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Acknowledgements: The authors thank G. Marciani and O. Jöris for comments on archaeology; C. Jeong, M. Spyrou and K. Prüfer for comments on genetics; M. O’Reilly for graphical support for Fig. 5 and Extended Data Fig. 9; the entire IT and laboratory teams at the Department of Archaeogenetics of MPI-SHH for technical assistance; M. Meyer and S. Nagel for support with single-stranded library preparation; K. Post, P. van Es, J. Glimmerveen, M. Medendorp, M. Sier, S. Dikstra, M. Dikstra, R. van Eerden, D. Duineveld and A. Hoekman for providing access to human specimens from the North Sea (The Netherlands); M. D. Garralda and A. Estalrrich for providing access to human specimens from La Riera (Spain); J. Górski and M. Zając for providing access to human specimens from Maszycka cave; C. Di Patti for providing access to human specimens from San Teodoro 2 (Italy); P. Blaževičius for providing access to the Donkalnis human remains and the new radiocarbon dates; the Italian Ministry of Culture and Soprintendenza Archeologia Belle Arti e Paesaggio for the Provinces of Verona, Rovigo, and Vicenza for granting access to the human remains of Tagliente 2; F. Fontana, who carries out investigations of the Riparo Tagliente site (Italy); the Friuli Venezia Giulia Superintendency for providing access to the human tooth Pradis 1; and the Soprintendenza Archeologia Belle Arti e Paesaggio for the Provinces of Barletta-Andria-Trani and Foggia for providing access to the Paglicci human remains. This project has received funding by the European Research Council under the European Union’s Horizon 2020 research and innovation programme under grant agreements no. 803147-RESOLUTION (to S.T.), no. 771234-PALEoRIDER (to W.H.), no. 864358 (to K.M.), no. 724703 and no. 101019659 (to K.H.). K.H. is also supported by the Deutsche Forschungsgemeinschaft (DFG FOR 2237). E.A. has received funding from the Van de Kamp fonds. PACEA co-authors of this research benefited from the scientific framework of the University of Bordeaux’s IdEx Investments for the Future programme/GPR Human Past. A.G.-O. is supported by a Ramón y Cajal fellowship (RYC-2017-22558). L. Sineo, M.L. and D.C. have received funding from the Italian Ministry of University and Research (MUR) PRIN 2017 grants 20177PJ9XF and 20174BTC4R_002. H. Rougier received support from the College of Social and Behavioral Sciences of CSUN and the CSUN Competition for RSCA Awards. C.L.S. and T. Saupe received support from the European Union through the European Regional Development Fund (project no. 2014-2020.4.01.16-0030) and C.L.S. received support from the Estonian Research Council grant PUT (PRG243). S. Shnaider received support from the Russian Science Foundation (no. 19-78-10053)., Modern humans have populated Europe for more than 45,000 years1,2. Our knowledge of the genetic relatedness and structure of ancient hunter-gatherers is however limited, owing to the scarceness and poor molecular preservation of human remains from that period3. Here we analyse 356 ancient hunter-gatherer genomes, including new genomic data for 116 individuals from 14 countries in western and central Eurasia, spanning between 35,000 and 5,000 years ago. We identify a genetic ancestry profile in individuals associated with Upper Palaeolithic Gravettian assemblages from western Europe that is distinct from contemporaneous groups related to this archaeological culture in central and southern Europe4, but resembles that of preceding individuals associated with the Aurignacian culture. This ancestry profile survived during the Last Glacial Maximum (25,000 to 19,000 years ago) in human populations from southwestern Europe associated with the Solutrean culture, and with the following Magdalenian culture that re-expanded northeastward after the Last Glacial Maximum. Conversely, we reveal a genetic turnover in southern Europe suggesting a local replacement of human groups around the time of the Last Glacial Maximum, accompanied by a north-to-south dispersal of populations associated with the Epigravettian culture. From at least 14,000 years ago, an ancestry related to this culture spread from the south across the rest of Europe, largely replacing the Magdalenian-associated gene pool. After a period of limited admixture that spanned the beginning of the Mesolithic, we find genetic interactions between western and eastern European hunter-gatherers, who were also characterized by marked differences in phenotypically relevant variants.

