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New taxonomic, biological and faunistic data are presented for some species of sawflies occurring in southern Europe and North Africa. Pamphilius mediterraneus sp. nov. is described from Italy (Sicily). It belongs to the alternans sub-group of the alternans species group. Pristiphora omalos sp. nov. is described from Greece (Crete). It belongs to the depressa species group. New junior synonyms are Calameuta tazzekae Lacourt, 1991 of Calameuta gaullei (Konow, 1896), and Emphytus leucostomus Costa, 1890 of Allantus cingulatus (Scopoli, 1763). Periclista rufiventris Zombori, 1979 is moved from the subgenus Periclista to the subgenus Neocharactus. Ametastegia (Protemphytus) persica Khayrandish, Talebi & Blank, 2015 is recorded for the first time in Europe, from Italy (Sicily), and a key to West Palaearctic Protemphytus species provided. Allantus enslini (Forsius, 1918) is more widespread in southern Europe than previously recorded. An identification key distinguishes it from its closer West Palaearctic relatives. New hostplant records are for Halidamia affinis (Fallén, 1807) (Rubia peregrina), Janus compressus (Fabricius, 1793) (Sorbus torminalis), and Macrophya albicincta (Schrank, 1776) (Centranthus ruber). New distribution records are given for a few other species.

Heptamelus viitasaarii Liston, Mutanen & Prous sp. nov. Description. Female. Figs 23, 24A-C, 26A, B Colour. Fig. 26A, B. Black. Pale are: palpi, tegula, legs (except for arioli and extreme inner tip of metatibia); in fore wing base and apex of costa, apex of subcostal, base of stigma (Fig. 23A), R1, 1A, 2A+3A [color differences between veins of hind wing not clearly definable]; margins of median excision of abdominal tergum 1, medio-distal part of terga 2-4 (-6) [pale area on tergum 3 is the largest of these], all sterna, more or less the downturned parts of terga 2-8, more or less hypopygium and ovipositor valvifer 2. Wing membranes hyaline. Head. Pedicel about as long as scape. Head in lateral view with widest point on temple about equal to length of pedicel (Fig. 23E). Malar space slightly shorter than diameter of anterior ocellus. Lowest part of gena densely sculptured, without shining interspaces between the ill-defined pits (Fig. 23E). Clypeus densely pitted, weakly shiny; anterior margin widely emarginate, to about 0.4 of its length (Fig. 23D). Setae on upper head as long, or longer than, diameter of anterior ocellus (Fig. 23E). Thorax. Whole pronotum except for small antero-ventral area dull, with sculpture (Fig. 23H). Pits on upper mesepisternum large and well-defined (Fig. 23H). Mesoscutellum with large, scattered pits anteriorly and laterally, interspaces shiny; medially and posteriorly nearly without pits and entirely unsculptured (Fig. 23F). Anterior of mesoscutellar appendage densely pitted, with numerous setae, only small posterior area unpitted (Fig. 23F). Abdomen. Anterior of tergum 1 densely setose. Terga 2 and 3 glabrous. Terga increasingly setose from tergum 4 to apex of abdomen. Weak surface sculpture on tergum 2 becoming stronger on more distal terga. Sawsheath in dorsal view broad, with blunt tip (Fig. 24C). In lateral view exposed length of cercus about 0.5 × as long as exposed upper length of valvula 3 (Fig. 23G). Lancet (Fig. 24B, C): 13-14 serrulae; apical serrulae short and high. Body length: 4.5-8.0 mm Male. Figs 24D, 26C Similar to female, except for: red-brown antennal flagellum with basal two antennomeres more or less black; metatibiae completely pale; abdomen completely dark with small, obscure pale markings medially on terga 3-5. Penis valve: Fig. 24D. Body length: 4.5-5.5 mm Variability: 5-6 flagellomeres, depending on whether or not the distal one is subdivided; the proportions of this antennomere are thus highly variable. Number of pits on upper mesepisternum variable. The scape and pedicel of females may be more or less pale. Type material. Holotype ♀ (ZMUO.044130). Type locality: Finland, Karelia borealis, Kitee Otravaara, YKJ grid coordinates: 6868:3663 [61.8877N 30.1096E]; 04.08.2019 [larva] in Athyrium filix-femina (L.), Rearing 380/2019, leg. M. Mutanen. Labels white, printed, except for red holotype label. Deposited in the ZMUO. Paratypes. Total: 92♀, 6♂. Finland [leg. M. Mutanen and in ZMUO unless otherwise stated]: Karelia borealis, Kitee: Otravaara [Type locality]; 17♀ (including ZMUO.044126, ZMUO.044127, ZMUO.044128, ZMUO.044129, ZMUO.044131, ZMUO.044132, ZMUO.044133, ZMUO.044134, ZMUO.044135, ZMUO.044136), 04.08.2019 [larvae] in Athyrium filix-femina (L.), Rearing 380/2019. 7♀ (including ZMUO.034957, ZMUO.034958, ZMUO.034959), Otravaara, YKJ grid coordinates: 6868:3663 [61.888°N, 30.110°E], 19.06.2018. 30♀ (including ZMUO.058509, ZMUO.058510, ZMUO.058511, ZMUO.058512, ZMUO.058513, ZMUO.058514, ZMUO.058515, ZMUO.058516, ZMUO.058517, ZMUO.058518, ZMUO.058519, ZMUO.058520), Otravaara, YKJ grid coordinates: 6868:3663 [61.888°N, 30.110°E], 24.08.2020, leg. M. Mutanen and M. Prous. Host: Athyrium filix-femina, rearing 174/2020. 1♀ (ZMUO.046037), 15.06.2020. ZMUO and SDEI. Otravaara, YKJ grid coordinates: 6868:3664 [61.887°N, 30.129°E]; 8♀ (including ZMUO.034859, ZMUO.034860), 19.06.2018. 1♀ (ZMUO.034995), 18.06.2018. Pajarinmäki, YKJ grid coordinates: 68899:36670 [62.078°N, 30.197°E]; 1♀ (ZMUO.032885), at light, 11.07.2017 [end date of about 5-day period]. 1♀ (ZMUO.033391), at light, 18.07.2017 [end date of about 7-day period]. 2♀ (including ZMUO.034899), 20.06.2018. 1♂ (ZMUO.061917), 14.06.2021, leg. A. Liston, M. Mutanen, M. Prous. 1♂ (ZMUO.061893), 15.06.2021, leg. N. Kiljunen, A. Liston, M. Mutanen, M. Prous. 4♂ 8 ♀♀ (including ZMUO.060946, ZMUO.060947, ZMUO.060948, ZMUO.060949, ZMUO.060950, ZMUO.060951, ZMUO.060952, ZMUO.060953, ZMUO.060954, ZMUO.060955, ZMUO.058165), 18.06.2021, leg. N. Kiljunen, A. Liston, M. Mutanen, M. Prous. 2 larvae from Matteuccia struthiopteris (L.) (ZMUO.060557, ZMUO.060558) and 2 larvae from Athyrium filix-femina (L.) (ZMUO.060561, ZMUO.060562), 26.07.2021, leg. M. Mutanen, M. Prous. Papinniemi, YKJ grid coordinates: 6883:3656 [62.025°N, 29.990°E]; 6♀ (including ZMUO.034761, ZMUO.034762), 18.06.2018, ZMUO and SDEI. 1♀ (ZMUO.040497), 12.06.2019. Puhos, YKJ grid coordinates: 6889:3653 [62.080°N, 29.938°E]; 2♀ (including ZMUO.034968), 20.06.2018. Hiidensaari, YKJ grid coordinates: 6891:3668 [62.092°N, 30.226°E]; 1♀ (ZMUO.034908), 20.06.2018. Potoskavaara, YKJ grid coordinates: 6893:3670 [62.109°N, 30.266°E]; 1♀ (ZMUO.046430), 28.06.2020, leg. Tupu Vuorinen. Satulavaara, YKJ grid coordinates: 6877:3672 [61.964°N, 30.289°E]; 3♀ (including ZMUO.061792, SDEI-GISHym14061), 14.06.2021, leg. A. Liston, M. Mutanen and M. Prous. ZMUO and SDEI. Karelia australis: Imatra Kaikkallio, YKJ grid coordinates: 6778:3594 [61.104°N, 28.752°E]; 1♀ (ZMUO.058146), 16.06.2021, leg. A. Liston, M. Mutanen, N. Kiljunen, M. Prous. Joutseno Kuurmanpohja Sunninmäki [61.071°N, 28.731°E]; 1♀ (ZMUO.016815), 06.06.2015, leg. Jussi Vilen, coll. Matti Viitasaari. Tavastia australis: Kangasala Keisarinharju, YKJ grid coordinates: 6818:3347 [61.444°N, 24.141°E]; 1♀ (ZMUO.061964), 08.06.2021. Other material. Finland: Nylandia, Vantaa, YKJ grid coordinates: 669:37, 17.06.2021, 1♀, leg. Miikka Friman. Etymology. Named after Matti Viitasaari, who first recognised a very large female of this species as probably not belonging to Heptamelus ochroleucus or H. dahlbomi. Habitat. Damp, shady places in woodland. Biology. Many female specimens have been reared from larvae in Athyrium filix-femina (L.) Roth. (Fig. 25). Larvae found abundantly in 2021 in Matteucia struthiopteris (L.) Tod. in Kitee, Finland, were provisionally identified as H. viitasaarii by genetic comparison (identical to adults), as later confirmed when females started to emerge in 2022. The larvae feed inside the rachis, eating alternate portions nearly completely, or only partly (Fig. 25B, C). This results internally in a “ladder-like” appearance, which is externally clearly visible, especially against the light (Fig. 25A). This feeding habit is, however, possibly widespread in Heptamelus: at least H. dahlbomi larvae feed in the same way. Larvae of H. viitasaarii in M. struthiopteris regularly overwinter in a chamber made within the fertile fronds at the base of the stem. Strangely, no males have so far been reared, and the few males so far netted are all from a single locality (within an area of less than 1km2). It was observed that adults reared from plants of different size varied very much in body size, and that the smallest individuals had been feeding on the smallest plant, with correspondingly thin stalks. Distribution. Finland (North and South Karelia, Tavastia, and Helsinki area), Russia (Moscow). The record from Moscow was published by Vikberg (2017) as Heptamelus ochroleucus. Differential diagnosis. The characters which are most useful for the identification of European Heptamelus species are presented below in a key. The extensively pitted and setose mesoscutellar appendage distinguishes H. viitasaarii from the two other European species, as well as H. magnocularis Malaise, 1931 (Malaise 1931b) from the Russian Far East (see Liston et al. 2018). Note that all body parts of H. viitasaarii are conspicuously more densely setose, and pitted, than most other Heptamelus species which we have so far examined. From the females of Heptamelus dahlbomi, H. ochroleucus, H. magnocularis, H. montanus Togashi, 1961, and H. takeuchii Togashi, 1961 (the latter two from Japan), H. viitasaarii differs in its much shorter and higher apical serrulae of the lancet (Fig. 24A, B). Heptamelus viitasaarii has a wider sawsheath in dorsal view than either H. dahlbomi or H. ochroleucus. Heptamelus japonicus Togashi, 1961, only known in the male sex, differs from H. viitasaarii in its entirely pale thorax, abdomen and fore wing pterostigma. The male of H. takeuchii differs from H. viitasaarii in its more shallowly emarginate and smoother clypeus, and the pale abdominal sternum 9. The male of H. montanus resembles that of H. viitasaarii in coloration, but as in females of these species, the clypeus of H. montanus is much less deeply emarginate (approx. to about 0.2 of its length) and the setae on the upper head shorter (mostly shorter than the diameter of the anterior ocellus)., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Vikberg, V, 2017. Two sawflies new for the fauna of Russia, one being new for Europe and the western Palaearctic region (Hymenoptera: Symphyta: Heptamelidae and Tenthredinidae). Sahlbergia 23 (1): 17 - 19","Malaise, R, 1931b. Entomologische Ergebnisse der schwedischen Kamtchatka Expedition 1920-1922. (35. Tenthredinidae). Arkiv foer Zoologi 23(2 [A8]): 1-68. [Separatum]","Liston, AD, Vardal, H, Prous, M, 2018. New and poorly-known sawflies (Hymenoptera, Tenthredinoidea) from Sweden, with taxonomic notes on Palaearctic Heptamelus species described by Swedish authors. Entomologisk Tidskrift 139 (2): 119 - 131"]}

