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Dichelonyx linearis (Gyllenhal, 1817) Wickham 1911; Cornell 1972; Worthington and Larsen 2010; ISIC; IWUC; DAVC; ELFC; MJPC. Collection dates: April–June. Collected at light; located mating pairs on basswood leaves (Tilia americana) after sunset. County records (Figure 102): Allamakee, Benton, Boone, Buchanan, Chickasaw, Clayton, Dallas, Delaware, Fayette, Floyd, Fremont, Hardin, Howard, Johnson, Lee, Linn, Polk, Story, Van Buren, Winnebago, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 30, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Cornell, J. F., Jr. 1972. A taxonomic review of the beetle genus Dichelonyx Harris in North America (Scarabaeidae: Melolonthinae). PhD. Dissertation. Oregon State University. 253 p.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Ataenius spretulus Haldeman, 1848 Cartwright 1974; Stebnicka and Lago 2005; Worthington and Larsen 2010; ISIC; ELFC; PKLC. Collection dates: March–September. Collected at light and in flight during day. County records (Figure 77): Allamakee, Appanoose, Black Hawk, Dallas, Decatur, Fremont, Lee, Lucas, Osceola, Polk, Scott, Story, Taylor, Van Buren, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 24, DOI: 10.5281/zenodo.5353792, {"references":["Cartwright, O. L. 1974. Ataenius, Aphotaenius, and Pseudoataenius of the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). Smithsonian Contribution to Zoology 154: 1 - 106.","Stebnicka, Z. T., and P. K. Lago. 2005. The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19 (1 - 2): 55 - 83.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Phyllophaga congrua (LeConte, 1856) Travis 1934; Luginbill and Painter 1953; ISIC; DAVC; ELFC. Collection dates: March, May, September. Collected at light. Travis (1934) reported 44 specimens from Iowa indicating it was infrequent to rare in the state. Specimen label data: Missouri Valley, Harrison Co., Iowa, 20 May 1932 [14], G. C. Decker (ISIC); Super 8 Motel, Decatur Co., Iowa, 31 March 2012 [2], lights, ELF (ELFC). County records (Figure 120): Decatur, Harrison, Taylor., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 34, DOI: 10.5281/zenodo.5353792, {"references":["Travis, B. V. 1934. Phyllophaga of Iowa. Iowa State Journal of Science 8 (2): 313 - 365.","Luginbill, P., and H. R. Painter. 1953. May beetles of the United States and Canada. United States Department of Agriculture Technical Bulletin 1061: 1 - 102."]}

Tomarus gibbosus (DeGeer, 1774) Putnam 1876; Wickham 1911; Hardy 1991; Ratcliffe and Cave 2017; ISIC; IWUC; ELFC; DAVC. Collection dates: April–August. Collected at light. County records (Figure 175): Adair, Bremer, Cherokee, Dallas, Decatur, Delaware, Dickinson, Franklin, Fremont, Guthrie, Harrison, Henry, Jefferson, Johnson, Lee, Linn, Louisa, Mahaska, Marion, Monona, Monroe, Muscatine, Osceola, Page, Plymouth, Polk, Pottawattamie, Poweshiek, Scott, Story, Taylor, Van Buren, Wapello, Winnebago, Winneshiek, Woodbury., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 48, DOI: 10.5281/zenodo.5353792, {"references":["Putnam, J. D. 1876. List of Coleoptera found in the vicinity of Davenport, Iowa. Proceedings of the Davenport Academy of Natural Sciences 1: 169 - 173.","Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Hardy, A. 1991. A catalog of Coleoptera of America north of Mexico. Family: Scarabaeidae. Subfamilies Rutelinae and Dynastinae. Volume 34, part 2. United States Department of Agriculture Handbook 529: 1 - 57.","Ratcliffe, B. C., and R. D. Cave. 2017. The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae: Dynastinae. Bulletin of the University of Nebraska State Museum 30: 1 - 298."]}

