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34 results on '"Iankoshvili, Giorgi"'

1. The impact of insect herbivory on biogeochemical cycling in broadleaved forests varies with temperature

Author

Hwang, Bernice C., Giardina, Christian P., Adu-Bredu, Stephen, Barrios-Garcia, M. Noelia, Calvo-Alvarado, Julio C., Dargie, Greta C., Diao, Haoyu, Duboscq-Carra, Virginia G., Hemp, Andreas, Hemp, Claudia, Huasco, Walter Huaraca, Ivanov, Aleksandr V., Johnson, Nels G., Kuijper, Dries P. J., Lewis, Simon L., Lobos-Catalán, Paulina, Malhi, Yadvinder, Marshall, Andrew R., Mumladze, Levan, Ngute, Alain Senghor K., Palma, Ana C., Petritan, Ion Catalin, Rordriguez-Cabal, Mariano A., Suspense, Ifo A., Zagidullina, Asiia, Andersson, Tommi, Galiano-Cabrera, Darcy F., Jiménez-Castillo, Mylthon, Churski, Marcin, Gage, Shelley A., Filippova, Nina, Francisco, Kainana S., Gaglianese-Woody, Morgan, Iankoshvili, Giorgi, Kaswamila, Mgeta Adidas, Lyatuu, Herman, Mampouya Wenina, Y. E., Materu, Brayan, Mbemba, M., Moritz, Ruslan, Orang, Karma, Plyusnin, Sergey, Puma Vilca, Beisit L., Rodríguez-Solís, Maria, Šamonil, Pavel, Stępniak, Kinga M., Walsh, Seana K., Xu, Han, and Metcalfe, Daniel B.

Published
2024
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5. The impact of insect herbivory on biogeochemical cycling in broadleaved forests varies with temperature

Author

Hwang, Bernice, Giardina, Christian P., Adu-Bredu, Stephen, Barrios-Garcia, M. Noelia, Calvo-Alvarado, Julio C., Dargie, Greta C., Diao, Haoyu, Duboscq-Carra, Virginia G., Hemp, Andreas, Hemp, Claudia, Huasco, Walter Huaraca, Ivanov, Aleksandr V., Johnson, Nels G., Kuijper, Dries P. J., Lewis, Simon L., Lobos-Catalán, Paulina, Malhi, Yadvinder, Marshall, Andrew R., Mumladze, Levan, Ngute, Alain Senghor K., Palma, Ana C., Petritan, Ion Catalin, Rordriguez-Cabal, Mariano A., Suspense, Ifo A., Zagidullina, Asiia, Andersson, Tommi, Galiano-Cabrera, Darcy F., Jiménez-Castillo, Mylthon, Churski, Marcin, Gage, Shelley A., Filippova, Nina, Francisco, Kainana S., Gaglianese-Woody, Morgan, Iankoshvili, Giorgi, Kaswamila, Mgeta Adidas, Lyatuu, Herman, Mampouya Wenina, Y.E., Materu, Brayan, Mbemba, M., Moritz, Ruslan, Orang, Karma, Plyusnin, Sergey, Puma Vilca, Beisit L., Rodríguez-Solís, Maria, Šamonil, Pavel, Stępniak, Kinga M., Walsh, Seana K., Xu, Han, Metcalfe, Daniel B., Hwang, Bernice, Giardina, Christian P., Adu-Bredu, Stephen, Barrios-Garcia, M. Noelia, Calvo-Alvarado, Julio C., Dargie, Greta C., Diao, Haoyu, Duboscq-Carra, Virginia G., Hemp, Andreas, Hemp, Claudia, Huasco, Walter Huaraca, Ivanov, Aleksandr V., Johnson, Nels G., Kuijper, Dries P. J., Lewis, Simon L., Lobos-Catalán, Paulina, Malhi, Yadvinder, Marshall, Andrew R., Mumladze, Levan, Ngute, Alain Senghor K., Palma, Ana C., Petritan, Ion Catalin, Rordriguez-Cabal, Mariano A., Suspense, Ifo A., Zagidullina, Asiia, Andersson, Tommi, Galiano-Cabrera, Darcy F., Jiménez-Castillo, Mylthon, Churski, Marcin, Gage, Shelley A., Filippova, Nina, Francisco, Kainana S., Gaglianese-Woody, Morgan, Iankoshvili, Giorgi, Kaswamila, Mgeta Adidas, Lyatuu, Herman, Mampouya Wenina, Y.E., Materu, Brayan, Mbemba, M., Moritz, Ruslan, Orang, Karma, Plyusnin, Sergey, Puma Vilca, Beisit L., Rodríguez-Solís, Maria, Šamonil, Pavel, Stępniak, Kinga M., Walsh, Seana K., Xu, Han, and Metcalfe, Daniel B.

