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Acanthotingis deltoides, sp. nov. (Figs. 1A–D, 2B, 4) urn:lsid:zoobank.org:act: A15AB0FC-61BF-4DF5-BEF8-878509EFDA11 Description. Head with five spines; frontal spines erect, short; occipital spines extremely long, porrect, nearly as long as first antennal segment, down-curved; medial spine long, porrect, nearly as long as occipital spines. Eyes small, red. Antennae moderately long; scape (segment I) infuscate, slender at base, greatly expanded on middle third, then constricted on apical third; pedicel (segment II) tan, shorter, one third length of scape; basiflagellomere (segment III) tan, slender, extremely long, more than four times longer than scape; distiflagellomere (segment IV) short, clavate, covered with fine hairs, black, basally tan. Bucculae tan, biseriate, closed apically projecting anteriorly beyond apex of frons and posteriorly over prosternum. Rostrum moderately long, extending to mesocoxae, yellow ochraceous, tip black. Pronotum finely punctate, yellow ochraceous; paranota biseriate, each anterolateral margin with two prominent spines, one expressed anteriorly and the other laterally, irregular spines and spinules along lateral margins; pronotal hood extremely minute, low, porrect; tricarinate; median carina low, but arched in middle, uniseriate except biseriate near middle; lateral carina not as tall as median carina, uniseriate, subparallel.Triangular pronotal posterior projection unicolorous with pronotum, rugose on basal third, then gradually areolate on apex. Hemelytra constricted, extending far beyond apex of abdomen, with spines along lateral margins; costal areas biseriate, with a few intercalary cells near hemelytral constriction; subcostal areas uniseriate at base, biseriate at middle, cells infuscate along subcostal extensions; rm veins elevated above cubital veins in lateral view; discoidal cells narrow, triseriate, each extending to middle of hemelytra; sutural areas triseriate at widest. Rostral laminae slightly elevated, uniseriate, subparallel on pro- and mesothorax, crescentic on metathorax. Legs yellow, slender; tarsi red fuscous. Abdomen narrow, ochraceous, beset with few scattered hairs; abdominal spiracular peritremes laterally projected. Measurements. (n = 1) Length: 2.27, width at widest: 1.20, length of antennal segments one through four, respectively: 0.22, 0.10, 1.21, and 0.33. Type material. Holotype: PANAMA: Colon Province. PNAR. Lago Gatun, malaise 99/100 e. 300 ft., 9̊ 20’ 08” N 79̊ 50’ 52” W, A. Gillogly 4–13-VII-1999 (♂ TAMU). Holotype will be deposited in the TAMU Holotype collection. Etymology. This species is named for its general triangular (delt) appearance. Plant associations. No plant associations are known. Comments. This species was listed as Acanthopharsa deltoides in an unpublished thesis (Knudson 2018)., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on pages 336-338, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Knudson, A. H. (2018) The Tingidae (Hemiptera: Heteroptera) of Southern Central America (with an emphasis on Costa Rica). Master's thesis, North Dakota University, Fargo, North Dakota, 314 pp."]}

Stephanitis (Stephanitis) parana Drake & Hambleton, 1944 Stephanitis parana Drake & Hambleton, 1944, 128–129. Stephanitis parana: Drake & Maa, 1953: 101 (checklist); Drake & Ruhoff, 1965: 355 (catalog); Neal & Schaefer, 2000: 101 (note). Geographic distribution. Pará, Brazil (Drake & Hambleton 1944). Plant associations. Unrecorded.

