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Ischnosoma quadriguttatum quadriguttatum (Champion, 1923) Figs 28, 45. NEPAL: Baglung: Dhaulagiri, Baglung Lekh, 10���15 km NW Baglung, 2350���2550 m, 28��18'36''N, 83��31'56''E, 10.���12.V.2004, leg. J. Schmidt, 10 exs. (cKle, cSch); Bhojpur: Valley NW Phedi, 1900 m, 25.V.1997, leg. W. Schawaller, 1 ex. (SMNS); Dailekh: Talpokhari S Dailekh, 1800 m, 29.V.1988, leg. W. Schawaller, 1 ex. (SMNS); Dhawalagi: Annapurna South Himal, SE of Narcheng, left riverside of Bele Khola, 2300 m, 28��30'40''N, 83��41'33''E, 25./ 26.V.2001, leg. G. Hirthe, 1 ex. (cHir); Dolakha: Rolwaling Himal, Lapchi Kang, range below Ting Sang La, 2300���2500 m, 1.IX.1999, leg. J. Schmidt, 1 ex. (NME); Dolpa���Mustang���Myagdi: right banks of Lethe Khola near Lethe, 2400 m, 5.���7.V.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); S slope, Dhaulagiri mountains, N Banduk, 2400���2600 m, 28��28'35''N, 83��35'05''E, 8.V.2009, leg. J. Schmidt, 8 exs. (NME); Dhaulagiri mountains, upper Marang Khola valley., 28��29'50''N, 83��27'37''E, 2500���2700 m, 16.V.2009, leg. J. Schmidt, 1 ex. (NME), 1 ex. (cSch); Dhaulagari Himal, S-slope, N Banduk village, 1900���2300 m, 28��27'22''N, 83��35'13''E to 28��28'07''N, 83��35'10''E, 6.V.2009, leg. J. Schmidt, 4 exs. (NME), 1 ex. (cSch); SW Dhaulagiri Himal, Dhara Khola Vali, 28��30'36''N, 83��18'16''E, 1900 m, 21.���22.V.2012, leg. J. Schmidt, 6 exs. (NME); S slope, Dhaulagari mountains, above Pathlekharka, 2500���2700 m, 28��32'19''N, 83��29'25''E, 12.V.2009, leg. J. Schmidt, 2 exs. (NME); SW Dhaulagiri, Maraini, 2400���2800 m, 11.V.2012, 28��31'07''N, 83��15'49''E, leg. J. Schmidt, 2 exs. (NME); Gorkha: Darondi Khola, above Barpak, 3300��� 3000 m, 11.VIII.1983, rhododendron forest, leg. J. Martens & W. Schawaller [243], 1 ex. (SMNS); Darondi Khola, below Barpak to Doreni, 1100��� 900 m, 12.VIII.1983, forest remnants, leg. J. Martens & W. Schawaller [246], 1 ex. (SMNS); Buri Gandaki, Nyak to lower Chuling Khola valley, 2450���2870 m, 2.VIII.1983, Pinus excelsa, mountain pasture, leg. J. Martens & W. Schawaller [227], 1 ex. (SMNS); same data, but 2270���2450 m, 1 ex. (SMNS); Ilam: road Ilam to Birtamode, Karphok, 1700 m, bushes, agricultural area, 2.IV.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); Mai Pokhari, 2100���2200 m, 9.��� 10.IV.1988, Castanopsis forest remnant, leg. J. Martens & W. Schawaller [319], 1 ex. (SMNS); Kaski: above Dhumpus, Berlese, 2100 m, 8.���10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Dhumpus, Laubwald, 2100 m, 8.���10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Pothana, 2000 m, 27.��� 29.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); Pokhara, souther bank of Phewa Lake, creek, sifted, 800���900 m, 8.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Annapurna region, Khare to Pothana, rhododendron forest, 1950���2000 m, 10. V. 2001, leg. G. Hirthe, 1 ex. (cHir); Annapurna mountains, Tadapani to Chuile, 2450��� 2550 m, 4.V.1999, leg. C. Kr��ger & G. Hirthe, 1 ex. (cHir); Lalitpur: Mount Phulchoki, 1800���2000 m, 25.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); above Godawari, Kunako Khola, sifted, 1750 m, 6.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Lamjung: Nawronkot near Larjung, Thakhola area, 2900���3000 m, leg. H. Franz, 1 ex. (NHMW); Manang: Annapurna mountains, Bagarchhap���Temang, 2400���2700 m, 17.IV.1999, leg. C. Kr��ger & G. Hirthe, 1 ex. (cHir); Panchthar: Paniporua, 2300 m, 16.���20.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [328], 1 ex. (SMNS); Parbat: Annapurna mountains, Bhickok to Deurali, 2100 m, 7.V.1999, leg. C. Kr��ger & G. Hirthe, 3 exs. (cHir, cSch); Annapurna region, West Mardi Himal, above Deurali, 2100���2200 m, 28��18'57''N, 83��49'59''E, 10. V. 2001, leg. G. Hirthe, 4 exs. (cHir, cSch); Rasuwa: Bagmati province, Gul Ghanjyang, 2600 m, 6.IV.1981, leg. I. L��bl & A. Smetana, 1 ex. (MHNG); Sankhuwasabha: above Pahakhola, 2600���2800 m, 31.V.���3.VI.1988, Quercus semecarpifolia, Rhododendron, leg. J. Martens & W. Schawaller [404], 3 ex. (SMNS); Arun Valley, between Mure and Hurure, 2050���2150 m, 9.���17.VI.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [412], 2 ex. (SMNS); N Furure, floodplain near creek, 27��30'44''N, 87��16'14''E, 2100 m, 7.XII.1998, sifted, leg. M. Hartmann, 1 ex. (NME); Arun Valley, W above Tashigaon, 2500���2700 m, 26.��� 27.V.2014, 27��36'33''N, 87��13'28''E, leg. J. Schmidt, 1 ex. (NME); Arun Valley, near Tashigaon, 2200���2400 m, 12.+ 28.���29.V.2014, 27��37'N, 87��14'E, leg. J. Schmidt, 2 ex. (NME); Sindhupalchok: Barahbise region, Ting ��� Sang ��� La, leg. H. Franz, 1 ex. (NHMW); Solukhumbu: below Pangum, 2500 m, 14.���15.V.1997, leg. W. Schawaller, 3 exs. (SMNS); Umg. Sutje near Lughla, Khumbu, leg. H. Franz, 2 exs. (NHMW); Surkhet: Bheri Khola bridge, 500 m, 24.���25.V.1998, leg. W. Schawaller, 1 ex. (SMNS); N Surkhet, 1600���2000 m, 28.V.1998, leg. W. Schawaller, 1 ex. (SMNS); Syangja: Manaslu mountains, Dudh Pokhari Lekh, upper Dardi Khola valley, 2600��� 2300 m, 15.���17.IV.2003, leg. J. Schmidt, 2 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, upper Phulinagiri Madi, 2500 m, 19.���21.IV.2003, leg. J. Schmidt, 10 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, below Helam Pokhari, 2000 m, 22.IV.2003, leg. J. Schmidt, 1 ex. (NME); Manaslu mountains, S of Bara Pokhari, 2300 m, 8.IV.2003, leg. J. Schmidt, 24 exs. (NME, cSch); Manaslu mountains, Bara Pokhari lekh, Chandi Khola valley, 11.���12.IV.2003, 2000���2300 m, leg. J. Schmidt, 1 ex, (NME); Manaslu mountains, S of Bara Pokhari, 2000 m, 7.IV.2003, leg. J. Schmidt, 1 ex. (NME); Bara Pokhari Lekh, 2500 m, 2.IV.1999, leg. G. Hirthe & C. Kr��ger, 2 exs. (cHir, cSch); Taplejung: confluence of Kabeli and Tada Khola, 1000 m, 23.���25.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [344], 5 exs. (SMNS); ascent to pasture Lassetham from Omje Khola, 2400���3150 m, 6.V.1988, mixed Quercus - Tsuga - Rhodonendron forest, leg. J. Martens & W. Schawaller [357], 1 ex. (SMNS). Remarks. I. quadriguttatum quadriguttatum is widespread in Southeast Asia. In Nepal, it is the most common species and present everywhere (Fig. 45). A second subspecies, I. quadriguttatum japonicum Kocian, 2003 was described from Japan., Published as part of Kocian, Mat���� & Sch��lke, Michael, 2018, New species and records of Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) from Nepal, pp. 501-522 in Zootaxa 4442 (4) on pages 516-518, DOI: 10.11646/zootaxa.4442.4.1, http://zenodo.org/record/1304822, {"references":["Champion, G. C. (1923) Some Indian Coleoptera (10). The Entomologist's Monthly Magazine, 59, 43 - 53, 77 - 80.","Kocian, M. (2003) Monograph of the world species of the genus Ischnosoma (Coleoptera: Staphylinidae). Acta Universitatis Carolinae. Biologica, 47, 3 - 151."]}

