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The present checklist of the oribatid mites compiles all known published records of Ecuador with 479 species belonging to 232 genera and 89 families; 180 of these species were described from Ecuador. Oribatid mites were found in 15 of the 24 provinces that conform the Ecuadorian territory. In the provinces of Zamora Chinchipe, Galápagos, Loja, Cotopaxi, Pichincha and Napo, 95% of the species were recorded. 23.6% of these species are known only from Ecuador, 47.8% are distributed in Neotropical region, the remaining are Pantropical, semicosmopolitan and cosmopolitan. This high number of species makes Ecuador the country with the second largest number of species records in Latin America after Brazil.

Licneremaeus sp. Records in Ecuador: Galápagos. Habitat: Litoral, arid, transition and moist zone. References: Schatz (1998): 385. Remarks: Licneremaeus sp. is included here, as it represents the first record of this genus from Ecuador., Published as part of Revelo-Tobar, Harol, 2022, Checklist of Oribatid mites (Acari: Oribatida) of Ecuador, pp. 1-96 in Zootaxa 5210 (1) on page 70, DOI: 10.11646/zootaxa.5210.1.1, http://zenodo.org/record/7345248, {"references":["Schatz, H. (1998) Oribatid mites of the Galapagos Islands - Faunistics, ecology and speciation. Experimental and Applied Acarology, 22, 373 - 409. https: // doi. org / 10.1023 / A: 1006097928124"]}

This paper presents a list of 78 species/subspecies of 50 genera and 27 families of soil-inhabiting oribatid mites (Oribatida) collected on Sibuyan Island, Philippines; of these, two species and one genus are recorded in the Oriental region for the first time, and 14 species/subspecies and two genera are recorded in the Philippine fauna for the first time. Trachyoribates insularis sp. n. (Haplozetidae) is described on the basis of adults.

Licneremaeus prodigiosus Schuster, 1958 Licneremaeus prodigiosus Schuster 1958, p. 223, fig. 1. Parco Naturale delle Alpi Marittime: PAL, PRATO single records. General distribution: Austrian Alps; Central, South, South-East Europe; southwestern Palaearctic; Mediterranean. New record for Italy. Habitat preferences: silvicolous; xerophilous. Remarks: Body length 180���200 ��m. A comparison with material from the ���Hundsheimer Berge��� (Lower Austria, near the site of the original description, Schatz & Fischer 2007) exhibits the species identity of the specimens from Cuneo. Licneremaeus giustii Bernini, 1973, known from the Eolian Islands, southern Italy, is morphologically very similar, especially in the pattern of the notogastral areolae. Morphological differences between L. prodigiosus and L. giustii are given in the posterior arrangement of these areolae, also in the number of notogastral setae (12 pairs in L. prodigiosus, 13 pairs in L. giustii), and in the length of the adanal setae (very short in L. prodigiosus, longer in L. giustii) (Bernini 1973). The specimens from Cuneo correspond in all mentioned features to L. prodigiosus, although the notogastral areolae are weakly expressed in the present specimens., Published as part of Schatz, Heinrich, Fortini, Lorenzo, Fusco, Tommaso, Casale, Francesca, Jacomini, Carlo & Giulio, Andrea Di, 2021, Oribatid mites (Acari, Oribatida) from " Parco Naturale delle Alpi Marittime " (Piedmont, Italy), pp. 501-540 in Zootaxa 5082 (6) on page 525, DOI: 10.11646/zootaxa.5082.6.1, http://zenodo.org/record/5797624, {"references":["Schuster, R. (1958) Beitrag zur Kenntnis der Milbenfauna (Oribatei) in pannonischen Trockenboden. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Abteilung I, Wien, 167, 221 - 235.","Schatz, H. & Fischer B. M. (2007) Hornmilben (Acari: Oribatida) von den Hundsheimer Bergen (Niederosterreich, Osterreich). Berichte des naturwissenschaftlich-medizinischen Vereins in Innsbruck, 94, 63 - 77.","Bernini, F. (1973) Notulae Oribatologicae VII. Gli Oribatei (Acarida) dell'isolotto di Basiluzzo (Isole Eolie). Lavori della Societa Italiana di Biogeografia, Nuova Serie, 3 (1972), 355 - 480. https: // doi. org / 10.21426 / B 63110608"]}

Oribatid mites were investigated in six sites in the Alpi Marittime Natural Park (Prov. Cuneo, Piedmont, northern Italy) which is part of the NATURA 2000 network. The selected locations cover 4 forested (fir, ash, 2 beech forests) and 2 grassland sites (mountain meadow, xeric grassland). Altogether 173 described oribatid species belonging to 51 families were encountered, complemented by one undetermined species (Scheloribates (Topobates) cf. carpathicus), added together 174 species. Among them 11 species are new records for Italy. A comparison with other Italian alpine provinces shows high similarity of the oribatid species composition from the investigated sites with the provinces of Bolzano, Trento, Sondrio. A total of 37 species from the Marittime Alps has also been recorded in the southern part of the nearby Swiss canton Ticino. The majority of the species in the investigated sites are known as silvicolous; xerophilous species were mainly found in the xeric grassland site at lower altitude. High species similarities are observed between the forested sites, which also show high species numbers (up to 96 spp. found in the fir forest), lowest similarities are found between the open grassland sites and the forested sites, as well as between the grassland sites themselves.

