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Ancyropilus exossuarius sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 56A289C5-8C26-45DD-B2D2-C9E6F31AD5AD (Figs 29, 68, 104, 140; Map 1) HOLOTYPE ♀: Queensland: Qld, Plevna Downs. Tompilly Hill base (PD6). 187m 26.725° Sx 142.651°E 16Sept-2Oct2008. Malaise Lambkin, Mackenzie, Starick. Eucalyptus 16278 (QM, type reg. T258546). PARATYPES (10): Queensland: Qld: 26.67° Sx 142.577°E 2.5km WNW Plevna Downs HS (PD3 M) 133m. Gidgee 15 Mar-3Apr 2009. Malaise. R. Mackenzie. 17317 (1 ♀, QM); Qld: Plevna Downs. 12.7km SSE HS (PD8). Malaise 26.786° Sx 142.648°E 16Sept-2Oct2008. Starick, Lambkin, Mackenzie. 145m Eucalyptus 16285 (1 ♁, 2, QM); Qld: 26.541° Sx 142.516°E Plevna Downs, 2.3km SSW Arima (PD2 M) Malaise. 13 Jan-3 Apr 2008. 132m. R.Mackenzie. Gidgee 17284 (1 ♁, QM); Qld: 26.728° Sx 142.651°E Plevna Downs, Tompilly Hill Base (PD6 M) 187m 24Apr-24Jul 2008. G Turner, R Mackenzie. Malaise. Gidgee. 17320. (1 ♀, 1, QM); Qld: 28.813° Sx 144.462°E Currawinya NP. 4km NW HQ (CW3M) dense Mulga. 10-26Sep 2008. 145m. A. Townsend & J. Burke. Malaise trap. 17227 (1 ♀, 1, QM); Qld: 24.135° Sx 143.201°E 3.5km SE HS Noonbah Stn (NB1M) 182m Malaise. Eastern Dead Finish scrub 22Sep-20Oct2008. 17261 Lambkin, Starick, Emmott (1, QM). Diagnosis. Pronotum rounded to weakly tuberculate, disc reflective in appearance, moderately punctate and weakly punctate-rugulose with impunctate areas, central impression not sulcate, tumescent areas either side of central impression indistinct; elytra dark with yellow-orange maculae (each elytron with a small faint humeral macula, a large central fasciate macula which converges slightly towards the suture, plus an apical macula which meets the external and sutural margins), punctation without nodules, 8 th stria beginning anterior to half way between base and fascia (between 2 nd and 7 th punctation of 7 th stria), all ten striae reaching apical macula, basal and apical punctation similar in diameter, often more than one seta across interstrial width (particularly basal half); femora yellow and brown, tarsi with three ventral tarsal pads. Ancyropilus exossuarius sp. nov. is superficially similar in appearance to Notopilo cambageicola sp. nov. (due to pronotal shape, general colouration and elytral maculation), though is easily recognised by the elongation of the elytra (length to width ratio of about 3:1 or longer) and elytral punctation, which lack nodules and do not differ significantly in diameter from the base to apex. Description. Habitus: Fig. 140. Total length: 5.9–8 mm (holotype, 8 mm). Head: Vertex and frons black, clypeus reddish-brown, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown; eyes separated by about 0.53–0.71 eye widths (holotype, 0.65); vertex and upper part of frons punctate-rugulose, lower part of frons (above epistomal suture) transversely rugulose with occasional punctation near eye margins; clypeus with lateral punctation; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 2 times (labium) the length of inside edges; antennae reaching near base of pronotum; eyes and cranium with long erect pale setae, frons and vertex with denser shorter medially-directed setae. Prothorax: Black to brownish, pronotal arch and basal collar often paler than disc, prosternum brownish-black (if pronotum black) or paler brown (if pronotum brown); pronotum about 1.2–1.35 times longer than wide (holotype, 1.24), sides rounded to weakly tuberculate, middle wider than pronotal arch; subapical depression v-shaped, central impression reasonably deep though obscure, not sulcate; disc with a smooth, reflective, general appearance, punctate near middle, punctate-rugulose laterally, a glabrous stripe basally from central impression, shorter semi-glabrous areas either side from base, a weak ovoid partly glabrous tumescence either side of central impression; lateral impression conspicuous, obscurly-shaped, positioned above weak lateral tubercle; disc with fine short multi-directional setae and sparser long erect setae. Pterothorax: Ventrites brown, with fine pale posteriorly- or medially-directed setae; elytra dark brown with yellow-orange markings (each elytron with a small faint humeral macula, a large central fasciate macula which converges slightly towards the suture, plus an apical macula which meets the external and sutural margins), length to width ratio 2.94–3.16:1 (holotype, 3.05:1); 8 th stria beginning anterior to half way between base and fascia (between 2 nd and 7 th punctation of 7 th stria), all ten striae reaching apical macula; punctation circular, without nodules, basal and apical punctation similar in diameter, punctures within striae close together (creating an appearance of neat straight rows of punctures); epipleurae extending into apical maculae; interstriae with short fine setae (usually more than one per puncture), plus erect setae only slightly longer or of similar length (one every two or three punctures), striae often with more than one seta across interstrial width (mostly in basal half); hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 crossvein small (but not completely absent). Legs: Approximately basal three-fifths of femora yellow, remaining parts of femora brown (dark brown then paler brown near joint); tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; meso- and metafemora slender, profemora weakly swollen. Abdomen: Ventrites brown. Male genitalia: Tegmen (Fig. 29) with tegbase slightly wider than parameroid lobes, slightly narrowed between middle and robust parameroid lobes, apex tapered to a digitiform process, dorsal sinus about one-fifth tegmen length, inner margins weakly converging internally, internal limit curved, ventral sinus longer and broader than dorsal sinus, length of apodeme about one-fifth tegmen length; median lobe as in Fig. 68; pygidium as in Fig. 104. Variation. In one male specimen from Plevna Downs (QM code 17284) the spaces between punctures in the longitudinal plane are proportionally larger than in other specimens of the type series, giving the impression that the striae are less orderly. Etymology. The specific epithet exossuarius (from Latin ‘ex’ meaning from and, Latin ‘ossuarius’ meaning place for the bones of the dead) refers to the type locality Plevna Downs, an area of central Queensland known for its rich fossil deposits. Biology. Specimens were collected from January to July and in September and October using Malaise traps in eucalypt and gidgee habitats. Distribution (Map 1). Ancyropilus exossuarius sp. nov. is known from three locations in western Queensland, from as far north as 24ºS, to near the NSW border.

brigalowae species group Diagnosis. Members of the brigalowae species group have at least some elytral interstices with several setae across between puncture rows which, in some species (e.g., A. monteithi sp. nov.), results in the elytra appearing vested with a thick setal mat of short hairs; most elytral striae reaching apices. Remarks. Groupings based on elytral fascia shape (rounded or square at suture) and pronotal form (laterally rounded or angulate) did not correlate with tegminal characteristics. Further attempts to recognise natural groupings within the brigalowae species group based on morphology were, therefore abandoned., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 12, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Monilonotum rufiventre (Gerstmeier) comb. nov. (Fig. 155; Map 2) Olesterus rufiventris Gerstmeier, 1990: 25. HOLOTYPE (♁): New South Wales: Tooloom NSW, 19.1.26 (QM). PARATYPES (2): Queensland: Brisbane, J.S. Rodston, 17.2.41 (2 ♀, QM); Brisbane, H. Hacker, 25.11.13 (1, QM). Other material examined (1): Queensland: SEQ: 25°27′ Sx 150°02′E Boggomoss No. 8, via Taroom. 12 Nov 1996 Qld. Mus.survey. Day hand collection 039 / QM Reg. No. T31633 / Olesterus rufiventris Gerstmeier 1990 Det. J.S. Bartlett 2011 (1, QM). Diagnosis. Among its congeners, M. rufiventre comb. nov. is recognisable by its small size (5.2–6.5 mm), its entirely back elytra, its orange thoracic and abdominal ventrites, and by its puncture-associated pronotal texturing which is conspicuous when compared to the smooth pronota of several small undescribed Monilonotum species. Habitus as in Fig. 155. Remarks. Gerstmeier (1990) provides a detailed description of ‘ Olesterus rufiventris ’.

Ancyropilus hornensis sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 91850EDB-374C-48AE-BB1E-28B7B0E321E2 (Figs 35, 74, 110, 141; Map 1) HOLOTYPE ♁: Northern Territory: Horn Islet Pellew Group, N.T. 15-21 Feb. 1968 B. Cantrell (QM, type reg. T258549). PARATYPES (2): Northern Territory: Horn Islet Pellew Group, N.T. 15-21 Feb. 1968 B. Cantrell (1 ♁, QM); Horn Islet Sir Edward Pellew Group, N.T. 15-21.ii.1968 B. Cantrell (1 ♀, QM). Diagnosis. Pronotum rotund (sub-globose); elytra dark with orange humeral fasciate and apical maculations (humeral maculae less well-defined), 8 th stria beginning near anterior margin of fascia or within fascia, striae reaching apical maculae, interstrial setae not dense as in other Ancyropilus species; femora yellow basally, brown apically, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 141. Total length: 7.3–9.8 mm (holotype, 8 mm). Head: Vertex and frons black to dark brown, clypeus reddish black or brown, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown; eyes separated by about 0.54–0.65 eye widths (holotype, 0.54); vertex and upper part of frons punctate-rugulose, lower part of frons (above epistomal suture) transversely rugulose with occasional punctation near eye margins; clypeus relatively heavily punctate; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillary) and 2 times (labial) the length of inside edges; antennae not reaching base of pronotum; eyes and cranium with long erect pale setae, frons and vertex with denser shorter medially-directed setae. Prothorax: Dark brown, pronotal arch and basal collar generally paler than disc, prosternum dark to pale brown; pronotum robust (sub-globose), about 1.14–1.25 times longer than wide (holotype, 1.14), sides rounded to subtly angulate, middle wider than pronotal arch; subapical depression v-shaped, central impression obsolete; disc moderately punctate though with a smooth reflective general appearance, punctures more dense laterally, a medial glabrous stripe in basal half, base with paired paramedical glabrous tumescences, anterior paramedical tumescent areas partly glabrous; disc with fine short multi-directional setae and sparser long erect setae. Pterothorax: Ventrites dark orange-brown, with fine pale posteriorly- or medially-directed setae; elytra dark brown with yellow-orange markings (each elytron with a weakly defined humeral macula running obliquely towards but not reaching the suture, a broad median fascia that crosses the suture and is broadest laterally—anteriorly meeting the humal macula in the female paratype, plus a trigonal apical macula), length to width ratio 2.62–2.72:1 (holotype, 2.72:1); 8 th stria beginning near anterior margin of median fascia, or within fascia (after about 7–10 punctures from base of the 7 th stria), all ten striae reaching apical macula, or 6 th and/or 10 th striae terminating before apical macula; punctation circular, without nodules, diameter of apical punctation similar or slightly smaller than diameter of basal punctures, striae separated by approximately one puncture diameter; epipleurae extending into apical maculae; interstrial setae generally arranged single-file along striae (two setae across strial width observed in striae posterior of median fascia); intrafoveal setae shorter than puncture diameter in the two observed males and as long as puncture diameter in the only known female specimen. Legs: Approximately basal half of profemora and basal three-fifths of mesoand metafemora yellow, remaining parts of femora brown; tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; meso- and metafemora slender, profemora weakly swollen. Abdomen: Ventrites brown. Male genitalia: Tegmen (Fig. 35), widest near middle, slightly narrowed between middle and parameroid lobes (the apices of which are tapered to a slender, pronounced, slightly inwards-directed digitiform process), dorsal sinus just under one-third tegmen length, ventral sinus about as long as dorsal sinus, apodeme very short, about one-fifthteenth tegmen length; median lobe as in Fig. 74; pygidium as in Fig. 110. Etymology. This species is named after the type locality of Horn Islet, of the Sir Edward Pellow group of islands located in the south-west corner of the Gulf of Carpentaria, Northern Territory. Biology. All three adult specimens were collected in mid-February. Distribution (Map 1). Ancyropilus hornensis sp. nov. is so far known only from Horn Islet, a small rocky landmass about 2 km long, near the larger Centre Island., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 25, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Ancyropilus gen. nov. ZooBank registration: urn:lsid:zoobank.org:act: AD29FEC9-897B-414B-B9BC-672F416BF2DE Gender. Masculine. Type species. Ancyropilus monteithi sp. nov. Diagnosis. Intrafoveal seta short but visible under magnification; ninth elytral stria present; lateral rims of elytral punctures without internal nodules; most elytral striae reaching apices; ventral pad absent from all basitarsi; tegmen ventral sinus equal to, or longer than, dorsal sinus, anterior margin of pre-parameroid area weakly sclerotised; spicular fork V-shaped; median lobe with conspicuous rearward-directed apico-lateral spine. Description. Eyes coarsely-facetted, strongly emarginate above supra-antennal elevations, separated by 0.36– 0.96 eye widths; antennal scape with carina bordering each side of flattened rear face; sensory face of terminal maxillary palpomere approximately 1–2 times longer than inside margin; sensory face of terminal labial palpomere approximately 1.9–3 times longer than inside margin; genae and submentum wrinkled; pronotum 1.1–1.35 times longer than wide, middle broader than anterior pronotal arch (most species) or as broad as pronotal arch (A. monteithi sp. nov. and A. simplex sp. nov.), disc generally heavily punctate (smoother in appearance only in A. exossuarius sp. nov.), central discal impression often obscure, rarely distinctly sulcate; elytra 2.27–3.16 times longer than wide at humeri, punctures without nodules, eighth stria variable, beginning from base to within transverse maculate fascia, all or most striate reaching apical macula, interstriae of most species with more than one seta across strial width (only A. hornensis sp. nov. with rows of singular setae), short, fine, intrafoveal seta present; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 short (most species) or substantially extended (observed only in A. noonbahensis and A. simplex), never completely absent; tarsi each with three well-developed ventral pads; spicular fork V-shaped; tegmen ventral sinus equal to, or longer than, dorsal sinus; median lobe with conspicuous rearward-directed apico-lateral spine. Etymology. The generic epithet Ancyropilus (from Greek ‘ankyra’ meaning anchor, and the genus Opilo) refers to the backward-directed apico-lateral spines of the median lobe evident in all dissected males. Remarks. Conspicuous rearward-directed lateral spines of the median lobe are found in Ancyropilus, Culcipternulus and Infectostriatus, though the spines are less pronounced in the latter genus. Putatively, mutual synapomorphies of Ancyropilus, Culcipternulus and Infectostriatus in relation to eachother could be the loss of elytral puncture nodules in Ancyropilus and Culcipternulus versus the reduction in number of elytral striae in Infectostriatus, plus development of ventral basitarsal pads in Culcipternulus versus reduction in length of tegminal ventral sinus in Ancyropilus and Infectostriatus. Ancyropilus hornensis sp. nov., with unique parameroid lobe apices and reduced interstrial vestiture, is on the other hand retained within Ancyropilus as synapomorphic character development appears to be unidirectional. Key to species of Ancyropilus 1 Each elytral interstice with a neat singular row of setae......... hornensis species group ( Ancyropilus hornensis sp. nov.) - At least some elytral interstices with several setae across interval width (elytra often appearing densely setose as a result)............................................................................... 2 (brigalowae species group) 2 (1) Elytral apices with a small pale macula isolated from outer and sutural margins, or apical macula completely lacking...... 3 - Elytral apices with macula complete to outer and sutural margins............................................... 4 3 (2) Legs entirely black; apical macula completely lacking (Fig. 146)......................... Ancyropilus simplex sp. nov. - Legs not entirely black; apical macula not meeting margins (Fig. 144)............... Ancyropilus noonbahensis sp. nov. 4 (2) Femora predominantly yellow, only extreme base black; lateral bulge of pronotum shallow (Fig. 143).............................................................................................. Ancyropilus monteithi sp. nov. - At least basal half of femora black, and/or pronotum clearly rounded or angulate laterally............................ 5 5 (4) Elytral fascia rounded before or at suture................................................................... 6 - Elytral fascia straight or angulate and crossing suture......................................................... 7 6 (5) Elytra between base and fascia two-tone brown and black (Fig. 147)....................... Ancyropilus tricolor sp. nov. - Elytra between base and fascia unicoloured black (Fig. 138).......................... Ancyropilus brigalowae sp. nov. 7 (5) Elytral fascia extending along suture for a short distance (Fig. 142)......................... Ancyropilus labris sp. nov. - Elytral fascia not extending along suture................................................................... 8 8 (7) Eighth elytral stria beginning between fascia and base.............................. Ancyropilus exossuarius sp. nov. - Eighth elytral stria beginning within fascia, or at anterior margin of fascia........................................ 9 9 (8) Elytra without humeral maculation (Fig. 139)........................................ Ancyropilus emmotti sp. nov. - Elytra with humeral maculation (Fig. 145).................................... Ancyropilus packsaddlensis sp. nov., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 11, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Platynotum bulli sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 1891666B-32FD-4515-82AE-7DFDC6E533C8 (Figs 17, 61, 97, 133, 176; Map 2) HOLOTYPE: Western Australia: Marloo Stn., Wurarga, W.A. 1931-1941. A. Goerling (ANIC). PARATYPES (3): Northern Territory: Stuart H’way, N.T., 296km S of Tennant Creek, 29 Nov. 1972, D.H.Colless (1, ANIC); A. Douglas leg., 26.ix.1964, 25 mi. S. Barrow Cr., N. Territory (1, WAM E88357); Zircon Patch, Harts Range, N.T., 5.10.03., R.M. Bull (1 ♀, JSBC). Western Australia: Australia, WA06/158, 16km n. Leinster, 27.84103S, 120.58300E, 504m, 1.3.2006, M. Baehr (1 ♁, WAM E113543, ex RGCM). Diagnosis. Pronotum tuberculate laterally, disc flattened, impressed in middle, not heavily punctate (most densely punctate laterally); elytra dark with poorly-defined orange humeral, fasciate and apical maculations (apical maculae extremely faint on one specimen), the transverse fascia meeting at the suture, punctation with small lateral nodules (more easily observed at middle fascia), 8 th stria beginning near base, all striae terminating near apex, interstrial setae mostly in single rows (occasionally more than one seta across interstrial width); femora yellow and brown, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 176. Total length: 11–13 mm. Head: Cranium dorsally black, cranium ventrally, clypeus and supra-antennal elevations reddish-black, anteclypeus semi-transparent pale orange, labrum, palpi and antennae darker orange or brownish; vertex moderately punctate, frons smooth medially, punctate laterally; anterior clypeal margin concave; eyes separated by 0.91–1.12 eye widths; genae wrinkled; submentum smooth; terminal maxillary palpomeres with exterior margins about 1.6 (females and males) times the length of inside edges; terminal labial palpomeres with exterior margins about 2.2 (females and males) times the length of inside edges; antennae not reaching base of pronotum; long and short, erect or medially-directed, orange setae, those of vertex and frons fine. Prothorax: Reddish-black; pronotum 1.18–1.26 times longer than wide, anterior part as wide as middle, sides conspicuously constricted between, subapical depression well-defined, v-shaped; disc dorso-ventrally compressed (appearing pressed in) (Fig. 17), central discal impression a linear sulcus; disc appearing smooth, though punctate and/or semi-rugulose in parts; yellowish fine multi-directional setae and thicker long erect setae. Pterothorax: Ventrites orange or brown, with orange posteriorly-directed setae; elytra dark brown, each elytron with ill-defined orange-brown maculate areas (a large basal macula, a fasciate band near middle and some specimens with an apical spot); length to width ratio 2.63–2.78:1; punctation with small lateral nodules (most easily observed at middle fascia), all striae terminating near apex, 8 th stria complete at base; interstriae about as wide as corresponding punctures, with very fine semi-erect setae (sometimes>1 per punctation) and less frequent longer erect setae, setae mostly in singular rows, occassionaly more than one seta across interstrial width; epipleurae extending into apical curve. Legs: Femora except extremities yellow, tibiae, tarsi and femoral extremities brown, ventral tarsal pads yellowish; femora swollen, profemora more than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 61) reasonably slender, slightly constricted in middle, parameroid lobes arcing laterally before almost meeting apically, tegminal arms apparently formed as an extension of a more heavily sclerotised H-shaped internal ‘frame’, dorsal sinus about one-fifth of tegmen length, broad internally, narrowed apically, ventral sinus about as long as dorsal sinus, tegminal arms tapering sharply to meet apodeme near basal one-seventh; median lobe (Fig. 97) with membranous apex; pygidium as in Fig. 133. Etymology. This species is named after Richard Bull in appreciation for the small, but interesting collection of clerids (the paratype from Zircon Patch among them) that he gifted to the first author. Biology. Adults have been collected in October, November and March. Distribution (Map 2). This species is known from four arid zone localities, two each in the Northern Territory and Western Australia., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 66, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Ancyropilus noonbahensis sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: A18F8F55-4DCA-47F5-88F8-B4DAF5546A30 (Figs 31, 70, 106, 144; Map 1) HOLOTYPE ♁: Queensland: Qld: 4km SE HS Noonbah Station (NB2 M), 198m, 24.137° Sx 143.207°E, 19Jan-7Feb2009, Malaise, A. Emmott, Mulga, 17253 (QM, type reg. T258548). PARATYPE: Queensland: same data as holotype (1 ♀, QM). Diagnosis. Pronotum robust, disc very densely punctate except narrow longitudinal stripe from central impression to pronotal furrow lacking punctation; elytra blackish-brown with orange fasciate and apical maculations, fasciae rounded before suture, humeral maculae absent, punctation lacking nodules, 8 th stria almost complete, only slightly shorter than 7 th stria, all striae complete to apical maculae; femora yellow basally, brown apically, only profemora slightly swollen; tarsi with three ventral tarsal pads. Ancyropilus noonbahensis sp. nov. is most similar to Ancyropilus simplex sp. nov. though differs from it by the tegmen with parameroid lobes entirely chitonised externally and U-shaped dorsal sinus, the femoral bases yellow, and the elytral apices clearly marked with a small yellow macula. Description. Habitus: Fig. 144. Total length: 7.2–8 mm (holotype, 7.2 mm). Head: Cranium black (submentum and gula paler in holotype only), clypeus and supra-antennal elevations reddish-black, anteclypeus semi-transparent orange, labrum and pre-ultimate palpomeres orange, terminal palpomeres and antennae darker orange or brownish; vertex and frons with dense network of irregularly-shaped punctation which becomes transversely rugulose before epistomal suture, clypeus more punctate than rugulose basally, smooth apically; anterior clypeal margin broadly concave; eyes separated by 0.69–0.72 eye widths; genae wrinkled; terminal maxillary palpomeres with exterior margins about 2 (females and males) times the length of inside edges; terminal labial palpomeres with exterior margins about 2.5 (females and males) times the length of inside edges; antennae almost reaching base of pronotum; long and short, erect or medially-directed, pale setae. Prothorax: Blackish-brown; pronotum 1.14–1.22 times longer than wide, widest at the middle, sides almost angular in male holotype, more rounded in female paratype, lateral impression obscure, subapical depression v-shaped, not well-defined due distribution of punctation, central discal impression open; disc very densely punctate except a conspicuous slender impunctate longitudinal line from central sulcus to pronotal collar, pronotal arch and collar less densely punctate; pale fine multi-directional setae and thicker long erect setae. Pterothorax: Ventrites black or brownish, metaventrite conspicuously punctate, with short pale posteriorly-directed setae; elytra blackish-brown with orange maculations (each elytron with a small pre-apical macula and a broad fasciate band rounded at the suture); length to width ratio 2.61–2.79:1; punctation large, circular, anterior of fascia, smaller posterior of fascia, without internal nodules, all striae terminating near apex, 8 th stria beginning near base; interstriae about as wide as corresponding punctures, with rows of fine erect setae (> 1/ puncture) and longer erect setae every few punctures; epipleurae extending into apical curve; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 substantialy extended basad of CuA 1 crossvein. Legs: Basal third of profemora and half or just less than half of meso- and metafemora yellow, apical two-thirds of profemora and half or slightly more than half of meso- and metafemora dark brown, tibiae and tarsi brown, ventral tarsal pads brownish-yellow; profemora slightly swollen, other femora more slender. Abdomen: Ventrites orange-brown. Male genitalia: Tegmen (Fig. 31) sinuate between equally broad base and parameroid lobes, parameroid lobes slightly bent inwards, apically with broad acumination, dorsal sinus slightly longer than one-quarter tegmen length, externally wide open, internal shape sub-spatulate, interrupted by opening with short membranous lobes which bear a row of setae, ventral sinus of similar length, apodeme about one-fifth tegmen length; median lobe as in Fig. 70; pygidium as in Fig. 106. Etymology. The specific epithet, noonbahensis, refers to Noonbah Station, a property in western Queensland where the type specimens were collected. Biology. The type series was taken in a single Malaise trap set up over the period 19 January to 7 February in mulga (Acacia) scrub. Distribution (Map 1). Ancyropilus noonbahensis sp. nov. is known only from Noonbah Station in Western Queensland.