Ischnosoma quadriguttatum quadriguttatum (Champion, 1923) Figs 28, 45. NEPAL: Baglung: Dhaulagiri, Baglung Lekh, 10–15 km NW Baglung, 2350–2550 m, 28°18'36''N, 83°31'56''E, 10.–12.V.2004, leg. J. Schmidt, 10 exs. (cKle, cSch); Bhojpur: Valley NW Phedi, 1900 m, 25.V.1997, leg. W. Schawaller, 1 ex. (SMNS); Dailekh: Talpokhari S Dailekh, 1800 m, 29.V.1988, leg. W. Schawaller, 1 ex. (SMNS); Dhawalagi: Annapurna South Himal, SE of Narcheng, left riverside of Bele Khola, 2300 m, 28°30'40''N, 83°41'33''E, 25./ 26.V.2001, leg. G. Hirthe, 1 ex. (cHir); Dolakha: Rolwaling Himal, Lapchi Kang, range below Ting Sang La, 2300–2500 m, 1.IX.1999, leg. J. Schmidt, 1 ex. (NME); Dolpa–Mustang–Myagdi: right banks of Lethe Khola near Lethe, 2400 m, 5.–7.V.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); S slope, Dhaulagiri mountains, N Banduk, 2400–2600 m, 28°28'35''N, 83°35'05''E, 8.V.2009, leg. J. Schmidt, 8 exs. (NME); Dhaulagiri mountains, upper Marang Khola valley., 28°29'50''N, 83°27'37''E, 2500–2700 m, 16.V.2009, leg. J. Schmidt, 1 ex. (NME), 1 ex. (cSch); Dhaulagari Himal, S-slope, N Banduk village, 1900–2300 m, 28°27'22''N, 83°35'13''E to 28°28'07''N, 83°35'10''E, 6.V.2009, leg. J. Schmidt, 4 exs. (NME), 1 ex. (cSch); SW Dhaulagiri Himal, Dhara Khola Vali, 28°30'36''N, 83°18'16''E, 1900 m, 21.–22.V.2012, leg. J. Schmidt, 6 exs. (NME); S slope, Dhaulagari mountains, above Pathlekharka, 2500–2700 m, 28°32'19''N, 83°29'25''E, 12.V.2009, leg. J. Schmidt, 2 exs. (NME); SW Dhaulagiri, Maraini, 2400–2800 m, 11.V.2012, 28°31'07''N, 83°15'49''E, leg. J. Schmidt, 2 exs. (NME); Gorkha: Darondi Khola, above Barpak, 3300– 3000 m, 11.VIII.1983, rhododendron forest, leg. J. Martens & W. Schawaller [243], 1 ex. (SMNS); Darondi Khola, below Barpak to Doreni, 1100– 900 m, 12.VIII.1983, forest remnants, leg. J. Martens & W. Schawaller [246], 1 ex. (SMNS); Buri Gandaki, Nyak to lower Chuling Khola valley, 2450–2870 m, 2.VIII.1983, Pinus excelsa, mountain pasture, leg. J. Martens & W. Schawaller [227], 1 ex. (SMNS); same data, but 2270–2450 m, 1 ex. (SMNS); Ilam: road Ilam to Birtamode, Karphok, 1700 m, bushes, agricultural area, 2.IV.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); Mai Pokhari, 2100–2200 m, 9.– 10.IV.1988, Castanopsis forest remnant, leg. J. Martens & W. Schawaller [319], 1 ex. (SMNS); Kaski: above Dhumpus, Berlese, 2100 m, 8.–10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Dhumpus, Laubwald, 2100 m, 8.–10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Pothana, 2000 m, 27.– 29.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); Pokhara, souther bank of Phewa Lake, creek, sifted, 800–900 m, 8.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Annapurna region, Khare to Pothana, rhododendron forest, 1950–2000 m, 10. V. 2001, leg. G. Hirthe, 1 ex. (cHir); Annapurna mountains, Tadapani to Chuile, 2450– 2550 m, 4.V.1999, leg. C. Krüger & G. Hirthe, 1 ex. (cHir); Lalitpur: Mount Phulchoki, 1800–2000 m, 25.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); above Godawari, Kunako Khola, sifted, 1750 m, 6.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Lamjung: Nawronkot near Larjung, Thakhola area, 2900–3000 m, leg. H. Franz, 1 ex. (NHMW); Manang: Annapurna mountains, Bagarchhap–Temang, 2400–2700 m, 17.IV.1999, leg. C. Krüger & G. Hirthe, 1 ex. (cHir); Panchthar: Paniporua, 2300 m, 16.–20.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [328], 1 ex. (SMNS); Parbat: Annapurna mountains, Bhickok to Deurali, 2100 m, 7.V.1999, leg. C. Krüger & G. Hirthe, 3 exs. (cHir, cSch); Annapurna region, West Mardi Himal, above Deurali, 2100–2200 m, 28°18'57''N, 83°49'59''E, 10. V. 2001, leg. G. Hirthe, 4 exs. (cHir, cSch); Rasuwa: Bagmati province, Gul Ghanjyang, 2600 m, 6.IV.1981, leg. I. Löbl & A. Smetana, 1 ex. (MHNG); Sankhuwasabha: above Pahakhola, 2600–2800 m, 31.V.–3.VI.1988, Quercus semecarpifolia, Rhododendron, leg. J. Martens & W. Schawaller [404], 3 ex. (SMNS); Arun Valley, between Mure and Hurure, 2050–2150 m, 9.–17.VI.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [412], 2 ex. (SMNS); N Furure, floodplain near creek, 27°30'44''N, 87°16'14''E, 2100 m, 7.XII.1998, sifted, leg. M. Hartmann, 1 ex. (NME); Arun Valley, W above Tashigaon, 2500–2700 m, 26.– 27.V.2014, 27°36'33''N, 87°13'28''E, leg. J. Schmidt, 1 ex. (NME); Arun Valley, near Tashigaon, 2200–2400 m, 12.+ 28.–29.V.2014, 27°37'N, 87°14'E, leg. J. Schmidt, 2 ex. (NME); Sindhupalchok: Barahbise region, Ting – Sang – La, leg. H. Franz, 1 ex. (NHMW); Solukhumbu: below Pangum, 2500 m, 14.–15.V.1997, leg. W. Schawaller, 3 exs. (SMNS); Umg. Sutje near Lughla, Khumbu, leg. H. Franz, 2 exs. (NHMW); Surkhet: Bheri Khola bridge, 500 m, 24.–25.V.1998, leg. W. Schawaller, 1 ex. (SMNS); N Surkhet, 1600–2000 m, 28.V.1998, leg. W. Schawaller, 1 ex. (SMNS); Syangja: Manaslu mountains, Dudh Pokhari Lekh, upper Dardi Khola valley, 2600– 2300 m, 15.–17.IV.2003, leg. J. Schmidt, 2 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, upper Phulinagiri Madi, 2500 m, 19.–21.IV.2003, leg. J. Schmidt, 10 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, below Helam Pokhari, 2000 m, 22.IV.2003, leg. J. Schmidt, 1 ex. (NME); Manaslu mountains, S of Bara Pokhari, 2300 m, 8.IV.2003, leg. J. Schmidt, 24 exs. (NME, cSch); Manaslu mountains, Bara Pokhari lekh, Chandi Khola valley, 11.–12.IV.2003, 2000–2300 m, leg. J. Schmidt, 1 ex, (NME); Manaslu mountains, S of Bara Pokhari, 2000 m, 7.IV.2003, leg. J. Schmidt, 1 ex. (NME); Bara Pokhari Lekh, 2500 m, 2.IV.1999, leg. G. Hirthe & C. Krüger, 2 exs. (cHir, cSch); Taplejung: confluence of Kabeli and Tada Khola, 1000 m, 23.–25.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [344], 5 exs. (SMNS); ascent to pasture Lassetham from Omje Khola, 2400–3150 m, 6.V.1988, mixed Quercus - Tsuga - Rhodonendron forest, leg. J. Martens & W. Schawaller [357], 1 ex. (SMNS). Remarks. I. quadriguttatum quadriguttatum is widespread in Southeast Asia. In Nepal, it is the most common species and present everywhere (Fig. 45). A second subspecies, I. quadriguttatum japonicum Kocian, 2003 was described from Japan.