Stephanitis (Stephanitis) Stål, 1873 (Figs. 3A–D, 4) Tingis: Laporte, 1833: 48. (not Fabricius, 1803) Stephanitis Stål, 1873: 119, 123. Tingis: Lethierry & Severin, 1896: 12. (not Fabricius, 1803) Cadamustus Distant, 1903: 47. (syn. by Horváth, 1906a) Maecenas Kirkaldy, 1904: 280. (unnecessary new name for Tingis Lethierry & Severin, 1896; syn. by Horváth, 1906a) Calliphanes Horváth, 1906b: 34. (unnecessary new name for Stephanitis: Champion, 1898; syn. by Drake & Maa, 1953) Mokanna Distant, 1910a: 111. (syn. by Horváth, 1912) Type species: Stephanitis Stål, 1873: Acanthia pyri Fabricius, 1775 by subsequent designation (Horváth, 1906b). Cadamustus Distant, 1903: Cadamustus typicus Distant, 1903 by indication. Maecenas Kirkaldy, 1904: Acanthia pyri Fabricius, 1775 by subsequent designation (Drake and Ruhoff 1960). Calliphanes Horváth, 1906b: Tingis mitrata by original designation. Mokanna Distant, 1910a: Mokanna princeps Distant, 1910a, by original designation. Diagnosis. Stephanitis can be separated from all other known genera by the following combination of characters: macropterous, pubescent or hairless above, without spinules on the veins of the areolae of the pronotum and hemelytra. Head with five cephalic spines, antennae slender, basiflagellomere, nearly half as long as body. Bucculae may or may not be contiguous apically, but comprised of at least two rows of areolae on lateral margins. Pronotal hood moderately globose or laterally compressed, almost completely covering head dorsally. Paranota wide, and raised posteriorly, reflexed and curved onto posterior dorsum of pronotum, lateral margins may be armed with spinules. Pronotum tricarinate, lateral carinae sometimes greatly reduced, median carinae usually as tall as pronotal hood (except S. joceliae) and extending to apex of posterior triangular projection. Costal area wide, at least biseriate; subcostal areas subvertical, at times forming a rounded tumidity with discoidal areas; discoidal areas short, less than one third the length of hemelytra, rm vein weak or absent in some species; sutural areas biseriate to triseriate. Legs long, slender. Ostiolar peritreme well developed., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 338, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Stal, C. (1873) Enumeratio Hemipterorum. vol. 3. Kongliga Svenska Vetenskaps Akademiens Handlingar, 11 (2), 1 - 163.","Laporte, F. L. N. C. (1833) Essai d'une classification systematique de l'ordre des hemipteres (hemipteres heteropteres, Latr.). Magasin Zool o gie, 2, 1 - 88.","Fabricius, J. C. (1803) Systema rhyngotorum, secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae, 314 pp. https: // doi. org / 10.5962 / bhl. title. 11644","Lethierry, L. & Severin, G. (1896) Catalogue general des Hemipteres. Vol. 3. F. Hayez, Bruxelles, 275 pp.","Distant, W. L. (1903) Contributions to a knowledge of the Rhynchota. Annales de la Societe Entomologique de Belgique, 47, 43 - 65.","Horvath, G. (1906 a) Synopsis Tingitidarum regionis Palaearcticae. Annales historico-naturales Musei nationalis hungarici, 4, 1 - 118.","Kirkaldy, G. W. (1904) Bibliographical and nomenclatorial notes on the Hemiptera. No. 3. The Entomologist, 37 (498), 279 - 283. https: // doi. org / 10.5962 / bhl. part. 2885","Horvath, G. (1906 b) A new gall-inhabiting bug from Bengal. Entomologists' Monthly Magazine 42, 33 - 34.","Champion, G. C. (1898) Notes on American and other Tingitidae, with descriptions of two new genera and four new species. Transactions of the Entomology Society of London, 1, 55 - 64.","Drake, C. J. & Maa, T. (1953) Chinese and other Oriental Tingoidea (Hemiptera). Quarterly Journal of the Taiwan Museum, 6 (2), 87 - 101.","Distant, W. L. (1910 a) The fauna of British India, including Ceylon and Burma. Rhynchota. Vol. V. Heteroptera: Appendix. Taylor and Francis, London, 362 pp.","Horvath, G. 1912. Species generis Tingitidarum Stephanitis. Annales Historico-Naturales Musei Nationalis Hungarici, 10, 319 - 339.","Fabricius, J. C. (1775) Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensbvrgi et Lipsiae, 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510","Drake, C. J. & Ruhoff, F. A. (1960) Lace-bug genera of the world (Hemiptera: Tingidae). Proceedings of the United States National Museum, 54, 1 - 107. https: // doi. org / 10.5479 / si. 00963801.112 - 3431.1"]}

