1), à 26, 3 km à l’ouest-nord-ouest d’Iracoubo (obs. M. Boudrie, 25 Feb 2018!); savanes Roches Blanches, vers le PK 170 de la RN 1, à 12,7 km à l’ouest d’Iracoubo (obs. M. Boudrie, 25 Feb 2018!). Note: Like Palhinhaea cernua, populations of this species and variety are concentrated along the coastal zone in French Guiana, and more widespread southwards in Suriname and in Guyana, due to the presence of large savannas and open areas., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 126-128, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

3.2. Palhinhaea cernua (L.) Franco & Vasc. Collections studied from the Guianas: 229 (GU: 70; SU: 50; FG: 109) Guyana: Roraima, 1852-1853, M.R. Schomburgk s.n. (NY); s.loc., 1868, R.H. Schomburgk I 395 (BR, P, US); s.loc, s.d. (prob. 1864-1872), Appun 195 (K); Essequibo, Mazaruni [as ‘Massaruni’], s.d. (prob. 1864-1872), Appun 7094 (P); Orcalla, Corentyne, Sep 1879, im Thurn s.n. (P?); s.loc., s.d. (prob. 1850-1870), Lesquereux s.n. (MU); Macouria Creek, Essequibo R., Demerara, 1897, Jenman s.n. (NY); Essequibo R. beyond Bartica, s.d., Jenman s.n. (K); s.loc., s.d., Jenman 2329 (K), 2330 (K); along the Berbice-Rupununi cattle trail, Berbice or Demerara County, 1919, Abraham 92 (NY); Kartabo, junction of Mazaruni and Cuyuni rivers, 11 Dec 1919, Hitchcock 17227 (NY, US); Potaro Landing, 27 Jun to 9 Jul 1921, Gleason 262 (K, NY, US); Rockstone, 15 Jul to 1 Aug 1921, Gleason 595 (K, NY, US); Upper Rupununi, near Dadanawa, 14 Jun 1922, de la Cruz 1497 (F, MO, NY, UC, US); junction of Mazaruni and Cuyuni rivers, Jun 1922, Graham 119 (NY); between Demerara and Berbice rivers, 15-19 Jul 1922, de la Cruz 1607 (F, MO, NY, US), Upper Mazaruni R., 22 Sep - 6 Oct 1922, de la Cruz 2331 (F, K, MO, NY, UC, US); Upper Mazaruni R., 22 Sep - 6 Oct 1922, de la Cruz 2339 (F, K, MO, NY, UC, US); Bartica, 5 Oct 1922, Lang & A. C. Persaud s.n. (F); along the Upper Mazaruni R., 14 Nov 1922, Leng 116 (NY); Malali, Demerara R., Oct-Nov 1922, de la Cruz 2664 (F, MO, NY, UC, US); Kamakusa, Upper Mazaruni R., 23-29 Nov 1922, de la Cruz 2789 (F, K, MO, NY, UC, US); Pomeroon R., 17-24 Dec 1922, de la Cruz 3107 (F, MO, NY, UC, US); Amakura R, NW District, 23-30 Mar 1923, de la Cruz 3505 (F, MO, NY); Comaka, Demerara R., May 1923, C. A. Persaud 289 (ASU); Mazaruni R. (Settlement), 3 Aug 1933, Mell & Mell 216 (MICH); s.loc., s.d. (prob. 1929), Richards 401 (K); s.loc., s.d. (prob. 1937), Sandwith 1015 (K); Potaro R., Garraway Stream, 25 Apr 1944, Maguire & Fanshawe 22969 (F, K, MO, NY, P, U, VEN); 1/ 2 mile W of Atkinson field, 19 Mar 1955, Irwin 386 (US); s.loc., s.d. (prob. 1957), Cook 228 (K); Berbice R., 12 Apr 1958, Harrison 835 (BRG not found, K); Upper Mazaruni R. Basin, near mouth of Partang R., Imbaimadai Savannas, 14 Jun 1960, Tillett S. S. & Tillett C. L. 43834 (AAU, NY); Port Kaituma, distr. NW, Aug 1967, Davis 250 (BRG not found, K, NY); Pakaraima Mts., Kamarang, 450 m, 15 Nov 1979, Maas & Vestra 4468 (BRG not found, K, MO, P, U); s.loc., 17 Oct 1981, Maas et al. 5821 (U); Tiperu quarry, R. B., Mazaruni R., 1 Apr 1986, A. C. Persaud 266 (F); U. Demerara-Berbice, Mabura Hill area, 6 Jun 1986, Pipoly & Boyan 7595 (BRG not found, CAY, F, NY, P, U, US); Cuyuni-Mazaruni, Pakaraima Mts., near Imbaimadai airstrip, 15 Jun 1986, Pipoly & Alfred 7862 (BRG not found, CAY, NY, U, US); U. Demerara-Berbice, Mabura Hill 2, 16 Sept 1986, Pipoly & Boyan 8532 (BRG not found, CAY, F, NY, US); Potaro-Siparuni, Kaieteur National Park, vicinity of Tukeit, 27 Jan 1987, Pipoly & Gharbarran 10059 (AAU, CAY, FDG not found, NY, P, U, US); Cuyuni-Mazaruni, unnamed peak S of Chinoweing Village, 700 m, 23 Feb 1987, Pipoly et al. 10556 (AAU, NY); East Berbice-Corentyne, Torani Canal, 5-12 km W of Canje R., 12-17 km E of Berbice R., 19 Apr 1987, Pipoly et al. 11691 (CAY, FDG not found, MSC, NLU, NY, U, US, VDB); Potaro-Siparuni, Kaieteur Falls, vicinity of Kaieteur airstrip, 8 Oct 1987, Kvist et al. 81 (AAU, BRG not found, CAY, U, US); Cuyuni-Mazaruni, Kartabo, Willems Timber Concession, 25 Jan 1989, Hahn & Tiwari 5185 (CAY, U, US); Potaro-Siparuni, Kaieteur National Park, 5 May 1989, Gillespie et al. 1241 (NY, US); East Berbice-Corentyne, Southern end of Long Savanna, upper Canje R., 31 Oct 1989, Gillespie et al. 2592 (CAY, NY, U, US); Berbice-Corentyne, from the banks of the Corentyne R. and Potoko R., 24 Apr 1990, McDowell & Gopaul 2464 (CAY, NY, U, US); Berbice-Corentyne, about 0.5 km W of Corentyne R., near Oreal, 26 Apr 1990, McDowell & Gopaul 2535 (NY, US); Potaro-Siparuni, along Potaro-Konawa road, between Golden Star - Mahadia cutoff and creek, 8 Oct 1990, McDowell 3410 (CAY, NY, U, US); Potaro-Siparuni Region, from camp on Elizabeth Creek 1 km along road, 16 Oct 1990, McDowell & Gopaul 3581 (F, US); Demerara-Mahaica, near Timehri airport, 22 Jan 1992, Hoffman & Capellaro 775 (CAY, U, US); Cuyuni-Mazaruni Region, Pakaraima Mts., Kurupung R., Kamarau Falls, 20 Jul 1992, Hoffman 2105 (BBS, U, US); Potaro-Siparuni, Pakaraima Mts, Upper Ireng R., 675 m, 13 Jan 1993, Henkel et al. 731 (CAY, MO, NY, U, US); U. Demerara-Berbice, Berbice savanna near Takama Army Base, 14 Aug 1993, Henkel et al. 2486 (CAY, NY, U, US); Potaro-Siparuni, Kamama village, 10 km North of Orinduik, 10 Mar 1995, Tiwari et al. 1458 (NY); Kanuku Mts, Moco Moco R., 17 Jul 1995, Jansen-Jacobs et al. 4544 (AAU, U); Potaro-Siparuni Region, Iwokrama rain forest reserve, along Kurupukari-Annai road, 75 m, 20 Nov 1995, Clarke 558 (AAU, NY, US); U. Demerara-Berbice, Tropenbos field station, 3 km S of Mabura Hill, 1 Jan 1996, Clarke 739 (NY, U, US); Cuyuni-Mazaruni Region, Holitipu, trail between camp, airstrip and surrounding areas, 1100 m, 7 Feb 1996, Clarke 1063 (US); Potaro-Siparuni Region, Kurupukari-Annai