Search

7. Drosera katangensis Taton — Fig. 9 Drosera katangensis Taton (1945c) 309;G.C.C. Gilbert (1951) 550,A.S.Rob. et al. (2017) 1422. — Type: H. Homblé 773 (holo BR), DRC, Lualaba. Plateau de la Manika, Nov. 1912. Caulescent herb. Stems up to 15 cm long, erect, densely glandular (Fig. 9b). Leaves alternate, sparsely dispersed, basal leaves smaller; stipules c. 2 mm long, apex lacerated into setaelike segments; lamina oblong-spathulate, 5–15 by 1.5–5 mm, abaxially pilose (Fig. 9a); petiole 7–20 mm long, densely pilose/ hirsute. Inflorescences cymose, with up to 8 flowers; peduncles 1, up to 20 cm long, erect at the base, densely pilose/hirsute (often recurved hairs); bracts linear to lanceolate, c. 5 mm long, glandular; pedicels 2–3 mm long, pilose. Sepals 5, oblong, c. 8 by 2 mm, apex obtuse, adaxially densely pilose/hirsute. Petals 5, obovate, c. 12 by 4 mm, pink or pinkish. Stamens 5; filaments c. 6 mm long; anthers c. 1.5 mm long, sagittate. Pistil: ovary ovoid; styles 3, bipartite to the base, branches 2.5–3 mm long, ascending. Seeds fusiform, black. Distribution — DRC (Fig. 9c). Habitat & Ecology — Wet plains. Additional specimens (see Fig. 9d for province map). DRC, Haut- Katanga, 63 km au N du Poste de Katshupa, Plateau des Kundelungu, alt. 1685 m, 28 Jan. 1967, F. Malaisse 4986 (BR); Lusinga, Plateau des Kibara, 14 Apr. 1969, S. Lisowski et al. 4659 (BR). Note — So far, this species has been collected only from Katanga Province (DRC), but it is likely to occur in neighbour- ing regions. The herbarium specimens (F. Malaisse 4986 and S. Lisowski et al. 4659) share characters with D. flexicaulis and D. elongata (elongated stem, erect leaves) as well as with D. bequaertii (erect base of the peduncle, hairiness of leaves and peduncle). Since only few specimens have been collected so far, this species is not well studied. The shared characters with D. flexicaulis, D. elongata and D. bequaertii indicate that D. katangensis could be a hybrid of the above mentioned species (Robinson et al. 2017). The specimen Homblé 774 in particular could support that thesis. While three of the four plants are easily identified as D. flexicaulis, one plant looks similar to D. katangensis showing the erect base of the inflorescence and similar leaf shape and arrangement but it has only sparsely pilose petioles and stem. The examination of fresh material and molecular genetic analysis might shed further light on the taxonomic status of D. katangensis., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on page 10, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Taton A. 1945 c. Drosera katangensis. Bulletin du Jardin Botanique de l'Etat a Bruxelles: 309.","Gilbert G. 1951. Droseraceae. In: Flore du Congo Belge et du Ruanda-Urundi. Institut national pour l'etude agronomique du Congo Belge.","Robinson A, Gibson R, Gonella P, et al. 2017. Drosera of the World - volume 3, Latin America, Africa. Redfern Natural History Productions, Poole."]}

8. Drosera madagascariensis DC. — Fig. 10 Drosera madagascariensis DC. (1824) 318; Diels (1906) 98; G.C.C. Gilbert (1951) 554; Keay (1954) 121; J.R. Laundon (1959) 5; Oberm. (1970) 200; J.R. Laundon (1978) 68; Troupin (1978) 307; Akoègn. et al. (2006) 539; A.S.Rob. et al. (2017) 1426. — Type: Lambert s.n. (holo G-DC), Madagascar, 1819. Drosera congolana Taton (1945b) 310; G.C.C. Gilbert (1951) 552. — Type: J. Bequaert 7191 (holo BR), DRC, Leopoldville, 29 Mar. 1915 Perennial, caulescent herb. Stems up to 25 cm long, rarely acau- lescent (see Note), c. 1 mm diam, only apical part of the stem self-supporting, glabrous or sparsely pilose (Fig. 10b). Leaves evenly spaced along the stem, alternate to spirally arranged, occasionally clustered, erect, older leaves reflexed; stipules oblong, 2–6(–8) by c. 1 mm, membranous, lacerated apically, ferrugineous; lamina obovate, elliptic or spathulate, 0.4–1.1 (–3) cm by 2–7 mm, bearing tentacles adaxially and around the margins, almost glabrous to pilose abaxially (Fig. 10a); petiole linear, 0.6–2(–3) cm by 0.3–1 mm, glabrous to pilose. Inflorescences cymose, with 2–15 flowers, axillary; peduncles 1–3 together; 9–55 cm long, arising laterally, ascending, basally particularly curved, glabrous, glandular or pilose; bracts linear to spathulate, 2–3 mm long, sparsely pilose, caducous; pedicels 1–10 mm long, glandular or pilose. Sepals 5, elliptic to oblonglanceolate, 4–7 by 1–2 mm, connate at the base, apex acute or obtuse, glandular and/or pilose. Petals 5, obovate, 5–12 by 3.5–6 mm, pink or purple. Stamens 5; filaments 2–5 mm long. Pollen in tetrads, tetrads c. 45 µm diam, single grains c. 30 µm diam, echinate (Fig. 10e), echini broadly conical, c. 2 mm long, of medium density with short clavate sculptural elements between them. Pistil: ovary subglobose to ovoid, c. 2 by 1–1.5 mm, glabrous; styles 3, c. 2 mm long, 2-partite to the base, apices clavate, white to pink. Seeds fusiform, 0.7–1 by c. 0.2 mm, brownish black, apiculate, testa reticulate. Distribution — Widespread in tropical Africa, South Africa, Madagascar (Fig. 10c). Habitat & Ecology — Perennially wet habitats (swamps, marshes, river and lake shores, perennially wet places in seasonally dry wet-savannahs). Additional