Key to the adults of known species of Licneremaeus of the world Bomfim et al. (2017) provided a key to the genus Licneremaeus but this key shows a minor mistake. They includ-ed L. novus along with the group of mites having 13 pairs of notogastral setae, but they have 12 pairs of notogastral setae only. Therefore, we corrected this mistake and updated the key with the present newly described species. 1. Notogaster with 12 or 13 pairs of setae, c 2 absent............................................................ 2 - Notogaster 14 pairs of setae, c 2 present [Brazil]......................... L. braziliensis Bomfim, Silva & Oliveira, 2017 2. Notogaster with 12 pairs of setae......................................................................... 3 - Notogaster with 13 pairs of setae......................................................................... 5 3. Single areola between the field delimited by the bases of setae c 1; single areola between the field delimited by c 1 and da.... 4 - Two areolae between the field delimited by the bases of seate c 1 ; two areolae between the field delimited by c 1 and da [Caucasia, Portugal, Spain]..................................................... L. novus Karppinen & Shtanchaeva, 1987 4. Three closely placed areolae between the field delimited by the bases of dm and da; no reticulations on prodorsum; ad 1 postanal [Austria, Spain]............................................................ L. prodigiosus Schuster, 1958 - Single areola between the field delimited by the bases of dm and da; prodorsum with reticulations; ad 1 para-anal [Fiji, Vietnam, Madagascar].................................................................. L. polygonalis Hammer, 1971 5. Areolae present on notogaster........................................................................... 6 - Areolae absent on notogaster [Chile]........................................... L. altiplanicus Covarrubias, 2005 6. Notogaster with more than 32 areolae..................................................................... 7 - Notogaster with less than 32 areolae..................................................................... 10 7. Transverse bump present on prodorsum.................................................................... 8 - Transverse bump absent on prodorsum [Brazil, Cuba].......................... L. cubanus Balogh & Mahunka, 1980 8. Sensillus clavate; no areolae between/anterior to the bases of setae da [Venezuela]........... L. exornatus Grandjean, 1931 - Sensillus flattened; areolae present between or anterior to the bases of setae da.................................... 9 9. Transverse ridge connecting the bases of ro present; two contiguous areolae between the setae c 1 [Italy]................................................................................................... L. giustii Bernini, 1973 - Transverse ridge connecting the bases of ro absent; single areolae between setae c 1 [Tunisia].. L. embeyisztini Mahunka, 1980 10. Number of areolae on notogaster ≥ 20.................................................................... 11 - Number of areolae on notogaster ≤ 17.................................................................... 12 11. Discoidal sensillus [Brazil, Guatemala]............................................ L. discoidalis Willmann, 1930 - Fusiform sensillus [Brazil, Japan, Komodo, New Guinea, Peru]......................... L. novaeguineae Balogh, 1968 12. Median areola present between the field delimited by the bases of c 1 and da...................................... 13 - Median areola sabsent between the field delimited by the bases of c 1 and da...................................... 16 13. Areolae present on ventral plate; curved hump present at the level of dp......................................... 14 - Areolae absent on ventral plate; curved hump at the level of dp present or absent.................................. 15 14. 14 pairs of areolae on notgaster; areolae present between the field delimited by the bases of da and dm; sensillus clavate [India].................................................................................. L. indicus sp.nov. - 10 pairs of areolae on notogaster; areolae between the field delimited by the bases of da and dm absent; sensillus flattened [England, USA, Vietnam]....................................................... L. licnophorus (Michael, 1882) 15. Curved hump present at the level of dp; areolae between the field delimited by the bases of da and dm present [Java]..................................................................................... L. linieatus Hammer, 1979 - Curved hump absent at the level of dp; areolae between the field delimited by the bases of da and dm absent [Mauritius]............................................................................. L. semiareolatus Mahunka, 1978 16. Median crista present between c 1 and da; bases of setae le connected by a transverse bump [Tanzania]................................................................................................. L. cristatus Mahunka, 1984 - No median crista between c 1 and da; bases of setae le not connected by transverse bump [Ethiopia]................................................................................................. L. costulatus Mahunka, 1982, Published as part of Arun, A. & Ramani, N., 2020, Two remarkable new species of oribatid mites (Acari: Oribatida) from Acacia litter of Kerala, India, pp. 539-558 in Zootaxa 4877 (3) on page 556, DOI: 10.11646/zootaxa.4877.3.7, http://zenodo.org/record/4570503, {"references":["Bomfim, G. S., Silva, J. G. & Oliveira, A. R. (2017) A new species of Licneremaeus (Oribatida: Licneremaeidae) from Brazil, with an identification key to the known species of the genus. Systematic and Applied Acarology, 22, 1872 - 1883. https: // doi. org / 10.11158 / saa. 22.11.7","Karppinen, E. & Shtanchaeva, U. Ya. (1987) New oribatid mites (Acarina, Oribatei) from the Caucasus area. Annales Entomologici Fennici, 53, 61 - 65.","Hammer, M. (1971) On some oribatids from Viti Levu, Fiji Islands. Biologiske Skrifter udgivet af Det Kongelige Danske Videnskabernes Selskab, 16 (6), 1 - 60.","Covarrubias, R. (2005) A new Altiplanic species from Chile: Licneremaeus altiplanicus n. sp. (Acari, Oribatida, Licneremaeidae). Acta Entomologica Chilena, 29 (1), 7 - 12.","Balogh, J. & Mahunka, S. (1980) New data to the knowledge of the oribatid fauna of Neogaea (Acari) V. Acta Zoologica Academiae Scientiarum Hungaricae, 26 (1 - 3), 21 - 59.","Grandjean, F. (1931) Le genre Licneremaeus Paoli (Acariens). Bulletin de la Societe Zoologique de France, 56, 221 - 250.","Bernini, F. (1973) Notulae Oribatologicae VII. Gli Oribatei (Acarida) dell'isolotto di Basiluzzo (Isole Eolie). Lavori Della Societa Italiana Di Biogeografia, 3, 355 - 480. https: // doi. org / 10.21426 / B 63110608","Mahunka, S. (1980) Oribatids from Tunisian soils (Acari: Oribatida). I. Folia Entomologica Hungarica, 41 (1), 123 - 124.","Willmann, C. (1930) Neue Oribatiden aus Guatemala. Zoologischer Anzeiger, 88 (9 - 10), 239 - 246.","Balogh, J. (1968) New oribatids (Acari) from New Guinea. Acta Zoologica Academiae Scientiarum Hungaricae, 14 (3 - 4), 259 - 285.","Michael, A. D. (1882) Further notes on British Oribatidae. Journal of the Royal Microscopical Society, 2, 1 - 18. https: // doi. org / 10.1111 / j. 1365 - 2818.1882. tb 00175. x","Hammer, M. (1979) Investigations on the oribatid fauna of Java. Biologiske Skrifter udgivet af Det Kongelige Danske Videnskabernes Selskab, 22 (9), 1 - 79.","Mahunka, S. (1978) Neue und interessante Milben aus dem Genfer Museum. XXVII. A first survey of the oribatid (Acari) fauna of Mauritius, Reunion and the Seychelles I. Revue Suisse de Zoologie, 85 (1), 177 - 236. https: // doi. org / 10.5962 / bhl. part. 82228","Mahunka, S. (1984) Oribatids from eastern part of the Ethiopian region (Acari). V. Acta Zoologica Hungarica, 30 (1 - 2), 87 - 136.","Mahunka, S. (1982) Oribatids from eastern part of the Ethiopian region (Acari). I. Acta Zoologica Academiae Scientiarum Hungaricae, 28 (3 - 4), 293 - 336."]}