Phymatoceropsis sibiricola (Zhelochovtsev, 1939) comb. nov. Figs 28, 29 Notes. Rhadinoceraea sibiricola Zhelochovtsev, 1939: 159-160. ♀. Holotype. Type locality: Russia, Krasnoyarsk Krai, Vostotschnoe (35 km SE of Minussinsk). In the original description, Zhelochovtsev (1939) compared R. sibiricola with " Rh. japonica Malaise" [Phymatoceropsis japonica (Malaise, 1931) (Malaise 1931a)], and stated that they are "closely allied". Comparison of female P. sibiricola specimens from Finland (Fig. 28) with a Japanese female of P. japonica in the SDEI collection confirmed their similarity. As mentioned by Zhelochovtsev, P. japonica has much longer antennae, but the differences described in the structure of the frontal area and mesopleura seemed less clear. The host plant of P. sibiricola, as discovered by Vikberg in Finland (see below), is Sambucus racemosa (Adoxaceae), since verified several times by Mutanen. The host plant of P. japonica is Sambucus sieboldiana Blume (Okutani 1956), which is sometimes treated as a synonym or subspecies of S. racemosa L. Summary of records in Finland [not all more recent records are listed]. Uusimaa: Mäntsälä, Saari, 67368:34139 [60.732°N, 25.423°E], 1♀ 18.05.2010, leg. Iiro Kakko (in his collection in Hämeenlinna). South Häme: Janakkala, Laurinmäki, 67526:3695 [60.862°N, 24.599°E], 1♀ 01.06.2009, 1♀ 26.05.2010, 1♀ 31.05.2010, leg. V.Vikberg (VVT). Janakkala, Hangastenmäki, 67525:3692 [60.861°N, 24.594°E], 1♀ 21.05.2011, 2♀ 22.05.2011, 2♀ 24.05.2011, 1♀ 31.05.2011, 1♀ 16.05.2012, 1♂ 21.05.2013, 1♀ 29.05.2013; in 2014-2021 females were collected every year and a few males altogether, leg. V.Vikberg (VVT). One male and one female collected in 2013 were photographed by Pekka Malinen http://id.luomus.fi/GL.7655 (♂), http://id.luomus.fi/GL.7656 (♀). North Häme: Jyväskylä. Vikberg identified two females in photographs taken in two different years by Raimo Pelkonen. The last year was 2019, but the first year much earlier. South Karelia: Imatra, Räikkölä, 61.121°N, 28.790°E, 1♀ 30.05.2021, 1♀ 01.06.2021, leg. M. Mutanen & M. Prous. Karelia borealis: Kitee, Pajarinmäki, 62.075°N, 30.186°E, 1♀ 22.05.2016, leg. M. Mutanen; 1♀ 06.06.2016, leg. M. Mutanen. Pajarinmäki, 62.078°N, 30.191°E, 1♀ 14.05.2018, leg. M. Mutanen; 1♀ 15.05.2018, leg. M. Mutanen; 1♀ 4♂ 20.06.2018, leg. M. Mutanen; 24♀ 1♂ 17.06.2020, leg. M. and I. Mutanen; 1♀ 18.5.2021, leg. M. Mutanen & M. Prous. Papinniemi, 62.025°N, 29.990°E, 1♀ 16.05.2018, leg. M. Mutanen. Puuteniemi, 62.167°N, 29.970°E, 1♀ 1♂ 15.05.2018, leg. M. Mutanen. Potoskavaara, 62.109°N, 30.266°E, 1♀ 15.05.2018, leg. T. Vuorinen. Puhos, 62.089°N, 29.958°E, 2♀ 18.06.2018, leg. M. Mutanen. Puhos, 62.089°N, 29.939°E, 1♀ 05.06.2019, leg. M. Mutanen. Tyynelä, 62.080°N, 30.321°E, 1♀ 13.05.2019, leg. M. Mutanen. Vanhahovi, 62.169°N, 29.974°E, 2♀ 15.05.2019, leg. M. Mutanen. Tavastia australis: Hämeenlinna, Hattelmalanharju, 60.971°N, 24.475°E, 1♀ 15.05.2013, leg. M. Raekunnas (ZMUO). Biology. Oviposition trial by Vikberg (3/2011 VV): three females captured in Hangastenmäki in 2011 were used in this experiment. Adoxa, Sambucus racemosa, Lonicera xylosteum and Galium sp. were offered to them. They laid eggs only on Sambucus racemosa leaves. The egg is laid through the upperside of the leaf into the tissue of the underside, usually one to two eggs per leaflet, but in one small leaflet eight eggs were counted. Oviposition occurred on 21.05.-24.05. The first larvae were observed on the morning of 28.05. Later, 28 larvae were counted. Five feeding instars were observed and after finishing feeding there was an "extra moult". Prepupae were seen on 07.06-08.06. 1st and 2nd instar larvae are gray, with a brown head. 5th instar larvae are 13-14 mm long; head width ca. 1.6 mm (Fig. 29). Head brownish, behind the eye a black fleck which is curved backwards and upwards. Body dorsally dark gray with greenish hue; ventrally whitish gray. Black suprastigmal flecks on thoracic segment 3 and abdominal segments 1-9.

Calameuta variabilis (Mocsáry, 1886) comb. nov. Fig. 5 Cephus quadriguttatus Costa, 1882: 198. ♂. Syntypes. Type locality: Sardinia. Synonymy with C. haemorrhoidalis auct. by Ghigi (1905). Primary homonym of Cephus quadriguttatus Westwood, 1874. syn. nov. Cephus variabilis Mocsáry, 1886a: 101, 103. ♀, ♂. Syntypes. Type locality: Kecskemétnél, Herkulesfürdõ, Nagyváradnál, Szilágymegyében Tasnádon, Tasnád-Szántó, and S.-A.- Ujhelynél (Mocsáry 1886b). Lectotype, ♀, designated below. Cephus quadriguttulatus Dalla Torre, 1894: 412. Replacement name for C. Cephus quadriguttatus Costa. syn. nov. Cephus quadrisignatus Costa, 1894: 252. Replacement name for C. Cephus quadriguttatus Costa. syn. nov. Cephus haemorrhoidalis var. signifer Konow, 1896b: 317-318. ♀. Holotype. Type locality: Syria, Akbes. syn. nov. Cephus pseudotabidus Kokujev, 1910: 136-137. ♀, ♂. Syntypes. Type locality: Transcaspia. Synonymy with C. haemorrhoidalis auct. by Gussakovskij (1935). syn. nov. Cephus diversipes Ghigi, 1915: 308-309. ♂. Holotype. Type locality: Greece, Rhodos. Synonymised with C. haemorrhoidalis auct. by Gussakovskij (1935). syn. nov. Trachelus syriacus Pic, 1917: 1-2. ♀. Syntypes. Type locality: Syria, Alexandrette. Synonymised with C. haemorrhoidalis auct. by Gussakovskij (1935). syn. nov. Calameuta haemorrhoidalis: Gussakovskij (1935), Benson (1946), and most subsequent authors. Type material examined and taxonomic notes. Lectotype of Cephus variabilis Mocsáry, 1886, hereby designated: ♀, id nr.017651 HNHM Hym.coll. (Fig. 5A); labels (Fig. 5A). Type locality: Romania, Herkulesfürdõ (HNHM). Paralectotypes (all HNHM): 5♂ and 2♀ also belong to the type series and have been labelled as paralectotypes: details of their sexes and localities were given by Zombori (1978). Note that the type series is heterogeneous, and contains specimens of Calameuta punctata as well as C. variabilis. Zombori noted the heterogeneity of the type series, but did not publish a lectotype designation, although the specimens were labelled by him as lectotype and paralectotypes. The types were found by Z. Vas in the HNHM, grouped as stated by Zombori. We do not follow Zombori’s intention according to his labels, but designate the female from Herculesfürdö (= Baile Herculane, Romania) (id nr. 017651 HNHM Hym. coll., labelled by Z. Vas) as lectotype (Fig. 5A). Thus, Calameuta variabilis (Mocsáry, 1886) comb. nov. can be used as a valid name for Calameuta haemorrhoidalis sensu Gussakovskij et auct. Cephus atripes Stephens, 1835 has sometimes been listed as a synonym of C. variabilis, e,g. by Taeger et al. (2010, under Calameuta haemorrhoidalis). The type specimen, or specimens, is probably lost. The description is short, and does not state the sex of the described specimens(s). de Dalla Torre (1894) treated C. atripes as a synonym of C. pygmeus (Linnaeus, 1767) and Konow (1905a) as a synonym of C. haemorrhoidalis. The color pattern described by Stephens does not fit very well with either of these, in either sex, nor with any other known north-west European cephid species. Stephens’ name is best treated as a species inquirendae. Biology and distribution. According to Macek et al. (2020, under Calameuta haemorrhoidalis) the host plants of C. variabilis are various Poaceae, including cereals such as rye (Secale cereale) and wheat (Triticum). According to our personal experiences, adults occur in dry places, mostly on or near wild grasses. The taxon to which the name C. variabilis is now applied has so far been found only in the West Palaearctic, and has an essentially Mediterranean distribution, summarized by Gussakovskij (1935) as comprising southern Europe, Crimea, Caucasus, western Turkmenia (Kopet-Dagh) and Syria. In Central Europe, it reaches at least as far north as Hungary (Zombori 1978), but specimens recorded from Austria and the Czech Republic (see Taeger et al. 2006) should be checked., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Ghigi, A, 1905. Catalogo dei Tenthredinidi del Museo zoologico di Napoli con osservazioni critiche e sinonimiche. Annuario del Museo Zoologico della R. Universita di Napoli, N. S. 1[1904](21): 1-28.","Mocsary, S, 1886b. A magyarorszagi fa-ronto darazsok. V. Rovartani Lapok 3: 113 - 120","Gussakovskij, VV, 1935. Insectes Hymenopteres, Chalastogastra 1. In: Stackelberg, AA, Ed., Faune de l'URSS. Academie des Sciences de l'URSS, Moscou, Leningrad 2(1): 1 - 453","Benson, RB, 1946. Classification of the Cephidae (Hymenoptera Symphyta). Transactions of the Royal Entomological Society of London 96 (6): 89 - 108, DOI: https://doi.org/10.1111/j.1365-2311.1946.tb00445.x","Zombori, L, 1978. New Sawfly Species in the Hungarian Fauna (Hymenoptera, Symphyta), IV. Annales Historico-Naturales Musei Nationalis Hungarici 70: 259 - 264","Taeger, A, Blank, SM, Liston, AD, 2010. World Catalog of Symphyta (Hymenoptera). Zootaxa 2580 (1): 1 - 1064, DOI: https://doi.org/10.11646/zootaxa.2580.1.1","de Dalla Torre, CG, 1894. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 1: Tenthredinidae incl. Uroceridae (Phyllophaga & Xylophaga). Guilelmus Engelmann, Lipsiae, [[6] pp.+ pp. I-VIII +] 1-459. https://doi.org/10.5962/bhl.title.10348","Konow, FW, 1905a. Hymenoptera. Fam. Lydidae. Genera Insectorum 27: 1 - 27","Macek, J, Roller, L, Benes, K, Holy, K, Holusa, J, 2020. Blanokridli Ceske a Slovenske republiky II. Siropasi. Academia, Praha","Taeger, A, Blank, SM, Liston, AD, 2006. European Sawflies (Hymenoptera: Symphyta) - A Species Checklist for the Countries. In: Blank, SM, Schmidt, S, Taeger, A, Eds., Recent Sawfly Research: Synthesis and Prospects. Goecke and Evers, Keltern: 399 - 504"]}

Dolerus timidus (Klug, 1818) sp. rev. Figs 13C, D, 14A, 14D, 14E Tenthredo (Dolerus) timida Klug, 1818: 300. ♀, ♂. Syntypes. Published type locality: “Deutschland” [Germany]. Lectotype designated below. Tenthredo (Dolerus) dubia Klug, 1818: 299-300. ♀, ♂. Syntypes. Type locality: Germany. Lectotype designated below. Primary homonym of Tenthredo dubia Ström, 1768 [= Tenthredo (Tenthredella) livida Linnaeus, 1758]. Dolerus dubius var. atratus Enslin, 1913: 159. ♂. Syntypes (assumed). Type locality: Switzerland. Lectotype designated below. Type material examined and taxonomic notes. Tenthredo (Dolerus) timida Klug, 1818. Lectotype ♂ hereby designated, labelled: “14199” [printed catalog no.], "Var. T. timida Kl. [Klug’s handwriting]", "GBIF-GISHym 2317", "Zool. Mus. Berlin" [printed], " Lectotypus [printed] ♂ Tenthredo (Dolerus) timida Klug, 1818 [handwritten in block letters] M.Heidemaa design. [printed]" [red], " Dolerus timidus (Klug, 1818) M.Heidemaa det." [printed]. Good condition, penis valve dissected, pinned with the specimen (Fig. 14A). Paralectotypes: 4♂ 5♀ (GBIF-GISHym: 2312, 2318, 2320-2326). All in ZMHB. Tenthredo (Dolerus) dubia Klug, 1818. Lectotype ♂ hereby designated, labelled: “14198” [catalog no.], " Dolerus pratensis (L.) ♂ [handwritten] det. F. Koch [printed]", "GBIF-GISHym 2315" [printed], "Zool. Mus. Berlin" [printed], " Lectotypus [printed] ♂ Tenthredo (Dolerus) dubia Klug, 1818 [handwritten in block letters] M.Heidemaa design. [printed]" [red], " Dolerus timidus (Klug, 1818) M.Heidemaa det." [printed], Condition satisfactory, genitalia dissected, penis valves on a slide pinned with the specimen (they fit D. timidus). Paralectotypes: 1♂, 1♀ ("GBIF-GISHym 2313", “… 2314"). All in ZMHB. Dolerus dubius var. atratus Enslin, 1913. Lectotype ♂ hereby designated, labelled: “Helvet.[ia]”, “Type” [pink, printed], " Dolerus ♂ Dolerus dubius Dolerus dubius var atratus [handwritten] Dr. Enslin det. [printed]", "Sammlung Dr. Enslin" [printed], "GBIF-GISHym 3102", "LECTOTYPE ♂ Dolerus dubius var. atratus Enslin, 1913 designated M. Heidemaa 2022" [red, printed]. Some damage: left flagellum missing, abdomen glued to a card together with the dissected penis valves and pinned with the specimen. ZSM. This melanic color form was identified as D. timidus based on its penis valve structure and the sculpture of the metepimeron., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080