specimens (see Fig. 10d for province map). ANGOLA, Bié, Silva Porto – Andulo, km 35, S12°09' E16°45', alt. 1700 m, 15 Mar. 1973, P. Bamps et al. 4153 (BR); Cusseque – TFO core site, plain, alluvial plain, fire in last dry season, S13°42' E17°6', alt. 1515 m, 29 Oct. 2011, R. Revermann 132779 (HBG), M. Finckh 132768 (HBG); Cuando Cubango, Baixo Cubango, 179 km von Cuangar in Richtung Caiundo, sumpfiges Gelände am Rio Cubango, Moorboden, alt. 1070 m, 29 Jan. 1952, H. Hess 52/509 (ZT); Menongue, Vila Serpa Pinto, pr. da confluencia do Camumbé com o Cuebe, alt. 1420 m, 12 Feb. 1960, E.J. Mendes 2482 (COI); Cuito- Cuanavale, confluencia dos rios Tchiengo com o Cuito, alt. 1280 m, 3 Mar. 1960, E.J. Mendes 2811 (COI); Cuito-Cuanavale, Longa, Cuango, vale do Cuango, alt. 1370 m, 18 Mar. 1960, E.J. Mendes 3185 (BR, P); Huambo, Mission de Humabo, Nov. 1942, C. Tisserant A310 (COI); Chianga, alt. 1700 m, 11 June 1962, Teixeira & Andrada 6446 (COI); Arredores de Nova Lisboa, Chiva,alt. 1700 m, 19 Mar.1971, M. de Silva 3480 (BR, COI); Huíla, Serra de Chella, aux environs de Huíla, alt. 1850 m, 1937, H.Humbert 16686 (P); Entlang einem Bache auf dem Hochplateau von Bimbe bei Humpata, alt. 2300 m, 9 Sept. 1950, H. Hess 50/188 (ZT); Lubango, Chipia, ao km 7 da picada para Missao do Munhino, 11 May 1971, A. Borges 248 (BR, COI); Lunda, Vila Henrique de Carvalho, rio Chicapa, alt. 1100 m, 16 Apr. 1937, A.W. Exell & F.A. Mendon ҫa 636 & 644 (COI); Malanje, Quimbango, Congolo, Reserca da Palanca Negra Gigante (ou Reserva do Luando), 7 June 1970, G. Barbosa et al. 11963 (COI); Uíge, near small lagoon, through which water flows, S6°1' E15°24',alt. 803 m, 25 Feb.2017, T. Lautenschläger 2017-02-139 & 2017-02-111 (DR). – DRC, Equateur, Ilema (Terr.Coquilhat- ville), 21 Mar. 1958, Èvrard 3748 (BR); Haut-Katanga, Le long de la route Mitwaba – Manono,à 11 km de Mitwaba – Sermikat, 21 Apr.1997, Symoens 3612 (BR); Ituri, Nioka (Mahagi), alt. 1700 m, 29 July 1960, D. Froment 758 (BR, P); Kinshasa, Route Soo-Ndunu, Maluku, 5 May 1971, H. Breyne 2190 (BR); Kongo Central, Binza, Route de Ngidinga à Kimvula, Territoire de Madimba, 15 Feb. 1960, P. Compere 1479 (BR); Kilenfu-Leg-Boko, S5°01' E15°11', 10 Feb. 2014, Nsimundele 2723 (BR); Kwango, Nto Mbombo, terr. Popokabaka, 9 Jan. 1959, L. Pauwels 1208 (BR); Tangi, alt. 700 m, Jan. 1930, J. Lebrun 127 (BR); Lualaba, Shaba, Kolwezi – Luena, km 114, Riv. Mutendele, 20 Jan.1986, Bamps & Malaisse 8222 (BR); Environs de Kolwezi, vallée de la Kanamwamfwe, 23 Aug. 2004, Malaisse & Kisimba 134 (BR); Sud-Kivu, Massif du Kahuzi, Env. du 43 km de la route Bukavu-Walikale, alt. 2200 m, 22 Apr. 1970, J. Lambinon 78/228 (BR); Tanganyika, Kilunga (Marungu), Apr. 1945, L. Dubois 1416 (BR); 3 km a l’W. de Kasiki, Plateau des Marungu, alt. 2000 m, 12 June 1969, S. Lisowski et al. 6139 (BR); Mare Mufufu, Plateau des Marungu, alt. 1900 m, 14 June 1969, S. Lisowski et al. 6606 (BR). Note — A very variable species with stems occasionally suppressed and dwarf forms (these show often suborbicular to elliptic leaves and a reddish colour). Hybridisation with D. elongata and D. affinis is possible. Plants in South Africa with suppressed stems (D. madagascariensis var. major Burtt Davy) were synonymized with D. curvipes Planch. and plants from southern Africa were synonymised with D. ramentacea Burch. ex DC. Both D. curvipes and D. ramentacea are now mostly accepted as separate taxa (Robinson et al. 2017). The distinct diagnostic features of D. madagascariensis (reflexed leaves, strongly curved base of inflorescence; Fig. 10) make most of the specimens easy to determine. However, the species is known to be highly variable in height, stem length and leaf size and shape depending on the habitat and available nutrients (Robinson et al. 2017). The dwarf form of D. madagascariensis (e.g., H. Breyne 4636) can be easily mistaken for D. burkeana or D. pilosa. To distinguish between these species leaf shape, seed shape and morphology and pollen morphology could be used. Furthermore, relatively large and robust specimens occur. Three large specimens are doubtfully determined as D. madagascariensis (partly H. Hess 50/188, A.W. Exell & F.A. Mendon ҫa 644 and 636) showing very large obovate to elliptic laminae, densely pilose stems, scapes and leaves and more or less erect to strongly curved inflorescences., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on pages 10-11, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Diels L. 1906. Droseraceae. In: Engler A (ed), Das Pflanzenreich IV. 112. Engelmann, Leipzig.","Gilbert G. 1951. Droseraceae. In: Flore du Congo Belge et du Ruanda-Urundi. Institut national pour l'etude agronomique du Congo Belge.","Keay RWJ. 1954. Flora of West Tropical Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1959. Droseraceae. In: Hubbard CE, Milne-Redhead E (eds), Flora of Tropical East Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1978. Droseraceae. In: Launert E (ed), Flora Zambesiaca. Flora Zambesiaca Managing Committee, London.","Troupin G. 1978. Flore du Rwanda. Musee Royal de l'Afrique Centrale, Tervuren.","Taton A. 1945 b. Drosera congolana. Bulletin du Jardin Botanique de l'Etat a Bruxelles: 310.","Robinson A, Gibson R, Gonella P, et al. 2017. Drosera of the World - volume 3, Latin America, Africa. Redfern Natural History Productions, Poole."]}

