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Middle Stone Age (MSA) technologies first appear in the archaeological records of northern, eastern and southern Africa during the Middle Pleistocene epoch. The absence of MSA sites from West Africa limits evaluation of shared behaviours across the continent during the late Middle Pleistocene and the diversity of subsequent regionalized trajectories. Here we present evidence for the late Middle Pleistocene MSA occupation of the West African littoral at Bargny, Senegal, dating to 150 thousand years ago. Palaeoecological evidence suggests that Bargny was a hydrological refugium during the MSA occupation, supporting estuarine conditions during Middle Pleistocene arid phases. The stone tool technology at Bargny presents characteristics widely shared across Africa in the late Middle Pleistocene but which remain uniquely stable in West Africa to the onset of the Holocene. We explore how the persistent habitability of West African environments, including mangroves, contributes to distinctly West African trajectories of behavioural stability. Results - Stratigraphy - Geochronology - Palaeoecology - Stone tool technology Discussion Methods - Excavation - Sedimentology - Geochronology - Palaeoecology - Stone tools

Levallois points are a prominent part of many Levantine Middle Paleolithic assemblages. They are either produced intentionally or incidentally by the Levallois core reduction technique and are of a generally similar shape, although the degree to which they were used as specialized tools has been questioned. Here, we examine Levallois points using geometric morphometric analyses to assess the range of shape variation in this artifact type. We then compare Levallois point shape variation and symmetry to a sample of Late Pleistocene-aged Folsom projectile points from North America. Folsom are highly standardized projectile points that current evidence suggests were primarily used for hunting ancient bison. Our results indicate that Levallois points are more variable and asymmetrical than Folsom and therefore more generalized than Folsom. Differences in manufacturing technique, hafting, and delivery system when used as weapons are posited as playing roles in the differences in shape and symmetry that we document. Introduction Materials and methods - Point samples - Geometric morphometric procedures - Statistical analyses Results - Levallois point shape variation - Comparison of Levallois points with Folsom - Symmetry of Levallois and Folsom points Discussion

South Asia hosts the world’s youngest Acheulean sites, with dated records typically restricted to sub-humid landscapes. The Thar Desert marks a major adaptive boundary between monsoonal Asia to the east and the Saharo-Arabian desert belt to the west, making it a key threshold to examine patterns of hominin ecological adaptation and its impacts on patterns of behaviour, demography and dispersal. Here, we investigate Palaeolithic occupations at the western margin of the South Asian monsoon at Singi Talav, undertaking new chronometric, sedimentological and palaeoecological studies of Acheulean and Middle Palaeolithic occupation horizons. We constrain occupations of the site between 248 and 65 thousand years ago. This presents the first direct palaeoecological evidence for landscapes occupied by South Asian Acheulean-producing populations, most notably in the main occupation horizon dating to 177 thousand years ago. Our results illustrate the potential role of the Thar Desert as an ecological, and demographic, frontier to Palaeolithic populations. Results - Sediment sequence. - Luminescence dating. - Palaeoenvironmental studies. - Lithic artefacts. Discussion Methods

Homo sapiens have adapted to an incredible diversity of habitats around the globe. This capacity to adapt to different landscapes is clearly expressed within Africa, with Late Pleistocene Homo sapiens populations occupying savannahs, woodlands, coastlines and mountainous terrain. As the only area of the world where Homo sapiens have clearly persisted through multiple glacial-interglacial cycles, Africa is the only continent where classic refugia models can be formulated and tested to examine and describe changing patterns of past distributions and human phylogeographies. The potential role of refugia has frequently been acknowledged in the Late Pleistocene palaeoanthropological literature, yet explicit identification of potential refugia has been limited by the patchy nature of palaeoenvironmental and archaeological records, and the low temporal resolution of climate or ecological models. Here, we apply potential climatic thresholds on human habitation, rooted in ethnographic studies, in combination with high-resolution model datasets for precipitation and biome distributions to identify persistent refugia spanning the Late Pleistocene (130–10 ka). We present two alternate models suggesting that between 27% and 66% of Africa may have provided refugia to Late Pleistocene human populations, and examine variability in precipitation, biome and ecotone distributions within these refugial zones. 1. Introduction 2. The role of refugia in African palaeoanthropology 3. Exploring potential human refugia in Late Pleistocene African landscapes (a) Identifying precipitation refugia (b) Biome changes within precipitation refugia (c) Open and forested landscapes within precipitation refugia (d) A case study of Late Pleistocene human occupation of eastern African refugia 4. Discussion

