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Licneremaeus sp. Records in Ecuador: Galápagos. Habitat: Litoral, arid, transition and moist zone. References: Schatz (1998): 385. Remarks: Licneremaeus sp. is included here, as it represents the first record of this genus from Ecuador., Published as part of Revelo-Tobar, Harol, 2022, Checklist of Oribatid mites (Acari: Oribatida) of Ecuador, pp. 1-96 in Zootaxa 5210 (1) on page 70, DOI: 10.11646/zootaxa.5210.1.1, http://zenodo.org/record/7345248, {"references":["Schatz, H. (1998) Oribatid mites of the Galapagos Islands - Faunistics, ecology and speciation. Experimental and Applied Acarology, 22, 373 - 409. https: // doi. org / 10.1023 / A: 1006097928124"]}

Licneremaeus prodigiosus Schuster, 1958 Licneremaeus prodigiosus Schuster 1958, p. 223, fig. 1. Parco Naturale delle Alpi Marittime: PAL, PRATO single records. General distribution: Austrian Alps; Central, South, South-East Europe; southwestern Palaearctic; Mediterranean. New record for Italy. Habitat preferences: silvicolous; xerophilous. Remarks: Body length 180���200 ��m. A comparison with material from the ���Hundsheimer Berge��� (Lower Austria, near the site of the original description, Schatz & Fischer 2007) exhibits the species identity of the specimens from Cuneo. Licneremaeus giustii Bernini, 1973, known from the Eolian Islands, southern Italy, is morphologically very similar, especially in the pattern of the notogastral areolae. Morphological differences between L. prodigiosus and L. giustii are given in the posterior arrangement of these areolae, also in the number of notogastral setae (12 pairs in L. prodigiosus, 13 pairs in L. giustii), and in the length of the adanal setae (very short in L. prodigiosus, longer in L. giustii) (Bernini 1973). The specimens from Cuneo correspond in all mentioned features to L. prodigiosus, although the notogastral areolae are weakly expressed in the present specimens., Published as part of Schatz, Heinrich, Fortini, Lorenzo, Fusco, Tommaso, Casale, Francesca, Jacomini, Carlo & Giulio, Andrea Di, 2021, Oribatid mites (Acari, Oribatida) from " Parco Naturale delle Alpi Marittime " (Piedmont, Italy), pp. 501-540 in Zootaxa 5082 (6) on page 525, DOI: 10.11646/zootaxa.5082.6.1, http://zenodo.org/record/5797624, {"references":["Schuster, R. (1958) Beitrag zur Kenntnis der Milbenfauna (Oribatei) in pannonischen Trockenboden. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Abteilung I, Wien, 167, 221 - 235.","Schatz, H. & Fischer B. M. (2007) Hornmilben (Acari: Oribatida) von den Hundsheimer Bergen (Niederosterreich, Osterreich). Berichte des naturwissenschaftlich-medizinischen Vereins in Innsbruck, 94, 63 - 77.","Bernini, F. (1973) Notulae Oribatologicae VII. Gli Oribatei (Acarida) dell'isolotto di Basiluzzo (Isole Eolie). Lavori della Societa Italiana di Biogeografia, Nuova Serie, 3 (1972), 355 - 480. https: // doi. org / 10.21426 / B 63110608"]}

Key to the adults of known species of Licneremaeus of the world Bomfim et al. (2017) provided a key to the genus Licneremaeus but this key shows a minor mistake. They includ-ed L. novus along with the group of mites having 13 pairs of notogastral setae, but they have 12 pairs of notogastral setae only. Therefore, we corrected this mistake and updated the key with the present newly described species. 1. Notogaster with 12 or 13 pairs of setae, c 2 absent............................................................ 2 - Notogaster 14 pairs of setae, c 2 present [Brazil]......................... L. braziliensis Bomfim, Silva & Oliveira, 2017 2. Notogaster with 12 pairs of setae......................................................................... 3 - Notogaster with 13 pairs of setae......................................................................... 5 3. Single areola between the field delimited by the bases of setae c 1; single areola between the field delimited by c 1 and da.... 4 - Two areolae between the field delimited by the bases of seate c 1 ; two areolae between the field delimited by c 1 and da [Caucasia, Portugal, Spain]..................................................... L. novus Karppinen & Shtanchaeva, 1987 4. Three closely placed areolae between the field delimited by the bases of dm and da; no reticulations on prodorsum; ad 1 postanal [Austria, Spain]............................................................ L. prodigiosus Schuster, 1958 - Single areola between the field delimited by the bases of dm and da; prodorsum with reticulations; ad 1 para-anal [Fiji, Vietnam, Madagascar].................................................................. L. polygonalis Hammer, 1971 5. Areolae present on notogaster........................................................................... 6 - Areolae absent on notogaster [Chile]........................................... L. altiplanicus Covarrubias, 2005 6. Notogaster with more than 32 areolae..................................................................... 7 - Notogaster with less than 32 areolae..................................................................... 10 7. Transverse bump present on prodorsum.................................................................... 8 - Transverse bump absent on prodorsum [Brazil, Cuba].......................... L. cubanus Balogh & Mahunka, 1980 8. Sensillus clavate; no areolae between/anterior to the bases of setae da [Venezuela]........... L. exornatus Grandjean, 1931 - Sensillus flattened; areolae present between or anterior to the bases of setae da.................................... 9 9. Transverse ridge connecting the bases of ro present; two contiguous areolae between the setae c 1 [Italy]................................................................................................... L. giustii Bernini, 1973 - Transverse ridge connecting the bases of ro absent; single areolae between setae c 1 [Tunisia].. L. embeyisztini Mahunka, 1980 10. Number of areolae on notogaster ≥ 20.................................................................... 11 - Number of areolae on notogaster ≤ 17.................................................................... 12 11. Discoidal sensillus [Brazil, Guatemala]............................................ L. discoidalis Willmann, 1930 - Fusiform sensillus [Brazil, Japan, Komodo, New Guinea, Peru]......................... L. novaeguineae Balogh, 1968 12. Median areola present between the field delimited by the bases of c 1 and da...................................... 13 - Median areola sabsent between the field delimited by the bases of c 1 and da...................................... 16 13. Areolae present on ventral plate; curved hump present at the level of dp......................................... 14 - Areolae absent on ventral plate; curved hump at the level of dp present or absent.................................. 15 14. 14 pairs of areolae on notgaster; areolae present between the field delimited by the bases of da and dm; sensillus clavate [India].................................................................................. L. indicus sp.nov. - 10 pairs of areolae on notogaster; areolae between the field delimited by the bases of da and dm absent; sensillus flattened [England, USA, Vietnam]....................................................... L. licnophorus (Michael, 1882) 15. Curved hump present at the level of dp; areolae between the field delimited by the bases of da and dm present [Java]..................................................................................... L. linieatus Hammer, 1979 - Curved hump absent at the level of dp; areolae between the field delimited by the bases of da and dm absent [Mauritius]............................................................................. L. semiareolatus Mahunka, 1978 16. Median crista present between c 1 and da; bases of setae le connected by a transverse bump [Tanzania]................................................................................................. L. cristatus Mahunka, 1984 - No median crista between c 1 and da; bases of setae le not connected by transverse bump [Ethiopia]................................................................................................. L. costulatus