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Lucilina colantonii n. sp. (Fig. 15) Type material. Holotype: MZB 50549, intermediate valve, width 2.7 mm (Figs 15 A–C). Paratype: RGM.1356860, intermediate valve, width 3.9 mm (Figs 15 D–F). Type locality. Hurghada, Egypt (St. 16). Other material examined. Egypt, Hurghada: St. 13: 2 intermediate valves (BD 213); St. 14: 2 intermediate valves (BD 214); St. 16: 1 intermediate valve (BD 215). Recent material examined. Sudan, Shab Runi Reef (P078): 2 intermediate valves, Figs 15 J–L (BD 216, MZB 50551); Sudan, Sanganeb Reef (D08): 1 intermediate valve (MZB 60260); Egypt, Gulf of Aqaba (P447): 4 intermediate valves, Figg 15 G–I (BD 217; MZB 50550); Egypt, Gulf of Aqaba, Blue Hole (P447A): 1 intermediate valve (BD 218). Maximum width of the intermediate valves: 5.5 mm. Type stage. Late Pleistocene, last interglacial MIS5e. Etymology. This species honours the marine geologist Paolo Colantoni (1934–2015) who first introduced M.T. to the study of recent and fossil molluscs of the Gulf of Aqaba. Diagnosis. Intermediate valve rectangular, moderately elevated, semicarinate in anterior profile, posterior margin almost straight at both sides of prominent apex, almost triangular, pleural areas with longitudinal regular striae of small, roundish granules, jugal area large, smooth, lateral areas not raised, with irregular radial striae of similar granules. Articulamentum white, apophyses small, triangular, jugal sinus wide, one slit, teeth finely grooved on upper side. Description. Intermediate valve rectangular, L/W ratio = 0.53–0.63, anterior margin convex, moderately elevated (H/W = 0.36–0.41), semicarinate in anterior profile, side margins slightly rounded, posterior margin almost straight at both sides of prominent apex, almost triangular, forming an angle of ca 120–130°, pleural areas with longitudinal and enough regular striae of small, roundish granules, ca 10–12 on each side, with reduced spaces between the striae of granules, jugal area large, smooth, lateral areas not raised, with 6–7 irregular radial striae of similar granules, ocelli scarcely visible in anterior third. Granules of longitudinal striae entirely covered by aesthetes of the same dimensions, not regularly arranged. Articulamentum white, apophyses small, triangular, jugal sinus wide, one slit, slit rays not indicated, teeth finely grooved on upper side. Remarks. Only poorly preserved and often incomplete intermediate valves are present in the studied material. The species is similar to Lucilina aegyptiaca n. sp., from which it differs in several characters (Tab. 3). The distribution of the aesthetes on the granules of the longitudinal striae is unusual, they are of the same di- mensions and arranged without a particular order, in contrast to the norm (generally with one or more central macroaesthetes and several microaesthetes arranged along the edge of the granule). Numerous valves of Lucilina colantonii n.sp. were identified in extant Red Sea material still unpublished, mixed in samples initially labeled L. sueziensis and L. perligera. Two such valves are illustrated for comparison (Fig. 15 G–L). No significant differences appear between extant and fossil valves. The extant valves are well preserved and show additional characters not preserved in the fossil state, namely the presence of longitudinal striae of small roundish granules with a diameter up to 60 µm, and ocelli with a diameter of 33–37 µm. Distribution. Late Pleistocene: Egypt (Hurghada: this study). Present-day (this study): Sudan, Sha’ab Rum Reef, Sanganeb Reef; Egypt, Gulf of Aqaba (Blue Hole).

Genus Lucilina Dall, 1882 Type species. Chiton confossus Gould, 1846 (= Chiton lamellosus Quoy & Gaimard, 1835), by subsequent designation (Pilsbry 1893). Remarks. Following the suggestions of some authors (e.g., Strack 2003; Schwabe 2004, 2006; Schwabe et al. 2008; Schwabe & Pittman 2014), the genus Lucilina is here used at full genus level, and not as a subgenus of Tonicia Gray, 1847. The species of Tonicia are geographically restricted to the South Eastern Pacific region of Latin America with a single species in the Caribbean, while members of Lucilina are widespread throughout the tropical Indo-Pacific. Lucilina is well represented in the Indian Ocean by six species, L. sueziensis (Reeve, 1847), L. perligera Thiele, 1909, L. pectinoides (Sykes, 1903), L. ceylonica (Leloup, 1936), L. carnosa (Kaas, 1979) and L. indica (Leloup, 1981), of which only the first two are known from the Red Sea. Distribution. Widespread in the Indian Ocean, the Indo-Pacific, Australia, the Central Pacific Ocean and Japan. The fossil record extends back to the middle Oligocene of France, Aquitaine Basin (Dell���Angelo et al. 2018a), the Neogene of west Pacific islands (Ladd 1966; Schwabe et al. 2008) and the Pleistocene of the Red Sea (Issel 1869)., Published as part of Dell'Angelo, Bruno, Landau, Bernard M., Sosso, Maurizio & Taviani, Marco, 2020, Late Pleistocene Red Sea Mollusca: 1. Polyplacophora, pp. 401-449 in Zootaxa 4772 (3) on page 415, DOI: 10.11646/zootaxa.4772.3.1, http://zenodo.org/record/3819654, {"references":["Dall, W. H. (1882) On the genera of chitons. Proceedings of the United States National Museum, 4, 279 - 291. https: // doi. org / 10.5479 / si. 00963801.4 - 228.279","Gould, A. A. (1846) On the shells collected by the United States Exploring Expedition, commanded by Charles Wilkes, U. S. N. Proceedings of the Boston Society of Natural History, 2 (14), 141 - 145. https: // doi. org / 10.5962 / bhl. part. 3318","Quoy, J. R. C. & Gaimard, J. P. (1835) Voyage de decouvertes de l' \" Astrolabe \". Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1928 - 1829, sous le commandement de M. J. Dumont d'Urville. Zoologie 3, Mollusques, Genre Oscabrion. J. Tastu, Paris, 43 pp., pls. 73 - 75. [pp. 369 - 411]","Pilsbry, H. A. (1892 - 1894) Monograph of Polyplacophora. In: Tryon, G. W., Manual of Conchology. Vol. 15 & 16. Academy of Natural Sciences, Philadelphia, pp. 1 - 128, pls. 1 - 30 (1892) + pp. i-xxxiv, 129 - 350, pls. 31 - 68 (1893) & pp. 1 - 64, pls. 1 - 10 (1893) + pp. 65 - 133, pls. 11 - 17 (1894).","Strack, H. L. (2003) Class Polyplacophora. In: Hylleberg, J. & Kilburn, R. N. (Eds.), Tropical Marine Mollusc Programme. Marine Molluscs of Vietnam. Phuket Marine Biological Center Special Publication, 28, pp. 12 - 15.","Schwabe, E. (2004) The Polyplacophora (Mollusca) collected during the First International Marine Biodiversity Workshop for Rodrigues (western Indian Ocean), with the description of a new species. Journal of Natural History, 38, 3143 - 3173. https: // doi. org / 10.1080 / 00222930410001695114","Schwabe, E., Sirenko, B. I. & Seeto, J. (2008) A checklist of Polyplacophora (Mollusca) from the Fiji Islands. Zootaxa, 1777 (1), 1 - 52. https: // doi. org / 10.11646 / zootaxa. 1777.1.1","Schwabe, E. & Pittman, C. (2014) New records of chitons (Mollusca: Polyplacophora) from Hawaii, including the description of a new species of Weedingia Kaas, 1988. Venus, 72, 49 - 64.","Gray, J. E. (1847) A list of the genera of Recent Mollusca, their synonyms and types. Proceedings of the Zoological Society of London, 15, 129 - 206. https: // doi. org / 10.1111 / j. 1469 - 7998.1847. tb 00164. x","Reeve, L. (1847) Conchologia iconica, or illustrations of the shells of molluscous animals. Monograph of the genus Chiton. Vol. 4. Reeve, Brothers, London, 64 pp., 28 pls. [pp. 21 - 84]","Thiele, J. (1909) Revision des Systems der Chitonen. Zoologica, 22, 1 - 132, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 11245","Sykes, E. R. (1903) Report on the Polyplacophora collected by Professor Herdman, at Ceylon, in 1902. Ceylon Pearl Oyster Fisheries, Supplementary Reports, 4, 177 - 180, pl. 1.","Leloup, E. (1936) Descriptions de deux especes d'amphineures provenant de Trincomali (Ceylon). Proceedings of the Malacological Society of London, 22 (2), 51 - 61.","Kaas, P. (1979) The Chitons (Mollusca: Polyplacophora) of Mozambique. Annals of the Natal Museum, 23, 855 - 879.","Leloup, E. (1981) Chitons de Tulear, Reunion, Maurice et Tahiti. Bulletin Institut Royale des Sciences Naturelles de Belgique, 53, 1 - 46, 4 pls.","Dell'Angelo, B., Lesport, J. - F., Cluzaud, A. & Sosso, M. (2018 a) The Oligocene to Miocene chitons (Mollusca: Polyplacophora) of the Aquitaine Basin, southwestern France, and Ligerian Basin, western France. Part 1: Leptochitonidae, Hanleyidae, Ischnochitonidae, Chitonidae, Spinochitonidae fam. nov. and Schizochitonidae. Bollettino Malacologico, 54, 1 - 47.","Ladd, H. S. (1966) Chitons and Gastropods (Haliotidae through Adeorbidae) from the western Pacific islands. Geological Survey Professional Paper, 531, 1 - 98, 16 pls. https: // doi. org / 10.3133 / pp 531","Issel, A. (1869) Malacologia del Mar Rosso. Ricerche zoologiche e paleontologiche. Parte terza. Catalogo delle conchiglie fossili raccolte sulle spiagge emerse del Mar Rosso. Biblioteca Malacologica, Pisa, 61 pp. [pp. 243 - 303]"]}