Onthophagus orpheus pseudorpheus Howden and Cartwright, 1963 Howden and Cartwright 1963; ISIC; CASC (Paratype); DAVC. Collection dates: April–June. Specimen label data: Fish Farm Mounds, Allamakee Co., Iowa, 25 May 2003, horse dung, DAV (DAVC); Shimek State Forest, Lee Co., Iowa, 26 June 2005, horse dung, DAV (DAVC). County records (Figure 85): Henry, Johnson, Story, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 26, DOI: 10.5281/zenodo.5353792, {"references":["Howden, H. F., and O. L. Cartwright. 1963. Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum 114 (3467): 1 - 135."]}

Ataenius platensis (Blanchard, 1846) ELFC. Collection dates: July. Specimen label data: First known Iowa specimen and NEW STATE RECORD: Timberhill Savanna, Decatur Co., Iowa, 19 July 2011, UV light, ELF (ELFC). This is the only specimen from Iowa. County record: Decatur., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 23, DOI: 10.5281/zenodo.5353792

Hoplia modesta Haldeman, 1843 Wickham 1911; Boyer 1940; Hardy 1977; Harpootlian 2001; ISIC; USNM; DAVC; ELFC. Collection dates: June. Collected from dogwood flowers (Cornus spp.). County records (Figure 111): Allamakee, Black Hawk, Dallas, Dickinson, Henry, Iowa, Johnson, Lee, Linn, Polk, Story, Van Buren., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 32, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Boyer, L. B. 1940. A revision of the species of Hoplia in America North of Mexico (Coleoptera: Scarabaeidae). Microentomology 5 (1): 1 - 31.","Hardy, A. R. 1977. A revision of the Hoplia of the Nearctic realm (Coleoptera: Scarabaeidae). California Department of Food and Agriculture, Occasional Papers in Entomology 23: 1 - 48.","Harpootlian, P. J. 2001. Scarab beetles (Coleoptera: Scarabaeidae) of South Carolina. Biota of South Carolina. Volume 2. Clemson University; Clemson, SC. 157 p."]}

Lucanus capreolus (Linnaeus, 1764) Wickham 1911; ISIC; IWUC; DAVC; ELFC; PKLC; MJPC. Collection dates: June–August. Collected at light. County records (Figure 38): Adair, Allamakee, Appanoose, Benton, Boone, Calhoun, Cerro Gordo Clayton, Dallas, Decatur, Delaware, Des Moines, Fremont, Hardin, Henry, Humboldt, Iowa, Jackson, Jefferson, Johnson, Keokuk, Kossuth, Lee, Linn, Louisa, Lucas, Monona, Monroe, Muscatine, Page, Polk, Sac, Scott, Story, Union, Van Buren, Wapello, Warren, Washington, Webster., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 13, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40."]}

Anomala flavipennis Burmeister, 1844 DAVC; ELFC. Collection dates: June–July. Collected at light. Specimen label data: First known Iowa specimens and NEW STATE RECORD: Waubonsie State Park, Fremont Co., Iowa, 3 July 2003, UV light, DAV (DAVC); Shimek State Forest, Van Buren Co., Iowa, 4 July 2004, UV light, ELF (ELFC). County records (Figure 164): Fremont, Pottawattamie, Van Buren., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 44, DOI: 10.5281/zenodo.5353792

Ataenius abditus Haldeman, 1848 Putnam 1876; Wickham 1911; Worthington and Larsen 2010; ISIC; IWUC; ELFC; DAVC. Collection dates: April–August. Collected under debris on sandbar and sifted from sandbar along a wooded stream. County records (Figure 73): Clayton, Dallas, Des Moines, Dickinson, Henry, Howard, Johnson, Keokuk, Lee, Muscatine, Story, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 23, DOI: 10.5281/zenodo.5353792, {"references":["Putnam, J. D. 1876. List of Coleoptera found in the vicinity of Davenport, Iowa. Proceedings of the Davenport Academy of Natural Sciences 1: 169 - 173.","Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Serica mystaca Dawson, 1922 Dawson 1922; ISIC; ELFC; DAVC; PKLC. Collection dates: April–July. Collected at light. County records (Figure 156): Appanoose, Boone, Dallas, Fremont, Johnson, Linn, Polk, Story, Van Buren, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 42, DOI: 10.5281/zenodo.5353792