Archaeological evidence of the use of animal hides in burials have recently become known in the Late Iron Age and the Middle Ages (800—1300 AD) in Finland and Karelia (Russia). The purpose of the article is to interpret the ritual of wrapping the dead in hides of Eurasian elk and reindeer. For the first time, the authors use a combination of innovative methods of archaeozoological analysis to study the remains of fur, and the method of ethnoarchaeological reconstruction to study the burial rite. Wrapping bodies in hides of wild animals is a widespread long-term tradition, which was practiced in Northern Europe by the Stone Age hunters. This funerary rite was supposed to ensure the “transformation” of a person into an ancestor animal. Reminiscences of hunting religious beliefs were preserved in the ideology of Northern Europe after the transition to agriculture.

During the Bronze Age (ca. 1800-500 BC), cremation became the dominant burial practice in Finland. These Bronze Age (BA) cremations have been excavated from burial cairns in Finland for more than 150 years. This work provides new information on human osteology from this previously understudied area in northeastern Europe. The results cover the earliest period in Finnish prehistory for which large-scale human osteological assemblages are available. In this article, we present a comprehensive human osteological study on the curated bone collections of the National Museum of Finland and other provincial museums, and provide new radiocarbon datings of the cairns. The results show that the cairns were most often used for single burials, without accompanying animals or artefacts. Double and multiple burials became more common in the Iron Age. Porotic hyperostosis was a common finding. Signs of osteoarthritis are more common in the central Satakunta region, where Iron Age burial practices first emerged already during the Late Bronze Age. These observed changes can be connected to changes in subsistence strategies that also enabled permanent settlements.

Supplementary audiovisual material: https://youtu.be/K-EE9YY6pHY https://youtu.be/gRpxQOx9ugs In the Late Mesolithic graves of Yuzhniy Oleniy Ostrov, Northwest Russia, large amounts of Eurasian elk (Alces alces) incisors have been found. These teeth, for the most part fashioned into portable pendants, seem to have formed decorative sets for the garments or accessories of the deceased. This article examines both the technologies associated with these artefacts and their uses, as well as reflecting on the sensorial experiences generated by them. Osteological analysis of a sample of 100 specimens indicates that all types of incisors were used for making the pendants. Traceological analysis indicates that the teeth were modified by scraping, grooving, grinding, and retouching. Traces of wear consist of general wear and distinctive pits or pecks on the perimeters of the crowns. These traces indicate that the pendants were worn before their deposition in the graves, in such a way that they were in contact with both soft and solid materials. The pattern of pits or pecks has until now been unreported in the traceological literature. In experiments, a similar pattern emerged when pendants of fresh elk incisors were hung in rows and bunches and struck against one another. These strokes created a rattling sound. Thus, the elk incisors of Yuzhniy Oleniy Ostrov appear to provide insight into previously unattainable sonic experiences and activities of Mesolithic hunter-gatherers, as well as the early history of the instrument category of rattles.

Burial practices in the large area of the north-eastern European and especially the western Russian forest zone are not as well-known as contemporary sites in, for example, western or southern Europe. To fill this gap in our knowledge of north-western Russian Stone Age hunter-gatherer burial traditions and to better understand the similarities and differences in burial traditions between these areas, a new Russian-Finnish collaboration was launched in late 2016. The main aim of this collaboration is to systematically investigate previously excavated mortuary materials from north-western Russia and study them in the broader context of hunter-gatherer funerary practices in north-eastern Europe. As a first case study, we revisited the burial finds from the Stone Age site of Kubenino, a multi-period site with at least six inhumation burials. Burials have been excavated here since the 1930s, but only three have been published so far. Here we describe all the Kubenino burials and present the results of new analyses, namely the osteological study of bone artefacts, the traceological study of flint artefacts, and new radiocarbon determinations. According to the novel radiocarbon data, the Kubenino burials date to the emergence of the Neolithic, which in this region is marked by the appearance of the first pottery-using hunter-gatherers. As our new analyses show, this transition towards the Neolithic is not only present in the everyday life of these hunter-gatherer groups but also resulted in subtle changes in the funerary realm. These changes include, for example, the intentional breakage of grave goods and the introduction of new artefact forms for animal raw materials that were present already in pre-pottery burials.

In this paper we describe the results of small-scale archaeological fieldwork projects carried out i North Karelia, Finland, in 2003–2014, which ended up having a far greater research impact than anyone could have anticipated. The projects yielded a multitude of new and relevant information, especially on the Early Mesolithic, including the earliest radiocarbon dates for human occupation in Eastern Fennoscandia. Results of this research have been published in a variety of venues. We use this opportunity to present the first summary of these results and to emphasize that even with discontinuous short-term funding it is possible to carry out ambitious and influential research. Over the years, Karelia has been one of the main focus areas of Mika Lavento’s research. We want to honour Mika’s 60th birthday by providing an overview of the results of our studies conducted in the northern parts of this large geographical and administrative area.