Stephanitis joceliae, sp. nov. (Figs. 3A–D, 4) urn:lsid:zoobank.org:act: AF4C7A38-7326-4626-8D65-15D8CFB30AF2 Diagnosis. Stephanitis joceliae can be separated from all other members of the genus, by the bucculae which project beyond the apex of the head; the long scape (0.37–0.40); by the ear shaped paranota which project apically beyond the head and pronotal hood; by the low, uniseriate median carina that is no more than half the height of the pronotal hood; and by the discoidal cell which is open apically. Description. Generally flat, wings and paranota broadly expanded, veins infuscate, areolae hyaline. Head with moderately long, downcurved spines, adpressed to head; median spine extremely long, adpressed to head, extending to clypeus; frontal spines shorter, adpressed to head, nearly extending to clypeus. Antennae moderately long, when directed posteriorly extending to wing base; scape (segment I) long, tan, beset with a few scattered long hairs; pedicel (segment II) extremely short, tan; basiflagellomere (segment III) extremely long, two- and one-half times longer than scape, tan; distiflagellomere (segment IV) long, about one half times as long as preceding, fuscous basally, black on apical half, beset with long fine hairs. Bucculae contiguous apically, extended beyond apex of head, biseriate; rostrum moderately long, extending to metacoxae. Pronotum punctate, ochraceous; tricarinate; median carina half as tall as pronotal hood, gradually sloping posteriorly, uniseriate; lateral carinae nearly as tall as median carina, uniseriate, subparallel, rounded in lateral view; pronotal hood narrow, moderately low, subtruncate anteriorly, comprised of two rows of cells along lateral margins, beset with short fine hairs; paranota slightly reflexed upwards, auriform, broadly expanded laterally and projecting anteriorly beyond apex of head, quadriseriate at widest; posterior pronotal projection areolate, extending to third abdominal tergite. Hemelytra broadly expanded; costal veins with minute hairs along lateral margins, hairs on basal fourth emerging from thickened projections; costal areas of hemelytra quadriseriate at base, five rows of areolae wide; subcostal areas biseriate on basal third, uniseriate throughout subcostal extensions; discoidal cells triseriate, open behind; sutural areas uniseriate. Metathoracic wings extremely small, not extending to middle of abdomen. Thorax dark ochraceous, venter beset with fine white wax; rostral laminae subparallel on pro- and mesothorax, slightly converging posteriorly, uniseriate. Legs slender, tan; femora long; tibiae slightly longer than femora; tarsi long, slender. Each ostiolar peritreme expanded laterally in posterior margin. Abdomen light red, narrow; pygophore as long as two preceding abdominal segments; parameres slender, short, not reaching anterior edge of pygophore in repose. Measurements. (n = 4; holotype in parentheses) Length: 3.20–3.57 (3.55), width at widest: 2.16–2.85 (2.85); height of hood: 0.32–0.36 (0.36); length of antennal segments one through four, respectively: 0.37–0.40 (0.40), 0.09–0.12 (0.12), 1.00–1.10 (1.10), and 0.61–0.70 (0.70). Type material. Holotype: PANAMA: Chiriqui Pr.: Resve. La Fortuna, Est. Biologica. El. 3900’, 8˚ 43’ 18” N 82˚14’ 17”W, 7–9-I-2001, Yellow pan traps. M. Yoder (1 ♂ TAMU). Paratypes: PANAMA: Chiriqui Pr.: Resve. La Fortuna, Est. Biologica. El. 3900’ N8˚ 43’ 18” W 82˚ 14’ 17”, 7–9-VIII-1999, ypt. 99/ 079, J. Schaffner & J. B. Woolly (1 ♂ TAMU); Bocas del Toro: Trail off Oleoducto. 4km W Fortuna Hwy. 5-VIII-1999, el. 3640 ft, 08˚ 47’ 14” N 82˚ 12’ 50” W, J. B. Wolly 99/083 (1 ♀ TAMU); Colclé: 7.2 km NE. El Copé. 730m 08°37’N, 80° 35’ W, #140, 20 V –7 VI 1995, J. Ashe, R. Brooks, flt. Intercept trap (1 ♂ SEMC). All types will be deposited in their respective collections. Etymology. This species is named in honor of eminent Brazilian heteropterist Jocelia Grazia in recognition of her many contributions to heteropteran systematics, especially of the Pentatomidae. Plant associations. No plant associations are known. Comments. This species was listed as Neobyrsa panamensis in an unpublished thesis (Knudson 2018).