Ataenius imbricatus (Melsheimer, 1846) ELFC. Collection dates: July. Specimen label data: First known Iowa specimen and NEW STATE RECORD: Yellow River State Forest, Allamakee Co., Iowa, 9 July 2005, UV light, ELF (ELFC). This is the only Iowa specimen. County record: Allamakee., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 23, DOI: 10.5281/zenodo.5353792

Dialytes striatulus (Say, 1825) Wickham 1911; Gordon and Skelley 2007; MCZC; ELFC; DAVC; MJPC. Collection dates: June–August. Gordon and Skelley (2007) designated a specimen labeled “ IOWA ” as the Neotype to replace Say’s destroyed type: “ Jackson Co., Maquoketa Caves St. Pk., 29 vii 1968, S. Beck, Ber#147, forest litter u. carrion/ Neotype (red paper)/ M. C. Z. Neotype 32754/ Dialytes striatulus (Say) Des. F. Chalumeau ‘82/ Neotype Trox striatulus Say, des. P. Skelley 2006.” Also collected at lights. County records (Figure 56): Fremont, Jackson, Johnson., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 17, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Gordon, R. D., and P. E. Skelley. 2007. A monograph of the Aphodiini inhabiting the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). The American Entomological Institute Memoirs 79: 1 - 580."]}

Dyscinetus morator (Fabricius, 1798) Wickham 1911; Evans 2014; Ratcliffe and Cave 2017; ISIC; IWUC; UMRC; CASC; DAVC; ELFC. Collection dates: April–July. Specimen label data: Nevada, Story Co., Iowa, 3 April 1902 [2], [no collector], determined H. F. Howden 1953 (ISIC); Ottumwa, Wapello Co., Iowa, 29 May 1937, Floyd Andre, determined H. F. Howden 1953 (ISIC); Soldier, Monona Co., Iowa, 1 June 1971, light trap, T. Orwig (ELFC); Shimek State Forest, Van Buren Co., Iowa, 9 June 2016 [2], UV light, ELF (ELFC). County records (Figure 172): Fremont, Jackson, Johnson, Henry, Lee, Monona, Muscatine, Polk, Story, Tama, Van Buren, Wapello., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 47, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Evans, A. V. 2014. Beetles of eastern North America. Princeton University Press; Princeton, NJ. 560 p.","Ratcliffe, B. C., and R. D. Cave. 2017. The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae: Dynastinae. Bulletin of the University of Nebraska State Museum 30: 1 - 298."]}

Cyclocephala borealis Arrow, 1911 Ratcliffe 1991; Ratcliffe and Paulsen 2008; Evans 2014; Ratcliffe and Cave 2017; ISIC. Collection dates: June. Ratcliffe and Cave (2017), following a rather exhaustive survey of U.S. collections, did not include Iowa records. However, C. borealis does occur in neighboring counties in western Illinois indicating a possibility that the species might occur in eastern Iowa. Recently, one of us (ELF) discovered one specimen in the collection at Iowa State University. Specimen label data: Clinton, Clinton Co., Iowa, 14 June 1949, D. R. Nicholson, determined H. F. Howden 1953 (ISIC). Only one specimen from Iowa. County record: Clinton., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 46, DOI: 10.5281/zenodo.5353792, {"references":["Ratcliffe, B. C. 1991. The scarab beetles of Nebraska. Bulletin of the University of Nebraska State Museum 12: 1 - 333.","Ratcliffe, B. C., and M. J. Paulsen. 2008. The scarabaeoid beetles of Nebraska. Bulletin of the University of Nebraska State Museum 22: 1 - 570.","Evans, A. V. 2014. Beetles of eastern North America. Princeton University Press; Princeton, NJ. 560 p.","Ratcliffe, B. C., and R. D. Cave. 2017. The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae: Dynastinae. Bulletin of the University of Nebraska State Museum 30: 1 - 298."]}