5. Drosera flexicaulis Welw. ex Oliv. — Fig. 7 Drosera flexicaulis Welw.ex Oliv.(1871) 403; Diels (1906) 98; G.C.C. Gilbert (1951) 554;A.S.Rob. et al. (2017) 1398. — Lectotype (designated by Diels (1906) 98): F.M.J. Welwitsch 1181 (lecto LISU), Angola, Huíla, Lopollo, Dec. 1859. Perennial herb, caulescent. Stems up to 25 cm long, erect, slender, glabrous (Fig. 7b). Leaves alternate, clustered somewhat into a rosette towards the apex, all leaves including lower leaves mostly erect or ascending (sometimes reflexed); stipules 1–2 mm long, connate at the base, apex lacerated, ferrugine- ous; lamina lanceolate, narrowly spathulate-ovate or spathulate, 3–15 by 1–5 mm, bearing tentacles adaxially and around the margins, glabrous or sparsely pilose abaxially (Fig. 7a); petioles 0.5–3.2 cm long, slender, glabrous or sparsely pilose. Inflorescences cymose, with 3–13 flowers; peduncles 1–4 together, 8–21(–30) cm long, ascending, glabrous;bracts filiform to linear, c. 2 mm long, caducous; pedicels 1–10 mm long, erect, glabrous or sparsely pilose. Sepals 5, oblong to lanceolate, 4–7 by 1–2 mm, connate at the base, sparsely pilose to pilose. Petals 5, obovate, 5–8 by 2.5–4 mm, white or purple. Stamens 5, filaments 4–5 mm long. Pollen in tetrads, tetrads c. 45–52 µm diam, single grains c. 25–30 µm diam, echinate (Fig. 7e), echini conical, c. 2 mm long, in low density, with short clavate sculptural elements between echini. Pistil: ovary subglobose to ovoid, c. 2 by 1 mm, glabrous; styles 3, c. 2.5 mm long, 2-partite to the base. Seeds fusiform, 0.7–0.9 by c. 0.2 mm, black, shiny, testa reticulate. Distribution — Angola, DRC, Zambia (Fig. 7c). Habitat & Ecology — Grows on sand and peat, in grassy swamps, seasonally dry wet-savannah. Additional specimens (see Fig.7d for province map). ANGOLA, Bié, junto à margem do rio Chitembo,alt. 1500 m, 31 Oct.1966, Teixeira et al. 10901 (BR); Cusseque – TFO core site, hilly landscape, fire in last dry season, S13°42' E17°5', 19 Oct. 2012, M. Finckh 135547 (HBG); Cusseque – TFO core site, plain:alluvial plain,fire in last dry season, S13°41' E17°6',alt. 1515 m, 29 Oct. 2011, M. Finckh 132756 (HBG); Cuando Cubango, Menongue, Vila Serpa Pinto, pr. da confluencia do Cambumbé com o Cuebe, alt. 1420 m, 12 Feb. 1960, E.J. Mendes 2484 (BR, COI); Menongue, andados 14 km de Vila Serpa Pinto para Longa, vale do Cambumbé,alt. 1430 m, 26 Feb.1960, E.J. Mendes 2754 (COI); Cuanza Sul, am Longa oberh. Minnesera, alt. 1230 m, 1 Feb.1900, H. Baum 687 (HBG); Lunda Sul, Saurimo, 25 Oct.1932, R.G.N. Young 1167 (COI). – DRC, Haut-Katanga, 6 km à l’WNW de la touree Oee de la Lut., Plateau des Kundelungu, alt. 1680 m, 25 Mar. 1969, S. Lisowski et al. 3789 (BR); près de la touree Oee de la Lut., Plateau des Kundelungu,alt. 1670 m, 26 Mar. 1969, S. Lisowski et al. 3552 (BR); 2 km a l’E des tourees de la Lut., Plateau des Kundelungu, alt. 1600 m, 8 Jan. 1971, S. Lisowski et al. 13079 (BR); Lualaba, Plateau de la Manika (environs Katentania), Nov. 1912, Homblé 774 (BR); village Matoki, Plateau de la Manika, alt. 1400 m, 20 Jan. 1969, S. Liswoski et al. 5 (BR); Kisoté, Plateau de la Manika, alt. 1500 m, 5 Apr. 1969, S. Lisowski et al. 4181 (BR); 2 km à l’W de la riviere Musonai vers le village Djoni, Plateau de la Manika, alt. 1420 m, 23 Mar. 1969, S. Lisowski et al. 5651 (BR). Note — This species has been treated as a synonym of D. affinis in several Floras (Laundon 1959, 1970, 1978). After morphological analysis this work follows the Flora of Tropical Africa (Oliver 1871) and treats D. flexicaulis as a separate species. Both species share the same habitat and have overlapping distributions.Additionally, hybrids of D. flexicaulis and D. affinis occur, which show characters of both parent species (Robinson et al. 2017)., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on pages 7-9, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Diels L. 1906. Droseraceae. In: Engler A (ed), Das Pflanzenreich IV. 112. Engelmann, Leipzig.","Gilbert G. 1951. Droseraceae. In: Flore du Congo Belge et du Ruanda-Urundi. Institut national pour l'etude agronomique du Congo Belge.","Laundon JR. 1959. Droseraceae. In: Hubbard CE, Milne-Redhead E (eds), Flora of Tropical East Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1970. Droseraceae. In: Wittnich Carisso L (ed), Conspectus Flo- rae Angolensis. Instituto Botanico de Coimbra & British Museum, Lissabon.","Laundon JR. 1978. Droseraceae. In: Launert E (ed), Flora Zambesiaca. Flora Zambesiaca Managing Committee, London.","Oliver D. 1871. Leguminosae to Ficoideae. In: Flora of Tropical Africa. Reeve & Co., London.","Robinson A, Gibson R, Gonella P, et al. 2017. Drosera of the World - volume 3, Latin America, Africa. Redfern Natural History Productions, Poole."]}