Dolerus junci (Stephens, 1835) nom. rev. Dosytheus Junci [sic!] Stephens, 1835: 84-85. Sex not explicitly stated. Syntypes. Published type localty: "Devonshire, and near Windsor" (England). Lectotype designated below. Dolerus Busaei [sic!] Snellen van Vollenhoven, 1858: 277. Sex not given. Lectotype ♂ designated by Thomas (1987). Type locality: “Haarlem” (Netherlands). syn. nov. Dolerus cothurnatus auct. nec Serville, 1823 (e.g. Benson 1952, Zhelochovtsev 1994, Lacourt 2020). Dolerus thargitai Zombori, 1994: 185-187. ♀. Holotype. Type locality: Csíkszentkirály, Borsáros, Kérújfürdő: Tolvajos patak (Romania). syn. nov. Remarks. Similarly to the melanic color form of D. yukonensis (see below), which was described as a distinct species, D. scoticus, by Cameron (1881a), the melanic form of D. junci was also described as a distinct species, D. thargitai, by Zombori (regarded as a synonym of D. yukonensis / D. scoticus until now). The melanic color form of D. junci has also been recorded from central Europe (Germany) and southern Europe (Italy, Switzerland: see, e.g. Pesarini 2012), but is not known from Fennoscandia. The males of the melanic forms of D. junci (01.06.1984) and D. yukonensis (24.05.1984) have been collected at the same locality at nearly the same time (Germany: Federsee, Brackenhof, leg. E. Jansen, CEJ). In addition to their different penis valve structure and other characters used in the keys by Benson (1952) and Zhelochovtsev (1994), D. junci and D. yukonensis differ in the shape and sculpture of the mesoscutellar appendage: almost unsculptured and without a distinct longitudinal keel in the former, and clearly sculptured and with a distinct keel in the latter. Most likely, the melanic color form of D. junci does not occur within the geographic range of D. incisus, but they can also be distinguished by the shape and the sculpture of the mesoscutellar appendage: meshed (nearly matt) and long, with a distinct median longitudinal keel (D. incisus) versus short and almost unsculptured (more or less glossy) and without a distinct median keel (D. junci). Type material examined and taxonomic notes. Dosytheus junci Stephens, 1835. Lectotype ♀ hereby designated, labelled: "Type H.T. [holotype]" [round with wide red margin, printed; not a holotype but a syntype!], "B.M. TYPE HYM. [printed] 1.575.", "B.M. TYPE HYM. [printed] Dosytheus Junci Stephens 1835.", " Junci.", "Stephens Coll. 53-46" [printed], " Dolerus palustris, Klug." [printed], " Lectotypus 2012 Dosytheus junci [printed]♀ [symbol handwritten] Stephens, 1835 M.Heidemaa des. [printed]", " Dolerus junci Stephens, 1835 M.Heidemaa det." [printed]. Condition: legs partly destroyed, left midleg missing, the apical flagellomeres missing, apex of left forewing broken. BMNH. The original description of D. junci by Stephens (1835) refers to syntypes ("Devonshire and near Windsor"). Although Kirby (1882) mentioned a "Type of D. junci ", this cannot be regarded as a lectotype designation, even though he was apparently only able to locate a single specimen. Dolerus busaei Snellen van Vollenhoven, 1858. Photos of the lectotype and of the penis valve were checked. Left midleg and two segments of the right antenna missing; the genital capsule is glued on a card. RMNH. This nominal taxon was previously treated as a synonym of D. cothurnatus auct. Dolerus cothurnatus auct. nec Serville, 1823. Haris and later Lacourt (2000) studied the syntypes (3 ♂) of D. cothurnatus Serville, but not their genitalia. Based on their penis valve structure, none of the syntypes fits D. cothurnatus auct.: the two syntypes (among them the one labelled as lectotype) represent D. germanicus and the third one is a male of D. aericeps. The name D. cothurnatus by Serville is placed here as a junior synonym of D. germanicus, by designating its lectotype (see also under D. germanicus), and the name Dolerus junci (Stephens, 1835) has to be used for D. cothurnatus auct. nec Serville. Dolerus thargitai Zombori, 1994. The holotype and some paratypes (2♀, 4♂) were studied. HNHM.

Abia nitens (Linnaeus, 1758) Tenthredo nitens Linnaeus, 1758: 556. ♂ [because the conspicuous dark dorsal patches on abdomen are mentioned]. Syntypes (assumed). Type locality: Europe. Lectotype designated by Malaise and Benson (1934). LSUK. Images of the lectotype (LINN 2402) in dorsal and lateral view are available (The Linnean Society of London 2022). Tenthredo sericea Linnaeus, 1767: 921. Sex not stated [but probably female, because dark dorsal patches on abdomen are not mentioned]. Syntypes (assumed). Type locality: Leipzig. syn. nov. Abia sericea: de Dalla Torre (1894), Konow (1905b), Enslin (1917), Liston and Späth (2006), Taeger et al. (2010). Abia dorsalis Costa, 1859: 5-6. ♀. Holotype [not examined]. Type locality: Italy, Cape Miseno near Naples. syn. nov. Notes. Abia nitens was successively mentioned as occurring in Sweden by Fallén (1807), Dahlbom (1836), and Thomson (1871), who all placed Tenthredo sericea as its synonym. Thomson’s description of leg color indicates that he had before him specimens of the species called by Taeger (1998) A. sericea, or perhaps A. candens [not distinguished from the former in Thomson’s time]. Influenced by the continued, widespread use of the name Abia nitens in Scandinavian literature, Taeger et al. (2006) and ArtDatabanken (2015) included Sweden within the range of A. brevicornis [as A. nitens auct.]. In fact, no Swedish specimens of A. brevicornis have been located in the MZLU or NHRS collections, and there are no published records from other Fennoscandian countries. Abia brevicornis is restricted to extremely dry, summer-warm sites, where its larval hosts occur (Scabiosa spp.) (Liston and Späth 2006). Probably it has a strongly continental distribution: its most north-westerly known localities in Europe are in Central Germany, whereas it is not definitely known in France (Noblecourt 2020), and a single old record from Spain needs confirmation. Although a presence of A. brevicornis in southern Sweden cannot be ruled out, we consider it likely that all references to A. nitens auct. in Sweden relate to either A. nitens [= Abia sericea] or A. candens., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Malaise, R, Benson, RB, 1934. The Linnean Types of Sawflies (Hymenoptera, Symphyta). Arkiv foer Zoologi 26(4 [A20]): 1-14. https://doi.org/10.1080/00222933408654888","2022. . http://linnean-online.org","de Dalla Torre, CG, 1894. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 1: Tenthredinidae incl. Uroceridae (Phyllophaga & Xylophaga). Guilelmus Engelmann, Lipsiae, [[6] pp.+ pp. I-VIII +] 1-459. https://doi.org/10.5962/bhl.title.10348","Konow, FW, 1905b. Hymenoptera. Fam. Tenthredinidae. Genera Insectorum 29: 1 - 176","Enslin, E, 1917. Die Tenthredinoidea Mitteleuropas VI. Deutsche Entomologische Zeitschrift (Beiheft 6): 539-662.","Liston, AD, Spaeth, J, 2006. On the Biology of Abia nitens (Linne, 1758): a Thermophile Sawfly with a Diurnal Larval Feeding-Pattern (Hymenoptera: Symphyta: Cimbicidae). In: Blank, SM, Schmidt, S, Taeger, A, Eds., Recent Sawfly Research: Synthesis and Prospects. Goecke and Evers, Keltern: 129 - 138","Taeger, A, Blank, SM, Liston, AD, 2010. World Catalog of Symphyta (Hymenoptera). Zootaxa 2580 (1): 1 - 1064, DOI: https://doi.org/10.11646/zootaxa.2580.1.1","Fallen, CF, 1807. Foersok till uppstaellning och beskrifning a de i Sverige fundne Arter af Insect-Slaegtet Tenthredo Linn. Kongliga Vetenskaps Academiens nya Handlingar 28 (3): 179 - 209","Dahlbom, G, 1836. Prodomus Hymenopterologiae Scandinavicae. C. F. Berling, Lund, DOI: https://doi.org/10.5962/bhl.title.67771","Thomson, CG, 1871. Hymenoptera Scandinaviae (Tenthredo et Sirex Lin.). H. Olsson, Lund 1: 1 - 342","Taeger, A, 1998. Bestimmungsschluessel der Keulhornblattwespen Deutschlands (Hymenoptera: Cimbicidae). In: Taeger, A, Blank, SM, Eds., Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme. Goecke and Evers, Keltern: 193 - 205","Taeger, A, Blank, SM, Liston, AD, 2006. European Sawflies (Hymenoptera: Symphyta) - A Species Checklist for the Countries. In: Blank, SM, Schmidt, S, Taeger, A, Eds., Recent Sawfly Research: Synthesis and Prospects. Goecke and Evers, Keltern: 399 - 504","ArtDatabanken (2015) Roedlistade arter i Sverige 2015. [Introduction pp 1-17; Hymenoptera pp 92-97]. ArtDatabanken SLU, Uppsala, 1-209.","Noblecourt, T, 2020. Liste systematique des Hymenopteres Symphytes de France (2020-2). Office National des Forets, Laboratoire National d'Entomologie Forestiere, Quillan"]}

Phymatoceropsis gracilicornis (Zaddach, 1859) comb. nov. Notes. Selandria gracilicornis Zaddach, 1859: 34-35. ♀. Holotype. Type locality [see Introduction, p. 7]: probably Königsberg [= Russia, Kaliningrad]. Blennocampa gracilicornis: André (1881). Monophadnus gracilicornis: Kirby (1882). Rhadinoceraea gracilicornis: Konow (1886a). Dicrostema gracilicornis: Benson (1952). Paracharactus gracilicornis: Smith (1969). The host plant of P. gracilicornis is Adoxa moschatellina L. (Adoxaceae) (Chambers 1947).

Dolerus anthracinus (Klug, 1818) Fig. 10A Notes. Tenthredo (Dolerus) anthracina Klug, 1818: 302. ♀, ♂. Syntypes. Type locality: Germany, Gartz in Pommern. Lectotype ♂ hereby designated, labelled: “14205” [collection catalog no., printed], " Anthracina Kl." [Klug’s handwriting], “Kl.” [Klug’s handwriting], "GBIF-GISHym 2308" [printed], "Zool. Mus. Berlin" [printed], " Lectotypus ♀ Tenthredo (Dolerus) anthracina Klug, 1818 M.Heidemaa des. '22 [handwritten in block letters]" [red], " Dolerus anthracinus (Klug, 1818) M.Heidemaa det. [printed] not nitens Zadd." [handwritten in block letters]. ZMHB., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080

Pristiphora coactula (Ruthe, 1859) Nematus coactulus Ruthe, 1859: 307-308. ♀. Holotype. Type locality: Iceland. Lygaeonematus (Lygaeotus) trochantericus Lindqvist, 1952: 101-102. ♀. Holotype (http://id.luomus.fi/GL.7708) and paratypes (♀, ♂). Type locality: Finland, Utsjoki, Outakoski. syn. nov. Notes. The nuclear sequence data obtained for this study revealed three main clusters within the Pristiphora carinata group: P. carinata, P. coactula, and P. borea + P. groenblomi + P. albilabris (Fig. 30). No nuclear sequence data are yet available for P. breadalbanensis (Cameron, 1882b) and P. lativentris (Thomson, 1871). A nearly perfect match morphologically to the L. trochantericus holotype is ZMUO.035514, which falls within the P. coactula cluster based on nuclear DNA (Fig. 30). There are two main clusters based on COI sequences, one of which contains only P. borea (Konow, 1904) and P. groenblomi (Lindqvist, 1952) and the other one all species (Fig. 31). Within the COI cluster containing all species (Fig. 31), P. borea, P. groenblomi, and P. albilabris (Boheman, 1852) (Betula feeders) tend to separate from P. coactula (Salix) and P. carinata (Hartig, 1837) (Vaccinium). Based on the specimens having nuclear data, the species (mainly females) of the Pristiphora carinata group may be separated by the following key, although it might not always work for all specimens, particularly P. coactula and P. borea. Excluded from the key are the (sub)arctic species P. breadalbanensis and P. lativentris. Pristiphora Pristiphora lativentris may have somewhat different serrulae from the other species (almost papilliform, see fig. 215 in Prous et al. 2017). The identity of Pristiphora breadalbanensis (most similar to P. borea and P. coactula) needs further research to confirm if characters (e.g. structure of median mesoscutal lobes) mentioned by Benson (1958) to separate this species are reliable. [table omitted] Examples of lancets of P. borea, P. carinata, and P. coactula are shown in Fig. 32A-C, but more specimens need to be examined to check if there are any consistent differences between the species. Morphological differences between the males of P. borea, P. carinata, and P. coactula are not clear. Externally, it seems that P. coactula tends to be paler (clypeus, pronotal angles, tegula, metafemur, and sternum 9 completely or mostly pale) than P. borea and P. carinata. The dorso-apical margin of the paravalva of P. borea (Fig. 34B) may be more strongly inclined basally compared to P. carinata and P. coactula, but differences between the penis valves of the latter two species are not clear (Fig. 34A, D, E). The most distinctive penis valve in the P. carinata group seems to belong to P. albilabris (Fig. 34F), which has the most distinctly inclined dorso-apical margin of paravalva. Overall shape of penis valve of P. groenblomi (Fig. 34C) is most similar to P. borea, but it may be larger., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Prous, M, Kramp, K, Vikberg, V, Liston, A, 2017. North-Western Palaearctic species of Pristiphora (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research 59: 1 - 190, DOI: https://doi.org/10.3897/jhr.59.12656.figure3","Benson, RB, 1958. Hymenoptera, Symphyta. Handbooks for the Identification of British Insects 6(2c): 139-258."]}