Pardalosus sayi (Gordon, 2006) MJPC. Collection dates: November. Specimen label data: First nine known Iowa specimens and NEW STATE RECORD: Waubonsie State Park, Fremont Co., Iowa, 11 November 2009, in leaf litter of steps at picnic shelter, MJP (MJPC). Only nine specimens from Iowa. County record: Fremont., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on pages 20-21, DOI: 10.5281/zenodo.5353792

Dellacasiellus concavus (Say, 1823) Wickham 1911; Gordon 1983; Worthington and Larsen 2010; Connior et al. 2014; ISIC; IWUC; UMRC; FSCA; DAVC; ELFC; PKLC. Collection dates: May–June, August. Collected at lights, on pocket gopher mounds, and sifted from pocket gopher mounds (Geomys bursarius) late afternoon to dusk. County records (Figure 54): Allamakee, Black Hawk, Des Moines, Harrison, Henry, Howard, Kossuth, Linn, Johnson, Osceola, Polk, Pottawattamie, Story, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 17, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Gordon, R. D. 1983. Studies on the genus Aphodius of the United States and Canada (Coleoptera: Scarabaeidae). VII. Food and habitat; distribution; key to eastern species. Proceedings of the Entomological Society of Washington 85 (4): 633 - 652.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90.","Connior, M. B., P. W. Kovarik, S. Chordas III, H. W. Robison, and P. E. Skelley. 2014. Coleoptera (Histeridae, Leiodidae, and Scarabaeidae) inhabiting the burrows of Baird's Pocket Gopher (Rodentia: Geomyidae: Geomys breviceps) in Arkansas. Insecta Mundi 0389: 1 - 27."]}

Copris fricator fricator (Fabricius, 1787) Putnam 1876; Wickham 1911; Worthington and Larsen 2010; ISIC; IWUC; CCMV; USNM; DAVC; ELFC; PKLC. Collection dates: April–October. Collected at lights and from dung. County records (Figure 90): Adair, Allamakee, Appanoose, Black Hawk, Boone, Buchanan, Carroll, Cherokee, Clayton, Dallas, Decatur, Delaware, Des Moines, Dickinson, Floyd, Fremont, Henry, Howard, Iowa, Jefferson, Johnson, Keokuk, Kossuth, Lee, Linn, Louisa, Lucas, Mahaska, Monona, Monroe, Muscatine, Osceola, Page, Polk, Pottawattamie, Poweshiek, Scott, Story, Taylor, Union, Van Buren, Wapello, Warren, Washington, Wayne, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 27, DOI: 10.5281/zenodo.5353792, {"references":["Putnam, J. D. 1876. List of Coleoptera found in the vicinity of Davenport, Iowa. Proceedings of the Davenport Academy of Natural Sciences 1: 169 - 173.","Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Phyllophaga quercus (Knoch, 1801) Wickham 1911; Jaques 1926; Travis 1934. Collection dates: no date of collection given with literature record. No specimens located in collections examined. County records (Figure 146): Allamakee, Story, Wapello., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 40, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Jaques, H. E. 1926. A preliminary survey of May beetles (Phyllophaga spp.) in Iowa. Proceedings of the Iowa Academy of Science 33: 337 - 339.","Travis, B. V. 1934. Phyllophaga of Iowa. Iowa State Journal of Science 8 (2): 313 - 365."]}

Diplotaxis basalis Fall, 1909 ISIC; IWUC. Collection dates: April–June. Specimen label data: First report from Iowa and NEW STATE RECORD: Experiment Station, Ames, Story Co., Iowa, 22 April 1897, [no collector] (ISIC); Ames, Story Co., Iowa, 29 May 1916, R. L. Webster (ISIC); Ottumwa, Wapello Co., Iowa, 29 May 1937 [2], Floyd Andre (ISIC). County records (Figure 105): Boone, Davis, Henry, Jefferson, Lee, Louisa, Scott, Story, Wapello, Washington., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 30, DOI: 10.5281/zenodo.5353792

