582 results on '"Kovařík, František"'
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2. Additions to the distribution of Sudanese scorpions
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Siyam, Manal, Dunlop, Jason A., Kovařík, František, Mohammad, Abubakr, and Pensoft Publishers
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Distribution ,East Africa ,Identification ,Public health ,Scorpiones - Published
- 2023
3. The evolutionary history of the relict scorpion family Iuridae of the eastern Mediterranean
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Parmakelis, Aristeidis, Dimitriadou, Domniki, Gkigkiza, Evdokia, Karampatsou, Lydia, Stathi, Iasmi, Fet, Victor, Yağmur, Ersen A., and Kovařík, František
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- 2022
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4. Scorpiops tangae sp. n. (Scorpiones: Scorpiopidae) from Laos.
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Kovařík, František, Šťáhlavský, František, and Stockmann, Mark
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A new species Scorpiops tangae sp. n. is described from Laos, fully illustrated with color photos showing its morphology and habitus. Scorpiops tangae sp. n. is the only species of the genus with a combination of the following five characters: chelal trichobothrium Eb3 located in distal half of manus between trichobothria Dt and Est; patella of pedipalp with 11 ventral and 17 external trichobothria; fingers of pedipalps undulate in male; pedipalp movable finger with ca. 45 IAD; and chela length to width ratio 3.7 in male. In addition to morphology and hemispermatophore, we also present the information about the karyotype of S. tangae sp. n. (2n=105). [ABSTRACT FROM AUTHOR]
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- 2024
5. Uroplectes ebogo sp. n. (Scorpiones: Buthidae) from Cameroon.
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Kovařík, František, Šťáhlavský, František, and Govorov, Valeriy
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SCORPIONS , *MORPHOLOGY , *PUNCTUATION , *SPECIES , *PHOTOGRAPHS - Abstract
A new species Uroplectes ebogo sp. n. is described from Cameroon, fully illustrated with color photos showing its morphology and habitus. Uroplectes ebogo sp. n. is the only species of the genus with a combination two characters: metasoma without carinae and metasoma V and telson without punctuation. In addition to morphology and hemispermatophore, we also present the information about the karyotype of Uroplectes ebogo sp. n. (2n=24). [ABSTRACT FROM AUTHOR]
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- 2024
6. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXXIII. Three new species of Gint from Ethiopia and Somaliland (Buthidae).
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Kovařík, František, Abdirahman Elmi, Hassan Sh, and Šťáhlavský, František
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BIOLOGICAL classification , *KARYOTYPES , *ARACHNIDA , *SPECIES , *MORPHOLOGY - Abstract
Three new species, Gint sahil sp. n. from Somaliland and Gint abshiri sp. n. and G. derbiae sp. n. from Ethiopia are described and compared with other species of the genus. Additional information is provided on the taxonomy and distribution of the genus Gint, fully complemented with color photos of specimens of the new species, as well as of their habitats. Furthermore, alongside the analyses of external morphology and hemispermatophores, we have provided description of the karyotype of G. abshiri sp. n. This species exhibits karyotype with 2n=27. Included is distribution map and a key for the genus Gint. [ABSTRACT FROM AUTHOR]
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- 2024
7. New records of Mesobuthus mesopotamicus (Penther, 1912) in Iraq and Mesobuthus faiki sp. n. from Turkey (Scorpiones: Buthidae).
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Yağmur, Ersen Aydın, Kovařík, František, Fet, Victor, Hussen, Fenik Sherzad, Kurt, Ridvan, Al-Khazali, Azhar Mohammed, Kachel, Hamid Saeid, and Al-Fanharawi, Ali Abdulhamza
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SCORPIONS , *SPECIES - Abstract
New locality records for Mesobuthus mesopotamicus (Penther, 1912) are given from Iraq. The populations from southeastern Turkey, earlier identified as M. mesopotamicus by Kovařík et al. (2022) are examined and compared with the specimens from type locality of M. mesopotamicus. These populations are described as a new species, Mesobuthus faiki sp. n. Detailed illustrations of both species and a map are given. [ABSTRACT FROM AUTHOR]
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- 2024
8. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXXII. Barbaracurus hofereki sp. n. from Djibouti.
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Kovařík, František
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ARACHNIDA , *SCORPIONS , *TEETH , *SPECIES , *FEMALES - Abstract
Barbaracurus hofereki sp. n. from Djibouti is described and compared with other species of the genus. B. hofereki sp. n. is the only species of the genus with pedipalp movable finger with 8 rows of granules and pectinal tooth count 27-28 in female. Also, its pedipalp chela with very narrow manus (chela length/width ratio 6.07 in female), narrower than in all other African species of the genus. A map of the distribution of the genus in the Horn of Africa and the Arabian Peninsula is provided. [ABSTRACT FROM AUTHOR]
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- 2024
9. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXXI. Two new genera from Somaliland: Sanaag gen. n. and Sahil gen. n. (Buthidae).
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Kovařík, František
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ARACHNIDA , *SCORPIONS , *SPECIES - Abstract
Two new monotypic buthid genera are described: Sanaag gen. n. and Sahil gen. n. from Somaliland, both belonging to the 'Buthus' group. Sanaag gen. n. (type species Gint maidensis Kovařík et al., 2018) differs from Gint Kovařík et al., 2013 in the structure of its hemispermatophore, which has a large, tall, subtriangular hook-like basal lobe, and in the shape of its telson which is rather bulbous. Sahil gen. n. (type species Sahil elmii sp. n.) differs from all morphologically similar small-sized genera of the Horn of Africa in having the ventral aspect of cheliceral fixed finger with two denticles, and the movable finger of the pedipalp with 7 subrows of denticles. [ABSTRACT FROM AUTHOR]
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- 2024
10. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXX. Parabuthus (Buthidae) (Part III), with description of three new species from Somaliland and occurrence of Parabuthus eritreaensis Kovařík, 2003.
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Kovařík, František, Lowe, Graeme, Elmi, Hassan Sh Abdirahman, and Šťáhlavský, František
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ARACHNIDA , *SPECIES distribution , *KARYOTYPES , *CHROMOSOMES , *SPECIES , *SCORPIONS - Abstract
A new record of an adult female of Parabuthus eritreaensis Kovařík, 2003 in Somaliland confirms true distribution of this species, already discussed in Kovařík et al. (2016: 19-21). Three new species are described from Somaliland, P. dorisae sp. n., P. evae sp. n., and P. quincyae sp. n.. The hemispermatophore of P. dorisae sp. n. is illustrated and described. In addition to the analyses of external morphology and hemispermatophore, we have provided descriptions of the karyotypes of P. dorisae sp. n. and P. quincyae sp. n. Despite the presence of multivalents (CVIII and CXIV), both species exhibit karyotypes with 2n=16 and chromosomes that gradually decrease in length, with the exception of the first chromosome, which is longer than the following chromosomes. A map of distribution of Parabuthus species in the Horn of Africa is included. [ABSTRACT FROM AUTHOR]
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- 2024
11. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae)
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Kovařík, František and BioStor
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- 2003
12. Pandinus (Pandinops) pococki sp. n. from Somalia, and Pandinus pugilator, a junior synonym of Pandinus (Pandinops) bellicosus comb. n. (Scorpiones, Scorpionidae)
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Kovařík, František and BioStor
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- 2000
13. Revision of family Chaerilidae (Scorpiones), with descriptions of three new species
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Kovařík, František and BioStor
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- 2000
14. In Memoriam Richard F. Ayrey (1948 -- 2024).
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DeBoer-Ayrey, Melinda, Myers, Brandon T., Bryson Jr., Robert W., Webber, Michael M., Graham, Matthew R., Lowe, Graeme, Fet, Victor, and Kovařík, František
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PSEUDOSCORPIONS - Abstract
This document is a memorial issue honoring Richard F. Ayrey, a US scorpiologist who passed away in 2024. The article provides a brief biography of Ayrey, highlighting his passion for nature and scorpions. It also mentions his contributions to the field, including the description of two new genera and 19 species of scorpions. Ayrey was well-respected in the scientific community and his work was published in various journals. The article includes personal anecdotes from colleagues and friends who remember Ayrey fondly. The text is a collection of personal reflections and anecdotes about a scorpion taxonomist named Rich Ayrey. The authors express their admiration for Rich's passion, enthusiasm, and dedication to studying scorpions. They recount memorable experiences they had with Rich, including discovering new scorpion species and going on field trips together. The authors also mention the scorpion taxa that Rich described and named after himself. Overall, the text pays tribute to Rich's contributions to the field of scorpion taxonomy and his impact on those who knew him. This document appears to be a collection of figures and references related to the work of Rich Ayrey, a researcher in the field of scorpion biology. The figures include maps and photographs of scorpion species and their type localities. The references list various publications by Ayrey, describing new species of scorpions and contributing to the field of scorpion biology. The document also includes personal reflections from individuals who knew Ayrey, highlighting his dedication [Extracted from the article]
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- 2024
15. Revision of the genus Olivierus in Xinjiang, China, with comments on Mesobuthus thersites (Scorpiones: Buthidae).
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Tang, Victoria, Zhenbang Liu, Graham, Matthew R., Fet, Victor, Kovařík, František, and Šťáhlavský, František
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SCORPIONS ,NUMBERS of species ,DIFFERENTIAL diagnosis - Abstract
The genus Olivierus Farzanpay, 1987 in Xinjiang Uygur Autonomous Region, China, is revised based on recently collected topotypes and other populations from 12 localities. Brief differential diagnoses are provided, with colored illustrations and photos in vivo habitus, emphasizing the key characters. Chinese appellations, conservation status, and documentation of behavior and post-envenomation symptoms are also included. Only two species are now recognized for this genus in Xinjiang: O. longichelus (Sun & Zhu, 2010) and O. przewalskii (Birula, 1897), based on both morphological and molecular evidence. The two species exhibit extensive distribution in Xinjiang (China) while also occurring in adjacent countries. Three new synonyms are proposed: Mesobuthus bolensis Sun et al., 2010 = Olivierus longichelus (Sun & Zhu, 2010), syn. n.; Mesobuthus karshius Sun & Sun, 2011 = Olivierus longichelus (Sun & Zhu, 2010), syn. n.; Olivierus tarabaevi Fet et al., 2021 = Olivierus longichelus (Sun & Zhu, 2010), syn. n. Two species, Olivierus extremus (Werner, 1936) and O. hainanensis (Birula, 1904), are likely synonymous with O. martensii (Karsch, 1879). Should future examination confirm this assumption, the total number of species in genus Olivierus would be reduced to 16. [ABSTRACT FROM AUTHOR]
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- 2024
16. Butheoloides cimrmani sp. n. from Ghana (Scorpiones: Buthidae)
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Kovařík, František and BioStor