road, mi 28, Iwokrama National Rainforest Reserve, 30 Sep 1996, Clarke 2665 (US); Barima-Waini Region, Waini R., Kwebana, vicinity of abandoned sawmill, 22 Oct 2001, Hollowell & Hinds 601 (NY, US); Potaro-Siparuni Region, new Ayanganna airstrip vicinity, 16 Jun 2003, Clarke 9934 (U, US); Cuyuni-Mazaruni Region, Pakaraima Mts, Mazaruni R., just above ABC Falls, 22 Feb 2004, Redden & Lyle 2955 (MO, NY, US); Potaro-Siparuni Region, Garraway stream, 102 miles on Bartica-Potaro road, 0.25 mile NE of base camp 1, 13 Mar 2004, Redden et al. 2258 (NY, US); Cuyuni-Mazaruni Region, Ataro R., below falls of Wayalayeng village, 20 Jun 2004, Clarke et al. 12249 (NY, US); Forest behind Isseneru village, on trail to Apiqué shop, 27 Sep 2006, Redden et al. 4317 (NY, US); Essequibo Islands W, Demerara, Bonasika Bauxite site 2, forest between Demerara and Essequibo R., 23 Jun 2010, Redden et al. 6797 (NY, US); Potaro-Siparuni, Mt. Ayanganna, East Slope, between new airstrip and banks of Potaro, along trail leading to miners camp, 695 m, 14 Mar 2014, Radosavljevic et al. 133 (U, US-3 sheets). Suriname: Paramaribo, 1837, Splitgerber 200 (U); s.loc., 1843, Hostmann & Kappler 333 (P); in sylvis humidis prope urb. Paramaribo, May 1844, Kappler 333 (B, NO, P, UC); s.loc., s.d. (prob. 1844), Hostmann s.n. (P); s.loc., s.d., Hostmann 333 (MO); s.loc., s.d., Hostmann [‘ Hortman. ’] 333 (P); in virgultis, s.d., Hostmann & Kappler 333 (P); s.loc., s.d., Kappler 333 (RB, 2 sheets); s.loc., s.d. (prob. 1844), Hostmann 372 (U); Boven Commewijne R., s.d. (prob. 1847), Focke 192 (U); s.loc., s.d. (prob. 1847), Focke 372 (U); Paramaribo, s.d. (prob. 1849-1855), Wullschlägel 662 (BR, P), Wullschlägel 663 (BR); Zanderij I, Sep 1914, Kuijper s.n. (U); forest at Zanderij, 31 May 1916, Samuels 240 (F, GH, K, NY, P, U, US-without number); Spoorban Sectie, 3 Aug 1920, Pulle 138 (U); Zanderij I, savanna, 22 Jul 1933, Lanjouw 193 (U); Weyneweg, between Albina and Moengo tapoe, 14 Aug 1933, Lanjouw 432 (U); Zanderij Is, Railway PK 70, Sectie O, 3 Jun 1944, Maguire & Stahel 23627 (BR, F, K, MO, NY, P, S, U, UC, US, VEN, WTU); vicinity Sectie 0, Railway at Km 68, 19 Oct 1944, Maguire & Stahel 25071 (GH, NY, U); Zanderij I, savanna at airstrip, 9 Sep 1948, Lanjouw & Lindeman 251 (U); Km 12 along Molugo road, 25 Sep 1948, Lanjouw & Lindeman 471 (U, US); Zanderijweg, Republiek, May 1950, Geijskes 173 (BBS); Dam, along railway, 9 Nov 1950, Florschütz & Florschütz 159 (U); Jodensavanne-Mapane kreek area (Suriname R.), 20 Dec 1954, Lindeman 6934 (U); Zanderij, Pontjikreek, new road to Matta, 9 Dec 1955, Jonker-Verhoef & Jonker 145 (U); Copieweg, 21 May 1958, Dirven 734 (U); Zanderij: Troelie, between Zanderij and Mata, 27 Nov 1960, Kramer & Hekking 2208 (BBS, U, VEN); Brownsweg, railway at Km 115-116, along trail to Mazaronitop, 1 km from Brownsweg, 6 Jan 1961, Kramer 2581 (U); Kappel Savanna, 10 km S of Tafelberg, 14 Feb 1961, Kramer & Hekking 3244 (U, US); Jodensavane, Suriname R., 19 Dec 1963, Schultz 9938 (BBS); trail Brownsweg-Mazaroni top, 200 m, 11 Feb 1963, Wessels Boer 638 (U); between Coppename R. and Emma Mts., 13 May 1963, Wessels Boer 1367 (U); W bank of Marataka at Saparrakreek, 9 May 1965, Maas & Tawjoeran 3222 (BBS, U); Savanne W. van Brinckheuvel, Nature Reserve Brinckheuvel (Saban Pasi Sav), 30 Aug 1967, Teunissen & Wildschut 11520 (U, WIS); s.loc., s.d. (prob. 1967), Teunissen & Wildschut s.n. (K); Brokopondo road near lake, 16 Aug 1970, de Gier 78 (U), de Gier 79 (U); Brokobaka, Sep 1974, Rudelman 1747 (BBS), Rudelman 1748 (BBS); Lely Mountains, SW plateaus, 550 m, 19 Sep 1975, Lindeman et al. 66 (U); Zanderij, 5 Mar 1976, Jansma 15556 (BBS); Zanderij Savanna, SE of airport, near and around perimeter road, 30 Sep 1979, Thomas 2407 (NY), Thomas 2420 (NY); s.loc., s.d. (prob. 1979), Bierhorst 58 S (VT); Between Berg en Dal and Brownsberg, 10 Jul 1982, Croat 53872 (BBS, MO); Inselberg Talouakem, Massif des Tumuc-Humac, 650 m, 10 Aug 1993, de Granville et al. 12209 (BBS, CAY, US); Botopasi, 10 Apr 1997, van Troon & Derveld 452 (BBS); s.loc., 24 Aug 1999, Klooster et al. 8 (U); Sipaliwini, around Rudi Kappel savanna, near airstrip, 325 m, 25 Aug 2013, Aguirre-Santoro & Michelangeli 1912 (MO, NY, US). French Guiana: Île de Cayenne, 1788, Stoupy s.n. (P-LA); Montsinéri, Oct 1830, Leprieur 173 (P, 2 sheets); s.loc., 1833, Leprieur s.n. (NO); s.loc., 1836, Leprieur s.n. (P); s.loc., 1838, Leprieur s.n. (P, 2 sheets); s.loc., 1840, Leprieur s.n. (P in herb. Richard); s.loc., s.d., Leprieur s.n. (P in herb. Richard); s.loc., s.d., Leprieur s.n. (F); s.loc., s.d., Leprieur s.n. (P in herb. Cosson); s.loc., s.d., Leprieur s.n. (P in herb. Bory de Saint-Vincent); s.loc., 1842, Mélinon 343 (F, P); Karouany, 1855, Sagot 748 (P, 4 sheets); Acarouany, Jun 1855, Sagot s.n. (P); Maroni, 1857, Sagot s.n. (P); Roura, 1858, Sagot 748 (P); Roura, 1858, Sagot s.n. (P); île Portal, Maroni, s.d., Sagot s.n. (P); s.loc. s.d., Sagot s.n. (P, 2 sheets); Cayenne, s.d. (prob. 1866), Jelski s.n. (US); rivière de Kourou, Grand Bois, Mar 1877, Crevaux s.n. (P); Cayenne, 1893, Beaududuit 16 (P), 17 (P), s.n. (P); Saint-Laurent-du-Maroni, près de la pépinière, Sep 1896, Soubirou s.n. (P); Matoury, région de Cayenne, Feb 1901, Lemée s.n. (P); s.loc., 1902, Geay 3284 (P); Saint-Jean, 18 Mar 1914, Benoist 942 (P); Badoeul [= Baduel], vicinity of Cayenne, 18 Jul 1921, Broadway 912 (NY, US); Saint-Laurent-du-Maroni, Oct 1927, Le Moult 3 (U); camp Lorrain, bassin du Maroni, 1 Aug 1948, Service forestier BAFOG s.n. (CAY, P); Maroni distr., near Decanville, camp Lorrain, 5 Aug 1948, Service forestier BAFOG 4016 (U); Saint-Laurent-du-Maroni, camp Lorrain, Aug 1948, Service forestier BAFOG 4015 (P); Saint-Laurent-du-Maroni, camp