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Sycorax sinuosa Curler sp. nov. (Figs. 19���21) DIAGNOSIS. Male terminalia: parameres with posterior lobes digitiform, directed medially; dorsomedial process of paramere short, not extending close to apices of aedeagus; medial processes of parameres sinuous, bulbous apically; aedeagus with apices acuminate. Female terminalia: spermathecal ducts inflated, ovoid, lightly annulated apically; sternum 10 triangular. DESCRIPTION. Male (Figs. 19���21): Measurements (n = 2), head width 0.28 mm, head length 0.20 mm, palpus length 0.09 mm, wing length 1.34 mm, wing width 0.55 mm, R 2 + 3 length 0.38 mm, R 3 length 0.28 mm, gonocoxite length 0.12 mm, gonostylus length 0.09 mm, gonostylus spine length 0.05 mm. Head elliptical in frontal view, wider than long. Antennae: scape and pedicel sub-spherical, only flagellomere 1 present in one specimen, with ascoids. Wing: radial fork with R 2 + 3 about 1.25 times longer than R 3. Terminalia: gonostylus with subterminal seta closer to apical spine; ejaculatory apodeme laterally compressed; aedeagus with apices narrow, blade-like, acuminate; parameres with posterior lobes digitiform, directed medially; dorsomedial process of paramere slightly wider than ejaculatory apodeme, slightly longer than wide, not extending close to apices of aedeagus; medial processes of parameres sinuous, bulbous apically, extending to apices of aedeagus; epandrium with setae alveoli conspicuous. Female: head as in male except flagellomere 1 approximately 1.5 times the length of flagellomere 2, ascoids present only on flagellomeres 1���8; mouthparts undeveloped. Terminalia: spermathecal ducts inflated, ovoid, lightly annulated apically; spermathecae spherical; genital fork about 1.5 times longer than broad, darkened with pigment; sternum 10 triangular. TYPE MATERIAL. Holotype [male]: New Caledonia. NORD PROVINCE: R��serve speciale de faune de l���Aoupini��, in alpine forest, 21 �� 10 ��� 40 ���S 165 �� 18 ��� 10 ���E, 700m, 30.xi��� 9.xii. 2000, coll. E. Schlinger, M. Irwin and J. Boutin, Malaise trap; deposited MNHN. Specimen dissected, mounted on micro-slide. Paratypes [1 male, 1 female]: same data as holotype; deposited in LACM. ETYMOLOGY. The species epithet sinuosa from Latin is in reference to the sinuous shape of the medial processes of the parameres. BIONOMICS. Unknown. DISTRIBUTION. Collected from one location in New Caledonia. COMMENTS. Males of Sycorax sinuosa are similar to those of S. spina, but can be separated by the shape of the dorsomedial and medial processes of the parameres. S. sinuosa is the only species of Sycorax known from New Caledonia that has a short dorsomedial process with sinuous medial processes. The single female specimen of S. sinuosa is tenuously associated with the male based on similarities of the head and wing. Ascoids are present only on some flagellomeres in the female. It is clear that ascoids are absent from the remaining flagellomeres and have not simply been broken or removed. All specimens of this species were collected in a location where there are no streams nearby., Published as part of Curler, Gregory R. & Jacobson, Amanda J., 2012, New species of Psychodidae (Diptera) from Australasia, with a checklist of the world species of Bruchomyiinae and Sycoracinae, pp. 43-65 in Zootaxa 3552 on pages 53-54, DOI: 10.5281/zenodo.282855

Sycorax dispar Satchell, 1950 (Figs. 30���31) DIAGNOSIS. Male terminalia: ejaculatory apodeme dorsoventrally compressed, clavate; parameres with posteromedial process digitiform, posterior lobes sinuous with pointed hooks apically; aedeagus with branches sickle-shaped, each with a single dens laterally. REDESCRIPTION. Male (Figs. 30���31): Wing: radial fork about 2 / 3 as long as medial fork. Terminalia: gonostylus with subterminal seta as long as apical spine, inserted near the base of style; ejaculatory apodeme dorsoventrally compressed, clavate; aedeagus with branches sickle-shaped, each with a single dens laterally; parameres with posterior lobes sinuous with pointed hooks apically; dorsomedial process of paramere digitiform, extending slightly beyond apices of aedeagus. TYPE MATERIAL. Holotype [adult male]: New Zealand. NORTH ISLAND: Nihotupu, 23.ii. 1923, coll. A.L. Tonnoir; deposited NZAC. DISTRIBUTION. Collected from one location in northern