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- 2003
17. The first record of Orthochirus glabrifrons (Kraepelin, 1903) (Scorpiones: Buthidae) from the United Arab Emirates.
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Kovařík, František, Yağmur, Ersen Aydın, Ullrich, Alexander, and Buzás, Balázs
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SCORPIONS , *SOUND recordings - Abstract
Orthochirus glabrifrons (Kraepelin, 1903) (Scorpiones: Buthidae) was described from Oman (Muscat). Here, we summarize known localities from Oman as well as records from the United Arab Emirates, which is the new country record for this species. Illustrations of morphology of both sexes are given together with a map of distribution. A lectotype of Orthochirus glabrifrons (Kraepelin, 1903) is designated. Paraorthochirus kaspareki Lourenço & Huber, 2000 and Paraorthochirus kinzelbachi Lourenço & Huber, 2000 are synonymized with Orthochirus glabrifrons (Kraepelin, 1903), syn. n. [ABSTRACT FROM AUTHOR]
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- 2023
18. Figure 2 from: Siyam M, Dunlop JA, Kovařík F, Mohammad A (2023) Additions to the distribution of Sudanese scorpions. Zoosystematics and Evolution 99(1): 45-53. https://doi.org/10.3897/zse.99.90875
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Siyam, Manal, primary, Dunlop, Jason A., additional, Kovařík, František, additional, and Mohammad, Abubakr, additional
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- 2023
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19. Figure 3 from: Siyam M, Dunlop JA, Kovařík F, Mohammad A (2023) Additions to the distribution of Sudanese scorpions. Zoosystematics and Evolution 99(1): 45-53. https://doi.org/10.3897/zse.99.90875
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Siyam, Manal, primary, Dunlop, Jason A., additional, Kovařík, František, additional, and Mohammad, Abubakr, additional
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- 2023
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20. Additions to the distribution of Sudanese scorpions
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Siyam, Manal, primary, Dunlop, Jason A., additional, Kovařík, František, additional, and Mohammad, Abubakr, additional
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- 2023
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21. Figure 1 from: Siyam M, Dunlop JA, Kovařík F, Mohammad A (2023) Additions to the distribution of Sudanese scorpions. Zoosystematics and Evolution 99(1): 45-53. https://doi.org/10.3897/zse.99.90875
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Siyam, Manal, primary, Dunlop, Jason A., additional, Kovařík, František, additional, and Mohammad, Abubakr, additional
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- 2023
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22. Lychas jakli sp n (Scorpiones: Buthidae) from Indonesia
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Kovařík, FrantIšek
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Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kovařík, FrantIšek (2023): Lychas jakli sp n (Scorpiones: Buthidae) from Indonesia. Euscorpius 367: 1-8, {"references":["FET, V. & G. LOWE. 2000. Family Buthidae C. L. Koch, 1837. Pp. 54-286 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder. Catalog of the Scorpions of the World (1758-1998). New York: The New York Entomological Society, 689 pp.","KOCH, C. L. 1845. Die Arachniden. Nurnberg: C. H. Zech'sche Buchhandlung, 11, 174 pp.","KOVARIK, F. 2009. Illustrated Catalog of Scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp.","KOVARIK, F. 2019. TaXonomic reassessment of the genera Lychas, Mesobuthus, and Olivierus, with descriptions of four new genera (Scorpiones: Buthidae). Euscorpius, 288: 1-27.","KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated Catalog of Scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas and Sassanidotus. Prague: Clairon Production, 400 pp.","LOURENCO, W. R. 2020. A new species of Lychas C. L. Koch, 1845 from Ko Tao Island, Thailand (Scorpiones: Buthidae). Revista Iberica de Aracnologia, 36: 59-64.","SOLEGLAD, M. E. & W. D. SISSOM 2001. Phylogeny of the family Euscorpiidae Laurie, 1896 (Scorpiones): a major revision. 25-111 in: Scorpions 2001 In Memoriam Gary A. Polis. Editorss Fet & Selden. British Arachnological Society, 2001, 404 pp.","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81(12): 297-316.","STUNDLOVA, J., F. STAHLAVSKY, V. OPATOVA, J. STUNDL, F. KOVARIK, P. DOLEJS & J. SMID. 2022. Molecular data do not support the traditional morphologybased groupings in the scorpion family Buthidae (Arachnida: Scorpiones). Molecular Phylogenetics and Evolution, 173(2022) 107511: 1-5 and Supplementary Information. https://www.sciencedirect.com/science/ article/pii/S1055790322001245","TIKADER, B. K. & D. B. BASTAWADE. 1983. Scorpions (Scorpionida: Arachnida). In The Fauna of India, Vol. 3. (Edited by the Director). Calcutta: Zoological Survey of India, 671pp.","VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaXie en Arachnologie, Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum National d'Histoire Naturelle Paris, 140: 857-958."]}
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- 2023
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23. Barbaracurus kabateki Kovařík & Lowe & Šťáhlavský & Just 2022, sp. n
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Kovařík, František, Lowe, Graeme, Šťáhlavský, František, and Just, Pavel
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Barbaracurus ,Arthropoda ,Barbaracurus kabateki ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Barbaracurus kabateki sp. n. (Figures 1–33, 101–103, 107, Table 1)
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- 2022
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24. Barbaracurus KovarIk, Lowe et Stahlavsky 2018
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Kovařík, František, Lowe, Graeme, Šťáhlavský, František, and Hurre, Ali Abdi
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Barbaracurus ,Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Key to species of Barbaracurus 1. Pedipalp movable finger without an external accessory granule midway along most proximal granule row......... 2 – Pedipalp movable finger with an external accessory granule midway along most proximal granule row...................... 3 2. Base color uniformly yellow or orange, without any darker markings; sternite VII with very weak carination.......................................................... B. prudenti (Lourenço, 2013) – Base color yellow with brown spots on carapace, dark stripes on tergites, dark pedipalp patella and metasoma V; sternite VII with 4 well developed carinae.......................................................................... B. ugartei ( KovařÍk, 2000) 3. Pedipalp movable finger with 6 rows of granules........... 4 – Pedipalp movable finger with 7 rows of granules........... 7 4. Pedipalp chela with narrower manus, chela length/ width ratio 4.3–5.6, finger margins weakly undulate at base, not leaving gap when closed (figs. 81, 84, 86 in KovařÍk et al., 2015)............................................................................... 5 – Pedipalp chela with broader manus, chela length/ width ratio 3.4–4.2, finger margins strongly undulate at base, leaving wide gap when closed.......................................... 6 5. Total length 22.5 mm in male, 32.1–32.25 mm in females; pectines with 16–17 teeth in both sexes; distributed in Ethiopia and Somalia..... B. subpunctatus ( Borelli, 1925) – Total length 31.25 mm in male, 38.80 mm in female; pectines with 25–27 teeth in both sexes; distributed in Somaliland.............................. B. feti KovařÍk et al., 2019 6. Telson vesicle pyriform in lateral profile, deeper anteriorly (figs. 83–84 in KovařÍk et al., 2018b); telson length/ depth ratio 2.75–2.89; pedipalp movable finger of female very weakly undulate at base; hemispermatophore basal lobe a weak carina (figs. 27, 35 in KovařÍk et al., 2018b)............................................................. B. somalicus (Hirst, 1907) – Telson vesicle symmetric in lateral profile (figs. 76–77 in KovařÍk et al., 2015); telson length/ depth ratio 2.60–2.73; pedipalp movable finger of female moderately undulate at base; hemispermatophore basal lobe a well developed scoop-like lamina (figs. 26, 34 in KovařÍk et al., 2018b)................................. B. sofomarensis ( KovařÍk et al., 2015) 7. Pedipalp chela with broader manus (Figs 53–55, and figs. 58, 60 in KovařÍk et al., 2018b), chela length/ width ratio 4.28–5.43 (♀), 3.2–3.42 (♂); proximal margins of pedipalp fingers of male strongly undulate, leaving gap with fingers closed (Fig. 54, and fig. 59 in KovařÍk et al., 2018b)...... 8 – Pedipalp chela with narrower manus (Figs. 7–9, figs. 62, 64, 66, 68, 210, 212 in KovařÍk et al., 2018b), chela length/ width ratio 4.07–6.12 (♀, ♂); proximal margins of pedipalp fingers of male weakly undulate, not leaving gap with fingers closed (Fig. 8, and figs. 62–67 in KovařÍk et al., 2018b)........................................................................ 9 8. Pectines with 17–19 teeth (♀, ♂); telson more bulbous, length/ depth ratio 2.27–2.37 (♀, ♂); distributed in Africa (Eritrea)............................. B. zambonellii (Borelli, 1902) – Pectines with 22–25 (♂) 19–23 (♀) teeth; telson less bulbous, length/ depth ratio 2.48–2.70 (♀, ♂); distributed in Arabian Peninsula............................................................................................... B. yemenensis KovařÍk et al., 2018 9. Metasoma V length/ width ratio 2.71 (♂); pedipalp chela length/ width ratio 4.07 (♂); hemispermatophore basal lobe obsolete, reduced to a weak ridge (Figs. 102–103); found in Saudi Arabia......................................... B. kabateki sp. n. – Metasoma V length/ width ratio 2.40–2.47 (♂); pedipalp chela length/ width ratio 4.24–4.70 (♂); hemispermatophore basal lobe a well developed scoop-like lamina (figs. 24– 25, 32–33 in KovařÍk et al., 2018b); found in Oman. – Telson less slender (figs. 85–87 in KovařÍk et al., 2018b), length/ depth ratio 2.70–2.72 (♂, ♀); not found in northern Oman....................... B. winklerorum KovařÍk et al., 2018, Published as part of Kovařík, František, Lowe, Graeme, Šťáhlavský, František & Just, Pavel, 2022, The genus Barbaracurus in Saudi Arabia (Scorpiones: Buthidae), with description of a new species., pp. 1-26 in Euscorpius 365 on page 23, DOI: 10.5281/zenodo.7616748, {"references":["KOVARIK, F., G. LOWE, M. SEITER, J. PLISKOVA & F. STAHLAVSKY. 2015. Scorpions of Ethiopia (Arachnida: Scorpiones), Part II. Genus Babycurus Karsch, 1886 (Buthidae), with description of two new species. Euscorpius, 196: 1 - 31.","KOVARIK, F., G. LOWE, F. STAHLAVSKY & A. A. HURRE. 2019. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XX. Barbaracurus feti sp. n. from Somaliland (Buthidae). Euscorpius, 280: 1 - 11.","KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2018 b. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41."]}
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- 2022
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25. Orthochirus katerinae sp. n. (Scorpiones: Buthidae) from Saudi Arabia
- Author
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Kovařík, FrantIšek and Just, Pavel
- Subjects
Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kovařík, FrantIšek, Just, Pavel (2022): Orthochirus katerinae sp. n. (Scorpiones: Buthidae) from Saudi Arabia. Euscorpius (Oxford, England) 362 (1): 1-9, DOI: http://doi.org/10.5281/zenodo.7399385, URL: http://dx.doi.org/10.5281/zenodo.7399384, {"references":["ALQAHTANI, A. R. & A. BADRY. 2021. A contribution to the scorpion fauna of Saudi Arabia, with an identification key (Arachnida: Scorpiones). Journal of King Saud University-Science, 33: 1-13.","ALQAHTANI, A. R., B. ELGAMMAL, K. I. GHALEB & A. BADRY. 2019. The scorpion fauna of the southwestern part of Saudi Arabia. Egyptian Academic Journal of Biological Sciences Zoology, 11(1): 19-29.","KARSCH, F. 1892.Arachniden von Ceylon und von Minikoy, gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift, 36(1891): 267-310.","KOVARIK, F. 2009. Illustrated Catalog of Scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp.","KOVARIK, F., V. FET, & E. A. YAGMUR. 2020. Further review of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) fromAsia: taXonomic position of O. melanurus, O. persa, O. scrobiculosus, and description of siX new species. Euscorpius, 318: 1-73.","KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated Catalog of Scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas and Sassanidotus. Prague: Clairon Production, 400 pp.","LOURENCO, W. R. 2007. Litter size in micro-buthoid scorpions (Chelicerata, Scorpiones). Boletin Sociedad Entomologica Aragonesa, 40: 473-477.","SOLEGLAD, M. E. & W. D. SISSOM. 2001. Phylogeny of the family Euscorpiidae Laurie, 1896 (Scorpiones): a major revision. Pp. 25-111 in: Fet, V. & P.A. Selden (eds). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society, 404 pp.","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81(12): 297-316.","VACHON, M. 1963. De l`utilite, en systematique d`une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle Paris, 35(2): 161-166.","VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaXie en Arachnologie, Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum National d´Histoire Naturelle Paris, 140: 857-958.","VACHON, M. 1979. Arachnids of Saudi Arabia, Scorpiones. Fauna Saudi Arabia, 1: 30-66."]}
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- 2022
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26. Orthochirus katerinae Kovařík & Just 2022, sp. n
- Author
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Kovařík, FrantIšek and Just, Pavel
- Subjects
Arthropoda ,Orthochirus ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Orthochirus katerinae ,Taxonomy - Abstract
Orthochirus katerinae sp. n. (Figures 1–27, Table 1) http://zoobank.org/urn:lsId:zoobank.org:act:D0609699- D022-4A3C-B277-59EF092E8152 Orthochirus innesi, ssp.?: Vachon, 1979: 53–55, figs. 36, 38. Orthochirus scrobiculosus (? in part): Alqahtani et al., 2019: 23, fig. 3i. Orthochirus sp. (? in part): Alqahtani & Badry, 2021: 7, fig. 4. TYPE LOCALITY AND TYPE DEPOSITORY. Saudi Arabia, Hajrah District, 5 km NW of At-Tinah, 20.1107947°N 40.8434278°E, 1400 m a. s. l.; FKCP. FIgure 1. Orthochirus katerinae sp. n., paratype male in vivo habitus. TYPE MATERIAL EXAMINED. Saudi Arabia, Hajrah District, 5 km NW of At-Tinah, 20.1107947°N 40.8434278°E, 1400 m a. s. l., 24–25 October 2022, 12♂ (holotype and paratypes) 2♂ juveniles (paratypes) leg. F. Kovařík & P. Just. ETYMOLOGY. The specific epithet is a patronym honoring Kateřina Rosová, a zoologist who assisted us during many field trips, helped to collect the material and supported us in our research. DIAGNOSIS (♂). Total length 26–30 mm in males. Trichobothrium d 2 on dorsal surface of pedipalp femur absent. Pectinal teeth number 16–20 in males. Movable finger of pedipalps with 7–8 rows of denticles, 8–9 ID and 7 OD. Dorsal carinae on pedipalp patella developed and smooth. Pedipalp femur granulated. Metasoma V dorsal surface mesially finely granulated. Metasoma I–II with 10 carinae, metasoma III with 8 carinae, metasoma IV–V with 2 dorsolateral carinae; complete ventrolateral carinae present also on metasoma V. Ventral carinae of metasoma I–III consist of small granules irregularly in wide row. Metasoma IV–V ventrally and laterally with fine punctation developed, spaces among punctae rather smooth in central part and partly granulated mainly on margins; metasoma I–III ventrally and laterally granulated and bumpy with punctation reduced. Tergites roughly to finely granulated. Sternite VII densely granulated, with four irregularly granulated carinae present. Pedipalp, metasoma and telson glabrous. Moderate to strong tibial spurs present on legs III and IV. Tarsomere I of legs I–III with 3–5 long setae. Ratio length/width of metasoma V 1.13–1.16 in males. Pedipalp femur length/width ratio 3.30–3.32 in males. DESCRIPTION. Total length of adult males 26–30 mm. Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps are given in Table 1. For habitus, see Figures 1–3. ColoratIon (Figs. 1–3). Carapace, tergites, and metasoma black. Femur of pedipalps black and patella of pedipalps black to reddish black, chela of pedipalps yellow. Femur of legs black and patella of legs black to reddish black, other segments of legs yellowish brown. Sternites black with yellow median area in posterior margin of sternites IV–VI, mainly on sternite V. Telson black to reddish brown. Mesosoma and carapace (Figs. 21–22). Mesosoma with a median reduced carina and is roughly to finely granulated. Carapace without carinae, roughly granulated eXcept smooth interocular area. Seventh sternite densely granulated and with four irregularly granulated carinae, the other sternites densely granulated but smooth in middle and posteriorly. Pectinal teeth number 16–20 (4 X 16, 5 X 17, 10 X 18, 4 X 19, 1 X 20) in males. Metasoma and telson (Figs. 15–20). Metasoma I–II with 10 granulated carinae. Metasoma III lacks lateral and metasoma IV lacks lateral and ventromedian carinae. Ventrolateral carinae are developed on metasoma I–III, V and indicated on metasoma IV, dorsolateral carinae are present on all metasomal segments. Ventral carinae of metasoma I–III consist of small granules irregularly in wide row. Metasoma I–IV and partly metasoma V are granulated laterally, metasoma III–V laterally punctate; granulation present on dorsal surfaces of metasoma I and V and absent on metasoma II–IV eXcept several solitary granules mainly on metasoma II. Fine punctation on metasoma IV–V ventrally developed, spaces among punctae rather smooth in central part and partly granulated mainly on margins. Entire metasoma and telson glabrous. Telson without punctation and granulation. PedIpalps (Figs. 4–14). Trichobothrium d 2 on dorsal surface of pedipalp femur absent; trichobothrium e 1 is situated at level with d 4. Femur of pedipalps with five granulated carinae and is granulated. Patella has seven smooth carinae, and the chela has smooth carinae which may be discernible throughout the length of the fiXed finger. The entire pedipalps are glabrous, with several short setae only. Movable fingers with 7–8 rows of denticles, 8–9 ID and 7 OD. Legs (Figs. 23–26). Moderate to strong tibial spurs present on legs III and IV. Femur with four partly granulated carinae; patella with five rather smooth carinae; tibia smooth. Patella with only a few setae. Tibia with several short setae. Tarsomere I of legs I–III with 3–5 long setae, legs IV without these setae. Tarsomeres I–II of all legs internal with two rather irregular rows of setae. Measurements. See Table 1. AFFINITIES. The described features distinguish O. katerinae sp. n. from all other species of the genus. Vachon (1979, fig. 36) cited several widely spread populations from Saudi Arabia as “ Orthochirus innesi Simon 1910, ssp.?” but did not further pursue their taXonomy. In reality, the type locality of O. innesi is in Egypt (Djebel Mokattam). Vachon probably assumed similarity with O. innesi negebensis Shulov & Amitai, 1960 described from Israel (Wadi Nafha, Central Negev), which is currently considered as valid species, Orthochirus negebensis (see, e.g., Lourenço, 2007: 477). Validity of O. katerinae sp. n. is confirmed also by our DNA analysis (paper in preparation), which identified these taXa as related but separate species. Morphologically, O. katerinae sp. n., based on the males, is characterized mainly by the total length of 26–30 mm; pectinal teeth number 16–20; movable finger of pedipalps with 7–8 rows of denticles, 8–9 ID and 7 OD; tarsomere I of legs I–III with 3–5 long setae; ratio length/width of metasoma V 1.13–1.16; pedipalp femur length/width ratio 3.30–3.32 and granulation present on dorsal surfaces of metasoma I and V and absent on metasoma II–IV eXcept several solitary granules mainly on metasoma II.