Eastern Africa has played a prominent role in debates about human evolution and dispersal due to the presence of rich archaeological, palaeoanthropological and palaeoenvironmental records. However, substantial disconnects occur between the spatial and temporal resolutions of these data that complicate their integration. Here, we apply high-resolution climatic simulations of two key parameters, mean annual temperature and precipitation, and a biome model, to produce a highly refined characterisation of the environments inhabited during the eastern African Middle Stone Age. Occupations are typically found in sub-humid climates and landscapes dominated by or including tropical xerophytic shrubland. Marked expansions from these core landscapes include movement into hotter, low-altitude landscapes in Marine Isotope Stage 5 and cooler, high-altitude landscapes in Marine Isotope Stage 3, with the recurrent inhabitation of ecotones between open and forested habitats. Through our use of high-resolution climate models, we demonstrate a significant independent relationship between past precipitation and patterns of Middle Stone Age stone tool production modes overlooked by previous studies. Engagement with these models not only enables spatiotemporally explicit examination of climatic variability across Middle Stone Age occupations in eastern Africa but enables clearer characterisation of the habitats early human populations were adapted to, and how they changed through time. - Results -- Middle and Late Pleistocene climates of MSA occupations. -- Classifying biomes and ecotones at MSA occupations. -- Characterising MSA environments throughout the Middle to Late Pleistocene. -- Phased habitability models. -- Exploring the relationship between climate and Middle Stone Age occupations. - Conclusion - Materials and methods -- Refining the climatic parameters. -- Cluster analyses. -- Biome classification. -- Phased climate suitability models. -- Mantel tests and multiple matrix regression.

Recent archaeological investigations in Sri Lanka have reported evidence for the exploitation and settlement of tropical rainforests by Homo sapiens since c. 48,000 BP. Information on technological approaches used by human populations in rainforest habitats is restricted to two cave sites, Batadomba-lena and Fa-Hien Lena. Here, we provide detailed study of the lithic assemblages of Kitulgala Beli-lena, a recently excavated rockshelter preserving a sedimentary sequence from the Late Pleistocene to the Holocene. Our analysis indicates in situ lithic production and the recurrent use of the bipolar method for the production of microliths. Stone tool analyses demonstrate long-term technological stability from c. 45,000 to 8,000 years BP, a pattern documented in other rainforest locations. Foraging behaviour is characterised by the use of lithic bipolar by-products together with osseous projectile points for the consistent targeting of semi-arboreal/arboreal species, allowing for the widespread and recurrent settlement of the wet zone of Sri Lanka. Introduction Mobility and hunter-gatherer tool-kits Kitulgala Beli-lena rockshelter Materials and methods Results - Late Pleistocene (c. 45,000–31,000 cal BP) - Terminal Pleistocene (17,157–11,314 cal BP) - Holocene (10,577–8,029 cal BP) Summary Discussion

An exclusive connection between Homo sapiens and Nubian Levallois technology has been posited, but remains to be demonstrated1 . Our re-evaluation of the fossil and lithic material from Shukbah Cave confounds such assumptions due to the identification of a Neanderthal molar tooth alongside Nubian Levallois cores and points at the site2 . Hallinan and colleagues3 question this fnding, instead supporting the use of Nubian Levallois technology as a fossile directeur to track expansions of Homo sapiens. We tackle these critiques, highlighting the problematic foundations in the assertion that Nubian Levallois technology is a unique, discrete entity, resulting in its misuse to support simplistic culture-historical narratives., peer-reviewed

Studies of modern carnivore accumulations of bone (i.e., neo-taphonomy) are crucial for interpreting fossil accumulations in the archaeological and paleontological records. Yet, studies in arid regions have been limited in both number and detailed taphonomic data, prohibiting our understanding of carnivore bone-accumulating and -modifying behavior in dry regions. Here, we present a taphonomic analysis of an impressive carnivore-accumulated bone assemblage from the Umm Jirsan lava tube in the Harrat Khaybar region, Saudi Arabia. The size and composition of the bone accumulation, as well as the presence of hyena skeletal remains and coprolites, suggest that the assemblage was primarily accumulated by striped hyena (Hyaena hyaena). Our findings (1) identify potentially useful criteria for distinguishing between accumulations generated by different species of hyenas; (2) emphasize the need for neo-taphonomic studies for capturing the full variation in carnivore bone-accumulating and modifying behavior; (3) suggest that under the right settings, striped hyena accumulations can serve as good proxies for (paleo)ecology and livestock practices; and (4) highlight the potential for future research at Umm Jirsan, as well as at the numerous nearby lava tube systems. We encourage continued neo-taphonomic efforts in regions important in human prehistory, particularly in arid zones, which have received little research attention. Introduction - Study area Material and methods Results - Species representation - Skeletal part representation - Bone surface modifications - Bone fragmentation and fracture patterns - Mortality profiles Discussion - Accumulator(s) of the Umm Jirsan assemblage - Striped hyena feeding behavior at Umm Jirsan - Implications for prehistory and paleoecology Conclusion