The remains of a semi-subterranean house were discovered at the Kharlovka 1-6 site on the Barents Sea coast of the Kola Peninsula, Russia. Despite suffering from erosion, features such as a double stone-boarded hearth and chimney pipe together with diagnostic finds allow us to attribute the house to the so-called Gressbakken type. The artefact assemblage includes stone tools, bone and antler tools and adornments, asbestos-tempered pottery, an amber pendant and a copper tubular bead. Numerous faunal remains are primarily represented by marine species of mammals and birds. Radiocarbon dates on charcoal point to the period 2600–2300 calBC, which is also supported by the artefact types, elevation of the cultural layer and construction type of the house. When accounting for the early dates of Kharlovka 1-6 and several other dwellings, it can be assumed that the Gressbakken house tradition emerged around 2600–2300 calBC. This date is somewhat earlier than previously thought.

In this paper, we introduce the first evidence of the use of human bone for making pendants in Northeast Europe. Twelve of the 37 studied pendants made of long bone splinters turned out to be human bone. Here, we present the ZooMS (Zooarchaeology by Mass Spectrometry) identifications of artefacts and their traceological analysis, and we discuss their implications for the archaeology of Mesolithic burial practices. Our results indicate that the raw material for some of the items was in a fresh or semi-fresh state before making pendants. They were used before they were placed into the graves, and most likely in the same ways as animal bone pendants. This is the first study that has found the use of human bone as raw material in Russian Karelia and the first time that the ZooMS method has been applied to archaeological materials from this region. Together with previous human bone artefact finds from the European Mesolithic period, the bone pendants from Yuzhniy Oleniy Ostrov indicate that the tradition of using human bone as raw material may have been widespread.

Levanluhta, an Iron Age water burial site in Finland, and its material consisting of commingled skeletal remains and artifacts, has been studied by several researchers over the past 100 years, resulting in multiple interpretations of the people and the site. Previous skeletal analyses have concluded that the majority of the individuals represented in the remains were females and children and were of relatively short stature, so possibly nutritionally deprived. This study re-analyzed the commingled adult human remains with updated methods. The methods applied in this study to estimate sex and stature were based on more representative European reference samples than the previously applied methods. The methods included morphology, osteometrics, and computed tomography (CT) scans. Our results indicated that depending on the reference data, the majority of the individual adult bones including os coxae (73%, n = 45) and long bones (humerus 83%-89%, n = 52; radius 72%-89%, n = 47; ulna 50%-65%, n = 58; femur 92%-100%, n = 25; tibia 77%-85%, n = 26) were classified as females based on their size and morphology. The cross-sectional bone properties of humerii, femora, and tibiae visualized using CT scanning also supported these findings. However, the cranial morphology did not show as clear female-biased sex ratio as other methods (42% females, 33% males, 24% undetermined, n = 33). In females, the mean stature based on the tibia (155.3 cm, n = 10) was within the range of the coeval European females and did not necessarily indicate nutritional deprivation, which is in line with previously published stable isotope findings from the site. The mean stature based on the tibia suggested that the Levanluhta males were short (164.0 cm, n = 3), but final interpretations were limited due to the small number of male individuals. The current study affirmed that the Levanluhta skeletal assemblage was female biased and gave new insights into interpretation of the stature.

Yuzhniy Oleniy Ostrov in Karelia, northwest Russia, is one of the largest Early Holocene cemeteries in northern Eurasia, with 177 burials recovered in excavations in the 1930s; originally, more than 400 graves may have been present. A new radiocarbon dating programme, taking into account a correction for freshwater reservoir effects, suggests that the main use of the cemetery spanned only some 100-300 years, centring on ca. 8250 to 8000 cal BP. This coincides remarkably closely with the 8.2 ka cooling event, the most dramatic climatic downturn in the Holocene in the northern hemisphere, inviting an interpretation in terms of human response to a climate-driven environmental change. Rather than suggesting a simple deterministic relationship, we draw on a body of anthropological and archaeological theory to argue that the burial of the dead at this location served to demarcate and negotiate rights of access to a favoured locality with particularly rich and resilient fish and game stocks during a period of regional resource depression. This resulted in increased social stress in human communities that exceeded and subverted the 'normal' commitment of many hunter-gatherers to egalitarianism and widespread resource sharing, and gave rise to greater mortuary complexity. However, this seems to have lasted only for the duration of the climate downturn. Our results have implications for understanding the context of the emergence-and dissolution-of socio-economic inequality and territoriality under conditions of socio-ecological stress.