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27. The genus Barbaracurus in Saudi Arabia (Scorpiones: Buthidae), with description of a new species
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Kovařík, František, Lowe, Graeme, Šťáhlavský, František, and Just, Pavel
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Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kovařík, František, Lowe, Graeme, Šťáhlavský, František, Just, Pavel (2022): The genus Barbaracurus in Saudi Arabia (Scorpiones: Buthidae), with description of a new species. Euscorpius 365: 1-26, DOI: http://doi.org/10.5281/zenodo.7616748, {"references":["KOVARIK, F., G. LOWE, M. SEITER, J. PLISKOVA & F. STAHLAVSKY. 2015. Scorpions of Ethiopia (Arachnida: Scorpiones), Part II. Genus Babycurus Karsch, 1886 (Buthidae), with description of two new species. Euscorpius, 196: 1-31.","KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2018b. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1-41.","KOVARIK, F., G. LOWE, F. STAHLAVSKY & A. A. HURRE. 2019. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XX. Barbaracurus feti sp. n. from Somaliland (Buthidae). Euscorpius, 280: 1-11.","KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.","KOVARIK, F., F. STAHLAVSKY.T. KORINKOVA, J. KRAL & T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1-12.","KOVARIK, F., R. TERUEL & G. LOWE. 2016. Two new scorpions of the genus Chaneke Francke, Teruel et Santibanez-Lopez, 2014 (Scorpiones: Buthidae) from southern Mexico. Euscorpius, 218: 1-20.","KOVARIK, F. & S. WHITMAN. 2005. Cataloghi del Museo di Storia Naturale dell'Universita di Firenze - sezione di zoologia \"La Specola\" XXII. Arachnida Scorpiones. Tipi. Addenda (1998-2004) e checklist della collezione (Euscorpiinae esclusi). Atti della Societa Toscana di Scienze Naturali, Memorie, serie B, 111 (2004): 103-119.","KRAEPELIN, K. 1913. Neue Beitrage zur Systematik der Gliederspinnen.III.A. Bemerkungen zur Skorpionenfauna Indiens. B. Die Skorpione, Pedipalpen und Solifugen Deutsch-Ostafrikas. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 30: 123-196.","LAMORAL, B. H. 1976. Akentrobuthus leleupi, a new genus and species of humicolous scorpion from eastern Zaire, representing a new subfamily of the Buthidae. Annals of the Natal Museum, 22(3): 681-691.","LOWE, G. 2000. A new species of Babycurus (Scorpiones: Buthidae) from northern Oman. Entomological News, 111: 185-192.","ROSSI, A., 2019. On two poorly known scorpions from Yemen: Compsobuthus manzonii (Borelli, 1915) and Babycurus borellii sp. n. (Scorpiones: Buthidae). Arachnida - Rivista Aracnologica Italiana, 2018, 16: 35-44 (dated February 2018 but published/accessible in March 2019).","SAN MARTIN, P. R. 1968. Estudio preliminar sobre una nueva quetotaxia en escorpiones (Microtityus rickyi - Buthidae). MorfologIa y accion mecanica. Caribbean Journal of Science, 8(3-4): 173-180.","SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64-160 in POLIS, G. A. (Ed.) The Biology of Scorpions. Stanford, California: Stanford University Press.","SISSOM, W. D. 1994. Descriptions of new and poorly known scorpions of Yemen (Scorpiones: Buthidae, Diplocentridae, Scorpionidae). Fauna of Saudi Arabia, 14: 3-39.","SOLEGLAD, M. E. & V. FET 2003. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1-34.","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297-316.","TERUEL, R. & T. M. RODRIGUEZ-CABRERA. 2020. Revision of the genus Tityopsis Armas, 1974 (Scorpiones: Buthidae). Part 1. General updates and description of four new species. Euscorpius, 304: 1-40.","VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161-166.","VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3e serie, 140 (Zoologie, 104): 857-958.","VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Serie D Sciences Naturelles, 281 (21): 1597-1599."]}
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28. Orthochirus Karsch 1892
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Kovařík, FrantIšek and Just, Pavel
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Arthropoda ,Orthochirus ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Orthochirus Karsch, 1892 (Figures 1–27, Table 1) http: //zoobank. org/urn: lsId: zoobank. org: act: 79D5A7E7- F14F -4D64-90ED -89BF47023562 Orthochirus Karsch, 1892: 306; Kovařík et al., 2020: 1–73, figs. 1–352, tables 1–4 (complete reference and synonyms list until 2020). DIAGNOSIS. Total length of adults 22–55 mm. Tergites I–VI with weak median or lateral carinae, or carinae indistinct. Patellar trichobothrium d 3 located between dorsomedian and dorsointernal carinae. Dorsal trichobothria of femur arranged in beta-configuration. Trichobothrium d 2 of pedipalp femur absent or present on dorsal surface. Chelicerae with typical buthid dentition (Vachon, 1963), ventral aspect of fiXed finger with two denticles. Tibial spurs present on legs III and IV; in all legs, mid-ventral aspect of tarsomere II sparsely setose with 1 or 2 rows of short spiniform setae. Pectines with fulcra, densely hirsute. ProXimal dentate margins of pedipalp fingers not strongly undulate; movable fingers of pedipalps with 7–10 rows of denticles and 2–5 subterminal denticles. Carapace strongly trapezoidal, lacking distinct carinae; in lateral view distinctly inclined downward from median eyes to anterior margin; 5 pairs of lateral eyes. First and second metasomal segments with carinae. Metasoma posteriorly widened; metasomal segments IV and V ventrally punctate. Telson elongate with subaculear tubercle absent, aculeus robust, as long as or longer than vesicle; hemispermatophore capsule with 3 laminate lobes + 1 hook-like basal lobe; spiracles slit-like., Published as part of Kovařík, FrantIšek & Just, Pavel, 2022, Orthochirus katerinae sp. n. (Scorpiones: Buthidae) from Saudi Arabia, pp. 1-9 in Euscorpius (Oxford, England) (Oxford, England) 362 (1) on page 1, DOI: 10.5281/zenodo.7399385, {"references":["KARSCH, F. 1892. Arachniden von Ceylon und von Minikoy, gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift, 36 (1891): 267 - 310.","KOVARIK, F., V. FET, & E. A. YAGMUR. 2020. Further review of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) fromAsia: taXonomic position of O. melanurus, O. persa, O. scrobiculosus, and description of siX new species. Euscorpius, 318: 1 - 73.","VACHON, M. 1963. De l`utilite, en systematique d`une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle Paris, 35 (2): 161 - 166."]}
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29. Reanalysis of Teruelius and Grosphus (Scorpiones Buthidae with descriptions of two new species
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Lowe, Graeme and KOvAřík, FRANtIšEk
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Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Lowe, Graeme, KOvAřík, FRANtIšEk (2022): Reanalysis of Teruelius and Grosphus (Scorpiones Buthidae with descriptions of two new species. Euscorpius 356: 1-105, DOI: http://doi.org/10.5281/zenodo.7475051, {"references": ["HEINGARD, M., N. TURETZEK, N-M. PRPIC & R. JANSSEN. 2019. FoxB, a new and highly conserved key factor in arthropod dorsal-ventral (DV) limb patterning. Evolution and Development, 10: 28, 1-16.", "JANSSEN, R., N. M. FEITOSA, W. G. M. DAMEN & N-M. PRPIC. 2008. The T-box genes H15 and optomotor-blind in the spiders Cupiennius salei, Tegenaria atrica and Achaearanea tepidariorum and the dorsoventral axis of arthropod appendages. Evolution and Development, 10: 2, 143-154.", "KAPUR, J. N., P. K. SAHOO & A. K. C. WONG. 1985. A new method for gray-level picture thresholding using the entropy of the histogram. Computer Vision, Graphics and Image Processing, 29 (3): 273-285. doi: 10.1016/0734- 189x(85)90125-2", "KLOOCK, C. T. 2008. A comparison of fluorescence in two sympatric scorpion species. Journal of Photochemistry and Photobiology B: Biology, 91: 132-136.", "KLOOCK, C. T. 2009. Reducing scorpion fluorescence via prolonged exposure to ultraviolet light. Journal of Arachnology, 37: 368-370.", "KLOOCK, C. T., K. ABRAHAM & R. REYNOLDS. 2010. Ultraviolet light detection: a function of scorpion fluorescence. Journal of Arachnology, 38: 441-445.", "KOVARIK, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societas Zoologicae Bohemoslovenicae, 59: 187-207.", "KOVARIK, F. 2007a. Leiurus nasheri sp. nov. from Yemen (Scorpiones, Buthidae). Acta Societatis Zoologicae Bohemicae, 71: 137-141.", "KOVARIK, F. 2007b. A revision of the genus Hottentotta Birula, 1908, with descriptions of four new species (Scorpiones, Buthidae). Euscorpius, 58: 1-107.", "KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.", "KOVARIK, F. 2018. Notes on the genera Buthacus, Compsobuthus, and Lanzatus with several synonymies and corrections of published characters (Scorpiones: Buthidae). Euscorpius, 269: 1-12.", "KOVARIK , F & LOWE, G. 2022. Review of Orthochiroides Kovarik, 1998 with description of a new species (Scorpiones Buthidae). Euscorpius, 349: 1-42.", "KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Euscorpius, 232: 1-38.", "KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI & A. A. HURRE. 2018a. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1-82.", "KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2018b. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with descriptions of Barbaracurus gen. n. and two new species from Oman and Yemen Euscorpius, 267: 1-41.", "KRAEPELIN, K. 1891. Revision der Skorpione. I. Die Familie der Androctonidae. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 8: 1-144.", "KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In: F. DAHL (Ed.). Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8 (Arachnoidea): 1-265.", "KRAEPELIN, K. 1900. Ueber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1(4): 1-17.", "KRAEPELIN, K. 1908.Die sekundaren Geschlechtscharaktere der Skorpione, Pedipalpen und Solifugen. Mitteilungen aus dem Naturhistorischen Museum (2. Beiheft zum Jahrbuch der Hamburgischen wissenschaftlichen Anstalten), 25: 181-225.", "KUHL, F. P. & G. R. GIARDINA. 1982. Elliptic Fourier features of a closed contour. Computer Graphics and Image Processing, 18: 236-258.", "LAMORAL, B. H. 1976. Akentrobuthus leleupi, a new genus and species of humicolous scorpion from eastern Zaire, representing a new subfamily of the Buthidae. Annals of the Natal Museum, 22 (3): 681-691.", "LAMORAL, B. H. 1979. The scorpions of Namibia (Arachnida: Scorpionida). Annals of the Natal Museum, 23 (3): 497-784.", "LANKESTER, E. R. 1885. Notes on certain points of the anatomy and generic characters of Scorpions. Transactions of the Zoological Society, London, 11 (10): 372-384.", "LAWRENCE, R. F. 1938. The arachnida of the Transvaal Museum expedition to South Rhodesia, November- December, 1937. Scorpions and Solifugae. Annals of the Transvaal Museum, 19 (2): 289-296.", "LAWRENCE, R. F. 1954. Fluorescence in Arthropoda. Journal of the Entomological Society of South Africa, 17: 167-170.", "LAWRENCE, R. F. 1961. A new forest-living scorpion from the Transvaal. Annals and Magazine of Natural History, (13) 4: 123-126.", "LAWRENCE, R. F. 1966. New and little known scorpions and solifuges from the Namib Desert and South West Africa. Scientific Papers of the Namib Desert Research Station, 29: 1-11.", "LI, S., K. C. KIM, S. ZONG, Y. LIU, J. KANG, C. PAK, Z. CAO & Y. WU. 2022. Dynamic synthesis and transport of fluorescent substances from moulting scorpions. Journal of Photochemistry & Photobiology, B: Biology, 234, 112511. https://doi.org/10.1016/j. jphotobiol.2022.112511", "LOPEZ-CABRERA, D., G. RAMOS-ORTIZA, E. GONZALEZ-SANTILLAN & R. ESPINOSA-LUNA. 2020. Characterization of the fluorescence intensity and color tonality in the exoskeleton of scorpions. Journal of Photochemistry & Photobiology, B: Biology, 209, 111945: 1-9.", "LORIA, S. F. & L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS ONE 9 (12): e112913. doi:10.1371/journal.pone.0112913.", "LOURENCO, W. R. 1995. Description de trois noveaux genres et quatre nouvelles especes de scorpions Buthidae de Madagascar. Bulletin du Museum National d'Histoire naturelle, Paris (Zoologie, Biologie et Ecologie Animale), (4), 17A (1-2): 95-106.", "LOURENCO, W. R. 1996a. Premier cas connu d'un subfossile de scorpion dans le copal de Madagascar. Comptes Rendus de l'Academie des Sciences, Paris, Ser. IIa, 323: 889-891.", "LOURENCO, W. R. 1996b. Faune de Madagascar. 87. Scorpions (Chelicerata, Scorpiones). Paris: Museum National d'Histoire Naturelle. 102 pp.", "LOURENCO, W. R. 1996c. Origins and affinities of the scorpion fauna of Madagascar. Pp. 441-455 in: LOURENCO, W. R. (Ed.). Biogeographie de Madagascar. Paris, Orstom.", "LOURENCO, W. R. 1996d. Microcharmus hauseri, nouvelle espece de scorpion de Madagascar (Scorpiones, Buthidae). Revue Suisse de Zoologie, 103 (2): 319-322.", "LOURENCO, W. R. 1998a. Une nouvelle famille est necessaire pour les microscorpions humicoles de Madagascar et d'Afrique. Comptes Rendus de l 'Academie des Sciences (Paris), 321: 845-848.", "LOURENCO, W. R. 1998b. Description of a new species of scorpion from the reserve Speciale d'Anjanaharibe-Sud, Madagascar. Pp. 69-72 in: GOODMAN, S.M. (Ed.). A floral and faunal inventory of the reserve Speciale d'Anjanaharibe-Sud, Madagascar, with reference to elevational variation\" Fieldiana Zoology, new series, 90 (1495). Chicago Natural History Museum.", "LOURENCO, W. R. 1999a. A new species of Grosphus Simon (Scorpiones, Buthidae ), the first record of an intertidal scorpion from Madagascar. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 12 (156): 183-188.", "LOURENCO, W. R. 1999b. On the phylogenetic position of the genus Birulatus Vachon, 1973 (Scorpiones, Buthidae) and redescription of Birulatus haasi. Zoology in the Middle East, 18: 109-113.", "LOURENCO, W. R. 1999c. Complementary notes on the phylogenetic position of the genus Lychasioides Vachon, 1973, and redescription of Lychasioides amieti (Chelicerata, Scorpiones, Buthidae). Revue Arachnologique, 13 (1): 7-13.", "LOURENCO, W. R. 1999d. Un modele de distribution geographique presente par les scorpions du genre Microcharmus Lourenco, avec la description d'une nouvelle espece. Comptes Rendus de l' Academie des Sciences, ser. 3, Sci. Vie, 322 (10): 843-846.", "LOURENCO, W. R. 2000a. Un nouveau genre de Scorpion malgache, maillon possible entre les Microcharmidae et les Buthidae. Comptes Rendus de l'Academie des Sciences, series III, Sciences de la vie, 323 (10): 877- 881.", "LOURENCO, W. R. 2000b. More about the Buthoidea of Madagascar, with special references to the genus Tityobuthus Pocock (Scorpiones, Buthidae). Revue Suisse de Zoologie, 107 (4): 721-736.", "LOURENCO, W. R. 2002. Nouvelles donnees sur la morphologie et la biogeographie des Microcharmus Lourenco avec confirmation de la validite des Microcharmidae (Chelicerata, Scorpiones). Biogeographica, 78(1): 35-47.", "LOURENCO, W. R. 2003a. Scorpiones, scorpions. Pp. 575- 579 in: GOODMAN, S. & BENSTEAD, J. (Eds.). The Natural History of Madagascar. Chicago, Illinois: The University of Chicago Press.", "LOURENCO, W. R. 2003b. New taxonomic considerations on some species of the genus Grosphus Simon, with description of a new species (Scorpiones, Buthidae). Revue Suisse de Zoologie, 110 (1): 141-154.", "LOURENCO, W. R. 2003c. Humicolous buthoid scorpions: a new genus and species from French Guiana. Comptes Rendus Biologies, 326 (12): 1149-1155.", "LOURENCO, W. R. 2004a. Humicolous microcharmid scorpions: a new genus and species from Madagascar. Comptes Rendus Biologies, 327: 77-83.", "LOURENCO, W. R. 2005. Diversity and endemism in the scorpion fauna of Madagascar. Biogeographica, 80: 43- 64.", "LOURENCO, W. R. 2009. A new sub-fossil scorpion of the genus Microcharmus Lourenco, 1995 from Malagasy copal (Scorpiones, Microcharmidae). Boletin Sociedad Entomologica Aragonesa, 44: 135-137.", "LOURENCO, W. R. 2012. Fluorescence in scorpions under UV light; can chaerilids be a possible exception? Comptes Rendus Biologies, 335: 731-734.", "LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in south-western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631-645.", "LOURENCO, W. R. 2020. A remarkable new species of Leiurus Ehrenberg, 1828 from the north deserts of Mali (Scorpiones: Buthidae). Revista Iberica de Aracnologia, 37: 147-152.", "LOURENCO, W. R. 2021. 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30. A revision of the genus Mesobuthus Vachon 1950 with a description of 14 new species (Scorpiones Buthidae)
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Kovařík, František, Fet, Victor, Gantenbein, Benjamin, Graham, Matthew R., Aydin, Ersen, Yağmur, Šťáhlavský, František, Nikita, Poverennyi, and Novruzov, Nizami E.
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Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kovařík, František, Fet, Victor, Gantenbein, Benjamin, Graham, Matthew R., Aydin, Ersen, Yağmur, Šťáhlavský, František, Nikita, Poverennyi, Novruzov, Nizami E. (2022): A revision of the genus Mesobuthus Vachon 1950 with a description of 14 new species (Scorpiones Buthidae). Euscorpius 348: 1-189, DOI: http://doi.org/10.5281/zenodo.7162849, {"references": ["ADILARDI, R. S., A. A. OJANGUREN-AFFILASTRO, D. A. MARTI & L. M. MOLA. 2020. Chromosome puzzle in the southernmost populations of the medically important scorpion Tityus bahiensis (Perty 1833) (Buthidae), a polymorphic species with striking structural rearrangements. Zoologischer Anzeiger, 288: 139-150.", "AITCHISON, J. E. T. 1888. The botany of the Afghan Delimitation Commission. Transactions of Linnean Society of London, Botany, 3(1): 1-139.", "ANIKIN, V. V. 1997. [Rare species of Insecta and Arachnida in marginal structures of the steppe zone of the Saratov Province]. 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LOWE. 2000. Family Buthidae. Pp. 54-286 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder (eds.). Catalog of the Scorpions of the World (1758-1998). New York: New York Entomological Society, 690 pp.", "FET, V., M. E. SOLEGLAD & S. L. ZONSTEIN. 2011. The genus Akrav Levy, 2007 (Scorpiones: Akravidae) revisited. Euscorpius, 134: 1-49.", "FET, V., M. E. SOLEGLAD, S. L. ZONSTEIN, I. NAAMAN, S. LUBATON, B. LANGFORD & A. FRUMKIN. 2017. The second record of a relict Akrav israchanani Levy, 2007 (Scorpiones: Akravidae) from Levana Cave, Israel. Euscorpius, 247: 1-12.", "FOMICHEV, A. A. 2011. A new locality of Mesobuthus eupeus thersites (C. L. Koch, 1839) (Scorpiones: Buthidae) in East Kazakhstan. Euscorpius, 136: 1-3.", "GANTENBEIN, B., V. FET, I. A. GANTENBEIN-RITTER & F. BALLOUX. 2005. Evidence for recombination in scorpion mitochondrial DNA (Scorpiones: Buthidae). Proceedings of the Royal Society: Biological Sciences, 272(1564): 697-704.", "GANTENBEIN, B., V. FET & A. V. GROMOV. 2003. The first DNA phylogeny of four species of Mesobuthus Vachon, 1950 (Scorpiones: Buthidae) from Eurasia. Journal of Arachnology, 31: 412-420.", "GHARAKHLOO, M. M., S. A. HEYDARABAD & E. A. YAGMUR. 2018. The scorpion fauna of West Azerbaijan Province in Iran (Arachnida: Scorpiones). Biharean Biologist, 12(2): 84-87.", "GRAHAM, M. R., E. A. MYERS, R. KAISER & V. FET. 2019. Cryptic species and co-diversification in sand scorpions from the Karakum and Kyzylkum deserts of Central Asia. Zoologica Scripta, 48(6): 801-812.", "GRAHAM, M. R., V. OLAH-HEMMINGS & V. FET. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a longstanding component of Central Asian biogeography (Scorpiones: Buthidae). Zoology in the Middle East, 55: 95-110.", "GRAHAM, M. R., D. A. WOOD, J. A. HENAULT, Z. J. VALOIS & P. E. CUSHING. 2017. Ancient lakes, Pleistocene climates and river avulsions structure the phylogeography of a large but little-known rock scorpion from the Mojave and Sonoran deserts. Biological Journal of the Linnean Society, 122(1): 133-146.", "GROMOV, A. V. 2001. The northern boundary of scorpions in Central Asia. Pp. 301-306 in Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society.", "GROMOV, A. V. & Ye. Ye. KOPDYKBAEV. 1994. [The fauna of the scorpions and solpugids (Arachnida: Scorpiones, Solifugae) in Kazakhstan]. Selevinia, 2(2): 19-23 (in Russian).", "HEDDERGOTT, M., M. STUBBE, W. STUBBE, P. STEINBACH & A. STUBBE. 2016. Geographical distribution of the genus Mesobuthus (Scorpiones: Buthidae) in Mongolia. Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, 13: 147-164.", "HUSSEN, F. S. & S. T. AHMED. 2020. New data of scorpion fauna, include two new records with identification key of scorpion species (Arachnida: Scorpiones) in Iraq. Plant Archives, 20 (2): 6711-6725.", "KACHEL, H. S, A. M. AL-KHAZALI, F. S. HUSSEN & E. A.YAGMUR. 2020. Checklist and review of the scorpion fauna of Iraq (Arachnida: Scorpiones). Arachnologische Mitteilungen, 61(1): 1-10.", "KALTSAS, D., I. STATHI & V. FET. 2008. Scorpions of the Eastern Mediterranean. Pp. 209-246 in Makarov, S.E. & R.N. Dimitrijevic (eds.). Advances in Arachnology and Developmental Biology. Papers dedicated to Prof. Dr. Bozidar Curcic. Vienna-Belgrade-Sofia, 517 pp.", "KARATAS, AY. & M. COLAK. 2005. Scorpions of Gaziantep Province, Turkey (Arachnida: Scorpiones). Euscorpius, 30: 1-7.", "KARATAS, A., V. FET & M. M. GHARKHELOO. 2012. On the type locality of Mesobuthus vesiculatus (Pocock, 1899) (Scorpiones: Buthidae). Euscorpius, 147: 1-3.", "KARATAS,A. & M. M. GHARKHELOO.2012. Redescription of Mesobuthus vesiculatus (Pocock, 1899) (Scorpiones: Buthidae) based on specimens from Iran. Turkish Journal of Zoology, 36(5): 576-584.", "KARATAS, A., M. M. GHARKHELOO & M. UCAK. 2012. Contribution to the distribution of the scorpions of Iran. Zoology in the Middle East, 55: 111-120.", "KARATAS, A. & A. KARATAS. 2001. First record of Mesobuthus eupeus (C. L. Koch, 1839) from central Anatolia (Scorpiones: Buthidae). Pp. 297-299 in Fet, V. & P. A. Selden (eds.) Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks, UK.", "KARATAS, A. & A. KARATAS. 2003. Mesobuthus eupeus (C.L. Koch, 1839) (Scorpiones: Buthidae) in Anatolia. Euscorpius, 7: 1-7.", "KARATAS, A., M. UCAK, A. KARATAS & O. EROGLU. 2019. Records of chromosomal data of some scorpions (Arachnida: Scorpiones) from Turkey. Hacettepe Journal of Biology and Chemistry, 47: 1-6.", "KESSLER, K. F. 1874. O russkikh skorpionakh [On Russian scorpions]. Trudy Russkago Entomologicheskago Obshchestva v S.-Peterburge [Transactions of the Russian Entomological Society in St. Petersburg], (1): 3-27 (in Russian) [date on the volume cover, 1876; separatum first available in 1874].", "KOCH, C. L. 1839a. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung, 5(6): 125-158, figs. 418-419.", "KOCH, C. L. 1839b. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung, 6(3): 51-53, figs. 466-467.", "KOCH, L. 1878. Kaukasische Arachnoideen. Pp. 36-71 in Schneider, O. (ed.) Naturwissenschaftliche Beitrage zur Kenntnis der Kaukasuslander,auf Grund seiner Sammelbeute herausgegeben. Dresden: Gesellschaft \"Isis\", 3.", "KOVARIK, F. 2002. A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany. Serket, 8(1): 1-23.", "KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.", "KOVARIK, F. 2019. Taxonomic reassessment of the genera Lychas, Mesobuthus, and Olivierus, with descriptions of four new genera (Scorpiones: Buthidae). Euscorpius, 288: 1-27.", "KOVARIK, F. & P. BERON. 2015. A checklist of the scorpions (Arachnida, Scorpiones) in the collections of the National Museum of Natural History (Sofia). Historia naturalis bulgarica, 22: 37-44.", "KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.", "KOVARIK, F., F. STAHLAVSKY, T. KORINKOVA, J. KRAL & T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1-12.", "KOVARIK, F. & S. WHITMAN. 2005. Cataloghi del Museo di Storia Naturale dell'Universita di Firenze - sezione di zoologia \"La Specola\" XXII. Arachnida Scorpiones. Tipi. Addenda (1998-2004) e checklist della collezione (Euscorpiinae esclusi). Atti della Societa Toscana di Scienze Naturali, Memorie, serie B, 111 (2004): 103-119.", "KOVARIK, F., E. A. YAGMUR, V. FET & S. NAVIDPOUR. 2011. On two subspecies of Mesobuthus eupeus (C. L. Koch, 1839) in Turkey (Scorpiones: Buthidae). Euscorpius, 109: 1-15.", "KRYZHANOVSKY, O. L. 1965. Sostav i proiskhozhdenie nazemnoi fauny Srednei Azii [The Composition and Origin of the Terrestrial Fauna of Middle Asia]. Moscow & Leningrad: Nauka, 419 pp. (in Russian; English translation: New Delhi, 1980).", "KUMAR, S., G. STECHER, M. LI, C. KNYAZ & K. TAMURA. 2018. MEGA X: molecular evolutionary genetics analysis across computing platforms. Molecular Biology and Evolution, 35(6): 1547.", "LARSSON, A. 2014. AliView: a fast and lightweight alignment viewer and editor for large datasets. Bioinformatics, 30(22): 3276-3278 (Nov 15). doi: 10.1093/bioinformatics/btu531.", "LIECHTENSTEIN, H. 1823. Naturhistorische Anhang. Pp. 113-150 in: Eversmann, E. Reise von Orenburg nach Buchara. Berlin: E. H. G. Christiani.", "LIPSKY, V. I. 1902. Gornaya Bukhara: rezul'taty tryokhletnikh puteshestvii v Srednyuyu Aziyu v 1896, 1897 i 1899 godu. Chast' 1. Gornaya Bukhara: Gissarskaya ekspeditsiya, 1896 g. [The Mountainous Buchara: the results of three years of travels to Central Asia in 1896, 1897 and 1899. Part 1. The Mountainous Buchara: the Gissar Expedition, 1896]. St. Petersburg (in Russian).", "MAKINO, S. 1956. A review of the chromosome numbers in animals. Tokyo: Hokurykan.", "MAYR, E. 1969. Principles of Systematic Zoology. New York: McGraw-Hill Book Co., xi + 428 pp.", "MAYR, E. 1971. Prinstipy zoologicheckoi sistematiki [Principles of Systematic Zoology]. Moscow: Mir, 454 pp. (in Russian).", "MIRSHAMSI, O. 2012. Ecological niche modeling of two scorpion species Mesobuthus eupeus (C. L. Koch, 1839) and M. phillipsii (Pocock, 1889) from the Iranian Plateau and Zagros region (Arachnida: Scorpiones). Euscorpius, 154: 1-10.", "MIRSHAMSI, O., A. SARI, E. ELAHI & S. HOSSEINIE. 2010. Phylogenetic relationships of Mesobuthus eupeus (C. L. Koch, 1839) inferred from COI sequences (Scorpiones: Buthidae). Journal of Natural History, 44(47): 2851-2872.", "MIRSHAMSI, O., A. SARI, E. ELAHI & S. HOSSEINIE. 2011a. Mesobuthus eupeus (Scorpiones: Buthidae) from Iran: A polytypic species complex. Zootaxa, 2929: 1-21.", "MIRSHAMSI, O., A. SARI & S. HOSSEINIE. 2011b. History of study and checklist of the scorpion fauna (Arachnida: Scorpiones) of Iran. Progress in Biological Sciences, 1(2):16-28.", "MLIKOVSKY, J. & S. FRAHNERT. 2009. Type specimens and type localities of birds (Aves) collected by Eduard Eversmann and Christian Pander during the Negri Expedition to Bukhara in 1820-1821. Zootaxa, 2297: 15-26.", "MORADI, M., E. A. YAGMUR, P. M. GHARAKHLOO & F. AHMADI. 2015. Scorpion fauna of Zanjan Province, Iran (Arachnida: Scorpiones). Journal of Applied Biological Sciences, 9(1): 11-14.", "MORITZ, M. & S.-CH. FISCHER. 1980. Die Typen der Arachniden-Sammlung des Zoologischen Museums Berlin. III. Scorpiones. Mitteilungen aus dem Zoologischen Museum in Berlin, 2(36): 309-326.", "NAVIDPOUR, S., M. EZATKHAH, F. KOVARIK, M. E. SOLEGLAD & V. FET. 2011. Scorpions of Iran (Arachnida, Scorpiones). Part VII. Kerman Province. Euscorpius, 131: 1-32.", "NAVIDPOUR, S., V. FET, F. KOVARIK & M. E. SOLEGLAD. 2008a. Scorpions of Iran (Arachnida, Scorpiones). Part III. Ilam Province. Euscorpius, 69: 1-29.", "NAVIDPOUR, S., V.FET, F. KOVARIK& M. E. SOLEGLAD. 2012. Scorpions of Iran (Arachnida, Scorpiones). Part VIII. Fars Province. Euscorpius, 139: 1-32.", "NAVIDPOUR, S., F. KOVARIK, M. E. SOLEGLAD & V. FET. 2008b. Scorpions of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province. Euscorpius, 65: 1-41.", "NAVIDPOUR, S., F. KOVARIK, M. E. SOLEGLAD & V. FET. 2008c. Scorpions of Iran