Abstract

Herbivorous insects alter biogeochemical cycling within forests, but the magnitude of these impacts, their global variation, and drivers of this variation remain poorly understood. To address this knowledge gap and help improve biogeochemical models, we established a global network of 74 plots within 40 mature, undisturbed broadleaved forests. We analyzed freshly senesced and green leaves for carbon, nitrogen, phosphorus and silica concentrations, foliar production and herbivory, and stand-level nutrient fluxes. We show more nutrient release by insect herbivores at non-outbreak levels in tropical forests than temperate and boreal forests, that these fluxes increase strongly with mean annual temperature, and that they exceed atmospheric deposition inputs in some localities. Thus, background levels of insect herbivory are sufficiently large to both alter ecosystem element cycling and influence terrestrial carbon cycling. Further, climate can affect interactions between natural populations of plants and herbivores with important consequences for global biogeochemical cycles across broadleaved forests.

Published
2024
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8. Into the unknown: the first barcode-assisted checklist of Psocoptera (Insecta, Psocodea) of Georgia with a census on country species richness

Author

Seropian, Armen, primary, Arsenashvili, Eka, additional, Bulbulashvili, Natalia, additional, Shubashishvili, Anano, additional, Iankoshvili, Giorgi, additional, Todua, Mariami, additional, Ananiashvili, Ana, additional, Japarashvili, Shota, additional, Chkhartisvhili, Tinatin, additional, Memishishi, Aleksi, additional, Balkhamishvili, Sopio, additional, Chitadze, Beka, additional, Karalashvili, Elisabeth, additional, Mumladze, Levan, additional, Hein, Nils, additional, and Rulik, Björn, additional

Published
2023
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9. Into the unknown: the first barcode-assisted checklist of Psocoptera (Insecta, Psocodea) of Georgia with a census on country species richness

Author

Seropian, Armen, Arsenashvili, Eka, Bulbulashvili, Natalia, Shubashishvili, Anano, Iankoshvili, Giorgi, Todua, Mariami, Ananiashvili, Ana, Japarashvili, Shota, Chkhartishvili, Tinatini, Memishishi, Aleksi, Balkhamishvili, Sopio, Chitadze, Beka, Karalashvili, Elisabeth, Mumladze, Levan, Hein, Nils, and Rulik, Björn

Subjects
Insecta, Arthropoda, CaBOL, faunistics, Animalia, Biodiversity, South Caucasus, Biota, Psocodea
Abstract

This checklist reports 47 species of Psocoptera from 15 families and three suborders from Georgia, of which 31 species are recorded for the first time, increasing the known fauna of the country by more than 65%. Of these, 37 species have been barcoded, representing 210 Barcode Identification Numbers (BINs). An additional 14 species are expected to occur in Georgia but remain undiscovered, meaning that only ≈ 77% of the fauna is currently documented. Barcodes, comments on distributions, and images of voucher specimens are given followed by a map of the sampling sites.

Published
2023

10. New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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11. Figure 1 from: Japarashvili S, Bulbulashvili N, Seropian A, Chkhartishvili T, Iankoshvili G, Chitadze B, Balkhamishvili S, Arsenashvili E, Todua M, Memishishi A (2023) New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa. Caucasiana 2: 63-70. https://doi.org/10.3897/caucasiana.2.e98998

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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12. Figure 4 from: Japarashvili S, Bulbulashvili N, Seropian A, Chkhartishvili T, Iankoshvili G, Chitadze B, Balkhamishvili S, Arsenashvili E, Todua M, Memishishi A (2023) New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa. Caucasiana 2: 63-70. https://doi.org/10.3897/caucasiana.2.e98998

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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13. Figure 5 from: Japarashvili S, Bulbulashvili N, Seropian A, Chkhartishvili T, Iankoshvili G, Chitadze B, Balkhamishvili S, Arsenashvili E, Todua M, Memishishi A (2023) New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa. Caucasiana 2: 63-70. https://doi.org/10.3897/caucasiana.2.e98998