Licneremaeus indicus sp. nov. (Figs. 14–28) Diagnosis. Rostral tectum without areolae. Sensillus discoidal with fine barbs. Notogaster with elevations or bumps. At most 14 areolae present on the notogaster in dorsal view. Thirteen pairs of notogastral setae present. Porose areae Aa and A 2 present. Pygidium at the posterior end of notogaster. Epimeral setation: 2-1-3-1. Ventral plate fully areolate. Measurements. Body length 101 (holotype), 98–103 (15 paratypes); body width 59 (holotype), 58–62 (15 paratypes). Integument. Cuticle greyish brown in color. Body covered with abundant cerotegument in the form of small rounded granules. Microareolar patterns seen on notogaster, other than the 14 areolae. Prodorsum (Figs. 14, 20–21, 23). Rostrum pointed apically. Lamellar (le, 8), rostral (ro, 10) and interlamellar (in, 8) setae simple, covered with granules/cerotegument and slightly curved. Length of setae le and ro shorter than the distance between the bases of their counterparts. Seta in inserted near to dorsosejugal suture, below the level of anterior tip of the notogaster. Bothridial cup widely open at the postero-lateral borders of prodorsum, sensillus (ss, 18), with clavate head and a short stalk, bearing thick, blunt spines. Transverse hump present in front of anterior most tip of dorsosejugal suture. Prodorsal surface bears chitinous lines. Seta le arises from the transverse ridge joining, two strong chitinous lines. Pedotectum I present, pedotectum II absent. Notogaster (Figs. 14, 20, 22, 24). Dorsosejugal suture highly convex, extending beyond interlamellar region and incomplete, being interrupted medially, towards the anteriormost portion. Notogaster cuneate anteriorly and oval posteriorly. Thirteen pairs of small, smooth, slightly curved setae present on notogaster, often covered with cerotegument giving a leaf like appearance. Setal length vary from 4–8 as the following; c 3, da (4); c 1, dm, dp, la (5); p 1, p 2, h 1, h 2, h 3 (6); lm (7); lp (8); c 2 and p 3 absent. Lyrifissures im, ip and ips present, im more or less horizontally placed between lm and lp; ip and ips visible in ventral view. Porose area Aa present closer to seta la; A 2 located posteriorly, above the level of seta h 2. An array of cuticular elevations and bumps present on notogaster. The first bump located at the region between setae da and c 1, surrounding an areolae; the second bump central in position, located between setae da and dm and it extends laterally on either sides, towards seta lm; the third bump located below the second one, curved and U-shaped, extending posteriorly, to slightly below the level of seta dp, encompassing an areolae. It extends laterally also on either side, as a transverse bump towards the insertion of seta lp. Three areolae present, between the field delimited by the bases of dm, la and lm. Apart from the above three areolae encompassed by the bumps, a single areola detected anteromedially on the notogaster, between setae c 1, of either side. Posterior border of notogaster produced in to a pygidium. Gnathosoma (Figs. 15, 25). Chelicera (20) with small fixed and movable digits, seta cha (8) small and smooth, chb inconspicuous. Cheliceral teeth feebly developed. Trägardh’s organ (Tg) narrowly triangular. Mentum and submentum with wrinkles. Subcapitular setae h (5), m (6), a (4), or (2) smooth. Rutellum with three weakly developed dendrites. Palpal (24) surface striated. Coxisternal region (Figs. 15, 26). With six pairs of short, smooth, and tapered epimeral setae (1a, 1b, 2a, 3a, 3b, and 4b), of approximately equal length. Epimeral setal formula 2-1-2-1 (4a absent). Epimeres I and II fused together and epimeres III and IV fused together and the fused epimeres delimited by apodemes. Coxisternal area margined by wrinkled ridges. Median depression seen on the fused epimeral plates, I and II as well as III and IV. Ventral plate (Figs. 15, 27–28). Covered with polygonal sculptures, except the regions adjacent to the genital and anal apertures and the region between these two plates. Anterior border of genital plates broader than the posterior border and the anterior margin of each genital plate located next to the apodeme delimiting the epimeres III/IV. Genital plates bear less prominent wrinkles and five pairs of small, smooth setae of length ranging from 2–6; g 1 (6), g 2 (4), g 3 (3), g 4 (4) and g 5 (3). Aggenital seta (ag) absent, anal plates broader postero-laterally, slightly below the median level. Two pairs of anal setae (an 1 & an 2, 2–3) and two pairs of adanal setae (ad 1 & ad 2, 5–7) present, all setae small, smooth, seta ad 3 absent, ad 1 located posterior to the anal plate, ad 2 inserted laterally, slightly below the middle level of anal plate. Integument of anal plate not smooth, but with feebly developed elevated areas. Legs (Figs. 16–19). All legs tridactylous, with the median claw thicker than the lateral ones. Claws bear small spines, set dorsally and with longitudinal striations. Femora of all legs with wrinkles, other segments smooth. Chaetotaxy of legs including solenidia: leg I: 1-3-3(1)-3(1)-14(1), leg II: 1-3-1(1)-2(1)-12(1), leg III: 2-3-1-2-10, and leg IV: 1-2-0-3-10 (see Table 2). LegTrochanterFemurGenuTibiaTarsusI v’ d, bv ”, v’ l’, (v), σ l’, (v), φ (ft), (tc), (it), (p), (u), (a), (pv), ωII v ’ d, bv ”, v ’ v ’’, σ l ’, (v) ft ’, (tc), (it), (p), (u), (a), pv ’, ωIII l ’, v ’ d, bv ”, v ’ l ’ l ’, v ’’ (ft), tc ’, (it), (p), (u), pv ’ IV v’ bv”, v’ - l’, (v) (ft), tc’, (it), (p), (u), pv’ Materials examined. Holotype and 15 paratypes collected from densely accumulated litter samples of A. auriculiformis, at the Calicut University Campus (11° 8’ 6.468’’ N 75° 53’ 18.5028’’ E), Malappuram (Dt.), Kerala, India, on 1.iv.2019, coll. Arun, A. Type deposition. The holotype and ten paratypes are deposited in the collections of Zoological Survey of India, Western Ghat Regional Centre, Kozhikode, Kerala, India; five paratypes are kept in the Acarology Collections, Division of Acarology, Department of Zoology, University of Calicut, Malappuram, Kerala, India. Etymology. The species epithet, “ indicus ” is derived from the name of country from where the type specimens were collected, duly considering the status as being the first species of the Family Licneremaeidae, described from India. Ecology. Large number of adults and various life stages of Licneremaeus indicus sp. nov. were found to inhabit the litter samples of A. auriculiformis. Hundreds of specimens were found to occupy all layers of litter samples, indicating their possible role in the degradation of highly recalcitrant Acacia litter. Remarks. Licneremaeus indicus sp. nov., differs from all its congeners by the possession of 14 areolae on the notogaster, clavate sensillus, presence of three notogastral bumps encompassing one areola each and by the presence of cuticular ridges on the rostral tectum between setae le and ro. The new species shows most resemblance to the oriental species, L. licnophorus in the number of notogastral setae, presence of curved bump at the level of seta dp, and polygonal sculptures on the ventral plate. However, it is distinct from L. licnophorus in the presence of areolae between the field delimited by the bases of setae da and dm, clavate nature of sensillus (flattened sensillus in L. licnophorus) and presence of pygidium on notogaster (absent in L. licnophorus).

Two new oribatid mite species viz. Papillacarus (Vepracarus) acaciensis sp. nov. and Licneremaeus indicus sp. nov. belonging to the respective oribatid families, Lohmanniidae and Licneremaeidae are described and illustrated. Specimens of both species were collected from litter of Acacia auriculiformis Benth. (Leguminosae) growing in different localities of the Calicut University Campus, Malappuram Dt. of Kerala. The family Licneremaeidae is recorded for the first time from India. Identification keys to all known species of the nominative subgenus Vepracarus and the genus Licneremaeus are also provided.