6. Drosera indica L. — Fig. 8 Drosera indica L. (1753) 282;Oliv. (1871) 402; Diels (1906) 77;G.C.C. Gilbert (1951) 551; Keay (1954) 122; J.R. Laundon (1959) 2, (1970) 25; Oberm. (1970) 201; J.R. Laundon (1978) 64;Akoègn.et al. (2006) 539;A.S.Rob. et al. (2017) 1418. — Lectotype (designated by Laundon (1959) Droseraceae 2): P. Hermann s.n. (lecto BM), India. Synonyms in Africa only: Drosera minor Schumach. (1827) 167. — Type: C.F. Schumacher s.n. (holo C, not seen), Guinea. Slender, annual, caulescent herb (Fig. 8b). Stems 3–50 cm long, erect or matted, canaliculate, (sparsely) glandular, green. Leaves alternate to spirally arranged, evenly spaced, erect, old leaves reflexed; stipules absent, a pair of setae-like hairs at the base of the leaves; lamina linear to filiform, 1–10 cm by 0.4–4 mm, apex long acuminate, adaxial surface and margins bearing tentacles, abaxially glabrous, green (Fig. 8a); petiole 0.1–1.5 cm long, glandular. Inflorescences cymose, with 3–20 flowers, recurved; peduncles 0.5–15 cm long, arising laterally from the stem, glandular; bracts linear to spathulate, c. 2.5 mm long, apex more or less dentate; pedicels 0.2–2 cm long, glandular. Sepals 5, ± lanceolate, 2–5 by 1–2 mm, apex acute or obtuse, glandular. Petals 5, spathulate to obovate, 4–8 by 2.5–6 mm, pink to purple, mauve (in Asia also white). Stamens 5; filaments 3–5 mm long, white or pink; anthers yellow. Pollen in tetrads, tetrads c. 50–55 µm diam, single grains c. 25 µm diam, long echinate (Fig. 8e), echini narrowly conical, 2–3 µm long, occurring in medium density, with short clavate sculptural elements between echini. Pistil: ovary subglobose to ovoid, 1.5–3 by c. 2 mm, glabrous; styles 3, bipartite to the base, branches c. 2.5 mm long, ascending. Seeds globose to ovoid, 0.4–0.5 by c. 0.3 mm, apiculate, black, testa reticulate (Fig. 8f). Distribution — Widespread in tropical Africa, Asia and Aus- tralia (Fig. 8c). Habitat & Ecology — Wide range of seasonally wet habitats (e.g., swamps, pools, rivers and lake shores). Additional specimens (see Fig. 8d for province map). ANGOLA, Benguela, Tchiyaka, 20 km nördlich von Quingenge, gefunden auf nasser Felsplatte, alt. 1750 m, 13 Feb. 1955, J.B. Damann s.n. (sine nummero) (ZT); Huíla, Humpata, alt. 1800 m, 5 Apr. 1937, J. Gossweiler 11104 (COI); Entre lá da Bandeira e Humpata, alt. 2000 m, 6 May 1937, A.W. Exell & F.A. Mendon ҫa 2599 (COI); Lubango, entre Palaca e o Perimetro Florestral, alt. 1960 m, 15 Apr. 1960, E.J. Mendes 3613 (COI); Moxico, Cameia National Park, Biodiversity Observatory, S11°31' E20°54', alt. 1126 m, 3 May 2016, Finckh & Zigelski 143317 (HBG). – DRC, Haut-Katanga/Haut-Lomami, Parc National de l’Upemba, Entre la riv. Kanonya et la riv. Lukange, 19 Feb. 1949, G.F. de Witte 5544 (BR); Haut-Uele, Mai-Kuku, 6 km au-delà Gombari, alt. 1200 m, 16 July 1937, J. Louis 4540 (BR); Parc National de la Garamba, route Dungu – Bagbele (Ukwa), km 17 de Bagbele, 25 Aug.1952, G. Troupin 1991 (BR); Ituri, Entre Jaradje et Aba, July 1931, J. Lebrun 3429 (BR, P); Kasai-Central, Samusambu, 65 km de Sandoo, Lulua, 6 June 1932, Overlaet 1252 (BR); Kinshasa, Kinshasa, 17 May 1915, Bequaert 7612 (BR); Congo-Kinsuka, près Leopoldville, July 1944, Coutreaux 1011 (BR); Kinsuka, 28 May 1947, E. Jans 490 (BR); Ile des Mimosas, Ngaliema, 1 June 1983, H. Breyne 4624 (BR); Kongo Central, Gombe Matadi, Jan.1950, H. Callens 2594 (BR); Tshopo, Niomgaza (Uili-Nepoko), June 1931, J. Lebrun 3174 (BR)., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on page 9, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Diels L. 1906. Droseraceae. In: Engler A (ed), Das Pflanzenreich IV. 112. Engelmann, Leipzig.","Gilbert G. 1951. Droseraceae. In: Flore du Congo Belge et du Ruanda-Urundi. Institut national pour l'etude agronomique du Congo Belge.","Keay RWJ. 1954. Flora of West Tropical Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1959. Droseraceae. In: Hubbard CE, Milne-Redhead E (eds), Flora of Tropical East Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1978. Droseraceae. In: Launert E (ed), Flora Zambesiaca. Flora Zambesiaca Managing Committee, London."]}

1. Drosera affinis Welw. ex Oliv. — Fig. 3 Drosera affinis Welw. ex Oliv. (1871) 402; Diels (1906) 88; J.R. Laundon (1959) 4; (1970) 27; (1978) 66; A.S.Rob. et al. (2017) 1318. — Type: F.M.J. Welwitsch 1183 (holo LISU), Angola, Distr. Huilla, Lopolo, Jan.1860. Perennial herb, caulescent. Stems mostly short, up to 10 cm high (Fig. 3b). Leaves up to 9 cm long (petiole and lamina), alternate to spirally arranged, clustered, densely arranged, mostly erect, old leaves reflexed; stipules 0.3–1 cm long, ferrugineous, apex lacerated; lamina narrowly spathulate or obovate, 1.5–3 by 0.2–0.5 cm, apex obtuse, gradually narrowing from the apex downwards, bearing tentacles adaxially and around the mar- gins, glabrous or sparsely pilose abaxially (Fig. 3a); petiole 3–7 cm long (2–5 times as long as lamina), slender, glabrous or sparsely pilose. Inflorescences cymose, with 3–13 flowers; peduncles 1–3 together; 8–35 cm long, arising laterally from the stem, slightly curved at the base or erect, glabrous, canaliculate; bracts linear to elliptic, 3–5 mm long, caducous; pedi- cels 2–10 mm long, glabrous or sparsely pilose (sometimes glandular). Sepals 5, oblong to lanceolate, 3–7 by 1.5–2 mm, connate at the base, sparsely pilose. Petals 5, elliptic to ob- ovate, 5–8 mm long, apex obtuse, pale pink to purple. Stamens 5, filaments c. 5 mm long and pink. Pollen in tetrads, yellow, tetrads c. 