Mahunka, 1982, Published as part of Arun, A. & Ramani, N., 2020, Two remarkable new species of oribatid mites (Acari: Oribatida) from Acacia litter of Kerala, India, pp. 539-558 in Zootaxa 4877 (3) on page 556, DOI: 10.11646/zootaxa.4877.3.7, http://zenodo.org/record/4570503, {"references":["Bomfim, G. S., Silva, J. G. & Oliveira, A. R. (2017) A new species of Licneremaeus (Oribatida: Licneremaeidae) from Brazil, with an identification key to the known species of the genus. Systematic and Applied Acarology, 22, 1872 - 1883. https: // doi. org / 10.11158 / saa. 22.11.7","Karppinen, E. & Shtanchaeva, U. Ya. (1987) New oribatid mites (Acarina, Oribatei) from the Caucasus area. Annales Entomologici Fennici, 53, 61 - 65.","Hammer, M. (1971) On some oribatids from Viti Levu, Fiji Islands. Biologiske Skrifter udgivet af Det Kongelige Danske Videnskabernes Selskab, 16 (6), 1 - 60.","Covarrubias, R. (2005) A new Altiplanic species from Chile: Licneremaeus altiplanicus n. sp. (Acari, Oribatida, Licneremaeidae). Acta Entomologica Chilena, 29 (1), 7 - 12.","Balogh, J. & Mahunka, S. (1980) New data to the knowledge of the oribatid fauna of Neogaea (Acari) V. Acta Zoologica Academiae Scientiarum Hungaricae, 26 (1 - 3), 21 - 59.","Grandjean, F. (1931) Le genre Licneremaeus Paoli (Acariens). Bulletin de la Societe Zoologique de France, 56, 221 - 250.","Bernini, F. (1973) Notulae Oribatologicae VII. Gli Oribatei (Acarida) dell'isolotto di Basiluzzo (Isole Eolie). Lavori Della Societa Italiana Di Biogeografia, 3, 355 - 480. https: // doi. org / 10.21426 / B 63110608","Mahunka, S. (1980) Oribatids from Tunisian soils (Acari: Oribatida). I. Folia Entomologica Hungarica, 41 (1), 123 - 124.","Willmann, C. (1930) Neue Oribatiden aus Guatemala. Zoologischer Anzeiger, 88 (9 - 10), 239 - 246.","Balogh, J. (1968) New oribatids (Acari) from New Guinea. Acta Zoologica Academiae Scientiarum Hungaricae, 14 (3 - 4), 259 - 285.","Michael, A. D. (1882) Further notes on British Oribatidae. Journal of the Royal Microscopical Society, 2, 1 - 18. https: // doi. org / 10.1111 / j. 1365 - 2818.1882. tb 00175. x","Hammer, M. (1979) Investigations on the oribatid fauna of Java. Biologiske Skrifter udgivet af Det Kongelige Danske Videnskabernes Selskab, 22 (9), 1 - 79.","Mahunka, S. (1978) Neue und interessante Milben aus dem Genfer Museum. XXVII. A first survey of the oribatid (Acari) fauna of Mauritius, Reunion and the Seychelles I. Revue Suisse de Zoologie, 85 (1), 177 - 236. https: // doi. org / 10.5962 / bhl. part. 82228","Mahunka, S. (1984) Oribatids from eastern part of the Ethiopian region (Acari). V. Acta Zoologica Hungarica, 30 (1 - 2), 87 - 136.","Mahunka, S. (1982) Oribatids from eastern part of the Ethiopian region (Acari). I. Acta Zoologica Academiae Scientiarum Hungaricae, 28 (3 - 4), 293 - 336."]}

Licneremaeus indicus sp. nov. (Figs. 14–28) Diagnosis. Rostral tectum without areolae. Sensillus discoidal with fine barbs. Notogaster with elevations or bumps. At most 14 areolae present on the notogaster in dorsal view. Thirteen pairs of notogastral setae present. Porose areae Aa and A 2 present. Pygidium at the posterior end of notogaster. Epimeral setation: 2-1-3-1. Ventral plate fully areolate. Measurements. Body length 101 (holotype), 98–103 (15 paratypes); body width 59 (holotype), 58–62 (15 paratypes). Integument. Cuticle greyish brown in color. Body covered with abundant cerotegument in the form of small rounded granules. Microareolar patterns seen on notogaster, other than the 14 areolae. Prodorsum (Figs. 14, 20–21, 23). Rostrum pointed apically. Lamellar (le, 8), rostral (ro, 10) and interlamellar (in, 8) setae simple, covered with granules/cerotegument and slightly curved. Length of setae le and ro shorter than the distance between the bases of their counterparts. Seta in inserted near to dorsosejugal suture, below the level of anterior tip of the notogaster. Bothridial cup widely open at the postero-lateral borders of prodorsum, sensillus (ss, 18), with clavate head and a short stalk, bearing thick, blunt spines. Transverse hump present in front of anterior most tip of dorsosejugal