Ischnosoma quadriguttatum quadriguttatum (Champion, 1923) Fig. 70. CHINA: Fujian: Kuatun [Guadun] (2300 m) [elevation wrong], 27 ° 40 'N, 117 ° 40 'E, 4.III.1938, 6.III.1938, 27.III.1938, 7.IV.1938, 24.V. 1938, leg. J. Klapperich, 5 exs. (MAKB, cSch); Kuatun [Guadun], 26.III. 1946, leg. Tschung sen., 2 exs. (NHMW); Wuji Shan Nat. Res., Sangan env., 900 m, 30.V.– 12.VI. 2001, leg. P. Hlaváč & J. Cooter, 6 exs. (cSch, cKoc); Gansu: S-Gansu, N Chengxian, W-Qingling Shan, 34 ° 10 ' 20 ''N, 105 ° 42 ' 19 ''E, 1830 m, stream valley with secondary deciduous forest, moist litter sifted, 29.VII. 2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu, N Chengxian, W-Qingling Shan, 34 ° 10 ' 17 ''N, 105 ° 42 ' 56 ''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII. 2012, leg. V. Assing, 6 exs. (cAss, cSch); S-Gansu, mountains SE Longnan, 33 ° 13 ' 20 ''N, 105 ° 15 ' 10 ''E, 2170 m, N-slope with shrubs and scattered coniferous trees, litter sifted, 31.VII. 2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu, S Longnan, Min Shan, 33 °05' 24 ''N, 104 ° 45 ' 13 ''E, 1500 m, N-slope, macchia, litter and moss sifted, 6.VIII. 2012, leg. V. Assing, 2 exs. (cAss); S-Gansu, N Chengxian, W-Qingling Shan, 34 °08' 16 ''N, 105 ° 46 ' 42 ''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII. 2012, leg. V. Assing, 1 ex. (cAss); S-Gansu [CH 12 -04] W Qinling Shan, 43 km N Chengxian, 34 °08' 16 ''N, 105 ° 46 ' 42 ''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII. 2012, leg. M. Schülke, 4 exs. (cSch); S-Gansu [CH 12 -04c] W Qinling Shan, 43 km N Chengxian, 34 °08' 16 ''N, 105 ° 46 ' 42 ''E, 1760 m, S-slope, secondary deciduous forest margin, litter between rocks sifted, 28.VII. 2012, leg. M. Schülke, 3 exs. (cSch); S-Gansu [CH 12 -05] W Qinling Shan, 47 km N Chengxian, 34 ° 10 ' 17 ''N, 105 ° 42 ' 56 ''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII. 2012, leg. M. Schülke, 3 exs. (cSch); S-Gansu [CH 12 -06] W Qinling Shan, 47 km N Chengxian, 34 ° 10 ' 20 ''N, 105 ° 42 ' 19 ''E, 1830 m, creek valley with secondary deciduous forest, moist litter sifted, 29.VII. 2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu [CH 12-17] Min Shan, 38 km SSW Longnan, 33 °05' 24 ''N, 104 ° 45 ' 13 ''E, 1500 m, N-slope, macchia, litter and moss sifted, 6.VIII. 2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu [CH 12 -08] Mts. 38 km SE Longnan, 33 ° 11 ' 20 ''N, 105 ° 14 ' 24 ''E, 2030 m, N-slope with scree, moss, fern roots and litter sifted, 31.VII. 2012, leg. M. Schülke, 1 ex. (cSch); S-Gansu Mts. 38 km SE Longnan, 2130 m, 33 ° 11 ' 16 ''N, 105 ° 14 ' 24 ''E W-slope with scree, scrubs and trees, litter, soil and moss sifted, 7.VIII. 2012, leg. D.W. Wrase [18 B], 1 ex. (cSch); Guizhou: Leishan Co., SE Kaili, NE Leishan, Leigong Shan, E-slope, 2.5 km E of pass, 26 ° 23.39 'N, 108 ° 13.33 'E, 1600 m, 23./ 24.VI. 2001, leg. H. Schillhammer [17 a], 1 ex. (NHMW); Hainan: Bawangling Nat. Forest Park, 12,3 km SE of Baotie, 19 ° 5.20 'N, 109 ° 11.80 'E, 1050 m, 8.V. 2011, sifting moist accumulation of leaf litter along stream in primary forest, leg. M. Fikáček, 1 ex. (NMP); Hubei: SE-Hubei, Mufu Shan, Jiugongshan forest park, 29.4 N, 114.6 E, 1000 m, 3./ 5./ 18.VI. 2002, leg. J. Turna, 1 ex. (NHMW); W-Hubei, Daba Shan, pass E of mt. Da Shennongjia, 12 km MW Muyuping, 31 ° 30 'N, 110 ° 21 'E, creek valley, 1950–2000 m, mixed deciduous forest, moss, dead wood and mushrooms sifted, 19.VII. 2001, leg. M. Schülke [CH 01- 13 c], 2 exs. (cSch); W-Hubei, Daba Shan, creek valley 8 km NW Muyuping, 31 ° 28 'N, 110 ° 22 'E, 1540 m, edge of small creek, 18.VII. 2001, leg. D.W. Wrase, 2 exs. (cSch); same data, but "deciduous forest, sifted from moss, leg. M. Schülke [CH 01- 16 a]", 2 exs. (cSch); Shaanxi: Qinling Shan, Taibai Shan, above Houzhenzi, 1300–1700 m, 9.VI.– 3.VII. 1998, leg. P. Jäger & J. Martens, 1 ex. (SMNS); Qinling Shan, pass at road Zhouzhi–Foping, 105 km SW Xian, N-slope, 1880 m, 33 ° 44 'N, 107 ° 58 'E, 4.VII. 2001, leg. A. Smetana [C 92], 2 exs. (cSme); same data, but " 1700 m, 33 ° 46 'N, 107 ° 58 ''E, 3.VII. 2001 [C 91]", 1 ex. (cSme); same data, but " 1990 m, small creek valley, mixed deciduous forest with bamboo and small meadows, 2./ 4.VII. 2001, leg. D.W. Wrase [01]", 1 ex. (cSch); same data but: "litter, dead wood and mushrooms sifted, 2./ 4.VII. 2001, leg. M. Schülke [CH 01-01]", 2 exs. (cSch); S-Shaanxi [CH 12-31], Micang Shan, 40 km SW Hanzhong, 32 ° 52 ' 25 ''N, 106 ° 37 ' 11 ''E, 1530 m, N-slope, mixed secondary forest, litter and moss sifted, 16.VIII. 2012, leg. M. Schülke, 4 exs. (cSch); S-Shaanxi [CH 12-30], Micang Shan, 33 km S Hanzhong, 32 ° 44 ' 44 ''N, 106 ° 52 ' 46 ''E, 1360 m, stream valley, forest margin, litter and soil sifted, 15.VIII. 2012, leg. M. Schülke, 2 exs. (cSch); S-Shaanxi [CH 12-32], Micang Shan, 42 km S Hanzhong, 32 ° 40 ' 43 ''N, 106 ° 48 ' 33 ''E, 1090 m, stream valley, shady S-slope, secondary mixed forest, litter, grass, and herbs along path sifted, 17.VIII. 2012, leg. M. Schülke, 2 exs. (cSch); S-Shaanxi [CH 12-28] Micang Shan, 34 km S Hanzhong, 32 ° 44 ' 22 ''N, 106 ° 51 ' 55 ''E, 1460 m, W-slope, deciduous forest margin with bamboo, litter, grass, and moss sifted, 14.VIII. 2012, leg. M. Schülke, 1 ex. (cSch); S-Shaanxi, Micang Shan 42 km S Hanzhong, 1090 m, 32 ° 40 ' 52 ''N / 106 ° 49 ' 16 ''E, NW-slope, margin of mixed forest with rocks, litter, moss and soil sifted) 14.VIII. 