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31. Review of Orthochiroides Kovařík 1998 with description of a new species (Scorpiones: Buthidae)
- Author
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Kovařík, František and Lowe, Graeme
- Subjects
Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kovařík, František, Lowe, Graeme (2022): Review of Orthochiroides Kovařík 1998 with description of a new species (Scorpiones: Buthidae). Euscorpius 349: 1-42, {"references":["ACOSTA, L. E., D. M. CANDIDO, E. H. BUCKUP & A. D. BRESCOVIT. 2008. Description of Zabius gaucho (Scorpiones, Buthidae), a new species from southern Brazil, with an update about the generic diagnosis. Journal of Arachnology, 36: 491-501.","BIRULA, A. A. 1917. Chlenistobryukhie paukoobraznye Kavkazskogo Kraya. Part I. Scorpiones. Zapiski Kavkazskogo Muzeya, 5: 1-253.","CHEUNG, C., L. DE VANTIER & K. VAN DAMME. 2006. Socotra. A Natural History of the Islands and Their People. Odyssey.","CULEK, M. 2013. Geological and morphological evolution of the Socotra Archipelago (Yemen) from the biogeographical view. Journal of Landscape Ecology, 6 (3):84-108.","DUPRE, G. 2007. Conspectus genericus scorpionorum 1758- 2006 (Arachnida: Scorpiones). Euscorpius, 50: 1-31.","EL-HENNAWY, H. K. 1992. A catalogue of the scorpions described from the Arab countries (1758-1990) (Arachnida: Scorpionida). Serket, 2(4): 95-153.","FET, E. V., D. NEFF, M. R. GRAHAM & V. FET. 2003. Metasoma of Orthochirus (Scorpiones: Buthidae): are scorpions evolving a new sensory organ? Revista Iberica de Aracnologia, 8: 69-72.","FET, V. & F. KOVARIK. 2020. New scorpion taxa (Arachnida: Scorpiones) described in the journal \"Euscorpius\" in 2002-2020. Euscorpius, 300: 1-31.","FET, V. & G. LOWE. 2000. Family Buthidae C. L. Koch, 1837, pp. 54-286 in: Fet, V., Sissom, W. D., G. Lowe & M. E. Braunwalder. 2000. Catalog of the Scorpions of the World (1758-1998). The New York Entomological Society, New York, 689 pp.","FET, V. & M. E. SOLEGLAD. 2005. Contributions to Scorpion Systematics. I. On Recent Changes in HighLevel Taxonomy. Euscorpius, 31: 1-13.","FET, V., M. E. SOLEGLAD & G. LOWE. 2005. A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1-40.","FRANCKE, O. F. 1977. Taxonomic observations on Heteronebo Pocock (Scorpionida: Diplocentridae). Journal of Arachnology, 4: 95-113.","GOLOBOFF, P. A. 1993. Estimating character weights during tree search. Cladistics, 9: 83-91.","GOLOBOFF, P. A., J. S. FARRIS, M. KALLERSJO, B. OXELMAN, M. J. RAMIREZ & C. A. SZUMIK. 1993. Improvements to resampling measures of group support. Cladistics, 19: 324-332.","GOLOBOFF, P. A. & S. A. CATALANO. 2016. TNT version 1.5, including a full implementation of phylogenetic morphometrics. Cladistics, 32: 221-238.","HIRST, S. 1925. On some scorpions from Morocco, with the description of a new genus and species. Annals and Magazine of Natural History, 9 (15): 414-416.","HULA, V. & J. NIEDOBOVA. 2020. The Mediterranean Recluse Spider Loxosceles rufescens (Dufour, 1820): a new invasive for Socotra Island (Yemen). Rendiconti Lincei. Scienze Fisiche e Naturali, https://doi.org/10.1007/ s12210-020-00925-7.","KARSCH, F. 1891. Arachniden von Ceylon und von Minikoy gesammelt von den Herren Doctoren P. und S. Sarasin. Berliner Entomologische Zeitschrift, 36 (2): 267-310.","KOVARIK, F. 1996. Baloorthochirus becvari gen. et sp. n. from Pakistan, and taxonomic position of Orthochirus luteipes (Scorpiones: Buthidae). Acta Societatis Zoologicae Bohemoslovenicae, 60: 177-181.","KOVARIK, F. 1998. Three new genera and species of Scorpiones (Buthidae) from Somalia. Acta Societatis Zoologicae Bohemicae, 62: 115-124.","KOVARIK, F. 2000. The Socotra Island and its scorpions. Akvarium terarium, 43(7): 63-67 (in Czech).","KOVARIK, F. 2002. A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany. Serket, 8(1): 1-23.","KOVARIK, F. 2003. Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones), with a key and descriptions of three new species. Acta Societatis Zoologicae Bohemicae, 67: 133-159.","KOVARIK, F. 2004. Revision and taxonomic position of genera Afghanorthochirus Lourenco & Vachon, Baloorthochirus Kovarik, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kovarik, Pakistanorthochirus Lourenco, and Asian Orthochirus Karsch, with descriptions of twelve new species (Scorpiones, Buthidae). Euscorpius, 16: 1-33.","KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp.","KOVARIK, F., V. FET & M. SIYAM. 2020a. Taxonomic position of Orthochirus olivaceus (Karsch, 1881),the type species of the genus Orthochirus Karsch, 1892 (Scorpiones: Buthidae). Euscorpius, 319: 1-15.","KOVARIK, F., V. FET & E. A YAGMUR. 2020b. Further review of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) from Asia: taxonomic position of O. melanurus, O. persa, O. scrobiculosus, and description of six new species. Euscorpius, 318: 1-73.","KOVARIK, F. & G. LOWE. 2012. Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae). Euscorpius, 138: 1-25.","KOVARIK, F. & G. LOWE. 2019. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVIII. Gint banfasae sp. n. from Somaliland (Buthidae). Euscorpius, 2729: 1-14.","KOVARIK, F. & G. LOWE. 2020. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXIV. Leiurus (Buthidae), with description of Leiurus gubanensis sp. n. Euscorpius, 309: 1-19.","KOVARIK, F. & G. LOWE. 2021. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXVI. Records of Hottentotta polystictus (Pocock, 1896), with descriptions of H. haudensis sp. n. and H. nigrimontanus sp. n. (Buthidae) from Somaliland. Euscorpius, 330: 1-28.","KOVARIK F., G. LOWE, P. JUST, A. I. AWALE, H. SH A. ELMI & F. STAHLAVSKY. 2018. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XV. Review of the genus Gint Kovarik et al., 2013, with description of three new species from Somaliland (Scorpiones, Buthidae). Euscorpius, 259: 1-41."]}
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32. Orthochiroides insularis
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Kovařík, František and Lowe, Graeme
- Subjects
Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Orthochiroides ,Buthidae ,Orthochiroides insularis ,Taxonomy - Abstract
Orthochiroides insularis (Pocock, 1899) (Figures 1–4, 82, 87–94, 111–112, 119, 130–133, 140–142, 166, 205–211; Tables 1, 3) Butheolus insularis Pocock, 1899: 8–9; Pocock, 1903: 180– 181; Kraepelin, 1903: 565; Francke, 1977: 112; Vachon, 1979: 237. Orthochirus insularis: Birula, 1917: 215. Orthochirus bicolor insularis: Levy & Amitai, 1980: 94; ElHennawy, 1992: 129; Fet & Lowe, 2000: 194; Kovařík, 2000: 64–66. Orthochiroides insularis: Kovařík, 2004: 23. TYPE LOCALITY AND TYPE REPOSITORY. Socotra, Mt. Raggit; BMNH. TYPE MATERIAL EXAMINED. Yemen, Socotra Island, Hadibu Plain (Mt. Raggit, 1000 feet), 1♀ (holotype), leg. Grant and Forbes, BMNH No. 1899.7.4.180. OTHER MATERIAL EXAMINED (FKCP). Yemen, Socotra Island, Qalansiyah env., Ditwah (lagoon), 23 m a. s. l., 12°41'42"N 53°30'08"E, 9.XII.2003, 1♂ (Figs. 1–2, 82, 87–90, 111), leg. D. Král; Gubbah village env., 7 m a. s. l., 12°36'35"N 53°46'56"E, 23.XI.2003, 1♀ (Figs. 3–4, 91–94, 112) 1juv., leg. D. Král; Qaariah village env., 11 m a. s. l., 12°38'05"N 54°12'39"E, 28.XI.2003, 1♂, leg. J. Farkač. DIAGNOSIS (ADULT ♂ ♀). Total length of adults 22–32 mm. Petite ‘trichobothria’ d 2 on dorsal surfaces of pedipalp femur and patella reduced or absent. Chela smooth, carinae E and D1 weak or obsolete on manus. Pectinal tooth count: 18–19 (♂), 16–18 (♀). Movable finger of pedipalp chela with 7 rows of median denticles, flanked by 7 ID and 7 OD. Metasoma IV–V laterally and ventrally punctate, without carinae except for smooth, rugose dorsosubmedian carinae, and ventrolateral carinae on metasoma V which are anteriorly smooth and posteriorly denticulate. Metasoma III either laterally and ventromedially punctate (♂), or laterally punctate and ventromedially granulate-reticulate (♀). Punctae small with diameters less than the mean distance between their centroids, together occupying much less area than smooth surfaces between them. Dorsal surfaces of all metasomal segments smooth, without punctae or granules. Aculeus much shorter than vesicle, aculeus L/telson L ~0.20. Color uniformly brown to black. Femur, patella and chela manus of pedipalp brown to black; chela fingers, and tibia and tarsomeres of legs yellowish brown to green. Tergites roughly granulated. Sternites IV–V with lateral surfaces matte, almost smooth. Sternite VII matte, almost smooth, with four distinct granulated carinae. Pedipalp, metasoma and telson glabrous, only metasoma V may have dorsolateral rows of setae, which can also occur on dorsal surface of telson. Moderately developed tibial spurs present on legs III and IV. Tarsomere I of legs I–III with 4–6 long setae in both sexes. DISTRIBUTION. Yemen, north part of Socotra Island (Fig. 119). COMMENTS. Fet & Lowe (2000: 194) listed “? Orthochirus socotrensis ” cited by Francke (1977) as a possible lapsus and synonym of Orthochirus bicolor insularis. However, Francke (1977: 112) cited the presence of three endemic species of scorpion on Socotra Island: Hemiscorpion socotranus Pocock, 1899, Butheolus insularis Pocock, 1899 and Orthochirus socotrensis (Pocock). He omitted Buthus socotrensis Pocock, 1889 (= Buthotus socotrensis = Hottentotta socotrensis), also known from Socotra Island. It is possible that Francke cited ‘ Orthochirus socotrensis ’ in error, instead of Buthotus socotrensis, as Pocock did not describe Orthochirus socotrensis (see also Vachon, 1979: 237)., Published as part of Kovařík, František & Lowe, Graeme, 2022, Review of Orthochiroides Kovařík 1998 with description of a new species (Scorpiones: Buthidae), pp. 1-42 in Euscorpius 349 on page 4, {"references":["FRANCKE, O. F. 1977. Taxonomic observations on Heteronebo Pocock (Scorpionida: Diplocentridae). Journal of Arachnology, 4: 95 - 113.","BIRULA, A. A. 1917. Chlenistobryukhie paukoobraznye Kavkazskogo Kraya. Part I. Scorpiones. Zapiski Kavkazskogo Muzeya, 5: 1 - 253.","KOVARIK, F. 2000. The Socotra Island and its scorpions. Akvarium terarium, 43 (7): 63 - 67 (in Czech).","KOVARIK, F. 2004. Revision and taxonomic position of genera Afghanorthochirus Lourenco & Vachon, Baloorthochirus Kovarik, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kovarik, Pakistanorthochirus Lourenco, and Asian Orthochirus Karsch, with descriptions of twelve new species (Scorpiones, Buthidae). Euscorpius, 16: 1 - 33."]}
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33. Mesobuthus zarudnyi sp. n. from Azerbaijan (Scorpiones: Buthidae)
- Author
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Novruzov, Nizami E., Kovařík, František, and Fet, Victor
- Subjects
Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Novruzov, Nizami E., Kovařík, František, Fet, Victor (2022): Mesobuthus zarudnyi sp. n. from Azerbaijan (Scorpiones: Buthidae). Euscorpius 347: 1-9, DOI: http://doi.org/10.5281/zenodo.7162669, {"references":["(BIRULA, A. A.) BYALYNITSKII-BIRULYA, A. A. 1917. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskago Muzeya (Memoires du Musee du Caucase), Tiflis: Imprimerie de la Chancellerie du Comite pour la Transcaucasie, A(5), 253 pp. (in Russian; published August 1917) (in Russian). English translation: Byalynitskii-Birulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. 1. Scorpions. Jerusalem: Israel Program for Scientific Translations, 170 pp.","FET, V. 1989. A catalogue of scorpions (Chelicerata: Scorpiones) of the USSR. Rivista del Museo Civico di Scienze Naturali \"Enrico Caffi\", 13(1988): 73-171.","FET, V. & G. LOWE. 2000. Family Buthidae. Pp. 54-286 in: Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder (eds.). Catalog of the Scorpions of the World (1758-1998). New York: New York Entomological Society, 690 pp.","KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.","KOVARIK, F. 2019. Taxonomic reassessment of the genera Lychas, Mesobuthus, and Olivierus, with descriptions of four new genera (Scorpiones: Buthidae). Euscorpius, 288: 1-27.","KOVARIK, F. & V. FET. 2006. Taxonomic position of the genus Sassanidotus Farzanpay, 1987 (Scorpiones: Buthidae). Euscorpius, 39: 1-9.","KOVARIK, F., et al. 2022.A revision of the genus Mesobuthus Vachon, 1950, with a description of 14 new species (Scorpiones: Buthidae). Euscorpius, 348 (in press).","KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.","NOVRUZOV, N. E. 2017. [Notes on night activity and trophic behaviour of Mesobuthus eupeus (C.L. Koch, 1839) (Arachnida, Scorpiones, Buthidae) in the Eastern Azerbaijan]. Eurasian Entomological Journal, 16(1): 67-72 (in Russian).","NOVRUZOV, N. E. 2019. [Distribution and morphometric variability of Mesobuthus eupeus (C.L. Koch, 1839) (Arachnida, Scorpiones, Buthidae) in Eastern Azerbaijan]. Eurasian Entomological Journal, 18(1): 27-38 (in Russian).","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297-316.","VACHON, M. 1950. Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28(2): 152-216.","VACHON, M. 1952. Etudes sur les scorpions. Institut Pasteur d'Algerie, Alger, 1-482. (published 1948-1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25-90, 162-208, 288-316, 441-481. 1949, 27: 66-100, 134-169, 281-288, 334-396. 1950, 28: 152-216, 383- 413. 1951, 29: 46-104).","VACHON, M. 1958. Scorpionidea (Chelicerata) de l'Afghanistan. (The 3d Danish Expedition to Central Asia. Zoological Results 23). Videnskabelige Meddelelser fra Dansk Naturhistrorisk Forening i KObehavn, 120: 121-187.","VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3e ser. n°140, Zool. 140: 857-958.","VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus de l'Academie des Sciences, Paris, D, 281: 1597-1599.","YUSUBOV, E. B. 1978. [Scorpions of the Baku and Absheron archipelagoes]. Uchenye zapiski AGU. Seriya biologicheskaya, 3: 49-50 (in Russian).","YUSUBOV, E. B. 1985. Skorpiony (Arachnidae, Scorpiones) Azerbaijana [Scorpions of Azerbaijan]. Avtoreferat [PhD digest]. Leningrad: Leningrad State University, 20 pp. (in Russian)."]}
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34. Hottentotta polystictus
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Kovařík, František and Lowe, Graeme
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Hottentotta ,Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Hottentotta polystictus ,Taxonomy - Abstract
Hottentotta polystictus (Pocock, 1896) (Figures 63–70, 113) Buthus polystictus Pocock, 1896: 178. Hottentotta polystictus: Kovařík & Ojanguren, 2013: 171–172, 318, 338–339, figs. 1069–1072, 1206–1216 (complete reference list until 2013); Kovařík & Mazuch, 2015: 23, figs. 112–131, table 4; Kovařík & Lowe, 2021: 3–9, figs. 1–45, 137–138, 140–143, table 1. TYPE LOCALITY AND TYPE REPOSITORY. Somalia, Goolis Mountains, inland of Berbera; BMNH. TYPE MATERIAL EXAMINED. Somaliland, Goolis Mountains, inland of Berbera, 2♀ 1im. (holotype and paratypes, fig. 84 in Kovařík, 2007: 55), leg. E. Lort Phillips, BMNH No. 1895.6.1.46-7. DJIBOUTI MATERIAL EXAMINED (FKCP). Djibouti, Arta Province, Arta, 11.5286°N 42.8508°E, 690 m a. s. l., 16.I.2014, 1juv. (No. 8622), leg. J. Lips; Djibouti Province, Goubetto, 11.4182°N 42.9059°E, 560 m a. s. l., 26.I.2022, 1♀ (No. 25588), leg. J. Lips; Tadjourah Province, Day, 11.7534°N 42.6560°E, 1680 m a. s. l., 30.I.2022, 1♀ 1juv. (No. 25607), leg. J. Lips; Tadjourah Province, Medeho, 11.9086°N 43.1356°E, 30 m a. s. l., 20.1.2022, 1♀ (Figs. 63–70, No. 25619), leg. J. Lips; Tadjourah Province, Randa, 11.8469°N 42.6578°E, 910 m a. s. l., 31.I.2022, 1juv. (No. 25584), leg. J. Lips; Barra Yer (Petit Barre), 11.31°N 42.71°E (11°18'33.56"N 42°42'39.17"E), 585 m a. s. l., I. 2017, 3juvs., leg. R. Štarha. DISTRIBUTION. Djibouti, Ethiopia, Somaliland (Fig. 113; Kovařík & Lowe, 2021: 25, fig. 138)., Published as part of Kovařík, František & Lowe, Graeme, 2022, Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXVIII Scorpions of Djibouti, pp. 1-31 in Euscorpius 357 on page 5, DOI: 10.5281/zenodo.7163285, {"references":["POCOCK, R. I. 1896. Report upon the scorpions, spiders, centipedes, and millipedes obtained by Mr. and Mrs. E. Lord Philips in the Goolis Mountains inland of Berbera, N. Somaliland. Annals and Magazine of Natural History, 6 (18): 178 - 186."]}
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35. Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXVIII Scorpions of Djibouti
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Kovařík, František and Lowe, Graeme
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Arthropoda ,Arachnida ,Scorpiones ,Hemiscorpiidae ,Animalia ,Biodiversity ,Buthidae ,Scorpionidae ,Taxonomy - Abstract
Kovařík, František, Lowe, Graeme (2022): Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXVIII Scorpions of Djibouti. Euscorpius 357: 1-31, {"references":["LOURENCO, W. R., B. DUHEM & J. L. CLOUDSLEY- THOMPSON. 2012. Scorpions frm Ennedi, Kapka and Tibesti, the mountains of Chad, with descriptions of nine new species (Scorpiones: Buthidae, Scorpionidae). Arthropoda Selecta, 21 (4): 307-338.","LOURENCO, W. R. & E-A. LEGUIN. 2011. Further considerations on the species of the genus Orthochirus Karsch, 1891 from Africa, with description of three new species (Scorpiones: Buthidae). Euscorpius, 123: 1-19.","LOURENCO, W. R. & J.-X. QI. 2006. Further considerations on the genus Butheolus Simon, 1882 and description of one new species from Saudi Arabia (Scorpiones, Buthidae). Zoology in the Middle East, 37: 91-97.","LOURENCO, W. R. & E. A. SADINE. 2021. The genus Orthochirus Karsch, 1891, in Algeria with description of a new species (Scorpiones, Buthidae). Bulletin de la Societe entomologique de France, 126 (2): 175-181.","LOURENCO W. R. & YTHIER E. 2021. A particular new species of Orthochirus Karsch, 1891 from Somalia (Scorpiones: Buthidae). Serket, 17 (4): 335-349.","LOWE, G. 2010. New picobuthoid scorpions (Scorpiones: Buthidae) from Oman. Euscorpius, 93: 1-53.","LOWE, G., & F. KOVARIK. 2019. Review of Grosphus Simon, 1880, with description of Teruelius gen. n., a new buthid genus from Madagascar (Scorpiones: Buthidae). Euscorpius, 281: 1-128","LOWE, G., & F. KOVARIK. 2022. Reanalysis of Teruelius and Grosphus (Scorpiones: Buthidae) with descriptions of two new species. Euscorpius, 356: 1-105.","LOWE, G., F. KOVARIK, M. STOCKMANN & F. STAHLAVSKY 2018. Review of Microbuthus (Scorpiones, Buthidae) from Oman and Yemen. Euscorpius, 263: 1-22.","MAGIN, C. & C. BURDETTE. 2013-2020. Eastern Africa: Somalia. WWF Ecoregions. https://www.worldwildlife. org/ecoregions/at1307 (accessed 16 January, 2022); offline May 2022, archived at: https://web.archive.org/ web/20220501000000*/https://www.worldwildlife.org/ ecoregions/at1307 (accessed 24 July 2022)","PAVESI, P. 1885. Aracnidi raccolti dal conte Bouturlin ad Assab e Massaua. Bolletino della Societa Entomologica Italiana, 17: 197-200.","POCOCK, R. I. 1896. Report upon the scorpions, spiders, centipedes, and millipedes obtained by Mr. and Mrs. E. Lord Philips in the Goolis Mountains inland of Berbera, N. Somaliland. Annals and Magazine of Natural History, 6(18): 178-186.","POCOCK, R. I. 1901. On a new species of the genus Parabuthus. Bollettino dei Musei di Zoologia ed Anatomia Comparata, 16(382): 1.","ROSSI, A. 2014a. Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic and Djibouti (Scorpiones: Scorpionidae). Onychium, 10(2013): 10-31.","ROSSI, A. 2014b. The fragmented peri-Saharan distribution of the subgenus Pandinurus Fet, 1997 with the description of a new species from Chad (Scorpiones, Scorpionidae, Pandinus). Serket, 14 (1): 6-14.","ROSSI,A. 2014c. New data on the rare species Pandinus nistriae Rossi, 2014 (Scorpiones: Scorpionidae). Arachnides, 72: 3-12.","ROSSI, A. 2015a. Sui sottogeneri di Pandinus Thorell, 1876 con revisione del genere Pandinurus Fet, 1997 stat. n. e descrizione di sette nuove specie e tre nuovi sottogeneri (Scorpiones: Scorpionidae). Onychium, 11: 10-66.","ROSSI, A. 2015b. Due ulteriori nuove specie del sottogenere Pandiborellius Rossi, 2015 dal Corno d'Africa (Scorpiones, Scorpionidae). Arachnida, Rivista Aracnologica Italiana, 2: 2-36.","ROSSI, A. 2015c. Ulteriori commenti sulla tassonomia dei generi Pandinus e Pandinurus, con la definizione di un nuovo sottogenere ed una nueva specie di Pandinurus dalla Somalia (Scorpiones: Scorpionidae). Arachnida, Rivista Aracnologica Italiana, 4: 41-55.","ROSSI, A. 2017. Complementi alla fauna del Corno d'Africa: famiglia Buthidae C. L. Koch, 1837 (Scorpiones), con la descrizione di tre nuove specie. Arachnida - Rivista Aracnologica Italiana, 9: 2-11 (2016).","SCHMITT, T., U. FRITZ, M. DELFINO, W. ULRICH & J. C. HABEL. 2021. Biogeography of Italy revisited: genetic lineages confirm major phylogeographic patterns and a pre-Pleistocene origin of its biota. Frontiers in Zoology, (2021) 18:34. https://doi.org/10.1186/s12983-021- 00418-9","SHULOV, A. & P. AMITAI. 1960. Observations sur les scorpions Orthochirus innesi E. Simon 1910, ssp. negebensis nov. Archives de l'Institut Pasteur de l'Algerie, 38 (1): 117-129.","SISSOM, W. D. 1994. Descriptions of new and poorly known scorpions of Yemen (Scorpiones: Buthidae, Diplocentridae, Scorpionidae). Fauna of Saudi Arabia, 14: 3-39.","SISSOM, W. D., G. A. POLIS & D. D. WATT. 1990. Field and laboratory methods. Pp. 445-461 in POLIS, G.A. (ed.). The Biology of Scorpions. Stanford University Press, Stanford, CA.","SOLEGLAD, M. E. & W. D. SISSOM. 2001. Phylogeny of the family Euscorpiidae Laurie, 1896: a major revision. Pp. 25-112 in: Fet, V. & P. A. Selden (eds.). Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks: British Arachnological Society.","SOUSA P., M. A. ARNEDO & D. J. HARRIS. , 2017. Updated catalogue and taxonomic notes on the Old- World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae). ZooKeys, 686: 15-84.","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81(12): 297-316.","VACHON, M. 1952. Etudes sur les scorpions. Institut Pasteur d'Algerie, Alger, 1-482. (published 1948-1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25-90, 162-208, 288-316, 441-481. 1949, 27: 66-100, 134-169, 281-288, 334-396. 1950, 28: 152-216, 383- 413. 1951, 29: 46-104).","VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en Arachnologie, Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum National d´Histoire Naturelle Paris, 140: 857-958.","VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus Hebdomadaires des Seances de l´Academie des Sciences, Paris, ser. D, 281: 1597-1599.","WERNER, F. 1916.Uber einige Skorpione und Gliederspinnen des Naturhistorischen Museum in Wiesbaden. Jahrbucher des Nassauischen Verein fur Naturkunde, 69: 79-97.","ZAMBRE, A. M., Z. A. MIRZA, R. V. SANAP, R. UPADHYE & S. M. MAQSOOD JAVED. 2011. A new species of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) from Maharashtra, India. Euscorpius, 107: 1-12."]}
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36. Gint gaitako Kovarik, Lowe, Pliskova & Stahlavsky 2013
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Just, Pavel, Šťáhlavský, František, Kovařík, František, and Štundlová, Jana
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Gint gaitako ,Gint ,Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
GINT GAITAKO All specimens exhibit 2 n = 30 chromosomes, which decrease in length from 4.64% to 1.80% of DSL (Supporting Information, Fig. S2A; Table S2). The post-pachytene cells show 13 bivalents and one quadrivalent (chromosomes 1, 18, 25 and 30) (Fig. 2M; Supporting Information, Fig. S2A). The pair of 18S rDNA sites is located in the terminal region of the third largest bivalent (chromosomes 6 and 7) (Fig. 2N; Supporting Information, Fig. S2A)., Published as part of Just, Pavel, Šťáhlavský, František, Kovařík, František & Štundlová, Jana, 2022, Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae), pp. 885-901 in Zoological Journal of the Linnean Society 196 (2) on page 891, DOI: 10.1093/zoolinnean/zlac049, http://zenodo.org/record/7184492
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37. Gint gubanensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky 2018
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Just, Pavel, Šťáhlavský, František, Kovařík, František, and Štundlová, Jana
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Gint ,Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Gint gubanensis ,Taxonomy - Abstract