Lorrain, Aug 1948, Service forestier BAFOG 4082 (P); Saint-Laurent-du-Maroni, crique Balaté, 2 Dec 1948, Service forestier BAFOG 4368 (P); village Central Inipi (Camopi), 23 Feb 1949, Aubert de la Rüe s.n. (P); village Ratier, Bassin R. Sikini, 15 Mar 1949, Aubert de la Rüe s.n. (P); crique Margot, bassin du Maroni, 28 Apr 1950, Service forestier BAFOG 5078 (CAY, P); Suzini, Feb 1951, Service forestier BAFOG 3704 (P); route de Mana, 3 Feb 1955, Service forestier BAFOG 7013 (U); Saint-Laurent-du-Maroni, 21 Aug 1961, Schnell 11330 (P); saut Sabbat, bassin de la Mana, 12 Aug 1962, Hallé 523 (CAY, P, U, US); ancien village de Bélizon, bassin de la Comté, 20 Jul 1965, Oldeman 1440 (AAU, CAY, P); Fourgassié, pk 1, route forestière ́Rougier», dite route de Cheuvreuil, plaine et montagne de Kaw, 13 Jun 1969, de Granville 247 (AAU, CAY, P); route du Tour de l’Île, plantation ONF, Montsinéry-Tonnégrande, 19 Aug 1969, de Granville 266 (CAY, P); ancien aérodrome du Galion, entre la route du Tour de l’Île et le Galion, Montsinéry-Tonnégrande, région de Cayenne, 13 Jan 1974, Descoings & Luu-D-Vinh 20127 (CAY, MPU, P), Descoings & Luu-D-Vinh 20143 (CAY, MPU, P); Port Inini, région de Cayenne, 15 Jan 1974, Descoings & Luu-D-Vinh 20219 (MPU, P); route de Saint-Jean-du-Maroni, Saint-Laurent-du-Maroni, 15 Jan 1974, Descoings & Luu-D-Vinh 20368 (CAY, MPU, P); lac Lallouette, île de Cayenne, 2 Mar 1976, Raynal-Roques & Tirel 18355 (P); piste de Saint-Elie, Sinnamary, 1 Jun 1976, Moretti 480 (CAY, P); rivière de Cayenne, île de Cayenne, 23 Feb 1977, Poncy 23 (P, U); Saül, 28 Jan 1978, Profizi 48 (MPU); savane Boeuf, Saül, 28 Jan 1978, Clavel & Profizi, s.n. (MPU); crique Anguille - 2, Montsinéry-Tonnégrande, région de Cayenne, 3 Nov 1978, Leeuwenberg 11684 (CAY, WAG, Z); piste de Saint-Elie, Sinnamary, 1 Jun 1980, de Foresta 245 (CAY); piste de Saint-Elie, Sinnamary, 21 Oct 1981, Billiet & Jadin 1094 (BR, CAY, U); bourg de Tonnégrande, région de Cayenne, 23 Mar 1982, Grenand 2070 (CAY, P); route de l’Est, R. N. 2, pk 24 environ, 18 Apr 1982, Fournet 164 (CAY); Rorota, bord du sentier, Ile de Cayenne, 12 Mar 1986, Kodjoed 68 (CAY); route du Tour de l’Île, D 5, au PK 30-30,5, région de Cayenne, 3 Apr 1986, Cremers 9448 (AAU, CAY, P); vicinity of the village, Saül, 7 Jun 1986, Mori 18398 (NY); piste Balaté, bassin du Maroni, 19 Nov 1986, Skog & Feuillet 7495 (AAU, CAY, NY, P, U, US); route du Galion, R. N. 2, entre les PK 15 et 16, région de Cayenne, 13 Dec 1986, Cremers 9489 (B, CAY, INPA, MG, MO, MY, NY, P, SJRP, U, US); Saint-Laurent-du-Maroni, bassin du Maroni, 15 Jun 1987, Fleury 290 (CAY, US); mont Grand Matoury - 2, région de Cayenne, 21 Jul 1987, Hahn 3530 (CAY, NY, P, US); bac de Roura, région de Cayenne, 12 Sep 1987, Weitzman & Hahn 267 (NY, US); piste de Saint-Elie, Sinnamary, 15 Sep 1987, Hahn 3788 (CAY, NY, US); bord de la piste de Roura au bac de Stoupan, Roura, plaine et montagne de Kaw, 31 Dec 1987, Cremers et al. 9654 (B, CAY); montagne Cacao, piste Coralie, bassin de la Comté, 14 Feb 1988, Feuillet 4514 (US); Montsinéry, piste de saut Léodate, région de Cayenne, 25 Feb 1988, Feuillet 4585 (US); piste de Saint-Elie, pk 14, parcelle ARBOCEL, Sinnamary, 1 Apr 1988, Larpin 289 (CAY); piste de Saint-Elie, Sinnamary, 6 Apr 1988, Philippe 246 (CAY); Dégrad Saramaka, Kourou, 9 Jun 1989, Cremers & Hoff 10645 (CAY); savane Bordelaise, route du Tour de l’Île, région de Cayenne, 7 Sep 1989, Cremers 10754 (CAY, P, UC); piste de Saint-Elie, pk 14 à 17, station ECEREX, Sinnamary, 30 Oct 1989, Hoff 5752 (AAU, CAY, STR); piste de Saint-Elie, Sinnamary, 5 Feb 1990, D. E. A. Paris VI 95 (CAY); piste de Saint-Elie, Sinnamary, 15 Mar 1990, Grazzioli 1 (CAY, MPU); piste de Saint-Elie, Sinnamary, 18 Mar 1990, Escoubeyrou 12 (CAY, MPU); chemin de fer Gare-Tigre - Saint-Elie - pk 6 à pk 11, bassin du Sinnamary, 7 Apr 1990, Hoff et al. 6548 (AAU, CAY); piste de Saint-Elie, Sinnamary, 1 Feb 1991, Delavaux 30 (CAY); mont Baduel, Cayenne, 27 Mar 1992, Cremers & Hoff 12958 (CAY); Digue Yiyi, région littorale, 26 May 1992, Toriola-Marbot 121 (CAY); route nationale R. N. 1, savane Yiyi, région littorale, 25 Jun 1992, Toriola-Marbot 297 (AAU, CAY, US); grands pripris de Yiyi, région littorale, 16 Jul 1992, Toriola-Marbot 414 (CAY); piste du Village Eskol, Roura, plaine et montagne de Kaw, 16 Sep 1992, Le Goff & Hoff 250 (CAY); crique Gabrielle, plaine et montagne de Kaw, 17 Sep 1992, Bordenave 193 (AAU, CAY, P); piste de Saint-Elie, Sinnamary, 28 Apr 1994, Albertini 54 (CAY), 56 (CAY), 57 (CAY); montagne de la Trinité, bassin de la Mana, 400 m, 3 Jul 1994, Larpin 1060 (CAY); Dégrad des Cannes, Rémire-Montjoly, 1 Jul 1997, Andrieux & Rousset 22 (CAY); secteur de Saint-Elie - 1, environs du village, bassin du Sinnamary, 130 m, 4 Mar 1997, Boudrie 2743 (CAY); Maripasoula, bassin du Maroni, 28 Jun 1997, Fleury 1179 (CAY, US); bord de la route D 5, à l’est de la rivière des Cascades, à 4,5 km au SSW de Tonnégrande (obs. M. Boudrie, 1 May 1999!); piste Citron à Boeuf-Mort – 3, région de Paul-Isnard (obs. M. Boudrie, 16 Jul 1999!); décharge de Rémire-Montjoly, 10 Jan 2000, Héquet 919 (CAY); piste de Saint-Elie, pk 14 à 17, station de Recherche IRD, Sinnamary, 23 Mar 2000, Prévost 3829 (CAY, MO, P, U, US); montagne Nivré, secteur de Dorlin, à 60 km à l’ENE de Maripasoula (obs. M. Boudrie, 5 Jul 2000!); vallée du Petit Inini, secteur de Dorlin, à 60 km à l’ENE de Maripasoula (obs. M. Boudrie, 6 Jul 2000!); pont sur la crique Yiyi - R. N. 1, région littorale, 9 May 2001, Chaix 480 (CAY); savane Lambert, à 3 km à l’WSW de Montsinéry (obs. M. Boudrie, 24 Jun 2001!); massif Dékou-Dékou - 1, région de Paul-Isnard, 300 m, 12 Aug 2001, Boudrie 3622 (CAY); piste de Saint-Elie, pk 18, Sinnamary, 19 Mar 2002, Tepe 582 (CAY, MU); route nationale RN 4 ́ La Matourienne », face au centre pénitentiaire, Rémire-Montjoly, 6 Oct 2002, David et al. 68 (CAY); Placer Trésor - 2, plaine et montagne de Kaw, 30 Jan 2003, Ek et al. 1458 (CAY, U); Petite Montagne Tortue, bassin de l’Approuague, 26 Feb 2003, Kennedy et al. 5565 (CAY, UBC, UCR); projet de route d’Apatou, entre 0 et 1 km à l’est de la crique Sparouine, 5 m, 23 Sep 2003, Bordenave et al. 7320 (CAY); village de Papaïchton, Centre d’Enfouissement Technique, bassin du Maroni, 23 Sep 2004, Tostain 424 (CAY); piste de Paul-Isnard, à 30 km au SSE de Saint-Laurent-du-Maroni (obs. M. Boudrie, 2 Apr 2005!); piste de Bon Espoir, à 40 km au SSE de Saint-Laurent-du-Maroni (obs. M. Boudrie, 2 Apr 2005!); vallée de la crique Bonne Entente, à 55 km au SSE de Saint-Laurent-du-Maroni (obs. M. Boudrie, 13 Apr 2005!); montagnes Plomb – 2, bassin du Sinnamary, 4 Jul 2005, Delnatte 22 (CAY); piste de Saint-Elie, pk 10, Sinnamary, 18 Sep 2005, Delnatte & Girod 163 (CAY); savanes de Karouabo, région littorale, 11 May 2007, de Granville 17447 (CAY, P); montagne des Chevaux, carrière SCC, Roura, 13 May 2007, Tostain 592 (CAY, NY, P, US); bord du sentier du bagne des Annamites, à 8 km au sud de Montsinéry (obs. M. Boudrie, 26 Oct 2007!); Maripasoula, Centre d’Enfouissement, Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 106-110, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

Boudrie, Michel, Cremers, Georges, Øllgaard, Benjamin (2023): Lycopodiaceae of the Guianas: List of exsiccata and distribution maps. Phytotaxa 601 (2): 103-136, DOI: 10.11646/phytotaxa.601.2.1, URL: http://dx.doi.org/10.11646/phytotaxa.601.2.1

4.7. Phlegmariurus killipii (Herter) B.Øllg. Collections studied from the Guianas: 7 (GU: 2: SU: 1; FG: 4) Guyana: U. Takutu-U. Essequibo, Wassarai Mts., summit of highest peak, 14 km S of S. Kassikaityu R., 1135 m, 8 Sep 1999, Clarke et al. 8281 (NY, US); U. Takutu-U. Essequibo, Wassarai Mts., summit of unnamed peak, 10.5 km S of S. Kassikaityu R., 900 m, 15 Sep 1999, Clarke et al. 8633 (US). Suriname: Wilhelmina Mts, Julianatop, 810 m, 17 Jul 1981, Werkhoven 16720 (BBS). French Guiana: Massif des Tumuc-Humac, monts Tumuc-Humac, 700 m, 2 Aug 1972, de Granville 1097 (CAY); montagnes Bellevue de l’Inini, bassin de l’Inini, 800 m, 19 Aug 1985, Cremers et al. 9030 (CAY); montagnes Bellevue de l’Inini, bassin de l’Inini, 14 Apr 1986, Feuillet 3863 (CAY); monts Atachi Bakka, bassin du Maroni, 700 m, 18 Jan 1989, Cremers et al. 10301 (CAY)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 116, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

List of Lycopodiaceae from the Guianas List of specimens by collectors In the list below, after the collector’s name, the first number (not between parentheses) is the specimen number and the numbers between parentheses correspond, according to the usual presentation in the fascicles of the Flora of the Guianas, to the number of the taxa following the order in our article (see table above), the first one being the genus, second one the species., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 130, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.14. Phlegmariurus tenuicaulis (Underw. & F.E.Lloyd) B.Øllg., Phytotaxa 57: 19. 2012.— Fig. 1. Lycopodium tenuicaule Underw. & F.E.Lloyd, Bull. Torrey Bot. Club 33: 113. 1906.— Urostachys tenuicaulis (Underw. & F.E.Lloyd) Herter, Repert. Spec. Nov. Regni Veg. 19: 165. 1923.— Huperzia tenuicaulis (Underw. & F.E.Lloyd) B.Øllg., Opera Bot. 92: 169. 1987. Type:— DOMINICA. Mt. Diablotin, 1903, Lloyd 878 (holotype NY! [144031]; isotype US-fragm.! [00135542]). Lycopodium linifolium L. var. gracile L’Herm. ex Fée, Mém. Foug. 11: 129. 1866. Type: GUADELOUPE. Sources de la rivière Saint-Louis, Feb. 1845, L’Herminier s.n. (holotype P! [P00559125]; poss. isotypes P! [P00559126], [P00559127], [P00559128], [P00559130], NY! [1754003]). A lax, slender pendulous epiphyte with delicate scattered leaves, the stems usually 20–30 cm long, loosely 2 to 4 times dichotomous. Stems smooth, often reddish, 0.3-0.7 mm thick excl. leaves, shoots 1–1.8 cm wide incl. leaves. Leaves paired to alternate, 1–2 mm apart, spreading to patent, the lamina usually twisted to a vertical position, narrowly elliptic to oblanceolate, slightly asymmetrical to subfalcate, with acute apex, 6–9 × 1–1.5 mm, the margins entire, midrib slender, long-decurrent. Sporangia 1–3 mm wide. Distribution: Greater Antilles (Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico), Lesser Antilles (Guadeloupe, Dominica, Martinique), where it grows at high elevations, around 1000 m (Proctor, 1989). This is the first record for continental South America. Habitats: Montane forests, 460–1600 m elev. Note: This species was overlooked in our previous article (Øllgaard et al., 2020) because the Pipoly specimen had not been found at that time. Collection studied from the Guianas: 1 (GU: 1) Guyana: Cuyuni-Mazaruni Region, at headwaters of Kangu R., W branch, ± 4 km NW of E peak of Mt. Ayanganna, 700 m, 5 Mar 1987, Pipoly et al. 11042 (FDG not found, NY [3835301!], US [00648152!])., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 122-123, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564, {"references":["Proctor, G. R. (1989) Ferns of Puerto Rico and the Virgin Islands. Memoirs of The New York Botanical Garden 53: 1 - 389.","Ollgaard, B., Boudrie, M. & Cremers, G. (2020) The Lycopodiaceae of Guyana, Suriname, and French Guiana. Phytotaxa 433 (2): 101 - 134. https: // doi. org / 10.11646 / phytotaxa. 433.2.3"]}