35–55 µm diam, single grains c. 23–30 µm diam, echinate (Fig. 3e), echini ± conical, c. 2 µm long, medium density of echini, short gemmate to clavate sculptural elements between echini. Pistil: ovary subglobose, glabrous; styles 3, c. 2.5 mm long, 2-partite to the base, arms filiform. Seeds fusiform, 0.7–0.9 by c. 0.2 mm, testa reticulate. Distribution — Angola, Congo, DRC, Malawi, Mozambique, Namibia, Tanzania, Zambia, Zimbabwe (Fig. 3c). Habitat & Ecology — Wet meadows, swamps, perennially wet places in seasonally dry wet-savannahs. Additional specimens (see Fig.3d for province map). ANGOLA, Bié, Camacupa, Cuemba, quedas, alt. 1200 m, 6 Oct. 1965, Teixeira & Matos 8995 (BR); Huíla, Huíla, Oct. 1898, E. Dekindt 466 (BR); Na margem di rio Nene, a montante da ponte, junto da Estrada de Huíla para Jau, 17 Dec. 1955, E.J. Mendes 1144 (COI); Sà da Bandeira, na subida para o miradouro da Tundavale, nos lugares enchrcados, 13 Oct. 1969, F. Murta & M. Silva 799 (COI); Lunda Sul, Dala, margem do rio Chiumbe,1927, Carrisso & Mendon ҫa 564 (COI); Dala, rio Chiumbe, lugares húmidos, alt. 1200 m, 23 Apr. 1937, A.W. Exell & F.A. Mendon ҫa 1087 (COI); Moxico, quedas de Dala, 17 Aug. 1965, R. Mendes dos Santos 1658 (COI). – DRC, Haut-Katanga, Kaziba, Terr. Manono, Parc national de le Upemba, alt. 1500 m, 16 Feb. 1948, G.F. de Witte 3383 (BR)., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on pages 4-5, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Diels L. 1906. Droseraceae. In: Engler A (ed), Das Pflanzenreich IV. 112. Engelmann, Leipzig.","Laundon JR. 1959. Droseraceae. In: Hubbard CE, Milne-Redhead E (eds), Flora of Tropical East Africa. Crown Agents for Oversea Governments and Administrations, London."]}

2. Drosera bequaertii Taton — Fig. 4 Drosera bequaertii Taton (1945a) 308;G.C.C. Gilbert (1951) 553;J.R. Laundon (1970) 27; (1978) 64; A.S.Rob. et al. (2017) 1334. — Type: J. Bequaert 476 (holo BR), DRC, 8 June 1912. Drosera compacta Exell & J.R.Laundon (1955a) 44. — Type: A.W. Exell & F.A. Mendon ҫa 1030 (holo COI), Angola, Lunda Sul, Rio Luachimo, entre Sila Henrique de Carvalho e Dala, ‘Tenga’, 24 Apr. 1937. Perennial herb, shortly caulescent. Stems 3–4 (up to 7) cm long, 1–2.5 mm diam, densely pilose (Fig. 4b). Leaves alternate, clustered at the apex and erect, basally more widely spaced and more or less reflexed; stipules 2–4.5 mm long, membra- nous, ferrugineous, lacerated at the apex; lamina obovate to spathulate, 4–12 by 3–6 mm, the adaxial surface and margins bearing tentacles, abaxially pilose (Fig. 4a); petiole 5–12 by 0.5–1 mm, flat, densely pilose. Inflorescence cymose, with 2–8 flowers; peduncles 1 or 2 together, 10–17 cm long, completely erect, basally glandular and pilose to densely pilose, apically glandular; bracts almost linear, 2–3 mm long, glandular or pi- lose; pedicels 2–7 mm long, glandular. Sepals 5, lanceolate to ovate, 3–4.5 by 1.5–2 mm, connate at the base, occasionally apex serrate, glandular. Petals 5, obovate, 5–7 by c. 4 mm, bright pink. Stamens 5, filaments 3–4 mm long. Pistil: ovary subglobose, 2–3 mm diam, glabrous; styles 3, c. 2 mm long, bipartite to the base, branches ascending, with broadened apices. Seeds suborbicular to ovoid, c. 0.3 by 0.2 mm (Fig. 4e), black, shining, testa smooth. Distribution — Angola, DRC, Zambia (Fig. 4c). Habitat & Ecology — Grassy wetlands, upland plateaus. Additional specimens (see Fig. 4d for province map). ANGOLA, Moxico, Cameia National Park, Biodiversity Observatory, S11°31' E20°55', alt. 1126 m, 3 May 2016, M.Finckh & Zigelski 143303 (HBG). – DRC, Haut-Katanga, Cuv. de Kabiashia, rivière Kandale, alt. 1000 m, 24 June 1968, J. Leonard 4617 (BR). Note — Drosera compacta is widely accepted as a synonym of D. bequaertii. Both type specimens show many similarities but differ in leaf arrangement (densely vs more openly arranged) and scape indumentum (basally densely pilose vs glandular and pilose). Since this species is only infrequently documented (Robinson et al. 2017) and only few specimens were studied, no comment can be made on separating both taxa. Here, a more detailed analysis is required., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on page 5, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Taton A. 1945 a. Drosera bequaertii. Bulletin du Jardin Botanique de l'Etat a Bruxelles: 308.","Gilbert G. 1951. Droseraceae. In: Flore du Congo Belge et du Ruanda-Urundi. Institut national pour l'etude agronomique du Congo Belge.","Laundon JR. 1970. Droseraceae. In: Wittnich Carisso L (ed), Conspectus Flo- rae Angolensis. Instituto Botanico de Coimbra & British Museum, Lissabon.","Robinson A, Gibson R, Gonella P, et al. 2017. Drosera of the World - volume 3, Latin America, Africa. Redfern Natural History Productions, Poole."]}