Marine Isotope Stage (MIS) 5, ~ 130 to 71 thousand years ago, was a key period for the geographic expansion of Homo sapiens, including engagement with new landscapes within Africa and dispersal into Asia. Occupation of the Levant by Homo sapiens in MIS 5 is well established, while recent research has documented complementary evidence in Arabia. Here, we undertake the frst detailed comparison of Levallois core technology from eastern Africa, Arabia, and the Levant during MIS 5, including multiple sites associated with Homo sapiens fossils. We employ quantitative comparisons of individual artefacts that provides a detailed appraisal of Levallois reduction activity in MIS 5, thereby enabling assessment of intra- and inter-assemblage variability for the frst time. Our results demonstrate a pattern of geographically structured variability embedded within a shared focus on centripetal Levallois reduction schemes and overlapping core morphologies. We reveal directional changes in core shaping and fake production from eastern Africa to Arabia and the Levant that are independent of diferences in geographic or environmental parameters. These results are consistent with a common cultural inheritance between these regions, potentially stemming from a shared late Middle Pleistocene source in eastern Africa., peer-reviewed

The long-standing debate concerning the integrity of the cultural taxonomies employed by archaeologists has recently been revived by renewed theoretical attention and the application of new methodological tools. The analyses presented here test the integrity of the cultural taxonomic division between Middle and Later Stone Age assemblages in eastern Africa using an extensive dataset of archaeological assemblages. Application of a penalized logistic regression procedure embedded within a permutation test allows for evaluation of the existing Middle and Later Stone Age division against numerous alternative divisions of the data. Results suggest that the existing division is valid based on any routinely employed statistical criterion, but that is not the single best division of the data. These results invite questions about what archaeologists seek to achieve via cultural taxonomy and about the analytical methods that should be employed when attempting revise existing nomenclature. Introduction Methods - Data Analyses - Weighted Binary log-F Regression - Permutation Analysis Results - Initial Analysis - Permutation Analysis Discussion Conclusions

The origin and evolution of hominin mortuary practices are topics of intense interest and debate1–3. Human burials dated to the Middle Stone Age (MSA) are exceedingly rare in Africa and unknown in East Africa1–6. Here we describe the partial skeleton of a roughly 2.5- to 3.0-year-old child dating to 78.3 ± 4.1 thousand years ago, which was recovered in the MSA layers of Panga ya Saidi (PYS), a cave site in the tropical upland coast of Kenya7,8. Recent excavations have revealed a pit feature containing a child in a flexed position. Geochemical, granulometric and micromorphological analyses of the burial pit content and encasing archaeological layers indicate that the pit was deliberately excavated. Taphonomical evidence, such as the strict articulation or good anatomical association of the skeletal elements and histological evidence of putrefaction, support the in-place decomposition of the fresh body. The presence of little or no displacement of the unstable joints during decomposition points to an interment in a filled space (grave earth), making the PYS finding the oldest known human burial in Africa. The morphological assessment of the partial skeleton is consistent with its assignment to Homo sapiens, although the preservation of some primitive features in the dentition supports increasing evidence for non-gradual assembly of modern traits during the emergence of our species. The PYS burial sheds light on how MSA populations interacted with the dead. Primary and intentional deposit Burial versus funerary caching Taxonomic assessment Implications for human cultural evolution Online content

The African Middle Stone Age (MSA, typically considered to span ca. 300–30 thousand years ago [ka]), represents our species’ first and longest lasting cultural phase. Although the MSA to Later Stone Age (LSA) transition is known to have had a degree of spatial and temporal variability, recent studies have implied that in some regions, the MSA persisted well beyond 30 ka. Here we report two new sites in Senegal that date the end of the MSA to around 11 ka, the youngest yet documented MSA in Africa. This shows that this cultural phase persisted into the Holocene. These results highlight significant spatial and temporal cultural variability in the African Late Pleistocene, consistent with genomic and palaeoanthropological hypotheses that significant, long-standing inter-group cultural differences shaped the later stages of human evolution in Africa. Introduction Results Discussion Materials and methods - Fieldwork - OSL dating methodology - Lithic analysis