Lucilina aqabaensis n. sp. (Fig. 13) Type material. Holotype: MZB 50545, a tail valve, St. 9, width 6.5 mm (Figs 13 A–F). Type locality. Gulf of Aqaba, Saudi Arabia (Ash Shaykh Humayd: this study): St. 9. Type stage. Late Pleistocene, last interglacial MIS5e. Etymology. The name refers to the Gulf of Aqaba, source of the material here described. Diagnosis. Tail valve semioval, mucro posterior, obtuse, antemucronal slope slightly convex, postmucronal slope perpendicular. Tegmentum finely granulose, antemucronal and postmucronal areas with concentric growth lines, characterised by the presence of groups of fused granules, irregularly arranged in concentric striae. Articulamentum white, apophyses well developed, jugal sinus wide, 10 slits, teeth slightly forwardly directed, finely grooved on upper side. Description. Tail valve semioval, length less than half width (L/W = 0.42), front margin straight, hind margin angularly rounded, mucro posterior, obtuse, antemucronal slope slightly convex, postmucronal slope perpendicular. Tegmentum finely granulose, roundish granules irregularly dispersed, extending over entire surface of valve, both antemucronal and postmucronal areas with concentric growth lines, characterised by the presence of groups of fused granules, more frequent and larger in postmucronal area along hind margin, giving the impression of being irregularly arranged in concentric striae. Roundish granules small and close-set, diameter ca 48–60 µm, with 1 large central macroaesthete and up to 10 microaesthetes along margin. Articulamentum white, apophyses well developed, jugal sinus wide, provided with a short plate, 10 slits, slit rays not indicated, teeth relatively long, slightly forwardly directed, all teeth finely grooved on upper side, denticulated on outer edge, eaves narrow, solid. Remarks. We assign this unique tail valve to the genus Lucilina based on the presence of having grooved teeth on the articulamentum (and not a callus as in the genus Onithochiton Gray, 1847), and having the mucro subterminal (and not subcentral as in the genus Tonicia Gray, 1847) (Kaas et al. 2006: 294). We did not observe the presence of ocelli in the postmucronal area. The characters of this single tail valve are so distinctive that they warrant description as a new species. The peculiar granulose sculpture of this species is different from that of other Lucilina species known from the Red Sea (including those described herein), and Indian Ocean (Kaas et al. 2006). Distribution. Late Pleistocene: Saudi Arabia, Gulf of Aqaba (Ash Shaykh Humayd: this study).

Lucilina sueziensis (Reeve, 1847) (Fig. 9) Chiton sueziensis Reeve 1847, pl. 20, fig. 134. Chiton (Tonicia) sueziensis; Issel 1869, p. 235; Moazzo 1939, p. 220. Chiton (Tonicia) sp.; Issel 1869, p. 318 (ex Savigny pl. 3, fig. 7). Tonicia suezensis (sic); Sykes 1907, p. 34; Leloup 1960, p. 40, figs 6, 8, pl. 1, fig. 1; Leloup 1980, p. 12; Vine 1986, p. 125. Lucilina sueziensis; Lamy 1938, p. 88; Blatterer 2019, p. 52, pl. 4, fig. 10 a���n. Tonicia sueziensis; Ferreira 1983, p. 271, figs 25���28; Dekker & Orlin 2000, p. 7; Slieker 2000, p. 50, fig. 28; Abubakr 2004, p. 73; Dell���Angelo et al. 2004, p. 56. Tonicia (Lucilina) sueziensis; Kaas & Van Belle 1988, p. 124, figs 41���44; Van Belle & Wranik 1991, p. 375, fig. 21; Strack 1993, p. 19, pl. 5, fig. 1, pl. 6, figs 1���2; Bosch et al. 1995, p. 191; Kaas & Van Belle 1998, p. 182; Anseeuw & Terryn 2004, p. 15, figs 5, 50���54; Kaas et al. 2006, p. 315, fig. 129, map 158; Dinapoli & Janssen 2009, p. 20, pl. 11 a���d. Tonicia (Lucinila) (sic) sueziensis; Mohammadian 2011, p. 211. Non Chiton (Tonicia) sueziensis; Issel 1869, p. 317 (ex Savigny pl. 3, fig. 4) (= Acanthopleura vaillantii de Rochebrune, 1882, juv., fide Kaas et al. 2006). Type material. BMNH 1591.2.7.7, lectotype designed by Ferreira (1983: 272, figs 26���27). Type locality. Gulf of Suez. Material examined. Jordan, Aqaba (Yamanie) (St. 1): 5 valves (4 intermediate and 1 tail) (MZB 60254). Saudi Arabia, Gulf of Aqaba (Ash Shaykh Humayd: this study): St. 4: 4 intermediate valves (MZB 60255); St. 5: 3 intermediate valves (MZB 60256); St. 7: 5 valves (4 intermediate and 1 tail) (MZB 60257); St. 8: 7 valves (3 head and 4 intermediate), Fig. 9J (MZB 50534); St. 9: 4 valves (1 head, 1 intermediate and 2 tail), Fig. 9L (MZB 50535). Egypt, Hurghada: St. 12: 20 valves (2 head, 13 intermediate and 5 tail) (BD 198); St. 13: 108 valves (13 head, 70 intermediate and 25 tail), Figs 9 A���I (BD 199; MZB 50533 a���c; RGM.1356854); St. 14: 28 valves (3 head and 25 intermediate) (BD 200); St. 14bis: 17 valves (11 intermediate and 6 tail), Fig. 9K (BD 201; MZB 50533 d); St. 16: 46 valves (5 head, 35 intermediate and 6 tail) (BD 202); St. 17: 1 tail valve (BD 203). Maximum width: 3.8 / 5.7 / 7.5 mm. Description. Head valve semicircular, posterior margin widely V-shaped, straight to weakly convex at both sides of minute apical notch, front slope slightly convex, tegmentum coarsely sculptured with up to 20 radial rows of small, warty, V-shaped rugosities, interstices each with row of minute, round to oval, black ocelli. Intermediate valves rectangular, L/W = 0.33���0.40, front margin slightly sinuous, concave in wide central part, convex at pleurae, moderately elevated (H/W = 0.34���0.41), anterior profile subcarinated, side margins rounded, posterior margin concave at both sides of bluntly protruding apex, lateral areas not raised, sculptured like head valve, 2���4 radial rows, the anterior one with larger rugosities, defining the diagonal line, ocelli anterior on 1/3 to 1/2 of lateral area, pleural areas with up to 14 longitudinal, sinuous, slightly granulose wrinkles, fanning out anteriorly, diverging at sides, converging and gradually shortening towards wide, smooth jugal area. Tail valve semicircular, L/W = 0.51���0.56, front margin evenly convex, straight in jugal part, mucro subcentral, not prominent, antemucronal and postmucronal slopes straight, some valves with postmucronal slope steep, strongly convex, antemucronal area sculptured like central areas, postmucronal area like head valve. Articulamentum with apophyses wide, triangular, trapezoidal in tail valve, connected across wide, shallow sinus by short, pectinated, jugal plate, insertion plates rather long, slit formula 8���10/1/10-12, teeth strongly fluted on outside, eaves narrow, solid. Remarks. Lucilina sueziensis is a highly variable species, especially in regards to the sculpture of the tegmentum (Ferreira 1983; Strack 1993; Anseeuw & Terryn 2004). The mucro of the tail valve is normally in a subcentral position, with antemucronal and postmucronal slopes almost straight or slightly concave (Fig. 9I), but some tail valves show a variability in morphology, with the mucro in a more posterior position, and the antemucronal and postmucronal slopes much more convex (Figs 9 K���L). Ferreira (1983: 271) noted in the description of the tail valve ���mucro prominent, central to slightly posterior; postmucro sloping at 30���45�� (steeper in larger specimens)���, agreeing with the valve illustrated in Fig. 9I, but the lectotype selected and figured (Ferreira 1983: fig 27���28) shows a postmucronal slope more similar to the valves illustrated in Fig. 9 K���L. A similar species is Lucilina carnosa (Kaas, 1979) from Madagascar, Mozambique, R��union, Seychelles, Amirante, and Maldive Islands. That species was synonymised with L. sueziensis by Ferreira (1983), but differs in having stronger sculpture, the tail valve subtriangular, as well as various details of the girdle (Kaas et al. 2006), and the two are therefore not synonymous. The main characters of the intermediate valves of Lucilina sueziensis are reported in Tab. 3. Distribution. Late Pleistocene: Jordan, Aqaba, (Yamanie); Saudi Arabia, Gulf of Aqaba (Ash Shaykh Humayd: this study); Egypt (Hurghada: Issel, 1869; this study). Present-day: Indian Ocean: Red Sea, Gulf of Suez, Gulf of Aden, Somalia, Seychelles, Rodriguez, and Socotra Islands (Dinapoli & Janssen 2009; Kaas et al. 2006; Blatterer 2019)., Published as part of Dell'Angelo, Bruno, Landau, Bernard M., Sosso, Maurizio & Taviani, Marco, 2020, Late Pleistocene Red Sea Mollusca: 1. Polyplacophora, pp. 401-449 in Zootaxa 4772 (3) on pages 415-417, DOI: 10.11646/zootaxa.4772.3.1, http://zenodo.org/record/3819654, {"references":["Reeve, L. (1847) Conchologia iconica, or illustrations of the shells of molluscous animals. Monograph of the genus Chiton. Vol. 4. Reeve, Brothers, London, 64 pp., 28 pls. [pp. 21 - 84]","Issel, A. (1869) Malacologia del Mar Rosso. Ricerche zoologiche e paleontologiche. Parte terza. Catalogo delle conchiglie fossili raccolte sulle spiagge emerse del Mar Rosso. Biblioteca Malacologica, Pisa, 61 pp. [pp. 243 - 303]","Moazzo, P. G. (1939) Mollusques testaces marins du Canal de Suez. Memoires de l'Institut d'Egypte, 38, 1 - 287.","Sykes, E. R. (1907) Reports on the marine biology of the Sudanese Red Sea. 5. On the Polyplacophora or chitons. Journal of the Linnean Society of London, 31, 31 - 34. https: // doi. org / 10.1111 / j. 1096 - 3642.1907. tb 00450. x","Leloup, E. (1960) Amphineures du golfe d'Aqaba et de la Peninsule Sinai. Sea Fisheries Research Station, Haifa, Bulletin, 29, 29 - 55, pl. 1.","Leloup, E. (1980) Chitons de la Mer Rouge, du golfe de Suez et de la Mediterranee. Bulletin Institut Royale des Sciences Naturelles de Belgique, 52, 1 - 14, 2 pls.","Vine, P. (1986) Red Sea Invertebrates. IMMEL Publishing, London, 224 pp.","Lamy, E. (1938) Mission Robert Ph. Dollfus en Egypte. Societe Misr pour les Pecheries S. S. \" Al Sayad \", Resultats Scientifiques. 2 e Partie. VII Mollusca Testacea. Memoires presentees a l'Institut d'Egypte, 37, 1 - 90, 1 pl.","Blatterer, H. (2019) Mollusca of the Dahab region. Oberosterreichisches Landesmuseum, Linz, Austria, 480 pp.","Ferreira, A. J. (1983) Researches on the coast of Somalia. The chiton fauna (Mollusca Polyplacophora). Monitore Zoologico Italiano, 9, 249 - 297. https: // doi. org / 10.1080 / 00269786.1983.11758574","Dekker, H. & Orlin, Z. (2000) Check-list of Red Sea Mollusca. Spirula, 47 (Supplement), 1 - 46.","Slieker, F. J. A. (2000) Chitons of the world. An illustrated synopsis of recent Polyplacophora. L'Informatore Piceno Ed., Ancona, 154 pp.","Abubakr, M. M. (2004) The Republic of Yemen Marine Biotic Ecosystem (Resources-Habitats and Species). Ministry of Water and Environment, Environment Protection Authority, Aden, 128 pp.","Dell'Angelo, B., Bonfitto, A., Sabelli, B. & Taviani, M. (2004) Chitons (Mollusca, Polyplacophora) from bioclastic sands of the Ifaty-Tulear back reefs (Madagascar, Western Indian Ocean). Bollettino Malacologico, Supplment 5, 45 - 62.","Kaas, P. & Van Belle, R. A. (1988) Chitons (Mollusca: Polyplacophora) from the coasts of Oman and the Arabian Gulf. American Malacological Bulletin, 6, 115 - 130.","Van Belle, R. A. & Wranik, W. (1991) Chitons (Mollusca: Polyplacophora) from the Coasts of Yemen and Socotra Island. Fauna of Saudi Arabia, 12, 366 - 381.","Strack, H. L. (1993) The Polyplacophora of the Red Sea. Journal of the Malacological Society of Australia, 14, 1 - 40. https: // doi. org / 10.1080 / 00852988.1993.11923901","Bosch, D. T., Dance, S. P., Moolenbeek, R. G. & Oliver, P. G. (1995) Seashells of Eastern Arabia. Motivate Publishing, Dubai, 290 pp.","Kaas, P. & Van Belle, R. A. (1998) Catalogue of living chitons (Mollusca, Polyplacophora). Second, revised edition. Backuys Publishers, Leiden, 204 pp.","Anseeuw, B. & Terryn, Y. (2004) Intertidal chitons (Mollusca: Polyplacophora) from the coast of Jordan, Red Sea, with the description of a new species of Parachiton Thiele, 1909. Bollettino Malacologico, Supplement 5, 1 - 24.","Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1 - 5. E. J. Brill, Leiden-Boston, 463 pp.","Dinapoli, A. & Janssen, R. (2009) Polyplacophora (Mollusca) of the Socotra Arcipelago - Systematics and Biogeography. Fauna of Arabia, 24, 1 - 46.","Mohammadian, H. (2011) The molluscs in Iran. Shabpareh Publishing Company, Tehran, 229 pp. [in Arabic script]","Rochebrune, A. T. de (1882) Monographie des especes fossiles appartenant a la classe des Polyplaxiphores. Annales des Sciences Geologiques, 14, 1 - 74, pls. 1 - 3.","Kaas, P. (1979) The Chitons (Mollusca: Polyplacophora) of Mozambique. Annals of the Natal Museum, 23, 855 - 879."]}