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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14. Figure 3 from: Japarashvili S, Bulbulashvili N, Seropian A, Chkhartishvili T, Iankoshvili G, Chitadze B, Balkhamishvili S, Arsenashvili E, Todua M, Memishishi A (2023) New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa. Caucasiana 2: 63-70. https://doi.org/10.3897/caucasiana.2.e98998

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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15. Figure 6 from: Japarashvili S, Bulbulashvili N, Seropian A, Chkhartishvili T, Iankoshvili G, Chitadze B, Balkhamishvili S, Arsenashvili E, Todua M, Memishishi A (2023) New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa. Caucasiana 2: 63-70. https://doi.org/10.3897/caucasiana.2.e98998

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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16. Figure 2 from: Japarashvili S, Bulbulashvili N, Seropian A, Chkhartishvili T, Iankoshvili G, Chitadze B, Balkhamishvili S, Arsenashvili E, Todua M, Memishishi A (2023) New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa. Caucasiana 2: 63-70. https://doi.org/10.3897/caucasiana.2.e98998

Author

Japarashvili, Shota, primary, Bulbulashvili, Natalia, additional, Seropian, Armen, additional, Chkhartishvili, Tinatin, additional, Iankoshvili, Giorgi, additional, Chitadze, Beka, additional, Balkhamishvili, Sopio, additional, Arsenashvili, Eka, additional, Todua, Mariami, additional, and Memishishi, Aleksi, additional

Published
2023
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18. The farther, the closer: geographic proximity and niche overlap versus genetic divergence in Caucasian rock lizards.

Author

Tarkhnishvili, David and Iankoshvili, Giorgi

Subjects
*LIZARDS, *TIME measurements, *VICARIANCE
Abstract

Caucasian rock lizards (Darevskia) are a speciose group with 42 described species from the Caucasus Mountain region. We selected 11 species from Georgia and adjacent territories to explore whether their distribution depends on the distribution of congeneric taxa. We estimated two measures for each species pair: (1) niche overlap between all pairs of the studied species; (2) the extent of contact between the ranges of individual species. We then calculated the correlation between these measurements and the time of divergence between the species. The results showed that in sympatric and parapatric (but not allopatric) species, there is a highly significant correlation between the divergence time and the extent of the contact, as well as between the divergence time and the overlap of suitable habitats. The ranges of the evolutionarily closest species have the smallest extent of contact. Species separated later have overlapping ranges, although are separated by altitude. The most distant species, irrespective of their phenotype, are commonly found in the same location. We suggest that a gradual increase in the contact extent is compensated for by increasing the effectiveness of other mechanisms of niche shift or behavioural isolation and hence the frequency of potentially deleterious interspecific contacts is kept low. [ABSTRACT FROM AUTHOR]

Published
2023
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20. Figure 1 from: Arsenashvili E, Iankoshvili G, Japarashvili S, Bulbulashvili N, Seropian A (2022) Discovery of Hemilepistus (Hemilepistus) klugii (Brandt, 1833) (Isopoda, Oniscidea) in the Republic of Georgia with an overview of the ecology and biology of the subgenus. Caucasiana 1: 13-16. https://doi.org/10.3897/caucasiana.1.e86183

Author

Arsenashvili, Eka, primary, Iankoshvili, Giorgi, additional, Japarashvili, Shota, additional, Bulbulashvili, Natalia, additional, and Seropian, Armen, additional

Published
2022
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21. New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia with an update on the distribution of some other little studied taxa.

Author

Japarashvili, Shota, Bulbulashvili, Natalia, Seropian, Armen, Chkhartishvili, Tinatin, Iankoshvili, Giorgi, Chitadze, Beka, Balkhamishvili, Sopio, Arsenashvili, Eka, Todua, Mariami, and Memishishi, Aleksi

Subjects
INSECTS, SPECIES, HYDROPHILIDAE, BEETLES, MITOCHONDRIAL DNA
Abstract

First records of two species of beetles (Insecta, Coleoptera), namely Pocadius adustus Reitter, 1888 (Nitidulidae) and Byrrhus pustulatus (Forster, 1770) (Byrrhidae) are given for Georgia. In addition, an update on the occurrences of Necrobia violacea (Linnaeus, 1758) (Cleridae), Coelostoma orbiculare (Fabricius, 1775) (Hydrophilidae) and Lycoperdina succincta (Linnaeus, 1767) (Endomychidae) are also presented based on the specimens collected in Samtskhe-Javakheti region, determined by morphological and mtDNA barcoding methods. [ABSTRACT FROM AUTHOR]