The Middle Stone Age (MSA) corresponds to a critical phase in human evolution, overlapping with the earliest emergence of Homo sapiens as well as the expansions of these populations across and beyond Africa. Within the context of growing recognition for a complex and structured population history across the continent, Eastern Africa remains a critical region to explore patterns of behavioural variability due to the large number of well-dated archaeological assemblages compared to other regions. Quantitative studies of the Eastern African MSA record have indicated patterns of behavioural variation across space, time and from different environmental contexts. Here, we examine the nature of these patterns through the use of matrix correlation statistics, exploring whether differences in assemblage composition and raw material use correlate to differences between one another, assemblage age, distance in space, and the geographic and environmental characteristics of the landscapes surrounding MSA sites. Assemblage composition and raw material use correlate most strongly with one another, with site type as well as geographic and environmental variables also identified as having significant correlations to the former, and distance in time and space correlating more strongly with the latter. By combining time and space into a single variable, we are able to show the strong relationship this has with differences in stone tool assemblage composition and raw material use, with significance for exploring the impacts of processes of cultural inheritance on variability in the MSA. A significant, independent role for terrain roughness for explaining variability in stone tool assemblages highlights the importance of considering the impacts of mobility on structuring the archaeological record of the MSA of Eastern Africa. Introduction What explains difference in stone tool composition Datasets and methods Results - Simple Mantel tests - Multiple matrix regressions Discussion Conclusions

We study human origins from the related disciplines of archaeology, palaeoanthropology and allied geosciences, which are driven forward by making new discoveries in the field. Collectively, our scientific ecosystem informs climate change policy, heritage management and conservation practices. The pandemic has forced an inevitable pause on our work, but several scientific studies now indicate that the severe acute respiratory syndrome coronavirus 2 (SARS-CoV-2), the causative agent of coronavirus disease 2019 (COVID-19), may be here to stay. Critically for fieldwork and related travel considerations, the pandemic will increasingly display temporally and spatially fragmented peaks and troughs7. Complicating this is the fact that affected countries are not consistent in their public health decisions and application of World Health Organization (WHO) guidance, particularly in terms of easing containment measures. Regional and country-based variations in the availability and quality of appropriate healthcare also underline the importance of protecting at risk communities. [excerpt], peer-reviewed

Archaeobotanical evidence from two Early Historic sites in Sri Lanka, Kantharodai and Kirinda, is reported, providing significant evidence for agricultural diversity beyond the cultivation of rice. These data highlight the potential of systematic archaeobotanical sampling for macro-remains in tropical environments to contribute to the understanding of subsistence history in the tropics. Direct AMS radiocarbon dating confirms both the antiquity of crops and refines site chronologies. Both sites have Oryza sativa subsp. indica rice and evidence of rice crop-processing and millet farming. In addition, phytolith data provide complementary evidence on the nature of early rice cultivation in Sri Lanka. Both Kantharodai and Kirinda possess rice agriculture and a diverse range of cultivated millets (Brachiaria ramosa, Echinochloa frumentacea, Panicum sumatrense, and Setaria verticillata). Pulses of Indian origin were also cultivated, especially Vigna radiata and Macrotyloma uniflorum. Cotton (Gossypium sp.) cultivation is evident from Kirinda. Both sites, but in particular Kirinda, provide evidence for use of the seeds of Alpinia sp., in the cardamom/ginger family (Zingiberaceae), a plausible wild spice, while coconuts (Cocos nucifera) were also found at Kirinda.

The Middle Stone Age (MSA) of eastern Africa has a long history of research and is accompanied by a rich fossil record, which, combined with its geographic location, have led it to play an important role in investigating the origins and expansions of Homo sapiens. Recent evidence has suggested an earlier appearance of our species, indicating a more mosaic origin of modern humans, highlighting the importance of regional and inter-regional patterning and bringing into question the role that eastern Africa has played. Previous evaluations of the eastern African MSA have identified substantial variability, only a small proportion of which is explained by chronology and geography. Here, we examine the structure of behavioural, temporal, geographic and environmental variability within and between sites across eastern Africa using a quantitative approach. The application of hierarchical clustering identifies enduring patterns of tool use and site location through the MSA as well as phases of significant behavioural diversification and colonisation of new landscapes, particularly notable during Marine Isotope Stage 5. As the quantity and detail of technological studies from individual sites in eastern Africa gathers pace, the structure of the MSA record highlighted here offers a roadmap for comparative studies.