Licneremaeus indicus sp. nov. (Figs. 14���28) Diagnosis. Rostral tectum without areolae. Sensillus discoidal with fine barbs. Notogaster with elevations or bumps. At most 14 areolae present on the notogaster in dorsal view. Thirteen pairs of notogastral setae present. Porose areae Aa and A 2 present. Pygidium at the posterior end of notogaster. Epimeral setation: 2-1-3-1. Ventral plate fully areolate. Measurements. Body length 101 (holotype), 98���103 (15 paratypes); body width 59 (holotype), 58���62 (15 paratypes). Integument. Cuticle greyish brown in color. Body covered with abundant cerotegument in the form of small rounded granules. Microareolar patterns seen on notogaster, other than the 14 areolae. Prodorsum (Figs. 14, 20���21, 23). Rostrum pointed apically. Lamellar (le, 8), rostral (ro, 10) and interlamellar (in, 8) setae simple, covered with granules/cerotegument and slightly curved. Length of setae le and ro shorter than the distance between the bases of their counterparts. Seta in inserted near to dorsosejugal suture, below the level of anterior tip of the notogaster. Bothridial cup widely open at the postero-lateral borders of prodorsum, sensillus (ss, 18), with clavate head and a short stalk, bearing thick, blunt spines. Transverse hump present in front of anterior most tip of dorsosejugal suture. Prodorsal surface bears chitinous lines. Seta le arises from the transverse ridge joining, two strong chitinous lines. Pedotectum I present, pedotectum II absent. Notogaster (Figs. 14, 20, 22, 24). Dorsosejugal suture highly convex, extending beyond interlamellar region and incomplete, being interrupted medially, towards the anteriormost portion. Notogaster cuneate anteriorly and oval posteriorly. Thirteen pairs of small, smooth, slightly curved setae present on notogaster, often covered with cerotegument giving a leaf like appearance. Setal length vary from 4���8 as the following; c 3, da (4); c 1, dm, dp, la (5); p 1, p 2, h 1, h 2, h 3 (6); lm (7); lp (8); c 2 and p 3 absent. Lyrifissures im, ip and ips present, im more or less horizontally placed between lm and lp; ip and ips visible in ventral view. Porose area Aa present closer to seta la; A 2 located posteriorly, above the level of seta h 2. An array of cuticular elevations and bumps present on notogaster. The first bump located at the region between setae da and c 1, surrounding an areolae; the second bump central in position, located between setae da and dm and it extends laterally on either sides, towards seta lm; the third bump located below the second one, curved and U-shaped, extending posteriorly, to slightly below the level of seta dp, encompassing an areolae. It extends laterally also on either side, as a transverse bump towards the insertion of seta lp. Three areolae present, between the field delimited by the bases of dm, la and lm. Apart from the above three areolae encompassed by the bumps, a single areola detected anteromedially on the notogaster, between setae c 1, of either side. Posterior border of notogaster produced in to a pygidium. Gnathosoma (Figs. 15, 25). Chelicera (20) with small fixed and movable digits, seta cha (8) small and smooth, chb inconspicuous. Cheliceral teeth feebly developed. Tr��gardh���s organ (Tg) narrowly triangular. Mentum and submentum with wrinkles. Subcapitular setae h (5), m (6), a (4), or (2) smooth. Rutellum with three weakly developed dendrites. Palpal (24) surface striated. Coxisternal region (Figs. 15, 26). With six pairs of short, smooth, and tapered epimeral setae (1a, 1b, 2a, 3a, 3b, and 4b), of approximately equal length. Epimeral setal formula 2-1-2-1 (4a absent). Epimeres I and II fused together and epimeres III and IV fused together and the fused epimeres delimited by apodemes. Coxisternal area margined by wrinkled ridges. Median depression seen on the fused epimeral plates, I and II as well as III and IV. Ventral plate (Figs. 15, 27���28). Covered with polygonal sculptures, except the regions adjacent to the genital and anal apertures and the region between these two plates. Anterior border of genital plates broader than the posterior border and the anterior margin of each genital plate located next to the apodeme delimiting the epimeres III/IV. Genital plates bear less prominent wrinkles and five pairs of small, smooth setae of length ranging from 2���6; g 1 (6), g 2 (4), g 3 (3), g 4 (4) and g 5 (3). Aggenital seta (ag) absent, anal plates broader postero-laterally, slightly below the median level. Two pairs of anal setae (an 1 & an 2, 2���3) and two pairs of adanal setae (ad 1 & ad 2, 5���7) present, all setae small, smooth, seta ad 3 absent, ad 1 located posterior to the anal plate, ad 2 inserted laterally, slightly below the middle level of anal plate. Integument of anal plate not smooth, but with feebly developed elevated areas. Legs (Figs. 16���19). All legs tridactylous, with the median claw thicker than the lateral ones. Claws bear small spines, set dorsally and with longitudinal striations. Femora of all legs with wrinkles, other segments smooth. Chaetotaxy of legs including solenidia: leg I: 1-3-3(1)-3(1)-14(1), leg II: 1-3-1(1)-2(1)-12(1), leg III: 2-3-1-2-10, and leg IV: 1-2-0-3-10 (see Table 2). LegTrochanterFemurGenuTibiaTarsusI v��� d, bv ���, v��� l���, (v), �� l���, (v), �� (ft), (tc), (it), (p), (u), (a), (pv), ��II v ��� d, bv ���, v ��� v ������, �� l ���, (v) ft ���, (tc), (it), (p), (u), (a), pv ���, ��III l ���, v ��� d, bv ���, v ��� l ��� l ���, v ������ (ft), tc ���, (it), (p), (u), pv ��� IV v��� bv���, v��� - l���, (v) (ft), tc���, (it), (p), (u), pv��� Materials examined. Holotype and 15 paratypes collected from densely accumulated litter samples of A. auriculiformis, at the Calicut University Campus (11�� 8��� 6.468������ N 75�� 53��� 18.5028������ E), Malappuram (Dt.), Kerala, India, on 1.iv.2019, coll. Arun, A. Type deposition. The holotype and ten paratypes are deposited in the collections of Zoological Survey of India, Western Ghat Regional Centre, Kozhikode, Kerala, India; five paratypes are kept in the Acarology Collections, Division of Acarology, Department of Zoology, University of Calicut, Malappuram, Kerala, India. Etymology. The species epithet, ��� indicus ��� is derived from the name of country from where the type specimens were collected, duly considering the status as being the first species of the Family Licneremaeidae, described from India. Ecology. Large number of adults and various life stages of Licneremaeus indicus sp. nov. were found to inhabit the litter samples of A. auriculiformis. Hundreds of specimens were found to occupy all layers of litter samples, indicating their possible role in the degradation of highly recalcitrant Acacia litter. Remarks. Licneremaeus indicus sp. nov., differs from all its congeners by the possession of 14 areolae on the notogaster, clavate sensillus, presence of three notogastral bumps encompassing one areola each and by the presence of cuticular ridges on the rostral tectum between setae le and ro. The new species shows most resemblance to the oriental species, L. licnophorus in the number of notogastral setae, presence of curved bump at the level of seta dp, and polygonal sculptures on the ventral plate. However, it is distinct from L. licnophorus in the presence of areolae between the field delimited by the bases of setae da and dm, clavate nature of sensillus (flattened sensillus in L. licnophorus) and presence of pygidium on notogaster (absent in L. licnophorus)., Published as part of Arun, A. & Ramani, N., 2020, Two remarkable new species of oribatid mites (Acari: Oribatida) from Acacia litter of Kerala, India, pp. 539-558 in Zootaxa 4877 (3) on pages 550-556, DOI: 10.11646/zootaxa.4877.3.7, http://zenodo.org/record/4570503

Murvanidze, Maka, Cilbircioğlu, Cihan, Özdemir, Esengül, Inak, Emre (2020): Checklist of oribatid mites (Acari: Oribatida) of the Central Black Sea basin of Turkey with new records for the country. Persian Journal of Acarology 9 (3): 255-277, DOI: 10.22073/pja.v9i3.61909, URL: https://www.mendeley.com/catalogue/ea74271c-b6cc-3222-8dc5-fe9f0a25fa45/

