222 results on '"Hairapetian, Vachik"'
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2. First palynological evidence from the Upper Devonian of Armenia (northern Gondwanan margin): biostratigraphic implications
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Khachatryan, Sirush, Cascales-Miñana, Borja, Danelian, Taniel, Breuer, Pierre, Steemans, Philippe, Grigoryan, Araik, Gabrielyan, Ivan, Hairapetian, Vachik, Regnier, Sylvie, Kroeck, David Marius, and Serobyan, Vahram
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- 2024
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3. The Permian–Triassic boundary section at Baghuk Mountain, Central Iran: carbonate microfacies and depositional environment
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Heuer, Franziska, Leda, Lucyna, Moradi-Salimi, Hemen, Gliwa, Jana, Hairapetian, Vachik, and Korn, Dieter
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- 2022
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4. Stratigraphic and volcanic signatures of Miaolingian-Late Ordovician rift pulses in the Alborz Mountains, northern Iran
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Álvaro, J. Javier, Ghobadi Pour, Mansoureh, Sánchez-García, Teresa, Kebria-ee Zadeh, Mohammad-Reza, Hairapetian, Vachik, and Popov, Leonid E.
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- 2022
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5. Mid-Cretaceous biostratigraphy (ammonites, inoceramid bivalves and foraminifers) at the eastern margin of the Anarak Metamorphic Complex (Central Iran)
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Wilmsen, Markus, Berensmeier, Michaela, Fürsich, Franz Theodor, Schlagintweit, Felix, Hairapetian, Vachik, Pashazadeh, Behnam, and Majidifard, Mahmoud Reza
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- 2020
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6. The early elasmobranch Phoebodus : phylogenetic relationships, ecomorphology and a new time-scale for shark evolution
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Frey, Linda, Coates, Michael, Ginter, Michał, Hairapetian, Vachik, Rücklin, Martin, Jerjen, Iwan, and Klug, Christian
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- 2019
7. Integrated stratigraphy, facies analysis and correlation of the upper Albian–lower Turonian of the Esfahan area (Iran): Unravelling the conundrum of the so-called “Glauconitic Limestone”
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Hairapetian, Vachik, Wilmsen, Markus, Ahmadi, Amir, Shojaei, Ziba, Berensmeier, Michaela, and Majidifard, Mahmoud Reza
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- 2018
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8. Fusulinoids from the Carboniferous–Permian transition beds from the Abadeh Region (Sanandaj–Sirjan Zone, Iran)
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Fassihi, Shirin, Sone, Masatoshi, Hairapetian, Vachik, and Esfahani, Fariba Shirezadeh
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- 2019
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9. EARLY FRASNIAN THELODONT SCALES FROM CENTRAL IRAN AND THEIR IMPLICATIONS FOR TURINIID TAXONOMY, SYSTEMATICS AND DISTRIBUTION
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HAIRAPETIAN, VACHIK, BLOM, HENNING, and TURNER, SUSAN
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- 2016
10. Danburite Crystals from Iran.
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Hairapetian, Vachik and Namnabat, Elahe
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CRYSTALS , *MINERALS , *GEOCHEMISTRY , *BORATE minerals , *IRON ores , *DOLOMITE , *PYRITES - Abstract
The danburite crystals from Larak Island are most often doubly terminated and can be quickly distinguished from those from Hormuz, which are characterized by a well-developed {001} face and more modifications including the forms {120}, {110}, {100}, and {011}. Danburite crystals on rhyolitic matrix from Larak Island; specimen width 12 cm, largest crystal 2.5 cm. Vachik Hairapetian specimen and photo. In addition to the differences in crystal size, transparency, and color, crystals of danburite from Hormuz Island tend to develop as more complex clusters than those from Larak Island. Danburite crystals on rhyolitic matrix from Larak Island; specimen width 7.1 cm, largest crystal 1.3 cm. Vachik Hairapetian specimen and photo. [Extracted from the article]