The terminology used to describe Palaeolithic industries has an important impact upon our interpretation of past behaviour. In South Asia, the term Late Palaeolithic is employed to refer to Late Pleistocene microlithic industries, distinguishing them chronologically from Holocene Mesolithic industries, and technologically from preceding Middle Palaeolithic technologies. Historically, however, an intermediate technological stage between Middle Palaeolithic and microlithic industries has been recognised and called ‘Upper’ Palaeolithic. Examining whether these ‘Upper’ Palaeolithic industries fit contemporary definitions of Middle or Late Palaeolithic technologies, distinct diversity within one of these technologies or a transitional phase between the two is therefore necessary to reintegrate these ‘Upper’ Palaeolithic sites into current debate. This is a particularly timely issue as the connection between some Late Palaeolithic artefact types, particularly backed microliths, and the earliest modern human populations in South Asia no longer appears tenable, and thus a rush to identify the earliest appearance of microliths must give way to more detailed examinations of behavioural variability. This paper re-examines lithic assemblages from Buddha Pushkar, western India, originally reported as an ‘Upper’ Palaeolithic industry. An attribute study of metric and categorical variables recorded on stone tools is used to examine how flaking technology, raw material use and reduction intensity vary within and between these assemblages. The results indicate raw material choices had a marked impact on stone tool technologies, nested within a pattern of technological diversity in western India during the terminal Pleistocene that complement models of regional trajectories in the evolution of Late Palaeolithic technologies.

Examinations of modern human dispersals are typically focused on expansions from South, East or North Africa into Eurasia, with more limited attention paid to dispersals within Africa. The paucity of the West African fossil record means it has typically been overlooked in appraisals of human expansions in the Late Pleistocene, yet regions such as Senegal occur in key biogeographic transitional zones that may offer significant corridors for human occupation and expansion. Here, we report the first evidence for Middle Stone Age occupation of the West African littoral from Tiemassas, dating to ∼44 thousand years ago, coinciding with a period of enhanced humidity across the region. Prehistoric populations mainly procured raw material from exposed Ypresian limestone horizons with Levallois, discoidal and informal reduction sequences producing flake blanks for retouched tools. We discuss this mid-Marine Isotope Stage 3 occupation in the context of the site's unique, ecotonal position amongst Middle Stone Age sites across West Africa, and its significance for Later Stone Age colonization of near coastal forests in the region. The results also support previous suggestions for connections between Middle Stone Age populations in West Africa and the Maghreb, for which the coastline may also have played a significant role.

Climate change is frequently highlighted as a key driver of biological evolution and cultural innovation in our species. It is often seen as influencing behavioural plasticity and the development of buffering mechanisms, for example in the form of more efficient technology and subsistence strategies. However, such hypotheses are yet to be studied in detail in South Asia, despite improving Late Pleistocene palaeoenvironmental records in this region and its crucial position in human dispersals beyond Africa. Here, we review evidence for technological and behavioural innovation across three regions of South Asia: the Thar Desert (north-west India), the Jurreru River Valley (south-east India), and the lowland Wet Zone of Sri Lanka. Together these areas form an ecotone from hyper-arid desert to humid rainforest that show different dynamics in the Late Pleistocene, and particularly during the Last Glacial Maximum. The archaeological records from each of these areas demonstrate a distinct nature and tempo of cultural change, probably reflecting, to some extent, the influence of climate change on forming heterogeneous local environments. Overall, however, the mosaic environments of South Asia made it an attractive region for the persistence of our species and their gradual uptake of cultural innovations during the Late Pleistocene.

The Indian Ocean coastline is argued to have been a critical route of modern human dispersal from Africa, introducing Late Palaeolithic industries into South Asia, but a dearth of evidence has prevented a direct evaluation of this. Kachchh (Gujarat, India), located immediately east of the Indus Delta, is an important setting to appraise the Palaeolithic occupation of the western Indian coastline. Targeted survey of Late Pleistocene sediments there found widespread evidence for occupation of Kachchh during the Late Pleistocene: Middle Palaeolithic and possibly Late Acheulean lithic artifacts. The conspicuous absence of Late Palaeolithic industries in these Late Pleistocene deposits, with their presence only noted in Holocene contexts, does not support the model of modern human expansions into India with these industries via the coastal route. We evaluate these results in the context of current debates regarding Late Pleistocene hominin demography, adaptation, and expansions.

Late Pleistocene palaeoenvironments in the Thar Desert (India), located at the eastern extent of the Saharo-Arabian desert belt, have fluctuated considerably as a result of the varying range and intensity of the Indian summer monsoon. Phases of widespread Pleistocene aridity are well documented in the Thar Desert, but research focusing on humid proxies is critical to examine how the region may have facilitated population expansions across southern Asia. At Katoati, located on the northeast margin of the Thar Desert, the combination of field recording of sediment sections with detailed analyses (micromorphology, stable isotope, loss on ignition, magnetic susceptibility, and X-ray fluorescence) from an archaeological site identify a series of hominin occupations during phases of enhanced humidity between ~96 and 60 ka. A gradient of humidity on the eastern margin of the Thar Desert during the late Pleistocene is identified, with the periodic humidity evident at Katoati occurring more frequently and with longer duration towards the southern margin. This uneven distribution of humidity in the Thar Desert is likely to have strongly influenced when and where hominin populations could expand into and across the region.