This checklist of the oribatid fauna of Canada and Alaska (excluding Astigmata) includes 580 identified species in 249 genera and 96 families. The known fauna of Canada includes 556 identified species in 247 genera, and that of Alaska includes 182 species in 95 genera; 39 of the 42 oribatid superfamilies are represented. We further list ~ 300 species that are currently unidentified, and possibly undescribed. In addition, we list 42 genera that are represented only by unidentified and probably undescribed species. For each species we give combinations and synonymies, specific locations in Alaska and the Provinces and Territories of Canada, habitats, and biogeography. There are 182 identified species known for Alaska, 152 for Yukon, 122 for Northwest Territories, 58 for Nunavut; 210 for British Columbia, 213 for Alberta, 15 for Saskatchewan, 84 for Manitoba, 167 for Ontario, 210 for Québec, 110 for Nova Scotia, 77 for New Brunswick, 84 for Newfoundland and 6 for Prince Edward Island. The known fauna of Canada is smaller than that of Austria, and is approximately equivalent to that of the Czech Republic. As these countries are much smaller in size than Canada and less ecologically diverse, we consider the Canadian and Alaskan fauna are at most 25% known. The paucity of these data reflects the absence of taxonomic and faunistic studies on Oribatida in State, Provinces or Territories, and especially in the Canadian and Alaskan National Park systems and the hundreds of Provincial Parks. Despite the almost 90% increase in described species since the catalogue of Marshall et al. (1987), there is a need for focussed, coordinated research on Oribatida in the natural regions throughout Canada and Alaska, and for monographs on families and genera with large numbers of undescribed species, such as Brachychthoniidae, Damaeidae, Cepheidae, Liacaridae, Oppiidae, Suctobelbidae, Hydrozetidae, Phenopelopidae, Scheloribatidae, Haplozetidae and Galumnidae.

A checklist of the oribatid mite species reported in Brazil is presented, including all published records up to 2015. A total of 576 described species in 206 genera and 83 families is presented. Information includes the names by which each species was reported in the Brazilian literature, its general known distribution and by Brazilian States, references, and remarks, when needed. As with most countries, there was a slow early accumulation of knowledge but in recent decades the pace of description has been relatively high. A graphical overview of the number of described oribatid mite species from Brazil in different decades is given. The proportion contributed by each of the major oribatid groups is generally similar to that of the overall world fauna, with a composition that reflects the South American fauna and all of the Neotropics in general. There is a relatively low percentage of primitive mites (Palaeosomata, Enarthronota) other than Lohmanniidae and Mesoplophoridae, which are quite diverse. The Brachypylina comprises about 68% of the oribatid mite fauna. In the checklist, 41% of the species are known only from Brazil, 37% from the Neotropical region, 13.5% have a wider distribution in the global tropical and subtropical regions, and 8.5% are considered cosmopolitan or semicosmopolitan species. The number of descriptions of new species since 2000 from Brazil (73 spp.) and South America (230) is high, but the oribatid mite fauna of these countries remains poorly known. Only continued studies can determine if the high number of species known only from Brazil is an indication of high endemism.

Huilicheremaeus atypicus (Mahunka, 1984) Licneremaeus atypicus Mahunka, 1984 Neotropical region. Brazil: States of Par�� and S��o Paulo. References: Oliveira et al. (2005): 58, Franklin et al. (2006): 1013, Santos et al. (2008): 734., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 60, DOI: 10.5281/zenodo.437584, {"references":["Oliveira, A. R., Norton, R. A. & Moraes, G. J. de (2005) Edaphic and plant inhabiting oribatid mites (Acari: Oribatida) from Cerrado and Mata Atlantica ecosystems in the State of Sao Paulo, southeast Brazil. Zootaxa, 1049, 49 - 68.","Franklin, E., Santos, E. M. & Albuquerque, M. I. (2006) Diversity and distribution of oribatid mites (Acari: Oribatida) in a lowland rain forest in Peru and in several environments of the Brazilians states of Amazonas, Rondonia, Roraima and Para. Brazilian Journal of Biology, 66 (4), 999 - 1020.","Santos, E. M. R., Franklin, E. & Magnusson, W. E. (2008) Cost-efficiency of Subsampling Protocols to Evaluate Oribatid-Mite Communities in an Amazonian Savanna. Biotropica, 40 (6), 728 - 735. http: // dx. doi. org / 10.1111 / j. 1744 - 7429.2008.00425. x"]}

Licneremaeus discoidalis Willmann, 1930 Northern South America. Brazil: State of Par��. Reference: Ferreira et al. (2012): 13., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 60, DOI: 10.5281/zenodo.437584, {"references":["Ferreira, R. N. C., Franklin, E., Souza, J. L. P. & Moraes, J. (2012) Soil oribatid mite (Acari: Oribatida) diversity and composition in semi-deciduous forest fragments in eastern Amazonia and comparison with the surrounding savanna matrix. Journal of Natural History, 46, 1 - 14. http: // dx. doi. org / 10.1080 / 00222933.2012.707245"]}

Licneremaeus cubanus Balogh & Mahunka, 1980 Neotropical region. Brazil: State of Par��. References: Franklin et al. (2006): 1013, Santos et al. (2008): 734., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 60, DOI: 10.5281/zenodo.437584, {"references":["Franklin, E., Santos, E. M. & Albuquerque, M. I. (2006) Diversity and distribution of oribatid mites (Acari: Oribatida) in a lowland rain forest in Peru and in several environments of the Brazilians states of Amazonas, Rondonia, Roraima and Para. Brazilian Journal of Biology, 66 (4), 999 - 1020.","Santos, E. M. R., Franklin, E. & Magnusson, W. E. (2008) Cost-efficiency of Subsampling Protocols to Evaluate Oribatid-Mite Communities in an Amazonian Savanna. Biotropica, 40 (6), 728 - 735. http: // dx. doi. org / 10.1111 / j. 1744 - 7429.2008.00425. x"]}

Licneremaeus licnophorus (Michael, 1882) Holarctic and tropical regions. Brazil: State of Rio de Janeiro. Reference: Ermilov & Tolstikov (2015f): 50., Published as part of Oliveira, Anibal R., Argolo, Poliane S., De, Gilberto J., Norton, Roy A. & Schatz, Heinrich, 2017, A checklist of the oribatid mite species (Acari: Oribatida) of Brazil, pp. 1-89 in Zootaxa 4245 (1) on page 61, DOI: 10.5281/zenodo.437584, {"references":["Ermilov, S. G. & Tolstikov, A. V. (2015 f) Report on oribatid mites (Acari, Oribatida) from Brazil, with description of a new species of the genus Papillacarus. Acarina, 23 (1), 49 - 54."]}

Bolger, Thomas (2017): Oribatid mites (Acari: Oribatida) recorded from Ireland: Catalogue, historical records, species habitats and geographical distribution, combinations, variations and synonyms. Zootaxa 4328 (1): 1-174, DOI: https://doi.org/10.11646/zootaxa.4328.1.1