Lucilina perligera Thiele, 1909 (Fig. 10) Lucilina perligera Thiele 1909, p. 97, pl. 10, figs 51���52; Kilias 1995, p. 167; Dell���Angelo et al. 2010a, p. 15, figs 5A���B; Blatterer 2019, p. 52, pl. 4, fig. 9 a���k. Tonicia perligera; Leloup 1960, p. 39, fig. 4; Leloup 1980, p. 12; Dekker & Orlin 2000, p. 7; Slieker 2000, p. 50, pl. 13, fig. 27. Tonicia (Lucilina) perligera; Strack 1993, p. 20, pl. 4, figs 7���9; Kaas & Van Belle 1998, p. 142; Anseeuw & Terryn 2004, p. 15, figs 24���31, 48���49; Kaas et al. 2006, p. 319, fig. 131, map 18. Non Tonicia perligera; Ferreira 1983, p. 274 (= Lucilina sueziensis, fide Strack 1993, p. 21). Type material. Holotype ZMHU Moll. 102059. Type locality. Red Sea. Material examined. Egypt, Hurghada: St. 13: 1 head valve, width 4 mm (Figs 10 A���C) (MZB 50536); St. 14 bis: 1 head valve, width 3 mm, Fig. 10D (MZB 50537). Recent material examined. Israel, Eilat (P448A): 7 valves (3 head, 2 intermediate and 2 tail), Figs 10 E���I (BD 204; MZB 50538). Description. Head valve semicircular, posterior margin almost straight, front slope slightly convex, tegmentum sculptured with quincuncially dispersed, large, clearly separated, highly elevated, roundish granules, diameter ca 120���160 ��m, ocelli minute, black, randomly dispersed. Articulamentum worn and incomplete. Remarks. The present material most resembles Lucilina perligera Thiele, 1909. A detailed description of that species, characterised by the tegmentum sculptured with large, widely separated granules, was given by Kaas et al. (2006). Two head valves at hand have the same type of sculpture, but with the granules more widely spaced (Figs 10A, D). We report a Recent head valve of Lucilina perligera from the Gulf of Aqaba (Figs 10 E���F), with the denser sculpture of close-set granules more typical for this species. The granules have the same roundish/oval shape and are of comparable dimensions, diameter ca 120���160 ��m in the fossil valve (Fig.10B), ca 140���180 ��m in the living valve (Fig. 10F). The diameter of the granules is variable, Strack (1993: 21) reported a diameter of 130���150 ��m in the holotype, and up to 260 ��m in loose valves studied; Anseeuw & Terryn (2004: 16) gave a diameter up to 280 ��m. The only difference between the head valves of living and fossil specimens seems to be the smaller, wider-spaced granules on the tegmentum in the fossil ones. We also illustrate, for completeness and for comparison with the other species of Lucilina described here, an intermediate and a tail valve of L. perligera from the Gulf of Aqaba (Figs 10 G���I). Lucilina perligera resembles L. sueziensis (see above) in many respects, but can be easily differentiated by the sculpture of the tegmentum, which has large, widely spaced granules. This is the first report of this species as a fossil. Distribution. Late Pleistocene: Egypt, Hurghada (this study). Present-day: northern Red Sea and Maldive Islands (Dell���Angelo et al. 2010a; Kaas et al. 2006; Blatterer 2019)., Published as part of Dell'Angelo, Bruno, Landau, Bernard M., Sosso, Maurizio & Taviani, Marco, 2020, Late Pleistocene Red Sea Mollusca: 1. Polyplacophora, pp. 401-449 in Zootaxa 4772 (3) on pages 417-418, DOI: 10.11646/zootaxa.4772.3.1, http://zenodo.org/record/3819654, {"references":["Thiele, J. (1909) Revision des Systems der Chitonen. Zoologica, 22, 1 - 132, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 11245","Kilias, R. (1995) Polyplacophora-Typen und - Typoide (Mollusca) im Zoologischen Museum in Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 71, 155 - 170. https: // doi. org / 10.1002 / mmnz. 19950710118","Dell'Angelo, B., Gori, S., Baschieri, L. & Bonfitto, A. (2010 a) Chitons (Mollusca, Polyplacophora) from the Maldive islands. Zootaxa, 2673 (1), 1 - 38. https: // doi. org / 10.11646 / zootaxa. 2673.1.1","Blatterer, H. (2019) Mollusca of the Dahab region. Oberosterreichisches Landesmuseum, Linz, Austria, 480 pp.","Leloup, E. (1960) Amphineures du golfe d'Aqaba et de la Peninsule Sinai. Sea Fisheries Research Station, Haifa, Bulletin, 29, 29 - 55, pl. 1.","Leloup, E. (1980) Chitons de la Mer Rouge, du golfe de Suez et de la Mediterranee. Bulletin Institut Royale des Sciences Naturelles de Belgique, 52, 1 - 14, 2 pls.","Dekker, H. & Orlin, Z. (2000) Check-list of Red Sea Mollusca. Spirula, 47 (Supplement), 1 - 46.","Slieker, F. J. A. (2000) Chitons of the world. An illustrated synopsis of recent Polyplacophora. L'Informatore Piceno Ed., Ancona, 154 pp.","Strack, H. L. (1993) The Polyplacophora of the Red Sea. Journal of the Malacological Society of Australia, 14, 1 - 40. https: // doi. org / 10.1080 / 00852988.1993.11923901","Kaas, P. & Van Belle, R. A. (1998) Catalogue of living chitons (Mollusca, Polyplacophora). Second, revised edition. Backuys Publishers, Leiden, 204 pp.","Anseeuw, B. & Terryn, Y. (2004) Intertidal chitons (Mollusca: Polyplacophora) from the coast of Jordan, Red Sea, with the description of a new species of Parachiton Thiele, 1909. Bollettino Malacologico, Supplement 5, 1 - 24.","Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1 - 5. E. J. Brill, Leiden-Boston, 463 pp.","Ferreira, A. J. (1983) Researches on the coast of Somalia. The chiton fauna (Mollusca Polyplacophora). Monitore Zoologico Italiano, 9, 249 - 297. https: // doi. org / 10.1080 / 00269786.1983.11758574"]}