2012, leg. D.W. Wrase [27], 1 ex. (cSch); S- Shaanxi, Micang Shan 40 km SW Hanzhong, 1530 m, 32 ° 52 ' 25 ''N / 106 ° 37 ' 11 ''E, N-slope, mixed secondary forest, litter and moss sifted, 16.VIII. 2012, leg. D.W. Wrase [31], 1 ex. (cSch); S-Shaanxi, Qinling Shan, 42 km SW Meixian, 1875 m, 34 °01' 32 ''N / 107 ° 24 ' 13 ''E, N-slope, secondary deciduous forest near creek, litter & grass sifted, 26.VII. 2012, leg. D.W. Wrase [02A], 1 ex. (cSch); Micang Shan, 33 km S Hanzhong, 1360 m, 32 ° 44 ' 44 ''N / 106 ° 52 ' 46 ''E, stream valley, shady brookside with bamboo/decidious shrubs, litter/moss/soil sifted, 15.VIII. 2012, leg. D.W.Wrase [30 B], 1 ex. (cSch); S-Shaanxi Micang Shan, 42 km S Hanzhong, 32 ° 40 ' 43 ''N, 106 ° 48 ' 33 ''E, 1090 m, stream valley, shady S-slope, secondary mixed forest, litter, grass, and herbs along path sifted, 17.VIII. 2012, leg. V. Assing, 4 exs. (cAss, cSch); S-Shaanxi, Micang Shan, 34 km S Hanzhong, 32 ° 44 ' 22 ''N, 106 ° 51 ' 55 ''E, 1460 m, W-slope, deciduous forest margin with bamboo, litter, grass, and moss sifted, 14.VIII. 2012, leg. V. Assing, 1 ex. (cAss); S-Shaanxi, Micang Shan, 42 km S Hanzhong, 32 ° 40 ' 52 ''N, 106 ° 49 ' 16 ''E, 1090 m, NW-slope, mixed forest margin with rocks, litter, grass, and moss sifted, 14.VIII. 2012, leg. V. Assing, 5 exs. (cAss, cSch); S-Shaanxi Micang Shan, 30 km S Hanzhong, 32 ° 45 ' 56 ''N, 106 ° 53 ' 57 ''E, 1070 m, stream valley, litter and soil sifted, 15.VIII. 2012,, leg. V. Assing, 2 exs. (cAss); Shaanxi/Chongqing: Shaanxi/Chongqing border, pass 20 km SSE Zhenping, 1700–1800 m, 31 ° 44 'N, 109 ° 35 'E, small creek valley, young mixed forest, litter and moss sifted, 9./ 12.VII. 2001, leg. D.W. Wrase [07], 3 exs. (cSch); same data, but "leg. M. Schülke [CH 01-07], 1 ex.; same data, but "leg. A. Smetana [C 101], 1 ex. (cSme); Sichuan: Emei Shan, 29 ° 33.6 'N, 103 ° 20.6 'E, 1800–2400 m, sifting 11 –17, 27.VI.– 5.VII. 2009, leg. V. Grebennikov, 32 exs. (CNC, cSch); Emei Shan, 29 ° 32 ' 56 ''N, 103 ° 20 ' 28 ''E, 2310 m, sifting 38, 20.VI. 2010, leg. V. Grebennikov, 2 exs. (CNC); Emei Shan, Leidongping, 2500 m, 29 ° 32 'N, 103 ° 21 'E, 18.VII. 1996, leg. A. Smetana, J. Farkač & P. Kabátek, 1 ex. (cSme); Emei Shan, Wannian, 1050 m, 19.– 30.III. 1999, leg. W. Schawaller, 1 ex. (SMNS); C-Sichuan, Wenjiang Distr., Dujiangyan County, Qingcheng Shan, 56 km NW Chengdu, 30 ° 54 'N, 103 ° 33 'E, 975 m, creek bank, forest remnant, 18.VI. 1999, leg. M. Schülke, 1 ex. (cSch); same data, but "field ridge, leg. D.W. Wrase", 1 ex. (cSch); same data, but "leg. A. Pütz", 2 exs. (cPüt); Qingcheng Shan 65 km NW Chengdu, 37 ° 53 'N, 103 ° 53 'E, 18.V./ 3.– 4.VI. 1997, leg. A. Pütz, 4 exs. (cPüt); W- Sichuan, Ganzi Tibet. Auton. Prefecture, Luding, W Erlang Shan pass, 2600 m, 7 km SSE Luding, 29 ° 51 'N, 102 ° 15 'E, litter and mushrooms sifted, 29.VI. 1999, leg. M. Schülke, 1 ex. (cSch); same data, but 20.– 29.VI. 1999, leg. D.W. Wrase", 1 ex. (cSch); W-Sichuan, Ya'an Prefecture, Shimian County, Daxue Shan, road between Anshunchang and Wanba, 20 km WSW Shimian, 1500 m, 9.VII.1999, 3 exs. (cPüt); same data, but " 12 km W Shimian, 1300 m ", 1 ex. (cPüt); S-Sichuan, S Xichang, Luoji Shan, 2300–2500 m, litter sifted, 16.– 24.VII. 1996, leg. S. Kurbatov, 1 ex. (MHNG); Xiling Shan, 1600–2400 m, litter sifted, 30.VII.– 4.VIII. 1996, leg. S. Kurbatov, 1 ex. (MHNG); Yunnan: SW-Yunnan, Rehai hot springs, 6 km SW Tengchong, 24 ° 57.1 'N, 98 ° 26.2 'E, 1400 m, 5.VI. 2007, sifted detrius and leaves in grove margin above hot stream, along old path, leg. J. Hájek & J. Růžička [CH 11], 1 ex. (cSch); SW-Yunnan, Gaoligong Shan, 90 km W Baoshan, 26.– 28.V. 1995, leg. S. Becvar, 1 ex. (NHMW); W-Yunnan, above Dali, 2000–2200 m, 4.– 17.IV. 1999, leg. W. Schawaller, 1 ex. (SMNS); NW-Yunnan, E slope N Gaoligong Shan, 27 ° 46.8 'N; 98 ° 33.1 'E, 2000–3000 m, sifting 1 –7, 12.– 15.VI. 2009, leg. V. Grebennikov, 3 exs. (CNC); NW-Yunnan, E slope N Gaoligong Shan, 27 ° 45 ' 27.1 ''N; 98 ° 35 ' 34.5 ''E, 2600 m, sifting 26, 2.VI. 2010, leg. V. Grebennikov, 3 exs. (CNC); NW-Yunnan, E slope N Gaoligong Shan, 27 ° 45 ' 40.8 ''N; 98 ° 36 '03.2''E, 2536 m, sifting 27, 3.VI. 2010, leg. V. Grebennikov, 1 ex. (CNC); W-Yunnan, Nujiang Lisu Aut.Pref., Nu Shan, 7 km NNW Coajian, 25 ° 43 ' 29 ''N, 99 °07' 57 ''E, 2420 m, secondary pine forest with shrubs, litter and bark sifted, 11.VI. 2007, leg. M. Schülke [CH 07- 30], 5 exs. (cSch); same data, but "leg. D.W. Wrase [30]", 2 exs. (cSch); same data, but "leg. A. Pütz", 2 exs. (cPüt, cKoc); W-Yunnan, Baoshan Pref., mountain range 25 km S Tengchong, 1900 m, 24 ° 48 ' 28 ''N, 98 ° 32 '03''E, devastated primary deciduous forest, litter and fungi sifted, 2.VI. 2007, leg. M. Schülke [CH 07- 17], 2 exs. (cSch); same data, but "leg. D.W. Wrase [17]", 1 ex. (cSch); W- Yunnan, Nujiang Lisu Aut. Pref., Salween side valley 5 km S Fugong, road SS 228 km 223, under stones, moist embarkment, 8.VI. 2007, leg. A. Pütz [CH 07- 25], 1 ex. (cPüt); W-Yunnan, Baoshan Pref., mountain range 22 km S Tengchong, 1750 m, 24 ° 49 ' 29 ''N, 98 ° 29 ' 27 ''E, secondary forest, litter and dead wood sifted, 2.VI. 2007, leg. A. Pütz [CH 07- 18], 1 ex. (cPüt); W-Yunnan, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 2100–2200 m, 24 ° 51 ' 22 ''N, 98 ° 45 ' 36 ''E, deciduous forest, litter, wood and fungi sifted, 4.VI. 2007, leg. A. Pütz [CH 07- 14 A], 1 ex. (cPüt); same data, but "pitfall trap, 31.V.