Stephanitis (Stephanitis) farameae Drake & Hambleton, 1935 Stephanitis farameae Drake & Hambleton 1935: 151–152. Stephanitis farameae: Monte, 1939: 78 (note); 1941: 132 (catalog); Drake & Maa, 1953: 100) (checklist); Drake & Ruhoff, 1965: 355 (catalog). Geographic distribution. São Paulo and Minas Gerais, Brazil (Drake & Hambleton 1935, Monte 1939). Plant associations. Faramea montevidensis [Rubiaceae] (Drake & Hambleton 1935).

Stephanitis (Stephanitis) mitrata (Stål 1860) Tingis mitrata Stål, 1860: 64. Tingis mitrata: Walker, 1873: 181 (catalog); Lethierry and Severin, 1896: 12 (catalog). Stephanitis mitrata: Stål, 1873: 123 (catalog); Champion 1898: 58, pl. II figs. 3, 3a (note); Drake & Poor, 1937a: 309 (note); Drake and Hambleton, 1938: 66 (note); Monte, 1941: 133 (catalog); Drake & Maa, 1953: 101 (checklist); Drake & Ruhoff, 1965: 355 (catalog); Neal & Schaefer, 2000: 101 (note). Calliphanes mitratus: Horváth, 1906b: 34 (note). Geographic distribution. Rio de Janeiro and São Paulo, Brazil (Monte 1941). Plant associations. Merostachys sp. [Poaceae] (Drake & Hambleton 1938)., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 341, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Stal, C. (1860) Bidrag till Rio Janeiro-Traktens Hemipter-Fauna. I. Ofversigt Kongliga Svenska Vetenskaps Akademiens Handlingar, 2 (7), 1 - 84. [1858]","Walker, F. (1873) Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. Part VI. Printed for the Trustees of the British Museum, London, 210 pp.","Lethierry, L. & Severin, G. (1896) Catalogue general des Hemipteres. Vol. 3. F. Hayez, Bruxelles, 275 pp.","Stal, C. (1873) Enumeratio Hemipterorum. vol. 3. Kongliga Svenska Vetenskaps Akademiens Handlingar, 11 (2), 1 - 163.","Champion, G. C. (1898) Notes on American and other Tingitidae, with descriptions of two new genera and four new species. Transactions of the Entomology Society of London, 1, 55 - 64.","Drake, C. J. & Poor, M. (1937 a) The South American Tingitidae (Hemiptera) described by Stal. Memoirs of the Carnegie Museum, 11 (5), 301 - 312.","Drake, C. J. & Hambleton, E. J. (1938) Concerning Brazilian Tingitidae (Hemiptera). Part III. Revista Entomologica, Rio de Janeiro, 8 (1 - 2), 44 - 68.","Monte, O. (1941) Catalogo dos tingitideos do Brazil. Separata dos Arquivos de Zoologia Do Estado de Sao Paulo, 2, 3, 65 - 174.","Drake, C. J. & Maa, T. (1953) Chinese and other Oriental Tingoidea (Hemiptera). Quarterly Journal of the Taiwan Museum, 6 (2), 87 - 101.","Drake, C. J. & Ruhoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington, D. C., 634 pp. https: // doi. org / 10.5479 / si. 03629236.243.1","Horvath, G. (1906 b) A new gall-inhabiting bug from Bengal. Entomologists' Monthly Magazine 42, 33 - 34."]}

Stephanitis (Stephanitis) aucta Drake, 1942 Stephanitis aucta Drake 1942: 20–21. Stephanitis aucta: Drake & Maa, 1953: 100 (checklist); Drake & Ruhoff, 1965: 354 (catalog). Geographic distribution. Chapada, Bahia, Brazil (Drake 1942). Plant associations. Unknown., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 338, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Drake, C. J. (1942) New Tingitidae (Hemiptera). Iowa State College Journal of Science, 17 (1), 1 - 21.","Drake, C. J. & Maa, T. (1953) Chinese and other Oriental Tingoidea (Hemiptera). Quarterly Journal of the Taiwan Museum, 6 (2), 87 - 101.","Drake, C. J. & Ruhoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington, D. C., 634 pp. https: // doi. org / 10.5479 / si. 03629236.243.1"]}