Published
2023
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23. Figure 3 from: Barateli N, Tarkhnishvili D, Iankoshvili G, Kokiashvili L, Dvali N, Janiashvili Z (2021) Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa 34: 71-81. https://doi.org/10.3897/herpetozoa.34.e63072

Author

Barateli, Natia, primary, Tarkhnishvili, David, additional, Iankoshvili, Giorgi, additional, Kokiashvili, Luka, additional, Dvali, Nikoloz, additional, and Janiashvili, Zurab, additional

Published
2021
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24. Supplementary material 1 from: Barateli N, Tarkhnishvili D, Iankoshvili G, Kokiashvili L, Dvali N, Janiashvili Z (2021) Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa 34: 71-81. https://doi.org/10.3897/herpetozoa.34.e63072

Author

Barateli, Natia, primary, Tarkhnishvili, David, additional, Iankoshvili, Giorgi, additional, Kokiashvili, Luka, additional, Dvali, Nikoloz, additional, and Janiashvili, Zurab, additional

Published
2021
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25. Figure 4 from: Barateli N, Tarkhnishvili D, Iankoshvili G, Kokiashvili L, Dvali N, Janiashvili Z (2021) Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa 34: 71-81. https://doi.org/10.3897/herpetozoa.34.e63072

Author

Barateli, Natia, primary, Tarkhnishvili, David, additional, Iankoshvili, Giorgi, additional, Kokiashvili, Luka, additional, Dvali, Nikoloz, additional, and Janiashvili, Zurab, additional

Published
2021
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27. Figure 2 from: Barateli N, Tarkhnishvili D, Iankoshvili G, Kokiashvili L, Dvali N, Janiashvili Z (2021) Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa 34: 71-81. https://doi.org/10.3897/herpetozoa.34.e63072

Author

Barateli, Natia, primary, Tarkhnishvili, David, additional, Iankoshvili, Giorgi, additional, Kokiashvili, Luka, additional, Dvali, Nikoloz, additional, and Janiashvili, Zurab, additional

Published
2021
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28. Figure 1 from: Barateli N, Tarkhnishvili D, Iankoshvili G, Kokiashvili L, Dvali N, Janiashvili Z (2021) Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa 34: 71-81. https://doi.org/10.3897/herpetozoa.34.e63072

Author

Barateli, Natia, primary, Tarkhnishvili, David, additional, Iankoshvili, Giorgi, additional, Kokiashvili, Luka, additional, Dvali, Nikoloz, additional, and Janiashvili, Zurab, additional

Published
2021
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29. Supplementary material 2 from: Barateli N, Tarkhnishvili D, Iankoshvili G, Kokiashvili L, Dvali N, Janiashvili Z (2021) Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa 34: 71-81. https://doi.org/10.3897/herpetozoa.34.e63072

Author

Barateli, Natia, primary, Tarkhnishvili, David, additional, Iankoshvili, Giorgi, additional, Kokiashvili, Luka, additional, Dvali, Nikoloz, additional, and Janiashvili, Zurab, additional

Published
2021
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30. Precise paternal ancestry of hybrid unisexual ZW lizards (genus Darevskia: Lacertidae: Squamata) revealed by Z-linked genomic markers.

Author

Yanchukov, Alexey, Tarkhnishvili, David, Erdolu, Meriç, Şahin, Mehmet Kürşat, Candan, Kamil, Murtskhvaladze, Marine, Gabelaia, Mariam, Iankoshvili, Giorgi, Barateli, Natia, Ilgaz, Çetin, Kumlutaş, Yusuf, Matur, Ferhat, Çolak, Faruk, Arakelyan, Marine, and Galoyan, Eduard

Abstract

We genotyped multiple populations of all seven parthenogenetic species of rock lizards, genus Darevskia , as well as their putative sexual parental populations, using double digest RAD-sequencing genomic markers. Taking advantage of the conserved homology of the ZW/ZZ sex chromosomes among lacertid lizards, we aligned our short sequence reads to a reference Z chromosome assembly of the common wall lizard Podarcis muralis. This provided unique insight into the origin of all-female hybrid populations, which inherited their single Z chromosome exclusively from a paternal ancestor. The mapped Z-linked loci were used to construct a robust time-calibrated phylogeny. In each parent–offspring species pair, the geographically nearest population of the paternal species was identified as the most likely ancestor of the respective parthenogen, mirroring the trend observed previously on the maternal side in mitochondrial DNA. The estimated splits between the sampled paternal sexual ancestors and their daughter species occurred much earlier than suggested previously and during two narrow time periods: (1) the parthenogenetic Darevskia armeniaca , D. dahli , D. uzzelli and D. rostombekowi dated back to ~0.5 or ~0.9 Mya, depending on the calibration point used, while (2) D. bendimahiensis , D. sapphirina and D. unisexualis appear to have diverged ~1 or ~2 Mya. [ABSTRACT FROM AUTHOR]