New Zealand. COMMENTS. Satchell (1950) described four species of Sycorax from New Zealand. Of these four species, S. dispar is the only one that is very similar to, and could potentially be confused with some species collected from New Caledonia, especially S. webbi. S. dispar is redescribed and illustrated here in order to clarify differences between it and S. webbi, particularly the dorsoventally compressed ejaculatory apodeme in the former species compared to the laterally compressed ejaculatory apodeme in the latter species. Females of S. dispar were included in the type series, but not described in detail by Satchell (1950). Regardless, specimens were dissected and mounted in a way that made it impossible to examine pertinent characters of the female genitalia. Therefore, they were not redescribed here., Published as part of Curler, Gregory R. & Jacobson, Amanda J., 2012, New species of Psychodidae (Diptera) from Australasia, with a checklist of the world species of Bruchomyiinae and Sycoracinae, pp. 43-65 in Zootaxa 3552 on pages 60-61, DOI: 10.5281/zenodo.282855, {"references":["Satchell, G. H. (1950) The New Zealand Psychodidae: A study based upon the collection and manuscript notes of the late Dr. A. L. Tonnoir. Transactions of the Royal Entomological Society of London, 101 (5), 147 - 178."]}

Genus Sycorax Haliday in Curtis, 1839 Sycorax Haliday in Curtis, 1839: 745. Type species: Sycorax silacea Haliday in Curtis, 1839, by monotypy. Microdixa M��ller, 1927: 535. Type species: Microdixa scutigera M��ller, 1927, by original designation., Published as part of Curler, Gregory R. & Jacobson, Amanda J., 2012, New species of Psychodidae (Diptera) from Australasia, with a checklist of the world species of Bruchomyiinae and Sycoracinae, pp. 43-65 in Zootaxa 3552 on page 51, DOI: 10.5281/zenodo.282855, {"references":["Curtis, J. (1839) British entomology: being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Volume 16. London.","Muller, G. W. (1927) Microdixa nov. gen. der Dixidae und ihre Verwandlung. Zeitschrift fur Morphologie und Okologie der Tiere, A, 7, 535 - 542."]}

Sycorax furca Curler sp. nov. (Figs. 16���18) DIAGNOSIS. Male terminalia: parameres with posterior lobes tapered to acute apices, without apical hooks; dorsomedial process of paramere bipartite, with dorsal and ventral divisions; dorsal division digitiform, extending slightly beyond aedeagus; ventral division bifurcate apically, about equal in length to aedeagus; aedeagus with apices broadly triangular. DESCRIPTION. Male (Figs. 16���18): Measurements (n= 2), head width 0.30 mm, head length 0.21 mm, palpus length 0.08 mm, wing length 1.22 mm, wing width 0.51 mm, R 2 + 3 length 0.33 mm (0.32���0.34), R 3 length 0.28 mm (0.27���0.30), gonocoxite length 0.11 mm, gonostylus length 0.08 mm, gonostylus spine length 0.07 mm. Head broadly rounded in frontal view. Antennae: scape and pedicel sub-spherical, flagellomeres missing in all specimens. Wing: radial fork with R 2 + 3 about 1.5 times longer than R 3. Terminalia: gonostylus with subterminal seta closer to base of apical spine; ejaculatory apodeme laterally compressed; aedeagus with apices broadly triangular; parameres with posterior lobes straight, tapered to acute apices, without apical hooks; dorsomedial process of paramere bipartite, with dorsal and ventral divisions; dorsal division digitiform, extending slightly beyond aedeagus; ventral division bifurcate apically, about equal in length to aedeagus; epandrium with setae alveoli sparse, inconspicuous. Female: Unknown. TYPE MATERIAL. Holotype [male]: New Caledonia. SUD PROVINCE: 17 km NNE Noume��, Mt. Koghis, 22 �� 10 ��� 33 ���S 166 �� 30 ��� 17 ���E, 550m, 13���24.xi. 2000, coll. E. Schlinger and J. Boutin, Malaise trap; deposited MNHN. Specimen dissected, mounted on micro-slide. Paratypes [2 males]: same data as holotype; deposited USNM and LACM. ETYMOLOGY. The species epithet furca is a Latin word meaning ���fork���, in reference to the shape of the ventral division of the dorsomedial process of the parameres. BIONOMICS. Unknown. DISTRIBUTION. Collected from one location in New Caledonia. COMMENTS. Males of Sycorax furca are easily distinguished from those of other Sycorax known to occur in New Caledonia. The bifurcate dorsomedial process of the parameres and the broad triangular apices of the aedeagus are unique to this species. Specimens of S. furca were collected in a forested area within a botanical garden. The stream nearest this site is approximately 1 / 2 mile away., Published as part of Curler, Gregory R. & Jacobson, Amanda J., 2012, New species of Psychodidae (Diptera) from Australasia, with a checklist of the world species of Bruchomyiinae and Sycoracinae, pp. 43-65 in Zootaxa 3552 on pages 52-53, DOI: 10.5281/zenodo.282855