South Asia has a rich Palaeolithic heritage, and chronological resolution for this record has substantially improved over the past decade as a result of focused, interdisciplinary research at a number of key sites. Expanding the spatial diversity of dated Palaeolithic sites in South Asia grows increasingly important to examine how patterns of change through time vary within and between the region’s diverse habitats. Critically, alternate models of modern human dispersals into South Asia highlight the significance of either coastal or continental routes of dispersal, but currently no coastal Palaeolithic sites directly dating to the timeframe of human expansions are known. Our previous research in Kachchh was the first study to clearly identify the presence of Palaeolithic sites in Late Pleistocene landscapes in close proximity to the Indian Ocean coastline. Here, we present the first results of surface survey and test excavation at the site of Sandhav (Kachchh, India), approximately 25 km from the modern shoreline. We characterise the geomorphology of the landscape, highlighting multiple phases of alluvial aggradation and post-depositional carbonate formation, associated with Palaeolithic artefacts. To date, excavations have tested the uppermost Pleistocene deposit, yielding a small collection of fresh Middle Palaeolithic artefacts associated with a luminescence age dating to the first half of MIS 5 (∼114 ka), which provides a minimum age for Late Acheulean artefacts in underlying units. We discuss our findings in the context of debates surrounding the timing, lithic technologies, and ecologies associated with the expansions of modern humans into South Asia.

Microliths–small, retouched, often-backed stone tools–are often interpreted to be the product of composite tools, including projectile weapons, and efficient hunting strategies by modern humans. In Europe and Africa these lithic toolkits are linked to hunting of medium- and large-sized game found in grassland or woodland settings, or as adaptations to risky environments during periods of climatic change. Here, we report on a recently excavated lithic assemblage from the Late Pleistocene cave site of Fa-Hien Lena in the tropical evergreen rainforest of Sri Lanka. Our analyses demonstrate that Fa-Hien Lena represents the earliest microlith assemblage in South Asia (c. 48,000–45,000 cal. years BP) in firm association with evidence for the procurement of small to medium size arboreal prey and rainforest plants. Moreover, our data highlight that the lithic technology of Fa-Hien Lena changed little over the long span of human occupation (c. 48,000–45,000 cal. years BP to c. 4,000 cal. years BP) indicating a successful, stable technological adaptation to the tropics. We argue that microlith assemblages were an important part of the environmental plasticity that enabled Homo sapiens to colonise and specialise in a diversity of ecological settings during its expansion within and beyond Africa. The proliferation of diverse microlithic technologies across Eurasia c. 48–45 ka was part of a flexible human ‘toolkit’ that assisted our species’ spread into all of the world’s environments, and the development of specialised technological and cultural approaches to novel ecological situations.

Chronometrically dated Pleistocene records of human occupations of West Africa are rare but offer critical information with which to explore patterns of human origins and adaptation both within the region and more widely across the continent. A number of Middle Stone Age sites are known from the larger river valleys of West Africa, but recent work at the site of Tiemassas has highlighted the presence of Late Pleistocene occupations of the West African coastline. Tiemassas is the westernmost Middle Stone Age site known from Africa, and is located at the interface of Sudanian savannahs, Guinean forest-savannah mosaics and mangrove habitats. Research in the 20th century identified rich collections of MSA stone tools at the site, and our earlier excavations at Tiemassas have constrained an occupation to ca. 44 thousand years ago. Here, we present the results of further chronological dating and detailed analyses of lithic assemblages from new excavations at the site. We synthesize these new findings with the wider suite of evidence from the site to characterize MSA occupations of the West African coast to between 62 and 25 thousand years ago. Our results suggest considerable technological continuity in stone tool technologies at Tiemassas. Despite the immediate proximity of the coastline, there is little evidence at present to suggest direct engagement with coastal resources by MSA populations. Rather, the ecotonal position of the site, alongside more diffuse benefits of its coastal position, may have provided an attractive context for Middle Stone Age occupation.