Lucilina tilbrooki Milne, 1958 (Figs 13–15) Lucilina tilbrooki Milne 1958: 152, figs 1–4; Kaas et al. 2006: 346, fig.145, map 51 (bibliography and synonymy); Dell’Angelo et al. 2010: 38, figs 2A–L, 3A–D, 4D–E; Sirenko 2012: 80, pl. 13 C, figs 21, 22. Tonicia (Lucilina) tilbrooki; Kaas & Van Belle 1980: 133; 1998: 188. Type material. Holotype NMV P18471. Type locality. Queensland, Capricorn Group, Heron Island. Material examined. South China Sea, Hainan Island, stn. 5, 19°38’12’’N, 111°01′52’’E, intertidal, on stones, 2 spms, BL 4.0–6.0 mm, 14.11.2016, leg. B. Sirenko; stn. 15, 18°12’09’’N, 109°29′40’’E, intertidal, on old corals, 1 spm, BL 3.0 mm, 19.11.2016, leg. B. Sirenko; stn. 27, 18°18′03’’N, 109°13′55’’E, depth 2–5 m, on stones covered with red crustose algae, 1 spm, BL 9.5 mm, 24.11.2016, leg. B. Sirenko. Distribution. The species is known from Maldives Islands, South Vietnam, Hainan Island and Australia. Remarks. The studied specimen (BL 5.0 mm) has a radula 1.5 mm long with 35 transverse rows of mature teeth and 20 gills extending from valve III to valve VII. The slit formula of this specimen is 8/1/10. This is the first record of L. tilbrooki from Hainan Island., Published as part of Sirenko, Boris I. & Zhang, Junlong, 2019, Chitons (Mollusca: Polyplacophora) of Hainan Island and vicinity, South China Sea, pp. 1-40 in Zootaxa 4564 (1) on page 18, DOI: 10.11646/zootaxa.4564.1.1, http://zenodo.org/record/2588445, {"references":["Milne, K. L. (1958) Descriptions of new species of chitons (Mollusca, Loricata). Royal Society of New South Wales, 152 - 155.","Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Vols 1 - 5. E. J. Brill, Leiden-Boston, 463 pp.","Sirenko, B. I. (2012) Chitons (Mollusca, Polyplacophora) of Nha Trang Bay, South Vietnam. In: Britayev, T. A. & Pavlov, D. S. (Eds.), Benthic fauna of the Bay of Nha Trang, South Vietnam. 2. KMK Scientific Press Ltd, Moscow, pp. 56 - 122.","Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons. W. Backhuys. Rotterdam, 144 pp."]}

Lucilina amanda (Thiele, 1909) Lucilina amanda Thiele 1909: 97, pl. 10, figs 42–50; Bergenhayn 1933: 31; Is. Taki 1938: 404, 410, 412, pl. 15, fig. 9, pl. 33, figs 9–13, pl. 34, figs 5–9; 1962: 47; Huang & Xu 1964: 9; Iw. Taki 1964b: 412; Higo 1973: 8; Ma 2004: 6, pl. 2, fig. G; Xu 2008: 457. Lucilina interplicata Bergenhayn, 1933: 29, fig. 10, pl. 1, fig. 9; Is. Taki 1938: 409, 410, 412; 1962: 47; Iw. Taki 1964b: 412; Higo 1973: 8. Tonicia interplicata; Leloup 1973: 6. Tonicia (Lucilina) amanda; Leloup 1973: 8, 9, 18; Kaas & Van Belle 1980: 7; 1998: 18; Van Belle 1982: 474; Bullock & Harper 1994: 38, pls. 8E, 11D, E, 12 C, D; Kaas et al. 2006: 359, fig. 151, map 36 (bibliography and synonymy). Type material. Holotype (ZMHU). Type locality. Japan, off Tago, 75 m. Distribution. Japan, Hong Kong (Van Belle 1982) and Xisha Islands (Huang & Xu 1964). Remarks. This rare chiton was found sporadically in the studied area in Hong Kong and Xisha Islands., Published as part of Sirenko, Boris I. & Zhang, Junlong, 2019, Chitons (Mollusca: Polyplacophora) of Hainan Island and vicinity, South China Sea, pp. 1-40 in Zootaxa 4564 (1) on page 9, DOI: 10.11646/zootaxa.4564.1.1, http://zenodo.org/record/2588445, {"references":["Thiele, J. (1909) Revision des Systems der Chitonen. Teil. 1. Zoologica, 22, 1 - 132, figs. A-E, pls. 1 - 10.","Bergenhayn, J. R. M. (1933) Die Loricaten von Prof. Dr. Sixten Bocks Expedition nach Japan und den Bonin-Inseln 1914. Kungliga Svenska Vetenskapsakademiens Handlingar, 12, l- 58.","Taki, Is. (1938) Report on the biological survey of Mutsu Bay. 31. Studies on chitons of Mutsu Bay with general discussion on chitons of Japan. Science Reports of Tohoku Imperial Universit y, Series 4, Biology, 12, 323 - 423.","Huang, X. & Xu, F. (1964) Polyplacophora. In: Zhang, X., Qi, Z., Lou, Z., Huang, X., Ma, X., Liu, Y., Xu, F. (Eds.) Pictorial guide to animals of China. Vol. I. Science Press, Beijing, pp. 5 - 10.","Taki, Iw. (1964 b) Classification of the class Polyplacophora, with a list of Japanese chitons. Venus, 22, 401 - 414.","Higo, S. (1973) A catalogue of molluscan fauna of the Japanese Islands and the adjacent area. Class Polyplacophora. Biological Society of Nagasaki Prefecture, Nagasaki, 2 - 8.","Ma, X. (2004) Polyplacophora. In: Qi, Z. (Ed.), Seashells of China. China Ocean Press, Beijing. pp. 3 - 6.","Xu, F. (2008) Class Polyplacophora Blainville, 1867. In: Liu, R. (Ed.), Checklist of marine biota of China seas, pp. 456 - 459.","Leloup, E. (1973) Quelques especes du genre Tonicia Gray, 1847. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 49, 1 - 19.","Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons. W. Backhuys. Rotterdam, 144 pp.","Van Belle, R. A. (1982) Supplementary notes on Hong Kong chitons (Mollusca: Polyplacophora). Proceeding of the first international marine biology Workshop. Hong Kong, 469 - 483.","Bullock, R. C. & Harper, K. D. (1994) The Polyplacophora of Cape d'Aguilar, Hong Kong. In: Morton, B. (Ed.), The Malacofauna of Hong Kong and Southern China III. Hong Kong University Press, Hong Kong, pp. 3 - 54.","Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Vols 1 - 5. E. J. Brill, Leiden-Boston, 463 pp."]}

Lucilina tenuicostata Sirenko, 2016 (Fig. 3C) Lucilina tenuicostata Sirenko 2016b: 490, figs 9, 10, 13D, G. Type material. The holotype (ZISP 2 243). Type locality. Vietnam, Gulf of Tongkin, Cat Ba Islands, 20°47.090′N, 107°06.103′E, 1–1.5 m, SCUBA, on old shells. Material examined. South China Sea, Hainan Island, stn. 13, 18°12’09’’N, 109°29′40’’E, depth 2–3 m, on old corals, 1 spm, BL 9.0 mm, 19.11.2016, leg. B. Sirenko. Distribution. South China Sea, Beibu Gulf (Gulf of Tongkin) and Hainan Island. Remarks. This is the first record of L. tenuicostata from Hainan Island., Published as part of Sirenko, Boris I. & Zhang, Junlong, 2019, Chitons (Mollusca: Polyplacophora) of Hainan Island and vicinity, South China Sea, pp. 1-40 in Zootaxa 4564 (1) on page 18, DOI: 10.11646/zootaxa.4564.1.1, http://zenodo.org/record/2588445, {"references":["Sirenko, B. I. (2016 b) A supplement to the recent Chitonidae (Mollusca: Polyplacophora) of Vietnam. In: Adrianov, A. V. & Lutaenko, K. A. (Eds.), Biodiversity of the western part of the South China Sea. Dalnauka, Vladivostok, pp. 481 - 502."]}

Lucilina mariae (Kaas, Van Belle & Strack, 2006) Tonicia (Lucilina) mariae Kaas, Van Belle & Strack 2006: 336, fig. 140, map 41. Tonicia confossa Van Belle, 1980: 34, pl. 1, fig. 3 (non Gould 1846). Tonicia (Lucilina) sp.; Van Belle 1982: 474, fig. 2. Type material. Holotype (BMNH). Type locality. Hong Kong, Tolo Channel (Channel Rocks), 10 m. Distribution. Only known from the type locality., Published as part of Sirenko, Boris I. & Zhang, Junlong, 2019, Chitons (Mollusca: Polyplacophora) of Hainan Island and vicinity, South China Sea, pp. 1-40 in Zootaxa 4564 (1) on page 18, DOI: 10.11646/zootaxa.4564.1.1, http://zenodo.org/record/2588445, {"references":["Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Vols 1 - 5. E. J. Brill, Leiden-Boston, 463 pp.","Gould, A. A. (1846) On the shells collected by the United States Exploring Expedition, commanded by Charles Wilkes, U. S. N. Proceedings of the Boston Society of Natural History, 2, 141 - 145. https: // doi. org / 10.5962 / bhl. part. 3318","Van Belle, R. A. (1982) Supplementary notes on Hong Kong chitons (Mollusca: Polyplacophora). Proceeding of the first international marine biology Workshop. Hong Kong, 469 - 483."]}