Hepatitis B virus (HBV) has been infecting humans for millennia and remains a global health problem, but its past diversity and dispersal routes are largely unknown. We generated HBV genomic data from 137 Eurasians and Native Americans dated between ~10,500 and ~400 years ago. We date the most recent common ancestor of all HBV lineages to between ~20,000 and 12,000 years ago, with the virus present in European and South American hunter-gatherers during the early Holocene. After the European Neolithic transition, Mesolithic HBV strains were replaced by a lineage likely disseminated by early farmers that prevailed throughout western Eurasia for ~4000 years, declining around the end of the 2nd millennium BCE. The only remnant of this prehistoric HBV diversity is the rare genotype G, which appears to have reemerged during the HIV pandemic., The research was funded by the Max Planck Society, the European Research Council (ERC) under the European Union’s Horizon 2020 research and innovation program (771234–PALEoRIDER, to W.H.; 805268–CoDisEASe to K. Bos; 834616–ARCHCAUCASUS to S.H.), the Slovak Academy of Sciences and the European Union’s Seventh Framework Programme and Marie Curie Actions under the Programme SASPRO (1340/03/03 to P.C.R.), the ERA.NET RUS Plus–S&T programm of the European Union’s Seventh Framework Programme (277–BIOARCCAUCASUS to S.Re. and S.H.), the Werner Siemens Stiftung (“Paleobiochemistry”, to CW), the Award Praemium Academiae of the Czech Academy of Sciences (to M.E.), the Institute of Archaeology of the Czech Academy of Sciences (RVO 67985912, to M.Dobe.), the Russian Foundation for Basic Research (19-09-00354a, to M.K.K. and V.V.K.; 19-78-10053 to SSh), the German Research Foundation (DFG-HA-5407/4-1–INTERACT to W.H. and RE2688/2 to S.Re.), the French National Research Agency (ANR-17-FRAL-0010–INTERACT, to M.F.D., M.Ri., S.Ro., S.Sai., D.Bi., and P.Le.), the Wenner-Gren Dissertation Fieldwork Grant (9558 to S.Sab.), and the Ministry of Education and Science of the Republic of Kazakhstan (AP08856654 to L.B.D., L.M., and E.Kh. and AP08857177 to A.Z.B.).

More than 4300 Eurasian elk (Alces alces) incisors, most of them pendants, were found in 84 burials in the Late Mesolithic cemetery of Yuzhniy Oleniy Ostrov, Northwest Russia. We analysed the manufacture techniques of elk teeth (4014), in the collection of the Peter the Great Museum of Anthropology and Ethnography, St Petersburg. A striking observation is that the manufacture of these pendants is similar in all burials. Teeth were worked by carving one or several grooves around the root tip. In addition to grooved ones, a number of teeth were not worked at all. The uniformity of the chosen species, tooth and techniques indicates that strict norms prevailed in the pendant industry. Despite the overall similarity, our study shows some variation in making pendants. A groove can cut the whole circumference of the root, or several distinct grooves can mark opposite sides of the root. Sometimes the grooves are deep and made carefully, and sometimes they are weak and made hastily. A typology of various groove types was created. In many graves, one groove type dominates. We interpret that this inter-burial variation and domination of one type resulted from personal choice and taste based on practicality. Such variation could also be associated with kin identifiers, but we did not find clear support for that in our study. Our study indicates that the groove types as such had no connection with particular ornaments, garments or hanging positions.

The use of feathers in ritual costumes and everyday clothing is well described in ethnographic sources throughout the world. From the same sources we know that bird wings and feathers were loaded with meaning in traditional societies worldwide. However, direct archaeological evidence of prehistoric use of feathers is still extremely scarce. Hence, feathers belong to the ‘missing majority’: items that are absent from the archaeological record but which we can assume to have been of importance. Here we present microscopic analysis of soil samples from hunter-gatherer burial contexts which reveal the first direct evidence of the use of feathers in the Mesolithic period of north-eastern Europe.