Dolerus sanguinicollis (Klug, 1818) Figs 11E, F, 12B-D, F, H Note. Tenthredo (Dolerus) sanguinicollis Klug, 1818: 305. ♀ [not explicitly stated, but indicated by colour characters]. Syntypes. Published type locality: Austria. Lectotype ♀ hereby designated, labelled: “14222” [printed], "Austr. Kl." [Type locality: Austria, Carinthia, Klagenfurt], " Tenthredo sanguinicollis Kl.", " Lectotypus " [printed in block letters] ♀ 2014 [handwritten] Tenthredo (Dolerus) sanguinicollis Klug, 1818 [handwritten], "Des. M.Heidemaa" [red label], "GBIF-GISHym 2343", "Zool. Mus. Berlin" [printed], " Dolerus sangunicollis (Klug, 1818) M.Heidemaa det.". Minor damage. 2♀ (same collecting data) labelled as paralectotypes (GBIF-GISHym: 2344, 2345). All in ZMHB., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080

Dolerus possilensis Cameron, 1882 Notes. Dolerus possilensis Cameron, 1882a: 178-179. ♀. Syntypes (assumed). Type locality: Possil Marsh near Glasgow, Scotland. Lectotype ♀ hereby designated, labelled: "B.M. TYPE HYM. 1.573" [printed], " Dolerus possilensis Cam.", "Cameron 96-76. [printed] Possil" [handwritten], "B.M. TYPE HYM. [printed] Dolerus possilensis (Cameron 1882)", " Lectotypus [printed] Dolerus possilensis Cameron, 1882 [handwritten in block letters] M.Heidemaa design. [printed]" [red]. Most of the abdomen missing. Ovipositor preparation apparently lost, but probably figured in Cameron (1882a: plate XIX, fig. 8). BMNH. Probably a parthenogenetic species: no male has ever been recorded. The only known Fennoscandian record is from Sweden: 1♀ [labelled as Dolerus sp.] “Resarö Waxholm / 6 1915", leg. R. Malaise, NHRS. New to the Swedish fauna., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Cameron, P, 1882a. A Monograph of the British Phytophagous Hymenoptera. (Tenthredo, Sirex and Cynips, Linne.). Volume 1. Ray Society, London 340 pp. [21 plates]"]}

Dolerus gonager (Fabricius, 1781) Tenthredo gonagra Fabricius, 1781: 412. Sex not given. Syntypes (assumed). Type locality: "Habitat in Germaniae plantis." [Germany]. Lectotype designated below. Dolerus femoratus Eversmann, 1847: 24-25. ♀. Syntypes. Type locality: Kasan (Russia, Taterstan). Lectotype designated below. Dolerus magnicornis Eversmann, 1847: 25. ♂. Syntypes (assumed). Type locality: Russia, Orenburg Province. Lectotype designated below. Type material examined and taxonomic notes. Tenthredo gonagra Fabricius, 1781. Lectotype ♀ hereby designated, labelled: " Tenthredo gonagra "; "LECTOTYPE Tenthredo gonagra Fabricius, 1781 des. SMBlank 2008" [red, printed: paralectotypes labelled in corresponding way], " ZMUC-GISHym 1050", " Dolerus gonager (F.) [handwritten] det. S.M.Blank 2008 ✓" [printed], " Dolerus gonager (Fabricius, 1781) M.Heidemaa det." [printed]. Right flagellomeres 6-7, right hind tarsus missing. Paralectotype 1♀: " Tenthredo gonagra ", " ZMUC-GISHym 1051". ZMUC. The type specimens correspond with Zhelochovtsev’s definition of Dolerus (Poodolerus) gonager, which has red knees, superficially pitted lateral mesonotal lobes and backwards directed setae of the parallel-sided ovipositor sheath (similar to Zhelochovtsev 1994: fig. 165.2, but with some of the longest hairs curved distally). Dolerus femoratus Eversmann, 1847. Lectotype ♀ hereby designated, labelled: "Kas[an] 16.V.", " Dolerus femoratus. Kl." [with black frames], "DEI-GISHym 30068" [white, printed], "Lectotype Dolerus femoratus Eversmann 1847 vide A. Taeger 2014 des." [red, printed]. In good condition. Belongs to D. gonager. Paralectotypes: 2♀ (DEI-GISHym 30069, 30070), are D. puncticollis Thomson, 1871. All in ZIN. Dolerus magnicornis Eversmann, 1847. Lectotype ♂ hereby designated, labelled: “Orb” [Orenburg], " Dolerus magnicornis mihi" [with black frames], "DEI-GISHym 30067" [white, printed], "Syntype Dolerus magnicornis Eversmann 1847 vide A. Taeger 2014" [red, printed], Lectotype Dolerus magnicornis Eversmann 1847 M.Heidemaa des." [red, printed]. In good condition. Belongs to D. gonager. ZIN., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Zhelochovtsev, AN, 1994. [Translation of the book published in Russian in 1988]. In: Medvedev GS (Ed.) Keys to the Insects of the European Part of the USSR. Volume III Hymenoptera Part VI Symphyta. E.J. Brill, Leiden"]}

Dolerus varispinus Hartig, 1837 Fig. 16G, J, M, P, S Dolerus varispinus Hartig, 1837: 239. ♀, ♂. Syntypes. Lectotype ♂ designated by Blank and Taeger (1992). Type locality: Germany [implicit from title of Hartig’s work]. Dolerus rugosus Konow, 1884: 281. ♀, ♂. Syntypes. Type locality: Fürstenberg (Mecklenburg). Lectotype designated below. Junior primary homonym of Dolerus rugosus Freymuth, 1870. Dolerus rugosulus Dalla Torre, 1894: 16. A replacement name for D. rugosus Konow. Type material examined and taxonomic notes. Dolerus varispinus Hartig, 1837. Lectotype examined. ZSM. Dolerus rugosus Konow, 1884. Lectotype ♂ hereby designated, labelled: " Dolerus rugosus m [mihi]. fbg. 4.84" (Fürstenberg / Mecklenburg, IV.1884) [with a double black frame], " Dolerus rugosus D. T. [Dalla Torre]", " Syntypus " [red, printed]; " Lectotypus [printed] ♂ Dolerus rugosus Konow, 1884, design. M. Heidemaa and Viitasaari [printed] 1999 [handwritten]" [red label], "GBIF-GISHym 3770" [printed], "Coll. Konow" [printed], "Eberswalde coll. DEI" [printed], " Dolerus 1997 [printed] Dolerus varispinus Hartig [handwritten] M. Heidemaa det. [printed]". SDEI. The taxonomic identity of the lectotype is certain despite its poor condition, because the katepimeron of the mesopleuron is preserved., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Blank, SM, Taeger, A, 1992. Die von Th. Hartig und A. Foerster in der Gattung Dolerus beschriebenen Arten (Hymenoptera, Tenthredinidae). Entomofauna. Zeitschrift fuer Entomologie 13 (12): 213 - 232"]}

Dolerus pratensis (Linnaeus, 1758) Figs 13A, B, 14C, 14F, 15 Tenthredo pratensis Linnaeus, 1758: 556. Sex not given. Syntypes (assumed). Lectotype ♀ designated by Malaise and Benson (1934). Type locality: Europe. Tenthredo (Dolerus) deserta Klug, 1818: 300-301. ♂. Holotype. Published type locality: "In hiesiger Gegend gefunden" [meaning the environs of Berlin, Germany]. Type material examined and taxonomic notes. Tenthredo pratensis Linnaeus, 1758. Lectotype. LSUK. Images of the lectotype (LINN 2501) in dorsal and lateral view and the labels are available (The Linnean Society of London 2022), but we did not examine the specimen itself. Tenthredo (Dolerus) deserta Klug, 1818. Holotype (GBIF-GISHym 2316; Fig. 15). Good condition, genitalia dissected and pinned with the specimen. ZMHB., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Malaise, R, Benson, RB, 1934. The Linnean Types of Sawflies (Hymenoptera, Symphyta). Arkiv foer Zoologi 26(4 [A20]): 1-14. https://doi.org/10.1080/00222933408654888","2022. . http://linnean-online.org"]}

Claremontia confusa (Konow, 1886) sp. rev. Fig. 7A Blennocampa confusa Konow, 1886b: 82. ♀. Type locality: Germany. Neotype ♀ (ZMHB) designated by Koch (1988). Synonymy with Claremontia brevicornis proposed by Koch (1988), wherein he overlooked the priority of the name Claremontia brevicornis (Blank and Taeger 1998). Monophadnoides confusa: Benson (1952); treated as species distinct from M. puncticeps, cultivated Fragaria species recorded as hosts of larva. Biology. Substantiating the statement about the host plant by Benson (1952), the ZMUO specimens, collected in the Helsinki area, were reared from larvae feeding on cultivated Fragaria., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Koch, F, 1988. Die Gattung Claremontia Rohwer (Hymenoptera, Tenthredinidae). Deutsche Entomologische Zeitschrift 35 (4-5): 311 - 330, DOI: https://doi.org/10.1002/mmnd.4810350421","Blank, SM, Taeger, A, 1998. Comments on the taxonomy of Symphyta (Hymenoptera) (Preliminary studies for a catalogue of Symphyta, part 4). In: Taeger, A, Blank, SM, Eds., Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme. Goecke and Evers, Keltern: 141 - 174","Benson, RB, 1952. Hymenoptera, Symphyta. Handbooks for the Identification of British Insects 6(2b): 51-137."]}

Dolerus liogaster Thomson, 1871 Fig. 16A, C, E, H, K, N, Q Dolerus liogaster Thomson, 1871: 286. ♀, ♂. Syntypes. Type locality: Sweden. Lectotype designated below. Dolerus schneideri Kiaer, 1898: 62-63. ♀, ♂. Syntypes. Type locality: Tromsø. Lectotype designated below. Dolerus truncatus? Dolerus truncatus Lacourt, 1988: 233-235. ♂. Holotype. Type locality: France, Alpes-Maritimes, Vallée de la Minière. CTN. Type material examined and taxonomic notes. Dolerus liogaster Thomson, 1871. Lectotype ♂ hereby designated, labelled: "Lp. in." [?Lapland], "LECTOTYPUS [printed] Dolerus liogaster ♂ Thomson, 1871 [handwritten in block letters] design. 1999 M. Heidemaa and M. Viitasaari [printed]" [red], " Dolerus liogaster ♂ [handwritten] M.Heidemaa det. [printed]" [white]. Penis valves dissected, pinned with the specimen. Paralectotypes: 2♀, 1♂. All in MZLU. Dolerus schneideri Kiaer, 1898. Lectotype ♀ hereby designated, labelled: “Tromsö 22.6.77", "lget expl" [spelling doubtful], "Ex coll. H. Kiaer" [printed], “261” [yellow], "LECTOTYPUS [printed] Dolerus schneideri ♀ Kiaer, 1898 [handwritten in block letters] design. 1999 M. Heidemaa and Viitasaari" [printed (excl. year), red], " Dolerus liogaster ♀M. Heidemaa det." [printed label]. Ovipositor dissected and pinned with the specimen, right antenna missing. Paralectotypes: 2♀. All in ZMUN. The synonymy of D. schneideri with D. liogaster was first proposed by Lindqvist (1943). Treated here as a color form of D. liogaster, but still separated in the key, because the melanic color form of D. liogaster also has black legs, and D. schmidti has a color form with a red-marked thorax. Specimens resembling D. schneideri are also known from the mountain areas of Central Europe, e.g. from Switzerland at altitudes of 1500-1700 m (Benson 1961). Dolerus truncatus Lacourt, 1988. The holotype (CTN) was studied. Its clypeus emargination looks as if it is abnormally developed. The penis valve (Fig. 18) appears to be somewhat distorted, perhaps during preparation, but resembles the valve of D. liogaster. Tentatively, we treat D. truncatus as a junior synonym of D. liogaster., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Lindqvist, E, 1943. Zur Kenntnis einiger Tenthrediniden. Notulae Entomologicae 23: 72 - 76","Benson, RB, 1961. The sawflies (Hymenoptera Symphyta) of the Swiss National Park and surrounding Area. Ergebnisse der wissenschaftlichen Untersuchungen im Schweizerischen Nationalpark 7: 161 - 195"]}