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- 2023
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11. Fusulinoids from the Bashkirian–Moscovian transition beds of the Shahreza region in the Sanandaj–Sirjan Zone, Iran
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Fassihi, Shirin, Sone, Masatoshi, Hairapetian, Vachik, and Esfahani, Fariba Shirezadeh
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- 2017
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12. The Permian–Triassic boundary section at Baghuk Mountain, Central Iran: carbonate microfacies and depositional environment
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Heuer, Franziska, Leda, Lucyna, Moradi-Salimi, Hemen, Gliwa, Jana, Hairapetian, Vachik, Korn, Dieter, Museum Für Naturkunde, Leibniz Institute for Research On Evolution and Biodiversity, Berlin, Germany, Geology Department, Faculty of Science, University of Isfahan, Isfahan, Iran, Eurasia Department and Beijing Branch Office, German Archaeological Institute, Berlin, Germany, and Department of Geology, Islamic Azad University, Isfahan (Khorasgan) BranchIsfahan, Iran
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Extinction event ,Global and Planetary Change ,Ecology ,Permian ,biology ,Central Iran ,ddc:563 ,Microbialites ,Early Triassic ,Geochemistry ,Paleontology ,Geology ,biology.organism_classification ,Permian–Triassic boundary ,Claraia ,Sedimentary depositional environment ,chemistry.chemical_compound ,chemistry ,Sponges ,Carbonate microfacies ,Subaerial ,Carbonate ,Sedimentary rock ,Ecology, Evolution, Behavior and Systematics - Abstract
Sections at Baghuk Mountain, 45 km NNW of Abadeh (Central Iran), have excellent exposures of fossiliferous marine Late Permian to Early Triassic sedimentary successions. Detailed bed-by-bed sampling enables the analysis of microfacies changes of three successive rock units across the Permian–Triassic boundary. The Late Permian Hambast Formation is mainly the result of biogenic carbonate production. Its carbonate microfacies is dominated by biogen-rich and bioturbated nodular limestones, indicating a well-oxygenated aphotic to dysphotic environment. The biogen-dominated carbonate factory in the Permian ceased simultaneously with the main mass extinction pulse, which is marked by a sharp contact between the Hambast-Formation and the overlaying Baghuk Member (= ‘Boundary Clay’). The clay and silt deposits of the Baghuk Member with some carbonate beds show only a few signs of bioturbation or relics of benthic communities. The Early Triassic Claraia Beds are characterised by a partly microbially induced carbonate production, which is indicated by frequent microbialite structures. The depositional environment does not provide evidence of large amplitude changes of sea level or subaerial exposure during the Permian–Triassic boundary interval. The deposition of the Baghuk Mountain sediments took place in a deep shelf environment, most of the time below the storm wave base., Deutsche Forschungsgemeinschaft http://dx.doi.org/10.13039/501100001659, Museum für Naturkunde – Leibniz-Institut für Evolutions- und Biodiversitätsforschung (3498)
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- 2021
13. Correction to: Fusulinoids from the Carboniferous–Permian transition beds from the Abadeh Region (Sanandaj–Sirjan Zone, Iran)
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Fassihi, Shirin, Sone, Masatoshi, Hairapetian, Vachik, and Esfahani, Fariba Shirezadeh
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- 2019
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14. FRASNIAN (UPPER DEVONIAN) BRACHIOPODS FROM ARMENIA: BIOSTRATIGRAPHIC AND PALAEOBIOGEOGRAPHIC IMPLICATIONS.