– 4.VI. 2007, leg. D.W. Wrase, 1 ex. (cSch); W-Yunnan, Baoshan Pref., mountain range 14 km E Tengchong, 1850 m, 25 °00' 28 ''N, 98 ° 38 '07''E, secondary mixed forest, litter sifted, 1.VI. 2007, leg. A. Pütz [CH 07- 16], 1 ex. (cPüt); W-Yunnan, Baoshan Pref., Gaoligong Shan, 29 km ESE Tengchong, 24 ° 55 ' 37 ''N, 98 ° 45 '09''E, 2350 m, devastated deciduous forest, litter, wood and fungi sifted, 1.VI. 2007, leg. A. Pütz [CH 07- 15], 4 exs. (cPüt); same data, but "leg. D.W.Wrase", 1 ex. (cSch); Yunnan, Baoshan Pref., Gaoligong Shan, W Pass 35 km SE Tengchong, 2100 m, 24 ° 50 ' 18 ''N, 98 ° 45 ' 43 ''E, devastated primary deciduous forest, litter, wood and mushrooms sifted, 25.VIII. 2009, leg. M. Schülke [CH 09-06], 2 exs. (cSch, cKoc); S- Yunnan, Pu'er Pref., Ailao Shan, 37 km NW Jingdong, 24 ° 45 ' 12 ''N, 100 ° 41 ' 24.5 ''N, 2300 m, devastated forest remnant, litter and dead wood sifted, 13.IX. 2009, leg. M. Schülke [CH 09- 48], 3 exs. (cSch); W-Yunnan, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 2150 m, 24 ° 51 ' 22 ''N, 98 ° 45 ' 36 ''E, devastated primery deciduous forest, litter, wood and mushrooms sifted, 6.VIII. 2009, leg. M. Schülke [CH 09-08], 1 ex. (cSch); S-Yunnan, Lincang Pref., Xue Shan, 11 km ENE Lincang, 2510 m, 23 ° 55 '01''N, 100 ° 11 ' 17.5 ''E, secondary pine forest with Rhododendron, small cleft with water, litter & mushrooms sifted, 10.IX. 2009, leg. M. Schülke [CH 09- 39], 2 exs. (cSch); S-Yunnan, Lincang/Dali Pref., Wuliang Shan, old pass road, N pass, 24 ° 45 ' 16.4 ''N, 100 ° 29 ' 50.3 ''E, 2350 m, forest remnant & tea plantation litter, mushrooms and grass sifted, 16.IX. 2009, leg. M. Schülke [CH 09- 55], 1 ex. (cSch); S-Yunnan, Lincang Pref., Wuliang Shan, old pass road, W side, 24 ° 42 ' 58.6 ''N, 100 ° 29 '52.0''E, 2200 m, small creek valley with primary forest remnant, litter sifted, 16.IX. 2009, leg. M. Schülke [CH 09- 47 a], 1 ex. (cSch); S-Yunnan, Lincang Pref., Bangma Shan, E pass, 17 km NW Lincang, 2040 m, 23 ° 57 ' 31 ''N / 99 ° 56 ' 13 ''E, secondary pine forest with ferns, litter sifted, 9.IX. 2009, leg. D.W.Wrase [36], 1 ex. (cSch); C-Yunnan [23], mountain W Yuxi, 24 ° 27 ' 11 ''N, 102 ° 29 ' 58 ''E, 2250 m, secondary mixed forest, litter, roots, and moss sifted, 31.VIII. 2014, leg. V. Assing [23], 1 ex. (cAss); C-Yunnan [6], NE Kunming, 25 °08' 35 ''N, 102 ° 53 ' 49 ''E, 2320 m, mixed forest with alder, oak, and pine, litter and mushrooms sifted, 13.VIII. 2014, leg. V. Assing [06], 1 ex. (cAss); C-Yunnan [5], NE Kunming, 25 °08' 40 ''N, 102 ° 53 ' 48 ''E, 2290 m, mixed deciduous forest with scattered pine trees, litter and mushrooms sifted, 11.VIII. 2014, leg. V. Assing [05], 1 ex. (cAss); S-Yunnan [22 / 22 a], SE Pingbian, 22 ° 54 ' 31 ''N, 103 ° 41 ' 44 ''E, 2100 m, primary subtropical broad-leaved forest, litter sifted, 27./ 28.VIII. 2014, leg. V. Assing, 14 exs. (cAss, cSch); Yunnan [2 a], mountains NE Dongchuan, 26 ° 14 ' 10 ''N, 103 ° 12 ' 31 ''E, 2670 m, scree slope with bushes and herbs, litter sifted, 9.VIII. 2014, leg. V. Assing, 1 ex. (cAss); N-Yunnan, Nujiang Lisu Autonomous Prefecture, Gongshan County, N Gaoligong Shan, 2500m, 27 ° 45.404 'N, 98 ° 35.749 'E, litter & debries at snowfield sifted during rain, 19.VI. 2005, leg. M. Schülke [C 2005 - 12], 2 exs. (cSch); S-Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. Na Ban (NNNR), 22 °09.49'N, 100 ° 39.92 'E, forest EKL, 26.III. 2009, leg. L. Mengl, 1 ex. (NME); S-Yunnan, Xishuangbanna, 20 km NW Jinghong, near Man Dian (NNNR), pitfall trap, 22 °07.80'N, 100 ° 40.05 'E, 740 m, rubber plantation, 28.VI. 2008, leg. A. Weigel, 1 ex. (NME); Zheijiang: Tianmu Shan, pass 25 km NNW Linan, 30 ° 25 ' 40 ''N, 119 ° 35 ' 30 ''E, 620–820 m, creek valley with bamboo and mixed forest, litter sifted, 16.VI. 2007, leg. A. Pütz [CH 07- 37], 5 exs. (cPüt); same data, but "leg. D.W. Wrase", 3 exs. (cSch); West Tianmu Shan N.R. way to peak of immortals, 30 ° 20 ' 34 ''N, 119 ° 25 ' 51 ''E, 1100–1200 m, primary mixed forest, litter and moss sifted, 15.VI. 2007, leg. A. Pütz [CH 07- 36], 2 exs. (cPüt); Tianmu Shan, pass 25 km NNW Linan, 620–820 m, 30 ° 25 ' 40 ''N, 119 ° 35 ' 30 ''E, creek valley with bamboo and mixed forest, litter, sifted, 16.VI. 2007, leg. M. Schülke [CH 07- 37], 5 exs. (cSch); Zheijiang/Anhui: Zheijiang/Anhui border, pass W Linan, 30 ° 17 ' 59 ''N, 118 ° 52 ' 41 ''E, 625 m, walnut plantation, plant refuse and litter sifted, 19.VI. 2007, leg. M. Schülke, 1 ex. (cSch); TAIWAN: Ilan Hsien: 75 km, N crossing highway, 1100 m, 22.V. 2003, flight intercept trap, leg. Chun Lin Li, 2 exs. (NMB); Shen-Mi Lake, 24 ° 22 ' 43 ''N, 121 ° 44 ' 12 ''E, 1110 m, 10.V. 1995, leg. A. Smetana [T 177], 1 ex. (cSme); Shen-Mi Lake, 24 ° 22 ' 43 ''N, 121 ° 44 ' 12 ''E, 1110 m, 9.V. 1995, leg. A. Smetana [T 175], 4 exs. (cSme, cSch); Kaohsiung Hsien, Tengchih, 1400 m, 21.– 23. VII. 2000, leg. H. Sugaya, 2 exs. (KUM, cSchü); Tengchih, 1400 m, 25.– 30. IV. 2001, leg. H. Sugaya, 3 exs. (KUM, cSchü); Tengchih, 2100 m, 19.– 23. IV. 2001, leg. H. Sugaya, 6 exs. (KUM, cSchü); Tengchih, 1700 m, 2.– 3. IV. 2002, litter, leg. H. Sugaya, 2 exs. (KUM); Tona Forest Station, 1050 m, 27.– 30.IV. 1998 (flight intercept trap), leg. A. Smetana & L. Robillard [T 193], 1 ex. (cSch); Road above Tona Forest Station, km 16–17, 1700–1800 m, 28.IV. 1998, leg. A. Smetana [T 190]. 2 exs. (cSme); Nantou Hsien: Meifeng, 2100 m, 6. V. 2001, leg. H. Sugaya, 20 exs. (KUM, cSchü); Meimu, 1600 m, litter, 30. III. 2002, leg. H. Sugaya, 5 exs. (KUM); Tehuashe, 900 m, 4.– 5. V. 2001, leg. H. Sugaya, 1 ex. (KUM); Nanshanchi, 800 m, 27. X. 2000, leg. H. Sugaya, 2 exs. (KUM, cSchü); Meifeng, 2130 m, 4.V. 1998, leg. A. Smetana [T 199], 1 ex. (cSch); Pintung Hsien: Chialoshui, 100 m, 11. III. 2000, leg. M. Maruyama, 1 ex. (KUM); Taichung Hsien: Anmashan, 2150 m, 13.V. 1992, leg. A. Smetana [T 129], 1 ex. (cSme); Anmashan, 2225 m, 2.V. 1990, leg. A. Smetana [T 37], 1 ex (cSme); Anmashan, 2000 m, 24.– 26.VI. 2003, flight intercept trap, leg. Chun Lin Li, 4 exs. (NMB); Chipen- Wenchuan, 400 m, 6.– 8. XI. 2000, leg. H. Sugaya, 3 exs. (KUM, cSchü); Chipen-Wenchuan, 400 m, 24.– 27.IV. 2001, leg. H. Sugaya, 5 exs. (KUM, cSchü); Taitung Hsien: road no. 20, after Li-Tao (Lidau), km 174, 1300 m, mountain forest litter, 8.IV. 2007, leg. S. Vít, 2 exs. (cSch); W-Taitung, road no. 24, Jhihben, forest recreation area, 400 m, litter sifted, 12.IV. 2007, leg. S. Vít, 1 ex. (cSch); Taipeh: Noi Dong logging road, 850 m, 19.II. 2004, flight intercept trap, leg. Chun Lin Li, 3 exs. (MNB); Remarks. Ischnosoma quadriguttatum is one of the most widespread species of the genus, its distribution ranging from Pakistan to Indonesia. In China the species is very common in the central and southern parts of the country (Fig. 70). Records from Fujian, Gansu, Hainan, Hubei and Shaanxi are new province records.