suture. Prodorsal surface bears chitinous lines. Seta le arises from the transverse ridge joining, two strong chitinous lines. Pedotectum I present, pedotectum II absent. Notogaster (Figs. 14, 20, 22, 24). Dorsosejugal suture highly convex, extending beyond interlamellar region and incomplete, being interrupted medially, towards the anteriormost portion. Notogaster cuneate anteriorly and oval posteriorly. Thirteen pairs of small, smooth, slightly curved setae present on notogaster, often covered with cerotegument giving a leaf like appearance. Setal length vary from 4–8 as the following; c 3, da (4); c 1, dm, dp, la (5); p 1, p 2, h 1, h 2, h 3 (6); lm (7); lp (8); c 2 and p 3 absent. Lyrifissures im, ip and ips present, im more or less horizontally placed between lm and lp; ip and ips visible in ventral view. Porose area Aa present closer to seta la; A 2 located posteriorly, above the level of seta h 2. An array of cuticular elevations and bumps present on notogaster. The first bump located at the region between setae da and c 1, surrounding an areolae; the second bump central in position, located between setae da and dm and it extends laterally on either sides, towards seta lm; the third bump located below the second one, curved and U-shaped, extending posteriorly, to slightly below the level of seta dp, encompassing an areolae. It extends laterally also on either side, as a transverse bump towards the insertion of seta lp. Three areolae present, between the field delimited by the bases of dm, la and lm. Apart from the above three areolae encompassed by the bumps, a single areola detected anteromedially on the notogaster, between setae c 1, of either side. Posterior border of notogaster produced in to a pygidium. Gnathosoma (Figs. 15, 25). Chelicera (20) with small fixed and movable digits, seta cha (8) small and smooth, chb inconspicuous. Cheliceral teeth feebly developed. Trägardh’s organ (Tg) narrowly triangular. Mentum and submentum with wrinkles. Subcapitular setae h (5), m (6), a (4), or (2) smooth. Rutellum with three weakly developed dendrites. Palpal (24) surface striated. Coxisternal region (Figs. 15, 26). With six pairs of short, smooth, and tapered epimeral setae (1a, 1b, 2a, 3a, 3b, and 4b), of approximately equal length. Epimeral setal formula 2-1-2-1 (4a absent). Epimeres I and II fused together and epimeres III and IV fused together and the fused epimeres delimited by apodemes. Coxisternal area margined by wrinkled ridges. Median depression seen on the fused epimeral plates, I and II as well as III and IV. Ventral plate (Figs. 15, 27–28). Covered with polygonal sculptures, except the regions adjacent to the genital and anal apertures and the region between these two plates. Anterior border of genital plates broader than the posterior border and the anterior margin of each genital plate located next to the apodeme delimiting the epimeres III/IV. Genital plates bear less prominent wrinkles and five pairs of small, smooth setae of length ranging from 2–6; g 1 (6), g 2 (4), g 3 (3), g 4 (4) and g 5 (3). Aggenital seta (ag) absent, anal plates broader postero-laterally, slightly below the median level. Two pairs of anal setae (an 1 & an 2, 2–3) and two pairs of adanal setae (ad 1 & ad 2, 5–7) present, all setae small, smooth, seta ad 3 absent, ad 1 located posterior to the anal plate, ad 2 inserted laterally, slightly below the middle level of anal plate. Integument of anal plate not smooth, but with feebly developed elevated areas. Legs (Figs. 16–19). All legs tridactylous, with the median claw thicker than the lateral ones. Claws bear small spines, set dorsally and with longitudinal striations. Femora of all legs with wrinkles, other segments smooth. Chaetotaxy of legs including solenidia: leg I: 1-3-3(1)-3(1)-14(1), leg II: 1-3-1(1)-2(1)-12(1), leg III: 2-3-1-2-10, and leg IV: 1-2-0-3-10 (see Table 2). LegTrochanterFemurGenuTibiaTarsusI v’ d, bv ”, v’ l’, (v), σ l’, (v), φ (ft), (tc), (it), (p), (u), (a), (pv), ωII v ’ d, bv ”, v ’ v ’’, σ l ’, (v) ft ’, (tc), (it), (p), (u), (a), pv ’, ωIII l ’, v ’ d, bv ”, v ’ l ’ l ’, v ’’ (ft), tc ’, (it), (p), (u), pv ’ IV v’ bv”, v’ - l’, (v) (ft), tc’, (it), (p), (u), pv’ Materials examined. Holotype and 15 paratypes collected from densely accumulated litter samples of A. auriculiformis, at the Calicut University Campus (11° 8’ 6.468’’ N 75° 53’ 18.5028’’ E), Malappuram (Dt.), Kerala, India, on 1.iv.2019, coll. Arun, A. Type deposition. The holotype and ten paratypes are deposited in the collections of Zoological Survey of India, Western Ghat Regional Centre, Kozhikode, Kerala, India; five paratypes are kept in the Acarology Collections, Division of Acarology, Department of Zoology, University of Calicut, Malappuram, Kerala, India. Etymology. The species epithet, “ indicus ” is derived from the name of country from where the type specimens were collected, duly considering the status as being the first species of the Family Licneremaeidae, described from India. Ecology. Large number of adults and various life stages of Licneremaeus indicus sp. nov. were found to inhabit the litter samples of A. auriculiformis. Hundreds of specimens were found to occupy all layers of litter samples, indicating their possible role in the degradation of highly recalcitrant Acacia litter. Remarks. Licneremaeus indicus sp. nov., differs from all its congeners by the possession of 14 areolae on the notogaster, clavate sensillus, presence of three notogastral bumps encompassing one areola each and by the presence of cuticular ridges on the rostral tectum between setae le and ro. The new species shows most resemblance to the oriental species, L. licnophorus in the number of notogastral setae, presence of curved bump at the level of seta dp, and polygonal sculptures on the ventral plate. However, it is distinct from L. licnophorus in the presence of areolae between the field delimited by the bases of setae da and dm, clavate nature of sensillus (flattened sensillus in L. licnophorus) and presence of pygidium on notogaster (absent in L. licnophorus).

Licneremaeus indicus sp. nov. (Figs. 14���28) Diagnosis. Rostral tectum without areolae. Sensillus discoidal with fine barbs. Notogaster with elevations or bumps. At most 14 areolae present on the notogaster in dorsal view. Thirteen pairs of notogastral setae present. Porose areae Aa and A 2 present. Pygidium at the posterior end of notogaster. Epimeral setation: 2-1-3-1. Ventral plate fully areolate. Measurements. Body length 101 (holotype), 98���103 (15 paratypes); body width 59 (holotype), 58���62 (15 paratypes). Integument. Cuticle greyish brown in color. Body covered with abundant cerotegument in the form of small rounded granules. Microareolar patterns seen on notogaster, other than the 14 areolae. Prodorsum (Figs. 14, 20���21, 23). Rostrum pointed apically. Lamellar (le, 8), rostral (ro, 10) and interlamellar (in, 8) setae simple, covered with granules/cerotegument and slightly curved. Length of setae le and ro shorter than the distance between the bases of their counterparts. Seta in inserted near to dorsosejugal suture, below the level of anterior tip of the notogaster. Bothridial cup widely open at the postero-lateral borders of prodorsum, sensillus (ss, 18), with clavate head and a short stalk, bearing thick, blunt spines. Transverse hump present in front of anterior most tip of dorsosejugal suture. Prodorsal surface bears chitinous lines. Seta le arises from the transverse ridge joining, two strong chitinous lines. Pedotectum I present, pedotectum II absent. Notogaster (Figs. 14, 20, 22, 24). Dorsosejugal suture highly convex, extending beyond interlamellar region and incomplete, being interrupted medially, towards the anteriormost portion. Notogaster cuneate anteriorly and oval posteriorly. Thirteen pairs of small, smooth, slightly curved setae present on notogaster, often covered with cerotegument giving a leaf like appearance. Setal length vary from 4���8 as the following; c 3, da (4); c 1, dm, dp, la (5); p 1, p 2, h 1, h 2, h 3 (6); lm (7); lp (8); c 2 and p 3 absent. Lyrifissures im, ip and ips present, im more or less horizontally placed between lm and lp; ip and ips visible in ventral view. Porose area Aa present