5.2. Pseudolycopodiella iuliformis (Underw. & F.E.Lloyd) Holub Collections studied from the Guianas: 4 (GU: 4) Guyana: Upper Mazaruni Distr., N slope of Mount Roraima, 850 m, 16 Feb 1985, Renz et al. 14225 (U), 2300 m, 14265 (U); Cuyuni-Mazaruni, Mt. Ayanganna, easternmost peak, 1350 m, 11 Mar 1987, Pipoly et al. 11127 (CAY, FDG not found, MO, NY, US); Cuyuni-Mazaruni, 1 km N of tip of northern prow of Roraima, 2000 m, 25 Feb 1989, Hahn & Gopaul 5507 (P, US); Cuyuni-Mazaruni, Waukauyengtipu eastern summit, 1570 m, 10 Jul 1997, Clarke et al. 5579 (US). Note: Pseudolycopodiella iuliformis has recently been recorded from the Cordillera del Condor in Ecuador and Peru (Zapata et al., 2021)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 124, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564, {"references":["Zapata, N., Perez, A. J., Burgess, K., Rea, E., Neill, D. A., Cevallos, D., Persson, C. & Ollgaard, B. (2021) Pseudolycopodiella iuliformis (Lycopodiaceae, Lycopodielloideae) in Ecuador and Peru; a disjunct species between the Guyana Shield and the Cordillera del Condor. Revista Ecuatoriana de Medicina y Ciencias Biologicas 42: 133 - 139. https: // doi. org / 10.26807 / remcb. v 42 i 2.903"]}

4.9. Phlegmariurus myrsinites (Lam.) B.Øllg. Collections studied from the Guianas: 5 (GU: 5) Guyana: Roraima, 1842, Schomburgk s.n. (US); Roraima, 1842, Schomburgk 2 (BM); s.loc., s.d., M. R. Schomburgk 1181 (BR, P, U); base of Roraima, s.d., im Thurn 230 (K); forest slopes near Roraima, s.d., Jenman s.n. (NY) – holotype of Lycopodium roraimense Underw. & F.E.Lloyd., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 119, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

3.3. Palhinhaea pseudocurvata B.Øllg. Collections studied from the Guianas: 9 (GU: 9) Guyana: Cuyuni-Mazaruni, Mt. Ayanganna, eastern side on steep slopes, 1275 m, 12 Mar 1987, Pipoly et al. 11188 (CAY, FDG not found, MPU, NY, US); Cuyuni-Mazaruni, 2-5 km NW of northern prow of Roraima, 1500 m, 24 Feb 1989, Hahn & Gopaul 5482 B (CAY, NY, U, US); Cuyuni-Mazaruni, Pakaraima Mts, NE plateau of Mt. Ayanganna, 1575 m, 1 Nov 1992, Henkel & Hoffman 93 (US); Cuyuni-Mazaruni, Mt. Wokomung, Pakaraima Mts, 130 m, 20 Feb 1993, Henkel et al. 1525 (CAY, NY, U, US); Potaro-Siparuni Region, Mt. Ayanganna, east face, plateau above second of four escarpments, 1380 m, 17 Jun 2001, Clarke et al. 9265 (NY, US); Potaro-Siparuni Region, Mt. Ayanganna, camp on summit plateau, 2040 m, 23 Jun 2001, Clarke et al. 9479 (US); Potaro-Siparuni Region, Mt. Wokomung, Little Ayanganna, upper slopes of highest point on Mt. Wokomung massif, 1525 m, 5 Jul 2003, Clarke et al. 10545 (US); Potaro-Siparuni Region, Mt. Wokomung, summit, 1560 m, 10 Jul 2003, Clarke et al. 10745 (US); Cuyuni-Mazaruni, Tulameng Mt. along line trail toward summit, 1535 m, 20 Jul 2010, Wurdack et al. 5418 (CAY, MO, US)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 110, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.6. Phlegmariurus intermedius (Trevis.) B.Øllg. Collections studied from the Guianas: 3 (GU: 3) Guyana: Mt. Roraima, 1842-43, Schomburgk 31 (P); s.loc., s.d., M. R. Schomburgk 1192 (B, P, K); s.loc., s.d., Schomburgk s.n. (HBG). Note: This species shows a very fragmented range, with one locality in Guyana, linking the Atlantic Forest of SE Brazil and the Lesser Antilles. Compared to our previous article (Øllgaard et al., 2020: 118) we confirm that this species is also known from the Dominican Republic (Greater Antilles) by a collection from May 2019 (W. Testo 2310, AAU!, VT!)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 116, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564, {"references":["Ollgaard, B., Boudrie, M. & Cremers, G. (2020) The Lycopodiaceae of Guyana, Suriname, and French Guiana. Phytotaxa 433 (2): 101 - 134. https: // doi. org / 10.11646 / phytotaxa. 433.2.3"]}

4.3. Phlegmariurus capillaris (Sodiro) B.Øllg. Collections studied from the Guianas: 2 (GU: 2) Guyana: Mt. Roraima, upper slope, s.d., im Thurn 192 (K); Upper Mazaruni R. basin, Merume Mt, Partang R., 1140 m, 8 Jul 1960, Tillett et al. 44850 (NY)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 113, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.5. Phlegmariurus funiformis (Cham. ex Spring) B.Øllg. Collections studied from the Guianas: 8 (GU: 8) Guyana: Macouria Creek, Essequibo R., Demerara, 1897, Jenman s.n. (NY, US); Mazaruni R., 1897, Jenman s.n. (NY, US); Mazaruni R., Jan 1897, Jenman 7252 (BRG not found, GH); Mazaruni R., s.d., Jenman 2 (K); Mazaruni Station, 2 Aug 1946, de la Cruz 5363 (NY); Upper Mazaruni R. basin, Mt. Ayanganna, along trail near camp, 17 Aug 1960, Tillett & Tillett 45667 (NY); Mt. Latipu, Mazaruni R., 900 m, 22 Aug 1977, Maas & Boyan 2613 (BRG not found, U, Z); Cuyuni-Mazaruni, Camp along Waruma R. 2 km upstream of confluence with Kako R., 600 m, 10 Feb 1989, Hahn & Gopaul 5225 (P, U, US). Note: The populations of Guyana are at the eastern limit of the range of the species., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 115-116, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.15. Phlegmariurus wilsonii (Underw. & F.E.Lloyd) B.