Ischnosoma quadriguttatum quadriguttatum (Champion, 1923) Fig. 70. CHINA: Fujian: Kuatun [Guadun] (2300 m) [elevation wrong], 27 �� 40 'N, 117 �� 40 'E, 4.III.1938, 6.III.1938, 27.III.1938, 7.IV.1938, 24.V. 1938, leg. J. Klapperich, 5 exs. (MAKB, cSch); Kuatun [Guadun], 26.III. 1946, leg. Tschung sen., 2 exs. (NHMW); Wuji Shan Nat. Res., Sangan env., 900 m, 30.V.��� 12.VI. 2001, leg. P. Hlav��č & J. Cooter, 6 exs. (cSch, cKoc); Gansu: S-Gansu, N Chengxian, W-Qingling Shan, 34 �� 10 ' 20 ''N, 105 �� 42 ' 19 ''E, 1830 m, stream valley with secondary deciduous forest, moist litter sifted, 29.VII. 2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu, N Chengxian, W-Qingling Shan, 34 �� 10 ' 17 ''N, 105 �� 42 ' 56 ''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII. 2012, leg. V. Assing, 6 exs. (cAss, cSch); S-Gansu, mountains SE Longnan, 33 �� 13 ' 20 ''N, 105 �� 15 ' 10 ''E, 2170 m, N-slope with shrubs and scattered coniferous trees, litter sifted, 31.VII. 2012, leg. V. Assing, 3 exs. (cAss, cSch); S-Gansu, S Longnan, Min Shan, 33 ��05' 24 ''N, 104 �� 45 ' 13 ''E, 1500 m, N-slope, macchia, litter and moss sifted, 6.VIII. 2012, leg. V. Assing, 2 exs. (cAss); S-Gansu, N Chengxian, W-Qingling Shan, 34 ��08' 16 ''N, 105 �� 46 ' 42 ''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII. 2012, leg. V. Assing, 1 ex. (cAss); S-Gansu [CH 12 -04] W Qinling Shan, 43 km N Chengxian, 34 ��08' 16 ''N, 105 �� 46 ' 42 ''E, 1760 m, N-slope, secondary deciduous forest margin, sifted, 28.VII. 2012, leg. M. Sch��lke, 4 exs. (cSch); S-Gansu [CH 12 -04c] W Qinling Shan, 43 km N Chengxian, 34 ��08' 16 ''N, 105 �� 46 ' 42 ''E, 1760 m, S-slope, secondary deciduous forest margin, litter between rocks sifted, 28.VII. 2012, leg. M. Sch��lke, 3 exs. (cSch); S-Gansu [CH 12 -05] W Qinling Shan, 47 km N Chengxian, 34 �� 10 ' 17 ''N, 105 �� 42 ' 56 ''E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII. 2012, leg. M. Sch��lke, 3 exs. (cSch); S-Gansu [CH 12 -06] W Qinling Shan, 47 km N Chengxian, 34 �� 10 ' 20 ''N, 105 �� 42 ' 19 ''E, 1830 m, creek valley with secondary deciduous forest, moist litter sifted, 29.VII. 2012, leg. M. Sch��lke, 1 ex. (cSch); S-Gansu [CH 12-17] Min Shan, 38 km SSW Longnan, 33 ��05' 24 ''N, 104 �� 45 ' 13 ''E, 1500 m, N-slope, macchia, litter and moss sifted, 6.VIII. 2012, leg. M. Sch��lke, 1 ex. (cSch); S-Gansu [CH 12 -08] Mts. 38 km SE Longnan, 33 �� 11 ' 20 ''N, 105 �� 14 ' 24 ''E, 2030 m, N-slope with scree, moss, fern roots and litter sifted, 31.VII. 2012, leg. M. Sch��lke, 1 ex. (cSch); S-Gansu Mts. 38 km SE Longnan, 2130 m, 33 �� 11 ' 16 ''N, 105 �� 14 ' 24 ''E W-slope with scree, scrubs and trees, litter, soil and moss sifted, 7.VIII. 2012, leg. D.W. Wrase [18 B], 1 ex. (cSch); Guizhou: Leishan Co., SE Kaili, NE Leishan, Leigong Shan, E-slope, 2.5 km E of pass, 26 �� 23.39 'N, 108 �� 13.33 'E, 1600 m, 23./ 24.VI. 2001, leg. H. Schillhammer [17 a], 1 ex. (NHMW); Hainan: Bawangling Nat. Forest Park, 12,3 km SE of Baotie, 19 �� 5.20 'N, 109 �� 11.80 'E, 1050 m, 8.V. 2011, sifting moist accumulation of leaf litter along stream in primary forest, leg. M. Fik��ček, 1 ex. (NMP); Hubei: SE-Hubei, Mufu Shan, Jiugongshan forest park, 29.4 N, 114.6 E, 1000 m, 3./ 5./ 18.VI. 2002, leg. J. Turna, 1 ex. (NHMW); W-Hubei, Daba Shan, pass E of mt. Da Shennongjia, 12 km MW Muyuping, 31 �� 30 'N, 110 �� 21 'E, creek valley, 1950���2000 m, mixed deciduous forest, moss, dead wood and mushrooms sifted, 19.VII. 2001, leg. M. Sch��lke [CH 01- 13 c], 2 exs. (cSch); W-Hubei, Daba Shan, creek valley 8 km NW Muyuping, 31 �� 28 'N, 110 �� 22 'E, 1540 m, edge of small creek, 18.VII. 2001, leg. D.W. Wrase, 2 exs. (cSch); same data, but "deciduous forest, sifted from moss, leg. M. Sch��lke [CH 01- 16 a]", 2 exs. (cSch); Shaanxi: Qinling Shan, Taibai Shan, above Houzhenzi, 1300���1700 m, 9.VI.��� 3.VII. 1998, leg. P. J��ger & J. Martens, 1 ex. (SMNS); Qinling Shan, pass at road Zhouzhi���Foping, 105 km SW Xian, N-slope, 1880 m, 33 �� 44 'N, 107 �� 58 'E, 4.VII. 2001, leg. A. Smetana [C 92], 2 exs. (cSme); same data, but " 1700 m, 33 �� 46 'N, 107 �� 58 ''E, 3.VII. 2001 [C 91]", 1 ex. (cSme); same data, but " 1990 m, small creek valley, mixed deciduous forest with bamboo and small meadows, 2./ 4.VII. 2001, leg. D.W. Wrase [01]", 1 ex. (cSch); same data but: "litter, dead wood and mushrooms sifted, 2./ 4.VII. 2001, leg. M. Sch��lke [CH 01-01]", 2 exs. (cSch); S-Shaanxi [CH 12-31], Micang Shan, 40 km SW Hanzhong, 32 �� 52 ' 25 ''N, 106 �� 37 ' 11 ''E, 1530 m, N-slope, mixed secondary forest, litter and moss sifted, 16.VIII. 2012, leg. M. Sch��lke, 4 exs. (cSch); S-Shaanxi [CH 12-30], Micang Shan, 33 km S Hanzhong, 32 �� 44 ' 44 ''N, 106 �� 52 ' 46 ''E, 1360 m, stream valley, forest margin, litter and soil sifted, 15.VIII. 2012, leg. M. Sch��lke, 2 exs. (cSch); S-Shaanxi [CH 12-32], Micang Shan, 42 km S Hanzhong, 32 �� 40 ' 43 ''N, 106 �� 48 ' 33 ''E, 1090 m, stream valley, shady S-slope, secondary mixed forest, litter, grass, and herbs along path sifted, 17.VIII. 2012, leg. M. Sch��lke, 2 exs. (cSch); S-Shaanxi [CH 12-28] Micang Shan, 34 km S Hanzhong, 32 �� 44 ' 22 ''N, 106 �� 51 ' 55 ''E, 1460 m, W-slope, deciduous forest margin with bamboo, litter, grass, and moss sifted, 14.VIII. 2012, leg. M. Sch��lke, 1 ex. (cSch); S-Shaanxi, Micang Shan 42 km S Hanzhong, 1090 m, 32 �� 40 ' 52 ''N / 106 �� 49 ' 16 ''E, NW-slope, margin of mixed forest with rocks, litter, moss and soil sifted) 14.VIII. 2012, leg. D.W. Wrase [27], 1 ex. (cSch); S- Shaanxi, Micang Shan 40 km SW Hanzhong, 1530 m, 32 �� 52 ' 25 ''N / 106 �� 37 ' 11 ''E, N-slope, mixed secondary forest, litter and moss sifted, 16.VIII. 2012, leg. D.W. Wrase [31], 1 ex. (cSch); S-Shaanxi, Qinling Shan, 42 km SW Meixian, 1875 m, 34 ��01' 32 ''N / 107 �� 24 ' 13 ''E, N-slope, secondary deciduous forest near creek, litter & grass sifted, 26.VII. 2012, leg. D.W. Wrase [02A], 1 ex. (cSch); Micang Shan, 33 km S Hanzhong, 1360 m, 32 �� 44 ' 44 ''N / 106 �� 52 ' 46 ''E, stream valley, shady brookside with bamboo/decidious shrubs, litter/moss/soil sifted, 15.VIII. 