Gilpinia pallida (Klug, 1812) Lophyrus pallidus Klug, 1812: 54-55. ♀, ♂. Syntypes. Type locality: Sweden and northern Germany. ZMHB. Lophyrus catocalus Snellen van Vollenhoven, 1858: 276-277. ♀, ♂. Syntypes. Lectotype ♀ designated by Thomas (1987). Type locality: Netherlands, Groesbeek near Nijmegen. syn. nov. Gilpinia catocala: Thomas (1987) recombination. Gilpinia verticalis Gussakovskij, 1947: 154-155, 227-228. ♀, ♂. Syntypes. Type locality: localities in Poland, Latvia and Russia. Synonymy with G. catocala by Thomas (1987). Notes. Although we have not studied the lectotype of Lophyrus catocalus, the interpretation of its taxonomic status is possible after referring to Gussakovskij’s description of G. verticalis and the other publications cited above. We agree with Pschorn-Walcher (1982) and Viitasaari and Varama (1987) that the two forms are an expression of individual variability, linked to population differences with a geographical component. Accordingly, we treat Gilpinia catocala as a junior synonym of G. pallida., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Thomas, PLL, 1987. An annotated catalogue of primary types of Symphyta (Hymenoptera) in the Netherlands. Zooelogische Mededeelingen 61 (5): 61 - 78","Pschorn-Walcher, H, 1982. Unterordnung Symphyta, Pflanzenwespen. In: Schwenke, W, Ed., Die Forstschaedlinge Europas. Paul Parey, Hamburg und Berlin, 4: 4-196: 232 - 234","Viitasaari, M, Varama, M, 1987. Sahapistiaeiset 4. Havupistiaeiset (Diprionidae). Reports, University of Helsinki. Department of Agricultural and Forest Zoology 10: 1 - 79"]}

Cephalcia arvensis Panzer, 1802 Fig. 6 Cephalcia arvensis Panzer, 1802: vol. 86 pl. 9. ♀. Syntypes. Type locality: Germany [according to title of work]. Blank et al. (2009); Nomen protectum with respect to Psen lucorum Schrank, 1802, nomen oblitum. Cephalcia abietis var. intermedia Hellén, 1948: 40-41. ♀. Syntypes. Type locality: Russia, Karelia, Paanajärvi. Cephalcia intermedia: Vikberg (1982); raised to species rank. Van Achterberg and van Aartsen (1986); synonymy with C. lariciphila (Wachtl, 1898). syn. nov. Notes. Cephalcia intermedia has in recent years mostly been understood as a distinct species close to C. arvensis (e.g. Viitasaari 2002b, Taeger et al. 2006), but with more extensively dark-patterned adults, occurring in northern Europe and the central European mountains. Because of its dark coloration, specimens of C. intermedia can also be mixed up with C. lariciphila (Wachtl, 1898), but the host plant of C. intermedia is Picea, as in C. arvensis, not Larix as in C. lariciphila. The status of C. intermedia and the supposed differences to C. arvensis were discussed by Vikberg (1982), Shinohara (1985), Midtgaard (1987) and Viitasaari (2002b). The only differences between C. arvensis and C. intermedia are in coloration: mainly of the abdomen, and less so of the antennae and legs. Shinohara (1985) pointed out that an unbroken range of color variability occurs between pale C. arvensis and the dark specimens identified as C. intermedia. This is in accordance with our observations, as the amount of dark color on the male abdomen varies considerably even in a single locality (Fig. 6). Small differences in colouration of larvae are possibly not constant, as the larvae of C. intermedia used for comparison were offspring of a single female. At present, there seems to be no convincing evidence for treating C. intermedia as a species distinct from C. arvensis, and we therefore synonymize them., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Blank, SM, Taeger, A, Liston, AD, Smith, DR, Rasnitsyn, AP, Shinohara, A, Heidemaa, M, Viitasaari, M, 2009. Studies toward a World Catalog of Symphyta (Hymenoptera). Zootaxa 2254 (1): 1 - 96, DOI: https://doi.org/10.11646/zootaxa.2254.1.1","Vikberg, V, 1982. Notes on the taxonomy and the nomenclature of some mainly Fennoscandian sawflies (Hymenoptera, Symphyta). Notulae Entomologicae 62 (2): 61 - 65","van Achterberg, C, van Aartsen, B, 1986. The European Pamphiliidae (Hymenoptera: Symphyta), with special reference to the Netherlands. Zooelogische Verhandelingen 234: 1 - 98","Viitasaari, M, 2002b. The Northern European taxa of Pamphiliidae (Hymenoptera). In: Viitasaari, M, Ed., Sawflies (Hymenoptera, Symphyta) I. A review of the suborder, the Western Palaearctic taxa of Xyeloidea and Pamphilioidea. Tremex, Helsinki: 235 - 358","Taeger, A, Blank, SM, Liston, AD, 2006. European Sawflies (Hymenoptera: Symphyta) - A Species Checklist for the Countries. In: Blank, SM, Schmidt, S, Taeger, A, Eds., Recent Sawfly Research: Synthesis and Prospects. Goecke and Evers, Keltern: 399 - 504","Shinohara, A, 1985. Some records of European Pamphiliidae (Hymenoptera). Entomologist's Gazette 36 (2): 161 - 164","Midtgaard, F, 1987. The Norwegian Xyelidae and Pamphiliidae (Hymenoptera). Fauna Norvegica, Ser. B: Norwegian Journal of Entomology 34 (2): 125 - 130"]}

Macrophya infumata Rohwer, 1925 (Tenthredinidae, Tenthredininae) Fig. 38 Notes. Finland: Karelia borealis, Kitee: 1♂, Puhos [62.082°N, 29.935°E], 30.06.2017, leg. Juha Salokannel. 1♀, Pajarinmäki, 16.06.2020, YKJ grid coordinates: 68898:36667 [62.078°N, 30.191°E], leg. Jaakko Pohjoismäki and M. Mutanen. 3♂, Pajarinmäki, 18.06.2021, YKJ grid coordinates: 68898:36667 [62.078°N, 30.191°E], leg. A. Liston, M. Mutanen, N. Kiljunen, M. Prous. Approximately 30 larvae, on Sambucus racemosa, Pajarinmäki, 26.07.2021, YKJ grid coordinates: 68898:36667 [62.078°N, 30.191°E], leg. M. Mutanen and M. Prous. All specimens in ZMUO (Fig. 38). In Europe previously recorded from the Russian regions of Kirov and Perm (Zhelochovtsev and Zinovjev 1996), but in iNaturalist (2022) more western Russian records are to be found (Kursk, Moscow, and St Peterburg Regions). The species is widespread in the East Palaearctic, from W. Siberia to Sakhalin, Japan, and northern China (Mallach 1936; Yoshida 2017). Host plant is Sambucus racemosa, including the closely related Japanese S. sieboldiana (Sakurai et al. 2009). The larvae were beaten from lower branches of S. racemosa in a shady habitat., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Zhelochovtsev, AN, Zinovjev, AG, 1996. Spisok pilil'shhikov i rogohvostov (Hymenoptera, Symphyta) fauny Rossii i sopredel'nyh territorij. II. Entomologicheskoe Obozrenie 75 (2): 357 - 379","2022. . https://www.inaturalist.org/taxa/564975-Macrophya-infumata","Mallach, N, 1936. Dritter Beitrag zur Kenntnis der Blattwespenfauna Chinas. Bulletin of the Fan Memorial Institute of Biology, Zoology. Peiping 6: 217 - 222","Yoshida, H, 2017. Checklist of Japanese Symphyta. Ver. 3.11. (7.IV.2017). https://symphyta.jimdo.com/","Sakurai, A, Hamanishi, Y, Maeto, K, Naito, C, 2009. Molecular biogeography of two sibling species of the sawfly genus Macrophya (Hymenoptera: Tenthredinidae) in Japan. Zoological Science 26 (5): 325 - 329, DOI: https://doi.org/10.2108/zsj.26.325"]}

Dolerus aericeps Thomson, 1871 Dolerus aericeps Thomson, 1871: 285. ♀, ♂. Syntypes. Published type locality: “södra Sverige" [southern Sweden]. Lectotype designated below. Tenthredo Eglanteriae [sic!] Fabricius, 1793: 109-110. syn. nov., nomen oblitum. ♀. Syntypes (assumed). Published type locality: "Habitat in Germania. Dom. Smidt." [Germany]. Lectotype designated below. Tenthredo (Dolerus) palustris Klug, 1818: 296-297. ♀, ♂. Syntypes. Published type locality: “Deutschland”. Lectotype designated below. Primary homonym of Tenthredo (Allantus) palustris Klug, 1818. syn. nov. Dolerus bajulus Serville, 1823: 58. ♀. Syntypes (assumed). Type locality: “Paris”. Lectotype designated below. Type material examined and taxonomic notes. Dolerus aericeps Thomson, 1871. Lectotype ♀ hereby designated, labelled: “Båst[ad]” [Type locality: Sweden, Skåne, Båstad], " Dolerus aericeps " [blue line along upper margin], " MZLU 2013 416" [pale green, printed, loan record], " Lectotypus 2014 Dolerus aericeps ♀ Thomson, 1871 M.Heidemaa design." [printed, red], " Dolerus aericeps Thomson, 1871 M.Heidemaa det.". In excellent condition. MZLU. Paralectotypes: 2♀, 2♂. MZLU. Tenthredo eglanteriae Fabricius, 1793. Lectotype ♀ hereby designated, labelled: " eglan [...] riae ", "LECTOTYPE Tenthredo eglanteriae Fabricius, 1793 des. SMBlank 2008" [red, printed], " Dolerus eglanteriae (F.) = Dolerus bajulus Serv. [handwritten] det. S.M.Blank 2008 ✓" [printed], " ZMUC-GISHym 1014", " Dolerus aericeps Thomson, 1871 M.Heidemaa det." [printed]. Left hind tarsus and some other tarsomeres missing. ZMUC. Among 2♀ and 1♂ specimens preserved in the ZMUC collection under the name T. eglanteriae, only the above female can be regarded as a syntype. The scutellum of the other female is red, and the tip of the abdomen of the male black. Both characters conflict with Fabricius’ original description. Compared with the other two specimens, the pale parts of the legs of the lectotype are darkened. The lectotype agrees with the species which has for a long time been known as Dolerus aericeps Thomson, 1871 (Zhelochovtsev 1994), characterized for example by the laterally directed and distally curved setae of the ovipositor sheath. Lacourt (2000) used the name D. bajulus Serville, 1823 for this species, but Blank et al. (2009) presented arguments for the use of the name D. aericeps (nomen protectum) versus D. bajulus (nomen oblitum). Tenthredo eglanteriae was synonymized with T. germanica by Klug (1819) and with Athalia glabricollis Thomson, 1870 by Konow (1897a). Since Brullé (1846), T. eglanteriae (nomen oblitum) has never again been used as valid, while between 1971-2021 the name Dolerus aericeps (nomen protectum) was used as valid by more than 120 authors in over 150 publications (Article 23.9.1, ICZN 1999). One paralectotype male of D. cothurnatus Serville belongs to this species (see under D. germanicus). Tenthredo (Dolerus) palustris Klug, 1818. Lectotype ♂ hereby designated, labelled: “14190” [collection catalog no.], "cotypus von Dol. palustris Klg. = Dolerus aericeps Thoms. O.Conde det 1941" [handwritten by Otto Conde], "GBIF-GISHym 2342" [printed], " Lectotypus [printed] ♂ Tenthredo (Dolerus) palustris Klug, 1818 [handwritten in block letters] M.Heidemaa des. [printed]" [red], " Dolerus aericeps Thomson, 1871 M.Heidemaa det.", "Zool. Mus. Berlin" [printed]. Left forewing and metatarsi missing, half of the genital capsule glued on a paper card and pinned with the lectotype. Paralectotype ♂ ("GBIF-GISHym 2341") with same labelling (except " Paralectotypus "). All in ZMHB. Formerly placed as a synonym of D. cothurnatus auct. (= D. junci (Stephens, 1835)), e.g. by Taeger et al. (2010). Dolerus bajulus Serville, 1823. Lectotype ♀ hereby designated, labelled: " Dolerus bajulus, Lep.", "Lectotype [red, printed in block letters]", "Lectotype D. bajulus LEP. det.: H.Haris 1996" [white], " D. aericeps THOMS. ♀ det.: A.Haris", " Lectotypus [printed] ♀[handwritten] Dolerus bajulus Serville, 1823 [handwritten in block letters] M.Heidemaa design." [printed, red]. " Dolerus aericeps Thomson, 1871 M.Heidemaa det" [printed]. MNHN., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Zhelochovtsev, AN, 1994. [Translation of the book published in Russian in 1988]. In: Medvedev GS (Ed.) Keys to the Insects of the European Part of the USSR. Volume III Hymenoptera Part VI Symphyta. E.J. Brill, Leiden","Lacourt, J, 2000. Liste des especes de la famille des Tenthredinidae decrites par J. G. Audinet-Serville, en Mai 1823 et par A. L. M. Le Peletier Comte de Saint-Fargeau, en Aout 1823, avec designation de lectotypes [Hymenoptera, Symphyta]. Revue francaise d'Entomologie (N. S.) 22 (2-3): 77 - 108","Blank, SM, Taeger, A, Liston, AD, Smith, DR, Rasnitsyn, AP, Shinohara, A, Heidemaa, M, Viitasaari, M, 2009. Studies toward a World Catalog of Symphyta (Hymenoptera). Zootaxa 2254 (1): 1 - 96, DOI: https://doi.org/10.11646/zootaxa.2254.1.1","Klug, F, 1819. Die Blattwespen (Tenthredo Linn.) der Fabricischen Sammlung. Zoologisches Magazin (herausgegeben von C. R. W. Wiedemann) 1[1817-1819](3): 64-91.","Konow, FW, 1897a. Synonymische Bemerkungen zu bisher nicht oder unrichtig gedeuteten Tenthredo Arten bei Geoffroy, Fabricius, Schrank und Villers. Wiener entomologische Zeitung 16: 1 - 18, DOI: https://doi.org/10.5962/bhl.part.12830","Brulle, A, 1846. Tome Quatrieme. In: Lepeletier de Saint-Fargeau (Ed.) Histoire naturelle des Insectes. Hymenopteres. Librairie Encyclopedique de Roret, Paris, 680 + 16 pp. [plates 37-48]","ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition.International Commission on Zoological Nomenclature, London, 306 pp.","Taeger, A, Blank, SM, Liston, AD, 2010. World Catalog of Symphyta (Hymenoptera). Zootaxa 2580 (1): 1 - 1064, DOI: https://doi.org/10.11646/zootaxa.2580.1.1"]}