Published
2022
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31. Genotypic Similarities Among The Parthenogeneticdarevskiarock Lizards With Different Hybrid Origins

Author

Tarkhnishvili, David, Yanchukov, Alexey, Sahin, Mehmet Kursat, Gabelaia, Mariam, Murtskhvaladze, Marine, Candan, Kamil, Galoyan, Eduard, Arakelyan, Marine, Iankoshvili, Giorgi, Kumlutas, Yusuf, Ilgaz, Cetin, Matur, Ferhat, colak, Faruk, Erdolu, Meric, Kurdadze, Sofiko, Barateli, Natia, and Anderson, Cort L.

Abstract

Background The majority of parthenogenetic vertebrates derive from hybridization between sexually reproducing species, but the exact number of hybridization events ancestral to currently extant clonal lineages is difficult to determine. Usually, we do not know whether the parental species are able to contribute their genes to the parthenogenetic vertebrate lineages after the initial hybridization. In this paper, we address the hypothesis, whether some genotypes of seven phenotypically distinct parthenogenetic rock lizards (genusDarevskia) could have resulted from back-crosses of parthenogens with their presumed parental species. We also tried to identify, as precise as possible, the ancestral populations of all seven parthenogens. Results We analysed partial mtDNA sequences and microsatellite genotypes of all seven parthenogens and their presumed ansectral species, sampled across the entire geographic range of parthenogenesis in this group. Our results confirm the previous designation of the parental species, but further specify the maternal populations that are likely ancestral to different parthenogenetic lineages. Contrary to the expectation of independent hybrid origins of the unisexual taxa, we found that genotypes at multiple loci were shared frequently between different parthenogenetic species. The highest proportions of shared genotypes were detected between (i)D. sapphirinaandD. bendimahiensisand (ii)D. dahliandD. armeniaca, and less often between other parthenogens. In case (ii), genotypes at the remaining loci were notably distinct. Conclusions We suggest that both observations (i-ii) can be explained by two parthenogenetic forms tracing their origin to a single initial hybridization event. In case (ii), however, occasional gene exchange between the unisexual and the parental bisexual species could have taken placeafterthe onset of parthenogenetic reproduction. Indeed, backcrossed polyploid hybrids are relatively frequent inDarevskia, although no direct evidence of recent gene flow has been previously documented. Our results further suggest that parthenogens are losing heterozygosity as a result of allelic conversion, hence their fitness is expected to decline over time as genetic diversity declines. Backcrosses with the parental species could be a rescue mechanism which might prevent this decline, and therefore increase the persistance of unisexual forms.

Published
2020

34. Isolated population of the Middle Eastern Phoenicolacerta laevisfrom the Georgian Black Sea Coast, and its genetic closeness to populations from southern Turkey

Author

Tarkhnishvili, David, Gabelaia, Mariam, Kandaurov, Andrei, Bukhnikashvili, Alexander, and Iankoshvili, Giorgi

Abstract

The Lebanon Lizard (Phoenicolacerta laevis) occurs on the Levantine coast and scattered populations are found in isolated coastal habitats along the southern and south-western coast of Turkey. We found an isolated but dense population of this species at and around the castle of Anaklia on Georgia’s Black Sea Coast, near the mouth of the river Enguri. The analysis of mitochondrial cytochrome b sequence suggests closeness of this population to the populations in the provinces of Kahramanmaraş, Adana and Içel in southern Turkey, rather than to those in Lebanon and Israel. It is thought that the species was introduced from a Turkish region to Anaklia, which was an important trade location at the eastern Black Sea coast until the late 18thcentury. This is the first established location of this species outside the eastern Mediterranean area. Climate and competition with rock lizards of the genus Darevskiaare the most likely reasons preventing expansion of the species into the neighboring areas of Western Georgia.

Published
2017
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