9. Drosera pilosa Exell & J.R.Laundon — Fig. 11 Drosera pilosa Exell & J.R.Laundon (1956) 213;J.R. Laundon (1959) 4; (1970) 26; Troupin (1978) 305; A.S.Rob. et al. (2017) 1452. — Type: A. Stolz s.n. (holo BM), Tanzania, Tanganyika territory, Kyimbila District, North of Lake Nyasa, 1885. Acaulescent, perennial herb (Fig. 11b). Leaves in a basal rosette; stipules c. 3 mm long, connate at the base, apex lacer- ated; lamina elliptic, ovate or obovate, 4–13 by 2–8 mm, the adaxial surface and margins bearing tentacles, abaxially pilose (Fig.11a); petiole 3–15 by up to 2 mm, gradually broadening into the lamina, abaxially pilose. Inflorescences cymose, with 2 – 6 flowers; peduncles 1 or 2 together, 2–16 cm long, arising laterally from the rosette, ascending, canaliculate, pilose; bracts elliptic, 2–3 mm long; pedicels 1–7 mm long, pilose. Sepals 5, ± elliptic, 3–6 by 1.5–2.5 mm, apex acute, pilose. Petals 5, c. 6 mm long, red to purple. Stamens 5; filaments c. 4 mm long. Pollen in tetrads, tetrads c. 40–45 µm diam, single grains 30–35 µm diam, echinate (to baculate; Fig. 11e), echini broadly conical, 1–2 mm long, in high density with short clavate sculptural elements between them. Pistil: ovary subglobose, c. 1.5 by 2 mm; styles 3, c. 2 mm long, bipartite to the base. Seeds ovoid, 0.3–0.5 by c. 0.2 mm, black, testa smooth (Fig. 11f). Distribution — Angola, Burundi, Cameroun, DRC, Guinea, Kenya, Malawi, Ruanda, Sierra Leone, Tanzania (Fig. 11c). Habitat & Ecology — Montane species. Additional specimens (see Fig. 11d for province map). ANGOLA, Cuando Cubango, Cuito-Cuanavale, Longa, vale do Cuiriri, alt. 1360 m, 19 Mar. 1960, E.J. Mendes 3212 (COI). – DRC, Sud-Kivu, Environs de Kitwabaluzi, S2°53' E28°35', Dec.1942, Coutreaux 999 (BR); Kabare,alt. 2200 m, 9 Dec. 1956, Christiaensen 1953 (BR); Unknown province, Luemba, Sept. 1946, Kunet 32 (BR). Note — Very similar to D. burkeana but with pilose peduncle, pedicels and sepals., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on page 12, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Laundon JR. 1959. Droseraceae. In: Hubbard CE, Milne-Redhead E (eds), Flora of Tropical East Africa. Crown Agents for Oversea Governments and Administrations, London.","Troupin G. 1978. Flore du Rwanda. Musee Royal de l'Afrique Centrale, Tervuren."]}

3. Drosera burkeana Planch. — Fig. 5 Drosera burkeana Planch. (1848) 192; Oliv. (1871) 402; Diels (1906) 88; G.C.C. Gilbert (1951) 552; Keay (1954) 26; J.R. Laundon (1959) 2; (1970) 26; Oberm.(1970) 194;A.S.Rob. et al. (2017) 1338. — Type: J. Burke s.n. (holo K), South Africa, Nordwest, Macalisberg. Perennial herb (sometimes annual), acaulescent. Leaves in a basal rosette (Fig. 5b), held horizontally; stipules 2–3.5 by c. 1.5 mm, membranous, connate at the base, apex lacerated (3 or more teeth), ferrugineous; lamina suborbicular, obovate to spathulate, 2–10 by 2–9 mm, reddish, adaxial surface and margins bearing tentacles, abaxially pilose, rarely glabrous (Fig. 5a); petiole 2–25 by 0.5–1 mm, linear throughout, abruptly broadening into the lamina, reddish, pilose (or glabrous). Inflorescences cymose, with 3–12 flowers; peduncles 1–4 together, 4–33 cm long, arising laterally, ascending, glandular; bracts linear, spathulate to narrowly obovate, 1–2 mm long, glandular or glabrous; pedicels 2–12 mm long, glandular. Sepals 5, elliptic to obovate, 2–5.5 by 2–2.5 mm, apex acute or obtuse, serrated, surfaces glandular (especially adaxially). Petals 5, obovate, 5–7 by 2–4 mm, pink to purple or white. Stamens 5, filaments 3–5 mm long, same colour as petals, anthers yellow. Pollen in tetrads, yellow, tetrads c. 40–46 µm diam, single grains c. 30–35 µm diam, pollen grains echinate (Fig. 5e), echini narrowly conical, c. 2 mm long, high density of echini, with short gemmate to clavate sculptural elements between echini. Pistil: ovary subglobose to obovate, 1.5–3.5 by 1–2.5 mm, glabrous, green; styles 3, 4–5 mm long, bipartite to the base, branches slender, curved upwards. Seeds ovoid, 0.3–0.4 by 0.15–0.2 mm, black, testa reticulate. Distribution — Angola, Botswana, Burundi, Congo, DRC, Kenya, Lesotho, Madagascar, Malawi, Mozambique, Namibia, South Africa, Swaziland, Tanzania, Uganda, Zambia, Zimbabwe (Fig. 5c). Habitat & Ecology — Grows on loam, sand and peat, sea- sonally to perennially wet spots. Additional specimens (see Fig.5d for province map). ANGOLA, Benguela, Feuchte Wiese bei Humpata,alt. 1900 m, 9 Sept. 1950, H. Hess 50/ 254 (ZT); Cuanza Sul, zona de Cassongue estrada em direc ҫão de Kahumbi,após do rio Cuvele femea, Inselberg, alt. 1580 m, 1 Apr. 2007, T. Leyens 402 (FR); Cunene, Cuvelai,a cerca de 20 km para Cassinga, 22 July 1971, R. Santos 2918 (COI); Huambo, Huambo, 1923, H. Vanderyst 13125 (BR); près de la mission de Huambo, Aug. 1942, C. Tisserant A94 (COI); 1 km W der prot. Mission von Elende, Cuma, alt. 1350 m, 25 July 1961, J.B. Damann 3099 (ZT); Chianga, alt. 1700 m, 11 June 1962, Teixeira & Andrada 6445 (COI); Huíla, Serra de Chella, aux environs de Humpata, alt. 2000–2200 m, 1937, H. Humbert 16650 (P); Humpata, Chongorola, rio Nene, pâtanos, 20 June 1937, A.W. Exell & F.A. Mendon ҫa 2989 (COI); Sá da bandeira, Jau, Can- galongue, 3 Oct. 1965, C. Henriques 763 (COI); Lunda, 100 km norte de Muconda, 12 Aug. 1927, Carrisso & Mendon ҫa 221a (COI); Rio Caunhango junto da estrada Mucango a Casage, 16 Aug. 1927, Carrisso & Mendon ҫa 264 (COI). – DRC, Haut-Katanga, 6.3 km au NNE du Peste de Katshupa, Plateau des Kundelungu, alt. 1580 m, 3 Aug. 1966, F. Malaisse 4490 (BR); 7 km au N du Peste de Katshupa, Plateau des Kundelungu, alt. 1680 m, 1966-08-04, F. Malaisse 4504 (BR); 6 km à l’WNW de la