Notopilo xanthoimprocerus sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 332D1CD8-12CF-4547-8786-966A9BB0F98A (Figs 46, 84, 119, 160; Map 3) HOLOTYPE ♁: South Australia: S. Aust. 0.2km W Anvil Hole Native Well, Witjira NP, 26°21′27″S 135°42′27″E, pitfalls, 19-25 Nov 1995, Stony Desert Survey AL 03 (SAMA, database No. 25-021893). PARATYPES (3): South Australia: same data as holotype (1 ♀, SAMA); S. Aust. Andamooka, 4.7km SSE Stonwall Dam, 30°44′18″S 137°18′40″E, pitfalls, 11-5 Nov 1996, Stony Des. Surv. AN 002 (2 ♁, SAMA, database Nos. 25-021894, 25-021895). Diagnosis. Pronotum weakly rounded laterally, disc lightly punctate, appearing smooth; elytron yellowish with a brown semi-circular or triangular macula against the suture before the apical slope, punctation with lateral nodules, 8 th stria beginning between 3 rd and 6 th puncture of 7 th stria, most striae terminating near apical quarter, outer striae slightly before, interstriae with short semi-erect, and long erect, setae in single rows, intrafoveal setae long; legs entirely yellow, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 160. Total length: 6.3–7.8 mm (holotye, 6.75 mm). Head: Cranium yellowish or brown, if brown then may be paler ventrally and with frons paler anteriorly; clypeus orange, labrum and palpi yellow, antennae yellowish or yellow-brown; vertex and frons smooth, with only seta-associated punctation, frons broad, eyes separated by 1.3–1.55 eye widths (holotype, 1.3), a pair of shallow impressions between eyes (difficult to observe; most clearly visible on specimen with darker cranium), clypeus mostly smooth, anterior margin moderately concave; genae wrinkled; terminal maxillary palpomeres with exterior margins from 1.5 (females) to 2 (males) times the length of inside edges; terminal labial palpomeres with exterior margins from 2.3 (females) to 2.8 (males) times the length of inside edges; antennae long, reaching beyond base of pronotum, segments elongate, slender; setae pale, eyes with erect setae, cranium with long and short, erect or medially-directed, setae. Prothorax: Entirely yellow; pronotum 1.14–1.23 times longer than wide (holotype, 1.14), sides slightly rounded, middle about as wide as pronotal arch, subapical depression poorly defined, v-shaped; central impression shallow, open; disc appearing smooth, seta-associated punctation inconspicuous; yellowish fine multi-directional setae and thicker long erect setae. Pterothorax: Ventrites yellow, with short yellowish posteriorly-directed and erect setae; elytra yellow, each elytron with a brown semi-circular or triangular macula against the suture before the apical slope; length to width ratio 2.34–2.48:1 (holotype, 2.48:1); punctation with small lateral nodules (high magnification required), punctation appearing larger than actual size due to visibility of subcutaneous component, most striae terminating near apical quarter, outer striae slightly before, 8 th stria beginning between 3 rd and 6 th puncture of 7 th stria; interstriae with fine semi-erect, posteriorly-directed, setae (about every puncture) and longer, thicker, erect setae (every 2–3 punctures), intrafoveal setae spanning entire puncture diameter; epipleurae extending well into apical curve; hindwing with CuA and CuA 1 cross-veins complete, MP 3+4 absent basad of CuA 1 crossvein. Legs: Femora and tibiae yellow, tibial carinae dark for short distance at base, tarsi yellow or brown, ventral tarsal pads yellowish; profemora only slightly more swollen than other femora. Abdomen: Ventrites yellow or pale orange. Male genitalia: Tegmen (Fig. 46) sinuate between tegminal arms and parameroid lobes, the apices of the latter acuminate-digitiform, dorsal sinus about one-third tegmen length, apically very open, ventral sinus about 70% as long; median lobe as in Fig. 84; pygidium as in Fig. 119. 3+4 Etymology. The specific epithet, xanthoimprocerus (from Greek ‘xanthos’ meaning yellow, and Latin ‘improcerus’ meaning short or undersized), refers to this species being the shorter of the two, primarily yellowish, flightless species. Variation. The colour of the head varies from pale yellowish to brown. Biology. Adults have been collected in pitfall traps during November. Distribution (Map 3). Notopilo xanthoimprocerus sp. nov. is known from arid regions of South Australia. Remarks. One dissected male, from Algebuckina Water Hole, South Australia (non-type specimen, SAMA), differs from the three males of the type series in having a more expansive brown triangular elytral maculation and slightly differently proportioned tegminal components. Examination of further specimens is required before the taxonomic status of this ‘form’ can be determined.

Monilonotum bunyense sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: FFA603F4-F2C1-4ABF-B58D-43E5E0081E96 (Figs 39, 78, 151; Map 2) HOLOTYPE (sex not determined): Queensland. Bunya Mtns 10-12-25 H. Hacker (QM, type reg. T258557). PARATYPES (13): Queensland. Bunya Mts. Q. 22.12. 37 N. Geary 3000’ // K 304558 (1, AM); Bunya Mts. Q. 14.12. 37 N. Geary 2000’ // K 304557 (1, AM); Bunya Mts. Q. 2.1. 38 N. Geary 2000’ // K 304559 (1, AM); Bunya Mtns. Q. (26.50S 151.33E) 3 km. from summit on Kingaroy Rd., 6.i.70 light trap, in Nothofagus forest Britton, Holloway, Misko (3, ANIC); H. Elston Collection // Brisbane, Q. Oct. 1920 F.E. Wilson // K 304539 // Opilo congruus Newm. Id. By A.M. Lea variety (1, AM); National Pk. Q. H. Hacker Nov. 1920 // Opilo ID by A.M. Lea (1, QM); Nat. Park Q. (1, QM); Mt Tamborine 29.12.50 C.Oke S.Q. // COL-65680 // COL 65681 (2 specimens on same card-mount, NMV); SEQ: 27°27′ Sx 152°55′E Enoggera Reservoir site3, 16Oct-4Nov 1999 100m C.J. Burwell RF Malaise 50202 (1 ♀, QM). New South Wales. Tooloom, N.S.W. Jan 1926 H. Hacker (1 ♁, QM). Additional material. Queensland. Bundaberg Queensland / A. H. Elston Collection / K 304524 (1, AM). Diagnosis. Pronotum rounded laterally, disc without obvious punctation; elytra predominantly yellowish with dark brown maculate areas basally, post-basally and pre-apically, punctation without nodules, 8 th stria beginning near base, striae terminating just beyond elytral mid-length; femora yellow basally, pale brown apically, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 151. Total length: 8–10.5 mm (holotype, 9.9 mm). Head: Vertex, frons, clypeus, supra-antennal elevations and genae brown, gular region paler, anteclypeus semi-transparent orange, antennae, labrum and palpi orange-brown; eyes separated by 0.88–1.13 eye widths (holotype, 1.08); vertex and frons mostly smooth with only occasional small seta-associated punctations, frons weakly impressed centrally; genae wrinkled, submentum smooth; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 2 times (labium) the length of inside edges; antennae long, reaching beyond base of pronotum; eyes and most of cranium vested with erect pale setae, frons with slightly shorter mediallydirected setae. Prothorax: Dark brown laterally and disc either side of median-line, pronotal arch, pronotal collar and disc medially pale brown to yellowish; pronotum 1.14–1.27 times longer than wide (holotype, 1.21), sides evenly curved, middle slightly broader than lateral part of pronotal arch; subapical depression shallow v-shaped, disc without central sulcus, pronotum with weak lateral impression, surface smooth, almost impunctate; moderately distributed with long erect setae and shorter finer multi-directional setae. Pterothorax: Ventrites orange-brown to dark brown, vested with short, pale, posteriorly-directed, semidecumbent setae; elytra predominantly yellowish with dark brown maculate areas basally, post-basally and pre-apically (basal area to between 5–7 th puncture from base, post-basal area variable, often triangular, joined to basal area along lateral margin, the pre-apical area beginning immediately where punctures become obsolete); length to width ratio 2.75–2.95:1 (holotype, 2.81); 8 th stria beginning near base, all stria terminating abruptly just beyond elytral mid-length, at junction of larger yellow and brown areas, punctation without nodules, interstriae smooth, often narrower than diameter of punctures, epipleurae extending into apical curve, becoming obscure near apex; interstriae with posteriorly-directed, semi-erect setae about as long as distance between centres of adjacent punctures, plus similar setae of slightly longer length, intrafoveal setae extremely fine and short (most easily observable in dark areas of elytra). Legs: Femora yellow basally, brown apically (basal one-third of profemora and basal half of meso- and metafemora yellow), tibiae and tarsi brown, ventral tarsal pads yellowish or orange; apical two-thirds of profemora moderately swollen, other femora very slightly swollen apically. Abdomen: First ventrite yellowish, other ventrites brown. Male genitalia: Tegmen (Fig. 39) broadest at basal opening (tegminal struts), slightly narrowed before apical inward curve of parameroid lobes, apices pointed; dorsal sinus about one-third tegmen length, narrow internally, broadly opening apically; ventral sinus about half dorsal sinus length; apodeme about one-third tegmen length; median lobe as in Fig. 78. Etymology. This species is named after the type locality, Bunya Mountains in Queensland. Variation. A non-type specimen from Bundaberg has the dark brown colouration more broadly distributed on the prontal disc and elytral base. Biology. Specimens were collected in November, December and January, one by light trap, in Nothofagus forest. Distribution (Map 2). So far known from several localities in southeastern Queensland, plus Tooloom in New South Wales., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 31-32, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo katherinensis sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 98BFA4B9-7E6E-4678-A1F6-0405A5568093 (Figs 59, 95, 131, 174; Map 4) HOLOTYPE ♁: Northern Territory: Australia 07, NT35, Scotts Ck. 65km sw Katherine, Victoria Hwy., 14.55.45S, 131.52.66E 126m, 11.11.2007. M.Baehr (QM, type reg. T258554; ex RGCM). PARATYPES (3): Northern Territory: Australia 07, NT33, King R. 30km sw Katherine, Victoria Hwy., 14.42.43S, 132.04.44E 104m, 11.11.2007. M.Baehr (1 ♁, 1, RGCM); Australia 07, NT16, 10km ese Katherine, Stuart Hwy. 14.29.12S, 132.24.04E 100m, 5.11.2007, M.Baehr (1, RGCM); Tindal, N.T. 14.31S 132.22E 1-20 Dec. 1967 light trap W.J.M. Vestjens (1 ♀, ANIC). Diagnosis. Pronotal sides slightly angulate; disc extremely smooth; central impression conspicuous; tumescent areas either side of central impression indistinct; elytra dark with orange maculae (each elytron with a large central globular fasciate macula which is at least slightly curved at the suture plus an apical macula, which meets the external and sutural margins); some punctures with very small lateral nodules (easily misinterpreted as lacking nodules); 8 th stria beginning just anterior of the fascia; striae 1–8 or 1–9 reaching apical macula; striae with a single seta across interstrial width; femora yellow and brown; tarsi with three ventral tarsal pads. Description. Habitus: Fig. 174. Total length: 6–7.2 mm (holotype, 6.2 mm). Head: Vertex and frons black, clypeus and supra-antennal elevations reddish-brown, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown, venter dark reddish brown, gula paler; eyes separated by about 0.53–0.63 eye widths (holotype, 0.55); vertex and lower part of frons mostly smooth, vertex with only a few small punctures, lower part of frons with subtle transverse wrinkles, upper part of frons with some larger punctures; clypeus without strong punctation; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 2.2 times (labium) the length of inside edges; antennae reaching base of pronotum; eyes and cranium with long and short pale erect setae, frons and vertex with occassional long erect setae and shorter medially-directed setae. Prothorax: Disc black infused with reddish brown; pronotum about 1.17–1.19 times longer than wide (holotype, 1.17), sides slightly angulate, middle wider than pronotal arch; subapical depression deeply vshaped; central impression conspicuous; disc extremely smooth in general appearance, punctation sparse, shallow; preapical tumescence indistinct; lateral impression inconspicuous; disc with short multi-directional, and erect, setae. Pterothorax: Ventrites black-brown, with fine mostly posteriorly-directed setae; elytra black-brown with orange markings (each elytron with a large central globular fasciate macula which is at least slightly curved at the suture plus an apical macula, which meets the external and sutural margins), length to width ratio 2.52–2.64:1 (holotype, 2.55:1); 8 th stria beginning just anterior of the fascia, striae 1–8 or 1–9 reaching apical macula; punctation circular, some punctures with minute lateral nodules (easily misinterpreted as lacking nodules; best observed in punctures near base and half way between fascia and apex); epipleurae extending into apical maculae; interstriae with single rows of moderately long to very long, posteriorly-directed, setae, intrafoveal setae short; hindwing with CuA 3+4 cross-vein complete (CuA 1 cross-vein not observed), MP 3+4 absent basad of CuA 1 cross-vein. Legs: Approximately basal three-fifths of profemora, two-thirds of mesofemora and three-quarters of metafemora yellow, remaining apical parts of femora brown; tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; profemora slightly more swollen than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 59) narrowing between middle and the robust parameroid lobes, apex tapered to a short digitiform process, dorsal sinus just over one-third tegmen length, inner margins sub-parallel in middle, opening about twice as broad as innermost width, ventral sinus about half as long as dorsal sinus, apodeme about one-quarter tegmen length; median lobe as in Fig. 95; pygidium as in Fig. 131. Etymology. This species is named after the Katherine region of the Northern Territory. Biology. Adults were collected in November and December. Distribution (Map 4). Notopilo katherinensis sp. nov. is known from only three collecting localities between 10 and 65 km from Katherine, and from Tindal (near Katherine), in the Northern Territory.

Notopilo interfabulatus sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: A990BE5C-AD78-453B-9C11-A67E2D7B9359 (Figs 49, 87, 122, 163; Map 4) HOLOTYPE ♁: Queensland: 15.45S 144.15E GPS 2km NNW “Jowalbinna”, QLD 17 Jan. 1994 P. Zborowski E.D. Ewards, at light (ANIC). PARATYPES (9): Queensland: 17.01S 145.35E QLD Davies Creek Nat. Pk. 5–12 Mar. 1998 A.A. Calder (1 ♁, ANIC); n. Qld Tolga 3 ii 1986 J.D. Brown light trap (1 ♁, QDPC —ex QPIM); 17.39S 145.27E QLD Millstream Falls Nat. Pk. 28 Feb.1998 A.A. Calder / at light open forest (1 ♁, ANIC); 17.39S 145.27E QLD Millstream Falls Nat. Pk. 7 Mar. 1998 A.A. Calder / at light open forest (1 ♀, ANIC); n. Qld Mt. Mulligan, plateau 15-19.iv.1985 K.H. Halfpapp 700m at light (1 ♀, QDPC —ex QPIM); 15.45S 144.15E GPS 2km NNW “ Jowalbinna ”, QLD 17 Jan. 1994 P. Zborowski E.D. Ewards, at light (1, ANIC); n. Qld Tolga 20-xii-1985 J.D. Brown light trap (1, QDPC —ex QPIM); n. Qld Tolga 14.xi.86 J.D. Brown light trap (1, QDPC —ex QPIM); n. Qld 7 km NE of Tolga 7-21.xi.1988 Storey & De faveri light trap (1, QDPC —ex QPIM). Additional material (all males with tegmina and external morphology indistinguishable from those of the type series): Queensland: Iron Range, Cape York Pen., N. Qld 26 May-2 June, 1971 B. K. Cantrell (1 ♁, QM); 11.45S 142.35E QLD Heathlands 20-22 Jun. 1992 T.A. Weir at light (1 ♁, ANIC); Qld: 20°07.0′ Sx 146°37.8′E Rochford Scrub. Site 3. 280m. 11Dec2006 - 11Feb2007. S.Wright, Malaise 14592 vinescrub, lancewood / bendee (1 ♁, QM); Telegraph line Crossing, Jardine R., Cape York, N.Qld. 15-17.vi.1969 G. Monteith (1 ♁, QM). Diagnosis. Pronotum rotund, evenly rounded laterally, disc not heavily punctate; elytra dark with orange fasciate and apical maculations (humeral maculae absent), punctation with nodules, 8 th stria beginning between 1 st and 3 rd puncture of 7 th stria; femora bicoloured; ventral tegminal sinus about one-third the length of dorsal sinus, dorsal sinus margins weakly curving towards eachother (remaining well-separated, not almost touching). Extremely similar to N. lawnhillensis sp. nov. though clearly differentiated by length of ventral tegminal sinus, separation of dorsal sinus margins, and the position of the basal-most puncture of the 8 th elytral stria. Description. Habitus: Fig. 163. Total length: males 7.6–10.7 mm (holotype 7.7 mm); females 8–11.6 mm. Head: cranium black, clypeus and supra-antennal elevation with a reddish hue, anteclypeus, labrum, palpi and antennae orange to orange-brown; eyes separated by 0.58–0.7 eye widths (holotype 0.65); vertex with fine punctures, frons punctate-rugulose, clypeus mostly smooth medially, punctate laterally; genae and submentum heavily wrinkled; ratio of exterior to interior edges of terminal palpomeres approximately 1.8–2.2:1 (maxillae) and 2.5–3:1 (labium); antennae almost reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Blackish to dark-brown, pronotal arch and collar sometimes paler anteriorly; pronotum 1.21–1.29 times longer than wide (holotype 1.25), sides rounded to subtly angulate, middle slightly wider than anterior part; subapical depression v-shaped, central impression conspicuous though not groovelike, disc smooth in overall appearance, punctures shallow, sparse; short fine multi-directional setae and long erect setae. Pterothorax: Ventrites dark-brown, with short posteriorly-directed, and occasional long, setae; elytra black to brown with orange markings (each elytron with a large apical macula meeting external and sutural margins, and a transverse fascia crossing straight at the sutures near elytral mid-length); length to width ratio 2.33–2.64:1 (holotype 2.45); 8 th stria beginning after first, second or third puncture of 7 th stria (most commonly after the third), most striae reaching apical macula (striae 1, 9 and 10 sometimes not reaching macula), punctation with small lateral nodules (punctures with one lateral nodule present on inner rim and, variably, one lateral nodule present or absent on outer rim—best viewed at 50x or higher magnification), punctation posterior of fascia notably smaller than basal punctation, intrafoveal setae short; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 absent basad of CuA 1 crossvein. Legs: Femora yellow basally and black-brown apically (apical third of profemora and apical quarter of mesofemora and metafemora black-brown), tibiae and tarsi brown, tarsal pads yellowish; profemora slightly thicker than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 49) slender, subparallel or very weakly sinuate near middle, parameroid lobes evenly curved towards pointed or vaguely digitiform apices, dorsal sinus about one-third tegmen length, inner margins weakly curving towards eachother though remaining well-separated, ventral sinus one-third as long, apodeme about one-third tegmen length; median lobe as in Fig. 87; pygidium as in Fig. 122. Etymology. The specific epithet interfabulatus (from Latin ‘inter’ meaning amongst or amid, and ‘fabulae’ stories or myths) refers to this species inhabiting a region well-known for its large body of UNESCO-listed prehistoric rock paintings of figurative and abstract depictions of humans, animals and mythical figures called Quinkans. Biology. Specimens comprising the type series were collected from December to June, at light or by use of a Malaise trap. Millstream Falls specimens were taken from open forest. Distribution (Map 4). The type series of Notopilo interfabulatus sp. nov. was selected from specimens collected close to the type location of Jowalbinna near Laura far north Queensland. Male non-type specimens collected outside this region, morphologically indistinguishable from the holotype, suggest this species ranges along eastern Queensland at least from Rochford Scrub, south of Townsville, to the tip of Cape York Peninsula. The related species, N. lawnhillensis sp. nov. is more widely distributed across northern Australia.