The focus of this article is an analysis of tubular bone items made by cutting off the epiphyses of large bird ulnae, such as those found in burials 67 and 69 at Late Mesolithic Yuzhniy Oleniy Ostrov (Lake Onega, Northwest Russia). The bone tube from grave 69 was studied systematically, and tested to determine whether it could have been used as a sound instrument. In order to do so, we made a replica of the object, and tested its functions as a flute and a reed pipe. More generally, we propose alternatives to conventional and somewhat unimaginative interpretations of tubular bone artefacts. We raise awareness of their possible acoustic dimension, and discuss the outlook, structure, function and uses of simple bone flutes, whistles and reed pipes, highlighting their most common role as decoy whistles or animal calls. Blown instruments should be kept in mind when looking for explanations of tubular bone artefacts. Sound tools, and especially decoy whistles or animal calls, were common and necessary implements used everywhere and throughout all time periods, especially among hunters and fowlers, and should be considered a basic part of the bone tool assemblage.

Reindeer herding emerged among the indigenous Sámi of Northern Fennoscandia between ca. 800 and 1500 CE. While the details of the reindeer domestication process are still actively debated, it has been hypothesized that the transition to reindeer herding affected Sámi ritual practice, especially animal offerings given at various sacred sites. To explore this hypothesis, we analyzed DNA from reindeer bone samples dating to ca. 1200–1700 CE from Sámi offering sites located in present-day Northern Finland as well as from samples originating from ancient dwelling site in Southern Finland and Kola Peninsula, Russia. The results show that haplotypes related to wild Finnish forest reindeer (Rangifer tarandus fennicus) began to be replaced by haplotypes common in modern domesticated reindeer in the faunal assemblages from offering sites starting between 1400 and 1600 CE. The results suggest that, although the role of reindeer herding in the economy of the Sámi communities varied greatly, the transition to reindeer herding may have affected ritual practices, testifying to a shared way of reciprocating with the land and animals.

Background The Finnhorse was established as a breed more than 110 years ago by combining local Finnish landraces. Since its foundation, the breed has experienced both strong directional selection, especially for size and colour, and severe population bottlenecks that are connected with its initial foundation and subsequent changes in agricultural and forestry practices. Here, we used sequences of the mitochondrial control region and genomic single nucleotide polymorphisms (SNPs) to estimate the genetic diversity and differentiation of the four Finnhorse breeding sections: trotters, pony-sized horses, draught horses and riding horses. Furthermore, we estimated inbreeding and effective population sizes over time to infer the history of this breed. Results We found a high level of mitochondrial genetic variation and identified 16 of the 18 haplogroups described in present-day horses. Interestingly, one of these detected haplogroups was previously reported only in the Przewalski’s horse. Female effective population sizes were in the thousands, but declines were evident at the times when the breed and its breeding sections were founded. By contrast, nuclear variation and effective population sizes were small (approximately 50). Nevertheless, inbreeding in Finnhorses was lower than in many other horse breeds. Based on nuclear SNP data, genetic differentiation among the four breeding sections was strongest between the draught horses and the three other sections (FST = 0.007–0.018), whereas based on mitochondrial DNA data, it was strongest between the trotters and the pony-sized and riding horses (ΦST = 0.054–0.068). Conclusions The existence of a Przewalski’s horse haplogroup in the Finnhorse provides new insights into the domestication of the horse, and this finding supports previous suggestions of a close relationship between the Finnhorse and eastern primitive breeds. The high level of mitochondrial DNA variation in the Finnhorse supports its domestication from a large number of mares but also reflects that its founding depended on many local landraces. Although inbreeding in Finnhorses was lower than in many other horse breeds, the small nuclear effective population sizes of each of its breeding sections can be considered as a warning sign, which warrants changes in breeding practices. Electronic supplementary material The online version of this article (10.1186/s12711-019-0480-8) contains supplementary material, which is available to authorized users.

The wetland find in Levänluhta (western Finland) consists of unburnt, mixed up remains from almost 100 human individuals along with artefacts and animal bones. This spring site, a small lake at the time of use (ad300–800), has been investigated archaeologically from the late nineteenth century onwards. An impressive array of finds, including precious artefacts, is on display at the National Museum of Finland. However, the material has not previously been subjected to systematic research to clarify who these people were, and why they were buried in a small lake at a time when cremation was the prevailing burial tradition. Here we present the results of a multidisciplinary study that includes new analyses and interpretations of the finds and the site. Prestigious artefacts, peripheral location, and the fact that only a few males were found suggest this unusual burial site was a cemetery for socially or ideologically deviant members of the society.

Tooth pendants of European elk, Eurasian beaver and brown bear are the most common artefact type in graves at Late Mesolithic Yuzhniy Oleniy Ostrov on Lake Onega, northwestern Russia. In one burial...