closer to seta la; A 2 located posteriorly, above the level of seta h 2. An array of cuticular elevations and bumps present on notogaster. The first bump located at the region between setae da and c 1, surrounding an areolae; the second bump central in position, located between setae da and dm and it extends laterally on either sides, towards seta lm; the third bump located below the second one, curved and U-shaped, extending posteriorly, to slightly below the level of seta dp, encompassing an areolae. It extends laterally also on either side, as a transverse bump towards the insertion of seta lp. Three areolae present, between the field delimited by the bases of dm, la and lm. Apart from the above three areolae encompassed by the bumps, a single areola detected anteromedially on the notogaster, between setae c 1, of either side. Posterior border of notogaster produced in to a pygidium. Gnathosoma (Figs. 15, 25). Chelicera (20) with small fixed and movable digits, seta cha (8) small and smooth, chb inconspicuous. Cheliceral teeth feebly developed. Tr��gardh���s organ (Tg) narrowly triangular. Mentum and submentum with wrinkles. Subcapitular setae h (5), m (6), a (4), or (2) smooth. Rutellum with three weakly developed dendrites. Palpal (24) surface striated. Coxisternal region (Figs. 15, 26). With six pairs of short, smooth, and tapered epimeral setae (1a, 1b, 2a, 3a, 3b, and 4b), of approximately equal length. Epimeral setal formula 2-1-2-1 (4a absent). Epimeres I and II fused together and epimeres III and IV fused together and the fused epimeres delimited by apodemes. Coxisternal area margined by wrinkled ridges. Median depression seen on the fused epimeral plates, I and II as well as III and IV. Ventral plate (Figs. 15, 27���28). Covered with polygonal sculptures, except the regions adjacent to the genital and anal apertures and the region between these two plates. Anterior border of genital plates broader than the posterior border and the anterior margin of each genital plate located next to the apodeme delimiting the epimeres III/IV. Genital plates bear less prominent wrinkles and five pairs of small, smooth setae of length ranging from 2���6; g 1 (6), g 2 (4), g 3 (3), g 4 (4) and g 5 (3). Aggenital seta (ag) absent, anal plates broader postero-laterally, slightly below the median level. Two pairs of anal setae (an 1 & an 2, 2���3) and two pairs of adanal setae (ad 1 & ad 2, 5���7) present, all setae small, smooth, seta ad 3 absent, ad 1 located posterior to the anal plate, ad 2 inserted laterally, slightly below the middle level of anal plate. Integument of anal plate not smooth, but with feebly developed elevated areas. Legs (Figs. 16���19). All legs tridactylous, with the median claw thicker than the lateral ones. Claws bear small spines, set dorsally and with longitudinal striations. Femora of all legs with wrinkles, other segments smooth. Chaetotaxy of legs including solenidia: leg I: 1-3-3(1)-3(1)-14(1), leg II: 1-3-1(1)-2(1)-12(1), leg III: 2-3-1-2-10, and leg IV: 1-2-0-3-10 (see Table 2). LegTrochanterFemurGenuTibiaTarsusI v��� d, bv ���, v��� l���, (v), �� l���, (v), �� (ft), (tc), (it), (p), (u), (a), (pv), ��II v ��� d, bv ���, v ��� v ������, �� l ���, (v) ft ���, (tc), (it), (p), (u), (a), pv ���, ��III l ���, v ��� d, bv ���, v ��� l ��� l ���, v ������ (ft), tc ���, (it), (p), (u), pv ��� IV v��� bv���, v��� - l���, (v) (ft), tc���, (it), (p), (u), pv��� Materials examined. Holotype and 15 paratypes collected from densely accumulated litter samples of A. auriculiformis, at the Calicut University Campus (11�� 8��� 6.468������ N 75�� 53��� 18.5028������ E), Malappuram (Dt.), Kerala, India, on 1.iv.2019, coll. Arun, A. Type deposition. The holotype and ten paratypes are deposited in the collections of Zoological Survey of India, Western Ghat Regional Centre, Kozhikode, Kerala, India; five paratypes are kept in the Acarology Collections, Division of Acarology, Department of Zoology, University of Calicut, Malappuram, Kerala, India. Etymology. The species epithet, ��� indicus ��� is derived from the name of country from where the type specimens