2012, leg. D.W.Wrase [30 B], 1 ex. (cSch); S-Shaanxi Micang Shan, 42 km S Hanzhong, 32 �� 40 ' 43 ''N, 106 �� 48 ' 33 ''E, 1090 m, stream valley, shady S-slope, secondary mixed forest, litter, grass, and herbs along path sifted, 17.VIII. 2012, leg. V. Assing, 4 exs. (cAss, cSch); S-Shaanxi, Micang Shan, 34 km S Hanzhong, 32 �� 44 ' 22 ''N, 106 �� 51 ' 55 ''E, 1460 m, W-slope, deciduous forest margin with bamboo, litter, grass, and moss sifted, 14.VIII. 2012, leg. V. Assing, 1 ex. (cAss); S-Shaanxi, Micang Shan, 42 km S Hanzhong, 32 �� 40 ' 52 ''N, 106 �� 49 ' 16 ''E, 1090 m, NW-slope, mixed forest margin with rocks, litter, grass, and moss sifted, 14.VIII. 2012, leg. V. Assing, 5 exs. (cAss, cSch); S-Shaanxi Micang Shan, 30 km S Hanzhong, 32 �� 45 ' 56 ''N, 106 �� 53 ' 57 ''E, 1070 m, stream valley, litter and soil sifted, 15.VIII. 2012,, leg. V. Assing, 2 exs. (cAss); Shaanxi/Chongqing: Shaanxi/Chongqing border, pass 20 km SSE Zhenping, 1700���1800 m, 31 �� 44 'N, 109 �� 35 'E, small creek valley, young mixed forest, litter and moss sifted, 9./ 12.VII. 2001, leg. D.W. Wrase [07], 3 exs. (cSch); same data, but "leg. M. Sch��lke [CH 01-07], 1 ex.; same data, but "leg. A. Smetana [C 101], 1 ex. (cSme); Sichuan: Emei Shan, 29 �� 33.6 'N, 103 �� 20.6 'E, 1800���2400 m, sifting 11 ���17, 27.VI.��� 5.VII. 2009, leg. V. Grebennikov, 32 exs. (CNC, cSch); Emei Shan, 29 �� 32 ' 56 ''N, 103 �� 20 ' 28 ''E, 2310 m, sifting 38, 20.VI. 2010, leg. V. Grebennikov, 2 exs. (CNC); Emei Shan, Leidongping, 2500 m, 29 �� 32 'N, 103 �� 21 'E, 18.VII. 1996, leg. A. Smetana, J. Farkač & P. Kab��tek, 1 ex. (cSme); Emei Shan, Wannian, 1050 m, 19.��� 30.III. 1999, leg. W. Schawaller, 1 ex. (SMNS); C-Sichuan, Wenjiang Distr., Dujiangyan County, Qingcheng Shan, 56 km NW Chengdu, 30 �� 54 'N, 103 �� 33 'E, 975 m, creek bank, forest remnant, 18.VI. 1999, leg. M. Sch��lke, 1 ex. (cSch); same data, but "field ridge, leg. D.W. Wrase", 1 ex. (cSch); same data, but "leg. A. P��tz", 2 exs. (cP��t); Qingcheng Shan 65 km NW Chengdu, 37 �� 53 'N, 103 �� 53 'E, 18.V./ 3.��� 4.VI. 1997, leg. A. P��tz, 4 exs. (cP��t); W- Sichuan, Ganzi Tibet. Auton. Prefecture, Luding, W Erlang Shan pass, 2600 m, 7 km SSE Luding, 29 �� 51 'N, 102 �� 15 'E, litter and mushrooms sifted, 29.VI. 1999, leg. M. Sch��lke, 1 ex. (cSch); same data, but 20.��� 29.VI. 1999, leg. D.W. Wrase", 1 ex. (cSch); W-Sichuan, Ya'an Prefecture, Shimian County, Daxue Shan, road between Anshunchang and Wanba, 20 km WSW Shimian, 1500 m, 9.VII.1999, 3 exs. (cP��t); same data, but " 12 km W Shimian, 1300 m ", 1 ex. (cP��t); S-Sichuan, S Xichang, Luoji Shan, 2300���2500 m, litter sifted, 16.��� 24.VII. 1996, leg. S. Kurbatov, 1 ex. (MHNG); Xiling Shan, 1600���2400 m, litter sifted, 30.VII.��� 4.VIII. 1996, leg. S. Kurbatov, 1 ex. (MHNG); Yunnan: SW-Yunnan, Rehai hot springs, 6 km SW Tengchong, 24 �� 57.1 'N, 98 �� 26.2 'E, 1400 m, 5.VI. 2007, sifted detrius and leaves in grove margin above hot stream, along old path, leg. J. H��jek & J. Růžička [CH 11], 1 ex. (cSch); SW-Yunnan, Gaoligong Shan, 90 km W Baoshan, 26.��� 28.V. 1995, leg. S. Becvar, 1 ex. (NHMW); W-Yunnan, above Dali, 2000���2200 m, 4.��� 17.IV. 1999, leg. W. Schawaller, 1 ex. (SMNS); NW-Yunnan, E slope N Gaoligong Shan, 27 �� 46.8 'N; 98 �� 33.1 'E, 2000���3000 m, sifting 1 ���7, 12.��� 15.VI. 2009, leg. V. Grebennikov, 3 exs. (CNC); NW-Yunnan, E slope N Gaoligong Shan, 27 �� 45 ' 27.1 ''N; 98 �� 35 ' 34.5 ''E, 2600 m, sifting 26, 2.VI. 2010, leg. V. Grebennikov, 3 exs. (CNC); NW-Yunnan, E slope N Gaoligong Shan, 27 �� 45 ' 40.8 ''N; 98 �� 36 '03.2''E, 2536 m, sifting 27, 3.VI. 2010, leg. V. Grebennikov, 1 ex. (CNC); W-Yunnan, Nujiang Lisu Aut.Pref., Nu Shan, 7 km NNW Coajian, 25 �� 43 ' 29 ''N, 99 ��07' 57 ''E, 2420 m, secondary pine forest with shrubs, litter and bark sifted, 11.VI. 2007, leg. M. Sch��lke [CH 07- 30], 5 exs. (cSch); same data, but "leg. D.W. Wrase [30]", 2 exs. (cSch); same data, but "leg. A. P��tz", 2 exs. (cP��t, cKoc); W-Yunnan, Baoshan Pref., mountain range 25 km S Tengchong, 1900 m, 24 �� 48 ' 28 ''N, 98 �� 32 '03''E, devastated primary deciduous forest, litter and fungi sifted, 2.VI. 2007, leg. M. Sch��lke [CH 07- 17], 2 exs. (cSch); same data, but "leg. D.W. Wrase [17]", 1 ex. (cSch); W- Yunnan, Nujiang Lisu Aut. Pref., Salween side valley 5 km S Fugong, road SS 228 km 223, under stones, moist embarkment, 8.VI. 2007, leg. A. P��tz [CH 07- 25], 1 ex. (cP��t); W-Yunnan, Baoshan Pref., mountain range 22 km S Tengchong, 1750 m, 24 �� 49 ' 29 ''N, 98 �� 29 ' 27 ''E, secondary forest, litter and dead wood sifted, 2.VI. 2007, leg. A. P��tz [CH 07- 18], 1 ex. (cP��t); W-Yunnan, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 2100���2200 m, 24 �� 51 ' 22 ''N, 98 �� 45 ' 36 ''E, deciduous forest, litter, wood and fungi sifted, 4.VI. 2007, leg. A. P��tz [CH 07- 14 A], 1 ex. (cP��t); same data, but "pitfall trap, 31.V.��� 4.VI. 2007, leg. D.W. Wrase, 1 ex. (cSch); W-Yunnan, Baoshan Pref., mountain range 14 km E Tengchong, 1850 m, 25 ��00' 28 ''N, 98 �� 38 '07''E, secondary mixed forest, litter sifted, 1.VI. 2007, leg. A. P��tz [CH 07- 16], 1 ex. (cP��t); W-Yunnan, Baoshan Pref., Gaoligong Shan, 29 km ESE Tengchong, 24 �� 55 ' 37 ''N, 98 �� 45 '09''E, 2350 m, devastated deciduous forest, litter, wood and fungi sifted, 1.VI. 2007, leg. A. P��tz [CH 07- 15], 4 exs. (cP��t); same data, but "leg. D.W.Wrase", 1 ex. (cSch); Yunnan, Baoshan Pref., Gaoligong Shan, W Pass 35 km SE Tengchong, 2100 m, 24 �� 50 ' 18 ''N, 98 �� 45 ' 43 ''E, devastated primary deciduous forest, litter, wood and mushrooms sifted, 25.VIII. 2009, leg. M. Sch��lke [CH 09-06], 2 exs. (cSch, cKoc); S- Yunnan, Pu'er Pref., Ailao Shan, 37 km NW Jingdong, 24 �� 45 ' 12 ''N, 100 �� 41 ' 24.5 ''N, 2300 m, devastated forest remnant, litter and dead wood sifted, 13.IX. 2009, leg. M. Sch��lke [CH 09- 48], 3 exs. (cSch); W-Yunnan, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 2150 m, 24 �� 51 ' 22 ''N, 98 �� 45 ' 36 ''E, devastated primery deciduous forest, litter, wood and mushrooms sifted, 6.VIII. 2009, leg. M. Sch��lke [CH 09-08], 1 ex. (cSch); S-Yunnan, Lincang