Dolerus ferrugatus Serville, 1823 Dolerus ferrugatus Serville, 1823: 59. ♀. Syntypes. Type locality: Paris. Lectotype designated below. Dolerus brevicornis Thomson, 1871: 288. ♀, ♂. Syntypes. Published type locality: “södra Sverige" [southern Sweden]. Lectotype designated below. Primary homonym of D. brevicornis Zaddach, 1859. Dolerus thomsoni var. miricolor Konow, 1887a: 283. ♂. Syntypes (assumed). Type locality: Fürstenberg / Mecklenburg. Lectotype designated below. Type material examined and taxonomic notes. Dolerus ferrugatus Serville, 1823. Lectotype ♀ hereby designated, labelled: “Saint-Fargeau” [round], “Lectotype” [red, printed in block letters], "Lectotype D. ferrugatus LEP. ♀ det. A.Haris 1996", " D. ferrugatus LEP. ♀ det.: A.Haris", " Lectotypus [printed]♀ Dolerus ferrugatus Serville, 1823 [handwritten in block letters] M.Heidemaa design. [printed]" [red], " Dolerus ferrugatus Serville, 1823 M.Heidemaa det." [printed]. In good condition (the right flagellum missing). MNHN. Dolerus brevicornis Thomson, 1871. Lectotype ♀ hereby designated (type locality: Sweden, Öland), labelled: “Ö.” [Öland], " MZLU 2013 414" [green, printed (a loan record label)], " Lectotypus [printed] ♀ 2014 Dolerus brevicornis Thomson, 1871 [handwritten in block letters] M.Heidemaa des. [printed]" [red], " Dolerus ferrugatus Serville, 1823 M.Heidemaa det." [printed]. 2♀ (from Öland and Scania), 1♂ (the melanic form, from Lund) labelled as paralectotypes. All in MZLU. Dolerus thomsoni var. miricolor Konow, 1887. Lectotype ♂ hereby designated, labelled: "f. [Fürstenberg] 12/4. [18]84.", " Dolerus v. [var.] Dolerus miricolor Knw. [Konow] fbg. 12/4. 84. Typus" [with double black frame], " Dolerus ferrugatus Lep. f. miricolor Knw. Typus. ♂ det. OConde [Otto Conde] 1943", “Typus” [red, printed], “Holotypus” [red, printed], "Coll. Konow" [printed], "GBIF-GISHym 4215" [printed], " Lectotypus [printed] ♂ Dolerus thomsoni var. miricolor [handwritten in block letters] M.Heidemaa des. [printed]" [red], " Dolerus ferrugatus Serville, 1823 M.Heidemaa det." [printed]. Right flagellum missing, penis valves dissected (a microscope slide: Symphyta coll. Nr: 159). All in SDEI. The melanic colour form of D. ferrugatus, described by Konow as D. thomsoni var. miricolor, has sometimes been mixed up with D. pachycerus Hartig, 1837. Apart from their different penis valves, they can be separated by the shape and the structure of the mesoscutellar appendage: long, concave at the sides, and with a distinct longitudinal keel in D. pachycerus compared with D. ferrugatus., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080

Dolerus aericepsellus Heidemaa & Mutanen sp. nov. Figs 8, 9 Description. Holotype ♀. Figs 8A, B, 9B, C. Colour. Fig. 8A, B. Body black with terga 2-7 and sterna 2-7 orange. Terga 8-9 brownish black, 10 brownish basally but yellowish brown apically, apical third of protibiae brownish. Ventral margin of valvula 3 and cerci brownish. Wings clear. Measurements (mm). Body length: 5.0, distance from tegula to base of pterostigma: 3.1, head breadths: 1.46 (max. at eyes), 1.28 (max. behind eyes), 1.07 (min. behind eyes), head length (behind eyes): 0.21, max diameter of eye: 0.62, breadth of postocellar field: 0.36, OC: 0.96, OOCL: 0.35, OOL: 0.20, POL: 0.19. Length of: metafemur: 1.18, metatibia: 1.76, ovipositor sheath (from the base of valvifer 2 to the apex of valvula 3): 1.01. Length of flagellomeres: 1: 0.48, 2: 0.41, 3: 0.37, 4: 0.31, 5: 0.29, 6: 0.26, 7: 0.25. Large structures. Clypeal emargination almost half as deep as clypeal median length. Clypeus asymmetrical, its antero-lateral lobes round at apex, the left lobe longer than the right. Distance between antennal sockets 1.6 × as long as the malar space. Length of antennomere 3 about 1.24 × that of 4. Oblique furrow not outlined. Distance between cenchri about 1.1 × as long as width of cenchrus. Metatarsomere 1 about 1.75 × as long as 2. Mesoscutellar appendage with weakly outlined ridge medially. Setae. Metascutellum with few setae (about 5). Abdomen dorsally mostly pubescent from segment 7 to last one (laterally and ventrally from 2 to last one). Longest setae of valvula 3 in dorsal view markedly curved in apical third, and forming an angle of about 80-90° (Fig. 9B). Macrosculpture. Pits on head rather irregular in size and distribution, more distinct and separated on postocular and postocellar area. Density and size of pits on thorax vary, the largest (partly fused) on lateral sides of median mesoscutal lobes and on mesoscutellum. Pits on lateral lobes of mesoscutellum and near the median mesoscutal groove sparse and small. Pits on mesepisternum largest, partly isolated and round, partly polygonal and fused (Fig. 9C). Distinct pits on pectus nearly absent (Fig. 9C). Microsculpture. Meshes on mesoscutellar appendage extensive, on tergum 1 absent, on metepimeron (Fig. 9C) and on ventral surface of metacoxa absent. Sculpticells on anterior half of katepimeron of mesopleuron rib-like, irregular and of uneven height, and absent on metepimeron, pectus, and terga (all glossy). Male. Figs 8C, D, 9A, D-G. Closely resembles the female, but the anterior margin of tergum 2 blackish, basal 2/3 of hind tibia (with apical spurs), and base of metatarsomere 1 reddish yellow (Fig. 8C, D). Penis valve valviceps long and narrow (Fig. 9G). Measurements (mm). Body length (paratype male ZMUO.035740): 5.3, distance from tegula to base of pterostigma: 3.2, max. diameter of eye: 0.60, head breadth: 1.47 (max. at eyes), breath of postocellar field: 0.38, head length (behind eyes): 0.24, OC: 1.01, OOCL: 0.18, OOL: 0.29, POL: 0.21, metafemur: 1.25, metatibia: 1.64, metatarsus: 0.50. Length of flagellomeres: 1: 0.59, 2: 0.51, 3: 0.50, 4: 0.44, 5: 0.41, 6: 0.39, 7: 0.41. Genetic data. Three specimens with DNA barcode sequence data are available. They comprise a distinct cluster BIN BOLD:ABV8002, with 0.3% intraspecific variability. Based on sequences longer than 600 bp, the minimum distance to the closest relative D. aericeps (n=26) is 3.9%, but a few specimens identified as D. yukonensis from North America show a minimum distance of even less, 3.6%. Differential diagnosis. Adults of D. aericepsellus are most similar to D. aericeps Thomson, but are distinguished by the following characters: smaller body size (similar to D. elderi Kincaid, 1900, or smaller); very short, strongly narrowing postocular area; glossy metascutellum with few distinct pits and setae (3-6); pectus without distinct pits; and thorax black, including tegulae (based on the barcoded specimens: 1♀ 2♂). Lateral postocellar furrows very distinct in both sexes (Fig. 9F), like in D. aericeps and D. incisus. The thorax of female D. aericeps is usually largely red, whereas in the D. aericepsellus paratype female it is almost entirely black. Dolerus aericeps ab. nigricollis was described by Lindqvist (1943) from northern Finland close to the localities where D. aericepsellus was collected. We examined the type specimen of Dolerus aericeps ab. nigricollis deposited in the MZH and found that it is not conspecific with D. aericepsellus but with D. aericeps, although it has a nearly completely black thorax, with two obscure reddish flecks. Some other examined specimens of D. aericeps from northern Finland show reduced red markings on the thorax, suggesting that this character varies in D. aericeps. Type material. Holotype ♀. Type locality: Finland, Enontekiö, Pousu. Labelled [white, printed if not given otherwise]: "FIN Le Enontekiö Pousu 7651:3266 [68.846°N, 21.197°E] 27.6.2020 Marko Mutanen leg.", "Sawfly tissue 2020-1079 M.Mutanen" [greenish], “http://id.zmuo.oulu/fi/ZMUO.044566”, "Holotypus [block letters] ♀ Dolerus aericepsellus Heidemaa & Mutanen, 2022 [handwritten in block letters] des. M. Heidemaa" [red]. Good condition: wings spread, left fore and mid legs removed as tissue samples. ZMUO. Paratypes. Total: 1 ♀, 3 ♂: Finland: Lapponia inarensis, Utsjoki Pulmankijärvi, YKJ grid coordinates: 7761:3539 [69.928°N, 28.031°E], 1♂ (ZMUO.033245) 07.07.2017, leg. M. Mutanen, N. Mutanen, A. Mutanen, SDEI. 1♂ (ZMUO.035740) (Fig. 9A, D-G), same collection data as preceding, but 05.07.2018, ZMUO. Lapin Lääni, Nuorgam 17 km SSE, 69.944°N, 28.041°E, 1♀ (BC ZSM HYM 11450), leg. M. Kraus, ZSM. 1♂ (BC ZSM HYM 11451), same collection data as preceding, ZSM. Etymology. The name is an adjective derived from the species name of its closely similar sister species, D. aericeps Thomson, using the Latin diminutive suffix - ellus., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Lindqvist, E, 1943. Zur Kenntnis einiger Tenthrediniden. Notulae Entomologicae 23: 72 - 76"]}