Stephanitis (Stephanitis) blatchleyi Drake, 1925 Stephanitis blatchleyi Drake, 1925: 37. Leptobyrsa blatchleyi: Blatchley, 1926: 470 (note). Stephanitis blatchleyi: Drake & Maa, 1953: 100 (checklist); Drake & Ruhoff, 1965: 354 (catalog); Froeschner, 1988: 730 (catalog); Neal & Schaefer, 2000: 101 (note); Wheeler & Stoops, 2013: 673–675 (note). Geographic distribution. Florida, USA (Wheeler & Stoops 2013). Plant associations. Lyonia ferruginea [Ericaceae] (Wheeler & Stoops 2013) Lyonia fruticosa (Halbert 2018)., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 338, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Drake, C. J. (1925) Concerning some Tingitidae from the Gulf States (Heteroptera). Florida Entomologist, 9, 36 - 39. https: // doi. org / 10.2307 / 3492477","Blatchley, W. S. (1926) Heteroptera or true bugs of eastern North America, with especial references to the faunas of Indiana and Florida. Nature Publishing Co., Indianapolis, 1116 pp. https: // doi. org / 10.5962 / bhl. title. 6871","Drake, C. J. & Maa, T. (1953) Chinese and other Oriental Tingoidea (Hemiptera). Quarterly Journal of the Taiwan Museum, 6 (2), 87 - 101.","Drake, C. J. & Ruhoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington, D. C., 634 pp. https: // doi. org / 10.5479 / si. 03629236.243.1","Froeschner, R. C. (1988) Family Tingidae Laporte, 1807 (= Tingididae, Tingitidae): The Lace Bugs. In: Henry, T. J. & Froeschner, R. C. (Eds.), Catalog of the Heteroptera, or True Bugs, of Canada and the Continental United States. E. J. Brill, Leiden and New York, pp. 708 - 733. https: // doi. org / 10.1201 / 9781351070447 - 44","Wheeler, A. G. Jr. & Stoops, C. A. (2013) Stephanitis blatchleyi (Hemiptera: Tingidae): First host plant association for a rarely collected lace bug. Florida Entomologist, 96 (2), 673 - 675. https: // doi. org / 10.1653 / 024.096.0244","Halbert, S. E. (2018) Entomology Section. Anderson, P. J. & Hodges, G. S. (Eds.), Tri-ology, 57 (2), pp. 7 - 14. Available from: https: // www. fdacs. gov / ezs 3 download / download / 82775 / 2395058 / Media / Files / Plant-Industry-Files / Tri-ology / Triology % 2057 - 2 % 20 April % 20 June % 202018.1. pdf (accessed 20 October 2020)"]}

Acanthotingis apicicornis Monte, 1940 (Fig. 2A) Acanthotingis apicicornis Monte 1940, 15–16. Acanthotingis apicicornis: Drake & Ruhoff 1965: 59–60 (catalog). Geographic distribution. Known only from the type series taken in São Paulo, Brazil (Monte 1940). Plant associations. Rubiaceae (Monte 1940). Comments. The type specimens are presumed lost due to the fire that destroyed much of the Museu Nacional du Brazil, in Rio de Janeiro, 2 September 2018., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 336, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Monte, O. (1940) Descricao de um novo genero e uma nova especie de tingitideo (Hem.). Papeis Avulsos Departamento Zoologia, Sao Paulo, Brazil, 1, 13 - 16.","Drake, C. J. & Ruhoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington, D. C., 634 pp. https: // doi. org / 10.5479 / si. 03629236.243.1"]}