Trox scaber (Linnaeus, 1767) Wickham 1911; Vaurie 1955; ISIC; IWUC; CASC; DAVC; ELFC; MJPC. Collection dates: March–June. Specimens collected at light. Specimen label data: Hitchcock Nature Area, Pottawattamie Co., Iowa, 10 June 2008, light, ELF (ELFC). County records (Figure 22): Des Moines, Henry, Johnson, Kossuth, Linn, Monona, Pottawattamie, Story., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 9, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Vaurie, P. 1955. A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). Bulletin of the American Museum of Natural History 106 (1): 1 - 89."]}

Ataenius cognatus LeConte, 1858 Wickham 1911; Hendrickson 1930; IWUC. Collection dates: April–September. Specimen label data: Mt. Pleasant, Henry Co., Iowa, 8 August 1928, E. P. Meiners (IWUC); Clayton Co., Iowa, 4 August 1932, MWJ (IWUC); Wapello Co., Iowa, 3 May 1934, Albert, determined Marshall 1943 (IWUC). County records (Figure 74): Appanoose, Black Hawk, Boone, Buchanan, Clayton, Des Moines, Harrison, Henry, Howard, Iowa, Jasper, Johnson, Louisa, Page, Polk, Pottawattamie, Poweshiek, Scott, Sioux, Story, Union, Wapello., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 23, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Hendrickson, G. O. 1930. Studies on the insect fauna of Iowa prairies. Iowa State Journal of Science 4 (2): 49 - 175."]}

Euphoria pilipennis (Kraatz, 1883) Orozco 2012; USNM (two specimens labeled only “IA” in Smithsonian collection (M.J. Paulsen, pers. comm., 2012)). Collection dates: no date of collection given with literature record. County record: no specific locality given., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 49, DOI: 10.5281/zenodo.5353792, {"references":["Orozco, J. 2012. Monographic revision of the American genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae). The Coleopterists Society Monograph 11: 1 - 182."]}

Hoplia laticollis LeConte, 1856 Worthington and Larsen 2010; ELFC; DAVC. Collection dates: June–July. Specimens collected from dogwood flowers (Cornus spp.) and sweeping; one specimen from deer dung baited pitfall trap. Specimen label data: Fish Farm Mounds Wildlife Area, Allamakee County, Iowa, 2 July 2003, sweeping, ELF (ELFC). County records (Figure 110): Allamakee, Black Hawk, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 32, DOI: 10.5281/zenodo.5353792, {"references":["Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Omorgus scabrosus (Palisot de Beauvois, 1818) King 1914; DAVC; ELFC. Collection dates: June–July. Specimens collected at carcasses and at UV light. Specimen label data: Cedar Rapids, Linn Co., Iowa, 8 July 2000, mouse carcass, DAV (DAVC). County records (Figure 32): Fremont, Henry, Linn., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 11, DOI: 10.5281/zenodo.5353792, {"references":["King, I. N. 1914. The Coleoptera of Henry County, Iowa. Proceedings of the Iowa Academy of Science 21: 317 - 340."]}

Phyllophaga delata (Horn, 1887) ELFC; DAVC. Collection dates: May–June. Specimen label data: First two known Iowa specimens and NEW STATE RECORD: Sharon Bluffs State Park, Appanoose Co., Iowa, 20 May 2006 [2], UV light, ELF, DAV (ELFC, DAVC). This is a range extension northward of approximately 145 km (90 miles) from nearest known specimen collected in Missouri (Kristin B. Simpson, pers. comm., 2007). Only eighteen specimens from Iowa. County records (Figure 124): Appanoose, Van Buren., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 35, DOI: 10.5281/zenodo.5353792, {"references":["Horn, G. W. 1887. Revision of the species of Lachnosterna of America north of Mexico. Transactions of the American Entomological Society 14: 209 - 296."]}