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SEROBYAN, VAHRAM, DANELIAN, TANIEL, HAIRAPETIAN, VACHIK, GRIGORYAN, ARAIK, CRÔNIER, CATHERINE, RANDON, CARINE, and MOTTEQUIN, BERNARD
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An assemblage of seven brachiopod species belonging to the orders Rhynchonellida, Atrypida and Spiriferida are studied from three localities (Ertych, Djravank and Noravank) of Central Armenia. The examined material is recovered from shallow water nodular limestones and provides insights into the diversity of Frasnian brachiopods on that part of the northern margin of Gondwana preserved within the South Armenian Block. The revision of Atrypa (Planatrypa) ertichensis, a biostratigraphically significant species for the Frasnian of the Lesser Caucasus (Armenia and Nakhichevan), revealed the presence of frills, an ornamental feature rarely observed in Atrypa (Planatrypa) representatives and considered as unknown in this species. Taxonomic discussion also involves the selection of neotypes for Ripidiorhynchus gnishikensis and A. (P.) ertichensis. The newly described taxon, Angustisulcispirifer arakelyani n. gen., n. sp., appears to be one of the most biostratigraphically important species for the Frasnian of Armenia. The size variability of Cyphoterorhynchus koraghensis and Desquamatia (Seratrypa) abramianae is documented quantitatively for the first time and it shows a continuous and progressive growth without any distinct groupings; the former is a palaeobiogeographically important species for the Frasnian strata of the northern Gondwana margin. Pending the revision of the Pakistani and Iranian material ascribed to C. koraghensis, that may include several subspecies, a plaster cast of its lectotype from the Frasnian of Kuragh in Chitral (northwest Pakistan) and the holotype as well as one of the paratypes of Cyphoterorhynchus koraghensis interpositus from the Frasnian Bahram Formation of the Ozbak-Kuh region in eastern Iran are illustrated herein. Finally, a new Frasnian brachiopod zone, namely the Ripidiorhynchus gnishikensis–Angustisulcispirifer arakelyani assemblage Zone is here introduced for the studied sections. Although its base and top cannot be identified, it is constrained to the Frasnian based on conodonts identified in the Djravank section. It may be considered as a partly lateral equivalent of the Cyrtospirifer subarchiaci–Cyphoterorhynchus arpaensis brachiopod Zone established in Nakhichevan. [ABSTRACT FROM AUTHOR]
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- 2023
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15. Ammonites Biostratigraphy of the Sarcheshmeh Formation (Aptian); Hosein Abad section, northeast of the Kopet Dagh Basin.
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Bahrani, Maryam, Majidifard, Mahmoud Reza, Hairapetian, Vachik, Aleali, Seyed Mohsen, and Wilmsen, Markus
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Introduction Hosein Abad section, with a thickness of 90 meters, is located in the structural zone of KopetDagh near the border of the North Khorasan and Golestan provinces. Stratigraphically and paleontologically, it has been done on different formations of the KopetDagh basin, including the Sarcheshmeh Formation. However, detailed stratigraphic and fossil studies are still underway due to the stratigraphic features of the KopetDagh basin and various lateral changes in the formations in this basin. Are associated with importance. Biostratigraphy using Ammonites from Sarcheshmeh Formation in the Hosein Abad section has been studied. The KopetDagh Basin was formed as an intracontinental basin in northeastern Iran after the closure of the Hercynian Ocean that followed the Early Kimmerian orogeny (Berberian and King, 1981). From the Jurassic through Eocene, relatively continuous sedimentary succession forms five major transgressive-regressive sequences in the eastern KopetDagh. Fault-controlled subsidence of the KopetDagh Basin started in the late mid-Jurassic (and terminated in the Oligocene, causing the deposition of up to 10km thick sedimentary succession (Berberian and King, 1981). Materials and Methods This research has been conducted according to field, laboratory, and office studies. Through field studies, the stratigraphic features of the Sarcheshmeh Formation, including the material and thickness of the layers, were harvested. In addition, 21 samples of ammonites were collected from the base and top of the formation after washing. Then, samples were carefully examined to determine the genus and species. To determine the age and compatibility of biozones, Reboulet et al. (2009), biozonation, which has conducted a comprehensive study in the Mediterranean basin, has been used and equated. The study also identified biozones with other studies, such as Raisosadat et al. (2011) and Kotetishvili et al. (2000) have been compared in the KopetDagh and Caucasus basins (Mediterranean). Stratigraphy Sarcheshmeh Formation is 90 meters in thickness in the Hosein Abad section. From the base, this Formation consists of 40 m alternating grey to green marls and thin to medium muddy limestone. In this sequence, 10 m of medium-bedded limestone is placed, and 40 m of marl and muddy limestone are re-deposited. The