Stephanitis (Stephanitis) olyrae Drake & Hambleton, 1935 Stephanitis olyrae Drake & Hambleton, 1935: 150–151, fig. 5. Stephanitis olyrae: Monte, 1939: 78 (note); 1941: 133 (catalog); Drake & Maa, 1953: 101 (checklist); Drake & Ruhoff, 1965: 358 (catalog); Neal & Schaefer, 2000: 101 (note). Geographic distribution. Minas Gerais and São Paulo, Brazil (Drake & Hambleton 1935, Monte 1939). Plant associations. Olyra micrantha [Poaceae] (Drake & Hambleton 1935)., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 341, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006, {"references":["Drake, C. J. & Hambleton, E. J. (1935) New Brazilian Tingitidae (Hemiptera) Part II. Archivos do Instituto de Biologia, Sao Paulo, 6, 141 - 154.","Monte, O. (1939) Lista preliminar dos tingitideos de Minas Gerais. Revista Sociedade Brazileira de Agronomia, 2, 1, 63 - 87.","Drake, C. J. & Maa, T. (1953) Chinese and other Oriental Tingoidea (Hemiptera). Quarterly Journal of the Taiwan Museum, 6 (2), 87 - 101.","Drake, C. J. & Ruhoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington, D. C., 634 pp. https: // doi. org / 10.5479 / si. 03629236.243.1"]}

Key to the species of Acanthotingis Monte 1. Paranota with three to four stout spines; costal margins of hemelytra devoid of spines; subcostal area of hemelytra with four rows of small cells........................................................... Acanthotingis apicicornis Monte - Paranota with three or four spinules and a series of small spinules along posterior lateral margins; costal margin of hemelytra with a series of small spinules; subcostal area of hemelytra with two rows of cells......... Acanthotingis deltoides sp. nov., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, pp. 334-344 in Zootaxa 4958 (1) on page 336, DOI: 10.11646/zootaxa.4958.1.20, http://zenodo.org/record/4692006

Ambotingis senta (Drake and Hambleton 1945) Plate 1 figs a–d Monanthia senta Drake and Hambleton 1942: 329; Drake and Hambleton 1945: 356. Dictyla senta: Drake and Ruhoff 1960a: 51. Ambotingis senta: Drake and Ruhoff 1960b: 31, 33, fig. 2 [Not fig. 1, see comments]; Drake and Ruhoff 1960c: 80; Drake and Ruhoff 1965: 83, pl. 10; Ojeda and Neciosup 1974: 114. Diagnosis. Readily separated from its congener, by the sharp and projecting cephalic spines, by the more elevated and spiny paranota, by the variegated color of the hemelytra, and by spines along the lateral margins. Material examined. Type material examined: HolotypeƋ: Peru: Sullena, May, 10, 1942, E. J. Hambleton (USNM). Allotype♀: Sullena, May, 10, 1942, E. J. Hambleton (USNM). (Figure 1). Paratypes: Peru, Sullena, May 10, 1942, E. J. Hambleton (2♂ 2♀ USNM). Peru, Paita, July, 7, 1940, E. J. Hambleton (1 missing abdomen USNM). Other specimens. Ecuador, October, 19, 1944, E. J. Hambleton (4♂ 2♀ USNM). Ecuador: Machala, September, 27, 1944, E. J. Hambleton, (1♂ 2♀ USNM). Plant associations. The host plant for this species is unknown. Ojeda and Neciosup (1974) listed many host plant records for different tingid species found in Peru, but they did not list any host plants for A. senta. Drake and Hambleton (1941) stated that this species had been taken on a woody shrub, but they did not provide any specific information. Comments. In their original description, Drake and Ruhoff (1960b) inadvertently switched the captions of figure 1 and figure 2; they corrected this error later that same year (Drake and Ruhoff, 1960c). PLATE 1. Ambotingis senta. Fig. a. dorsal habitus of holotype (left) and allotype (right). Fig. b. Type labels. Fig. c. lateral habitus. Fig. d. venation of forewing., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2017, Review of the Genus Ambotingis Drake and Ruhoff (Hemiptera: Heteroptera: Tingidae), pp. 146-150 in Zootaxa 4269 (1) on page 147, DOI: 10.11646/zootaxa.4269.1.10, http://zenodo.org/record/581356, {"references":["Drake, C. J. & Hambleton, E. J. (1945) Concerning