Culcipternulus mareebensis sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 97EF3741-9818-47CA-8C44-7533069E1777 (Figs 8, 18, 36, 75, 111, 148; Map 1) HOLOTYPE (sex unknown): Queensland: Qld: 16°58′ Sx 145°26′E Emerald Hill, Mareeba. 6-7 Nov 2004. G.B. Monteith. Open forest. 11680 (QM, type reg. T258550). PARATYPES (3): Queensland: Water trap Bakerville 9- xii-1969 J.D. Brown (1 ♁, QDPC); 15.29S 145.16E Mt. Cook Nat. Pk. QLD 11-12 Oct. 1980 T. Weir (1, ANIC); Queensland. F.P. Dodd. 1904-27. // Opilo congruus Newm. aberr. script. S. Schenkling det. (1, NHML). Additional material. Western Australia: 14.37S 125.48E, 8km SW Walsh Pt., 17 May 1983, WA, I.Naumann, J.Cardale (1 ♁, ANIC); Nita Downs Station 19°05′S 121°41′E Western Australia January 1981 A.M. & M.J Douglas // mostly from swimming pool or vicinity of homestead light. Ex alcohol (1, WAM). Diagnosis. Middle of pronotum rounded, broader than anterior part, disc densely punctate; elytra dark brown with black areas bordering orange fasciate and apical maculae, punctation without nodules, 8 th stria beginning near base, most striae terminating before apical maculae; femora mostly brown though yellow at base, all tarsi with 4 ventral tarsal pads. Description. Habitus: Fig. 148. Total length: 12.2–15.8 mm (holotype 12.4 mm). Head: Cranium reddishbrown, supra-antennal elevations darker, anteclypeus, labrum, palpi and antennae also reddish-brown; eyes separated by 0.9–1.08 eye widths (holotype 0.9); vertex and frons dense with circular to irregular-shaped punctures (surface appearing weakly rugose in parts), frons weakly convex, clypeus smooth with a few circular punctures (seta-associated); genae wrinkled, submentum partly wrinkled (a narrow smooth strip medially); ratio of exterior to interior edges of terminal palpomeres about 2.5:1 (maxillae) and 3:1 (labium); antennae relatively short (compared to other species treated in this revision) not reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Reddish-brown; pronotum 1.16–1.23 times longer than wide (holotype 1.23), sides rounded, widest at middle; subapical depression v-shaped, terminating in a shallow non-sulcate central depression; disc dense with large irregular-shaped punctures, spaces between punctures rugose in appearance, a smooth transverse tumescence either side of central impression, a smooth longitudinal strip behind central impression; short fine multi-directional setae and long erect setae. Pterothorax: Ventrites reddishbrown to dark brown; metaventrite with numerous seta-associated punctures, most setae directed postero-medially; elytra brown and black with orange to yellowish fasciate and apical markings (each elytron with a large orange transverse hatchet-shaped fascia which is broadest laterally, and an orange apical macula which meets the outside margin; the brown areas between base and fascia and fascia and apex infused with black); elytra length to width ratio 2.48–2.55:1 (holotype 2.53); 8 th stria beginning near base; punctation without lateral nodules; punctures large at base, reducing in diameter towards apices (to about half diameter or smaller posterior of fascia), most striae ending before apical maculae; epipleurae virtually obsolete at apical maculae; interstriae with singular rows of fine posteriorly-directed semi-decumbent setae, and less frequent longer erect setae. Legs: Femora yellow basally and brown apically (apical two-thirds to three-quarters of profemora and a little more than the apical half of mesoand metafemora brown), tibiae and tarsi brown, all basitarsi with a small but distinct ventral tarsal pad (Fig. 18), tarsal pads yellowish; profemora slightly thicker than other femora. Abdomen: Ventrites brown. Male genitalia: Tegmen as in Figs 8, 36, tegminal plate with an incurved lateral sclerite at its broadest point, weakly sinuate towards parameroid lobes which neatly taper to a rounded (not digitiform) point distally, dorsal sinus weakly narrowed in middle, about one-quarter tegmen length, ventral sinus about three-quarters as long as dorsal sinus; median lobe as in Fig. 75; pygidium as in Fig. 111. Etymology. This species is named after its type locality, the northern Queensland town of Mareeba. Biology. Specimens have been collected in October, November and December. Distribution (Map 1). Queensland: Atherton Tablelands. Western Australia: Kimberley. Remarks. Specimens from Western Australia cannot be morphologically differentiated from the Queensland specimens constituting the type series. A male from Walsh Point (Mitchell Plateau, WA) was dissected and the tegmen found to differ slightly in proportion (though not in shape) to that of the male from Bakerville (Irvinebank, Qld). The absence of the blackish colouration above and below the fascia observed on the Walsh Point specimen does not correlate with geography as the elytral colouration of the other Western Australian specimen is more like those from Queensland. In the absence of strong evidence for species-status for the Western Australian specimens we consider them, and the Queensland specimens to be disjunct populations of a single species. The Western Australian specimens are not given paratype status in case strong evidence for species-status is presented at a later date., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 27-28, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

cambageicola species group Diagnosis. Hind wing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 absent; outer profile of tegminal parameroid lobes gradually sloping inwards towards apices; apical lobes of median lobe well-developed, with apico-medial sclerite and small apico-lateral acumination (synapomorphic within Notopilo). Remarks. The cambageicola species group of Notopilo contains four species: N. cambageicola sp. nov.; N. interfabulatus sp. nov.; N. lawnhillensis sp. nov.; N. tanybasilaris sp. nov..

Notopilo confusus sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 0D9DD6A0-B7F2-462A-8FC4-592AE092857D (Figs 21, 57, 129, 172, 185; Map 4) HOLOTYPE ♁: South Australia: Adelaide, SA Feb-June 1986, S.Rondonuwu (ANIC). PARATYPES (16): South Australia: Adelaide, SA Feb-June 1986, S.Rondonuwu (11, ANIC); Mt Lofty Rgs. S. H. Curnow // Ex. Coll. S.A. Mus (1, ANIC); A.H. Elston Collection // Adelaide S. Australia A.H. Elston // K 304527 // 74 Opilo congruus Newm. Id by A.M. Lea (1, AM); Adelaide // Museum Paris Coll. Castelnau Coll. Sedillot 1935 (1, MNHN); Gawler S. Australia. // Pascoe Coll. 93-60. (1 ♁, NHML); 29959 // Nov Holl S Austr // Fry Coll. 1900 100. (1, NHML). Additional material. Western Australia: De Boulay // Nov.Holl. Occid. // Fry Coll. 1905-100 (2, NHML *); W. Australia Champion Bay Duboulay // Museum Paris ex Coll. R.Oberthur (1 ♁, 1, MNHN); A.H. Elston Collection // Beverley, WA // K 304535 // 74 Opilo congruus Newm. (1, AM); Yanchep W. Aust., Ex TuART [?] 20.xi.1970, S.J. Curry // Agriculture (Dept) Western Australia 49861 (WADA); R.P. McMillan, Spencers Brook. Sept. 1948 (1, WAM E88346); WA Carlisle (Perth) 31°58′37″S 115°55′24″E 8 February 2000 B.P. Hanich // on outside flywire door at night (1, WAM E88389). * Note: two NHML specimens (see Fig. 185) were discovered to be part of the original syntype series of Opilo femoralis Westwood (see ‘comment on lectotype designation’ under Platynotum femorale stat. rev., comb. nov. below). Diagnosis. Pronotum evenly rounded laterally, disc not heavily punctate; elytra dark with orange fasciate and apical maculations (humeral maculae faint or absent), punctation with nodules, 8 th stria beginning before fascia, at least striae 3–8 reaching apical maculae (intrastrial-punctures may become widely spaced apart towards apex); femora yellow basally, brown apically, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 172. Total length: 7.1–9.6 mm (holotype 7.1 mm). Head (Fig. 21): Cranium black, clypeus and supra-antennal elevation with a reddish or orange hue, anteclypeus, labrum, palpi and antennae orange to orange-brown; eyes separated by 0.56–0.78 eye widths (holotype 0.75); vertex slightly to moderately punctate, frons well-punctated and slightly impressed above narrowest point, punctate-rugulose below, clypeus mostly smooth with a few punctations near hypostomal suture; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.8–2.2:1 (maxillae) and 2.5–3:1 (labium); antennae reaching near base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Blackish to reddish-brown, pronotal arch and collar sometimes paler; pronotum 1.16–1.36 times longer than wide (holotype 1.19), sides rounded to slightly tuberculate, middle about as wide as anterior part; subapical depression vshaped, disc with deep central sulcus (sulcus smooth, more open than linear) and a poorly-defined shallow sulcus on each side, surface with small seta-associated punctations, smooth in overall appearance; short fine multi-directional setae and long erect setae. Pterothorax: Ventrites orange-brown, with short posteriorly-directed, and occasional long, setae; elytra brown with orange markings (each elytron with a large apical macula and a transverse fascia crossing elytral mid-length which is narrowed or curved at the suture, small humeral maculae uncommon); length to width ratio 2.77–3.04:1 (holotype 2.88); 8 th stria beginning between first and sixth puncture of 7 th stria, at least striae 3–8 ending at or near apical macula, punctation with small lateral nodules (most prominent anterior of fascia), punctation posterior of fascia smaller than basal punctation and may be more widely spaced, epipleurae extending into apical maculae, interstriae with erect and angled setae of slightly differing lengths (the shorter semi-decumbent setae seen in other species has become more similar to the erect setae in this species), intrafoveal setae short; hindwing with CuA 3+4 and CuA 1 cross-veins complete (CuA 1 very thin), MP 3+4 absent basad of CuA 1 cross-vein. Legs: Femora yellow basally and brown apically (apical half of profemora, apical two-fifths of mesofemora and apical third of metafemora), tibiae and tarsi brown, ventral tarsal pads yellowish; profemora only slightly thicker than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 57) relatively broad, subparallel or wekly narrowing towards parameroid lobes, the latter tapering to a digitiform process, dorsal sinus about two-fifths tegmen length, narrow internally, wide open externally, ventral sinus one-third as long, apodeme about one-quarter tegmen length; pygidium as in Fig. 129. Etymology. The specific epithet confusus (Latin, meaning confounded, confused or having been brought into disorder) refers to the confusion of Westwood erroneously considering this species conspecific with Opilo femoralis Westwood. Biology. Notopilo confusus sp. nov. has been collected from February to June (SA) and in November (WA). Distribution (Map 4). Specimens are known from several localities between Geraldton in Western Australia and the type locality, Adelaide in South Australia., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 60-61, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo magnus sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: F9E93449-371C-4E30-8F0B-42BD9D2DF232 (Fig. 167; Map 5) HOLOTYPE ♀: Western Australia: Moora W.Australia L.J. Newman // Agriculture (Dept) Western Australia 49870 (WADA). Diagnosis. Antennae short, flagellomeres thick-set; dark with orange to yellowish fasciate and apical maculations (humeral maculae absent), fasciae meeting at suture, 8 th stria beginning near base, all striae terminating before apical maculae; femora entirely dark. Notopilo magnus sp. nov. is easily separated from N. reduncus stat. rev., nom. nov. by its antennae which are comparatively shorter and with thicker flagellomeres, and by the black area between the elytral fascia and apical maculae being no larger than the fascia. Description. Habitus: Fig. 167. Total length: 17.6 mm. Head: Vertex and frons black, clypeus, supra-antennal elevations and submentum reddish-black, gula orange, anteclypeus semi-transparent orange, labrum and antennae dark brown, palpi orange-brown; eyes separated by about 0.8 of an eye width (holotype); frons sculptured, base punctate and semi-rugulose, eyes bordered with denser punctation, deeply sulcate above narrowest point, bulging and partly smooth at and below narrowest point, laterally impressed either side at epipleural suture, clypeus uneven, laterally with small seta-associated punctures near supra-antennal elevations; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1 times (maxillae) and 2.5 times (labium) the length of inside edges; antennae relatively short, not reaching base of pronotum, flagellomeres moderately thickened; eyes and most of cranium vested with erect pale setae, frons with slightly shorter setae, central sulcus with some medially-directed setae. Prothorax: Blackish-brown, venter, pronotal collar and arch (marginally) more reddish; pronotum 1.2 times longer than wide, sides broadly rounded, widest at middle; subapical depression v-shaped, disc centrally sulcate (sulcus smooth, more open than linear, lateral sulci absent, surface smooth in appearance though with noticeable seta-associated punctures; well-distributed with long erect setae and fewer shorter finer setae. Pterothorax: Ventrites reddish-brown, vested with pale, short, posteriorly-directed, setae; elytra blackish with orange markings (each elytron with a large apical macula and a broad fascia which broadens marginally and at the suture), length to width ratio 2.6:1; 8 th stria beginning at base, striae 1–3 and 9–10 terminating within or at the posterior margin of fascia, striae 4–8 incrementally terminating between fascia and apical macula (4 the shortest, 7 or 8 the longest), punctation large up to anterior fascia margin, then punctures of inner and outer striae getting smaller, circular, without nodules, interstriae smooth, epipleurae extending into apical curve, interstriae with very fine short semi-reclinate setae (>1 per puncture) and longer thicker erect setae (Legs: Entirely dark brown, ventral tarsal pads orange; profemora only slightly thicker than other femora. Abdomen: Ventrites orange. Etymology. The specific epithet magnus (Latin, meaning large) refers to the large size of this species. It is only equaled in size by larger specimens of N. reduncus stat. rev., nom. nov. Distribution (Map 5). The unique Holotype was collected in Moora, Western Australia., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 53-54, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Ancyropilus labris sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 83E4BDA5-6482-4754-81FE-75BEC2A47566 (Figs 142; Map 1) HOLOTYPE (sex unknown): Western Australia: Kununurra W.A. 12.xii.1983 A.Postle (ANIC). PARATYPE: Western Australia: Wyndham W.A. 11 Jan 30 T. Campbell (1 ♀, ANIC). Diagnosis. Pronotum angulate (tuberculate) laterally, disc punctate-rugulose laterally, smoother above, either side of central impression strongly tumescent; elytra notably broader than pronotum, dark with orange humeral, fasciate and apical maculations, the transverse fascia widening broadly towards lateral margins and narrowly at suture (anteriorly running along suture longer than posteriorly), punctation without nodules, 8 th stria beginning at 6 th punctation of 7 th stria, all striae terminating at apical macula, more than one seta across interstrial width; femora predominantly yellow basally, black or brown apically, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 142. Total length: 7.45–8.1 mm (holotype, 7.45 mm). Head: Cranium black, clypeus and supra-antennal elevations with reddish hue, anteclypeus semi-transparent orange, labrum and antennae orangebrown, palpi orange; eyes separated by about 0.5 eye widths; frons narrow, more rugulose than punctate, upper part of frons partly glabrous, lower part (above epistomal suture) transversely rugulose; clypeus punctate laterally, smoother medially; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.4 times (maxillae) and 2.3 times (labium) the length of inside edges; antennae not reaching base of pronotum; eyes and most of cranium vested with long erect setae, frons and vertex with shorter medially-directed setae. Prothorax: Black or dark brown; pronotum about 1.15 times longer than wide, robust, sides angular (tuberculate), widest in middle; subapical depression deeply v-shaped, central impression small and deep, a weak impression above lateral tubercle, either side of central impression strongly tumescent, in middle at base weakly tumescent; disc conspicuously punctate-rugulose laterally, disc above, and pronotal arch, smoother with smaller punctation; welldistributed with very fine short setae (sometimes in tuft-like clusters around tumescent parts) and fewer long erect setae. Pterothorax: Sternites brown, with fine pale posteriorly-directed setae; elytra notably wider than pronotum, black or dark brown with orange markings, each elytron with a large apical macula, a broad transverse fascia which widens towards lateral margins and expands just before suture (anteriorly running along suture longer than posteriorly), and a small, less well-defined, humeral macula; length to width ratio 2.27–2.45:1 (holotype, 2.45:1); 8 th stria beginning near anterior margin of fascia, at about 6 th punctation of 7 th stria, all striae reaching apical macula; punctation circular, without nodules; epipleurae extending half way into apical macula; interstriae dense with very short, fine, setae (>3 per puncture, often several setae across interstrial width), plus slightly longer erect setae (Legs: Basal half of profemora and basal three-fifths of meso- and metafemora yellow, femora black or brown apically, tibiae and tarsi brown, ventral tarsal pads orange; profemora slightly swollen, other femora slender. Abdomen: Brown. Male genitalia: Unknown. Etymology. The specific epithet labris (from Greek ‘labrys’, a double-bladed axe) refers to the shape of the elytral fasciate maculae. Biology. Adults have been collected during December and January. Distribution (Map 1). Ancyropilus labris sp. nov. is known only from two specimens collected in the northeastern corner of Western Australia. Remarks. Most abdominal dissections made during the present revision were done so for the purpose of documenting male morphology. The relatively swollen abdomen of the A. labris holotype is far more suggestive of a female than a male. The integrity of the specimen was therefore maintained as the risk of causing damage to the specimen for no diagnostic gain was deemed high., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 16, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo brevistriatus sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 764B7F7D-3EF7-4FB2-AB9F-FC3120248F28 (Figs 54, 91, 126, 169; Map 3) HOLOTYPE ♁: Western Australia: WA: Diamond Tree State Forest (near Manjimup) 34°19′12″S 116°07′15″E 22 February 1995 M.J. Payne // under karri bark (WAM E88406). PARATYPES (18): Western Australia: Karridale, W. Austr., ll.1959 leg. H.Demarz (2, RGCM); K. G. Sound // K 304534 (1 ♀, AM); the “Gloucester Tree”, 4km SE Pemberton, WA, 3 jan 1986, C. Reid, under Eucalyptus bark (1, ANIC); Australia, WA06/181, 10km s. York, 31.97310S, 116.80792E, 190m, 7.3.2006, M.Baehr (1 ♁, WAM E113542, ex RGCM); Australia, WA06/194, 10km wsw. Walpole, 34.99679S, 116.65066E, 66m, 11.3.2006, M.Baehr (1, RGCM); SWA // H. J. Carter Coll. P. 20.4.22 // COL-65654 (1, on card with COL-65655, NMV); same data as previous except COL-65655 (1 on card with COL-65654, NMV); S WA, HIC, 12.13 // Agriculture (Dept) Western Australia 49866 (1, WADA); WA: Quinninup 34°26′S 116°15′E 29 December 1994 M.J. Payne // karri–marri–jarrah forest/ under karri bark (1, WAM E88405); WA: Quinninup 34°26′S 116°15′E 22 December 1995 M.J. Payne // under karri bark (1, WAM E88409); WA: Quinninup 34°26′S 116°15′E 12 December 1995 M.J. Payne (1, WAM E88408); WA: Quinninup 34°26′S 116°15′E 28 November 1995 M.J. Payne // under karri bark regen (1, WAM E88407); Denmark 34.57 S 117.21 E Western Australia 30 jan. 1988 R.P. McMillan (1, WAM E88350); Denmark 34°57′S 117°21E Western Australia December 1998 M.L. & S. Jones (1, WAM E88352); WA: Denmark 34°57′S 117°21′E, 10 January 1997 R.P. McMillan, at MV light at night (1, WAM E88351). Diagnosis. Pronotum rounded laterally, disc smooth with minimal punctation; elytra dark with orange fasciate and apical maculations, humeral maculae absent, punctation without obvious nodules, 8 th stria complete, striae terminating within fascia; femora yellow basally, brown apically, tarsi with three ventral tarsal pads. Notopilo brevistriatus sp. nov. is easily separated from the superficially similar N. variipes comb. nov. by the lack of obvious nodules within elytral punctation, the absence of elytral punctation posterior to fascia, the wider separation of the eyes and by its Western Australian distribution. Description. Habitus: Fig. 169. Total length: 12–16.62 mm (holotype 14.9 mm). Head: Vertex, frons, genae and submentum blackish, clypeus and supra-antennal elevations reddish-black, anteclypeus transparent yellow/ orange, antennae, labrum and palpi orange; eyes separated by 0.85–1.1 eye widths (holotype 0.85); vertex and frons mostly smooth with only occasional small seta-associated punctations, frons slightly raised above clypeus; genae and submentum wrinkled; exterior margins of terminal palpomeres about 2–2.2 times (maxillae) and 2.3–3 times (labium) the length of inside edges; antennae quite long, reaching beyond base of pronotum; eyes and most of cranium vested with erect pale setae, frons with slightly shorter medially-directed setae. Prothorax: Blackishbrown, venter, pronotal collar and arch sometimes paler; pronotum 1.2–1.3 times longer than wide (holotype 1.24), sides round, widest at middle; subapical depression deeply v-shaped, disc centrally sulcate (sulcus smooth, more open than linear) and with obscure lateral sulci, surface smooth, almost impunctate; moderately distributed with long erect setae and shorter finer multi-directional setae. Pterothorax: Ventrites reddish-brown, vested with short pale and occasional longer setae; elytra dark reddish-brown with orange markings (each elytron with a large apical macula and a transverse fascia), length to width ratio 2.95–3.12:1 (holotype 3.06); 8 th stria beginning near base, all striae terminating at or within fascia, punctation large, circular, apparently without nodules (vestigial nodules visible in some specimens as darker pigmented spots on lateral inside edges of punctures), interstices smooth, epipleurae extending into apical curve, interstriae with very fine short semi-reclinate setae (>1 per puncture) and longer thicker erect setae (Legs: Mostly yellow, tip of femora, base of tibiae and tarsi brown, ventral tarsal pads paler; femora slender (profemora thicker but not particularly swollen). Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 54) broadest apically, slightly narrowed between parameroid lobes and tegminal arms, dorsal sinus broad and opening apically, just less than one-third tegmen length, ventral very short, about two-thirds as long, apodeme about one-quarter tegmen length; median lobe as in (Fig. 91); pygidium as in Fig. 126. Etymology. The specific epithet brevistriatus (from Latin ‘brevis’ meaning short, and Latin ‘stria’ meaning line) refers to the shortened elytral puncture rows of this species which are not extended beyond the posterior margin of the transverse fascia. Variation. The dark area posterior of the elytral fascia is almost as dark as the pronotum in some specimens and about as light as the area anterior to the elytral fascia in other specimens. Biology. The specimens from Wilson Inlet were collected at light while the “Gloucester Tree” specimen was found under bark of Eucalyptus. Specimens have been collected during the period December to March. Distribution (Map 3). Notopilo brevistriatus sp. nov. is known only from southwest Western Australia., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 57-58, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