Øllg. Collection studied from the Guianas: 1 (SU: 1) Suriname: Emmaketen Mts., zuidlijn, 2 km van splitsing, 700 m, 4 Aug 1959, Daniels & Jonker 825 (U). Note: The population in Suriname is extremely isolated and is the easternmost extension of its range. Probably this species is undercollected in Guyana., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 123-124, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.4. Phlegmariurus dichotomus (Jacq.) W.H.Wagner Collections studied from the Guianas: 87 (GU: 11; SU: 10; FG: 66) Guyana: Demerara, Barima R., Mar 1896, Jenman s.n. (NY); Demerara, Icooroowa, Jenman s.n. (NY); s.loc. s.d., Jenman s.n. (E?); Rio Cuyuni, s.d., Sandwith 844 (K); s.loc., s.d. (prob. 1937), Sandwith 1193 p.p. (K); Upper Rupununi R., near Dadanawa, 14 Jun 1922, de la Cruz 1502 (F, GH, MO, NY, US); Rupununi Distr., between Kuyuwini Landing and Kassikaityu R., 150 m, 23 Oct 1992, Jansen-Jacobs et al. 3069 (U); Region U. Takutu-U. Essequibo, Essequibo R., W bank, 100 m S of mouth of Kuyuwini R., 150 m, 1 Oct 1993, Henkel et al. 3263 (NY, US); Rupununi Dist., Bushmouth Shea to Quitaro R., Bowl Creek, 200 m, 31 Aug 1995, Jansen-Jacobs et al. 4970 (AAU, NY, U, US); U. Takutu-U. Essequibo, Wassarai Mts., upper slopes of northern outlier, 0-0.5 km S of summit, 660 m, 1 Sep 1999, Clarke 8114 (U, US); Upper Essequibo R., Mora forest, 24 Oct 2001, Chanderbali 440 (BBS). Suriname: E rege Perica, s.d. (prob. 1849-1855), Wullschlägel 1872 (P); near Parwa, Feb 1940, Stahel 33 (NY, U); forest at Jandé-kreek, 1.5 hour downstream from Kabel, Suriname-rivier, 22 Jul 1953, Lindeman 4456 (U); Jodensavanne-Mapane kreek area (Suriname R.), vicinity of Camp 8, rainforest, 6 Jul 1955, Schultz 7256 (U); Zuidhelling Julianatop, Wilhelmina-geb., 700 m, 2 Aug 1963, Schultz 10277 (U); Wilhelmina Mts, Zuid R., 2 km above confl. with Lucie R., 235 m, 15 Sep 1963, Irwin et al. 55729 (NY); Brokopondo, 2 km S of Afobaka, forest along Sara Creek, Feb 1965, Donselaar 2124 (U); Lely Mountains, SW plateaus covered with ferrobauxite, 550 m, 28 Sep 1975, Lindeman et al. 496 (U); Brokopondo, Brownsberg Nature Park, Mazaronitop, 500 m, 7 Mar 2003, Christenhusz et al. 2614 (AAU); Marowijne Distr., Lely Mts, 625 m, 5 Dec 2004, Jansen-Jacobs et al. 6917 (U). French Guiana: Cayenne, januario in Avicenniis, s.d. (1781-1789), Richard s.n. (P); Cayenne, Sep in Sylvis, s.d. (1781-1789), Richard s.n. (P); Guyana centralis, Apr 1833, Leprieur s.n. (P); Guyane, rivière du Jam [= Jari?], 1834, Leprieur s.n. (P); s.loc., 1836, Leprieur s.n. (P); s.loc., 1849, Mélinon 108 (P); Saint-Jean, 23 Apr 1914, Benoist 1115 (P); Saint-Jean, 26 Apr 1914, Benoist 1140 (P); haut Oyapock, Crique Tamouri, affluent de la Camopi, Feb 1945, Aubert de la Rüe s.n. (P); s.loc., 5 Nov 1954, Black et al. 54-17507 (CAY); saut Fracas, sur la Mana, 25 Aug 1962, Hallé 676 (P); saut des Polissoirs, bassin de l’Yaloupi, 23 Apr 1970, de Granville 445 (CAY, P); pied du Toukouchipan, versant est, monts Tumuc-Humac, 450 m, 21 Aug 1972, Sastre 1749 (P); mont Galbao, région de Saül, 650 m, 11 May 1973, de Granville 1617 B (P); Saül, Roche Bateau, 23 Jan 1974, Hallé 2281 (MPU); layon Petit Boeuf Mort, région de Saül, 25 Jan 1974, Descoings & Luu 20544 (MPU); Saül, petit circuit Boeuf-Mort, 25 Jan 1974, Hallé 133 (MPU); sommet du Galbao, région de Saül, 16 Mar 1976, Hallé 2483 (MPU); mont Galbao, région de Saül, 650 m, 13 Dec 1976, Cremers 3947 (CAY); mont Galbao, région de Saül, 650 m, 14 Dec 1976, Cremers 3986 (CAY); crique Gabrielle, plaine et montagne de Kaw, 10 m, 2 Feb 1977, Cremers 4237 (CAY, P); saut Pier Kourou, bassin du Tampok, 27 Mar 1977, Cremers 4520 (CAY); fleuve Approuague, Riv. Arataye, saut Pararé, 19 Aug 1977, Sastre 5728 (P); station de l’ Arataye, bassin de l’Approuague, 2 Sep 1977, Sastre 5873 (P); île d’Antecume Pata, bassin du Maroni, 5 Nov 1977, Cremers 4921 (CAY); île d’Antecume Pata, bassin du Maroni, 20 Nov 1977, Cremers 5092 (CAY); fleuve Approuague, riv. Arataye, saut Pararé, 14 Oct 1978, Sastre 6202 (P); piste de Carbet Maïs vers le Pic Matécho, région de Saül, 14 Jul 1979, de Granville 3103 (CAY); Piste de Risquetout, région de Cayenne, 4 Dec 1979, de Granville 3203 (CAY); massif des Emerillons, région des Emerillons, 250 m, 16 Sep 1980, Cremers 6704 (CAY); massif des Emerillons, région des Emerillons, 300 m, 20 Sep 1980, Cremers 6733 (CAY); massif Lucifer, région de Paul-Isnard, 550 m, 9 Nov 1982, Billiet & Jadin 1643 (BR, CAY); massif Lucifer, région de Paul-Isnard, 550 m, 8 Feb 1983, Cremers 7909 (CAY); lieu-dit Citron, région de Paul-Isnard, 100 m, 12 Feb 1983, Cremers 7962 (CAY); saut Awali, bassin du Tampok, 9 Apr 1983, Feuillet 796 (CAY); pied de la falaise du second sommet, 1 km W de la DZ naturelle, 3-4 km ENE du Grand Croissant, bassin de l’Oyapock, 130 m, 5 Dec 1983, Feuillet 1183 (CAY, P); montagne de Kaw, plaine et montagne de Kaw, 100 m, 1 Apr 1984, de Granville 6709 (CAY, P); Trois-Sauts - village Wayampi, bassin de l’Oyapock, 16 Aug 1985, Prévost & Grenand 1950 (CAY); montagne La Fumée 1, région de Saül, 21 Nov 1985, Prévost 2102 (AAU, CAY); mont Galbao, région de Saül, 650 m, 15 Jan 1986, de Granville et al. 8737 (CAY, P, US); 2 km W of Saül, track Limonade, 180 m, 11 Aug 1986, Ek 482 (U); layon Limonade, région de Saül, 200 m, 22 Sep 1986, Ek & Montfoort 148 (CAY, U); montagne La Fumée 1, région de Saül, 200 m, 22 Jun 1988, Windisch 5276 (AAU, CAY); saut Gobaya Soula, bassin du Maroni, 125 m, 6 Jan 1989, Cremers et al. 10118 (CAY); monts Atachi Bakka, bassin du Maroni, 450 m, 10 Jan 1989, Cremers et al. 10181 (B, CAY, US); route de Bélizon, ca. 500 m S of Eaux Claires, Saül, 250 m, 6 Feb 1990, Mori & Gracie 21143 (NY); île d’Antecume Pata, bassin du Maroni, 135 m, 23 Mar 1991, Veth 102 (CAY, U); barrage de Petit-Saut, bassin du Sinnamary, 4 Apr 1994, Cremers 13275 (CAY); montagne de Kaw, plaine et montagne de Kaw, 300 m, 27 Nov 1994, Billiet & Jadin 6422 (BR, CAY); montagne de la Trinité, bassin de la Mana, 400 m, 12 Mar 1997, de Granville 13300 (CAY, P); savane-roche Dachine, 150 m, 8 Apr 1997, Cremers & Crozier 14812 (CAY); secteur de Saint-Elie - 2, ouest du village, bassin du Sinnamary, 200 m, 21 Jun 1998, Boudrie 3025 (CAY); route de Bélizon - entre Saül et le layon Grand Boeuf Mort, région de Saül, 250 m, 2 Jul 1998, Mori et al. 24815 (CAY); colline d’Augusta et environs, à 1 km à l’est de la vallée de la crique Elysée, à 5 km à l’WNW de Citron, région de Paul-Isnard (obs. M. Boudrie, 9 Jul 1999!); massif Dékou-Dékou - 2, région de