Family Licneremaeidae Grandjean, 1954 Licneremaeus Paoli, 1908 �� Licneremaeus licnophorus (Michael, 1882) �� Original name: Notaspis licnophorus Michael, 1882. �� In: Journal of the Royal Microscopical Society, (2), 10. �� Historical combinations/variations: Notaspis licnophora (Michael) Ewing, 1917. Irish Reference/s and Habitat/s: Machair fore dunes and Machair fixed (stable) dunes and Machair grassland (Arroyo and Bolger 2011); soil, moss in canopy and moss on soil in oak forests (Arroyo et al. 2012, 2013b). Counties: Clare, Donegal, Galway, Kerry, Laois, Mayo, Roscommon, Sligo and Wicklow., Published as part of Bolger, Thomas, 2017, Oribatid mites (Acari: Oribatida) recorded from Ireland: Catalogue, historical records, species habitats and geographical distribution, combinations, variations and synonyms, pp. 1-174 in Zootaxa 4328 (1) on page 118, DOI: 10.11646/zootaxa.4328.1.1, http://zenodo.org/record/1000543, {"references":["Arroyo, J, Kenny, J. & Bolger, T. (2012) A survey of the Oribatida and Mesostigmata (Acarina) of Irish forests. Bulletin of the Irish Biogeographical Society, 36, 33 - 59.","Arroyo, J., Kenny, J. & Bolger, T. (2013 b) Variation between mite communities in Irish forest types - importance of bark and moss cover in canopy. Pedobiologia, 56, 241 - 250. https: // doi. org / 10.1016 / j. pedobi. 2013.09.003"]}

Licneremaeus prodigiosus Schuster, 1958 Distribution in Georgia. W: Batumi Botanical Garden, Sataplia Reserve; E: Khashuri (Shtanchaeva & Sub��as 2010) Global distribution. Palaearctic Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 56, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Murvanidze, Maka, Mumladze, Levan (2016): Annotated checklist of Georgian oribatid mites. Zootaxa 4089 (1): 1-81, DOI: http://doi.org/10.11646/zootaxa.4089.1.1

Licneremaeus licnophorus (Michael, 1882) Distribution in Georgia. W: Ritsa Reserve, Musera; E: Khashuri (Shtanchaeva & Subías 2010) Global distribution. Holarctic Ecology. Moss, tree bark Licneremaeus novus Karppinen & Shtanchaeva, 1987 Distribution in Georgia. W: Ritsa Reserve, Saken (Shtanchaeva & Subías 2010) Global distribution. Mediterranean Ecology. Forest soils, Published as part of Murvanidze, Maka & Mumladze, Levan, 2016, Annotated checklist of Georgian oribatid mites, pp. 1-81 in Zootaxa 4089 (1) on page 56, DOI: 10.11646/zootaxa.4089.1.1, http://zenodo.org/record/257567

Akrami, Mohammad Ali (2015): An annotated checklist of oribatid mites (Acari: Oribatida) of Iran. Zootaxa 3963 (4): 451-501, DOI: http://dx.doi.org/10.11646/zootaxa.3963.4.1

Licneremaeus sp. Iran localities. Mazandaran province (Barimani & Kamali 1998).

Licneremaeus prodigiosus SCHUSTER, 1958 Z u r n d o r f e r E i c h e n w a l d: vereinzelt (2 Individuen). H a b i t a t b i n d u n g: inausgesprochenenxerothermenStandorten, Trockenwiesen. A l l g e m e i n e V e r b r e i t u n g: In Österreich in Trockenrasen in Kärnten (PERLINGER & SCHATZ 2009) und am Alpen-Ostrand gefunden: Burgenland: Seewinkel (locus typicus im Boden eines Trockenwäldchens nahe Illmitz, SCHUSTER 1958, 1959a, 1959b), Niederösterreich: Hundsheimer Berge (SCHATZ & FISCHER 2007). Ungarn: mehrere Fundorte in West- und Zentralungarn (MAHUNKA & MAHUNKA- PAPP 2004). Südliche Paläarktis: Süd-, Südosteuropa, Krim, Kazakhstan; "südliche Art", selten., Published as part of Schatz, H. & Hofstätter, S., 2014, Hornmilben (Acari, Oribatida) aus dem Europa-Schutzgebiet Zurndorfer Eichenwald (Burgenland, Österreich), pp. 1775-1793 in Linzer biologische Beiträge 46 (2) on page 1785, DOI: 10.5281/zenodo.5312338, {"references":["SCHUSTER R. (1958): Beitrag zur Kenntnis der Milbenfauna (Oribatei) in pannonischen Trockenboden. - Sitz. ber. Osterr. Akad. Wiss., Mathem. - naturwiss. Kl., Abt. I, Wien 167: 221 - 235.","SCHUSTER R. (1959 a): Okologisch-faunistische Untersuchungen an bodenbewohnenden Kleinarthropoden (speziell Oribatiden), des Salzlachengebietes im Seewinkel. - Sitz. ber. Osterr. Akad. Wiss., Mathem. - naturwiss. Kl., Abt. I, Wien 168 (1): 27 - 78.","SCHUSTER R. (1959 b): Der Indikationswert von Bodenmilben (Oribatei) fur die tiergeographische Beurteilung des Alpen-Ostrandes. - Verh. Dtsch. Zool. Ges., Munster / Westfalen: 363 - 369."]}