pascoei species group Diagnosis. Beetles ranging from 6.5 to 10.5 mm in length, with the pale transverse elytral fascia pigmented and flat, not formed as a callositous bulla. Remarks. The pascoei species group of Monilonotum contains four known species., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 31, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

variipes species group Diagnosis. Hind wing with CuA 1 cross-vein absent (synapomorphy within Notopilo); CuA 3+4 cross vein present or absent; tegmen with pre-apical broadening of parameroid lobes. Remarks. The variipes species group of Notopilo contains three species: N. brevistriatus sp. nov.; N. gerstmeieri sp. nov.; N. variipes (Chevrolat) comb. nov.., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 56, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

reduncus species group Diagnosis. Large beetles up to about 17 mm; legs unicoloured, dark; elytral punctures lacking internal nodules, striae gradually terminating between fascia and apical maculae (differentiated from the congruus species group externally by the dark legs, and internally by tegminal form—see below). Remarks. The tegmen of N. reduncus stat. rev., nom. nov. is unique in form due to complex dorsal and ventral sinus margins, the dorsal layers of which form an acute rear-ward projection. The incurvate RP vein of the hind wing of N. reduncus is also unique among species treated in this revision (synapomorphic within Notopilo). The non-nodulate elytral punctures, large body size and general habitus of N. magnus sp. nov. suggests close relatedness to N. reduncus, however, examination of the hind wing and dissection of the terminalia, required for confirmation, have not been undertaken to avoid damage to the unique holotype. Notopilo magnus is therefore listed here only tentatively., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 53, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Monilonotum pascoei (Gorham, 1876) comb. nov. (Figs 22, 41, 79, 114, 153; Map 2) Opilo pascoii Gorham, 1876: 71. Type material examined. LECTOTYPE (here designated): New South Wales: Type (printed on a white circular label with red border) // N.S. W. (handwritten on a green oval-shaped label) // Gorham Type (printed on red card) // Pascoe coll. 93-60 // Opilo pascoei Gorh. (handwritten) (NHML). Comment on lectotype designation. As Gorham (1876) made no reference to specimens in his description it cannot be assumed that the nominal species group name Opilo pascoei was based on a single specimen. The specimen labelled ‘type’ in the Natural History Museum, London (NHML) must therefore be considered a syntype (ICZN 1999: Article 73.2) and available for lectotype designation (ICZN 1999: Article 74.1). Other material examined (14 specimens): Victoria: A.H. Elston Collection // Beaconsfield, V F. E. Wilson 26.12.19 // K 304538 (1 ♀, AM); Victoria Redhill 13 Jan 1974 // O. pascoei Gorh. // E. Gowing-Scope collection BMNH(E) 2005-4 (1 ♀, NHML); Woori Yallock Vict. 13.1.26 // F. E. Wilson Collection // COL-65672 // COL-65673 (2 ♀ on same card-mount, NMV); 2657 // Nat. Mus. Victoria C. Frech’s Coll 5.11.08 // Opilo pascoei Gorh. Victoria // COL-65675 (1 ♀, NMV); Vic // COL-65671 (1 ♀, NMV); Kosciusko Vic 2.26 (1 ♁, ANIC); Macedon, Vic C. Oke // COL-65667 (1 ♁, NMV); Riddell, V. 1.1.27 C. Oke // COL-65668 // COL-65669 (2 ♁ on same cardmount, NMV). New South Wales: Mt Murray Rd Macquarie Pass NSW 19.i.86, Leptospermum // S.G. Watkins Collection Donated 2001 (1 ♀, ANIC); West Pymble, nr Sydney N.S.W. 22.i.1983 D.J. Scambler // K 304577 (1 ♀, AM); Australia: NSW Mt Keira (west flank) 27 Nov 1994 A. Sundholm; On flowers of Acmena smithi (1 ♁, ANIC). Australia Capital Territory: Orroral R. ACT 25 July 1982 K.R. Pullen // under bark Eucalyptus // Kim Pullen Collection (1 ♀, ANIC). Diagnosis. Pronotum rounded laterally, disc smooth, without conspicuous punctation; elytra predominantly dark brown with the entire basal third to two-fifths orange-brown and a whitish transverse outward sloping fasciae and a whitish triangular apical maculae, punctation without nodules, 8 th stria beginning just behind humeral tumescence, striae terminating at posterior margin of transverse fascia; coxae, trochantins and femoral base paler than majority of femora, tarsi with three ventral tarsal pads. Monilonotum pascoei is easily confused with M. sundholmi sp. nov. though may be recognised by the colouration of its coxae, trochantins and femoral bases, which are paler than the ventrites and remainder of legs; the elytral intervals (spaces between punctures) of M. pascoei are wider and more rounded than those of M. sundholmi sp. nov. which tend to be narrow and ridge-like. Description. Habitus: Fig. 153. Total length: 7.7–9.7 mm. Head: Vertex and frons black, clypeus, supraantennal elevations and genae black-brown, gular region paler, anteclypeus semi-transparent orange, antennae, labrum and palpi orange-brown; eyes separated by 1.2–1.6 eye widths; vertex and frons only lightly punctate; genae wrinkled, submentum smooth; exterior margins of terminal palpomeres about 1.7–2.3 times (maxillae) and 2.8–3 times (labium) the length of inside edges (the larger terminal maxillary palpomeres being those of males); antennae long, reaching to or slightly beyond base of pronotum; eyes and most of cranium vested with erect pale setae, frons with slightly shorter medially-directed setae. Prothorax: Dark brown except pronotal collar and pronotal projection whitish or pale yellow; pronotum 1.14–1.27 times longer than wide, sides evenly curved, middle slightly broader than lateral part of pronotal arch; subapical depression v-shaped, central impression of disc smooth, flowing from subapical depression, not sulcate, pronotum with weak lateral impression, surface smooth, almost impunctate, punctation consisting of sparse inconspicuous (shallow and smooth) setose impressions; moderately distributed with long erect setae and shorter finer multi-directional setae. Pterothorax: Ventrites orange, vested with short, pale, posteriorly-directed, semi-decumbent setae; elytra (Fig. 22) three-toned (each elytron with basal third to two-fifths orange-brown this area often sloping posteriorly toward suture, posterior three-fiths to two-thirds black-brown, interrupted by a whitish transverse fascia, which slopes slightly away from the suture, and a triangular apical macula); length to width ratio 2.55–2.86:1; 8 th stria beginning immediately behind humeral tumescence, all stria terminating abruptly at posterior margin of whitish transverse fascia, punctation without nodules, epipleurae extending into apical curve, becoming obscure near apex; interstriae with posteriorly-directed, semi-erect setae about as long as distance between centres of neighbouring punctures, plus erect setae of slightly longer length, very small intrafoveal setae seldomly observed (apparently absent from most punctures). Legs: Almost entirely orange-brown, hind coxae, trochanters and extreme base of metafemora paler (orange, yellow or whitish); all femora swollen in males (most notably the metafemora), profemora only swollen in females (other femora slender). Abdomen: Basal half of first ventrite pale (orange, yellow or whitish), other ventrites black-brown. Male genitalia: Tegmen as in Fig. 41; apex of parameroid lobes narrowing unevenly to a point, inner margin indented; dorsal sinus slightly shorter than onethird tegmen length, apical third gradually broadening with profile of parameroid lobes, inner two-thirds reasonably broad (in comparison with M. doddi and M. sundholmi), sub-parallel; ventral sinus about two-thirds as long as dorsal sinus; apodeme about one-third tegmen length; median lobe as in Fig. 79; pygidium as in Fig. 114. Variation. In contrast to females, males have noticeably swollen femora, larger terminal maxillary palpomeres and the slightly more bulging eyes. Biology. Plant associations include Eucalyptus, Leptospermum and Syzygium (Myrtaceae). An ACT specimen, collected in July under eucalypt bark was probably taking refuge over winter there; the collection times of other specimens (November to February) are likely more representative of when adults are active. Distribution (Map 2). Vic, ACT and NSW: all known localities fall within the Temperate Broadleaf and Mixed Forest terrestrial ecoregions as defined by the National Reserve System (DAWE 2012)., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 34-35, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["Gorham, H. S. (1876) Notes on the coleopterous family Cleridae, with descriptions of new genera and species. Cistula Entomologica, 2, 57 - 106.","ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp.","DAWE. (2012). National Reserve System: Terrestrial Ecoregions of Australia. Department of Agriculture, Water and the Environment (formerly Department of Sustainability, Environment, Water, Population and Communities), Canberra, Map. Available from: https: // www. awe. gov. au / agriculture-land / land / nrs / science / ibra / australias-ecoregions (accessed 6 March 2022)"]}

rufiventre species group Diagnosis. Smaller beetles to 7 mm, with the pale transverse elytral fascia formed as a callositous bulla. Remarks. The rufiventre species group of Monilonotum contains two described, and several undescribed species., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 36, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Culcipternulus gen. nov. ZooBank registration: urn:lsid:zoobank.org:act: 949D3490-731A-46C6-968D-9F11CE8FCCCC Gender. Masculine. Type species. Culcipternulus mareebensis sp. nov., by present designation. Diagnosis. Punctures of at least elytral striae 1–3 terminating before apical macula; interstriae with neat singular row of hairs; all basitarsi with a small but distinct ventral tarsal pad (Fig. 18); tegmen with sides gradually narrowing towards distal end of phallobasic apodeme, middle notably broader than parameroid lobes, ventral sinus shorter than dorsal sinus. MAP 1. Species distribution for Ancyropilus and Culcipternotus, with colour legend to National Reserve System Terrestrial Ecoregions (topographic layer image credit: NASA 2002). Description. Eyes coarsely-facetted, strongly emarginate above supra-antennal elevations, separated by 0.9– 1.08 eye widths; antennal scape with carina bordering each side of flattened rear face; sensory face of terminal maxillary palpomere approximately 2.5 times longer than inside margin; sensory face of terminal labial palpomere approximately 3 times longer than inside margin; genae wrinkled, submentum partly wrinkled; pronotum 1.16–1.23 times longer than wide, middle broader than anterior pronotal arch, disc with relatively heavy punctures, central discal impression not distinctly sulcate; elytra 2.48–2.55 times longer than wide at humeri, punctures without nodules, eighth stria beginning near base, most elytral striae terminating before apices, interstrial setae arranged in simple rows (not with several setae across strial width), intrafoveal setae present; hindwing with CuA 3+4 and CuA 1 cross-veins complete; tarsi each with four well-developed ventral pads (pads of basitarsi smaller, though wellformed); tegmen ventral sinus about three-quarters as long as dorsal sinus; median lobe with conspicuous rearwarddirected apico-lateral spine. Etymology. The generic epithet Culcipternulus (a combination of Latin ‘culcita’ meaning cushion or pillow, Greek ‘pterna’ meaning heel, and the diminutive suffix ‘-ulus’), refers to the small ventral pads present on all basitarsi, a condition unique among the genera treated in this revision. Remarks. Culcipternulus is represented in collections by only six specimens, potentially constituting two species seperable only by genetalic characters. One of these forms is described below, with formal designation of species-status for the other pending further investigation., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 26-27, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["NASA (2002) NASA Visible Earth. Blue Marble: Land Surface, Shallow Water, and Shaded Topography. EOS Project Science Office, NASA Goddard Space Flight Centre, Greenbelt. Available from: https: // visibleearth. nasa. gov / images / 57752 / bluemarble-land-surface-shallow-water-and-shaded-topography (accessed 25 April 2022)"]}

hornensis species group Diagnosis. Most elytral striae terminating near apical macula; interstriae with neat singular row of hairs; tegmen with narrowly digitiform parameroid lobes, ventral sinus equal in length to dorsal sinus, middle slightly broader than parameroid lobes., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 25, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo gen. nov. ZooBank registration: urn:lsid:zoobank.org:act: CE086160-0DDF-41F7-9052-C2EDC95FF754 Gender. Masculine. Type species. Opilo congruus Newman, 1842, by present designation. Diagnosis. Antennal scape with a carina bordering each side of the rear face; elytral intrafoveal setae short but visible under magnification; elytral interstrial setae arranged in simple rows; ninth elytral stria present; pronotum rotund, or if appearing flattened then it is widest anteriorly (not in the middle); basitarsi without a distinct ventral pad; tegmen with ventral sinus shorter than dorsal sinus; apical lobes of median lobe without large rearward-directed spine or barb (a minute acumination present only in N. congruus comb. nov. and species of the cambageicola species group). Description. Eyes coarsely-facetted, strongly emarginate above supra-antennal elevations, separated by 0.53– 2.0 eye widths; antennal scape with carina bordering each side of flattened rear face; sensory face of terminal maxillary palpomere approximately 1.0–2.3 times longer than inside margin; sensory face of terminal labial palpomere approximately 1.8–3.0 times longer than inside margin; genae and submentum wrinkled; pronotum 1.06–1.36 times longer than wide, broadest near middle or middle as wide as anterior pronotal arch, disc lightly to moderately punctate, median sulcus conspicuous or inconspicuous; elytra 1.4–3.15 times longer than wide at humeri, punctures with or without nodules, start of eighth stria variable (from base to near middle), at least some discal striae terminating between posterior margin of fascia and apices (most species) or all striae terminating close to fascia (variipes species group), interstrial setae in single rows, intrafoveal setae mostly short, less commonly spanning across entire puncture; hindwing with crossvein CuA 3+4 complete (most species) or absent (N. variipes comb. nov., N. xanthoprolatus sp. nov.), and crossvein CuA 1 complete (most species), incomplete (N. cambageicola sp. nov. and N. reduncus stat. rev., nom. nov.) or absent (variipes species group), MP 3+4 absent basad of CuA 1 crossvein; tarsomeres 2–4 each with a well-developed ventral pad; tegminal ventral sinus one-third as long to as long as dorsal sinus (never longer than dorsal sinus); apical lobes of median lobe generally plate-like, lacking conspicuous rearwards-directed apico-lateral spine (a minute lateral acumination present amongst members of the cambageicola species group and in N. congruus comb. nov.). Etymology. The generic epithet Notopilo is an informal combining of not (a shortened form of the Greek notios, meaning ‘southern’, in reference to Australia, the ‘great southern land’) and opilo (from the Latin opilio, meaning ‘shepherd’, though used here in reference to the genus Opilo Latreille to which three of the included species had been assigned for more than a century). The name Notopilo also implies that its constituent species are ‘not’ congeneric with Opilo mollis (Linnaeus), the type species of Opilo. Remarks. The monophyly of Notopilo is putatively supported by the complete absence of a remnant of hindwing vein MP 3+4 basad of crossvein CuA 1 (very short in other genera, but consistently absent only in the species of Notopilo). Sixteen of the twenty species assigned to Notopilo gen. nov. have been further assigned to one of five relatively homogenous species groups for which synapomorphies have been identified. Relationships of the four remaining species (see ‘unplaced to species group’ below) with each other and with the other Notopilo species remain undetermined. The decision not to assign genus status to each of the species groups was made because erecting several monotypic genera for the remaining species was deemed not to be a realistic solution. Consequentially, the relatively taxonomically conservative concept of Notopilo put forth here, has resulted in a genus that is less-obviously homogeneous than the other five genera proposed in this revision. Key to species of Notopilo (genitalia-based species groups not indicated) 1 Elytra with at least some punctures internally nodulate....................................................... 2 - Elytral punctation lacking internal nodules............................................................... 17 2 (1) Elytra with the three or four inner-most striae terminating within transverse fascia, outer striae extending slightly beyond fascia, at least some punctures nodulate; males with mat of short decumbent setae at striae 1–3 within, and just posterior of, fascia (Fig. 171).................................................... Notopilo variipes (Chevrolat) comb. nov. - All elytral striae extending beyond fascia, or elytra not fasciate; males without setal mat............................ 3 3 (2) Eyes separated by at least 1.2 eye widths; pronotal anterior margin as broad as, or broader than, middle................ 4 - Eyes separated by no greater than one eye width; pronotal anterior margin rarely as broad as middle, typically at least slightly narrower........................................................................................... 9 4 (3) Elytra colour predominantly pale (Figs 158, 160, 161)....................................................... 5 - Elytra colour predominantly dark........................................................................ 7 5 (4) Each elytron with a single pre-apical semi-circular or triangular macula at the suture (Fig. 160).............................................................................................. Notopilo xanthoimprocerus sp. nov. - Each elytron with a broad pre-apical band and a post-basal spot or fascia......................................... 6 6 (5) Each elytron with a post-basal fascia that runs along suture towards scutellum; elytra not particularly elongate (length to width ratio 2.71:1) (Fig. 158)........................................................... Notopilo eremosus sp. nov. - Each elytron with a post-basal spot that does not run along suture; elytra notably elongate (length to width ratio range 2.88– 3.13:1) (Fig. 161).......................................................... Notopilo xanthoprolatus sp. nov. 7 (4) Pronotal anterior margin conspicuously broader than middle; elytra elongate (length to width ratio range 2.92–2.95:1) (Fig. 157)............................................................................ Notopilo calicis sp. nov. - Pronotal anterior margin about as broad as middle or slightly narrower; elytra less elongate (length to width ratio range 2.4–2.6:1).......................................................................................... 8 8 (7) Legs entirely black; elytral pale maculae typically glossy and enamel-like in appearance (Fig. 156)................................................................................................ Notopilo beswickensis sp. nov. - Legs bicoloured; elytral pale maculae never glossy and enamel-like in appearance (Fig. 159)..................................................................................................... Notopilo tompricensis sp. nov. 9 (3) Elytral transverse fascia positioned posterior of elytral mid-length, with its anterior margin at the mid-point or nearer to the apices than the base; profemora entirely, or almost entirely, black (Fig. 170)............... Notopilo gerstmeieri sp. nov. - If elytral transverse fascia as above, then profemora basally pale.............................................. 10 10 (4) Humeral macula present and joined to elytral fascia along lateral margin proximal to epipleuron... Notopilo elstoni sp. nov. - Humeral macula, if present, not joined to elytra fascia along lateral margin...................................... 11 11 (10) Elytra lacking apical maculation, broad and short (length to width ratio 2:1), notably broader than pronotum (Fig. 175)..................................................................................... Notopilo obesus sp. nov. - Elytra more elongate (length to width ratio at least 2.3:1 or greater), apical macula present.......................... 12 12 (11) Elytral fascia conspicuously rounded at suture (Fig. 174); nodules of elytral punctures small and difficult to view, or apparently absent.................................................................... Notopilo katherinensis sp. nov. - Not as above, or if elytral fascia rounded at suture then nodules of elytral punctures easily viewable under magnification..................................................................................................... 13 13 (12) Length to width ratio of pre-fasciate area of single elytron ranging from 2.2:1 to 2.7:1 (Figs 165, 172)................ 14 - Length to width ratio of pre-fasciate area of single elytron ranging from 1.7:1 to 2:1 (e.g., Fig. 164).................. 15 14 (13) Elytral stria 10 (sometimes also 9) terminating before apical macula....................... Notopilo confusus sp. nov. - Elytral striae 9-10 terminating near or within apical macula........................... Notopilo tanybasilaris sp. nov. 15 (13) Elytral humeri maculate (whether distinct or faint, humeral macula never entirely absent), entire base never maculate (Fig. 162); nodules of elytral punctures distinct........................................ Notopilo cambageicola sp. nov. - Elytral humeri lacking maculation (as in Figs 163–165), or entire elytral base maculate; nodules of elytral punctures sometimes indistinct.......................................................................................... 16 16 (15) Eighth elytral stria beginning between the 4 th and 7 th puncture of the 7 th stria, elytral humeri not maculate (typical form, Queensland/NT to Pilbara into Gascoyne, WA) or elytral base entirely maculate (WA variety, south of Pilbara /Gascoyne border); length of ventral tegminal sinus about half the length of dorsal sinus (Fig. 50)...... Notopilo lawnhillensis sp. nov . - Eighth elytral stria beginning between the 1 st and 3 rd puncture of 7 th striae, elytral humeri never maculate; ventral tegminal sinus about one-third the length of dorsal sinus (Fig. 49); north Queensland (from Townsville region to tip of Cape York Peninsula)................................................................. Notopilo interfabulatus sp. nov . 17 (1) All elytra striae terminating within fascia (Fig. 169)................................. Notopilo brevistriatus sp. nov. - Elytra striae terminating posterior of fascia............................................................... 18 18 (17) Legs bicoloured.............................................................. Notopilo congruus comb. nov. - Legs black......................................................................................... 19 19 (18) Antennal flagellomeres slender.......................................... Notopilo reduncus stat. rev., nom. nov. - Antennal flagellomeres thick-set.................................................... Notopilo magnus sp. nov., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 36-38, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["Newman, E. (1842) List of insects collected at Port Phillip, South Australia, by Edmund Thomas Higgins, Esq. The Entomologist, 1 (23), 361 - 369."]}