Paul-Isnard, 250 m, 12 Apr 2001, Boudrie 3486 (CAY); sommet du Dékou-Dékou - 8, région de Paul-Isnard, 565 m, 17 Aug 2001, Boudrie 3654 (CAY); camp Voltaire, bassin de la Mana, 70 m, 17 Dec 2002, Perez-C. et al. 768 (CAY, QCA); sentier botanique de la Réserve Trésor, montagne de Kaw, à 14 km au sud de Roura (obs. M. Boudrie, 16 Feb 2003!); station de l’Arataï, Réserve Naturelle des Nouragues, 160 m, 23 Feb 2003, Boudrie & groupe FOG-2003 3887 (CAY); inselbergs du haut Marouini – A, inselberg de la D. Z., 180 m, 24 Jun 2004, de Granville & Crozier 16426 (CAY); monts Kotika, plateau latéritique sommital, 730 m, 21 Feb 2005, de Granville et al. 16912 (CAY, P, US); Réserve Naturelle de la Trinité, secteur Aïmara, 27 Jan 2006, Girod & Héquet 3484 (CAY); monts Kotika, plateau latéritique sommital, 730 m, 3 Sep 2007, Delnatte 1336 (CAY); crique Limonade, confluence avec la crique Cambrouze, région de Saül, alt. 180 m, 16 Feb 2008, Tostain et al. 1828 (CAY); massif Dékou-Dékou, sommet oriental, flanc nord, région de Paul-Isnard, 300 m, 3 Jun 2008, Delnatte 1901 (CAY); Abattis Kotika, bassin du Maroni, 85 m, 11 Jan 2012, Silland et al. 353 (CAY); monts Atachi Bakka 2, bassin de l’Inini, 170 m, 4 Feb 2012, Girault et al. 676 (CAY); canal Roy, marais de Kaw, à 2,6 km au nord-est du village de Kaw, commune de Régina (obs. M. Boudrie, 27 Jul 2012!); crique Grande Waki nord, bassin de la Waki, 140 m, 18 Aug 2012, Girault & Léotard 794 (CAY). Note: The large number of collections in French Guiana and the presence of this species in Brazil suggests that this species is under-collected in Guyana and in Suriname., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 113-115, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564, {"references":["Sastre, C., 1749 (4 - 4); 5728 (4 - 4); 5873 (4 - 4); 6202 (4 - 4)","Granville, J. - J. de, 247 (3 - 2); 266 (3 - 2); 445 (4 - 4); 1097 (4 - 7); 1617 A (4 - 13); 1617 B (4 - 4); 2243 (4 - 13); 3103 (4 - 4); 3203 (4 - 4); 6709 (4 - 4); 13300 (4 - 4); 17747 (3 - 2)","Billiet, F. & Jadin, B., 1044 (5 - 4 a); 1094 (3 - 2); 1479 (3 - 2?); 1643 (4 - 4); 1859 (5 - 4 a); 4482 (4 - 13); 4644 (5 - 4 a); 6422 (4 - 4)","Prevost, M. - F. & Grenand, P., 866 (4 - 8 b); 1950 (4 - 4)","Tostain, O. et al., 5973 (4 - 1); 1828 (4 - 4); 2271 (5 - 4 a)","Delnatte, C., 22 (3 - 2); 1336 (4 - 4); 1901 (4 - 4)"]}

5.5. Pseudolycopodiella tatei (A.C.Sm.) Holub Collections studied from the Guianas: 5 (GU: 2; SU: 2; FG: 1) Guyana: Mt. Roraima, at Philipp Swamp, 1570 m, 11 Nov 1927, Tate 350 (NY); Potaro-Siparuni, Pakaraima Mts., Upper Ireng R., 750 m, 3 Feb 1993, Henkel et al. 1158 A (CAY, NY, P, U, US). Suriname: Kappelsavane, S Tafelberg, 300 m, 25 Feb 1961, Kramer & Hekking 3040 p.p. (U p.p., US p.p.); W of Marataka, at Saparra-kreek, 8 May 1965, Maas & Tawjoeran 3218 (BBS, U). French Guiana: Savane incluse de Nancibo, Roura, plaine et montagne de Kaw, 15 m, 25 Sep 2011, Boudrie & Léotard 4515 (CAY). Note: A very rare species, showing a disjunct distribution (Guianas and Atlantic Forest)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 128-129, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

4.8a. Phlegmariurus linifolius (L.) B.Øllg. var. linifolius Collections studied from the Guianas: 1 (FG: 1) – however not confirmed for French Guiana (see note below). French Guiana: s.loc., Montagne donavit [= probably given to the mycologist Camille Montagne by Leprieur], 1859, Leprieur 12 (P). Note: Leprieur’s specimen from French Guiana is the only one for the Guianas. Contrary to what we have mentioned in our article from 2020, this specimen was in fact collected by Leprieur rather than by the mycologist C. Montagne and was given to C. Montagne by Leprieur. It is well known that Leprieur sent many specimens from French Guiana to Montagne for his studies, and Montagne never visited French Guiana (Hoff, 2000). In addition, there are some doubts about this collection because it appears that Leprieur, in the period 1850–1858, stayed in Martinique where P. linifolius var. linifolius is present. Therefore, it is possible that this specimen was collected in Martinique rather than in French Guiana. It is unclear whether the date 1859 is the date when the lycopod was collected, or when the specimen was given to Montagne., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 117-118, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564, {"references":["Hoff, M. (2000) Index of French Guiana Botanists and Plant Collectors. Herbier, Universite de Strasbourg, Institut de Botanique, Strasbourg, 356 pp."]}

3.4. Palhinhaea steyermarkii (B.Øllg.) Holub Collections studied from the Guianas: 5 (GU: 5) Guyana: Mt. Roraima, 2315 m, 2 Nov 1973, R . Persaud 162 (NY); Mt. Roraima, 2300 m, 16 Feb 1985, Renz et al. 14254 (AAU: photo ex U, U); Dist. Mazaruni-Potaro: Roraima, summit, La Proa Camp E of border, near Lake Gladys, 2800 m, 14 Apr 1988, Liesner 23285 (UC); Cuyuni-Mazaruni R., Pakaraima Mts, NE plateau of Mt. Ayanganna, 1600 m, 3 Nov 1992, Hoffman & Henkel 3170 (U, US); Potaro-Siparuni Region, Mt. Ayanganna, east face, edge of summit plateau, 2000 m, 24 Jun 2001, Clarke et al. 9548 (US). Additional specimens: Two specimens, collected by Ule in the Roraima massif, and determined by us as Palhinhaea steyermarkii, are mentioned below because it is uncertain if these collections were made on the Guyanian side: Roraima, 2300 m, Dec 1909, Ule 8416 (BONN-Nessel 200) and 8496 (BONN-Nessel 201, K, L)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 111, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