Empria improba (Cresson, 1880) Fig. 22A-E Emphytus improbus Cresson, 1880: 11. ♀, ♂. Syntypes. Type locality: Nevada, USA. Lectotype ♂ (type No. 365) designated by Smith (1979). ANSP. Tenthredo (Poecilostoma) hybrida Erichson in: Ménétriés in: Middendorff, 1851: 60-61. ♀. Syntypes (assumed). Primary homonym of Tenthredo (Tenthredo) hybrida Eversmann, 1847. Type locality: Udskoj Ostrog [Russia, Khabarovsk Krai, Udskoe]. Lectotype ♀ designated by Prous et al. (2011). ZIN. syn. nov. Poecilosoma plana Jakowlew, 1891: 31. ♀. Type locality: Irkutsk, Russia. ♀. Holotype. ZIN. syn. nov. Empria itelmena Malaise, 1931b: 23. ♀, ♂. Syntypes. Type locality: Kamtschatka, E[lisowo] [Russia, Kamchatka Krai]. Lectotype ♀ designated by Prous et al. (2011). NHRS. syn. nov. Empria camtschatica Forsius, 1928: 46-47. ♀. Holotype. Type locality: Russia, Kamchatka Krai, Bolsheretsk [Bolscheretsk]. MZH. syn. nov. Notes. The species boundaries between willow-feeding taxa of the Empria immersa group (E. immersa, E. camtschatica, E. plana, and E. improba) have proved to be difficult to elucidate (Prous et al. 2014, 2020). In Fennoscandia, two forms can commonly be found at the same time and place (Prous et al. 2014): E. immersa with a dark pterostigma and short antenna, and E. camtschatica with a pale pterostigma and long antenna. These two forms can also be distinguished by larval morphology (Fig. 22). Based on ex ovo rearings by M. Prous (two females from Sweden and Estonia) and ex larva rearings by Ponomarev (2022) of E. immersa, and ex ovo rearings by M. Prous of E. camtschatica (using two females from Sweden), the main difference seems to be in head coloration: E. immersa with an occipital fleck or stripe (Fig. 22F-M) and E. camtschatica with occipital and parietal stripes (Fig. 22A-E). An additional difference may be that glandubae (white conical warts) are more prominent in E. immersa than in E. camtschatica. Although based on limited specimen sampling, genome scale data (Prous et al. 2020) support E. immersa as a distinct species most consistently compared to the other species in the E. immersa group. In Fennoscandia, taxonomy is complicated by the presence of occasional specimens identifiable as E. plana, somewhat intermediate in morphology between E. immersa and E. camtschatica (pterostigma like E. camtschatica, saw intermediate). Genome scale data of one E. plana female from Sweden do not indicate affinity with E. immersa, but do show at least some affinity with E. camtschatica (Sweden), E. improba (Canada), and one other E. plana (Hokkaido, Japan) (see fig. 5 in Prous et al. 2020). Given the above, we synonymize E. plana and E. camtschatica with E. improba, because clear boundaries between these taxa cannot at present be drawn. Thus, in Europe, the specimens with dark pterostigma, short antennae and more prominent serrulae of the saw can be identified as E. immersa, and those with pale pterostigma, usually longer antennae, and less prominent serrulae as E. improba (see Prous et al. 2014). In North America, however, at least some E. improba specimens look externally more like E. immersa (dark pterostigma and metafemur), while the serrulae of the saw resemble E. camtschatica. Lacourt (2020) suggested that E. camtschatica could be a synonym of E. improba, but genetically these taxa are not necessarily closer to each other than they are to E. plana (Prous et al. 2020). If the circumscription of E. improba as proposed here is considered incorrect, then it remains unclear how many additional species should be recognized, and how these should be delimited. For example, in Europe the morphological distinction between E. camtschatica and E. plana is not clear, although these forms can be more reliably distinguished from E. immersa., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Smith, DR, 1979. Nearctic sawflies. IV. Allantinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture, Washington DC 1595: 1 - 172","Prous, M, Heidemaa, M, Shinohara, A, Soon, V, 2011. Review of the sawfly genus Empria (Hymenoptera, Tenthredinidae) in Japan. ZooKeys 150: 347 - 380, DOI: https://doi.org/10.3897/zookeys.150.1968","Prous, M, Blank, SM, Heibo, E, Loennve, OJ, Taeger, A, Vardal, H, Liston, A, 2014. Sawflies (Hymenoptera, Symphyta) newly recorded from Sweden. Entomologisk Tidskrift 135 (3): 135 - 146","Prous, M, Lee, KM, Mutanen, M, 2020. Cross-contamination and strong mitonuclear discordance in Empria sawflies (Hymenoptera, Tenthredinidae) in the light of phylogenomic data. Molecular Phylogenetics and Evolution 143 (106670): 1 - 16, DOI: https://doi.org/10.1016/j.ympev.2019.106670","Ponomarev, AG, 2022. Babochki i zhuki Moscovskoj oblasti. Published on the Internet. http://insectamo.ru","Lacourt, J, 2020. Symphytes d'Europe. Hymenopteres d'Europe 2. N. A. P. Editions, Verrieres-le-Buisson"]}

Calameuta punctata (Klug, 1803) Figs 3, 4 Astatus punctatus Klug, 1803: 55, plate VII figs 2a, b. ♀. Syntypes (assumed). The type material is considered to be lost or destroyed. Published type locality: Germany [implicit from title of Klug’s work]. Neotype designated below. Calameuta punctata: Zombori (1978). Astatus floralis Klug, 1803: 53-54, plate VI figs 5a, b. ♂. Syntypes (assumed). The type material is considered to be lost or destroyed. Published type locality: Germany [implicit from title of Klug’s work]. syn. nov. Astatus analis Klug, 1803: 54-55, plate VII fig. 1. ♀. Syntypes (assumed). The type material is considered to be lost or destroyed. Published type locality: Germany [implicit from title of Klug’s work]. syn. nov. Cephus filum Gussakovskij, 1935: 112, 358, 361. ♀, ♂. Syntypes. Type locality: Sarepta, Caucasus, and southern Siberia. ZIN. syn. nov. Calameuta filum: Benson (1946). Type material examined and taxonomic notes. To help resolve the taxonomic disagreements in the interpretation of these nominal taxa, and promote the future stability of nomenclature, a neotype is designated for Astatus punctatus: Astatus punctatus Klug, 1803. Neotype ♀ (DEI-GISHym21255, Fig. 4A-D), hereby designated. Germany, Brandenburg, Landkreis Märkisch-Oderland, Müncheberg, Trebnitz, 52.535°N, 14.204°E, damp meadow, swept from Alopecurus pratensis, 16.05.2015, leg. A. Liston (deposited in the SDEI). Labelling [printed on pale paper if not stated otherwise]: "Germany: Brandenburg; Landkreis Märkisch-Oderland, Müncheberg Trebnitz 16.05.2015 leg. A. D. Liston", “21255” [handwritten] with part of a leg gummed to card], “DEI-GISHym21255”, "♀ Calameuta punctata (Klug) [handwritten] det. A. Liston 2018", "NEOTYPE ♀ Astatus punctatus Klug, 1803 designated A. Liston 2022" [red]. Klug’s description states that abdominal segment 4 has obscure, paired dorsal spots; segment 5 four separate yellow spots, one pair laterally, the other dorsally; segments 6 and 7 with spots on their lower posterior margins; segment 8 immaculate; segment 9 completely yellow. Accordingly, we selected as neotype a specimen with small pale markings on terga 4-7 as well as 8-10 (Fig. 4A-D). The abdomen of a second female (DEI-GISHym21260) collected at the same place and time has fewer and less extensive pale markings and is thus intermediate in this respect to other female C. punctata specimens collected in Germany and all known Finnish and Estonian specimens, which have a completely black abdomen apart from terga 8-10 (Fig. 3A, C). Astatus floralis and A. analis have in the past generally been treated as synonyms of Calameuta haemorrhoidalis auct. [our Calameuta variabilis], e.g. by Konow (1905a). The opinion that A. floralis is a synonym of Cephus pygmeus (Linnaeus, 1767), as in de Dalla Torre (1894), cannot be accepted: Klug’s description of leg colour does not fit C. pygmeus. Our reason for placing A. floralis and A. analis as synonyms of Calameuta punctata rather than of C. variabilis is based primarily on one of the main characters which distinguishes C. punctata from C. variabilis: the structure of the maxillary palps. In the description of Astatus which precedes the descriptions of A. floralis, A. analis and A. punctatus, Klug characterized the genus thus: "Palpi[...]anteriores[...]sexarticulati, articulis duobus baseos cylindricis, aequalibus, tertio crassiori, longiori, subcylindrico, quarto longissimo, graciliori, quinto brevissimo, ultimo longitudine fere tertii subulato[...]". The described proportions of maxillary palpomeres 5 and 6 therefore fit C. punctata (Fig. 3D), not C. variabilis (Fig. 5B). Calameuta variabilis is unique in Calameuta in having maxillary palpomeres 5 and 6 of almost equal length (Gussakovskij 1935; Benson 1968; Zombori 1978). Zombori (1978) correctly identified Calameuta variabilis [which he called C. haemorrhoidalis] as a taxon distinct from C. punctata, and summarized the characters that distinguish them, but interpreted some of the names wrongly. Notably, Zombori (1978) did not mention the major contradiction in the morphology of the maxillary palps, as described by Klug, when he tentatively suggested that A. floralis and A. analis might be synonyms of Astatus haemorrhoidalis auct. ["the description of the latter two [Astatus floralis, Astatus analis] rather corresponds to the one given by Fabricius for C. haemorrhoidalis, accordingly, they are considered as synonyms of the latter name."]. Zombori’s main reason for doubting that Astatus analis was synonymous with Calameuta punctata, seems to have been the wording of Klug’s descriptions, which suggested that the thorax of Astatus analis is shinier than that of Calameuta punctata. Apart from this, Klug’s description of A. analis fits the darker forms within the rather wide range of variability in the female sex of C. punctata. Strangely, in his discussion of these names, Zombori (1978) does not mention Calameuta filum at all. The explicit collection data given by Klug (1803) for Astatus floralis ("Locus in editioribus argillosis; in floribus"), A. analis ("Locus in editioribus; in floribus"), and A. punctatus ("Locus in floribus") are, in part, not easy to interpret. Clearly, "in floribus" means that the specimens were collected from flowers. We think that "in editioribus argillosis" refers simply to the type of locality, i.e. an elevated place on clayey ground. This fits well with the type of sites at which C. punctata has recently been collected in Germany (see below). The synonymy of Calameuta filum with C. punctata can be proposed with a high degree of confidence. The characters described by Gussakovskij (1935) for the former are precisely those used by Zombori (1978) to characterize the latter. The same characters are also given by Viitasaari (1975) in his description of Finnish specimens identified as Calameuta filum, and which he compared with a syntype of that species. Viitasaari (1984) subsequently noted that Calameuta punctata sensu Zombori (1978) and C. filum are probably conspecific. Biology and distribution. The only recorded host plant of Calameuta punctata is Alopecurus pratensis L. (Vikberg 1978; Liston 2015), on which it is possibly monophagous. Accordingly, C. punctata occurs mostly in rather moist places. Its wider geographic range is not entirely clear, particularly because the identity of Calameuta pravei (Dovnar-Zapolskij, 1926) remains unresolved. This has been considered to be a valid species (e.g. Llorente and Gayubo 1990), or a synonym of C. punctata (e.g. Taeger et al. 2010). Calameuta pravei was recorded by Gussakovskij (1935) from Transcaucasia, Crimea, and the western Kopet-Dagh (Turkmenia). Llorente and Gayubo (1990) added records from Spain. Excluding these records, C. punctata is known from south-east Russia, Transcaucasia and south Siberia to Irkutsk (Gussakovskij 1935, as Cephus filum), and central and northern Europe (specimens examined by us). The distribution of C. punctata is therefore rather different from that of C. variabilis (see below), but their ranges overlap at least in parts of south-central Europe. Calameuta variabilis [as C. haemorrhoidalis auct.] has been stated to occur in Germany based on the mention of Germany as the type locality in the original descriptions by Fabricius (1781) of Tenthredo haemorrhoidalis and by Klug (1803) of Astatus punctatus, A. floralis and A. analis, coupled to an apparently faulty understanding of which taxa are represented by these names. Blank et al. (1998, 2001) listed C. haemorrhoidalis from Germany, only for Berlin-Brandenburg, dating the record respectively as "vor 1803" and “1802”. This refers to the type material of Klug’s species. Later, Liston et al. (2012) treated C. haemorrhoidalis auct. [C. variabilis] as extinct in Germany, and added C. punctata to the German list, based on recently collected specimens. Although it cannot be ruled out that C. variabilis once occurred in Germany, but has since disappeared, we think it more likely that in historical times only C. punctata ever occurred there, and propose in future to include only it in the list of German Symphyta. Fennoscandian and Estonian specimens which were previously identified as C. filum also belong to C. punctata. Based on COI sequences, C. punctata is split into two barcode clusters. Three specimens from Finland and one from Estonia are identical (BOLD:ACQ7596), but differ from two German specimens by 5.0-5.5% (no BIN assigned yet, GenBank accessions MW353981 and MW353982). The BOLD:ACQ7596 is closer to C. pallipes, differing by a minimum of 4.1%., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Zombori, L, 1978. New Sawfly Species in the Hungarian Fauna (Hymenoptera, Symphyta), IV. Annales Historico-Naturales Musei Nationalis Hungarici 70: 259 - 264","Benson, RB, 1946. Classification of the Cephidae (Hymenoptera Symphyta). Transactions of the Royal Entomological Society of London 96 (6): 89 - 108, DOI: https://doi.org/10.1111/j.1365-2311.1946.tb00445.x","Konow, FW, 1905a. Hymenoptera. Fam. Lydidae. Genera Insectorum 27: 1 - 27","de Dalla Torre, CG, 1894. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 1: Tenthredinidae incl. Uroceridae (Phyllophaga & Xylophaga). Guilelmus Engelmann, Lipsiae, [[6] pp.+ pp. I-VIII +] 1-459. https://doi.org/10.5962/bhl.title.10348","Gussakovskij, VV, 1935. Insectes Hymenopteres, Chalastogastra 1. In: Stackelberg, AA, Ed., Faune de l'URSS. Academie des Sciences de l'URSS, Moscou, Leningrad 2(1): 1 - 453","Benson, RB, 1968. Hymenoptera from Turkey, Symphyta. Bulletin of the British Museum (Natural History). Entomology series 22 (4): 111 - 207, DOI: https://doi.org/10.5962/bhl.part.9952","Klug, F, 1803. Monographia Siricum Germaniae atque generum illis adnumeratorum. F. Schueppel, Berolini, [i-xii+] 1-64. [+7 unpag] https://doi.org/10.5962/bhl.title.10284","Viitasaari, M, 1975. On the genera Janus Lepeletier and Calameuta Konow (Hym., Cephidae) in Eastern Fennoscandia. Annales Entomologici Fennici 41 (4): 113 - 120","Viitasaari, M, 1984. Sahapistiaeiset 3. Siricoidea, Orussoidea and Cephoidea. Reports, University of Helsinki. Department of Agricultural and Forest Zoology 6: 1 - 66","Vikberg, V, 1978. Calameuta filum (Gussakovskij) (Hymenoptera, Cephidae) found in South Haeme. Notulae Entomologicae 58 (4): 176 - 177","Liston, AD, 2015. New records and host plants of Symphyta (Hymenoptera) for Germany, Berlin and Brandenburg. Contributions to Entomology. Beitraege zur Entomologie 65 (2): 383 - 391, DOI: https://doi.org/10.21248/contrib.entomol.65.2.383-391","Llorente, G, Gayubo, SF, 1990. Estudio sobre la sinfitofauna del oeste espanol. I (Hymenoptera, Symphyta). EOS. Revista Espanola de Entomologia 65 (2): 23 - 29","Taeger, A, Blank, SM, Liston, AD, 2010. World Catalog of Symphyta (Hymenoptera). Zootaxa 2580 (1): 1 - 1064, DOI: https://doi.org/10.11646/zootaxa.2580.1.1","Fabricius, JC, 1781. Species Insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Tom. I. Carol. Ernest Bohn, Hamburgi et Kilonii, [pp. viii+] 552 pp. https://doi.org/10.5962/bhl.title.36509","Blank, SM, Boeve, J-L, Heitland, W, Jaenicke, M, Jansen, E, Koch, F, Kopelke, J-P, Kraus, M, Liston, AD, Ritzau, C, Schmidt, S, Taeger, A, 1998. Checkliste der Pflanzenwespen Deutschlands (Hymenoptera: Symphyta). In: Taeger, A, Blank, SM, Eds., Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme. Goecke and Evers, Keltern: 13 - 34","Blank, SM, Deters, S, Drees, M, Jaenicke, M, Jansen, E, Kraus, M, Liston, AD, Ritzau, C, Taeger, A, 2001. Symphyta. In: Dathe, HH, Taeger, A, Blank, SM, Eds., Verzeichnis der Hautfluegler Deutschlands (Fauna Germanica 4). Entomologische Nachrichten und Berichte, Beiheft 7: 8 - 27","Liston, AD, Jansen, E, Blank, SM, Kraus, M, Taeger, A, 2012. Rote Liste und Gesamtartenliste der Pflanzenwespen (Hymenoptera: Symphyta) Deutschlands. Stand Maerz 2011. In: Binot-Hafke, M, Balzer, S, Becker, N, Gruttke, H, Haupt, H, Hofbauer, N, Ludwig, G, Strauch, M, Eds., Rote Liste gefaehrdeter Tiere, Pflanzen und Pilze Deutschlands. Wirbellose Tiere Teil 1. Naturschutz und Biologische Vielfalt, Bonn-Bad Godesberg 70(3): 489 - 556"]}