Northeastern Siberia has been inhabited by humans for more than 40,000 years but its deep population history remains poorly understood. Here we investigate the late Pleistocene population history of northeastern Siberia through analyses of 34 newly recovered ancient genomes that date to between 31,000 and 600 years ago. We document complex population dynamics during this period, including at least three major migration events: an initial peopling by a previously unknown Palaeolithic population of ‘Ancient North Siberians’ who are distantly related to early West Eurasian hunter-gatherers; the arrival of East Asian-related peoples, which gave rise to ‘Ancient Palaeo-Siberians’ who are closely related to contemporary communities from far-northeastern Siberia (such as the Koryaks), as well as Native Americans; and a Holocene migration of other East Asian-related peoples, who we name ‘Neo-Siberians’, and from whom many contemporary Siberians are descended. Each of these population expansions largely replaced the earlier inhabitants, and ultimately generated the mosaic genetic make-up of contemporary peoples who inhabit a vast area across northern Eurasia and the Americas. Analyses of 34 ancient genomes from northeastern Siberia, dating to between 31,000 and 600 years ago, reveal at least three major migration events in the late Pleistocene population history of the region.

Publication of the rich medieval and early modern bone material excavated in Finnish Lapland over the past decades is long due. Bone assemblages can complete the picture of the life and livelihood of the early Saami societies presented in written sources. The data from three Saami dwelling sites and two market places, which are published here, support in many ways the written information, but show also some discrepancies between these two sources of information. The osteological data may raise more questions than they answer, but serve as an important source material for further research.

Paleodietary research is a complex field, which requires large sets of background information. Owing to increasing interest and activity in the field, a substantial amount of archaeological isotope baseline data exist for Northern Europe, consisting mainly of animal bone collagen δ13C, δ15N, and δ34S values. However, the data are scattered into dozens of publications written in multiple languages and less-accessible formats, making the data laborious to use. This article presents the first compilation work of this data, the open access dIANA database (Dietary Isotopic baseline for the Ancient North; https://www.oasisnorth.org/diana.html), aimed to support (paleo)dietary research in the Baltic Sea area. The database work is complemented with new analyses of archaeological and (pre-)modern domestic and wild fauna from Finland and Russia broadening the selection of analysed species in the database. We present and discuss data examples, which on one hand show existing spatiotemporal isotope patterns related to diet and differences in the environmental carbon sources and on the other, also visualize the current status of baseline research and the need for further analyses in the circum-Baltic area. Paleodietary research is a complex field, which requires large sets of background information. Owing to increasing interest and activity in the field, a substantial amount of archaeological isotope baseline data exist for Northern Europe, consisting mainly of animal bone collagen δ13C, δ15N, and δ34S values. However, the data are scattered into dozens of publications written in multiple languages and less-accessible formats, making the data laborious to use. This article presents the first compilation work of this data, the open access dIANA database (Dietary Isotopic baseline for the Ancient North; https://www.oasisnorth.org/diana.html), aimed to support (paleo)dietary research in the Baltic Sea area. The database work is complemented with new analyses of archaeological and (pre-)modern domestic and wild fauna from Finland and Russia broadening the selection of analysed species in the database. We present and discuss data examples, which on one hand show existing spatiotemporal isotope patterns related to diet and differences in the environmental carbon sources and on the other, also visualize the current status of baseline research and the need for further analyses in the circum-Baltic area.

Development of dental abnormalities due to improper occlusal wear is common among modern domestic horses. This phenomenon often is attributed to jaw conformation. Rostral mandibular hooks may develop in horses with underjet or mandibular prognathism, a condition where the lower jaw protrudes forward, beyond the upper jaw. Less abrasive diet, free of phytoliths and matrix-like plant fibers, also may promote enamel and focal overgrowths of equine dentition. Here we report a rostral mandibular hook in a lower premolar tooth of a medieval horse, found in a spring deposit in Levanluhta, Osthrobothnia, Finland. To our knowledge, this is the first such report from a medieval horse. (C) 2017 Elsevier Inc. All rights reserved.

Levanluhta is a unique archaeological site with the remains of nearly a hundred Iron Age individuals found from a water burial in Ostrobothnia, Finland. The strongest climatic downturn of the Common Era, resembling the great Fimbulvinter in Norse mythology, hit these people during the 6th century AD. This study establishes chronological, dietary, and livelihood synthesis on this population based on stable carbon and nitrogen isotopic and radiocarbon analyses on human remains, supported by multidisciplinary evidence. Extraordinarily broad stable isotopic distribution is observed, indicating three subgroups with distinct dietary habits spanning four centuries. This emphasizes the versatile livelihoods practiced at this boundary of marine, freshwater, and terrestrial ecosystems. While the impact of the prolonged cold darkness of the 6th century was devastating for European communities relying on cultivation, the broad range of livelihoods provided resilience for the Levanluhta people to overcome the abrupt climatic decline.