Pref., Xue Shan, 11 km ENE Lincang, 2510 m, 23 �� 55 '01''N, 100 �� 11 ' 17.5 ''E, secondary pine forest with Rhododendron, small cleft with water, litter & mushrooms sifted, 10.IX. 2009, leg. M. Sch��lke [CH 09- 39], 2 exs. (cSch); S-Yunnan, Lincang/Dali Pref., Wuliang Shan, old pass road, N pass, 24 �� 45 ' 16.4 ''N, 100 �� 29 ' 50.3 ''E, 2350 m, forest remnant & tea plantation litter, mushrooms and grass sifted, 16.IX. 2009, leg. M. Sch��lke [CH 09- 55], 1 ex. (cSch); S-Yunnan, Lincang Pref., Wuliang Shan, old pass road, W side, 24 �� 42 ' 58.6 ''N, 100 �� 29 '52.0''E, 2200 m, small creek valley with primary forest remnant, litter sifted, 16.IX. 2009, leg. M. Sch��lke [CH 09- 47 a], 1 ex. (cSch); S-Yunnan, Lincang Pref., Bangma Shan, E pass, 17 km NW Lincang, 2040 m, 23 �� 57 ' 31 ''N / 99 �� 56 ' 13 ''E, secondary pine forest with ferns, litter sifted, 9.IX. 2009, leg. D.W.Wrase [36], 1 ex. (cSch); C-Yunnan [23], mountain W Yuxi, 24 �� 27 ' 11 ''N, 102 �� 29 ' 58 ''E, 2250 m, secondary mixed forest, litter, roots, and moss sifted, 31.VIII. 2014, leg. V. Assing [23], 1 ex. (cAss); C-Yunnan [6], NE Kunming, 25 ��08' 35 ''N, 102 �� 53 ' 49 ''E, 2320 m, mixed forest with alder, oak, and pine, litter and mushrooms sifted, 13.VIII. 2014, leg. V. Assing [06], 1 ex. (cAss); C-Yunnan [5], NE Kunming, 25 ��08' 40 ''N, 102 �� 53 ' 48 ''E, 2290 m, mixed deciduous forest with scattered pine trees, litter and mushrooms sifted, 11.VIII. 2014, leg. V. Assing [05], 1 ex. (cAss); S-Yunnan [22 / 22 a], SE Pingbian, 22 �� 54 ' 31 ''N, 103 �� 41 ' 44 ''E, 2100 m, primary subtropical broad-leaved forest, litter sifted, 27./ 28.VIII. 2014, leg. V. Assing, 14 exs. (cAss, cSch); Yunnan [2 a], mountains NE Dongchuan, 26 �� 14 ' 10 ''N, 103 �� 12 ' 31 ''E, 2670 m, scree slope with bushes and herbs, litter sifted, 9.VIII. 2014, leg. V. Assing, 1 ex. (cAss); N-Yunnan, Nujiang Lisu Autonomous Prefecture, Gongshan County, N Gaoligong Shan, 2500m, 27 �� 45.404 'N, 98 �� 35.749 'E, litter & debries at snowfield sifted during rain, 19.VI. 2005, leg. M. Sch��lke [C 2005 - 12], 2 exs. (cSch); S-Yunnan, Xishuangbanna, 23 km NW Jinghong, vic. Na Ban (NNNR), 22 ��09.49'N, 100 �� 39.92 'E, forest EKL, 26.III. 2009, leg. L. Mengl, 1 ex. (NME); S-Yunnan, Xishuangbanna, 20 km NW Jinghong, near Man Dian (NNNR), pitfall trap, 22 ��07.80'N, 100 �� 40.05 'E, 740 m, rubber plantation, 28.VI. 2008, leg. A. Weigel, 1 ex. (NME); Zheijiang: Tianmu Shan, pass 25 km NNW Linan, 30 �� 25 ' 40 ''N, 119 �� 35 ' 30 ''E, 620���820 m, creek valley with bamboo and mixed forest, litter sifted, 16.VI. 2007, leg. A. P��tz [CH 07- 37], 5 exs. (cP��t); same data, but "leg. D.W. Wrase", 3 exs. (cSch); West Tianmu Shan N.R. way to peak of immortals, 30 �� 20 ' 34 ''N, 119 �� 25 ' 51 ''E, 1100���1200 m, primary mixed forest, litter and moss sifted, 15.VI. 2007, leg. A. P��tz [CH 07- 36], 2 exs. (cP��t); Tianmu Shan, pass 25 km NNW Linan, 620���820 m, 30 �� 25 ' 40 ''N, 119 �� 35 ' 30 ''E, creek valley with bamboo and mixed forest, litter, sifted, 16.VI. 2007, leg. M. Sch��lke [CH 07- 37], 5 exs. (cSch); Zheijiang/Anhui: Zheijiang/Anhui border, pass W Linan, 30 �� 17 ' 59 ''N, 118 �� 52 ' 41 ''E, 625 m, walnut plantation, plant refuse and litter sifted, 19.VI. 2007, leg. M. Sch��lke, 1 ex. (cSch); TAIWAN: Ilan Hsien: 75 km, N crossing highway, 1100 m, 22.V. 2003, flight intercept trap, leg. Chun Lin Li, 2 exs. (NMB); Shen-Mi Lake, 24 �� 22 ' 43 ''N, 121 �� 44 ' 12 ''E, 1110 m, 10.V. 1995, leg. A. Smetana [T 177], 1 ex. (cSme); Shen-Mi Lake, 24 �� 22 ' 43 ''N, 121 �� 44 ' 12 ''E, 1110 m, 9.V. 1995, leg. A. Smetana [T 175], 4 exs. (cSme, cSch); Kaohsiung Hsien, Tengchih, 1400 m, 21.��� 23. VII. 2000, leg. H. Sugaya, 2 exs. (KUM, cSch��); Tengchih, 1400 m, 25.��� 30. IV. 2001, leg. H. Sugaya, 3 exs. (KUM, cSch��); Tengchih, 2100 m, 19.��� 23. IV. 2001, leg. H. Sugaya, 6 exs. (KUM, cSch��); Tengchih, 1700 m, 2.��� 3. IV. 2002, litter, leg. H. Sugaya, 2 exs. (KUM); Tona Forest Station, 1050 m, 27.��� 30.IV. 1998 (flight intercept trap), leg. A. Smetana & L. Robillard [T 193], 1 ex. (cSch); Road above Tona Forest Station, km 16���17, 1700���1800 m, 28.IV. 1998, leg. A. Smetana [T 190]. 2 exs. (cSme); Nantou Hsien: Meifeng, 2100 m, 6. V. 2001, leg. H. Sugaya, 20 exs. (KUM, cSch��); Meimu, 1600 m, litter, 30. III. 2002, leg. H. Sugaya, 5 exs. (KUM); Tehuashe, 900 m, 4.��� 5. V. 2001, leg. H. Sugaya, 1 ex. (KUM); Nanshanchi, 800 m, 27. X. 2000, leg. H. Sugaya, 2 exs. (KUM, cSch��); Meifeng, 2130 m, 4.V. 1998, leg. A. Smetana [T 199], 1 ex. (cSch); Pintung Hsien: Chialoshui, 100 m, 11. III. 2000, leg. M. Maruyama, 1 ex. (KUM); Taichung Hsien: Anmashan, 2150 m, 13.V. 1992, leg. A. Smetana [T 129], 1 ex. (cSme); Anmashan, 2225 m, 2.V. 1990, leg. A. Smetana [T 37], 1 ex (cSme); Anmashan, 2000 m, 24.��� 26.VI. 2003, flight intercept trap, leg. Chun Lin Li, 4 exs. (NMB); Chipen- Wenchuan, 400 m, 6.��� 8. XI. 2000, leg. H. Sugaya, 3 exs. (KUM, cSch��); Chipen-Wenchuan, 400 m, 24.��� 27.IV. 2001, leg. H. Sugaya, 5 exs. (KUM, cSch��); Taitung Hsien: road no. 20, after Li-Tao (Lidau), km 174, 1300 m, mountain forest litter, 8.IV. 2007, leg. S. V��t, 2 exs. (cSch); W-Taitung, road no. 24, Jhihben, forest recreation area, 400 m, litter sifted, 12.IV. 2007, leg. S. V��t, 1 ex. (cSch); Taipeh: Noi Dong logging road, 850 m, 19.II. 2004, flight intercept trap, leg. Chun Lin Li, 3 exs. (MNB); Remarks. Ischnosoma quadriguttatum is one of the most widespread species of the genus, its distribution ranging from Pakistan to Indonesia. In China the species is very common in the central and southern parts of the country (Fig. 70). Records from Fujian, Gansu, Hainan, Hubei and Shaanxi are new province records., Published as part of Kocian, Mat���� & Sch��lke, Michael, 2016, New species and records of Ischnosoma Stephens (Coleoptera, Staphylinidae, Tachyporinae) from China, pp. 201-242 in Zootaxa 4105 (3) on pages 233-237, DOI: 10.11646/zootaxa.4105.3.1, http://zenodo.org/record/271275, {"references":["Champion, G. C. (1923) Some Indian Coleoptera (10). The Entomologist's Monthly Magazine, 59, 43 - 53, 77 - 80."]}