3.1. Palhinhaea camporum (B.Øllg. & P.G.Windisch) Holub Collections studied from the Guianas: 19 (GU: 10; SU: 9) Guyana: Parabaru savanna, watershed between Rupununi and Kuyuwini R., 15-17 Feb 1938, A. C. Smith 3072 (F, GH, K, MO, NY, P, S, U, US); Orinduik Falls, Ireng R., 15 Aug 1958, Harrison 1459 (BRG not found, K, NY); Upper Mazaruni River Basin, Ayanganna plateau, Haieka savanna, E side of Haieka, 2 mi E of Chinoweing Village, 740 m, 21 Aug 1960, Tillett et al. 45217 (AAU, NY); Ité swamp, 12 mi. N of Manari Cr., 1 mi W of north road, Rupununi, 100 m, 26 Sep 1963, Goodland 883 (COL, M, US); U. Takutu-U. Essequibo, Makatui savanna, ca. 3,5 km SW of Aishalton, 17 Nov 1982, Stoffers et al. 378 (CAY, U); U. Takutu-U. Essequibo, Kuyuwini R., Aishalton Landing, upstream 2-3 km, N & S banks, 200 m, 9 Sep 1983, Henkel et al. 2778 (CAY, U, US); Cuyuni-Mazaruni, vicinity of Chinoweing Village, 660 m, 21 Feb 1987, Pipoly et al. 10457 (AAU, CAY, FDG not found, NY, P, US); Cuyuni-Mazaruni, Utshe camp on savanna, 975 m, 22 May 1990, McDowell & Gopaul 2719 (CAY, NY, U, US); Rupununi District, Shea Rock, 160 m, 10 Feb 1994, Jansen-Jacobs et al. 3690 (CAY, NY, U); U. Takutu-U. Essequibo, Parabara Savanna, trail from Karaudarnau to Kuyuwini R., 270 m, 1 Jun 1997, Clarke 5103 (U, US). Suriname: Apikollo savanna, 29 Jan 1911, Hulk 65 (U); Boven-Sipaliwini, kamp IV, groote savanne, 23 Oct 1935, Rombouts 217 (U); Boven-Sipaliwini, kamp XIX, Morro Grande, 15 Feb 1936, Rombouts 476 (U, 2 sheets); Boven-Lawa, Palaime-savanne, 2 May 1952, Geijskes s.n. (U); s.loc., s.d., Geijskes s.n. (GH); R. Sipaliwini, rivulet Palaime, Palaime Savanna, 24 Feb 1963, Wessels Boer 773 (U); Sipaliwini savanna, foot of hill, 26 Aug 1966, Donselaar 3609 (BBS); W Sipaliwini savanna area on Brazilian frontier, N of 4- Gebroeders’ creek, 260 m, 6 Sep 1968, Oldenburger et al. 75 (BBS, U); small Sipaliwini savanna, 280 m, 30 Dec 1968, Oldenburger et al. 722 (U). Note: In Guyana and Suriname this species is restricted to the internal savannas, at middle elevation., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on pages 105-106, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564

5.4b. Pseudolycopodiella meridionalis (Underw. & F.E.Lloyd) Holub var. mesetarum (B.Øllg.) B.Øllg., Boudrie & Cremers, comb. nov. —Basionym: Lycopodiella caroliniana (L.) Pic.Serm. var. mesetarum B.Øllg., Nord. J. Bot. 23: 39–40. 2004).—Synonym: Pseudolycopodiella caroliniana (L.) Holub var. mesetarum (B.Øllg.) B.Øllg., Phytotaxa 57: 20. 2012. Type:— VENEZUELA. Territorio Federal Amazonas: Departamento Atabapo: Cerro Marahuaca-Huha. Cumbre altiplanicie no arbolada, más o menos plana, con piedras expuestas, río abajo, 03°40’N, 65°30’W, 2580 m, 31 Jan 1982, Steyermark et al. 125967 (AAU! holotype). This new combination, invalid in Øllgaard et al. (2020: 131), is hereby validated. Collection studied from the Guianas: 1 (GU: 1) Guyana: Mt. Roraima, summit, 26 Nov 1927, Tate 380 p.p. (NY p.p.)., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 128, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564, {"references":["Ollgaard, B., Boudrie, M. & Cremers, G. (2020) The Lycopodiaceae of Guyana, Suriname, and French Guiana. Phytotaxa 433 (2): 101 - 134. https: // doi. org / 10.11646 / phytotaxa. 433.2.3"]}

1.1. Lycopodiella longipes (Hook. & Grev.) Holub Collections studied from the Guianas: 21 (GU: 18; SU: 3) Guyana: Near Roraima, Demerara, 1897, Jenman s.n. (NY); between Rupununi and Kuyuwini R., Parabaru Savanna, 16 Feb 1938, Smith A.C. 3067 (GH, K, MO, NY, P, S, U, US); Upper Mazaruni River Basin, Merume Mountains, near mouth of Río Partang, 460 m, 14 Jun 1960, Tillett & Tillett 43835 (AAU, K, NY); Pakaraima Mountains, Kopinang Savanna, Kopinang Falls, 800 m, 30 Aug 1961, Maguire 46007 (NY, U, US, VEN); Pakaraima Mts, Kopinang Savanna, Kopinang Falls, 800 m, s.d., Maguire s.n. (K, US, VEN); Pakaraima Mts., Kako Amerindian village, Kako R., 13 Nov 1979, Maas & Vestra 4400 (U); Pakaraima Mts., Mt. Aymatoi, 1150 m, 15 Oct 1981, Maas et al. 5689 (U); U. Takutu-U. Essequibo, Aishalton Hospital, 19 Nov 1982, Stoffers et al. 462 (CAY, U); Cuyuni-Mazaruni, Pakaraima Mts, Imbaimadai Creek, W of Imbaimadai, 505 m, 22 Jun 1986, Pipoly & Alfred 7981 (AAU, B, BRG not found, CAY, NY, P, U, US); Cuyuni-Mazaruni, Chinoweing Village, Ayanganna Plateau, 660 m, 20 Feb 1987, Pipoly et al. 10371 (AAU, NY); Cuyuni-Mazaruni, vicinity of Chinoweing Village, 660 m, 21 Feb 1987, Pipoly et al. 10463 (AAU, CAY, FDG not found, NY, P, US); Cuyuni-Mazaruni Region, savanna 2-3 km W of Maipuri Falls, Karowrieng R., 600 m, 24 Dec 1989, Gillespie & Smart 2881 (NY, U, US); Cuyuni-Mazaruni, Pakaraima Mts, Imbaimadai, just NW of airstrip, 510 m, 16 May 1992, Kelloff et al. 796 (NY); Rupununi Distr., S of Aishailton, 150 m, 16 Feb 1997, Jansen-Jacobs et al. 5364 (U); Cuyuni-Mazaruni Region, Mt. Maringma summit, 2110 m, 17 Jun 2004, Clarke et al. 11763 (US); Cuyuni-Mazaruni, Kako R., 485 m, 6 May 2009, Wurdack et al. 4833 (CAY, NY, US); Cuyuni-Mazaruni, Imbaimadai, Mazaruni R., 515 m, 29 Jul 2010, Redden et al. 7113A (BRG not found, CAY, NY, TEX, US); Cuyuni-Mazaruni, below 1st escarpment of (four) of Kamasuka Mt., 0-1 mi SW of Ducking 1st camp, 640 m, 15 May 2012, Wurdack et al. 5638 (NY); Suriname: Boven-Sipaliwini, kamp IV, 23 Oct 1935, Rombouts 218 (U); Boven-Tapanahoni, Palaime savanne, 9 Mar 1954, Geijskes s.n. (U); W. Sipaliwini savanna N of ‘4- Gebroeders’ creek, on Brazilian border, 260 m, 6 Sep 1968, Oldenburger et al. 76 (BBS, U). Note: A rare species in the Guianas, at the eastern limit of its range in Suriname, located on large savannas of the interior., Published as part of Boudrie, Michel, Cremers, Georges & Øllgaard, Benjamin, 2023, Lycopodiaceae of the Guianas: List of exsiccata and distribution maps, pp. 103-136 in Phytotaxa 601 (2) on page 104, DOI: 10.11646/phytotaxa.601.2.1, http://zenodo.org/record/8129564