Infectostriatus absentis sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 93B88442-023B-4276-9296-3B77A04C5EF4 (Figs 15, 37, 76, 112, 149; Map 2) HOLOTYPE ♁: Queensland: Tolga, N. Qld, 26.iii.1984, J.D. Brown, light trap (QM, type reg. T258556). PARATYPES (13): Queensland: Coen, N.Q., 5/72 GB // J.G. Brooks Bequest, 1976 (1, ANIC); Cooktown, Queensland // Museum Paris ex Coll R. Oberthur (1, MNHN); Station Ck, 12km N Mt. Molloy, N Qld, 9 Jan. 1970, Walford-Huggins // ex collection A. Walford-Huggins // E. Gowing-Scope collection BMNH(E) 2005-4 (1, NHML); Parada, July ‘65 // M 296 (1♀, QDPC); Tolga, N. Qld, 20.v.1983, J.D. Brown, light trap (1, QDPC); n. Qld, Tolga, 8 iv 1986, J.D. Brown, Light trap (1, QDPC); Tolga, N. Qld, 1-v-1984, J.D. Brown, light trap (1♁, QDPC); n. Qld, Tolga, 8-15 iii 1985, J.D. Brown, light trap (1♁, QDPC); n. Qld, 7 km NE of Tolga, 5.1.1987, Storey & De Faveri, light trap (1, QDPC); n. Qld, 7 km NE of Tolga, Feb 1987, Storey & De Faveri, light trap (1, QDPC); n. Qld, 7 km NE of Tolga, Mar 1987, Storey & De Faveri, light trap (2♁, QDPC); n. Qld, 7 km NE of Tolga, Apr 1987, Storey & De Faveri, light trap (1, QDPC). Diagnosis. Infectostriatus absentis sp. nov. is most easily differentiated from I. differens sp. nov. by the absence of punctation between the seventh and tenth striae between the base and elytral fascia. Description. Habitus: Fig. 149. Total length: 6.7–9.7 mm. Head: Vertex, frons, genae and gula black, clypeus and supra-antennal elevation dark reddish-brown, anteclypeus, labrum, antennae and palpi orange-brown or chestnut-brown; frons narrow (eyes separated by 0.5–0.8 eye widths), weakly impressed above narrowest point, frontal sculpturing confused-foveolate, often transversely wrinkled below narrowest point, genae wrinkled; exterior margins of terminal palpomeres about 1.6 times (maxillae) and 3 times (labium) the length of inside edges; antennae reaching near base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Pronotum dark reddish-brown to reddish-black, darkest laterally, sometimes paler at anterior and posterior margins and ventrally; pronotum 1.24–1.3 times longer than wide, anteriorly slightly narrower than head, sides impressed above conspicuous lateral tubercle, subapical depression deeply v-shaped, disc weakly tuberculate behind prontal arch, sulcate centrally; surface smooth with only occasional small seta-associated punctures, well-vested in short fine multi-directional setae and occasional long erect setae. Pterothorax: Ventrites orange-brown, impunctate, moderately vested with short pale and occasional long setae; elytra (Fig. 15) brown with yellowish markings (a large apical macula, a broad ‘stepped’ central fascia and a variable irregular sub-basal macula—sometimes thin and running obliquely from the suture towards the fascia and joined to it along the suture, or sometimes broadly connected to the fascia and enclosing a small dark spot), length to width ratio 2.6–2.8:1, punctation with lateral bead-like nodules, 8 th and 9 th striae highly reduced, evident only within dark area between fascia and apical macula, 8 th stria beginning between the tenth and fifteenth puncture of the 7 th stria, the 9 th stria represented by a few punctures only, punctation relatively large until near apex; disc well-vested in very fine short and thicker erect setae, short setae on anterior and lateral perimeter of punctations and within interstriae, longer setae in rows following interstriae, epipleurae extending well into apical curve; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 crossvein small (but not completely absent). Legs: Femora bicoloured, basally yellow (basal half of profemora, basal three-quarters of meso- and metafemora), apically brown, tibiae and tarsi brown, ventral tarsal pads yellowish; profemora slightly swollen, other femora slender. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 37) robust, slightly narrowing before parameroid lobes, the latter angularly expanded laterally, bent inwards and blunt (not pointed) apically, dorsal sinus very wide, just over one-quarter tegmen length, slender transparent (sensory?) membrane apparent at broadest point, ventral sinus almost as long, tegminal arms sharply meeting apodeme, apodeme just over quarter tegmen length; median lobe as in Fig. 76; pygidium as in Fig. 112. Etymology. The specific epithet absentis (Latin, meaning absent or missing) refers to the total absence of basal punctures between the seventh and tenth elytral striae. Biology. Adults have been collected from January to July, many by light trapping. Distribution (Map 2). Atherton Tableland (Parada, Tolga, Station Creek), Cooktown and Coen in far north Queensland.

Ancyropilus simplex sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: FA9C15ED-CF77-4BF3-A033-E401928ACE3E (Figs 33, 72, 108, 146; Map 1) HOLOTYPE (sex not determined): Western Australia: 22.09S 118.07E GPS, 25km NNW of Wittenoom Gorge, WA, 4 Oct. 1995, D.C.F.Rentz & L.Lowe, Stop 92 (ANIC). PARATYPES (42): Western Australia: same data as holotype (1 ♀, 1, ANIC); 25 km NW Bandya Stn., W. Aust., 2.xi.1989, K.T. Richards // Agriculture (Dept.) Western Australia 49857 (1 ♁, WADA); Carnarvon WA., 25-II-77, M.S. Moulds // Opilo congruus Newm. A. Walford Huggins det. 1983 (1 ♀, QM); 107 miles SSE of Carnarvon, WA, 21 April 1968, I.F.B. Common & M.S. Upton (1 ♀, 4, ANIC); Australia, WA Menzies, 10 km North, 14 January 1989, M.S. Moulds // K 304506 (1, AM); WA, Pilbara region. Juna Downs Station, Great Northern Highway, approx. 8km S of Karijini Dr toff., -22:41:36, 118:42:19, 3- Jan-2006 to 8-Jan-2006, LTM sites, A. Donnelly & CVA Volunteer PILB038/11M (Malaise trap) // 1750 // K 246911 (1, AM); WA, Pilbara region. Hamersley station, Horseshoe Bore, 300m N of windmill, -22:29:10, 117:41:28; 16- Feb-2005, 20-Feb-2005, LTM sites, M. Bulbert & S. Ginn, PILB044/04M // 1135 // K 246910 (1, AM); WA, Pilbara region. Hamersley station, Horseshoe Bore, 300m N of windmill, -22:29:10, 117:41:28, 3-Jan-2006 to 8-Jan-2006, LTM sites, A, Donnelly & CVA volunteer, PILB044/11M (Malaise trap) // +2 // 1749 // K 246912 (1, WAM E113528, ex AM); WA, Pilbara region. Juna Downs Station, Juna Downs Rd to Packsaddle Bore approx. 5km E of homestead., -22:52:31, 118:31:49, 19-Nov-2004 to 24-Nov-2004, LTM sites, CVA volunteers, PILB039/03M (Malaise trap) // 1140 // K 246905 (1 ♁, AM); WA, Pilbara region. Juna Downs Station, Juna Downs Rd to Packsaddle Bore approx. 20km E of homestead, -22:51:30, 118:40:14, 3-Jan-2006 to 8-Jan-2006, LTM sites, A. Donnelly & CVA volunteer, PILB036/11M (Malaise trap) // 1753 // K 246898 (1, WAM E113529, ex AM); WA, Pilbara region. NanburraWittenoom Rd, approx 13km NE of Railway Rd. x-ing, nr fenceline, -22:26:8, 117:49:56, 18-Nov-2004 to 23-Nov-2004, LTM sites, CVA volunteers, PILB041/03M (Malaise trap) // 1137 // K 246908 (1, AM); same data as previous except: 1138 // K 246907 (1 ♁, WAM E113530, ex AM); same data as previous except: 1139 // K 246906 (1 ♁, WAM E113531, ex AM); WA, Pilbara region. Nanburra-Wittenoom Rd, approx 13km NE of Railway Rd. x-ing, nr fenceline, -22:26:8, 117:49:56, 28-Oct-2005 to 2-Nov-2005, LTM sites, CVA volunteers, PILB041/10M (Malaise trap) // 2030 // K 246904 (1, AM); same data as previous except: 2031 // K 246913 (1, WAM E113532, ex AM); WA, Pilbara region. Nanburra-Wittenoom Rd, approx 13km NE of Railway Rd. x-ing, nr fenceline, -22:26:8, 117:49:56, 13-April-2005 to 18-April-2005, M. Bulbert & G. Wood, CVA volunteers, PILB041/05M (Malaise trap) // 1136 // K 246909 (1, AM); 28.22S 122.37E Deeba Rock Hole 34km NEbyE of Laverton WA 12.xi.1977 T.A. Weir (1, ANIC); Marloo Stn. Wurarga, W.A. 1931-1941 A. Goerling (1 ♁, 3 ♀, 6, ANIC); 7.5 km SE of Banjiwarn HS (27°42′S 121°37′E) W. Aust. 22-28 Feb. 1980. T.F. Houston et al. 316-10 // at light at night // Western Australian Museum Dept. of Biological Survey Site. B W Camp (1, WAM E88383); 37 km SW Youanmi. Western Australia. 28.45 S 119.31 E. 13-14 March 1982. T.F. Houston & B. Hanich. 437-8 // at light at night (1, WAM E88384); Kathleen Valley 27°24′S 120°39′E Western Australia 1963 T. Moriarty (1, WAM E88386); WA Charles Darwin Reserve, 2.5 km N White Wells HS 29°33′39″S 116°57′52″E 4-5 March 2008 T.F. Houston 1254-2b // ex gas lantern / pitfall trap 10:00pm to dawn, mixed shrubland with emergent eucalypts and native pine (1, WAM E88387); Ca. 1 km WSW of Koora Retreat, Koorarawalyee, Western Australia. 31°15′54″S 120°00′50″E 26-27 December 2011 T.F. Houston 1390-1 // Ex fluoro-light/ pitfall trap operated 7:45 to 4 am (1, WAM E88388); Mogumber, 16.xii.55, Collector R.P. McMillan (1, WAM E88385); Wooramel River, West. Australia 25°47′S 115°58′E 14 April 1979, M. Peterson // at light at night (1, WAM E88380); Warnie River, 49 km E of Paynes Find, Western Australia, 12-13 March 1982, T.F. Houston & B. Hanich, 436-1 // at light at night (2, WAM E88374, E88375); Wannara Stn [Wannarra Station] WA, 13 Feb 2010, Light, SR Patterson (1, WAM E88401). Diagnosis. Pronotum robust, disc very densely punctate except narrow longitudinal stripe from central impression to pronotal furrow, and small obscure area either side of central impression, lacking punctation; elytron blackishbrown with pale orange fasciate maculation, fascia rounded before suture, distinct humeral and apical maculae absent (common) or small apical macula present (uncommon), punctation lacking nodules, 8 th stria beginning anterior to fascia, between 3 rd and 8 th punctation of 7 th stria, all striae almost complete to apex; femora entirely dark brown, only profemora slightly swollen; tarsi with three ventral tarsal pads. Ancyropilus simplex sp. nov. is most similar to A. noonbahensis sp. nov. though differs from it by the tegmen with base of parameroid lobes weakly chitonised externally and dorsal sinus V-shaped, the femora entirely black, and the apical elytral maculation absent (common) or only faintly represented (uncommon). Description. Habitus: Fig. 146. Total length: 7.3–13.9 mm (holotype, 8 mm). Head: Cranium black, clypeus and supra-antennal elevations reddish-black, anteclypeus semi-transparent orange, labrum, palpi and antennae orange to reddish-brown; vertex and frons with dense network of irregularly-shaped punctation, above clypeus with weak transverse rugulosity; clypeus variable, rugulose to punctate, anterior margin straight to weakly angulateconcave; eyes separated by 0.65–0.96 eye widths (holotype, 0.75); genae wrinkled; terminal maxillary palpomeres with exterior margins about 1.8 times the length of inside edges; terminal labial palpomeres with exterior margins about 2.5 times the length of inside edges; antennae almost reaching base of pronotum; setae pale, long erect setae on, and bordering, eyes and along clypeus, shorter medially-directed on frons, posteriorly-directed setae above eyes. Prothorax: Black to blackish-brown; pronotum 1.13–1.24 times longer than wide (holotype, 1.19), middle about as wide as pronotal arch, sides rounded, lateral impression obscure, subapical depression v-shaped, central discal impression shallow, indistinct; disc very densely punctate except a conspicuous slender impunctate longitudinal line from central impression to pronotal collar and small obscure area before pronotal arch either side of central impression, pronotal arch and collar less densely punctate; setae pale, dense fine multi-directional setae forming swirld around impunctate areas, thicker longer erect setae most dense laterally. Pterothorax: Ventrites black or brownish, metaventrite with seta-associated punctation, setae short pale posteriorly-directed; elytra blackish-brown with a single pale-orange maculation (each elytron with a broad fasciate band rounded at the suture), apices lacking maculation (common) or with a small pale macula (uncommon); length to width ratio 2.73–2.91:1 (holotype, 2.75); punctation circular, large anterior to fascia, smaller posterior of fascia, without internal nodules, all striae terminating near apex, 8 th stria beginning anterior to fascia, between 3 rd and 8 th punctation of 7 th stria (beginning closer to base in larger-sized specimens); interstriae about as wide as corresponding punctures (may appear narrower), with rows of fine semi-erect setae (> 1/puncture) and longer erect setae every few punctures, more than one seta often across interstrial width, epipleurae extending into apical curve; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 substantialy extended basad of CuA 1 crossvein. Legs: Femora entirely dark brown, tibiae and tarsi lighter brown, longitudinal tibial carina often darker than rest of tibia, ventral tarsal pads pale brown; profemora slightly swollen, other femora slenderer. Abdomen: Ventrites orange to orange-brown. Male genitalia: Tegmenal plate (Fig. 33) broadest at base, narrowing before parameroid lobes, base of parameroid lobes weakly chitonised externally, tapered and digitiform apically, dorsal sinus about one-quarter tegmen length, wide open externally, internal limit V-shaped, opening at mid-point with short membranous (setose?) lobes, ventral sinus of similar length, apodeme about one-quarter tegmen length; median lobe as in Fig. 72; pygidium as in Fig. 108. Etymology. The specific epithet, simplex (Latin, meaning single), refers to the single large orange maculation just posterior of the middle on each elytron. This is the only species of the genus in which most indivuals lack maculation on the elytral apices (a more-or-less distinct mark was observed on a single specimen). Variation. Each elytron of one specimen collected at Juna Downs Station in the Pilbara region has a more-orless distinct, small, pale, apical macula. Biology. Specimens have been collected from October to April, some using a Malaise trap. Distribution (Map 1). Ancyropilus simplex sp. nov. is known only from central Western Australia., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 19-21, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Platynotum gracile sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 9E4BF236-0C53-4633-ADD0-9FFB86900A4A (Figs 65, 101, 137, 180; Map 2) HOLOTYPE ♁: Queensland: 6km N Taroom, Qld, 25°36′S 149°46′E, 1 Oct 1991, 200m, G. Daniels, mv lamp (QM, type reg. T258559). PARATYPE (1): QLD: 24°49′Sx 149°45′E Brigalow Res.Stn. site 2. 13Oct2001 Pyrethrum Monteith. Burwell. 170m brigalow trunks 10222 (1 ♀, QM). Diagnosis. Pronotum sub-tuberculate laterally, disc smooth, punctation mostly seta-associated; elytra dark with orange fasciate and apical maculations fasciae positioned posterior of elytral mid-length (humeral maculae absent), fasciae meeting at the suture, punctation lacking nodules, 8 th stria beginning half way between base and fascia, striae 1–9 complete to apical maculae, stria 10 ending just before apical macula; femora yellow basally, brown apically (Description. Habitus: Fig. 180. Total length: 10.4 mm. Head: Cranium black, clypeus and supra-antennal elevations infused with reddish-black, anteclypeus semi-transparent orange, antennae, labrum and palpi orange to orange-brown; eyes separated by 0.62 eye widths; vertex with small punctures and fine transverse wrinkles basally and heavier punctation towards frons, frons with scattered punctation, heaviest laterally, least dense above clypeus, surface without conspicuous sulcate or elevated areas; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.8 times (maxillae) and 2.8 times (labium) the length of inside edges; antennae almost reaching base of pronotum; entire frons with fine pale medially-directed setae. Prothorax: Black, venter infused with deep reddish hue; pronotum 1.3 times longer than wide, sides sub-tuberculate, middle about as wide as pronotal arch; subapical depression deeply v-shaped, tapered towards a short deep central sulcus, lateral sulci obscure, disc smooth, punctation mostly seta-associated; numerous short fine multi-directional setae and occassional very long erect setae. Pterothorax: Black infused with reddish hue, vested with numerous short and occasional long setae; elytra blackishbrown with orange markings (each elytron with an apical macula and a transverse fascia just posterior of the elytral mid-length); length to width ratio 3.17:1; 8 th stria beginning half way between base and fascia, striae 1–9 reaching apical macula, tenth stria ending just before apical macula, punctation lacking internal nodules, basal punctation almost twice the diameter of those posterior of fascia, epipleurae extending to apical maculae, interstriae smooth, less than width of basal punctation, about as wide as corresponding punctations posterior of fascia, interstriae with fine short semi-erect setae (often> 1 per puncture) and longer erect setae (often 1 per every 2 punctures). Legs: Coxae brownish, slightly more than basal half of profemora and about basal two-thirds of meso- and metafemora yellow, slightly less than apical half of profemora and about apical one-third of meso- and metafemora dark brown, tibiae and tarsi pale brown, ventral tarsal pads yellowish; profemora very slightly swollen, other femora slender. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 65) slender, widest at bend of tegminal arms, evenly tapering towards parameroid lobes, the latter more heavily sclerotised and apparently associated with an internal frame-like structure, dorsal sinus short, about one-fifth tegmen length, narrow, ventral sinus about as long or slightly shorter, apodeme about one-quarter tegmen length, broadly spatulate at base; median lobe (Fig. 101) with apical thread-like process; pygidium as in Fig. 137. Etymology. The specific epithet gracile (Latin, meaning slender) refers to the long slender body shape of this species. Biology. One specimen was collected at light (m. v. lamp), the other by pyrethrum knockdown on brigalow trunks; both in October. Distribution (Map 2). Platynotum gracile sp. nov. is known only from two localities, one just north of, the other just south of, Isla Gorge National Park, south-central Queensland., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 78-79, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["Gunter, N. L., Leavengood, J. M., Bartlett, J. S., Chapman, E. G. & Cameron, S. L. (2013) A molecular phylogeny of the checkered beetles and a description of Epiclininae a new subfamily (Coleoptera: Cleroidea: Cleridae). Systematic Entomology, 38 (3), 626 - 636. https: // doi. org / 10.1111 / syen. 12019","Bartlett, J. S. (2021) A preliminary suprageneric classification for Clerinae (Coleoptera: Cleridae) based on molecular and morphological evidence, including a review of tegminal terminology. Annales Zoologici, Warszawa, 71 (4), 737 - 766. https: // doi. org / 10.3161 / 00034541 ANZ 2021.71.4.003"]}