GINT GUBANENSIS Karyotype of G. gubanensis comprises 2 n = 45 chromosomes, which decrease in length from 2.97% to 1.02% of DSL (Supporting Information, Fig. S2G; Table S2). Overall, 21 bivalents and a single trivalent, composed of chromosomes 7, 33 and 45, are detected in post-pachytene nuclei (Fig. 3I; Supporting Information, Fig. S2G). One pair of 18S rDNA loci is found on a terminal part of the heteromorphic chromosome pair (chromosomes 26 and 44). A single large 18S rDNA cluster is located on the overhanging part of chromosome 26 (the larger chromosome of the heteromorphic pair) (Fig. 3J; Supporting Information, Fig. S2G). MTDNA DIVERSITY AND PHYLOGENY The sequences for both 16S and COI were obtained from 22 Gint individuals. In G. calviceps, only the 16S gene fragment was successfully sequenced. The final alignment with a total length of 963 bp (16S: 358 bp; COI: 605 bp) consisted of 765 conserved sites, 194 variable sites and 172 parsimony-informative sites. Based on COI data, Gint exhibits genetic variation at both inter- and intraspecific levels, with the pairwise genetic distances ranging from 3.28% to 11.52% between species, and from 0.00% to 2.64% within species (Supporting Information, Tables S3, S4). The haplotype network reconstruction showed a presence of 15 COI sequence haplotypes in Gint (Supporting Information, (Fig. S3; Table S3). Within the species studied, G. banfasae shows the highest within-location haplotype diversity (four haplotypes per locality), but with minor pairwise genetic differences (p -distance: ≤0.66%). In this species, all individuals of the same cytotype have distinct haplotypes. The haplotypes of cytotype I individuals (S1531 and S1534) are separated from one another by four mutational steps. Cytotype II (S1532) shares its haplotype with the cytotype III individual (S1530), which is one mutational step distant from the haplotype of the cytotype III specimen (S1533) (Supporting Information, Fig. S3). Gint dabakalo, comprising three haplotypes, shows the highest intraspecific genetic variation with a p -distance of 0.17–2.64%. In this species, the genetic distance between individuals of cytotype I and cytotype II from the same locality is higher than between each of them and a specimen of cytotype III from a distinct locality (Supporting Information, Fig. S3; Table S1). Gint maidensis, a species with a stable karyotype 2 n = 34, is represented by three haplotypes with a p -distance of 0.33–0.50%. Gintgaitako presentstwohaplotypes, where one haplogroup, formed by five specimens, is distant from the other by one mutational step (p -distance: 0.17). In G. amoudensis, individuals of cytotype II, cytotype III and cytotype IV share the same haplotype, whereas the specimen of cytotype I differs from this haplogroup by eight point mutations (p -distance: 1.49%). Both BI and ML mtDNA analyses provide similar tree topologies and nodal supports for clades. Phylogenetic reconstruction depicts Gint as a monophyletic group (Fig. 5). Most of the Gint species form well-supported clades [bootstrap support (bs) ≥ 83, posterior probability (PP) ≥ 0.98]. The monophyly of the G. banfasae specimens remains unresolved due to low support values. Gint maidensis forms an early-branching clade, which is sister to the clade comprising all of the remaining Gint species. This clade is further subdivided into two groups: (1) a clade in which G. gubanensis is a sister-species to G. amoudensis, and (2) a clade consisting of G. gaitako, G. calviceps, G. dabakalo and G. banfasae. In the latter, G. gaitako forms a sister-lineage to the clade comprising G. calviceps, G. dabakalo and G. banfasae. The mutual relationships between the species in this group remain unresolved due to low support values. THE RECONSTRUCTION OF ANCESTRAL CHROMOSOME NUMBER AND POSITION OF 18S RDNA GENE CLUSTERS Maximum parsimony analysis inferred 2 n = 32–34 as the ancestral chromosome number for the genus Gint (Fig. 6A). Two independent increases of 2 n were detected within Gint: (1) the clade containing all cytotypes of G. amoudensis and G. gubanensis, and (2) cytotype III within the G. dabakalo clade. When compared with the ancestral 2 n, a tendency of decreasing of 2 n from 30 to 18 chromosomes can be observed in G. gaitako, G. dabakalo and G. banfasae. Concerning the position of 18S rRNA gene clusters, maximum parsimony inferred their terminal position as the ancestral state for the genus Gint (Fig. 6B). Two independent changes of the ancestral terminal position to a subterminal position were identified in G. amoudensis and in the clade containing G. dabakalo and G. banfasae., Published as part of Just, Pavel, Šťáhlavský, František, Kovařík, František & Štundlová, Jana, 2022, Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae), pp. 885-901 in Zoological Journal of the Linnean Society 196 (2) on page 893, DOI: 10.1093/zoolinnean/zlac049, http://zenodo.org/record/7184492
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38. Gint dabakalo Kovarik & Mazuch 2015
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Just, Pavel, Šťáhlavský, František, Kovařík, František, and Štundlová, Jana
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Gint ,Arthropoda ,Arachnida ,Gint dabakalo ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
GINT DABAKALO In this species, the differences in the diploid number, multivalent associations and the appearance of 18S rDNA-bearing chromosomes are detected at inter- and intrapopulation levels, resulting in three distinguished cytotypes. Specimen S 1305 from the locality Burao is assigned to cytotype I, and specimens S1527 and S1199 from N of Burao, Togdheer represent cytotypes II and III, respectively. Cytotype I (1 Ƌ; S1305) has 2 n = 23 chromosomes, which decrease in length from 7.90% to 2.55% of DSL (Supporting Information, Fig. S1D; Table S2). The post-pachytene cells show nine bivalents and one pentavalent that is composed of chromosomes 1, 4, 5, 10 and 23 (Fig. 2G; Supporting Information, Fig. S1D). The pair of 18S rDNA sites is located in the subterminal region of the largest bivalent (Fig. 2H). Cytotype II (1 Ƌ; S1527) exhibits 2 n = 24 chromosomes, which decrease in length from 7.65% to 2.95% of DSL (Supporting Information, Fig. S1E; Table S2). The post-pachytene nuclei show a configuration of eight bivalents, one trivalent (chromosomes 2, 12 and 17) and one pentavalent (chromosomes 1, 3, 4, 7 and 20) (Fig. 2I; Supporting Information, Fig. S1E). The pair of 18S rDNA sites is situated in the subterminal region of chromosomes 2 and 12 of the trivalent (Fig. 2J; Supporting Information, Fig. S1E). Cytotype III (1 Ƌ; S1199) has 2 n = 27 chromosomes, which decrease in length from 8.30% to 1.80% of DSL (Supporting Information, Fig. S1F; Table S2). In the post-pachytene spermatocytes, we observe eight bivalents, two trivalents and one pentavalent (Fig. 2K). The larger trivalent is composed of chromosomes 2, 9 and 10, the smaller trivalent is composed of chromosomes 4, 22 and 25, and chromosomes 1, 3, 5, 8 and 17 form a pentavalent (Supporting Information, Fig. S1F). The 18S rDNA sites are located in the subterminal region of the chromosomes 2 and 9 of the larger trivalent (Fig. 2L; Supporting Information, Fig. S1F)., Published as part of Just, Pavel, Šťáhlavský, František, Kovařík, František & Štundlová, Jana, 2022, Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae), pp. 885-901 in Zoological Journal of the Linnean Society 196 (2) on pages 890-891, DOI: 10.1093/zoolinnean/zlac049, http://zenodo.org/record/7184492
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39. Gint banfasae Kovarik & Lowe 2019
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Just, Pavel, Šťáhlavský, František, Kovařík, František, and Štundlová, Jana
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Gint ,Arthropoda ,Arachnida ,Scorpiones ,Gint banfasae ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
GINT BANFASAE In this species, we detected intrapopulation variability in diploid number and/or multivalent associations showing the presence of three cytotypes. The position of a single pair of 18S rDNA sites, as well as the 18S rDNA-bearing chromosome pair involved in the multivalent chain, are consistent in all cytotypes. Cytotype I (2 ƋƋ; S1531, S1534) has 2 n = 18 chromosomes, which decrease in length from 9.85% to 1.81% of DSL (Supporting Information, Fig. S1A; Table S2). In post-pachytene nuclei, the chromosomes form seven bivalents and one quadrivalent that is composed of the four larger chromosomes (Fig. 2A). A single pair of 18S rDNA sites is located in the subterminal region of the chromosomes 3 and 4 involved in the quadrivalent (Fig. 2B; Supporting Information, Fig. S1A). Cytotype II (1 Ƌ; S1532) exhibits 2 n = 18 chromosomes, which decrease in length from 9.79% to 1.77% of DSL (Supporting Information, Fig. S1B; Table S2). The post-pachytene cells show six bivalents and one hexavalent (chromosomes 1, 2, 3, 4, 15 and 18) (Fig. 2C; Supporting Information, Fig. S1B). A pair of 18S rDNA sites is located in the subterminal region of the chromosomes 3 and 4 involved in the hexavalent (Fig. 2D). Cytotype III (2 ƋƋ; S1530, S1533) has 2 n = 19 chromosomes, which decrease in length from 9.66% to 1.78% of DSL (Fig. 2E; Supporting Information, Fig. S1C; Table S2). Post-pachytene spermatocytes exhibit five bivalents, one trivalent (chromosomes 7, 17 and 18) and one hexavalent (chromosomes 1, 2, 3, 4, 14 and 19). A pair of 18S rDNA sites is situated in the subterminal region of the chromosomes 3 and 4 involved in the hexavalent (Fig. 2F)., Published as part of Just, Pavel, Šťáhlavský, František, Kovařík, František & Štundlová, Jana, 2022, Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae), pp. 885-901 in Zoological Journal of the Linnean Society 196 (2) on page 889, DOI: 10.1093/zoolinnean/zlac049, http://zenodo.org/record/7184492
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40. Gint maidensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky 2018
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Just, Pavel, Šťáhlavský, František, Kovařík, František, and Štundlová, Jana
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Gint ,Arthropoda ,Gint maidensis ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
GINT MAIDENSIS The karyotype of G. maidensis comprises 2 n = 34 chromosomes, which decrease in length from 4.22% to 1.87% of DSL (Supporting Information, Fig. S2B; Table S2). The post-pachytene nuclei exhibit 17 bivalents (Fig. 2O). No multivalent associations are found in this species. One pair of 18S rDNA sites is located in the terminal region of the largest bivalent (Fig. 2P)., Published as part of Just, Pavel, Šťáhlavský, František, Kovařík, František & Štundlová, Jana, 2022, Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae), pp. 885-901 in Zoological Journal of the Linnean Society 196 (2) on page 892, DOI: 10.1093/zoolinnean/zlac049, http://zenodo.org/record/7184492
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- 2022
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41. Gint amoudensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky 2018
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Just, Pavel, Šťáhlavský, František, Kovařík, František, and Štundlová, Jana
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Gint ,Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy ,Gint amoudensis - Abstract
GINT AMOUDENSIS T h e m o s t s t r i k i n g i n t r a s p e c i f i c v a r i a b i l i t y i s discovered in G. amoudensis, which exhibits four cytotypes differing in the diploid number, multivalent associations and in the number and position of 18S rDNA sites. The cytotypes II, III and IV occur sympatrically, while the cytotype I (S1325) is found in a remote locality Borama c. 150 km far from the other cytotypes. Cytotype I (1Ƌ; S1325) exhibits 2 n = 36 chromosomes, which decrease in length from 4.49% to 1.63% of DSL (Supporting Information, Fig. S2C; Table S2). The post-pachytene nuclei show 16 bivalents and one quadrivalent (chromosomes 1, 2, 9 and 15) (Fig. 3A; Supporting Information, Fig. S2C). Three 18S rDNA sites are detected (Fig. 3B), two of them are located in the subterminal region of the chromosomes 1 and 2 involved in the quadrivalent and one in the terminal position of chromosome 2 in the quadrivalent (Supporting Information, Fig. S2C). Cytotype II (1 Ƌ; S1291) has 2 n = 35 chromosomes, which decrease in length from 5.01% to 1.45% of DSL (Supporting Information, Fig. S2D; Table S2). The post-pachytene spermatocytes exhibit 14 bivalents, one trivalent (chromosomes 4, 24 and 35) and one quadrivalent (chromosomes 1, 5, 8 and 23) (Fig. 3C; Supporting Information, Fig. S2D). Three 18S rDNA loci are present (Fig. 3D), one pair is located on the chromosomes 1 (subterminal site) and 5 (terminal site) involved in the quadrivalent, and one single heterozygous locus is situated in the interstitial region of the third largest bivalent (chromosome 9; Fig. 3D; Supporting Information, Fig. S2D). Cytotype III (1 Ƌ; S1293) has 2 n = 36 chromosomes, which decrease in length from 4.71% to 1.52% of DSL (Supporting Information, Fig. S2E; Table S2). The post-pachytene cells show 15 bivalents and one hexavalent (chromosomes 1, 2, 5, 6, 11 and 24) (Fig. 3E; Supporting Information, Fig. S2E). One pair of 18S rDNA sites is located in the subterminal region of the chromosomes 1 and 2 of the hexavalent (Fig. 3F; Supporting Information, Fig. S2E). Cytotype IV (1 Ƌ; S1292) exhibits 2 n = 35 chromosomes, which decrease in length from 4.66% to 1.54% of DSL (Supporting Information, Fig. S2F; Table S2). The post-pachytene nuclei show a configuration of 13 bivalents, one trivalent (chromosomes 7, 26 and 35) and one hexavalent (chromosomes 1, 4, 5, 6, 10 and 23) (Fig. 3G; Supporting Information, Fig. S2F). One pair of 18S rDNA sites is located in the subterminal region of the chromosomes 1 and 4 involved in the hexavalent (Fig. 3H; Supporting Information, Fig. S2F)., Published as part of Just, Pavel, Šťáhlavský, František, Kovařík, František & Štundlová, Jana, 2022, Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae), pp. 885-901 in Zoological Journal of the Linnean Society 196 (2) on pages 892-893, DOI: 10.1093/zoolinnean/zlac049, http://zenodo.org/record/7184492
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42. Euscorpius sulfur sp. n. (Scorpiones: Euscorpiidae), a new cave scorpion from Albania and northwestern Greece.