The Thar Desert, located in western South Asia, marks a major global biogeographic boundary and a major adaptive threshold for the eastward expansions of modern humans from Africa across Asia. Examining the framework of palaeoenvironmental change in this region, both in terms of the regional manifestation of global climate change and the physical geography of the landscape, is therefore important to understand how modern humans first engaged with this significant shift in ecology. Here, we present evidence for the oldest chronometrically dated evidence for fluvial activity within this region, located at the Nal Quarry site in the central Thar Desert. We use luminescence dating of alluvial facies to demonstrate phases of fluvial activity at the site at ∼172–174, 140–150, 79–95 and 26 thousand years ago. This result substantially extends existing evidence for fluvial activity within the Thar Desert, as well as overlapping with evidence from the southern and eastern Thar desert indicating increased fluvial activity during the Last Interglacial (Marine Isotope Stage 5), whereas the cessation of fluvial deposition at Nal Quarry is contemporaneous with the onset of activity within the Ghaggar-Hakkra channel in the northern and western Thar Desert. Critically, the phases of fluvial activity identified at Nal overlaps with substantial behavioural change across South Asia, as well as the wider expansion of modern humans across the continent. This research illuminates a dynamic fluvial landscape that existed in the late Middle Pleistocene and early Late Pleistocene at a key threshold for modern human dispersals.

Sri Lanka has produced the earliest clear evidence for Homo sapiens fossils in South Asia and research in the region has provided important insights into modern human adaptations and cultural practices during the last ca. 45,000 years. However, in-depth multidisciplinary analyses of Late Pleistocene and Holocene sequences remain limited to just two sites, Fa Hien-lena and Batadomba-lena. Here, we present our findings from the reinvestigation of a third site, Kitulgala Beli-lena. New chronometric dating from the site confirms the presence of humans as early as ca. 45,000 cal. BP. in the island’s Wet Zone rainforest region. Our analyses of macrobotanical, molluscan, and vertebrate remains from the rockshelter show that this early human presence is associated with rainforest foraging. The Late Pleistocene deposits yielded evidence of wild breadfruit and kekuna nut extraction while the Holocene layers reveal a heavy reliance on semi-arboreal and arboreal small mammals as well as freshwater snails as a protein source. The lithic and osseous artefacts demonstrate that populations developed a sophisticated tool kit for the exploitation of their immediate landscapes. We place the rich Kitulgala Beli-lena dataset in its wider Sri Lankan context of Late Pleistocene foraging, as well as in wider discussions of our species’ adaptation to ‘extreme’ environments as it moved throughout Asia.

The ecological adaptations that stimulated the dispersal and technological strategies of our species during the Late Pleistocene remain hotly disputed, with some influential theories focusing on grassland biomes or marine resources as key drivers behind the rapid expansion and material culture innovations of Homo sapiens within and beyond Africa. Here, we present novel chronologically resolved, zooarchaeological taxonomic and taphonomic analysis, and stable isotope analysis of human and faunal tooth enamel, from the site of Panga ya Saidi (c. 78–0.4 ka), Kenya. Zooarchaeological data provide rare insights into the fauna associated with, and utilized by, Late Pleistocene-Holocene human populations in tropical coastal eastern Africa. Combined zooarchaeological and faunal stable isotope data provide some of the only dated, ‘on-site’ archives of palaeoenvironments beyond the arid interior of eastern Africa for this time period, while stable isotope analysis of humans provides direct snapshots of the dietary reliance of foragers at the site. Results demonstrate that humans consistently utilized tropical forest and grassland biomes throughout the period of site occupation, through a transition from Middle Stone Age to Later Stone Age technological industries and the arrival of agriculture in the region. By contrast, while coastal resources were obtained for use in symbolic material culture, there is limited evidence for consumption of marine resources until the Holocene. We argue that the ecotonal or heterogeneous environments of coastal eastern Africa may have represented an important refugium for populations during the increasing climatic variability of the Late Pleistocene and Holocene, and that tropical environments were one of a diverse series of ecosystems exploited by H. sapiens in Africa at the dawn of global migrations.

The Acheulean is the longest lasting cultural–technological tradition in human evolutionary history. However, considerable gaps remain in understanding the chronology and geographical distribution of Acheulean hominins. We present the first chronometrically dated Acheulean site from the Arabian Peninsula, a vast and poorly known region that forms more than half of Southwest Asia. Results show that Acheulean hominin occupation expanded along hydrological networks into the heart of Arabia from Marine Isotope Stage (MIS) 7 until at least ~190 ka ̶ the youngest documented Acheulean in Southwest Asia. The site of Saffaqah features Acheulean technology, characterized by large flakes, handaxes and cleavers, similar to Acheulean assemblages in Africa. These findings reveal a climatically-mediated later Acheulean expansion into a poorly known region, amplifying the documented diversity of Middle Pleistocene hominin behaviour across the Old World and elaborating the terminal archaic landscape encountered by our species as they dispersed out of Africa., peer-reviewed