Trox spinulosis spinulosis Robinson, 1940 Vaurie 1955; ELFC; DAVC. Collection dates: April–July. Specimens were collected at light, carnivore dung, and road kill. County records (Figure 24): Appanoose, Black Hawk, Bremer, Dallas, Decatur, Dickinson, Fremont, Hardin, Henry, Johnson, Lee, Linn, Madison, Mills, Monona, Muscatine, Page, Pottawattamie, Scott, Story, Van Buren, Woodbury., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on pages 9-10, DOI: 10.5281/zenodo.5353792, {"references":["Robinson, M. 1940. Studies in the Scarabaeidae II. Transactions of the American Entomological Society 66 (2): 141 - 160.","Vaurie, P. 1955. A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). Bulletin of the American Museum of Natural History 106 (1): 1 - 89."]}

Trox sordidus LeConte, 1854 Wickham 1911; King 1914; Vaurie 1955, 1958; IWUC; CASC. Collection dates: July. Specimen label data: Washington, Washington Co., Iowa, 8 July 1913, Mayhew, determined Cartwright 1938 (IWUC). County records (Figure 23): Henry, Johnson, Story, Washington., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 9, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","King, I. N. 1914. The Coleoptera of Henry County, Iowa. Proceedings of the Iowa Academy of Science 21: 317 - 340.","Vaurie, P. 1955. A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). Bulletin of the American Museum of Natural History 106 (1): 1 - 89.","Vaurie, P. 1958. New distribution records of North American Trox (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 12: 43 - 46."]}

Omorgus punctatus (Germar, 1824) Wickham 1911; ISIC; IWUC. Collection dates: May–July. Specimen label data: Mt. Pleasant, Henry Co., Iowa, 26 May 1913, light, Jaques (IWUC). County records (Figure 31): Des Moines, Fremont, Henry, Pottawattamie, Story., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 11, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40."]}

Anomala ludoviciana Schaeffer, 1906 Potts 1977; Hardy 1991; ISIC; IWUC; UMRC; USNM; DAVC; ELFC. Collection dates: April, June–July. Collected sweeping prairie and in flight over prairie. Specimen label data: Cedar Hills Sand Prairie State Preserve, Black Hawk Co., Iowa, 5 July 2003, sweeping, ELF (ELFC); Marietta Sand Prairie State Preserve, Marshall Co., Iowa, 13 June 2003, vervain flower (Verbena stricta), ELF (ELFC). County records (Figure 166): Black Hawk, Linn, Marshall, Monona, Pottawattamie, Story, Woodbury., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 45, DOI: 10.5281/zenodo.5353792, {"references":["Potts, R. W. L. 1977. Revision of the Scarabaeidae: Anomalinae 2. An annotated checklist of the Anomala of the United States and Canada. The Pan-Pacific Entomologist 53 (1): 34 - 42.","Hardy, A. 1991. A catalog of Coleoptera of America north of Mexico. Family: Scarabaeidae. Subfamilies Rutelinae and Dynastinae. Volume 34, part 2. United States Department of Agriculture Handbook 529: 1 - 57."]}

Onthophagus orpheus canadensis (Fabricius, 1801) Howden and Cartwright 1963; Worthington and Larsen 2010; ELFC; DAVC; PKLC; MJPC. Collection dates: April–July, September–October. Collected from deer dung, on dead toad, under leaves, and on a log. County records (Figure 83): Allamakee, Appanoose, Black Hawk, Bremer, Butler, Clayton, Dallas, Delaware, Dubuque, Fayette, Hardin, Johnson, Jones, Lee, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 25, DOI: 10.5281/zenodo.5353792, {"references":["Howden, H. F., and O. L. Cartwright. 1963. Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum 114 (3467): 1 - 135.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Dichelonyx kirbyi Brown, 1946 ELFC. Collection dates: May–June. Specimen label data: First known Iowa specimens and NEW STATE RECORD: Cedar Bend Park, Bremer Co., Iowa, 27 May 1998, under playground equipment, ELF, J. J. Freese (ELFC); Backbone State Park, Delaware Co., Iowa, 2 June 2006, UV light, ELF (ELFC). Only two specimens from Iowa. County records (Figure 101): Bremer, Delaware., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 30, DOI: 10.5281/zenodo.5353792