touree Oee de la Lut., Plateau des Kundelungu, alt. 1675–1680 m, 25 Mar. 1969, S. Lisowski et al. 3975 & 3453 (BR); Haut-Katanga/ Haut-Lomami, Parc National de l’Upemba,route automobile pour le Shinkulu,alt. 1450 m, 21 May 1948, G.F. de Witte 03862 (BR); Parc National de l’Upemba, 24 Sept. 1949, G.F. de Witte 07541 (BR); Lualaba, Dilolo, Sept. 1931, Overlaet 1239 (BR); Biano, S10°16' E26°03', alt. 1575 m, 23 May 1952, A. Schmitz 3918 (BR)., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on page 7, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Diels L. 1906. Droseraceae. In: Engler A (ed), Das Pflanzenreich IV. 112. Engelmann, Leipzig.","Gilbert G. 1951. Droseraceae. In: Flore du Congo Belge et du Ruanda-Urundi. Institut national pour l'etude agronomique du Congo Belge.","Keay RWJ. 1954. Flora of West Tropical Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1959. Droseraceae. In: Hubbard CE, Milne-Redhead E (eds), Flora of Tropical East Africa. Crown Agents for Oversea Governments and Administrations, London."]}

Drosera L. — Fig. 2 Drosera L. (1753) 281; Keay (1954) 120; Oberm. (1970) 189; J.R. Laundon (1978) 64; Kubitzki & C.Bayer (2003) 201; A.M. Ellison & Adamec (2018) 50. — Type: D. rotundifolia L. Carnivorous, annual or perennial herbs. Stems caulescent or acaulescent. Leaves simple, in a basal rosette or alternate to spirally arranged along the stem; stipules present or absent, membranous, often lacerated; petiole present or confluent with lamina; lamina variously shaped (Fig. 2), adaxial surface covered with short- and long-stalked glands (tentacles) and digestive glands that trap and digest insects. Inflorescences 1–many-flowered cymes, axillary or terminal, flowers bisexual. Sepals 5 (rarely 4–8); ± connate at the base. Petals 5 (rarely 4–8); free. Stamens 5 (rarely 4–8); free. Pollen in tetrads. Pistil: ovary (2–)3–5 carpels; styles (2–)3–5. Fruit a loculocidal capsule. Seeds small, numerous. Distribution — Over 240 species, almost worldwide. Habitat & Ecology — Swamps, sandy, seasonally and peren- nially wet and low-nutrient habitats. KEY TO THE ANGOLAN AND CONGOLESE DROSERA SPECIES 1. Stipules absent, lamina linear............. 6. D. indica 1. Stipules present, lamina lanceolate, spathulate, obovate, ovate, elliptic, suborbicular or orbicular.............. 2 2. Acaulescent, leaves in a basal rosette, leaves horizontal. 3 2. Caulescent, including short stems (one to several cm long, rarely appearing acaulescent), young leaves erect, older leaves variable (erect, perpendicular to stem, reflexed).. 5 3. Seeds fusiform, seed testa reticulate, peduncle strongly curved at the base, peduncle, pedicels and sepals glabrous, glandular or pilose, petals pink or purple. — Usually caules- cent, rarely dwarf forms with suppressed stems................................... 8. D. madagascariensis 3. Seeds ovoid, seed testa smooth or reticulate, peduncle curved at the base, peduncle, pedicels and sepals glandular or pilose, petals pink to purple or white.............. 4 4. Petiole narrow, abruptly broadening into the suborbicular to spathulate lamina, peduncle, pedicels and sepals glandular, petals white or purple, seed testa reticulate 3. D. burkeana 4. Petiole gradually broadening into the elliptic to obovate lamina, peduncle, pedicels and sepals usually pilose, petals pink to purple, seed testa smooth.......... 9. D. pilosa 5. Peduncle arising apically, erect throughout its whole length, stem very short (c. 4 cm long), densely pilose, seeds sub- orbicular to ovoid.................... 2. D. bequaertii 5. Peduncle arising laterally, curved or erect at the base, then ascending, stem short to very long (up to 90 cm), glabrous, glandular or pilose, seeds fusiform................. 6 6. Older (but still green) leaves reflexed towards the stem, old parts of the stem enclosed with old reflexed leaves, peduncle strongly curved at the base..... 8. D. madagascariensis 6. Older (but still green) leaves held erect or ± perpendicular to the stem (rarely reflexed), peduncle ± curved at the base 7 7. Stem up to 90 cm long, older (but still green) leaves per- pendicular to stem, lamina obovate to elliptic, peduncle and petioles pilose or glandular-pilose........ 4. D. elongata 7. Stem shorter (max. 25 cm long, sometimes longer), older (but still green) leaves ± erect, lamina ± spathulate, peduncles and petioles glabrous/sparsely pilose or densely pilose. 8 8. Peduncle and petioles densely pilose/hirsute, stem up to 15 cm long, densely glandular, lamina oblong-spathulate. ― Only known from Katanga, DRC.... 7. D. katangensis 8. Peduncle and petioles glabrous or sparsely pilose, stem short (up to 10 cm) or elongated (up to 25 cm), glabrous, lamina narrowly spathulate, spathulate or obovate..... 9 9. Stem short (up to 10 cm), leaves closely arranged, petioles very slender and 3–7 cm long............. 1. D. affinis 9. Stem elongated (up to 25 cm, sometimes longer), alternating between segments with evenly spaced leaves and segments with rosette-like clustered leaves, petioles max. 3.2 cm long.............................. 5. D. flexicaulis, Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on pages 3-4, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Keay RWJ. 1954. Flora of West Tropical Africa. Crown Agents for Oversea Governments and Administrations, London.","Laundon JR. 1978. Droseraceae. In: Launert E (ed), Flora Zambesiaca. Flora Zambesiaca Managing Committee, London.","Ellison AM, Adamec L (eds). 2018. Carnivorous plants - physiology, ecology, and evolution. Oxford University Press, Oxford."]}