were collected, duly considering the status as being the first species of the Family Licneremaeidae, described from India. Ecology. Large number of adults and various life stages of Licneremaeus indicus sp. nov. were found to inhabit the litter samples of A. auriculiformis. Hundreds of specimens were found to occupy all layers of litter samples, indicating their possible role in the degradation of highly recalcitrant Acacia litter. Remarks. Licneremaeus indicus sp. nov., differs from all its congeners by the possession of 14 areolae on the notogaster, clavate sensillus, presence of three notogastral bumps encompassing one areola each and by the presence of cuticular ridges on the rostral tectum between setae le and ro. The new species shows most resemblance to the oriental species, L. licnophorus in the number of notogastral setae, presence of curved bump at the level of seta dp, and polygonal sculptures on the ventral plate. However, it is distinct from L. licnophorus in the presence of areolae between the field delimited by the bases of setae da and dm, clavate nature of sensillus (flattened sensillus in L. licnophorus) and presence of pygidium on notogaster (absent in L. licnophorus)., Published as part of Arun, A. & Ramani, N., 2020, Two remarkable new species of oribatid mites (Acari: Oribatida) from Acacia litter of Kerala, India, pp. 539-558 in Zootaxa 4877 (3) on pages 550-556, DOI: 10.11646/zootaxa.4877.3.7, http://zenodo.org/record/4570503

Licneremaeus discoidalis Willmann, 1930 Northern South America. Brazil: State of Par��. Reference: Ferreira et al. (2012): 13., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 60, DOI: 10.5281/zenodo.437584, {"references":["Ferreira, R. N. C., Franklin, E., Souza, J. L. P. & Moraes, J. (2012) Soil oribatid mite (Acari: Oribatida) diversity and composition in semi-deciduous forest fragments in eastern Amazonia and comparison with the surrounding savanna matrix. Journal of Natural History, 46, 1 - 14. http: // dx. doi. org / 10.1080 / 00222933.2012.707245"]}

Licneremaeus cubanus Balogh & Mahunka, 1980 Neotropical region. Brazil: State of Par��. References: Franklin et al. (2006): 1013, Santos et al. (2008): 734., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 60, DOI: 10.5281/zenodo.437584, {"references":["Franklin, E., Santos, E. M. & Albuquerque, M. I. (2006) Diversity and distribution of oribatid mites (Acari: Oribatida) in a lowland rain forest in Peru and in several environments of the Brazilians states of Amazonas, Rondonia, Roraima and Para. Brazilian Journal of Biology, 66 (4), 999 - 1020.","Santos, E. M. R., Franklin, E. & Magnusson, W. E. (2008) Cost-efficiency of Subsampling Protocols to Evaluate Oribatid-Mite Communities in an Amazonian Savanna. Biotropica, 40 (6), 728 - 735. http: // dx. doi. org / 10.1111 / j. 1744 - 7429.2008.00425. x"]}

Licneremaeus licnophorus (Michael, 1882) Holarctic and tropical regions. Brazil: State of Rio de Janeiro. Reference: Ermilov & Tolstikov (2015f): 50., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 61, DOI: 10.5281/zenodo.437584, {"references":["Ermilov, S. G. & Tolstikov, A. V. (2015 f) Report on oribatid mites (Acari, Oribatida) from Brazil, with description of a new species of the genus Papillacarus. Acarina, 23 (1), 49 - 54."]}

Licneremaeus prodigiosus Schuster, 1958 Distribution in Georgia. W: Batumi Botanical Garden, Sataplia Reserve; E: Khashuri (Shtanchaeva & Sub��as 2010) Global distribution. Palaearctic Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 56, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Licneremaeus licnophorus (Michael, 1882) Distribution in Georgia. W: Ritsa Reserve, Musera; E: Khashuri (Shtanchaeva & Subías 2010) Global distribution. Holarctic Ecology. Moss, tree bark Licneremaeus novus Karppinen & Shtanchaeva, 1987 Distribution in Georgia. W: Ritsa Reserve, Saken (Shtanchaeva & Subías 2010) Global distribution. Mediterranean Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 56, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Licneremaeus sp. Iran localities. Mazandaran province (Barimani & Kamali 1998).