Lucilina lamellosa (Quoy & Gaimard, 1835) (Figs 3A, B, 5–12) Chiton lamellosus Quoy & Gaimard 1835: 386, pl. 74, figs 29–32. Tonicia (Lucilina) lamellosa; Kaas & Van Belle 1980: 71; 1998: 106; Kaas et al. 2006: 350 –352, fig. 147, maps 48, 61 (bibliography and synonymy). Tonicia lamellosa; Saito 2000: 21, pl. 10, fig. 12. Lucilina lamellosa; Schwabe 2006a: 102–104, figs 1C, 6A–F, 7; Schwabe et al. 2008: 22–24, fig. 5B (bibliography and synonymy); Sirenko 2012: 80, pls. 13 F; 16 A; Saito 2017: 735, pl. 7, fig. 9. Type material. Lectotype (MNHN 6058) designated by Strack (1986). Type locality. Tonga, Tongatapu (21°10’S, 175°13’W). Material examined. Lectotype (MNHN 6058); South China Sea, Hainan Island. MBM 100297, 55-610, 1 spm, forma typical, BL 12 mm, 8 spms, forma costata, BL 6.0–19.0 mm, 20.04.1955; 55-640, 3 spms, forma typica, BL 14.0–15.0 mm, 14 spms, forma costata, BL 8.0–22.0 mm, 23.04.1955; Xincun, depth 0.5 m, on old corals 6 spms, forma costata, BL 9.0–15.0 mm, 21.04.1958, leg. O. Scarlato; Xincun, depth 1.0 m, on old corals, 2 spms, forma costata, BL 9.0–10.0 mm, 21.04.1958, leg. O. Scarlato; Xisha Islands, 58M-1246, 4 spms, forma typica, BL 25.0–29.0 mm; 1 spm, forma typica, BL 34.0 mm, 23.05.1981; MBM 100328, 4 spms, forma typica, BL 16.0–28.0 mm, 1 spm, forma costata, BL 29.0 mm; Guangdong Province, 2 spms, forma costata, 10.0–15.0 mm; Submersed reef, R/ V Akademik Shirshov, 18°39’N, 111°40’E, depth 0.5–2.0 m, on old corals, 1 spm, forma typical, BL 15.0 mm, 25.07.1981, Distribution. The species has a wide distribution in the Indo-Pacific: Andaman Sea, Vietnam, South China including Guangdong Province, Hainan Island and Xisha Islands, Australia and Samoa as well. Remarks. While examining L. lamellosa, we found two forms: L. lamellosa forma typica (Figs 3A, 5–6) and the new form L. lamellosa forma costata (Figs 3B, 7–12). Both forms differ from each other in the sculpture of tegmentum of the pleural areas. Lucilina lamellosa forma typica has demonstrated the similarity of it with the lectotype (MNHN 6058) from Tonga. Both forms of L. lamellosa have a distinct diagonal ridge sculptured with concentric shingle-like ledges. In L. lamellosa forma typica, pleural areas are also sculptured with shingle-like ledges (Fig. 5D), whereas in L. lamellosa forma costata, the pleural areas are sculptured with longitudinal ribs (Fig. 7E). The latter form is similar to Lucilina nigropunctata (Carpenter, 1865) in its sculpture of pleural areas. L. lamellosa forma costata differs from L. nigropunctata in having subtriangular pits on the central area and in the absence of radiating rows of small tubercles in lateral areas. Moreover, forma costata has the tail valve wider than the head valve and the longitudinal ribs do not curve towards the jugum as in L. nigropunctata. Lucilina lamellosa forma costata is also similar to L. truncata (Sowerby, 1841) from the Philippines but differs from the latter in having longitudinal ribs on the pleural areas (vs. more or less concentrical, distinctly spaced, wavy grooves in L. truncata), subtriangular pits on the central area (pits are unknown in L. truncata). Both forms of L. lamellosa were found near Fujian Province, Hainan Island and Xisha Islands. The forma costata inhabits the coast of Guangdong Province. The examined specimen of L. lamellosa forma typica (BL 15.0 mm) has a radula of 4.2 mm in length and 27 gills extended from valve II to valve VIII. The slit formula of this specimen is 9/1/13. The radula of the specimen of L. lamellosa forma costata (BL 14.0 mm) is 4.5 mm long and its slit formula is 9/1/15. A smaller specimen of L. lamellosa forma costata (BL 9.0 mm) has a radula 3.5 mm long with 34 transverse rows of mature teeth and 20 gills extended from valve III to valve VII. The slit formula of this specimen is 9/1/14. The latter specimen is similar to L. mariae Kaas et al., 2006 in having longitudinal ribs on the pleural areas of the intermediate valves, but differs from it in having sparsely scattered subtriangular pits on the central area (vs. no pits in L. mariae). Because both forms differ from each other only in the sculpture of the tegmentum of the pleural area and because there are specimens with intermediate sculpture, we refrain from describing a new subspecies or a new species. We assign a registration number of the Zoological Institute RAS (ZISP 2301) to Lucilina lamellosa forma costata., Published as part of Sirenko, Boris I. & Zhang, Junlong, 2019, Chitons (Mollusca: Polyplacophora) of Hainan Island and vicinity, South China Sea, pp. 1-40 in Zootaxa 4564 (1) on pages 10-12, DOI: 10.11646/zootaxa.4564.1.1, http://zenodo.org/record/2588445, {"references":["Quoy, J. R. C. & Gaimard, J. P. (1835) Voyage de decouvertes de l' \" Astrolabe \". Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. Zoologie 3, Mollusques, Genre Oscabrion, Paris, 369 - 411.","Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons. W. Backhuys. Rotterdam, 144 pp.","Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Vols 1 - 5. E. J. Brill, Leiden-Boston, 463 pp.","Schwabe, E. (2006 a) On a small collection of chitons from Papua New Guinea (Mollusca: Polyplacophora). Venus, 65, 97 - 112.","Schwabe, E., Sirenko, B. & Seeto, J. (2008) A checklist of Polyplacophora (Mollusca) from the Fiji islands. Zootaxa, 1777, 1 - 52.","Sirenko, B. I. (2012) Chitons (Mollusca, Polyplacophora) of Nha Trang Bay, South Vietnam. In: Britayev, T. A. & Pavlov, D. S. (Eds.), Benthic fauna of the Bay of Nha Trang, South Vietnam. 2. KMK Scientific Press Ltd, Moscow, pp. 56 - 122.","Strack, H. L. (1986) Notes on New Caledonian chitons. I. The species described by St. - M. Souverbie. Basteria, 50, 193 - 198.","Carpenter P. P. (1865) Descriptions of two species of Chitonidae from the collection of W. Harper Pease, Esq. Proceedings of the Zoological Society of London, 33, 511 - 512. https: // doi. org / 10.1111 / j. 1469 - 7998.1865. tb 02380. x","Sowerby, G. B. (II.) (1841) Descriptions of several new species of chitones, brought by H. Cuming, Esq., from the Philippine Islands. Proceedings of the Zoological Society of London, 9, 61 - 62."]}

Lucilina rainfordiana Hull, 1924 Hull, A.F.B., 1924. New Queensland Loricates. Proceedings of the Royal Society of Queensland 36: 109���116 [115, 116], pl. 21, fig. 3. Paratypes: MA28563, one whole dried individual and one individual with intact valves and girdle but without soft parts, collected by A.F.B. Hull and E.H. Rainford on 17 September, 1923 from the type locality North Head, Port Denison, Queensland, Australia. C urrent name: Lucilina fortilarata (Reeve, 1847) ��� Schwabe (2009a)., Published as part of Wilma M. Blom, 2017, Fossil and Recent molluscan types in the Auckland War Memorial Museum. Part 2: Polyplacophora and Scaphopoda, pp. 71-76 in Records of the Auckland Museum 52 on page 74, DOI: 10.5281/zenodo.1488080, {"references":["Schwabe, E. 2009 a. Lucilina fortilirata (Reeve, 1847). In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 386291 on 22 June 2017."]}

Notes are given on nine chiton species, collected mainly during the 'Oceanic Reefs' Expedition to the Seychelles (1992-1993), with some biogeographical observations. Ischnochiton goudi (family Ischnochitonidae) is described as a new species.

Lucilina perligera Thiele, 1909 (Figures 5A–B) Lucilina perligera Thiele 1909: 97, pl. 10, figs 51–52. Tonicia perligera; Mastaller 1979: 26, 254, 259; Leloup 1980: 12. Tonicia (Lucilina) perligera; Slieker 2000: 50, pl. 13, fig. 27; Anseeuw & Terryn 2004: 15, figs 24–31, 48–49; Kaas et al. 2006: 319, fig. 131 (bibliography and synonymy). Type: Holotype ZMB Moll. 102059 (Kilias 1995: 167). Type locality: Red Sea. Material examined: M13: 1 intermediate valve. Remarks: Detailed descriptions of this species are given by Strack (1993) and Anseeuw & Terryn (2004). The investigated material comprises of an intermediate valve only, but the characteristic sculpture, formed by large, well separated granules, covering the entire tegmentum surface, is sufficient to identify the species. Lucilina perligera seems to be a rather rare species, and only a few living specimens are known (Anseeuw & Terryn 2004). It was considered endemic to the Red Sea, so this new record considerably extends the distribution of this species. Distribution: Lucilina perligera is reported from the northern Red Sea and the Maldive Islands., Published as part of Dell'Angelo, Bruno, Gori, Sandro, Baschieri, Leonardo & Bonfitto, Antonio, 2010, Chitons (Mollusca, Polyplacophora) from the Maldive islands, pp. 1-38 in Zootaxa 2673 (1) on page 15, DOI: 10.11646/zootaxa.2673.1.1, http://zenodo.org/record/5304551, {"references":["Thiele, J. (1909) Revision des Systems der Chitonen. Zoologica, 22, 1 - 132, pls 1 - 10.","Mastaller, M. (1979) Beitrage zur Faunistik und Oekologie der Mollusken und Echinodermen in den Korallenriffen bei Aqaba, Rotes Meer. Dissertation Ruhr-Universitat Bochum, 344 pp.","Leloup, E. (1980) Chitons de la Mer Rouge, du Golfe de Suez et de la Mediterranee. Bulletin Institut Royale des Sciences Naturelles de Belgique, 52 (5), 1 - 14, pls 1 - 2.","Slieker, F. J. A. (2000) Chitons of the world. An illustrated synopsis of recent Polyplacophora. L'Informatore Piceno Ed., Ancona, vi + 154 pp.","Anseeuw, B. & Terryn, Y. (2004) Intertidal chitons (Mollusca: Polyplacophora) from the coast of Jordan, Red Sea, with the description of a new species of Parachiton Thiele, 1909. Bollettino Malacologico, suppl. 5 (2003), 1 - 24.","Kaas, P., Van Belle, R. A. & Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1 - 5. Brill, Leiden- Boston, 463 pp.","Kilias, R. (1995) Polyplacophora-Typen und - Typoide (Mollusca) im Zoologischen Museum in Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 71, 155 - 170.","Strack, H. L. (1993) The Polyplacophora of the Red Sea. Journal of the Malacological Society of Australia, 14, 1 - 40."]}