4. Drosera elongata Exell & J.R.Laundon — Fig. 6 Drosera elongata Exell & J.R.Laundon (1955b) 42; J.R. Laundon (1970) 29; A.S.Rob. et al. (2017) 1384. — Type: A.W. Exell & F.A. Mendon ҫa 1268 (holo BM), Angola, Lunda Sul, Luma-Cassai, ‘Tenga’, 27 Apr. 1937. Slender, scrambling, caulescent herb. Stems up to 90 cm long, c. 1 mm diam, flexuous, pilose to densely pilose (Fig. 6b). Leaves evenly spaced along the stem, alternate to spirally arranged, semi-erect, older leaves perpendicular to stem or slightly reflexed; stipules 2–5 mm long, membranous, deeply lacer- ated apically, ferrugineous; lamina obovate to elliptic, 3–6 by 2–4 mm, bearing tentacles adaxially and around the margins, pilose abaxially (Fig. 6a); petiole 0.7–1.5 cm long, slender, pilose. Inflorescences cymose, with 2–9 flowers; peduncles 1 or 2 together, 9–25 cm long, arising laterally, ascending, pilose to glandular-pilose; bracts linear to oblanceolate, 2–3 mm long, pilose or glandular-pilose; pedicels 1–6 mm long, pilose to glandular-pilose. Sepals 5, lanceolate to elliptic, 3–5 by 1–2 mm, connate at the base, apex obtuse or acute, glan- dular-pilose. Petals 5, obovate, 4–6 by 2–3 mm, obtuse, pink. Stamens 5, filaments 2–4 mm long. Pollen in tetrads, tetrads c. 45–50 µm diam, single grains c. 30 µm diam, echinate (Fig. 6e), echini conical, c. 2 mm long, sometimes constricted in the middle, in low to medium density, with short gemmate to clavate sculptural elements between echini. Pistil: ovary c. 1 by 1.5 mm, glabrous; styles 3, 2-partite at the base. Seeds fusiform, 0.5–0.8 by c. 0.2 mm, black, testa smooth (Fig. 6f). Distribution — Angola, Burundi, Congo, DRC, Tanzania, Zambia (Fig. 6c). Habitat & Ecology — Perennially wet places in seasonally dry wet-savannah, upland plateaus. Additional specimens (see Fig. 6d for province map). ANGOLA, Lunda, R. Lalege, 14 Aug. 1927, Carrisso & Mendon ҫa 245a (COI); Uíge, Mucaba swampy area, S7°13' E15°5', alt. 1209 m, 23 Feb. 2017, T. Lautenschläger 2017-02-113 (DR). – DRC, Haut-Katanga, Parc National de l’Upemba, Kiluba, Luswahila, alt. 1880 m, 16 Jan. 1948, G.F. de Witte 03249 (BR); Parc National de l’Upemba, Kaziba, alt. 1300 m, 18 Apr. 1949, G.F. de Witte 06261 (BR); Haut-Katanga / Haut-Lomami, Parc National de l’Upemba, alt. 1680 m, 24 Aug. 1949, G.F. de Witte 07540 (BR). Note — Hybridisation with D. madagascariensis has been observed, the hybrids show characteristics of both species (Robinson et al. 2017)., Published as part of Rjosk, A., Neinhuis, C., Monizi, M. & Lautenschläger, T., 2022, Synopsis of the genus Drosera (Droseraceae) in Angola and the Democratic Republic of the Congo, pp. 1-14 in Blumea - Biodiversity, Evolution and Biogeography of Plants 67 (1) on page 7, DOI: 10.3767/blumea.2022.67.01.03, http://zenodo.org/record/7679465, {"references":["Laundon JR. 1970. Droseraceae. In: Wittnich Carisso L (ed), Conspectus Flo- rae Angolensis. Instituto Botanico de Coimbra & British Museum, Lissabon.","Robinson A, Gibson R, Gonella P, et al. 2017. Drosera of the World - volume 3, Latin America, Africa. Redfern Natural History Productions, Poole."]}

Isocitrate dehydrogenase (IDH) mutations are beginning to drive decisions on therapy for glioma patients. Here we sought to determine the impact of adjuvant treatment in patients with IDH-mutant, 1p/19q non-codeleted secondary high-grade astrocytoma (sHGA) WHO grades III/IV. Clinical data of 109 sHGA patients grades III/IV, in addition to IDH mutation-, 1p/19q-codeletion- and MGMT-promoter methylation status-were retrospectively analyzed. Survival analysis in relation to adjuvant treatment modalities and molecular profiling were performed. Out of 109 patients, 88 patients (80.7 %) harbored IDH mutations, 30 patients had a 1p/19q-codeletion (27.5 %) and 69 patients (63.3 %) exhibited a methylated MGMT-promoter status. At a median follow-up of 9.8 years, 62 patients (57 %) died. The postsurgical treatment included: radio-chemotherapy (RT-CT; 54.5 %), RT alone (19.3 %), and CT alone (22.7 %). The median overall survival (OS) in the entire group was 3.4 years (1.9-6.7 years). Patients who received RT-CT had a significantly longer OS compared with those who underwent RT alone (6.5 vs. 1.2 years, HR 0.35, CI 0.32-0.51, p = 0.011). In the IDH-mutant 1p/19q non-codeleted sHGA subgroup the RT-CT cohort had a significantly longer OS in comparison to the RT cohort (6.4 vs. 1.2 years, HR 2.7, CI 1.1-6.5, p = 0.022). In the stepwise multivariable Cox model for OS of all 88 IDH-mutant sHGA patients, survival was strongly associated with only one factor, namely, adjuvant RT-CT at diagnosis of a sHGA. This retrospective long-term study demonstrates that RT and CT (mostly PCV) significantly improves progression-free and overall survival in IDH-mutant secondary high-grade astrocytoma patients, regardless of 1p/19q-codeletion status.