Schatz, H., Hofstätter, S. (2014): Hornmilben (Acari, Oribatida) aus dem Europa-Schutzgebiet Zurndorfer Eichenwald (Burgenland, Österreich). Linzer biologische Beiträge 46 (2): 1775-1793, DOI: http://doi.org/10.5281/zenodo.5312338, {"references":["BALOGH J. (1943): Magyarorszag Pancelosatkai (Conspectus Oribateorum Hungariae). - Math. Termesz. Koszlem. 39 (5): 1-202.","BAYARTOGTOKH B., SCHATZ H., FISCHER B.M. & I.E. SMELYANSKY (2011): Occurrence of a mediterranean species in Central Europe and Asia, with notes on the generic status and biogeography of Simkinia and Hemileius (Acari: Oribatida). - Acarologia 51 (3): 359- 370.","BRUCKNER A. (1995): Cave-dwelling oribatid mites (Acarina, Cryptostigmata) from East Austria. - Verh. Zool.-Bot. Ges. Osterreich 132: 81-107.","FISCHER B.M., SCHATZ H. & M. MARAUN (2010): Community structure, trophic position and reproductive mode of soil and bark-living oribatid mites in an alpine grassland ecosystem. - Experimental and Applied Acarology 52 (3): 221-237.","FORSSLUND K.H. (1958): Notizen uber Oribatei (Acari) II. - Entomol. Tidskr. 79 (1-2): 75- 86.","FRANZ H. & M. BEIER (1948): Zur Kenntnis der Bodenfauna im pannonischen Klimagebiet Osterreichs. - Ann. nat. Mus. Wien 56: 440-549.","HOPPERGER M. & H. SCHATZ (2013): Hornmilben (Acari, Oribatida) von Castelfeder (Sudtirol, Italien). - Gredleriana 13: 71-98.","KRISPER G. & S. LAZARUS (2014): Bodenzoologische Untersuchungen an zwei Trockenrasen in der Steiermark - Erstnachweise von Hornmilben (Acari, Oribatida). - Mitteilungen des Naturwissenschaftlichen Vereines fur Steiermark 143: 121-130.","MAHUNKA S. (1987): A survey of the oribatids of Kiskunsag National Park (Acari: Oribatida). - In: MAHUNKA S. (ed.): Fauna of Kiskunsag National Park, vol. 2, Akademiai Kiado, Budapest: 346-397.","MAHUNKA S. & L. MAHUNKA- PAPP (2002): Oribatids (Acari: Oribatida) from the Ferto- Hansag National Park (NW Hungary). - The Fauna of the Ferto-Hansag National Park, Budapest: 199-229.","MAHUNKA S. & L. MAHUNKA- PAPP (2004): A catalogue of the Hungarian oribatid mites (Acari: Oribatida). - Pedozoologica Hungarica No 2. Hungarian Natural History Museum, Budapest, 363 pp.","MEYER E. (1994): Bodenzoologische Bestandserhebungen in Agrarlandschaften Osterreichs (Burgenland, Oberosterreich). - Mitt. Osterr. bodenkndl. Ges. 48 (4): 107-125.","MIHELCIC F. (1956): Oribatiden Sudeuropas V. - Zool. Anz. 157: 154-174.","MIHELCIC F. (1957): Die Oribatiden Zentralspaniens (Ein Beitrag zur Faunistik, Okologie und Verbreitung der Hornmilben des sommertrockenen Spaniens). - Verh. zool.-bot. Ges., Wien 97: 14-26.","MIKO L., WEIGMANN G. & R. NANNELLI (1994): Redescription of Protoribates lophotrichus (Berlese, 1904) (Acarina, Oribatida). - Redia 77 (2): 251-258.","PEREZ- INIGO C. (1993): Acari, Oribatei, Poronota. - In: RAMOS A. et al. (eds.): Fauna Iberica. Museo de Ciencias Naturales, Madrid 3: 320 pp.","PEREZ- INIGO C. (1997): Acari. Oribatei. Gymnonota I. - In: RAMOS A. et al. (eds.): Fauna Iberica. Museo de Ciencias Naturales, Madrid 9: 373 pp.","PERLINGER H. & H. SCHATZ (2009): Faunistik der Hornmilben (Acari: Oribatida) in ausgewahlten Trockenstandorten Karntens (Osterreich). - Carinthia II 199/119: 543- 552.","SCHATZ H. (1983): U.-Ordn.: Oribatei, Hornmilben. - Catalogus Faunae Austriae, Wien, Teil IXi: 118 pp.","SCHATZ H. (1995): Hornmilben in Trockenrasenboden des Virgentales (Osttirol, Osterreich) 2. Teil: Faunistik (Acari, Oribatida). - Ber. nat.-med. Ver., Innsbruck 82: 121-144.","SCHATZ H. (2004): Die Hornmilbenfamilie Brachychthoniidae (Acari, Oribatida) in Tirol (Osterreich). - Denisia, Linz 12: 343-355.","SCHATZ H. (2005): Hornmilben (Acari, Oribatida). GEO-Tag der Artenvielfalt 2005 in Tirol - Erhebungen im Naturpark Kaunergrat. - Ber. nat.-med. Ver., Innsbruck 92: 261-264.","SCHATZ H. (2008): Hornmilben (Acari: Oribatida) im Naturpark Schlern - Rosengarten (Sudtirol, Italien). - Gredleriana 8: 219-254.","SCHATZ H. & B.M. FISCHER (2007): Hornmilben (Acari: Oribatida) von den Hundsheimer Bergen (Niederosterreich, Osterreich). - Ber. nat.-med. Ver. Innsbruck 94: 63-77.","SCHATZ H. & B.M. FISCHER (2013): Die Hornmilben (Acari: Oribatida) der Jagdberggemeinden. - In: Naturmonografie Jagdberggemeinden. Dornbirn, inatura Erlebnis Naturschau: 18: 569-580.","SCHATZ H., BEHAN- PELLETIER V.M., OCONNOR B.M. & R.A. NORTON (2011): Suborder Oribatida van der Hammen, 1968. In: Zhang Z.-Q. (ed.): Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. - Zootaxa 3148: 141- 148.","SCHUSTER R. (1958): Beitrag zur Kenntnis der Milbenfauna (Oribatei) in pannonischen Trockenboden. - Sitz.ber. Osterr. Akad. Wiss., Mathem.-naturwiss. Kl., Abt. I, Wien 167: 221-235.","SCHUSTER R. (1959a): Okologisch-faunistische Untersuchungen an bodenbewohnenden Kleinarthropoden (speziell Oribatiden), des Salzlachengebietes im Seewinkel. - Sitz.ber. Osterr. Akad. Wiss., Mathem.-naturwiss. Kl., Abt. I, Wien 168 (1): 27-78.","SCHUSTER R. (1959b): Der Indikationswert von Bodenmilben (Oribatei) fur die tiergeographische Beurteilung des Alpen-Ostrandes. - Verh. Dtsch. Zool. Ges., Munster/Westfalen: 363-369.","SCHWALBE T. (1989): Oppiella signata, eine neue Art der Familie Oppiidae aus dem Osterzgebirge (Acari, Oribatei). - Dtsch. Entomol. Z., N.F. 36 (1-3): 99-101.","STARY J. (2000a): List of oribatid mites (Acari: Oribatida) of the Bohemia, Czech Republic. - Sbornik Prirodovedneho klubu v Uh. Hradisti 5: 129-154. [in Czech.]","STARY J. (2000b): List of oribatid mites (Acari: Oribatida) of the Moravia, Czech Republic. - Sbornik Prirodovedneho klubu v Uh. Hradisti 5: 155-173. [in Czech.]","STARY J. (2006): List of oribatid mites (Aari: Oribatida) of the Slovak Republic. - Sbornik prirodovedneho klubu v Uh. Hradisti 8: 21-38. [in Slovak]","SUBIAS L.S. (2014): Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (excepto fosiles). - (Originally published in Graellsia, 60 (numero extraordinario): 3-305 (2004), actualized pdf in February 2014, 577 pp.)","SUBIAS L.S. & A. ARILLO (2001): Acari, Oribatei, Gymnonota II. Oppioidea. - In: RAMOS A. et al. (eds.): Fauna Iberica. Museo de Ciencias Naturales, Madrid 15: 289 pp.","TARMAN K. (1977): The southern species of the Oribatid fauna in Yugoslavia. - Biol. Vestnik, Ljubljana 25: 63-73. [in Slowenisch]","TOPERCER E. (1980): Synusia and vertical distribution of Oribatei of Devinska Kobyla. - Acta Ecologica 8 (21): 61-94. [in Slowakisch]","WEIGMANN G. (2006): Hornmilben (Oribatida). - Die Tierwelt Deutschlands, 76. Teil. Goecke & Evers, Keltern, 520 pp.","WILLMANN C. (1951): Untersuchungen uber die terrestrische Milbenfauna im pannonischen Klimagebiet Osterreichs. - Sitz.ber. Osterr. Akad. Wiss., Mathem.-naturwiss. Kl., Abt. I, Wien 160: 91-176."]}