Lucilina Dall, 1882 Type species: Chiton confossus Gould, 1846 = Chiton lamellosus Quoy & Gaimard, 1835, subsequent designated by Pilsbry (1893). Distribution: Tropical and subtropical waters of the Indo-Pacific, Neogene ��� Recent., Published as part of Schwabe, Enrico, Sirenko, Boris I. & Seeto, Johnson, 2008, A checklist of Polyplacophora (Mollusca) from the Fiji islands, pp. 1-52 in Zootaxa 1777 on page 22, DOI: 10.5281/zenodo.274284, {"references":["Gould, A. A. (1846) Shells collected by the United States Exploring Expedition under the command of Charles Wilkes. Proceedings of the Boston Society of Natural History, 2, 141 - 145.","Quoy, J. R. C. & Gaimard, J. P. (1835) Voyage de decouvertes de l' \" Astrolabe \". Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d´Urville. Zoologie 3, Mollusques, Genre Oscabrion- Chiton, Linne, J. Tastu, Paris, 369 - 411, pls 73 - 75."]}

Lucilina novemrugata (Bergenhayn, 1930) comb. nov. (Figures 5 E, F) Chresonymy in Kaas et al. (2006: 355). Tonicia novemrugata Bergenhayn 1930 a: 33, pl. 2, figs 42���43, 50, pl. 3. figs 52���54, 62 ��� 65. Tonicia (Lucilina) novemrugata; Kaas & Van Belle 1980: 91; 1998: 132; Kaas et al. 2006: 355, fig. 149. Locus typicus: Kiribati (= Gilbert Islands) [0�� 36 ���S 174 �� 23 ���E]. Primary type: Syntype (SMNH 973). Material examined: ZISP: 1 adult spm, Vekai Island [17 �� 34 ���S 178 �� 49 ���W], Callisto St. 93, 0.5 m, leg. Boris I. Sirenko, 0 8 February 1975, wet. Habitat: No information available. Distribution: The species was considered to be endemic to Kiribati (Kaas et al. 2006: 357). Remarks: This is the first record of this species from Fiji. For the new combination see comments under the preceding species., Published as part of Schwabe, Enrico, Sirenko, Boris I. & Seeto, Johnson, 2008, A checklist of Polyplacophora (Mollusca) from the Fiji islands, pp. 1-52 in Zootaxa 1777 on page 26, DOI: 10.5281/zenodo.274284, {"references":["Kaas, P. & Van Belle, R. A. & Strack, H. L. (2006) Monograph of living chitons. (Mollusca: Polyplacophora) 6, Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae; Additions to Volumes 1 - 5, Leiden, The Netherlands: Koninklijke Brill NV. 463 pp.","Bergenhayn, J. R. M. (1930 a) Die Loricaten von Dr. Sixten Bocks Pazifik-Expedition 1917 - 1918, mit spezieller Berucksichtigung der Perinotumbildungen und der Schalenstruktur. Goteborgs Kungliga Vetenskaps - och Vitterhets-Samhalles Handlingar, (5) B, 1, 1 - 52, pls 1 - 3.","Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons (Mollusca: Polyplacophora), Dr. W. Backhuys Publisher, Rotterdam, 144 p."]}

Lucilina nigropunctata (Carpenter, 1865) comb. nov. (Figures 5 C, D) Chresonymy in Kaas et al. (2006: 353). Acanthopleura nigropunctata Carpenter 1865: 511; Kaas & Van Belle 1980: 90; Kaas et al. 2006: 353. Tonicia nigropunctata; Dautzenberg & Bouge 1933: 419; Richard 1985: 414; Kaas et al. 2006: 353. Tonicia seriegranulata Bergenhayn 1930 a: 30, pl. 2, figs 37���41, 44 ��� 49, 51, 51 a (holotype SMNH 975, type locality: Fiji Islands, Viti Levu, Bau, Station 9., small offshore reefs, dead corals); Kaas & Van Belle 1980: 118; 1998: 168; Kaas et al. 2006: 353. Lucilina tupauana Hull MS, Risbec 1946: 180, figs 126���129 (anatomy only) (syntype MNHN 6024, type locality: New Caledonia); Kaas & Van Belle 1980: 135; 1998: 191; Kaas et al. 2006: 353. ? Tonicia sowerbyi; Leloup 1973: 1 (specimen from Noumea). Tonicia seriogranulata [sic]; Leloup 1973: 6. Tonicia (Lucilina) nigropunctata; Kaas & Van Belle 1998: 131; Kaas et al. 2006: 353, fig. 148. Locus typicus: Tahiti [17 �� 38 ���S 149 �� 25 ���W]. Primary type: Lectotype (USNM 19297), designated by Ferreira (1986: 259), not seen. Type (s) without name-bearing function: One paralectotype (SBMNH 35041), two paralectotypes (NHM). Fiji records: Tonicia seriegranulata Bergenhayn 1930 a: 30, pl. 2, figs 37���41, 44 ��� 49, 51, 51 a. Material examined: ZISP: 2 adult spms, Vekai Island [17 �� 34 ���S 178 �� 49 ���W], Callisto St. 93, 0.5 m, leg. Boris I. Sirenko, 0 8 February 1975, wet; ZISP: 1 adult spm, Cicia Island [17 �� 45 ���S 179 �� 18 ���W], Callisto St. 84, 7 m, leg. Boris I. Sirenko, 0 6 February 1975, wet. Habitat: The species lives under dead coral skeletons in shallow water or among living corals. Distribution: The species is known from New Caledonia, Fiji and the Society Islands (Kaas et al. 2006). Remarks: The synonymy of Tonicia seriegranulata with Lucilina nigropunctata was suggested by Leloup (1973) and since then this idea has been accepted by some (e.g., Kaas & Van Belle 1980, 1998; Kaas et al. 2006), due to the identical structures of the valves, the girdle elements and the arrangements of the aesthetes. However, following the suggestions of some authors (e.g., Strack 2003, Schwabe 2004, 2006a) the genus Lucilina is here restored as a separate genus and not considered a subgenus of Tonicia Gray, 1847, differing from the latter mainly in having a multidentate second lateral tooth, whereas it is unicuspidate in Tonicia. Additionally the species of Tonicia are geographically restricted to the South Eastern Pacific region of Latin America with a single species in the Caribbean, while members of the genus Lucilina are widespread throughout the tropical Indo-Pacific. ��� Lucilina nigropunctata (Carpenter, 1865) (Figures 12 A���D) Fiji records: Lucilina sp. B Ladd 1966 partim: 23, pl. 1, fig. 17. Material examined: USNM 648218: 1 tail valve from the Ndalithoni Limestone of Vanua Mbalavu, Station 110 B [17 �� 14 ���S 178 �� 57 ���W] (probably Pliocene); USNM: 1 (of two) tail valve from the Miocene (Tertiary f) Suva Formation on Viti Levu, Station FB- 20. Remarks: Ladd (1966) mentioned a tail valve and at least one intermediate valve from the Ndalithoni Limestone of Vanua Mbalavu [17 �� 14 ���S 178 �� 57 ���W] (probably Pliocene), and two tail valves from the Miocene Suva Formation on Viti Levu. The species description is quite good and clearly separates the species from L. lamellosa, based on the distinct longitudinal grooves in the pleural and antemucronal areas of this species. Reexamination of the available material has shown that the two tail valves (of three), although showing variations in the number and size of the ribs on the antemucronal area, should be identified as L. nigropunctata (Figs 12 A���D). The taxonomic status of the single intermediate valve (Fig. 12 E) remains unclear. With some doubt we leave it as Lucilina (see below), although the size (19.2 mm in width) and the smooth surface are also characteristic of a species of Onithochiton. As we did not study the specimens of Onithochiton quercinus of Cernohorsky (1978: 42, see above), we can not say with certainty if a larger species besides the below-mentioned Onithochiton societatis Thiele, 1909 occurs in Fijian waters. Nevertheless it does not belong to the same species as the remaining valves Ladd mentioned., Published as part of Schwabe, Enrico, Sirenko, Boris I. & Seeto, Johnson, 2008, A checklist of Polyplacophora (Mollusca) from the Fiji islands, pp. 1-52 in Zootaxa 1777 on pages 24-26, DOI: 10.5281/zenodo.274284, {"references":["Kaas, P. & Van Belle, R. A. & Strack, H. L. (2006) Monograph of living chitons. (Mollusca: Polyplacophora) 6, Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae; Additions to Volumes 1 - 5, Leiden, The Netherlands: Koninklijke Brill NV. 463 pp.","Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons (Mollusca: Polyplacophora), Dr. W. Backhuys Publisher, Rotterdam, 144 p.","Dautzenberg, Ph. & Bouge, J. - L. (1933) Les mollusques testaces marins des etablissements francais de l'Oceanie. Journal de Conchyliologie, Paris, 77, 351 - 469.","Richard, G. (1985) Fauna and Flora. A first compendium of French Polynesian Sea-Dwellers. Amphineura, 413 - 414 In: Delesalle, B., Galzin, R. & Salvat, B. (eds) 5 th International Coral Reef Congress, Tahiti, 27 May - 1 June 1985. Volume 1. \" French Polynesian Coral Reefs \", 379 - 520.","Bergenhayn, J. R. M. (1930 a) Die Loricaten von Dr. Sixten Bocks Pazifik-Expedition 1917 - 1918, mit spezieller Berucksichtigung der Perinotumbildungen und der Schalenstruktur. Goteborgs Kungliga Vetenskaps - och Vitterhets-Samhalles Handlingar, (5) B, 1, 1 - 52, pls 1 - 3.","Risbec, J. (1946) Etudes anatomiques sur les amphineures de la Nouvelle-Caledonie. Journal de la Societe des Oceanistes, 2, 129 - 190.","Leloup, E. (1973) Quelques especes du genre Tonicia Gray, 1847. Bulletin Institut Royal des Sciences Naturelles de Belgique, 49 (6), 1 - 19.","Kaas, P. & Van Belle, R. A. (1998) Catalogue of living chitons (Mollusca, Polyplacophora). Second, revised edition, W. Backhyus Publishers, Leiden, 204 pp.","Ferreira, A. J. (1986) A revision of the genus Acanthopleura Guilding, 1829 (Mollusca: Polyplacophora). The Veliger, 28, 221 - 279.","Strack, H. L. (2003) Class Polyplacophora (formerly Amphineura), pp. 12 - 15. In: Hylleberg, J. & Kilburn, R. N. (eds) Tropical Marine Mollusc Programme. Marine Molluscs of Vietnam. Annotations, Voucher Material, and Species in need of Verification. Phuket Marine Biological Center Special Publication, 28, 1 - 300.","Schwabe, E. (2004) The Polyplacophora (Mollusca) collected during the First International Marine Biodiversity Workshop for Rodrigues (western Indian Ocean), with the description of a new species. Journal of Natural History, 38, 3143 - 3173.","Schwabe, E. (2006 a) On a small collection of chitons from Papua New Guinea (Mollusca: Polyplacophora). Venus, 65, 97 - 112.","Ladd, H. S. (1966) Chitons and Gastropods (Haliotidae through Adeorbidae) from the Western Pacific Islands. United States Geological Survey Professional Papers, 531, 1 - 98, 16 pls.","Cernohorsky, W. O. (1978) Report on the molluscan fauna of the Lau Group, Fiji Islands. Bulletin of the Royal Society of New Zealand, 17, 39 - 52."]}