Notopilo tanybasilaris sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 2B94F695-58FC-4F7E-9AE0-609E1CFC8806 (Figs 51, 88, 124, 165; Map 4) HOLOTYPE ♁: Queensland: QLD: 24°49′ Sx 149°45′E Brigalow Res.Stn. site 2. 12Oct2001 scrub/briga. Burwell. Monteith. 170m pry.trunks, logs.10209 (QM, type reg. T258553). PARATYPES (4): Queensland: same data as holotype (1, QM); QLD: 24°49′ Sx 149°45′E Brigalow Res.Stn. site 2. 13Oct2001 Pyrethrum Monteith.Burwell. 170m brigalow trunks 10222 (1, QM); QLD: 24°32′ Sx 150°36′E Mt Scoria. 200m. 23Mar2001. vine scrub GB Monteith. Pyrethrum-trunks&logs. 10055 (1 ♁, QM); CQ: 22°16′Sx 147°16′E Mazeppa NP, S end. 240m 18 Dec 2000. 9885 GB Monteith. Brigalow Pyrethrum-trunks &logs (1, QM). Diagnosis. Pronotal sides curvate, disc extremely smooth with isolated punctures; central impression short; tumescent areas either side of central impression indistinct; elytra dark with orange maculae (each elytron with a transverse band posterior to middle plus an apical macula meeting external and sutural margins); punctures with conspicuous lateral nodules; 8 th stria beginning near base; outer striae reaching apical macula; femora yellow and black or black-brown; tarsi with three ventral tarsal pads. Notopilo tanybasilaris sp. nov. is superficially similar to Notopilo cambageicola sp. nov., N. lawnhillensis sp. nov. and N. confusus sp. nov. It differs from N. cambageicola sp. nov. by the absence of humeral maculae, from N. lawnhillensis sp. nov. by its more elongate elytra and shorter ventral sinus of the tegmen and from N. confusus sp. nov. by its more rotund pronotum and the outer striae of its elytra reaching the apical maculae. Description. Habitus: Fig. 165. Total length: 6.7–9.1 mm (holotype, 9.1 mm). Head: Vertex and frons black, clypeus and supra-antennal elevations reddish-black, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown; eyes separated by about 0.6–0.7 eye widths (holotype, 0.65); vertex smooth or punctate behind eyes, frons with punctures connected by wrinkles, clypeus with isolated punctures; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.3 times (maxillae) and 2 times (labium) the length of inside edges; antennae reaching close to base of pronotum; eyes and cranium with long and short pale erect setae, frons with shorter medially-directed setae. Prothorax: Rotund in appearance, black; pronotum about 1.21–1.26 times longer than wide (holotype, 1.21), sides curvate, middle wider than pronotal arch; subapical depression deeply v-shaped; central impression short, not sharply defined; disc mostly nitid with small, shallow, isolated punctures; preapical tumescence weak; lateral impression inconspicuous; disc with short multi-directional, and long erect, setae. Pterothorax: Ventrites black or brown, with fine mostly posteriorly-directed setae; elytra black or dark brown with orange markings (each elytron with a broad transverse fascia just behind the elytral mid-length—its anterior and posterior margins either parallel or diverging towards suture—plus an apical macula meeting the external and sutural margins), length to width ratio 2.65–2.88:1 (holotype, 2.79:1); 8 th stria beginning near base, between 1 st and 3 rd puncture of 7 th stria; punctation circular, with conspicuous lateral nodules (nodules can make some punctures appear slightly elongate rather than circular), large in basal half then reduced to about half that diameter posterior of fascia; interstriae with single rows of long and short erect or semi-erect setae, intrafoveal setae shorter than half puncture diameter in basal punctures, occasionally longer than diameter of smaller apical punctures; epipleurae extending into apical maculae; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 absent basad of CuA 1 crossvein. Legs: Approximately basal three-fifths of profemora and two-fifths of meso- and metafemora yellow, remaining apical parts of femora black to black-brown; tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; profemora slightly more swollen than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 51) only slightly narrowed between middle and robust parameroid lobes, apices tapered to short inwardly-directed digitiform processes, dorsal sinus about one-third tegmen length, inner margins abruptly expanded before inner limit, ventral sinus about one-third as long as dorsal sinus, apodeme about one-third tegmen length; median lobe as in Fig. 88; pygidium as in Fig. 124. Etymology. The specific epithet tanybasilaris (from Greek ‘tanyo’ meaning stretched out, and Neo-Latin ‘basilaris’ at the base) refers to the elongate shape of the immaculate region of the elytra between the base and anterior margin of the transverse fascia. Biology. This species has been collected in October, December and March by spraying brigalow trunks and logs with pyrethrum. Distribution (Map 4). Notopilo tanybasilaris sp. nov. is known only from three localities within the brigalow belt of central Queensland., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 49-50, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo variipes (Chevrolat) comb. nov. (Figs 12, 56, 93, 128, 171, 183; Map 3) Opilo variipes Chevrolat, 1874: 285. Type material. LECTOTYPE ♁ (here designated): New South Wales: Sidney [illegible] // variipes Chv Type Sydney // TYPE // Museum Paris Coll. Chevrolat Coll. Sedillot 1935 (MNHN) (Fig. 183). Comment on lectotype designation. As Chevrolat (1874) made no reference to specimens in his description it cannot be assumed that the nominal species group name Opilo variipes was based on a single specimen (ICZN 1999: Recommendation 73F). The MNHN specimen labelled ‘type’ must therefore be considered a syntype (ICZN 1999: Article 73.2) and available for lectotype designation (ICZN 1999: Article 74.1). Condition of lectotype: A3–11 of right antenna missing, frass and debris attached to many longer setae, right side of metaventrite torn at exit point of pin, present pin not original; otherwise in good condition. Other material examined (63 specimens): Victoria. 37.59S 147.43E, Rotomah Is., 20km SE by S Bairnsdale, VIC, 20-23 Feb. 1987, D.C.F. Rentz & J. Balderson (1 ♀, ANIC); CE Cole, Melbourne, 9.3.18 // 11 // congruus // COL-65659 (1 ♁, NMV); Barton, Victoria, J.E.Dixon (1 ♀, NMV); Trentham, Victoria, E.T. Smith (1 ♁, NMV); Launching Place, Vic., C. Oke // COL-65652 (1 ♁, NMV); Na Na Goon, 21.1.[?], E. Smith (1 ♁, NMV); Victoia, Grampians // Schenkling det. // DEI Muncheberg Col —01931 (1 ♁, SDEI); Victoria 1880 // Museum Paris (Coll. C. H. Schill) H. Donckier 1909 (1 ♁, MNHN); Victoria Australie // Ex Musaeo E. Hintz // Museum Paris ex Coll R. Oberthur (1 ♁, MNHN); Australie Victoria // 10 // Opilo variipes Chev. // Museum Paris Coll. M. Pic (1 ♁, MNHN); Victoria Australie // Museum Paris Coll. M. Pic (1 ♁, MNHN); Australie Victoria // variipes Chev. // Ex Musaeo E. Hintz // Museum Paris 1952 Coll R. Oberthur (1 ♀, MNHN); Victoria // Museum Paris Coll. Castelnau Coll. Sedillot 1935 (1 ♁, MNHN). New South Wales. R.H. Mulder Collection // Lilyvale 17-11-1973 N.S. W.— R.H.M. // K 304493 (1 ♁, AM); Culoul Range 6.1.1979 // K 304496 (1 ♁, AM); K. K. Spence Collection // French’s Forest KKS xii 33 // K 304556 (1 ♁, AM); Australia: Federal Hwy N.S.W., ii.63, B.P. Moore (1 ♀, ANIC); 37.13S 149.43E NSW East Boyd NP 54km SE Bombala, 6 Dec. 2004 - 12 Jan. 2005 C. Lambkin, N. Starick // Anteaters Rd. Malaise Trap ANIC bulk sample 2614 (1 ♀, ANIC); 105km SW of Nowra NSW, on Nerriga-Nowra Rd., 19.i.71, woodlands, S. Misko & K. Pullen (2 ♁, ANIC); 35.30S 150.18E Kioloa SF, 15km NE Batemans Bay, NSW Jan. 87 M. G. Robinson flight interc. Trap (1 ♀, ANIC); Stockard Home, Combined St., Wingham NSW 23.xii.90 S. Watkins // S.G. Watkins Collection Donated 2001 // 233 (1 ♀, ANIC); 34.24S 150.50E Mt Keira scout camp, NSW c. 320m 4-5 Mar. 1981 Lawrence & Calder (1 ♀, ANIC); Narrara N.S.W, Oct 1936, F.E. Wilson // F. E. Wilson Collection // COL-65641 (1 ♁, NMV); Narara N.S.W, 27-11-46 AB (3 ♁, NMV); Australia, NSW, 5km nw Wollombi (sw Cessnock) 4.12.1991 leg. R. Gerstmeier (10 ♁, 3 ♀, RGCM); Australia, NSW, 15km sw Cessnock, 4.12.1990 leg. R. Gerstmeier (4 ♁, 3 ♀, RGCM); Australia, NSW, 12km sw Bulga 7.12.1990 leg. M. Baehr (1 ♀, RGCM). Queensland. SEQ: 27°27′ Sx 152°55′E Enoggera Reservoir site3, 4Nov 1999 Rainforest Monteith. Burwell. pyreth. On hoop pines. 7921 (1 ♀, QM); SEQ: 25°27′ Sx 151°23′E Gurgeena Plat. Rainforest 10 Oct-19 Dec 1998 G.Monteith & C.Gough intercept. 360m 7513 (1 ♀, QM); Queensland // Coll. E.W. Janson (1 ♀, MNHN). Unknown locality. 2926 // Notoxus congruus, New. —BM // Museum Paris Coll. Gorham 1914 (1 ♁, MNHN); Darling Riv. // Museum Paris Coll. Castelnau Coll. Sedillot 1935 (1 ♁, MNHN); Museum Paris Coll. Castelnau Coll. Sedillot 1935 (1 ♁, MNHN); Australie CH. French // Determin S. Schenkling // Oplio variipes Chevr. (1 ♁, IRSNB); Bayswater (1 ♁, NMV); [illegible—W Yalok?] 14-1-06 // 2652 (1 ♁, NMV); [no data] (1 ♁, NMV). Diagnosis. Pronotum rounded laterally, disc without obvious punctation; elytra dark with orange fasciate and apical maculations, humeral maculae absent, punctation with nodules (Fig. 12), 8 th stria absent anterior to fascia (beginning within it), striae terminating within, or just after, fascia, males with setal mat covering striae 1–4 within fascia; femora yellow basally, pale brown apically (much paler than elytra), tarsi with three ventral tarsal pads. Similar to N. brevistriatus sp. nov., though easily differentiated from it by having internally nodulate punctation and the outer striae extending beyond elytral fascia. Description. Habitus: Fig. 171. Total length: 10.5–16.7 mm (lectotype 11.2 mm). Head: Vertex, frons, genae and submentum blackish, clypeus and supra-antennal elevations reddish-brown, anteclypeus transparent yellow/ orange, antennae, labrum and palpi orange-brown; eyes separated by 0.53–0.75 eye widths (lectotype 0.66); vertex and frons mostly smooth with only occasional small seta-associated punctations, surface even, not sulcate; genae and submentum wrinkled; exterior margins of terminal palpomeres about 2.3 times (maxillae) and 2.7 times (labium) the length of inside edges; antennae reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with slightly shorter medially-directed setae. Prothorax: Blackish-brown, venter, pronotal collar and arch paler more reddish or brownish; pronotum 1.26–1.35 times longer than wide (lectotype 1.32), sides round, widest at middle; subapical depression deeply v-shaped, disc with deep central sulcus (sulcus smooth, more open than linear) and obscure lateral sulci, surface smooth, almost impunctate; moderately distributed with long erect setae and shorter finer multi-directional setae. Pterothorax: Ventrites brown to reddish-brown, vested with short pale and occasional long setae; elytra dark reddish-brown with orange markings (each elytron with a large apical macula and a transverse fascia which is broadest at the suture); length to width ratio 2.79–3.15:1 (lectotype 2.91); mostly 9-striate (8 th stria begins within transverse fascia), all stria terminating well before apex (1–3 or 4 within fascia or near, 4 or 5–10 just posterior of fascia), punctation with nodules (most visible within fascia at striae 4–10), very large and complete anterior to fascia, smaller with posterior edge indistinct posterior to fascia, interstriae smooth, epipleurae extending into apical curve, interstriae with very fine short semi-reclinate setae (often>1 per puncture) and longer thicker erect setae (Legs: Profemora pale brown, basal half of meso- and metafemora yellow, apical half pale brown, tibiae and tarsi pale brown, ventral tarsal pads paler; profemora very slightly swollen, other femora slender. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 56) moderately slender, middle sinuate, parameroid lobes conspicuously expanded laterally, the latter apically subdigitiform, dorsal sinus long, about one-third tegmen length, slightly narrowed at its half-length, terminally curved, ventral sinus shallow, about three-eighths the length of dorsal sinus, tegminal arms gradually meeting apodeme, apodeme a little less than quarter tegmen length; median lobe as in Fig. 93; pygidium as in Fig. 128. Variation. Colour of some specimens closer to black than brown; brown area of meso- and metafemora sometimes less than half of femoral length (possibly females only). Biology. In southeast Queensland Notopilo variipes has been collected in a flight intercept trap in rainforest at Gurgeena Plateau and by pyrethrum spraying Hoop Pines at Enoggera Reservoir; in New South Wales it was collected using flight intercept traps at Kioloa State Forest. Collection data indicates that adults are active from October to March. Distribution (Map 3). Throughout Victoria and New South Wales into south-east Queensland. Unplaced to species group Remarks. The following four species cannot be assigned to any of the five Notopilo species groups defined above, and we have not identified synapomorphies on which to base further species groups to which they may be assigned. All four species have CuA 3+4 and CuA 1 cross-veins complete., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 59-60, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["Chevrolat, A. (1874) Catalogue des Clerides de la collection de M. A. Chevrolat. Revue et Magasin de Zoologie Pure et Appliquee, Series 3, 2, 252 - 329.","ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp."]}

Platynotum culgoense sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 04250E2A-E14F-41F0-800B-82A7F3673012 (Figs 62, 98, 134, 177; Map 2) HOLOTYPE ♁: Queensland: Qld: 28.939° Sx 147.004°E Culgoa Floodplain NP 10km NE Toulby Gate (CG3) 16Sep-1Oct2008. Simpson Coward. 140m. Malaise. Brigalow 16325 (QM, type reg. T258558). PARATYPES (3): Queensland: Qld: Currawinya NP, 4kmNW HQ (CW3). 28.813° Sx 144.462°E Mulga 15Dec2007 - 8Feb2008. Lambkin, Townsend, Starick. Malaise 16249 (1 ♀, QM); Qld: Plevna Downs. 12.7km SSE HS (PD8). Malaise 26.786° Sx 142.648°E 16Sept-2Oct2008. Starick, Lambkin, Mackenzie. 145m Eucalyptus 16285 (1 ♀, QM); Qld: 3.7km S HS Noonbah Station (NB4 M). 186m 24.141° Sx 143.19°E 19Jan-7Feb2009. Malaise A.Emmott. Dense Gidgyea scrub. 17254 (1 ♀, QM). Diagnosis. Pronotum tuberculate laterally, disc flattened, impressed in middle, not heavily punctate (most densely punctate laterally); elytra dark with yellowish fasciate and apical maculations (humeral maculae absent), the transverse fascia curved before (not meeting at) the suture, punctation without nodules, 8 th stria beginning near base, all striae reaching apical macula, interstrial setae in single rows; femora yellow and brown, tarsi with three ventral tarsal pads. Platynotum culgoense sp. nov. differs from P. femorale comb. nov. and P. gracile sp. nov. by its flatter pronotal disc, complete 8 th elytral striae, the pre-sutural rounding-off of the elytral fascia and by its smaller size. Description. Habitus: Fig. 177. Total length: 6.6–7.1 mm (holotype 7 mm). Head: Cranium blackish to dark brown, submentum and gula region paler, clypeus and supra-antennal elevations infused with reddish-black, anteclypeus semi-transparent orange, antennae, labrum and palpi orange-brown; eyes separated by 1.14–1.26 eye widths (holotype 1.2); vertex with numerous well-spaced punctations laterally, partly impunctate medially, frons punctate mostly around eye margins with few medial punctations (largely smooth medially), impunctate area in upper part of frons slightly elevated; genae wrinkled; submentum smooth; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 3 times (labium) the length of inside edges; antennae almost reaching base of pronotum; vertex and lower part of frons with fine pale erect or medially-directed setae. Prothorax: Light to dark brown, pronotum 1.2–1.27 times longer than wide (holotype 1.26), sides with weak lateral tubercle behind middle; pronotal arch extremely short laterally, almost as wide as pronotum at tubercles; disc notably compressed and broadly impressed in middle; subapical depression broadly v-shaped and adjoining linear central impression, lateral impression obscure, positioned anterior of lateral tubercle; disc lightly punctate around central impression, heavily punctate-rugulose laterally, elevated areas less punctate or impunctate; disc with numerous short fine multidirectional setae and occassional long erect setae. Pterothorax: Ventrites brown to orange-brown, vested with short pale and occasional long setae; elytra dark or light brown with orange to yellowish markings (each elytron with an apical macula and a broad transverse fascia which is rounded-off before, and not meeting at, the suture); length to width ratio 2.75–2.93:1 (holotype 2.75); 8 th stria beginning near base, only slightly shorter than 7 th stria, all stria reaching apical macula, punctation lacking internal nodules, relatively uniform in size along full elytral length, punctation of outer striae often smaller than those of inner striae, epipleurae extending to apical maculae; interstriae with very fine short erect setae (often 1 per puncture) and longer thicker erect setae (about 1 per every 3 or 4 punctures), intrafoveal setae about half a puncture diameter in length. Legs: Coxae brownish, approximately basal half of profemora and about basal three-fifth to two-thirds of meso- and metafemora yellow, femora broan apically, demarcation of yellow and brown parts of profemora unclear on some specimens, tibiae and tarsi pale brown, ventral tarsal pads yellowish; profemora notably swollen, other femora moderately swollen; tibial spurs solid, particularly protibial spur. Abdomen: Ventrites orange or brownish. Male genitalia: Tegmen (Fig. 62) relatively slender, needlelike, widest at bend of tegminal arms, evenly tapering to parameroid lobes, the latter narrow acuminate and slightly bent laterally, dorsal sinus short, about one-sixth tegmen length, narrow internally, opening externally, ventral sinus similar in length, tegminal arms gradually meeting short (about one-eighth tegmen length) apodeme; median lobe as in (Fig. 98) with apical process; pygidium as in Fig. 134. Etymology. This species is named after the collecting locality of the male holotype, Culgoa Floodplains National Park, which is situated at the Queensland and New South Wales border at longitude 147° east. Biology. Specimens were collected from September to February by setting up Malaise traps in Eucalyptus and Acacia plant communities. Distribution (Map 2). Southern and central Western Queensland. Remarks. The species group name Notopilo culgoaensis, published in the 2009-2010 Bush Blitz survey report for north-western New South Wales and southern Queensland (ABRS 2014a, b) with no accompanying description and without reference to a name-bearing type, is deemed unavailable (ICZN 1999, Article 16.4). MAPS 4&5. Species distribution. (Map 4) Notopilo (cambageicola species group and unplaced species); (Map 5) Notopilo (congruus and reduncus species groups) (topographic layer image credit: NASA 2002)., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 66-73, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["ABRS (2014 a) North-western NSW and southern Qld 2009 - 2010. A Bush Blitz survey report. Australian Biological Resources Study, Canberra. Available from: https: // bushblitz. org. au / north-western-nsw-and-southern-qld- 2009 - 10 / (accessed 6 March 2022)","ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp.","NASA (2002) NASA Visible Earth. Blue Marble: Land Surface, Shallow Water, and Shaded Topography. EOS Project Science Office, NASA Goddard Space Flight Centre, Greenbelt. Available from: https: // visibleearth. nasa. gov / images / 57752 / bluemarble-land-surface-shallow-water-and-shaded-topography (accessed 25 April 2022)"]}

Monilonotum gen. nov. ZooBank registration: urn:lsid:zoobank.org:act: 36F952E9-96BE-41C9-BC6F-294E1E13FA00 Gender. Neuter. Type species. Opilo pascoii Gorham, 1876, by present designation. Diagnosis. Intrafoveal setae, if present, minute and not clearly visible under magnification (or apparently absent) (Fig. 22); basitarsi without a distinct ventral pad; ninth elytral stria present; inside lateral rims of elytral punctures without nodules; elytral interstriae setae arranged in simple rows; elytral striae terminating abruptly near mid-length; pronotal basal collar typically paler than pronotal disc; carinae of inside face of antennal scape often not well-defined; median lobe without rearward-directed spine or barb, distally digitiform. Description. Eyes facets intermediately coarse, strongly emarginate above supra-antennal elevations, separated by 0.88–1.62 eye widths; carinae of rear face of antennal scape short or weak (less conspicuous than most other genera treated here); sensory face of terminal maxillary palpomere 1.6 longer than inside margin; sensory face of terminal labial palpomere 1.0–2.5 times longer than inside margin; genae wrinkled, submentum smooth; pronotum 1.5–3.0 times longer than wide, middle slightly broader than pronotal arch, smooth, almost impunctate, median sulcus subtle or apparently absent, basal collar typically paler than disc; elytra 2.52–2.95 times longer than wide at humeri, punctures without nodules, eighth stria beginning immediately behind humeral tumescence, all stria terminating abruptly near elytral mid-length, interstrial setae in single rows, intrafoveal setae apparently absent in majority of punctures (an extremely short seta, difficult to observe even under high magnification, was occasionally found in some specimens); hindwing with CuA 3+4 and CuA 1 crossveins complete, MP 3+4 substantialy extended basad of CuA 1 crossvein; tarsi each with three well-developed ventral pads; tegmen ventral sinus half to two-thirds as long as dorsal sinus; median lobe without apico-lateral spine, with distal digitiform process; specular fork Y-shaped. Etymology. The generic epithet Monilonotum (from Latin ‘monile’ meaning necklace, and Latin ‘nota’ meaning mark and also suggestive of the pronotum), refers to the paleness of the pronotal collar which, in the majority of species of this genus, is characteristically contrasted against a darker pronotum. Remarks. Evidence for early divergence of Monilonotum from the other genera treated here can be found in the digitiform terminal process of the median lobe of Monilonotum which resembles that found in genera of Australian Opilonina such as Trogodendron Spinola and Olesterus Spinola, the extreme reduction of intrafoveal setae (putatively synapomorphic), the loss of the carina of the antennal scape (also putatively synapomorphic), plus the retention of plesiomorphic fused/non-separated state of the spicular fork (collectively more derived in the remaining genera of this revision). Key to species of Monilonotum 1 Pale transverse elytral fascia, if present, flat, not calositous or bullose (Figs 151–154).......... (pascoei species group) ... 2 - Pale transverse elytral fascia always present as a raised, calositous or bullose formation (Fig. 155).................................................................................................. (rufiventre species group)... 5 2 (1) Elytral colouration yellowish to pale brown and dark brown (Fig. 151).................. Monilonotum bunyense sp. nov. - Elytral colouration black, reddish-brown and white.......................................................... 3 3 (2) Each elytron with a small white fleck between transverse fascia and base; elytral apices without maculation (Fig. 152)............................................................. Monilonotum doddi (Schenkling) stat. rev., comb. nov. - Elytra between fascia and base without white fleck; elytral apices with whitish macula.............................. 4 4 (3) Metacoxae and metaventrite of similar colour; interstices anterior to elytral fascia thin and slightly raised, giving punctation a grid-like appearance within this area............................................ Monilonotum sundholmi sp. nov. - Metacoxae whitish or yellowish, metaventrite orange-brown; interstices anterior to elytral fascia smooth, punctation not gridlike in appearance within this area.................................... Monilonotum pascoei (Gorham) comb. nov. 5 (4) Elytral base red, sternum not red............................... * Monilonotum eburneocincta (Gorham) comb. nov. - Elytra entirely black; sternum orange............................. * Monilonotum rufiventre (Gerstmeier) comb. nov. * See notes below, under each species heading, for information about potentially superficially similar undescribed species.