Far northeastern Siberia has been occupied by humans for more than 40 thousand years. Yet, owing to a scarcity of early archaeological sites and human remains, its population history and relationship to ancient and modern populations across Eurasia and the Americas are poorly understood. Here, we analyze 34 ancient genome sequences, including two from fragmented milk teeth found at the ~31.6 thousand-year-old (kya) Yana RHS site, the earliest and northernmost Pleistocene human remains found. These genomes reveal complex patterns of past population admixture and replacement events throughout northeastern Siberia, with evidence for at least three large-scale human migrations into the region. The first inhabitants, a previously unknown population of “Ancient North Siberians” (ANS), represented by Yana RHS, diverged ~38 kya from Western Eurasians, soon after the latter split from East Asians. Between 20 and 11 kya, the ANS population was largely replaced by peoples with ancestry related to present-day East Asians, giving rise to ancestral Native Americans and “Ancient Paleosiberians” (AP), represented by a 9.8 kya skeleton from Kolyma River. AP are closely related to the Siberian ancestors of Native Americans, and ancestral to contemporary communities such as Koryaks and Itelmen. Paleoclimatic modelling shows evidence for a refuge during the last glacial maximum (LGM) in southeastern Beringia, suggesting Beringia as a possible location for the admixture forming both ancestral Native Americans and AP. Between 11 and 4 kya, AP were in turn largely replaced by another group of peoples with ancestry from East Asia, the “Neosiberians” from which many contemporary Siberians derive. We detect gene flow events in both directions across the Bering Strait during this time, influencing the genetic composition of Inuit, as well as Na Dene-speaking Northern Native Americans, whose Siberian-related ancestry components is closely related to AP. Our analyses reveal that the population history of northeastern Siberia was highly dynamic throughout the Late Pleistocene and Holocene. The pattern observed in northeastern Siberia, with earlier, once widespread populations being replaced by distinct peoples, seems to have taken place across northern Eurasia, as far west as Scandinavia.

The effects of bio-adaptation-based visual cues in conveying affective information was studied while conducting social meditation in a virtual reality (VR) environment. In a laboratory study, 22 dyads practiced a short duration, modified version of empathy-evoking compassion meditation. The immersive VR provided electroencephalography (EEG) -based real-time visual feedback, as changing color, on the user's brain activation; the breathing rate of the users was visualized as a movement cue. In addition, the synchronization of the feedback signals between the users was also visualized. Initial results suggest that the perceived affective interdependence, a component of social presence, was increased when both of the bio-adaptive visual feedbacks were presented to the users, compared to when there was no visual feedback. Also, the EEG-based visual feedback led to higher affective interdependence than the respiration based visual feedback. The findings suggest that affective contagion may occur by mediated adaptive cues that are based on physiological signals.

New radiocarbon dates from bones of the wild boar ( Sus scrofa) found in archaeological contexts in Finland indicate the presence of the species in Southern Finland c. 9000–8000 cal. BP, that is, before the Holocene thermal optimum. However, the number of wild boar was never large enough to support a permanent population. Reasons for the scarcity of wild boars even during the warmer periods may have been the thick snow cover, the growing human population near the distribution route, as well as hunting.

Recent archaeological studies reveal the importance of birds in prehistoric North-European hunter-fisher-gatherer burial practices. In this article I describe two examples of bird species at prehistoric hunter-gatherer burials: the Eurasian jay (Garrulus glandarius) at the Middle Neolithic Zvejnieki site in northern Latvia, and the osprey (Pandion haliaetus) at the Late Mesolithic Yuzhniy Oleniy Ostrov site in western Russia. I suggest that the bone finds and their archaeological contexts indicate a prehistoric ideology that can be interpreted as representing totemism and shamanism. The wing bones had a specific function and meaning, probably connected to protection, transformation or transport. The deposition of osprey legs may indicate that the power of this bird was particularly appreciated and re-mobilized in the burial.

The article by Overton and Hamilakis challenges so-called traditional zooarchaeology and works as a manifesto for a new social zooarchaeology, as the authors call it. This new social zooarchaeology moves beyond the thinking of animals as (purely) resources and instead reinstates their position as sentient and autonomous agents. The approach is fresh and evidence-based (e.g. Robb 2010). The sites and bone materials used as examples come from Late Mesolithic Denmark: the Ertebølle site Aggersund in Jutland and Ertebølle Grave 8 at Vedbæk, Sjælland. Both sites were excavated and analysed many years ago, but the bone material has been reanalysed and interpreted for this study.