Dolerus gessneri André, 1880 Fig. 19 Notes. Dolerus Gessneri [sic!] André, 1880: 273. Sex not given. Syntypes. Type locality: Switzerland. Lectotype ♀ hereby designated, labelled: “Suisse”, "DEI GISHym 89367" [printed], "coll. Desbrochers [handwritten] k. [kollektsija = coll.] A. Jakovleva [nomin. A. Jakovlev]" [printed], "SYNTYPUS [printed in red] Dolerus gessneri ANDRÉ, 1880 [handwritten] teste A. Taeger 20[printed]18 [handwritten]" [white with a red frame]. "Lectotype Dolerus gessneri André 1880 vide A. Taeger 2022 des." [red, printed]. Good condition, but slightly dusty. Paralectotypes: 2♂ (one without flagella) with the same data (except “♂” [printed], "DEI-GISHym 89365" and “D… 89366" and " Paralectotypus " instead of " Lectotypus "). All in ZIN. The color form of Dolerus gessneri with a red-banded abdomen was described as a distinct species, D. labiosus Konow, 1897 (Konow 1897b), but was later mostly treated as a synonym (sometimes also as a subspecies) of D. gessneri, because intermediate color forms with variable red markings on the abdomen occur. The specimens from the Russian Far East with completely black forelegs, which have sometimes been determined as D. gessneri, probably belong to some other Dolerus (subgen. Equidolerus) species. Dolerus gessneri, a Holarctic species, needs a revision, because COI barcoding data suggest that it could include more than one species, and some morphological characters correlate with the genetic data. There are three BIN clusters (BOLD:AAL2317, BOLD:ACE7304, BOLD:ACE3617) diverging by 1.8-2.3%. BOLD:AAL2317 is closest to D. timidus and BOLD:ACE7304 is closest to D. pratensis (minimum divergence in both cases 1.2%). All three BIN clusters are represented in Fennoscandia, two of which (Fig. 19) have been found in Finland., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Andre, E, 1880. Species des Hymenopteres d'Europe & d'Algerie. L'Auteur, Beaune (Cote-d'Or), 1[1879-1882](7): 237-300. [catalogue 29-36]","Konow, FW, 1897b. Neue palaearctische Tenthrediniden. Wiener entomologische Zeitung, 16 (6): 173 - 187"]}

Dolerus vulneratus Mocsáry, 1878 Notes. Dolerus vulneratus Mocsáry, 1878: 199. ♀. Syntypes (assumed). Type locality: Siberia. Lectotype ♀ hereby designated, labelled: “Siber[ia]”, " Dolerus vulneratus ♀ Mocs.", "Holotypus [printed with red] ♀ Dolerus vulneratus Mocsáry, 1878 Zombori, 1977 [handwritten]", " Lectotypus [printed] ♀ Dolerus vulneratus Mocsáry, 1878 M.Heidemaa des. ‘22” [handwritten, red]. " Dolerus vulneratus Mocsáry, 1878 M.Heidemaa det." [printed]. HNHM. The “holotype” label attached by Zombori has no nomenclatural significance, because this interpretation was never published. Moreover, the number of specimens was not given in the original description and the ♀ symbol does not necessarily indicate that Mocsáry had only one specimen, even if only one specimen was found in the author’s collection. Such assumptions about the status of specimens as holotypes should be avoided according to the Code (ICZN 1999: Articles 72.4.7, 73F). The larva is known through rearing and observations by Ponomarev (2022). Host plant: unidentified Poaceae spec., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition.International Commission on Zoological Nomenclature, London, 306 pp.","Ponomarev, AG, 2022. Babochki i zhuki Moscovskoj oblasti. Published on the Internet. http://insectamo.ru"]}

Fenusa ewaldi D. R. Smith, 2011 (Tenthredinidae, Blennocampinae) Figs 36, 37 Notes. A total of about 40♀ and numerous mines. All specimens leg. M. Mutanen, in ZMUO and SDEI: Finland: Ostrobottnia ouluensis, Oulu Linnanmaa kasvipuutarha [Oulu Botanic Garden], YKJ grid coordinates: 7219:3427 [65.064N, 25.460E]: adults, 26.06.2018. Larvae in mines on Rosa, 12.07.2019, Rearing 142/2019. Larvae in mines on Rosa, 07.08.2019, Rearing 376/2019. Adults, 06.06.2020. Larvae in mines on Rosa, 06.07.2020, Rearing: 30/2020. Ostrobottnia kajanensis, Kuhmo Ulvinsalo, YKJ grid coordinates: 7103:3665 [63.992°N, 30.382°E]: about 20 mines on Rubus chamaemorus, of which ca 8 inhabited, with one larva in alcohol and four females emerged, collected 10.08.2019, Rearing: 400/2019. Fenusa ewaldi was described from two females reared from leaf-mines on Rosa collected in Novosibirsk, West Siberia (Smith and Altenhofer 2011). Also reared from Rosa by Ponomarev (2022): Russia, Moscow oblast, Voynovo-Gora, 55.846°N, 39.063°E, 10.07.2020. Since at least 2018, leaf-mines of Fenusa ewaldi have been frequently found on Rosa, e.g. R. woodsii and R. acicularis in Oulu Botanic Garden, and numerous females reared (Fig. 36). At these three localities, cultivated Rosa species in parks and gardens were the hosts (Fig. 37A). Mines were found mostly on more sheltered rose bushes. The rearing of four females from leaf-mines on Rubus chamaemorus (above) was therefore unexpected, considering the habitat was shady, moist, natural old-growth mixed forest dominated by spruce (Fig. 37B)., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Smith, DR, Altenhofer, E, 2011. The second record of a sawfly leafminer on Rosa, a new species of Fenusa (Hymenoptera: Tenthredinidae). Proceedings of the Entomological Society of Washington 113 (1): 57 - 60, DOI: https://doi.org/10.4289/0013-8797.113.1.57","Ponomarev, AG, 2022. Babochki i zhuki Moscovskoj oblasti. Published on the Internet. http://insectamo.ru"]}

Dolerus incisus Goulet, 1986 Fig. 20A-C Notes. Finland: Regio kuusamoensis, Kuusamo Siikauopaja, YKJ grid coordinates: 7362:3612 [66.334°N, 29.511°E], 2♀ (ZMUO.035155, ZMUO.035156), 12.06.2018. 1♀ (ZMUO.045265), 20.06.2020. All specimens leg. M. Mutanen. ZMUO. First record in the Palaearctic. Recorded previously only in boreal North America: Alaska, Alberta, Yukon, and North West Territories (Goulet 1986). The genetic and morphological affinity of D. incisus to D. junci and D. yukonensis suggests that the host plant of D. incisus is also a species of Equisetum (Goulet 1986). The habitat in Finland is a shallow oxbow lake (meander) of the Oulanka river with plenty of Equisetum fluviatile, which is also the host of D. junci. This habitat matches perfectly with that given for D. incisus in North America by Goulet (1986). Several other Dolerus species occur syntopically, including the closely related D. junci Stephens (= D. cothurnatus auct., see above). The melanic form of D. incisus is recorded from Finland (all three are females, see above, Fig. 20A) and Norway (Heimdalsmunnen, alt 1150 m, leg. E. Heibo, CEH). Melanic females of D. subarcticus (recorded from Norway and Kamtschatka, see under D. subarcticus) can be distinguished by their postocellar furrows, the sawsheath (valvula 3) shape, and its apical setation (Fig. 20B). Melanic males can be separated most reliably by their penis valves (Fig. 20C, 20D). Only the melanic colour form of D. incisus has so far been recognized from Fennoscandian countries, whereas in Canada and the USA both the black color form and a form with red-banded abdomen and partly reddish legs are known. We studied paratypes of D. incisus: 2♀, 2♂, some of them dissected, CNC., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080, {"references":["Goulet, H, 1986. The genera and species of the Nearctic Dolerini (Symphyta: Tenthredinidae: Selandriinae): classification and phylogeny. Memoirs of the Entomological Society of Canada 135(S135): 1-208. https://doi.org/10.4039/entm118135fv"]}

Dolerus subarcticus Hellén, 1956 Fig. 20D Dolerus subarcticus Hellén, 1956: 99-100. ♀, ♂. Syntypes. Published type locality: 16 localities in Finnish Lapland, Russian Karelia and Kola Peninsula, and one locality in Siberia (Krasnojarsk oblast, Jenisseisk). Lectotype designated below. Dolerus willoughbyi Benson, 1956: 55-58. ♀, ♂. Holotype. Type locality: Abisko (Sweden). Type material examined and taxonomic notes. Dolerus subarcticus Hellén, 1956. Lectotype ♀ hereby designated, labelled (printed on pale paper if not stated otherwise): "Fennia, Li. Utsjoki, Outakoski, 29.6.1947 [type locality], leg. Hellén” [printed], " Lectotypus 2014, Dolerus subarcticus, ♀ Hellén, 1956, M.Heidemaa design." [red], " Dolerus subarcticus Hellén, 1956, M.Heidemaa det." [white]. In good condition. 10♀, 8♂ paralectotypes also examined: Ivalo, Kantalaks, Kilpisjärvi, Lutto, Muonio, Paanajärvi, Petsamo, Umba, Utsjoki (Outakoski). All in MZH. Dolerus willoughbyi Benson, 1956. 2♀, 2♂ paratypes studied. BMNH. The melanic (black) form (♂) and a nearly black form, with obscure reddish patches on abdominal terga 2-4 (♀), are known from Norway (near Hovet and near Geitryggtunnelen, leg. E. Heibo, CEH), and the melanic form also from Kamtschatka, Russia (a syntype female of D. pratensis var. totus determined by R. Malaise, in NHRS, see also under D. yukonensis)., Published as part of Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia & Prous, Marko, 2022, Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta), pp. 151-218 in Deutsche Entomologische Zeitschrift 69 (2) on page 151, DOI: 10.3897/dez.69.84080