Notopilo gerstmeieri sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 0B0DFD7E-52B7-463F-9E2D-1A2D1CC8D10E (Figs 4, 55, 92, 127, 170; Map 3) HOLOTYPE ♁: New South Wales: NSW 30km S of Glen Innes 28 Dec. 1982 J.Doyen coll. (ANIC). PARATYPES (15): New South Wales: Australia, NSW, 27km s Glen Innes 28.11.1990 leg. R. Gerstmeier (4, RGCM; 1 ♁, 1, JSBC); Australia, NSW, 15km n Armidale 28.11.1990 leg. R. Gerstmeier (1 ♀, 1, RGCM); Australia, NSW, Mt Kosciusko NP, Sawpit Ck. 1400m 11.- 12.12.1990 leg. M. Baehr (2, RGCM); Australia, NSW, 30km w Cooma 11.12.1990 Leg. M. Baehr (1 ♀, RGCM); Jindabyne 1/06 H.C. // Australie // Museum Paris E. Le Moult 1916 // Opilo congruus Newm. (2 specimens on one card, MNHN); Australie Sidney 12-II-1902 // Museum Paris coll R. Oberthur (1, MNHN); Tarago Rd Bungendore NSW 5.ii.1978 E.B. Britton (1, ANIC). Diagnosis. Pronotum broadly rounded laterally, disc extremely smooth, almost impunctate dorsally, punctation conspicuous only laterally, central impression shallow; elytra dark with orange fasciate and apical maculations (humeral maculations absent), fascia meeting at the suture (anterior and posterior margins parallel or angled towards lateral margins), punctation with conspicuous lateral nodules, 8 th stria beginning between base and half way towards fascia, at least striae 4–9 reaching apical maculae; profemora entirely, or almost entirely, brown, other femora yellow basally, brown apically, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 170. Total length: 8.1–12.6 mm (holotype 8.1 mm). Head: Cranium black to dark brown, genae and submentum dark brown, gula orange, clypeus and supra-antennal elevation more reddish, anteclypeus orange, labrum and palpi orange-brown, antennae darker brown; eyes separated by 0.76–0.97 eye widths (holotype 0.73); vertex and frons moderately punctate, frons less punctate above clypeus, surface at narrowest point uneven, clypeus with few punctations; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.7:1 (maxillae) and 2:1 (labium); antennae reaching near base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter finer medially-directed setae. Prothorax: Black to blackishbrown, pronotal arch may be infused with dark red anteriorly; pronotum 1.13–1.25 times longer than wide (holotype 1.23), sides rounded, middle wider than pronotal arch; subapical depression v-shaped, indistinct near middle, meeting shallow central impression; disc extremely smooth, conspicuously punctate laterally only; sparse short erect setae, even fewer long erect setae. Pterothorax: Ventrites blackish-brown, with numerous pale, posteriorly- to posteromedially-directed setae; elytra blackish to dark brown with orange markings (each elytron with a transverse fascia behind the middle which meets at the suture and narrows slightly towards the lateral margin, and large apical macula); length to width ratio 2.74–2.92:1 (holotype 2.78); 8 th stria beginning between base and half way towards fascia (between 2 nd and 6 th punctations of 7 th stria), at least striae 4–9 reaching apical macula (striae 1 and 10 not reaching apical macula, striae 2 and 3 variable), punctation with conspicuous lateral nodules, punctations posterior of fascia about half the size of those anterior to fascia, epipleurae extending into apical maculae, interstriae with very fine short semi-erect setae and less frequent longer thicker erect setae, not with several setae across interstical width; hindwing with CuA 3+4 cross-vein complete, CuA 1 cross-vein absent. Legs: Profemora entirely, or almost entirely, brown, basal one-third to two-fifths of meso- and metafemora yellow, apical two-thirds to three-fifths brown; tibiae and tarsi brown, ventral tarsal pads yellowish; meso- and metafemora slender, profemora slightly swollen. Abdomen: Ventrites orange to orange-brown. Male genitalia: Tegmen (Figs 4, 55) broadest near middle, narrowing before expanding again before the apically acuminate parameroid lobes, dorsal sinus about one-third tegmen length, sides internally parallel, diverging before opening, ventral sinus about half as deep, apodeme about one-quarter tegmen length; median lobe as in Fig. 92; pygidium as in Fig. 127. Variation. In dorsal view, the elytral fascia may appear straight (narrowing just before the lateral margins) or angled (narrowing towards the lateral margins from near the suture). Etymology. We dedicate this species to the Prof. Dr Roland Gerstmeier of Munich, Germany, prolific clerid beetle specialist plus friend and colleague of the first author. Biology. Specimens were collected during the period November to February. Distribution (Map 3). Notopilo gerstmeieri sp. nov. has been collected from regions along, or east of, the Great Dividing Range in New South Wales, from Mt Kosciuszko to Glen Innes., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 58-59, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Monilonotum eburneocincta (Gorham) comb. nov. Opilo eburneocinctus Gorham, 1878: 160. Status of type material. Syntypes not located. Remarks. Despite neither type material, nor specimens determined to be O. eburneocintus, being found by the first author in Australia or Europe, Gorham (1878) provides clues in his descrition of ‘ eburneocintus ’ as a smaller version of Opilo pascoei with a white ‘raised’ elytral fascia, a red elytral base and a sternum that is not red (as in O. pascoei). The first author is aware of numerous, miniature ‘ pascoei -like’ species with callositous, bullose (i.e., raised) elytral fasciae, though the ventrites of these species range from bright orange to reddish-orange. Despite being confident enough to transfer Opilo eburneocinctus to Monilonotum, we have decided not to describe any of the smaller species as new while the identity of Monilonotum eburneocinta comb. nov. remains unknown. Distribution (Map 2). New South Wales., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 36, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["Gorham, H. S. (1878) Descriptions of new genera and species of Cleridae, with notes on the genera and corrections of synonymy. Transactions of the Entomological Society of London, 1878 (2), 153 - 167. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 00871. x"]}

Ancyropilus packsaddlensis sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: E27871CE-9CEB-4109-B892-0C171352336E (Figs 32, 71, 107, 145; Map 1) HOLOTYPE ♁: New South Wales: Packsaddle, 111 mi. N Broken Hill, 21-24 Nov. 1969 // Malaise trap H.E. Evans, R.W. Matthews (ANIC). PARATYPES (4): Queensland: Qld: 26.67° Sx 142.577°E 2.5km WNW Plevna Downs HS (PD3 M) 133m. 25 Nov 2008 - 17 Dec 2009. R. Mackenzie. Pitfall. Gidgee. 17317 (1 ♀, QM); Qld: 26.67°Sx 142.577°E 2.5km WNW Plevna Downs HS (PD3 M) 131m. Gidgee. 15 Mar-3 Apr 2009. Malaise. R. Mackenzie. 17317 (1 ♀, 2, QM). Diagnosis. Pronotum angulate to almost rounded laterally, disc heavily punctate, glabrous areas in middle at base and either side of central impression; elytra dark with orange humeral, fasciate and apical maculations, fascia (with anterior and posterior margins approximately parallel) meeting at the suture, punctation lacking nodules, 8 th stria beginning within fascia, at least striae 3–9 reaching apical maculae; femora yellow basally, brown apically, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 145. Total length: 6–7.1 mm (holotype 6.2 mm). Head: Cranium black to dark brown, submentum and gula bright orange, clypeus and supra-antennal elevation more reddish, anteclypeus orange, labrum, palpi and antennae orange-brown; eyes separated by 0.73–0.83 eye widths (holotype 0.73); vertex punctate, upper half of frons densely punctate, lower part punctate-rugulose until just before clypeus, clypeus with few punctations; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.6:1 (maxillae) and 2.6:1 (labium); antennae not reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter finer medially-directed setae. Prothorax: Pronotal disc and arch variously dark and light brown (pronotal arch and a medial discal stripe often lighter than remainder of disc), basal collar and venter orange to yellowish; pronotum 1.2–1.24 times longer than wide (holotype 1.22), sides angulate to almost rounded, middle wider than pronotal arch; subapical depression v-shaped, meeting shallow central impression; disc densely punctate, a thin medial glabrous stripe from base to central impression (difficult to view in paler specimens), area either side of central impression weakly tumescent, partially glabrous; short fine multi-directional setae (forming subtle swirls adjacent to glabrous areas) and long erect setae. Pterothorax: Ventrites orange to yellowish, with numerous pale, posteriorly- to posteromedially-directed setae; elytra dark brown with orange markings (each elytron with a squarish humeral macula, a very broad transverse fascia which meets at the suture and large apical macula); length to width ratio 2.59–2.72:1 (holotype 2.63); 8 th stria beginning within fascia (between its anterior margin and its middle), all striae reaching apical macula or striae 1, 2 and 10 not reaching apical macula, punctation lacking nodules, punctations posterior of fascia almost half the size of those anterior to fascia, epipleurae extending into apical maculae, interstriae with very fine short semi-erect setae and less frequent longer thicker erect setae, several setae often across interstical width; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 crossvein small (but not completely absent). Legs: Basal two-thirds of profemora and basal three-quarters of meso- and metafemora yellow, all femora apically dark brown; tibiae and tarsi brown, ventral tarsal pads yellowish; meso- and metafemora slender, profemora slightly swollen. Abdomen: Ventrites orange to yellowish. Male genitalia: Tegmen (Fig. 32) broadest near middle, narrowing before expanding again before apically acuminate parameroid lobes, dorsal sinus about one-quarter tegmen length, wide open, sides slightly convergent, internal lateral opening with short membranous lobes bearing a brush-like row of very short setae, ventral sinus about as deep, apodeme about one-quarter tegmen length; median lobe as in Fig. 71; pygidium as in Fig. 107. Variation. The paler brown area of the pronotum may be distinct or obscure. Etymology. This species is named after the type locality of the male holotype, Packsaddle in north-western New South Wales. Biology. Specimens were collected using pitfall, and Malaise, traps during the periods November-December and March-April. Traps at Plevna Downs were placed in Gidgee (Acacia) plant communities. Distribution (Map 1). Ancyropilus packsaddlensis sp. nov. is known only from Packsaddle in north-western New South Wales and Plevna Downs in south-western Queensland. Remarks. Similarities in body form, and general shape of the tegmen, suggest Ancyropilus packsaddlensis sp. nov. and A. emmotti sp. nov. are closely related. The decision to describe them as distinct species was based on correlations between consistant differences in the depth of the dorsal sinus of the tegmen and elytral patternation., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 18-19, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Infectostriatus gen. nov. ZooBank registration: urn:lsid:zoobank.org:act: C11736E5-943C-41E5-8D14-DF24D7BF7B65 Gender. Masculine. Type species. Infectostriatus absentis sp. nov., by present designation. Diagnosis. Intrafoveal setae short but visible under magnification; basitarsi without a distinct ventral pad; ninth elytral stria absent (Fig. 15); inside lateral rims of elytral punctures with nodules; elytral interstriae with several setae distributed across interval width; elytral striae terminating near apices; apical lobes of median lobe with rearward-directed spine or barb. Description. Eyes coarsely-facetted, strongly emarginate above supra-antennal elevations, separated by 0.5– 0.8 eye widths; antennal scape with carina bordering each side of flattened rear face; sensory face of terminal maxillary palpomere 1.6 times longer than inside margin; sensory face of terminal labial palpomere 2.3–3.0 times longer than inside margin; genae wrinkled; pronotum 1.15–1.3 times longer than wide, broadest near middle, disc lightly to heavily punctate, median sulcus present; elytra 2.44–2.8 times longer than wide at humeri, punctures with nodules, eighth stria with post-basal/pre-apical interruption or beginning between tenth and fifteenth puncture of seventh stria, ninth stria absent, most marginal and discal striae terminating near apices, interstrial setae not in single rows (often several setae across interval width), intrafoveal setae present; hindwing with CuA 3+4 cross-vein complete, CuA 1 cross-vein complete, RP not incurvate; tarsomeres 2–4 each with a well-developed ventral pad; tegmen ventral sinus as long as, or longer than dorsal sinus; median lobe with apico-lateral spine. Etymology. The generic epithet Infectostriatus (from Latin ‘infectus’ meaning unfinished, and ‘stria’ meaning line or furrow) refers to the synapomorphic absence or reduction of the 8 th and 9 th elytral striae (i.e., intervals). Remarks. Infectostriatus gen. nov. contains two species from far north Queensland. Key to species of Infectostriatus 1 Base of elytra between 7 th and 10 th striae devoid of punctures (Fig. 15).................. Infectostriatus absentis sp. nov. - Base of elytra between 7 th and 10 th striae with short single row of punctures............. Infectostriatus differens sp. nov., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 28, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Infectostriatus differens sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 0872ADA2-B565-4E15-A5FA-954E06F63362 (Figs 6, 38, 77, 150; Map 2) HOLOTYPE ♁: Queensland: 13.53S 143.11E GPS, 6km NNE Coen, QLD, 13 Jan. 1994, P.Zborowski & E.D.Edwards, at light (ANIC). Diagnosis. Infectostriatus differens sp. nov. is most easily differentiated from I. absentis sp. nov. by the presence of punctation between the seventh and tenth striae between the base and elytral fascia. Description. Habitus: Fig. 150. Total length: 9.2 mm. Head: Vertex, frons, genae and gula black, clypeus and supra-antennal elevation black to dark reddish-brown, anteclypeus, labrum, antennae and palpi orangebrown or chestnut-brown; frons narrow (eyes separated by 0.52 eye widths), impressed above narrowest point, frontal sculpturing confused-foveolate, transversely wrinkled below narrowest point; genae wrinkled, wrinkles extending into submentum though not meeting in middle; exterior margins of terminal palpomeres about 1.6 times (maxillae) and 2.3 times (labium) the length of inside edges; antennae almost reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Pronotum black infused with a deep reddish hue; pronotum 1.15 times longer than wide, sides sub-tuberculate in middle, above each tubercle an oblique sulcus, subapical depression shallow v-shaped, disc with a central longitudinal sulcus; surface moderately well punctated, dorso-lateral punctation sometimes sub-rugulose, subtly tumescent posterior of subapical depression and surrounding central sulcus, well-vested in short fine multidirectional setae and long erect setae. Pterothorax: Ventrites black, barely punctate, moderately vested with short pale and occasional long setae; elytra blackish to brown with yellowish or orange markings (each elytron with a large apical macula, a broad ‘stepped’ central fascia which extends anteriorly between the suture and 5 th stria and ends just before base, plus a small humeral macula); length to width ratio 2.44:1; punctation with lateral bead-like nodules, 8 th and 9 th striae highly reduced, the 8 th evident within dark areas between fascia and apical macula and between fascia and base, the 9 th stria represented by fewer punctations between fascia and apical macula only, punctation relatively large until near apex, disc well-vested in very fine short and thicker erect setae, interstriae with alternating long erect setae and shorter ‘leaning’ setae, epipleurae extending well into apical curve; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 absent basad of CuA 1 crossvein. Legs: Profemora entirely black, meso- and metafemora with basal half orange and apical half black, tibiae and tarsi brown, ventral tarsal pads yellowish; profemora slightly swollen, other femora slender. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 38) relatively slender, middle weakly sinuate, broadest at ‘bend’ of tegminal arms, parameroid lobes angled inwards and blunt (not pointed) apically, dorsal sinus very wide, angular, just over one-quarter tegmen length, corner of broadest point with transparent (sensory?) membrane, ventral sinus almost as long, tegminal arms sharply meeting apodeme, apodeme just over quarter tegmen length; median lobe as in Fig. 77. Etymology. The specific epithet differens (Latin, meaning dissimilar) simply refers to the clear differences between this and the superficially very similar type species. Biology. The holotype was collected in January, at light. Distribution (Map 2). The holotype was collected near Coen in far north Queensland., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 30, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Platynotum foveosetosa sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: E1C4C71E-EEB0-458C-B70A-51E4F8D7721E (Figs 24, 64, 100, 136, 179; Map 2) HOLOTYPE (♁): Western Australia: WA: Weelhamby Lake, west MO11-2 lake floor. wet pitfalls, 29°11′01″S 116°27′32.2″E (GPS) 6 Oct 2002 – 9 Jan 2003 CALM Wheatbelt Biodiversity Survey (WAM E88396). PARATYPE (1): Western Australia: WA: Weelhamby Lake, west MO11-2 lake floor. wet pitfalls, 29°11′01.7″S 116°27′32.2″E (GPS) 9 Jan 2003 – 18 March 2003 CALM Wheatbelt Biodiversity Survey (1 ♀, WAM E88392). Diagnosis. Pronotum round laterally, disc punctate; elytra light brown with poorly defined orange to yellowish markings, punctation lacking nodules, 8 th stria beginning near base, punctures of similar diameter along elytral length, striate to apex (some striae terminating at apical macula), single seta of anterior puncture rim up to twice puncture diameter in length; legs yellow (except tibiae sometimes brownish and tarsi brown); tarsi with three ventral tarsal pads. Description. Habitus: Fig. 179. Total length: 7.7–10.2 mm (holotype, 7.7 mm). Head: Cranium dark brown (almost black), clypeus and supra-antennal elevations slightly paler, anteclypeus semi-transparent orange, antennae, labrum and palpi orange to orange-brown; eyes separated by 2–2.13 eye widths (holotype, 2.13); vertex and frons with dense network of irregular-shaped punctation, only anterior margin of clypeus impunctate, epistomal suture reasonably indistinct; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 2.5 times (labium) the length of inside edges; antennae almost reaching base of pronotum; head including eyes well-vested with long pale erect setae, frons also with numerous medially-directed setae. Prothorax: Dark brown to reddish-brown, venter light brown; pronotum 1.1–1.2 times longer than wide (holotype, 1.1), sides rounded, pronotal arch very slightly broader than middle; subapical depression v-shaped though not well-marked, a short sulcate central impression at apex of v-shape; lateral sulci apparently absent, disc reasonably heavily punctate, surface glabrous between individual punctures, base bi-tumescent (tumescence punctate), area either side of central impression flat (not tumescent); disc reasonably dense with long erect setae plus shorter multi-directional or anteriorly-directed setae. Pterothorax: Ventrites light brown, vested with numerous short and occasional long setae; elytra light brown with paler (orange to yellowish) indistinct markings (each elytron with a poorly-defined transverse fascia near middle and a poorly-defined apical macula which may or may not be joined together, the colouration/ patternation of the humeral area is even more vaguely-defined); length to width ratio 2.5–2.6:1 (holotype, 2.5:1); 8 th stria beginning near base (after first or second puncture of 7 th stria), some striae terminating or merging into others near start of apical macula, punctures lacking internal nodules, punctures of similar diameter along most of elytral length, epipleurae extending almost to apices, extreme apices weakly turned outward; interstriae smooth, at base about as wide as puncture diameter, interstriae with long erect setae and shorter setae with a slight posterior lean, (generally with several shorter setae for every long seta), intrafoveal setae very long, about twice the diameter of a puncture (Fig. 24). Legs: Coxae, femora and yellow, tibiae yellow or infused with brown, longitudinal tibial carina brown, tarsi brown, ventral tarsal pads yellowish; all femora of comparable thickness. Abdomen: Ventrites yellowish-brown. Male genitalia: Tegmen (Fig. 64) needle-like, broad at bend of tegminal arms, evenly tapering to parameroid lobes, the latter narrow acuminate and conspicuously bent laterally, dorsal sinus short, about threequarters tegmen length, ventral sinus shorter than dorsal sinus, tegminal arms gradually taperd toward apodeme and joined near spatulate base; median lobe (Fig. 100) simple with an apical process; pygidium as in Fig. 136. Etymology. The specific epithet foveosetosa (from Latin ‘fovea’ meaning hole and, Latin ‘seta’ meaning bristle), refers to the single rearwards projecting setae of the anterior rim of elytral punctures, which are notably longer than those of any other species treated in this revision. Biology. Collection in pitfall traps implies ground-dwelling behaviour. Distribution (Map 2). Platynotum foveosetosa sp. nov. is known only from Weelhamby Lake, near Koolanooka, Western Australia.