Despite occupying a central geographic position, investigations of hominin populations in the Arabian Peninsula during the Lower Palaeolithic period are rare. The colonization of Eurasia below 55 degrees latitude indicates the success of the genus Homo in the Early and Middle Pleistocene, but the extent to which these hominins were capable of innovative and novel behavioural adaptations to engage with mid-latitude environments is unclear. Here we describe new field investigations at the Saffaqah locality (206–76) near Dawadmi, in central Arabia that aim to establish how hominins adapted to this region. The site is located in the interior of Arabia over 500 km from both the Red Sea and the Gulf, and at the headwaters of two major extinct river systems that were likely used by Acheulean hominins to cross the Peninsula. Saffaqah is one of the largest Acheulean sites in Arabia with nearly a million artefacts estimated to occur on the surface, and it is also the first to yield stratified deposits containing abundant artefacts. It is situated in the unusual setting of a dense and well-preserved landscape of Acheulean localities, with sites and isolated artefacts occurring regularly for tens of kilometres in every direction. We describe both previous and recent excavations at Saffaqah and its large lithic assemblage. We analyse thousands of artefacts from excavated and surface contexts, including giant andesite cores and flakes, smaller cores and retouched artefacts, as well as handaxes and cleavers. Technological assessment of stratified lithics and those from systematic survey, enable the reconstruction of stone tool life histories. The Acheulean hominins at Dawadmi were strong and skilful, with their adaptation evidently successful for some time. However, these biface-makers were also technologically conservative, and used least-effort strategies of resource procurement and tool transport. Ultimately, central Arabia was depopulated, likely in the face of environmental deterioration in the form of increasing aridity., peer-reviewed

Despite occupying a central geographic position, investigations of hominin populations in the Arabian Peninsula during the Lower Palaeolithic period are rare. The colonization of Eurasia below 55 degrees latitude indicates the success of the genus Homo in the Early and Middle Pleistocene, but the extent to which these hominins were capable of innovative and novel behavioural adaptations to engage with mid-latitude environments is unclear. Here we describe new field investigations at the Saffaqah locality (206-76) near Dawadmi, in central Arabia that aim to establish how hominins adapted to this region. The site is located in the interior of Arabia over 500 km from both the Red Sea and the Gulf, and at the headwaters of two major extinct river systems that were likely used by Acheulean hominins to cross the Peninsula. Saffaqah is one of the largest Acheulean sites in Arabia with nearly a million artefacts estimated to occur on the surface, and it is also the first to yield stratified deposits containing abundant artefacts. It is situated in the unusual setting of a dense and well-preserved landscape of Acheulean localities, with sites and isolated artefacts occurring regularly for tens of kilometres in every direction. We describe both previous and recent excavations at Saffaqah and its large lithic assemblage. We analyse thousands of artefacts from excavated and surface contexts, including giant andesite cores and flakes, smaller cores and retouched artefacts, as well as handaxes and cleavers. Technological assessment of stratified lithics and those from systematic survey, enable the reconstruction of stone tool life histories. The Acheulean hominins at Dawadmi were strong and skilful, with their adaptation evidently successful for some time. However, these biface-makers were also technologically conservative, and used least-effort strategies of resource procurement and tool transport. Ultimately, central Arabia was depopulated, likely in the face of environmental deterioration in the form of increasing aridity.

The dispersal of Homo sapiens out of Africa has been extensively researched across several disciplines. Here we review the evidence for spatial and temporal variability in lithic (stone tool) technologies relative to the predictions of two major hypotheses: 1) that a single successful dispersal occurred 60–50 thousand years ago (ka), marked by a trail of geometric/microlithic technologies, and 2) that multiple dispersals occurred, beginning much earlier (probably in Marine Isotope Stage [MIS] 5), associated with Middle Palaeolithic technology in its early phase. Our results show that Late Pleistocene geometric/microlithic technologies exhibit significant temporal and regional differences between each other. These differences suggest independent, convergent origins for these technologies, which are likely to have been repeatedly re-invented. In contrast, we identify similarities between East African lithic technologies from MIS 8 onwards and Middle Palaeolithic assemblages as far east as India by MIS 5. That this constellation of technological features – particularly an emphasis on centripetal Levallois reduction reflecting interchangeable preferential and recurrent methods, along with particular retouched forms such as points – transcends ecologies and raw material types suggests that it is unlikely to entirely reflect technological convergence (analogy). Our results indicate an early onset of multiple dispersals out of Africa. The hypothesis of an early onset to successful dispersal is entirely consistent with the possibility of further subsequent (post-MIS 5) dispersals out of Africa. Testing such hypotheses through quantified comparative lithic studies and interdisciplinary research is therefore likely to significantly advance understanding of the earliest H. sapiens dispersals.