Lucilina lamellosa (Quoy & Gaimard, 1835) (Figure 5 B) Chresonymy in Kaas et al. (2006: 350). Chiton lamellosus Quoy & Gaimard 1835: 386, pl. 74, figs 29���32; Clessin 1903: 101, pl. 36, fig. 7; Iredale & Hull 1927: 132; Boss & Bieler 1991: 58; Gowlett-Holmes 2001: 43; Schwabe 2004: 3153; Dinapoli 2004: 58; Kaas et al. 2006: 314. Chiton confossus Gould 1846: 143 (holotype at USNM 30763, type locality: Fiji); Iredale & Hull 1926 b: 267; 1927: 130; A. G. Smith 1960: 166; Kaas & Van Belle 1980: 30; 1998: 50; Higo et al. 1999: 28; Slieker 2000: 141; Gowlett- Holmes 2001: 43; Schwabe 2004: 3153; Dinapoli 2004: 58; Kaas et al. 2006: 314. Chiton pictus Reeve 1847 (non de Blainville, 1825, nor Bean in Thorpe, 1844): pl. 15, sp. & f. 79 (the whereabout of the holotype is unknown, it is not at NHM [pers. comm. Roberto Portela Miguez], type locality: ���Torres Strait, Raines Island ��� it is herein restricted to Australia: Queensland: Great Barrier Reef: Raine Island [11.36���S 144 ��02���E]); Kobelt 1894: 26; Iredale & Hull 1926 b: 268; Kaas & Van Belle 1980: 100; 1998: 144; Boss & Bieler 1991: 58; Slieker 2000: 150; Gowlett-Holmes 2001: 43; Kaas et al. 2006: 350. Chiton insculptus Souverbie in Souverbie & Montrouzier 1866 (non A. Adams, 1852): 248, pl. 9, fig. 5 (holotype at MHNB, type locality: New Caledonia: ��le Art [19 �� 43 ���S 163 �� 39 ���E]; not seen); Leloup 1952: 61; Kaas & Van Belle 1980: 64; 1998: 96; Slieker 2000: 146; Gowlett-Holmes 2001: 43; Kaas et al. 2006: 350. Chiton montrouzieri Souverbie 1873: 287 (replacement name for Chiton insculptus Souverbie in Souverbie & Montrouzier, 1866; non C. insculptus A. Adams, 1852); Kaas & Van Belle 1980: 87; 1998: 127; Slieker 2000: 148; Gowlett- Holmes 2001: 43; Kaas et al. 2006: 350. Lucia confossa; Dall 1879: 298, pl. 3, fig. 29. Tonicia lamellosus (- a); Pilsbry 1893: 209, pl. 40, figs 8���11; Nierstrasz 1905 a: 97; Bergenhayn 1930 a: 33; Leloup 1952: 62; Richard 1985: 414; Gosliner et al. 1996: 124, fig. & spec. 421; Schwabe 1998: 94; Slieker 2000: 22, 96, 146, pl. 36, figs 24 ��� 24 a; Saito 2001: 21, pl. 10, fig. 12; Wells 2002: 113; Okusu et al. 2003: 284; Okusu 2003: 77; Schwabe 2006 a: 111. Tonicia confossa; Pilsbry 1893: 210, pl. 57, figs 33���35; Melvill & Standen 1899: 180; Nierstrasz 1905 a: 90, 109, 112, pl. 2, fig. 31, pl. 6, figs 165 A���D; 1905 b: 151; Horst & Schepman 1908: 525; Hedley 1910: 352; Iredale 1910: 158; Bergenhayn 1930 a: 33, 36, pl. 3, figs 66���69; Dautzenberg & Bouge 1933: 419; Ang 1967: 392, 407, pl. 8, figs 1���4; Richard 1985: 414; Strack 2003: 14; Kaas et al. 2006: 350. Tonicia picta; Pilsbry 1893: 211, pl. 40, figs 5, 6; Nierstrasz 1905 a: 96; Hedley 1910: 352; Iredale 1910: 158; Ashby 1918: 86; 1920: 291; Leloup, 1952: 61. Toniciopsis picta; Thiele 1893: 371, pl. 30, fig. 27; Cotton 1964: 88. Tonicia insculpta; Pilsbry 1894: 89, pl. 10, figs 10���11; Nierstrasz 1905 a: 97. Chiton (Tonicia) pictus; Clessin 1903: 26, pl. 10, fig. 8. Chiton (Tonicia) confossa; Couturier 1907: 175. Lucilina lamellosa; Thiele 1909 b: 97; Eernisse 1984: 259; Strack 2003: 14; Schwabe 2006 a: 102, figs 1 C, 6 A���F, 7; Vinther 2006: 17, figs 11, 13, pl. 14; Saito 2006 c: 122, fig. 6 B; Burghardt et al. 2006: 30. Lucilina shirleyi Iredale 1914 c (replacement name for Chiton pictus Reeve, 1847; non C. pictus de Blainville, 1825): 131; Iredale & Hull 1926 b: 268, pl. 38, figs 12, 14; 1927: 131, pl. 16, figs 12, 14; Mackay 1930: 293; Leloup 1952: 61; Joyce 1959: 239, fig. 5 b; Kaas & Van Belle 1980: 119; 1998: 170; Gowlett-Holmes, 2001: 43; Kaas et al. 2006: 350. Tonicia (Lucilina) confossa; Ashby 1918: 86; Thiele 1931: 22. Tonicia picata [sic]; Ashby 1924: 332. Tonicia (Lucilina) shirleyi; Ashby 1926: 384; 1928: 172. Lucilina insculpta; Risbec 1946: 176, fig. 34. Lucilina confossus (- a); Cotton 1964: 88, 101; Ladd 1966: 23; Saito 2006 c: 122. ? Onithochiton quercinus (Gould, 1846); Cernohorsky 1978: 42; Coleman 2003: 67, un-numbered text figure. Tonicia (Lucilina) lamellosa; Kaas & Van Belle 1980: 71; 1998: 106; Van Belle 1983: 132, pl. 12, fig. 6; Schwabe 1997: 23, text figs; 2005 a: 54, pl. 2, fig. 8; 2006 b: 20; Gowlett-Holmes 1998 b: 187; 2001: 43; Schwabe & Slieker 2001: 225; Strack 2003: 14; Kaas et al. 2006: 350, fig. 147. cf. Chiton; Birkeland et al. 1987: 45. ? Tonicia (Lucilina) sp.; Sirenko 2006: 40, fig. 11.17 Locus typicus: ��� Tonga Tabou��� (= Tonga, Tongatapu [21 �� 10 ���S 175 �� 13 ���W]). Primary type: Lectotype (MNHN 6058) designated by Strack (1986: 194). Fiji records: Chiton confossus Gould 1846: 143. Tonicia confossa; Bergenhayn 1930 a: 36, pl. 3, figs 66���69. ? Onithochiton quercinus; Cernohorsky 1978: 42. Material examined: ZSM Mol 20061636: 1 juvenile spm, Viti Levu, Laucala Bay, west side of the mole of the USP, 18 ��09.054'S 178 �� 27.231 'E, up coming tide, under stones, 0���0.5 m, leg. Andreas Altn��der, 20 August 2006, wet; ZSM Mol 20061615: 5 adult spms, Viti Levu, Laucala Bay, north side of Nukumbutho Island 18 �� 10.474 'S 178 �� 28.343 'E; lowest low tide, landward in calm water close to the beach, 0.5 m, under stones and dead corals, which were mainly covered with algae, leg. Enrico Schwabe, 21 August 2006, wet; ZSM Mol 20061618: 1 adult spm, 3 subadult and juvenile spms, Viti Levu, Laucala Bay, west side of the Nukumbutho channel of the fringing reef opposite the USP, 18 �� 10.899 'S 178 �� 27.971 'E, attached to pieces of dead corals, embedded in fine coral sand among dead coral boulders, strong currents, about 1 m, leg. Enrico Schwabe, 24 August 2006, wet; ZSM Mol 20061631: 6 adult spms, Viti Levu, Laucala Bay, north side of Makuluva Island 18 �� 11.250 'S 178 �� 30.992 'E, under pieces of dead corals, partly embedded in fine coral sand, 1���3 m, leg. Enrico Schwabe, Andreas Altn��der & Michael Schr��dl, 26 August 2006, wet; ZSM Mol 20061918: 1 adult spm, Viti Levu, Laucala Bay, reef flat opposite the Suva Harbour at the fringing reef, 18 �� 10.434 'S 178 �� 24.987 'E, at low tide, high amount of living corals, some sandy channels and patches, 1 m, under coral plates, leg. Andreas Altn��der, 29 August 2006, wet; ZSM Mol 20061625: 1 subadult spm + 2 juvenile spms, Viti Levu, Laucala Bay, west side of the Nukumbutho channel of the fringing reef opposite the USP, 18 �� 10.899 'S 178 �� 27.971 'E, out side of the fringing reef, in deep channels with strong currents, in pieces of dead corals encrusted with coralline red algae, about 4 m, leg. Enrico Schwabe, 29 August 2006, wet; ZSM Mol 20061634: 1 adult spm, Viti Levu, Laucala Bay, North of Makuluva Island, 18 �� 10.890 'S 178 �� 30.587 'E, sandy bottom with sea grass patches, under dead