Notopilo elstoni sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: 486B7112-8072-4C35-9B50-123DADEFF11A (Figs 58, 94, 130, 173; Map 4) HOLOTYPE ♁: South Australia: New Kalamurina St. Warburton R. S. Aust 10 mar. 1972. E. Matthews (SAMA). PARATYPES (4): South Australia: L. Callabonna A. Zietz // A. H. Elston Collection // K 304526 // 741 Opilo congruus Newm. var (1, AM); 27.33S 135.27E Oodnadatta S.Aust. 29 Nov 89 I.Bunic (1, ANIC); Australia, SA. Near Oodla Wirra 427m, 32°51′S 139°06′E, 15.xi.2000 // leg. A. Podlussany, I. Rozner, George Hangay. G &K Hangay Collection (1, ANIC); Australia, SA, Wobna Mound Springs, 3.- 4.1.2004, M. Baehr (1, RGCM). Other material examined. 27.21S 136.36E, attracted to light, Mierantana W.H., S.Aust. 23 Oct 89, I. Bunic (1 ♀, ANIC). Diagnosis. Pronotum rounded, disc moderately punctated though retaining a smooth reflective appearance, central impression extremely short and shallow, tumescent areas either side of central impression indistinct; elytra dark with orange maculae (each elytron with a humeral macula extending obliquely towards the suture but ending at the second stria, a large central fasciate macula which is curved at the suture and which is thinly joined to the humeral macula along the lateral margin, plus an apical macula which meets the external margins entirely and the sutural margin at the apex only), punctation with small nodules, 8 th stria beginning near base (between 2 nd and 4 th punctation of 7 th stria), some or all of striae 3–6 reaching apical macula, mostly a single seta across interstrial width (rarely two); femora yellow and brown, tarsi with three ventral tarsal pads. Description. Habitus: Fig. 173. Total length: 6.9–9.5 mm (holotype, 9.1 mm). Head: Vertex black, frons entirely black or reddish-brown from middle of eyes to clypeus, clypeus and supra-antennal elevations reddish-brown, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown; eyes separated by about 0.73–0.85 eye widths (holotype, 0.85); vertex with numerous shallow punctations (heavier behind eyes), upper part of frons more punctate- rugulose, lower part of frons (above epistomal suture) with light transverse to weaklycurved rugulosity and occasional punctation; clypeus with numerous punctations; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 2 times (labium) the length of inside edges (based on paratypes as the palpi of the holotype are damaged); antennae not reaching base of pronotum; eyes and cranium with long erect pale setae, frons and vertex with denser shorter medially-directed setae. Prothorax: Black to reddish-black, prosternum and basal collar orange-brown; pronotum 1.14–1.22 times longer than wide (holotype, 1.15), sides rounded, middle wider than pronotal arch; subapical depression deeply v-shaped, indistinct near middle; central impression indistinct, extremely shallow, short; disc convex, well-distributed with circular punctation (though retaining a smooth, reflective appearance), punctation heaviest laterally, subrugulose, punctation least dense medially, a glabrous stripe basally, a weak ovoid partly glabrous tumescence either side of central impression; lateral impression conspicuous, obscurly-shaped; with fine short multi-directional setae and sparsely with longer erect setae. Pterothorax: Ventrites orange, with fine pale posteriorly- or medially-directed setae; elytra dark brown with orange markings (each elytron with a humeral maculation extending obliquely towards the suture but ending at the second stria, a large central fasciate maculation which is curved at the suture and which is thinly joined to the humeral maculation along the lateral margin, plus an apical maculation which meets the external margins entirely and the sutural margin at the apex only), length to width ratio 2.5–2.6:1 (holotype, 2.5:1); 8 th stria beginning anterior of fascia, between 2 nd and 4 th punctation of 7 th stria, some or all of striae 3–6 reaching apical macula (at least 4 and 5); punctation circular, with small nodules (most easily viewed in dark area posterior of humeral maculae); epipleurae extending into apical maculae; interstriae with short fine setae (at least one per puncture), plus longer thicker erect setae (at least one every two punctures), striae mostly with a single seta across interstrial width (rarely two), intrafoveal setae short; hindwing with CuA 3+4 cross-vein complete (CuA 1 cross-vein not observed), MP 3+4 absent basad of CuA 1 cross-vein. Legs: Basal two-thirds of profemora and basal three-quarters of meso- and metafemora yellow, remaining parts of femora dark brown; tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; profemora slightly more swollen than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 58) broad, middle very weakly sinuate, apices of parameroid lobes digitiform, dorsal sinus wide, about onethird tegmen length, ventral sinus about half as long as dorsal sinus, apodeme just under one-quarter tegmen length; median lobe as in Fig. 94; pygidium as in Fig. 130. Variation. The apical elytral maculae of the paratype from Wobna Mound Springs do not extend to the extreme apex. A female specimen from Mierantana Water Hole, South Australia, conforms to the above description in most respects but differs only in being larger (11.1 mm long) and in having differently patterned elytra (the humeral macula spans the entire base and is connected to the central, fasciate, macula along the suture). In the absence of a male (for comparison of genitalia) we can only consider this to represent within-species variation. We are not designating it as a paratype of N. elstoni sp. nov. in case future discovery of a male with similar elytral markings proves otherwise. Etymology. We name this species after Albert Harold Elston (1890–1957), clerk and amateur entomologist from Adelaide, South Australia, within whose collection the Lake Callabonna paratype was originally contained. In ten papers published from 1919 to 1930, Elston proposed exactly 100 species group names for Australian beetles, mainly in the families Elateridae (51 names) and Cleridae (42 names). In terms of number of Australian clerids described, Elston was second only to Frenchman, Maurice Pic, who described 46 species, though in contrast to Pic, whose descriptions were often four or five lines in length, Elston’s descriptions were substantial and informative. Biology. Adult specimens have been collected in November, January and March. Distribution (Map 4). Notopilo elstoni sp. nov. is known from the northern, arid region of South Australia., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 61-63, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo obesus sp. nov. ZooBank registration: urn:lsid:zoobank.org:act: FF303390-F098-4E0C-9296-60CED2C88888 (Figs 5, 16, 60, 96, 132, 175; Map 4) HOLOTYPE: Queensland: Coopers Plains, 768 Boundary Road, grounds of CSR Ltd.; 30.ix.-3.x.2014; S. Collingwood; panel trap (alpha-pinene & ethanol lure) in Pinus sp. (QM, type reg. T258555). PARATYPES (3): Queensland: Camp Hill, carpark of White’s Hill Shopping Village, Samuel Street, 1.xii.2003, J.S. Bartlett, under lights at night (1 ♀, JSBC); -23.83738, 151.26048, Port of Gladstone, Barney Point, Gate LPG1075; 24.i.-7.ii.2017; J. Logan, Panel Trap, α-pinene+EtOH lure (1 ♁, QDPC). New South Wales: 11 March 2019; Blaxland Ridge, Blaxland Ridge Rd. at light; 3328′13.0″S, 15048′01.4″E; Vr.R. Bejšák-Colloredo-Mansfeld lgt. (1, VRBC). Diagnosis. Pronotum round laterally, disc punctate-rugulose,either side of central impression strongly tumescent; elytra notably broader than pronotum, each elytron dark with a large slightly angulate transverse fascia which meets at the suture and an obscure basal macula; punctation with small nodules; 8 th stria beginning near base; striae 3-7 terminating at apical slope (other striae shorter); femora entirely brown; tarsi with three ventral tarsal pads. Description. Habitus: Fig. 175. Total length: 7.9 mm. Head: Cranium black, clypeus and supra-antennal elevations with reddish hue, anteclypeus, labrum and palpi orange-brown, antennae brown; eyes separated by about 0.92 eye widths; upper part of frons punctate-rugulose, lower part with weak transverse wrinkles; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillae) and 2.5 times (labium) the length of inside edges; antennae almost reaching base of pronotum; eyes and most of cranium vested with long erect setae, frons and vertex with shorter medially-directed setae. Prothorax: Black, reddish-black in parts; pronotum about 1.11 times longer than wide, sides rounded, widest in middle; subapical depression deeply v-shaped; central impression deeply excavated, weakly sulcate; either side of central impression strongly tumescent; disc unevenly punctate-rugulose, pronotal arch smoother, spaces between individual punctures sub-nitid; numerous fine short multi-directional setae plus fewer long erect setae. Pterothorax: Sternites reddish-brown, with fine pale posteriorlydirected setae; elytra (Fig. 16) about 1.4 times wider than pronotum, blackish brown with orange markings (each elytron with a large, slightly angulate, transverse fascia which meets at the suture plus an obscure brown triangular maculation its outer-most margin extending from the humerus to inner corner of fascia; apices lacking maculae); length to width ratio 2:1; 8 th stria beginning near base; striae 3–7 reaching apical slope, other striae shorter; punctures with small lateral nodules (most clearly visible in basal punctures), relatively well-spaced; basal intervals at least one puncture width, wider towards apex (punctures gradually smaller towards apex); epipleurae terminating within apical curve; intervals with many short fine, and less frequent longer, setae (much of the discal setae rubbed off holotype), intrafoveal setae short; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 absent basad of CuA 1 cross-vein. Legs: Femora, tibiae and tarsi brown, ventral tarsal pads brownish yellow; profemora slightly more swollen than other femora; front and middle legs stout, hind legs more elongate. Abdomen: Orange-brown. Male genitalia: Tegmen (Fig. 60) subparallel, heavily sclerotised structural H-shaped ‘frame’ more apparent than in congeners, parameroid lobes with relatively thick terminal sub-digitiform processes, dorsal sinus deep and wide, about one-third tegmen length, internally with preapical membrane (potentially sensory in function), ventral sinus almost two-thirds as long as dorsal sinus, apodeme very short, spatulate, about one-tenth tegmen length; median lobe as in Fig. 96; pygidium as in Fig. 132. Etymology. The specific epithet, obesus (Latin, meaning fat) refers to the barrel-shaped form of this species which makes it instantly recognisable amongst its congeners. Biology. Specimens were captured in a static panel trap containing an alpha-pinene+ethanol lure, or collected at light. Distribution (Map 4). The four known specimens were collected in Queensland (Gladstone and Brisbane) and New South Wales (Blaxlands Ridge)., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 64, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

beswickensis species group Diagnosis. Anterior pronotal arch as broad or broader than middle and pronotal median sulcus mostly inconspicuous, shallow or apparently absent (synapomorphies within Notopilo); hind wing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 absent; outer profile of tegminal parameroid lobes gradually sloping inwards towards apices; apical lobes of median lobe well-developed, without apico-medial sclerite; eyes widely separated (frons width = 1.26–2.0 eye widths) (symplesiomorphy within Notopilo). Remarks. The beswickensis species group of Notopilo contains six species: N. beswickensis sp. nov.; N. calicis sp. nov.; N. eremosus sp. nov.; N. tompricensis sp. nov.; N. xanthoimprocerus sp. nov.; N. xanthoprolatus sp. nov.., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on page 38, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294

Notopilo congruus (Newman) comb. nov. (Figs 7, 11, 13, 52, 89, 125, 166, 181; Map 5) Opilo congruus Newman, 1842: 365. Type material. LECTOTYPE (here designated) ♁: Victoria: Type [circular label with red border] // Opilus congruus Newman Entomol 365 / 55.91 Pt. Philip [pale blue paper label, folded] // Opilus congruus Type Newm. Port Philip [white card label] // Opilus congruus Newman Entomol 365 [pale blue paper label, folded] (NHML) (Fig. 181). Comment on lectotype designation: As Newman (1842) made no reference to specimens in his description, it cannot be assumed that the nominal species group na me, Opilo congruus, was based on a single specimen (ICZN 1999: Recommendation 73F). The NHML ‘type’ must therefore be considered a syntype (ICZN 1999: Article 73.2) and available for lectotype designation (ICZN 1999:Article 74.1).We designate the aforementioned NHML specimen (Fig. 166) as the lectotype of Opilo congruus Newman for the purpose of fixing it as the sole name-bearing type of that nominal taxon. Condition of lectotype: Clean, left fifth pro-tarsomere and claw missing, numerous pronotal and elytral setae rubbed off. Comment on nomenclature: Several species have been erroneously erected under the genus Opilus Latreille (1806) rather than Opilo Latreille (1802) which has priority over the later, invalid, spelling. Following Article 11.9.3.2 of The Code (ICZN 1999), which states that “a species group name is deemed to have been published in combination with the correct original spelling of the generic name, even if it was actually published in combination with an emendation or incorrect spelling of the generic name”, the genus component of the original species group name spelling of Opilus congruus (see Newman 1842: 365) is deemed an incorrect subsequent spelling of the genusgroup name Opilo Latreille. The combination Opilo congruus Newman, 1842, is thefore recognised as the correct available name spelling. Other material examined. (87 specimens): Victoria: Ballarat 10/5/91 Froggatt // Opilo congruus Newm., Coll. & det. W.W. Froggatt // Opilo congruus Newm. Ballarat, V. (1, ANIC); Bedick Riv., Jan 1938, V., F.E. Wilson // O. congruus (1, NMV); Blackburn, V., T.G. Mills (1, NMV); 37.47S 148.44E VIC Cape Conran Coastal Park 24km SEW Orbost 12 Jan.-13 Feb. 2005 C.Lambkin, N. Starick / Malaise trap ANIC bulk sample 2623 heath under forest (1, ANIC); Australia: Chinaman I., Vict. 10.v.58, B.P. Moore // Opilo congruus Newm., Det. B.P. Moore (1, ANIC); Eltham, V., 22.11.18, F.E. Wilson // 428 // F. E. Wilson Collection // COL-65649 (1, NMV); Emerald, V., 4.7.20, C. Oke // COL-65650 (1, NMV); COL-65651 [on same pin as previous] (1, NMV); Frankston, 29.9.18/43 // C. E. Cole // congruus // COL-65657 (1, NMV); Frankston, V., 5.36, Goudie (1, NMV); Frankston, Victoria 5/36 // J.G. Brooks Bequest, 1976 (1, ANIC); Kilmore East, 2.4.20 (6, NMV); Kinglake West, V., 29.8.1954, A.N. (1, NMV); Australia, VIC, 20km s. Mansfield, 16.12.1990, leg. M. Baehr (1, RGCM); Melbourne, Deane (1, QM); F.T. Gully V. (1, QM); A.H. Elston Collection // Ararat V., H.W. Davey // K 304530 // 74 Opilo congruus Newm. (1, AM); Melbourne // Pascoe Coll. 93-60. (1 ♁, NHML); Moe, Vict. 14-4-1948, C.G.L.Gooding // C.G.L. Gooding Collection donated to A.N.I.C. 1979 (1,ANIC); Meyer, Nunawading, Vic, 13-8-58, Forest Hill (1,ANIC); Ringwood, V., R. Blackwood // K 304532 // Opilo congruus Newm. Id by A.H. Elston (1, AM); Tyres River, Vict. Oct 1969, T.Brown // C.G.L. Gooding Collection donated to ANIC 1979 (1, ANIC); Warburton District, Victoria (1, NMV); Warrandyte V., 2-3-97, LJ Cookson—9692 // Opilo congruus (Newman, 1842) det. JS Bartlett 2007 (1 ♀, JSBC); Victoria Australie // Museum Paris Coll. M. Pic (1 ♀, MNHN); Museum Paris Australie Victoria H. Rolle 1905 // Victoria // Opilo congruus Newm. Schenkling det. (1, MNHN); Victoria Austral // Ex Musaeo E. Hintz // Museum Paris 1952 Coll R. Oberthur (1, MNHN); Victoria Australie // Museum Paris Coll. M. Pic (3, MNHN); Victoria, Barton // Museum Paris Coll Gorham 1914 (1, MNHN); Victoria n. Holl. // Coll. E.W. Janson (2, MNHN). New South Wales: Armidale, N.S.W., 4-viii-1988, M.Coombs (1, QM); Australia, NSW, Aspley Gorge NP, 1- 2.12.1990, leg. M. Baehr (2, RGCM); Bega, N.S.W., Nov 1947, E. Smith (1, NMV); Blue Mtns., N. S. Wales, G. E. Bryant, Jan 1909 // G. Bryant Coll. 1919-147. (1, NHML); Blue Mountains, 1—05 H J C // congruus // COL-65642 (1, NMV); COL-65643 [on same pin as previous] (1, NMV); Bombala, N.S.W., 1929, Rev. A.J. Barrett // K 304554 // K60416 (1, AM); R.H. Mulder Collection // The Bonnet, 19-7-1959, N.S.W.—R.H.M. // K 304491 (1, AM); Australia, QLD [NSW], New England Ra. so Glen Innes, 10.12.1990, leg. Wachtel (1 ♁, 4, RGCM); Hornsby, C. Gibbons // K 304520 (1, AM); 35.30S 150.18E, Kioloa SF, 15km NE Batemans Bay, NSW, Dec.86, M.G. Robinson, flight interc. Trap (1, ANIC); Lane Cove, N.S.W., 28 Dec. 1948, N.W.Rodd. // K 304566 (1, AM); Michelago, N.S.W., viii.1938 // Brit Mus. 1947-431. (1, NHML); Mt Tomah Blue Mtns NSW 26.4. 82 N. W.Rodd // K 304495 (1, AM); National Pk., N. S. Wales, G. E. Bryant, 7.xii.08 // G. Bryant Coll. 1919-147. // Opilo congruus Newm. Compd. with Type (1, NHML); N S Wales, Strathfield (1, ZMHB); Sydney: coll. Luddermann // Schenkling det. // DEI Muncheberg Col—01941 (1 ♁, SDEI); K. K. Spence Collection // Sydney, 20 1 31, N. S. Wales, Dr. K. K. Spence // K 304555 (1, AM); nr. Sydney // Opilo congruus Newm // Pascoe Coll. 93-60. (1, NHML); Sydney // Museum Paris Coll. Castelnau Coll. Sedillot 1935 (1, MNHN); 28.53S 152.34E, 3km E of Tabulam, N.S.W., 23 Nov 1983, M.S.Harvey & D.C.F.Rentz (1, ANIC); N. S. Wales // Museum Paris 1952 Coll R. Oberthur (2, MNHN); N. S. Wales // Museum Paris Coll R. Oberthur (1, MNHN). Australian Capital Territory: Black Mt. Reserve, ACT, 28.vii.1971, S.Misko (1, ANIC); Blundell’s, F.C.T., G.A. Currie (1, ANIC); Australia, ACT, 10km w Cotter River, 9.12.1990, leg. M. Baehr (1, RGCM); Australia: ACT, Murrumbidgee R. near Canberra, 30-XI 1969 // Coll. H. Evans R.W. Matthews (1, ANIC). Queensland: Australia, QLD, Brisbane 1.1991 leg.Wachtel (1, RGCM); Broadwater, 20-7-25 (1, QDPC); Stanthorpe, Date 8-11-26 // Opilo congruus Newm. Id. by A. M. Lea (1, QDPC); Stanthorpe Dt., Q. Coll. H. Jarvis 11-19 (1 ♁, 1, QDPC); Tolga, N. Qld 22.x.85 J.D. Brown Light trap (1, QDPC); Tolga, N. Qld 31.x-6.xi.85 J.D. Brown Light trap (1, QDPC); Tolga, N. Qld 13-20.xi.1985 J.D. Brown Light trap (1, QDPC); Australia: n. Qld Tolga 14.xi.86 J.D. Brown light trap (1, QDPC); Australia: n. Qld Tolga 10-1 1986 J.D. Brown light trap (1, QDPC); Australia: n. Qld Tolga 3 xi 1985 J.D. Brown light trap (1, QDPC); Australia: n. Qld Tolga 16 xii 1985 J.D. Brown light trap (2, QDPC); Australia: n. Qld 7 km NE of Tolga 2-ii-1987 Storey & De faveri light trap (1, QDPC); Australia: n. Qld 7 km NE of Tolga 7-21.xi.1988 Storey & De faveri light trap (2, QDPC); Australia: n. Qld 7 km NE of Tolga Feb 1987 Storey & De faveri light trap (1, QDPC). Diagnosis. Pronotum evenly rounded laterally, disc not heavily punctate; elytra dark with orange fasciate and apical maculations (humeral maculae absent), fasciae meeting at suture, punctation lacking nodules (Fig. 13), 8 th stria beginning before fascia, striae 1–3 and 8–10 not reaching apical maculae; femoral bases yellow, apical half (or less) dark, tarsi with three ventral tarsal pads. Notopilo congruus comb. nov. can be separated from N. reduncus stat. rev., nom. n. by its bicoloured legs and from N. confusus sp. nov. by its elytral punctation lacking internal nodules. Redescription. Habitus: Fig. 166. Total length: 7–13.1 mm (lectotype 10.02 mm). Head: Vertex, frons (including supra-antennal elevation), genae and submentum dark brown (gula often slightly paler), clypeus and antennae chestnut-brown (clypeus partly black along epistomal suture), anteclypeus transparent yellow/orange, labrum and palpi orange; eyes separated by 0.65–0.85 eye widths (lectotype 0.75); frons subtly raised and semiglabrous medially (variable), dense with shallow punctation laterally, punctations more widely separated nearer to vertex; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.7:1 (maxillae) and 2.3:1 (labium); antennae reaching near base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Brown (slightly paler than frons), pronotal collar and arch paler; pronotum 1.15–1.26 times longer than wide (lectotype 1.2), sides round, widest just posterior of middle; subapical depression deeply v-shaped, disc with deep central sulcus (sulcus smooth, more open than linear), and baso-lateral semi-sulcate undulations, surface smooth in appearance, punctations small and occasional dorsally, slightly larger and denser laterally; moderately distributed with short fine multi-directional setae and long erect setae. Pterothorax: Ventrites brown, barely punctate, sparsely vested with short pale and occasional long setae; elytra brown (similar shade to pronotal disc) with orange markings (each elytron with a large apical macula and a narrow weakly curved transverse fascia at the mid-length which touches the suture); length to width ratio 2.67– 2.96:1 (lectotype 2.82); mostly 10-striate, 8 th stria beginning between first and fifth punctation of 7 th stria, striae 1–3 and 8–10 end between fascia and apical maculae, punctation lacking nodules, large and complete anterior to fascia, smaller with posterior edge indistinct posterior to fascia, epipleurae extending well into apical curve, interstriae with very fine short semi-reclinate setae (>1 per puncture) and longer thicker erect setae (Legs: Femora yellow basally, light brown apically (apical half of profemora and one-third of meso- and metafemora), tibiae and tarsi light brown, ventral tarsal pads yellowish; profemora very slightly swollen, other femora slender. Abdomen: Ventrites orange. Male genitalia: Tegmen (Fig. 7, 52) very broad at ‘bend’ of tegminal arms with parameroid lobes long, narrow and tapering to a point, dorsal sinus just over onethird tegmen length, gradually narrowing then expanded near extremity, internal limit flat (not rounded), ventral sinus a little less than half as long as dorsal sinus, tegminal arms gradually meeting apodeme, apodeme just over quarter tegmen length; median lobe as in Fig. 89; pygidium as in Fig. 125. Variation. Many specimens blacker than brown; the frons can be lightly (as in the lectotype) or more heavily punctated and with, or without (as in the lectotype), a shallow central impression; the degree in curvature of the tegminal parameroid lobes differs slightly between specimens (though the overall form and proportions of components of the tegmen are consistent). Biology. Collecting records for N. congruus span the entire year for Victoria and it has been collected in summer and winter in New South Wales and Queensland. Collecting methods include malaise and light trapping. Distribution (Map 5). Temperate broadleaf and mixed forest ecoregions of Victoria, New South Wales and southeast Queensland; tropical moist broadleaf forests of far north Queensland; Mediterranean environs of coastal South Australia. Absent from Western Australia and the Northern Territory. Remarks. Despite the abovementioned differences of the tegminal parameroid lobes and frontal sculpturing, the general form of the male tegmen, also elytral and pronotal form and sculpturing, are consistent in specimens from far north Queensland to southern Victoria. We therefore consider the specimens examined to constitute a single species., Published as part of Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, pp. 1-81 in Zootaxa 5220 (1) on pages 51-53, DOI: 10.11646/zootaxa.5220.1.1, http://zenodo.org/record/7441294, {"references":["Newman, E. (1842) List of insects collected at Port Phillip, South Australia, by Edmund Thomas Higgins, Esq. The Entomologist, 1 (23), 361 - 369.","ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp.","Latreille, P. A. (1806) Genera Crustaceorum et Insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plusimus explicata. Vol. I. A. Koenig, Paris, xviii + 296 pp. https: // doi. org / 10.5962 / bhl. title. 65741","Latreille, P. A. (1802) Histoire Naturelle, Generale et Particuliere des Crustaces et des Insectes. L'Imprimerie de F. Dufart, Paris. [unknown pagination] https: // doi. org / 10.5962 / bhl. title. 15764"]}