Sycorax tridentata Curler sp. nov. (Figs. 32���34) DIAGNOSIS. Male terminalia: parameres with posterior lobes rounded, hooked apically; dorsomedial process of paramere digitiform, bulbous apically, extending far beyond apices of aedeagus; aedeagus with branches tridentate posterolaterally. Female terminalia: spermathecal ducts inflated, nearly globular, not annulated. DESCRIPTION. Male (Figs. 32���34): Measurements, head width 0.27 mm, head length 0.20 mm, palpus length 0.09 mm, wings incomplete, R 2 + 3 length 0.31 mm, R 3 length 0.30 mm, gonocoxite length 0.09 mm, gonostylus length 0.07 mm, gonostylus spine length 0.04 mm. Head broadly rounded in frontal view, slightly more wide than long. Antennae: scape and pedicel sub-spherical, flagellomeres missing. Wing: radial fork relatively long; R 2 + 3 approximately equal in length to R 3. Terminalia: gonostylus with subterminal seta inserted approximately half way between base and apical spine; ejaculatory apodeme laterally compressed; vasa deferentia nearly as long as ejaculatory apodeme; testes ovate difficult to differentiate from vasa deferentia; ejaculatory duct funnel-shaped, expanding from base to apex; aedeagus with branches tridentate posterolaterally; parameres with thorn-like processes internally; posterior lobes tapered, rounded, hooked apically; dorsomedial process of paramere digitiform, bulbous apically, extending far beyond apices of aedeagus; epandrium with setae alveoli sparse, inconspicuous. Female: head as in male except with ascoids present on at least flagellomeres 1���6, remaining flagellomeres absent; mouthparts undeveloped. Wing as in male Terminalia: spermathecal ducts inflated, nearly globular, not annulated; spermathecae spherical; genital fork lightly sclerotized, with little pigment; sternite 10 bell-shaped in dorsal view. TYPE MATERIAL. Holotype [male]: New Caledonia. SUD PROVINCE: 17 km NNE Noume��, Mt. Koghis, 22 �� 10 ��� 33 ���S 166 �� 30 ��� 17 ���E, 550m, 13���24.xi. 2000, coll. E. Schlinger and J. Boutin, Malaise trap; deposited MNHN. Specimen dissected, mounted on micro-slide. Paratypes [2 females]: same location as holotype, 1���5.xii. 2000, coll. M. Irwin; deposited LACM. ETYMOLOGY. The species epithet tridentata is Latin, meaning ���three teeth���, in reference to the three toothlike projections on the distal branches of the aedeagus. BIONOMICS. Unknown. DISTRIBUTION. Collected from one location in New Caledonia. COMMENTS. Sycorax tridentata was collected from a location that is approximately �� mile away from the nearest stream. This species is described based on a single male specimen and two tenuously associated female specimens. The practice of describing a species based on an examination of only a single male specimen is generally not satisfactory. However, it is highly unlikely that additional material of S. tridentata will be collected in the foreseeable future, and there is no question that it is a distinct species. It is similar to S. webbi, but differs in the shape of the dorsomedial process of the parameres and the shape of the aedeagus. S. tridentata has a dorsomedial process with a bulbous apex while S. webbi has no bulbous apex. The presumed females of S. tridentata are similar to females of S. spina in the shape and pigmentation of the genital fork, but differ in having spermathecal ducts that are nearly globular as opposed to ovoid. Furthermore, females of S. tridentata are also similar to those of S. webbi in the globular shape of the spermathecal ducts but can be differentiated by comparison of the genital fork, which is darkly pigmented and inverted Y-shaped in S. webbi., Published as part of Curler, Gregory R. & Jacobson, Amanda J., 2012, New species of Psychodidae (Diptera) from Australasia, with a checklist of the world species of Bruchomyiinae and Sycoracinae, pp. 43-65 in Zootaxa 3552 on pages 56-58, DOI: 10.5281/zenodo.282855