coral pieces, 0.5��� 1 m, leg. Enrico Schwabe, 30 August 2006, wet; ZISP 431-1966: 1 adult spm, Viti Levu, Suva [18 ��08���S 178 �� 25 ���E], coral reefs, leg. S. Stepanjants, 0 4 March 1966, ���Kanopus���, wet; ZISP: 1 juvenile spm, Cicia Island [17 �� 45 ���S 179 �� 18 ���W], Callisto St. 83, 0.3 m, leg. Boris I. Sirenko, 0 6 February 1975, wet; UF 295858: 1 adult spm, Viti Levu, 4.5 km WSW of Mbukatatanoa Reefs 18 �� 10 ��� 44 ������S 178 �� 27 ��� 27 ������E, reef flat, reef crest, under rocks, leg. Lisa Kirkendale & Victor Bonito, 11 October 2001 (LAK-015B), wet; UF UF 289421: 1 adult spm, Viti Levu, 1 km W of Mbusa Bay [19 ��04���S 178 ��03���E], 0���1.5 m, fringing reef, undersides of rocks, in crevices, leg. Holthuis, B. & Gustav Paulay, 0 4 February 1989 (BFIJI- 434), wet; UF 292960: 1 adult spm, Viti Levu, Island owned by Fijian Resort 18 ��08��� 30 ������S 177 �� 25 ��� 24 ������E, 0���0.5 m, reef flat, under sand, leg. Lisa Kirkendale & Victor Bonito, 20 October 2001 (LAK- 028F), wet; ZSM Mol 20070109: 21 larvae to juveniles, Viti Levu, Laucala Bay, west side of the Nukumbutho channel of the fringing reef opposite the USP, 18 �� 10.899 'S 178 �� 27.971 'E, among algae from the reef pools, 0.5���1.5 m, leg. Michael Schr��dl, Enrico Schwabe & Andreas Altn��der, 24 August 2006, wet. Habitat: Juveniles of the species were regularly found within crevices of dead coral pieces and under algae, adults were found under pieces of dead corals and other rubble, which was usually slightly embedded in fine coral sand or at least big enough so not to be overturned by currents. Distribution: The species has a wide Indo-Pacific distribution from Andaman Sea in the West and North to Australia in the South and American Samoa in the East (Schwabe 2006 a). Remarks: Although Schwabe & Lozouet (2006) speculated that Cernohorsky���s (1978: 42) record of Onithochiton quercinus (Gould, 1846) probably refers instead to Onithochiton lyelli (Sowerby in Broderip & Sowerby, 1832) it is also possible that the specimen is L. lamellosa, by far the most common species in Fiji. A photo request from the Auckland museum, where the specimen should be deposited was so far ineffective., Published as part of Schwabe, Enrico, Sirenko, Boris I. & Seeto, Johnson, 2008, A checklist of Polyplacophora (Mollusca) from the Fiji islands, pp. 1-52 in Zootaxa 1777 on pages 22-24, DOI: 10.5281/zenodo.274284, {"references":["Quoy, J. R. C. & Gaimard, J. P. (1835) Voyage de decouvertes de l' \" Astrolabe \". Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d´Urville. Zoologie 3, Mollusques, Genre Oscabrion- Chiton, Linne, J. Tastu, Paris, 369 - 411, pls 73 - 75.","Kaas, P. & Van Belle, R. A. & Strack, H. L. (2006) Monograph of living chitons. (Mollusca: Polyplacophora) 6, Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae; Additions to Volumes 1 - 5, Leiden, The Netherlands: Koninklijke Brill NV. 463 pp.","Clessin, S. (1903 - 1904) Die Familie Chitonidae. Systematisches Conchylien Cabinet, ed. 2, Nurnberg, 6, 1 - 120 (1903), 121 - 136 (1904), pls 1 - 41.","Iredale, T. & Hull, A. F. B. (1927) A Monograph of the Australian Loricates, Royal Zoological Society of New South Wales, Sydney: i - xiv, 1 - 168, pls 1 - 21.","Boss, K. J. & Bieler, R. (1991) Johannes Thiele and his contributions to Zoology. Part 2. Genus-Group Names (Mollusca). Nemouria, 39, 1 - 77.","Gowlett-Holmes, K. L. (2001) Polyplacophora. pp. 19 - 84 In: Wells, A. & Houston, W. W. K. (eds). Zoological Catalogue of Australia. Vol. 17.2. Mollusca: Aplacophora, Polyplacophora, Scaphopoda, Cephalopoda. CSIRO Publishing, Melbourne, Australia, i - xii, 353 pp.","Schwabe, E. (2004) The Polyplacophora (Mollusca) collected during the First International Marine Biodiversity Workshop for Rodrigues (western Indian Ocean), with the description of a new species. Journal of Natural History, 38, 3143 - 3173.","Dinapoli, A. (2004) Polyplacophora der Insel Sokotra. Systematik und Biogeographie. Diplomarbeit, vorgelegt der Fakultat fur Biowissenschaften der Ruprecht-Karls-Universitat Heidelberg, 137 pp.","Gould, A. A. (1846) Shells collected by the United States Exploring Expedition under the command of Charles Wilkes. Proceedings of the Boston Society of Natural History, 2, 141 - 145.","Iredale, T. & Hull, A. F. B. (1926 b) A monograph of the Australian loricates (Phylum Mollusca-Order Loricata). VII. The Australian Zoologist, 4, 256 - 276, pl. 37.","Smith, A. G. (1960) Amphineura, pp. 141 - 176. In: Moore, R. C. (ed.) Treatise on invertebrate paleontology. I, Mollusca 1, University of Kansas Press, 374 pp.","Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons (Mollusca: Polyplacophora), Dr. W. Backhuys Publisher, Rotterdam, 144 p.","Higo, S., Callomon, P. & Goto, Y. (1999) Catalogue and bibliography of the marine shell-bearing Mollusca of Japan, Elle Scientific Publications, 749 pp.","Slieker, F. J. A. (2000) Chitons of the world. An illustrated synopsis of recent Polyplacophora, L'Informatore Piceno, i - vi, 154 pp.","Reeve, L. A. (1847 - 1848) Conchologia Iconica, or Illustrations of the Shells of Molluscous Animals. 4, Monograph of the genus Chiton, Monograph of the genus Chitonellus, London, pls 1 - 27 (1847), pl. 28 (1848).","Blainville, H. M. D. de (1825) Oscabrelle. Oscabrion. In: Levrault, E. (ed.) 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(1879) Report on the limpets and chitons of the Alaskan and Arctic regions, with descriptions of genera and subspecies believed to be new. Proceedings of the United States National Museum, 1, 281 - 344, pls 1 - 5.","Nierstrasz, H. F. (1905 a) Die Chitonen der Siboga-Expedition. Siboga Expeditie, 48, 1 - 112, pls 1 - 8.","Bergenhayn, J. R. M. (1930 a) Die Loricaten von Dr. Sixten Bocks Pazifik-Expedition 1917 - 1918, mit spezieller Berucksichtigung der Perinotumbildungen und der Schalenstruktur. Goteborgs Kungliga Vetenskaps - och Vitterhets-Samhalles Handlingar, (5) B, 1, 1 - 52, pls 1 - 3.","Richard, G. (1985) Fauna and Flora. A first compendium of French Polynesian Sea-Dwellers. Amphineura, 413 - 414 In: Delesalle, B., Galzin, R. & Salvat, B. (eds) 5 th International Coral Reef Congress, Tahiti, 27 May - 1 June 1985. Volume 1. \" French Polynesian Coral Reefs \", 379 - 520.","Gosliner, T. M., Behrens, D. W. & Williams, G. C. (1996) Coral reef animals of the Indo-Pacific. 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Notes are given on nine chiton species, collected mainly during the 'Oceanic Reefs' Expedition to the Seychelles (1992-1993), with some biogeographical observations. Ischnochiton goudi (family Ischnochitonidae) is described as a new species.