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Kovařík, František, Audy, Marek, Sarbu, Serban M., and Fet, Victor
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SCORPIONS , *SULFUR , *SPECIES , *PHOTOGRAPHS , *HABITATS , *CAVES , *BRONZE - Abstract
A new species of cave scorpion, Euscorpius sulfur sp. n. (Scorpiones: Euscorpiidae) from Albania and northwestern Greece is described, fully complemented with color photographs of both live and preserved specimens, as well as their habitats. [ABSTRACT FROM AUTHOR]
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- 2023
43. Two new species of the remarkable scorpion genus Megacormus Karsch, 1881 (Scorpiones: Euscorpiidae).
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Teruel, Rolando, Kovařík, František, Lowe, Graeme, and Šťáhlavský, František
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SCORPIONS , *SPECIES , *ADULTS - Abstract
Two new species of the very interesting euscorpiid scorpion genus Megacormus Karsch, 1881, are herein described from eastern Mexico. Both had been previously misidentified in the published literature, one as Megacormus segmentatus Pocock, 1900, the other as Megacormus granosus (Gervais, 1844), but the direct comparison of adults of both sexes to adult topotypes from all members of the genus currently accepted as valid taxa, revealed solid morphological characters that warrant their recognition as distinct taxa. Both new species seem also to be endemics with a small distribution area, but apparently not restricted to a single locality. This addition raises the known composition of the genus to eight species, all of them exclusively from mountain sites of eastern Mexico. [ABSTRACT FROM AUTHOR]
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- 2023
44. Molecular data do not support the traditional morphology-based groupings in the scorpion family Buthidae (Arachnida: Scorpiones)
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Štundlová, Jana, primary, Šťáhlavský, František, additional, Opatova, Vera, additional, Stundl, Jan, additional, Kovařík, František, additional, Dolejš, Petr, additional, and Šmíd, Jiří, additional
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- 2022
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45. Tracking the trends of karyotype differentiation in the phylogenetic context of Gint, a scorpion genus endemic to the Horn of Africa (Scorpiones: Buthidae)
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Just, Pavel, primary, Šťáhlavský, František, additional, Kovařík, František, additional, and Štundlová, Jana, additional
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- 2022
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46. Scorpions of the Horn of Africa (Arachnida: Scorpiones) Part XXVII Lanzatus huluul sp n from Somaliland (Buthidae)
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Kovařík, František and Lowe, Graeme
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Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kovařík, František, Lowe, Graeme (2021): Scorpions of the Horn of Africa (Arachnida: Scorpiones) Part XXVII Lanzatus huluul sp n from Somaliland (Buthidae). Euscorpius 344: 1-12, DOI: http://doi.org/10.5281/zenodo.7162582, {"references":["DUPRE, G. 2007. Conspectus genericus scorpionorum 1758- 2006 (Arachnida: Scorpiones). Euscorpius, 50: 1-31.","FET, V. & M. E. SOLEGLAD 2005. Contributions to scorpion sYstematics. I. On recent changes in high-level taxonomY. Euscorpius, 31: 1-13.","FET, V., M. E. SOLEGLAD & G. LOWE. 2005. A new trichobothrial character for the high-level sYstematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1-40.","KOVARIK, F. 2001. Lanzatus somalicus gen. n. et sp. n. (Scorpiones: Buthidae) from Somalia. Acta Societas Zoologicae Bohemicae, 65 (1): 41-44.","KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp.","KOVARIK, F. 2018. Notes on the genera Buthacus, Compsobuthus, and Lanzatus with several sYnonYmies and corrections of published characters (Scorpiones: Buthidae). Euscorpius, 269: 1-12.","KOVARIK, F. & G. LOWE. 2019. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVIII. Gint banfasae sp. n. from Somaliland (Buthidae). Euscorpius, 272: 1-19.","KOVARIK, F., G. LOWE, H. SH. A. ELMI & F. STAHLAVSKY. 2019. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXI. Parabuthus (Buthidae) (Part II), with description of five new species from Somaliland and Ethiopia. Euscorpius, 290: 1-63.","KOVARIK, F., G. LOWE, P. JUST, A.I. AWALE, H.S.A. ELMI & F. STAHLAVSKY. 2018. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XV. Review of the genus Gint Kovarik et al., 2013, with description of three new species from Somaliland (Scorpiones, Buthidae). Euscorpius, 259: 1-41.","KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Euscorpius, 232: 1-38.","KOVARIK, F. & L. NJOROGE. 2021. Somalibuthus sabae sp. n., a new buthid scorpion from KenYa (Scorpiones: Buthidae). Euscorpius, 332: 1-19.","KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, LYchas, and Sassanidotus. Prague: Clairon Production, 400 pp.","KOVARIK, F., M. E. SOLEGLAD & V. FET. 2007. A new species of scorpion in the \"Charmus\" group from India (Scorpiones: Buthidae). Boletin Sociedad Entomologica Aragonesa, 40: 201-209.","LOURENCO, W. R. 2001a. Un nouveau genre de Buthidae, probable vicariant geographique d' Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77(1): 15-20.","LOURENCO, W. R. 2001b. Taxonomic considerations on the genera Butheolus Simon, Nanobuthus Pocock and Neobuthus Hirst (Scorpions, Buthidae) with the description of a new species of Neobuthus from Ethiopia. Ecology of Desert Environments, 2001: 171-183.","PRENDINI, L. & W. C. WHEELER 2005. Scorpion higher phYlogenY and classification, taxonomic anarchY, and standards for peer review in online publishing. Cladistics, 21: 446-494.","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297-316.","VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et tYpes de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857-958.","VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Ser.D Sciences Naturelles, 281 (21): 1597-1599.","WILLIAMS, S.C. 1969. A new species of Syntropis from Baja California Sur, Mexico with notes on its biologY. The Pan-Pacific Entomologist, 45 (4): 285-291."]}
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47. Comments on the establishment of the new name Liochelidae Fet & Bechly, 2001 (Archnida, Scorpiones) as a substitute for Ischnuridae Simon, 1879. (Case 3120a; see BZN 58: 280-281)
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Lourenco, Wilson R, Kovařík, František, and BioStor
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- 2002
48. A new species of Pseudouroctonus from the Pinaleño Mountains southern Arizona (Scorpiones Vaejovidae)
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Ayrey, Richard F., Kovařík, František, and Myers, Brandon T.
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Arthropoda ,Vaejovidae ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Taxonomy - Abstract
Ayrey, Richard F., Kovařík, František, Myers, Brandon T. (2021): A new species of Pseudouroctonus from the Pinaleño Mountains southern Arizona (Scorpiones Vaejovidae). Euscorpius 338: 1-12, DOI: http://doi.org/10.5281/zenodo.5742153
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- 2021
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49. Somalibuthus sabae sp. n., a new buthid scorpion from Kenya (Scorpiones: Buthidae)
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Kovařík, František and Njoroge, Laban
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Arthropoda ,Arachnida ,Scorpiones ,Animalia ,Biodiversity ,Buthidae ,Taxonomy - Abstract
Kova����k, Franti��ek, Njoroge, Laban (2021): Somalibuthus sabae sp. n., a new buthid scorpion from Kenya (Scorpiones: Buthidae). Euscorpius 332: 1-19, DOI: http://doi.org/10.5281/zenodo.5742065, {"references":["FET, V., M. E. SOLEGLAD & G. LOWE. 2005. A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1-40.","HARADON, R. M. 1984. New and redefined species belonging to the Paruroctonus baergi group (Scorpiones, Vaejovidae). Journal of Arachnology, 12: 205-221.","KOVARIK, F. 1998. Three new genera and species of scorpions (Buthidae) from Somalia. Acta Societas Zoologicae Bohemoslovenicae, 62: 115-124.","KOVARIK, F. 2001. Lanzatus somalicus gen. n. et sp. n. (Scorpiones: Buthidae) from Somalia. Acta Societas Zoologicae Bohemoslovenicae, 65 (1): 41-44.","KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp.","KOVARIK, F. 2019. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXII. Two new species of Neobuthus from Somaliland (Buthidae). Euscorpius, 294: 1-16.","KOVARIK, F. & G. LOWE. 2012. Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae). Euscorpius, 138: 1-25.","KOVARIK, F. & G. LOWE. 2019. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVIII. Gint banfasae sp. n. from Somaliland (Buthidae). Euscorpius, 272: 1-19.","KOVARIK, F., G. LOWE, A.I. AWALE, H.S.A. ELMI & A.A. HURRE. 2018b. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae). Euscorpius, 271: 1-81.","KOVARIK, F., G. LOWE, P. JUST, A.I. AWALE, H.S.A. ELMI & F. STAHLAVSKY. 2018a. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XV. Review of the genus Gint Kovarik et al., 2013, with description of three new species from Somaliland (Scorpiones, Buthidae). Euscorpius, 259: 1-41.","KOVARIK, F., G. LOWE, J. PLISKOVA& F. STAHLAVSKY. 2013. A new scorpion genus, Gint gen. n., from the Horn of Africa (Scorpiones, Buthidae). Euscorpius, 173: 1-19.","KOVARIK, F., G. LOWE, J. PLISKOVA& F. STAHLAVSKY. 2016a. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n. Euscorpius, 209: 1-21.","KOVARIK, F., G. LOWE, K. B. RANAWANA, D.HOFEREK, V. A. SANJEEWA JAYARATHNE, J.PLISKOVA & F. STAHLAVSKY. 2016b. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n. Euscorpius, 220: 1-133.","KOVARIK, F., G. LOWE & F. STAHLAVSKY. 2016c. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Euscorpius, 232: 1-38.","KOVARIK, F. & T. MAZUCH. 2015. Review of the genus Gint Kovarik et al., 2013, with description of two new species from Somaliland and Somalia (Puntland) (Scorpiones: Buthidae). Euscorpius, 209: 1-23.","KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.","LORIA, S.F. & L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS ONE 9(12): e112913. doi:10.1371/journal.pone.0112913.","LOWE, G. 2018. The genera Butheolus Simon, 1882 and Xenobuthus gen. n. (Scorpiones: Buthidae) in Oman. Euscorpius, 261: 1-73.","LOWE, G. & F. KOVARIK. 2016. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part V. Two new species of Neobuthus Hirst, 1911 (Buthidae), from Ethiopia and Eritrea. Euscorpius, 224: 1-46.","LOWE, G. & F. KOVARIK. 2019. Review of Grosphus Simon, 1880, with description of Teruelius gen. n., a new buthid genus from Madagascar (Scorpiones: Buthidae). Euscorpius, 281: 1-128.","LOWE, G. & F. KOVARIK. 2021. Fetilinia dentator sp. n. from Pakistan (Scorpiones: Buthidae). Euscorpius, 328: 1-10.","LOWE, G., F. KOVARIK, M. STOCKMANN & F. STAHLAVSKY. 2018. Review of Microbuthus with Description of M. satyrus sp. n. (Scorpiones, Buthidae) from Oman and Yemen. Euscorpius, 263: 1-22.","LOWE, G., F. KOVARIK, M. STOCKMANN & F. STAHLAVSKY. 2019. Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae). Euscorpius, 277: 1-30.","LOWE, G., E. A. YAGMUR & F. KOVARIK. 2014. A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula. Euscorpius, 191: 1-129.","PRENDINI, L. 2003. Discovery of the male of Parabuthus muelleri, and implications for the phylogeny of Parabuthus (Scorpiones: Buthidae). American Museum Novitates, 3408: 1-24.","SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64-160 in POLIS, G. A. (Ed.) The Biology of Scorpions. Stanford University Press, Stanford, California.","SOLEGLAD, M. E. & V. FET. 2001. High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni). Euscorpius, 11: 1-175.","SOLEGLAD, M. E. & V. FET. 2003. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1-34.","STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297-316.","STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121-133.","VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161-166.","VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857-958.","VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Ser.D Sciences Naturelles, 281 (21): 1597-1599.","VOLSCHENK, E. S. 2005. A new technique for examining surface morphosculpture of scorpions. Journal of Arachnology, 33: 820-825."]}
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- 2021
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50. Three new species of Olivierus (Scorpiones: Buthidae) from Kazakhstan and Uzbekistan
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FET, VICTOR, primary, KOVAŘÍK, FRANTIŠEK, additional, GANTENBEIN, BENJAMIN, additional, and GRAHAM, MATTHEW R., additional
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- 2021
- Full Text
- View/download PDF
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