Diplotaxis blanchardi Vaurie, 1956 Vaurie 1956, 1960; ELFC; DAVC. Collection dates: May–June, September. Collected at light. County records (Figure 106): Dallas, Decatur, Fremont, Hardin, Henry, Linn, Lucas, Plymouth, Story, Van Buren., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 31, DOI: 10.5281/zenodo.5353792, {"references":["Vaurie, P. 1956. Diplotaxis of the eastern United States, with a new species and other notes (Coleoptera, Scarabaeidae). The Coleopterists Bulletin 10: 1 - 9.","Vaurie, P. 1960. A revision of the genus Diplotaxis (Coleoptera, Scarabaeidae, Melolonthinae). Part 2. Bulletin of the American Museum of Natural History 120 (2): 161 - 434."]}

Phyllophaga tristis (Fabricius, 1781) Putnam 1876; Wickham 1911; Travis 1934; Worthington and Larsen 2010; ISIC; IWUC; ELFC; DAVC; PKLC. Collection dates: April–June. Collected at light and under soil pushup on path. Travis (1934) reported this species as the fourth most common species in Iowa based on collections. County records (Figure 149): Allamakee, Appanoose, Bone, Bremer, Dallas, Decatur, Delaware, Des Moines, Dubuque, Franklin, Fremont, Henry, Jackson, Johnson, Kossuth, Linn, Louisa, Lucas, Madison, Pottawattamie, Ringgold, Scott, Story, Van Buren, Wapello, Washington, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 41, DOI: 10.5281/zenodo.5353792, {"references":["Putnam, J. D. 1876. List of Coleoptera found in the vicinity of Davenport, Iowa. Proceedings of the Davenport Academy of Natural Sciences 1: 169 - 173.","Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Travis, B. V. 1934. Phyllophaga of Iowa. Iowa State Journal of Science 8 (2): 313 - 365.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Bolbocerosoma farctum Fabricius, 1781 Wickham 1911; Hendrickson 1930; DAVC. Collection dates: August. Specimen label data: Geode State Park, Henry Co., Iowa, 13 August 2003, UV light, DAV (DAVC). This is the only specimen currently known from Iowa. All specimens in university collections previously determined to be Bolbocerosoma farctum where actually B. bruneri. County record: Henry., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 5, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Hendrickson, G. O. 1930. Studies on the insect fauna of Iowa prairies. Iowa State Journal of Science 4 (2): 49 - 175."]}

Trichiotinus piger (Fabricius, 1775) Wickham 1911; Howden 1968; Worthington and Larsen 2010; ISIC; IWUC; DAVC; ELFC. Collection record: May–July. Collected from dogwood (Cornus spp.), wild rose (Rosa spp.), common milkweed (Asclepias syriaca), and fleabane (Erigeron spp.) flowers. County records (Figure 187): Adams, Allamakee, Appanoose, Crawford, Dallas, Des Moines, Fremont, Harrison, Henry, Jackson, Johnson, Kossuth, Lee, Linn, Madison, Muscatine, Page, Pottawattamie, Story, Taylor, Union, Warren, Washington, Wayne, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 52, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Howden, H. F. 1968. A review of the Trichiinae of North and Central America (Coleoptera: Scarabaeidae). Memoirs of the Entomological Society of Canada 64: 1 - 77.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Trox capillaris Say, 1824 Vaurie 1955; Zidek 2013; ISIC; IWUC; WIRC; DAVC; ELFC; MJPC. Collection dates: April–June. Collected under old deer carcasses along highways. Specimen label data: Waubonsie Access, Fremont Co., Iowa, 12 June 2005 [2], deer carcass, ELF, DAV, determined MJP 2005 (DAVC, ELFC); south of Farmington, Van Buren Co., Iowa, 5 May 2011 [18], deer carcass, DAV, ELF (DAVC, ELFC). County records (Figure 18): Dallas, Fremont, Jefferson, Linn, Van Buren, Story., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 8, DOI: 10.5281/zenodo.5353792, {"references":["Vaurie, P. 1955. A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). Bulletin of the American Museum of Natural History 106 (1): 1 - 89.","Zidek, J. 2013. Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 0314: 1 - 38."]}