Neotropical Tingitidae (Hemiptera). Journal of the Washington Academy of Sciences, 35, 356 - 367.","Drake, C. J. & Hambleton, E. J. (1942) Two new Peruvian Tingitidae (Hemiptera). Iowa State College Journal of Science, 16, 329 - 330.","Drake, C. J. & Ruhoff, F. A. (1960 a) Lace-bug genera of the world (Hemiptera: Tingidae). Proceedings of the United States National Museum, 54, 1 - 107.","Drake, C. J. & Ruhoff, F. A. (1960 b) Tingidae: new genera, species, homonyms, and synonyms (Hemiptera). Great Basin Naturalist, 20, 29 - 38.","Drake, C. J. & Ruhoff, F. A. (1960 c) A necessary correction. Great Basin Naturalist, 20, 80.","Drake, C. J. & Rhuoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington D. C., 634 pp.","Ojeda, D. & Neciosup, M. A. (1974) Tingidae (Hemiptera) del Peru y sus Hospederos. Revista Peruana de Entomologia, 17, 114 - 115."]}

Ambotingis ainslei (Drake and Poor), NEW COMBINATION Plate 2: Figures a–d Monanthia ainslei Drake and Poor 1938: 1 –2, fig. 1. Dictyla ainslei: Drake and Ruhoff 1960a: 50 –51; Drake and Ruhoff 1965: 181. Diagnosis. Ambotingis ainslei can be separated from A. senta by the blunt, suberect cephalic spines, by the less elevated paranota with few spines, by the posteriorly incomplete median carina, by the tan or cream-colored dorsum, and by the long and ovate hemelytra that lack spines along the lateral margins. Redescription. General appearance ovate, tan to cream colored with a few fuscous markings, thorax and abdomen testaceous. Head testaceous, armed with five long, cream-colored, suberect, blunt spines, occipital pair slightly divergent, clypeal pair converging. Head short, slightly expanded beyond eyes. Bucculae closed anteriorly, produced slightly beyond apex of head. Antennae slender, segment one about as long as wide, fuscous; segment two, twice as long as wide, narrower than segment one, dark reddish brown; segment three long, cream colored, indistinctly pubescent with small white hairs; segment four slightly stout, about as long as segments one and two taken together, apex fuscous, armed with long, fine, white hairs. Antenniferous tubercles obsolete, modified into areolate flaps on bucculae. Bucculae triseriate, terminating on anterior margin of prosternum. Rostrum nearly extending to metasternum. Rostral groove welldeveloped, shallow on prosternum, deep on mesosternum, pro- and mesosternal laminae subparallel, metasternal laminae widely diverging. Pronotum mostly areolate, anteriorly with large punctations, feebly tricarinate, median carina posteriorly obsolete or incomplete, lateral carinae converging, carinae with small indistinct areolae. Paranota large, reflexed over pronotum, covering anterior two thirds of lateral carinae, meeting at median carina. Paranota each with two linear, cariniform processes. Median carina slightly elevated anteriorly; pronotal collar well developed, forming small tectiform hood, which does not cover head. Mesothorasic scent gland lacking. Hemelytra elongate, ovate, extending slightly beyond apex of abdomen; costal area biseriate, with large pentagonal areolae, subcostal area triseriate, distinctly narrowed in middle, but widely expanding on both sides of constriction, appearing yolk shaped. R+M vein elevated in middle of each hemelytron, fuscous. Discoidal area with external margin sinuate, abruptly widening to four to five cells at greatest width. Legs slender, cream colored, femora as long as tibiae; tarsi slender, segment one small, segment two