Licneremaeus prodigiosus SCHUSTER, 1958 Z u r n d o r f e r E i c h e n w a l d: vereinzelt (2 Individuen). H a b i t a t b i n d u n g: inausgesprochenenxerothermenStandorten, Trockenwiesen. A l l g e m e i n e V e r b r e i t u n g: In Österreich in Trockenrasen in Kärnten (PERLINGER & SCHATZ 2009) und am Alpen-Ostrand gefunden: Burgenland: Seewinkel (locus typicus im Boden eines Trockenwäldchens nahe Illmitz, SCHUSTER 1958, 1959a, 1959b), Niederösterreich: Hundsheimer Berge (SCHATZ & FISCHER 2007). Ungarn: mehrere Fundorte in West- und Zentralungarn (MAHUNKA & MAHUNKA- PAPP 2004). Südliche Paläarktis: Süd-, Südosteuropa, Krim, Kazakhstan; "südliche Art", selten., Published as part of Schatz, H. & Hofstätter, S., 2014, Hornmilben (Acari, Oribatida) aus dem Europa-Schutzgebiet Zurndorfer Eichenwald (Burgenland, Österreich), pp. 1775-1793 in Linzer biologische Beiträge 46 (2) on page 1785, DOI: 10.5281/zenodo.5312338, {"references":["SCHUSTER R. (1958): Beitrag zur Kenntnis der Milbenfauna (Oribatei) in pannonischen Trockenboden. - Sitz. ber. Osterr. Akad. Wiss., Mathem. - naturwiss. Kl., Abt. I, Wien 167: 221 - 235.","SCHUSTER R. (1959 a): Okologisch-faunistische Untersuchungen an bodenbewohnenden Kleinarthropoden (speziell Oribatiden), des Salzlachengebietes im Seewinkel. - Sitz. ber. Osterr. Akad. Wiss., Mathem. - naturwiss. Kl., Abt. I, Wien 168 (1): 27 - 78.","SCHUSTER R. (1959 b): Der Indikationswert von Bodenmilben (Oribatei) fur die tiergeographische Beurteilung des Alpen-Ostrandes. - Verh. Dtsch. Zool. Ges., Munster / Westfalen: 363 - 369."]}

Licneremaeus Paoli, 1908 Typ: Notaspis licnophorus Michael, 1882 Im Bearbeitungsgebiet nur eine Art: [Licneremaeus licnophorus (Michael, 1882)], Published as part of Weigmann, G. & Miko, L., 2006, Hornmilben (Oribatida) [pages 323 to 417], pp. 323-417 in Hornmilben (Oribatida) [Dahl, Tierwelt Deutschlands, Teil 76], Keltern :Goecke & Evers on page 333

Licneremaeus Paoli, 1908 Typ: Notaspis licnophorus Michael, 1882 Im Bearbeitungsgebiet nur eine Art: [Licneremaeus licnophorus (Michael, 1882)]

Licneremaeus licnophorus (Michael, 1882) [177a-c] Diagnose: Dorsalborsten m����ig kurz, fein; Cos vor NG-Rand mit Querbalken; hinter Lenticulus eine runde Macula, NG-Mitte zur Seite durch Bogenlinien begrenzt, vorn seitlich und hinten median gro��fl��chig eingebeult, mit grobknotigem Cerotegument; 13 ng, 2 Paar Ap. klein, unscheinbar; VP lateral mit Netzstruktur; neben G mit spitzem Zahn, der einem runden Zahn am Hinterrand der Epi 4 gegen��ber steht. K��rperl��nge 200-210 ��m. Syn., Tax.: Notaspis licnophorus Michael, 1882; Michael 1888. Eremaeus l.: Berlese 1896 (AMS). Licneremaeus l.: Paoli 1908 (B); Perez-Inigo 1993 (B). ��kologie: In Moos, an Baumst��mmen. Verbreitung: Pal��arktis., Published as part of Weigmann, G. & Miko, L., 2006, Hornmilben (Oribatida) [pages 323 to 417], pp. 323-417 in Hornmilben (Oribatida) [Dahl, Tierwelt Deutschlands, Teil 76], Keltern :Goecke & Evers on page 333

Licneremaeus licnophorus (Michael, 1882) Bestimmung nach SELLNICK (1928:33), WILLMANN (1931:126) L��nge 205-230 ��m, L��nge:Breite 1,65-1,80 (8 Ex.) Belegmaterial: Stadtwald Ettlingen, Moderbuchenwald, Streu am Fu�� eines Buchenstammes, XI/ 1988I. WUNDERLE leg., 8 Ex., LNK A0200. [[Einzelfund in der Bodenstreu im Oktober 1980]]., Published as part of Beck, L. & Woas, S., 1991, Die Oribatiden-Arten (Acari) eines suedwestdeutschen Buchenwaldes I., pp. 37-82 in carolinea 49 on page 63