Animal bones in human burials may reveal aspects of the relationship between animals and humans. This article describes the roles of birds in mortuary practices and in the ideology of Stone Age northern Europe. Bird bones from two large burial sites, Middle Neolithic Ajvide (Gotland, Sweden) and Mesolithic and Neolithic Zvejnieki (Latvia) are investigated with osteological methods. Beads and pendants were fashioned from the wing bones of waterbirds, and used in the decoration of the body or the burial dress. The jay was found in three Neolithic burials at Zvejnieki, and it may have been a totem animal for the Middle Neolithic people at Zvejnieki, and its wings or feathers were presumably attached to dresses and costumes for the dead. Bird remains in burials at Ajvide, Zvejnieki and some other Stone Age cemeteries may indicate similar features in the way of perceiving birds, especially the possible symbolic roles of waterbirds and wings. The findings are discussed from the perspective of the cosmology of historical hunter–gatherer (and herding) groups in modern Russia.

We have conducted a detailed taphonomic study of the avifauna of the Pitted Ware culture site of Ajvide on the Island of Gotland in the Baltic Sea, in order to investigate the fowling patterns and the taphonomic history of the bird remains. We have investigated cultural as well as natural modifications on the bird bones, fragmentation and fracture patterns, and performed a systematic surface modification study. No specific area for the deposition of bird remains or specific bird species was identified. All major anatomical parts of birds are present in the assemblage, but there is a dominance of specimens from the wing elements. Traces of cultural modification were observed on the bones, including cut marks, burning, modification (implements, beads, raw material), and gnawing marks. The bone surface modifications and fracture analysis indicate that the majority of the bird bones at Ajvide did not lie on the soil surface for an extended period of time before being deposited in the soil. Dry fractures increase while fresh fractures decrease towards the upper levels of the stratigraphy, indicating more extensive post-depositional destruction. This may partly be connected to modern agriculture, but also to later use of the settlement area as a burial ground. The Ajvide assemblage contains a variety of birds living in different biotopes. However, bird hunting was mainly focused on sea birds. Auks and ducks are the most common families in the assemblage. We find it likely that the Ajvide hunters conducted organised hunting expeditions to two nearby islands for the hunting of auks, while it was possible to hunt other birds such as ducks closer to the site. The presence of medullary bone and bones from subadult birds indicates a main hunting season in late spring and early summer. However, comparisons with modern migration patterns indicate that hunting may have occurred throughout the year. Of special palaeozoological interest is the find of gannet (Morus bassanus), which apparently in Neolithic times visited the Baltic area more regularly than today. Copyright © 2006 John Wiley & Sons, Ltd.

Tubular bone artefacts of different size and form are commonly found in the Middle Neolithic inhumation burials in Gotland and Öland, Sweden. These burials were made by the people of the so-called Scandinavian Pitted Ware Culture. Small tubular artefacts have commonly been interpreted as beads used in decoration as they, in many cases, appear in clusters at the head region and along the body. In 1998 at Ajvide, archaeologists discovered a grave (62), which contained a large number of grave goods, among them tubular bone artefacts of an extraordinary character. Based on their outlook, these single or two-pieced artefacts with or without pierced holes were interpreted as flutes. Their suitability for sound production, however, has never been studied systematically. In this article, we will discuss the presence and function of the different tubular bone artefacts found in grave 62. We will describe the finds, sort them tentatively and report possible ethnographical parallels. The article is intended as an introduction to the research project, which seeks to analyze the organological structures of the artefacts and reassess their interpretation as musical instruments. The find contexts of the artefacts, as well as, the grave entity with all other artefacts, will be studied from the perspective of music archaeology.

Historical sources, such as tax rolls and accounts, can provide information about mediaeval fishing and fish trade, but this subject can also be investigated through archaeological methods. Archaeological research on the mediaeval and early historical fishing in Finland has not been undertaken in any detail. Research from neighbouring areas, mainly Sweden and Estonia, has provided information about mediaeval fishing in northern Europe. This paper presents the results of a osteological examination of a sample of archaeological fish bones excavated in the Old Town of Helsinki in 1993. The sample derives from the remains of a cellar in a house, used in late-sixteenth or early-seventeenth century by a wealthy person, probably a merchant.