more than four times as long as segment one, apically fuscous, slightly flattened dorso-ventrally, with long fine hairs on ventral surface, pretarsi light fuscous, narrower than segment two, tarsal claws divergent, fuscous. Abdomen reddish brown, elongate oval; abdominal segments subequal in length, slightly rugose near middle; sub-genital plate smooth, hexagonal, broadest near middle; gonocoxae lighter in color; ninth paratergites deeply rugose. Measurements: (n = 1) Length 2.60, width across pronotum: 0.89, height of paranota: 0.62, width across hemelytra: 1.37, antennal segments one through four, respectively: 0.10, 0.09, 0.64, and 0.20. Material examined: Type material examined: Holotype ♀: Guatemala: Concepcion, elevation 1400 ft., C. N. Ainslie (USNM) Other specimens: Costa Rica: Guanacaste, La Pacifica near Cañas, May, 22–26, 1984, E. Riley, D. Rider & D. LeDoux (3♀ DARC). Guatemala: El Progresso, Guastatoya, 5km S jct. Rio Sanarate and CA 9, 6051m, 14̊50' 19"N – 04' 16"W, January, 9, 2007, J. R. Jones (1♀ TAMU). Plant associations. No biological information has been published or was indicated on any of the specimen labels. Comments. All specimens known of A. ainsliei are female., Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2017, Review of the Genus Ambotingis Drake and Ruhoff (Hemiptera: Heteroptera: Tingidae), pp. 146-150 in Zootaxa 4269 (1) on page 148, DOI: 10.11646/zootaxa.4269.1.10, http://zenodo.org/record/581356, {"references":["Drake, C. J. & Poor, M. E. (1938) Nine new American Tingitidae (Hemiptera). Bulletin of the Brooklyn Entomological Society, 33, 28 - 34.","Drake, C. J. & Ruhoff, F. A. (1960 a) Lace-bug genera of the world (Hemiptera: Tingidae). Proceedings of the United States National Museum, 54, 1 - 107.","Drake, C. J. & Rhuoff, F. A. (1965) Lacebugs of the World: A Catalog (Hemiptera: Tingidae). Bulletin 243. Smithsonian Institution, Washington D. C., 634 pp."]}

Genus Ambotingis Drake and Ruhoff 1960 Type Species: Monanthia senta Drake and Hambleton, by original designation. Diagnosis. Ambotingis can be separated from all other allied tingid genera by the combination of the following characters: Costal area of the hemelytra broadly expanded, with pentagonal to triangular cells. Pronotum strongly convex and punctate, but obscured by the greatly reflexed paranota. Each paranota exhibits two linear cariniform processes. Ostiolar peritremes absent. Geographic distribution. Guatemala, Costa Rica, Ecuador, Peru, Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2017, Review of the Genus Ambotingis Drake and Ruhoff (Hemiptera: Heteroptera: Tingidae), pp. 146-150 in Zootaxa 4269 (1) on page 146, DOI: 10.11646/zootaxa.4269.1.10, http://zenodo.org/record/581356

Key to the species of Ambotingis 1 Costal margins of hemelytra armed with spines; paranota strongly elevated with many spines........................................................ ........................................................................................................................................... Ambotingis senta (Drake and Hambleton) - Costal margins of hemelytra unarmed; paranota less elevated, with few spines................... Ambotingis ainsliei (Drake and Poor), Published as part of Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2017, Review of the Genus Ambotingis Drake and Ruhoff (Hemiptera: Heteroptera: Tingidae), pp. 146-150 in Zootaxa 4269 (1) on page 146, DOI: 10.11646/zootaxa.4269.1.10, http://zenodo.org/record/581356