Trox hamatus Robinson, 1940 UMRC; DAVC; ELFC; MJPC. Collection dates: May–July. Collected at lights and carnivore dung. Specimen label data: First report from Iowa and NEW STATE RECORD: Waubonsie State Park, Fremont Co., Iowa, 15 June 2003, carnivore dung, DAV (DAVC); Shimek State Forest, Van Buren Co., Iowa, 21 June 2003, UV light, DAV (DAVC); Yellow River Forest, Allamakee Co., Iowa, 20 May 2004, dung, ELF (ELFC). County records (Figure 20): Allamakee, Appanoose, Dallas, Fremont, Hardin, Henry, Lee, Pottawattamie, Van Buren, Warren., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 9, DOI: 10.5281/zenodo.5353792, {"references":["Robinson, M. 1940. Studies in the Scarabaeidae II. Transactions of the American Entomological Society 66 (2): 141 - 160."]}

Phyllophaga inversa (Horn, 1887) Wickham 1911; Travis 1934; Worthington and Larsen 2010; ISIC; IWUC; ELFC; DAVC; PKLC. Collection dates: May–June. Collected at light. County records (Figure 138): Appanoose, Bremer, Dallas, Davis, Decatur, Dickinson, Dubuque, Henry, Jefferson, Johnson, Louisa, Lucas, Monroe, Polk, Story, Wapello, Winneshiek., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 38, DOI: 10.5281/zenodo.5353792, {"references":["Horn, G. W. 1887. Revision of the species of Lachnosterna of America north of Mexico. Transactions of the American Entomological Society 14: 209 - 296.","Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Travis, B. V. 1934. Phyllophaga of Iowa. Iowa State Journal of Science 8 (2): 313 - 365.","Worthington, R. J., and K. J. Larsen. 2010. An annotated checklist of scarab beetles (Coleoptera: Scarabaeidae) from northeastern Iowa. The Great Lakes Entomologist 43 (1 - 4): 77 - 90."]}

Serica evidens Blatchley, 1910, 1919 (name change) Dawson 1932; ISIC; IWUC; USNM; DAVC; ELFC. Collection dates: April–July. Collected at light and in leaf litter of woodland. County records (Figure 154): Appanoose, Boone, Delaware, Fayette, Fremont, Henry, Johnson, Keokuk, Lee, Linn, Madison, Muscatine, Story., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 42, DOI: 10.5281/zenodo.5353792, {"references":["Blatchley, W. S. 1910. Coleoptera or beetles known to occur in Indiana. The Nature Publishing Co.; Indianapolis, Indiana. 1,386 p.","Blatchley, W. S. 1919. Change of name. The Canadian Entomologist 51: 153.","Dawson, R. W. 1932. New species of Serica (Scarabaeidae). VI. Journal of the New York Entomological Society 40 (4): 529 - 562."]}

Dichelonyx albicollis (Burmeister, 1855) DAVC. Collection dates: June. Specimen label data: First known Iowa specimen and NEW STATE RECORD: Cedar Rapids, Linn Co., Iowa, June 2003, UV light, J. O. Durbin (DAVC). Only one specimen from Iowa. County record: Linn., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 29, DOI: 10.5281/zenodo.5353792