Norton, Roy A., Ermilov, Sergey G. (2014): Catalogue and historical overview of juvenile instars of oribatid mites (Acari: Oribatida). Zootaxa 3833 (1): 1-132, DOI: http://dx.doi.org/10.11646/zootaxa.3833.1.1

Fischer, Barbara M., Schatz, Heinrich (2013): Biodiversity of oribatid mites (Acari: Oribatida) along an altitudinal gradient in the Central Alps. Zootaxa 3626 (4): 429-454, DOI: http://dx.doi.org/10.11646/zootaxa.3626.4.2

Licneremaeidae Grandjean, 1931 Licneremaeus licnophorus (Michael, 1882) Distribution: Palaearctic Habitat: in xeric habitats; in moss, also arboricolous Obergurgl area: pine forest at 2050 m (Zirbenwald), Published as part of Fischer, Barbara M. & Schatz, Heinrich, 2013, Biodiversity of oribatid mites (Acari: Oribatida) along an altitudinal gradient in the Central Alps, pp. 429-454 in Zootaxa 3626 (4) on page 444, DOI: 10.11646/zootaxa.3626.4.2, http://zenodo.org/record/216337

Migliorini, Massimo (2009): Oribatid mite (Arachnida: Oribatida) coenoses from SW Sardinia *. Zootaxa 2318: 8-37, DOI: 10.5281/zenodo.192029

LICNEREMAEIDAE Licneremaus licnophorus 1 1 HOL THER

52. Licneremaus licnophorus (Michael, 1882) Licneremaeus licnophorus: P��rez-Inigo 1993, 64, fig. 21 a���b. Material examined. C 75: 24.V. 2006 (1); 18.VII. 2006 (1). Geographic distribution. Holarctic. Notes. Licneremaeus licnophorus is a typical thermophilous element of Mediterranean soils., Published as part of Migliorini, Massimo, 2009, Oribatid mite (Arachnida: Oribatida) coenoses from SW Sardinia *, pp. 8-37 in Zootaxa 2318 on page 24, DOI: 10.5281/zenodo.192029, {"references":["Michael, A. D. (1882) Further notes on British Oribatidae. Journal of the Royal Microscopical Society, 2, 10.","Perez-Inigo, C. (1993) Acari. Oribatei, Poronota. In: Ramos, M. A. et al. (Eds), Fauna Iberica, vol. 3. Museo Nacional de Ciencias Naturales, CSIC, Madrid, 320 pp."]}

LICNEREMAEIDAE Licneremaus licnophorus 1 1 HOL THER, Published as part of Migliorini, Massimo, 2009, Oribatid mite (Arachnida: Oribatida) coenoses from SW Sardinia *, pp. 8-37 in Zootaxa 2318 on page 37, DOI: 10.5281/zenodo.192029

Licneremaeus Paoli, 1908 Typ: Notaspis licnophorus Michael, 1882 Im Bearbeitungsgebiet nur eine Art: [Licneremaeus licnophorus (Michael, 1882)], Published as part of Weigmann, G. & Miko, L., 2006, Hornmilben (Oribatida) [pages 323 to 417], pp. 323-417 in Hornmilben (Oribatida) [Dahl, Tierwelt Deutschlands, Teil 76], Keltern :Goecke & Evers on page 333

Licneremaeidae Grandjean, 1931 Diagnose: Ptm fehlen; mit Lenticulus; Cos vorhanden; ss groß, blattförmig; PtcII fehlt; NG-Vorderrand in der Mitte offen; 13 ng, 2 Paar kleine A.p.; G und AN nahe beieinander, 5 g, ohne ag, 2 an, 2 ad; B 3-krlg.

Licneremaeus Paoli, 1908 Typ: Notaspis licnophorus Michael, 1882 Im Bearbeitungsgebiet nur eine Art: [Licneremaeus licnophorus (Michael, 1882)]

Licneremaeus licnophorus (Michael, 1882) [177a-c] Diagnose: Dorsalborsten m����ig kurz, fein; Cos vor NG-Rand mit Querbalken; hinter Lenticulus eine runde Macula, NG-Mitte zur Seite durch Bogenlinien begrenzt, vorn seitlich und hinten median gro��fl��chig eingebeult, mit grobknotigem Cerotegument; 13 ng, 2 Paar Ap. klein, unscheinbar; VP lateral mit Netzstruktur; neben G mit spitzem Zahn, der einem runden Zahn am Hinterrand der Epi 4 gegen��ber steht. K��rperl��nge 200-210 ��m. Syn., Tax.: Notaspis licnophorus Michael, 1882; Michael 1888. Eremaeus l.: Berlese 1896 (AMS). Licneremaeus l.: Paoli 1908 (B); Perez-Inigo 1993 (B). ��kologie: In Moos, an Baumst��mmen. Verbreitung: Pal��arktis., Published as part of Weigmann, G. & Miko, L., 2006, Hornmilben (Oribatida) [pages 323 to 417], pp. 323-417 in Hornmilben (Oribatida) [Dahl, Tierwelt Deutschlands, Teil 76], Keltern :Goecke & Evers on page 333

Licneremaeidae Grandjean, 1931 Diagnose: Ptm fehlen; mit Lenticulus; Cos vorhanden; ss gro��, blattf��rmig; PtcII fehlt; NG-Vorderrand in der Mitte offen; 13 ng, 2 Paar kleine A.p.; G und AN nahe beieinander, 5 g, ohne ag, 2 an, 2 ad; B 3-krlg., Published as part of Weigmann, G. & Miko, L., 2006, Hornmilben (Oribatida) [pages 323 to 417], pp. 323-417 in Hornmilben (Oribatida) [Dahl, Tierwelt Deutschlands, Teil 76], Keltern :Goecke & Evers on page 333

Licneremaeus licnophorus (Michael, 1882) Bestimmung nach SELLNICK (1928:33), WILLMANN (1931:126) L��nge 205-230 ��m, L��nge:Breite 1,65-1,80 (8 Ex.) Belegmaterial: Stadtwald Ettlingen, Moderbuchenwald, Streu am Fu�� eines Buchenstammes, XI/ 1988I. WUNDERLE leg., 8 Ex., LNK A0200. [[Einzelfund in der Bodenstreu im Oktober 1980]]., Published as part of Beck, L. & Woas, S., 1991, Die Oribatiden-Arten (Acari) eines suedwestdeutschen Buchenwaldes I., pp. 37-82 in carolinea 49 on page 63

Licneremaeidae Vorkommen: Die einzige mitteleurop��ischen Art, Licneremaeus licnophorus (MICHAEL, 1882), wurde bisher nur zweimal in der Bodenstreu gefunden, davon ein Einzelfund im Oktober 1980 und 8 Tiere am Stammfu�� eines Buchenstammes im November 1988., Published as part of Beck, L. & Woas, S., 1991, Die Oribatiden-Arten (Acari) eines suedwestdeutschen Buchenwaldes I., pp. 37-82 in carolinea 49 on page 63

Licneremaeidae Vorkommen: Die einzige mitteleuropäischen Art, Licneremaeus licnophorus (MICHAEL, 1882), wurde bisher nur zweimal in der Bodenstreu gefunden, davon ein Einzelfund im Oktober 1980 und 8 Tiere am Stammfuß eines Buchenstammes im November 1988.