lower boundary of the Sarcheshmeh Formation is composed of thick limestone of the Tirgan Formation beds. Also, the shale and silt deposits of the Sanganeh Formation have covered the upper boundary of the Sarcheshmeh Formation. Biostratigraphy Based on different references, ten species belonging to 4 genera were identified, and the stratigraphic range chart of the ammonites in the Sarcheshmeh Formation, Hosein Abad section, was determined. These are: Pedioceras sp., Deshayesitesconsobrinus, Prodeshayesiteslestrangei, Deshayesitesaff. topleyi, Deshayesitesdeshayesi, Dufrenoyiaformosa, Deshayesitesgrandis, Dufrenoyia sp., Deshayesites sp., Prodeshayesitesgermanicus, Deshayesitesinvolutus, Prodeshayesitesaff. jacksoni, Dufrenoyiamackesoni, Deshayesitesmirabilis. For Sarcheshmeh Formation in the Hosein Abad section and based on the stratigraphical distribution of ammonites, three biozones are proposed: - Deshayesitesconsobrinus This zone is introduced locally for the first time in this study, and it can be considered a Range-zone characterized by the first and last appearance of Deshayesitesconsobrinus. The thickness of this zone is 27 meters, and it starts from the base oftheSarcheshmeh Formation. Accompanying species of this zone include: Pedioceras sp., Prodeshayesiteslestrangei, Deshayesitesaff. topleyi. The identified biological zone could be equivalent to the DeshayesitesWeissi biological zone introduced by Reboulet et al. (2009). According to a 2009 study by Reboulet et al., The relative age of this zone can be attributed to the former Aptian. - Deshayesitesdeshayesi This 40 m thick zone (layers 27 to 67 m) begins with the first presence of Deshayesitesdeshayesi index species and includes marl deposits and muddy limestone from Sarcheshmeh Formation. Fossils associated with this biological zone include: Dufrenoyiaformosa, Deshayesitesgrandis, Dufrenoyia sp., Deshayesites sp., Prodeshayesitesgermanic us, Deshayesitesinvolutus, Prodeshayesitesaff. jacksoni. According to a 2009 study by Reboulet et al., The relative age of this zone can be attributed to the upper parts of the former Aptian. - Dufrenoyiamackesoni This biozone is also proposed locally for the first time in this study as the first emergence of Dufrenoyiamackesoni. The only associated species identified in this biological zone is Deshayesitesmirabilis. This zone is 23 meters thick (layers 67 to 90 meters) and includes marl deposits and clay-limestone from Sarcheshmeh Formation. The identified biological zone could be equivalent to the Dufrenoyiafurcate biological zone introduced by Reboulet et al. (2009). According to a 2009 study by Reboulet et al., The relative age of this zone can be attributed to the late Early Aptian. 5-Comparison of identified biozones with some previous studies Due to the stratigraphic status of the biozones identified in the study of Kotetshivilii et al. (2000), the biological zone of Deshayesitesconsobrinus is equivalent to the upper part of the zone of Deshayesitesweissi-Prochelonicerasalbrechti-austriae. Furthermore, other zones separated in this study, including Deshayesitesdeshayesi and Dufrenoyiamackesoni, are comparable to the zones introduced by Deshayesitesdeshayesi and Dufrenoyiafurcata by Kotetshivilii et al. (2000). In the Mediterranean basin, Reboulet et al. (2009) equated the two habitats of Deshayesitestuarkyricus and Deshayesitesweissi with the habitat of Deshayesitesweissi-Prochelonicerasalbrechti-austriae. In this study, the Deshayesitesconsobrinus zone can be considered equivalent to the Deshayesitesweissi zone. The other zones introduced by Reboulet et al. (2009) align with the Kotetshivilii et al. (2000) study. The Dufrenoyiamackesoni biozone identified in this study can be compared to the Dufrenoyia sp. The Deshayesitesconsobrinus zone can also be considered equivalent to the Deshayesitesweissi zone. In this study, the Deshayesitesdeshayesi zone is similar to previous studies regarding stratigraphic status. Conclusion A stratigraphic section with a thickness of 90 m was selected to determine the relative age of the deposits of the Sarcheshmeh Formation in the Hosseinabad section. Based on the stratigraphy of ammonites, three biological zones, Deshayesitesconsobrinus, Deshayesitesdeshayesi, and Dufrenoyiamackesoni, were selected for Sarcheshmeh Formation, and the zones Deshayesitesconsobrinus and Dufrenoyiamackesoni were reported locally for the first time in this study. The zones introduced confirm late Late Aptian. Furthermore, a comparison of the biozones in this study with some previous studies in the KopehDagh and Mediterranean basins indicates a good correlation with other biological zones of the Aptian. [ABSTRACT FROM AUTHOR]
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- 2023
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16. A new large tetrapodomorph sarcopterygian from the Late Devonian of Iran
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Davesne, Donald, Mondéjar-Fernández, Jorge, Hairapetian, Vachik, Rücklin, Martin, Wendt, Jobst, and Clément, Gaël
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- 2015
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17. Revision of Middle Devonian conodont biostratigraphy in the Negheleh section, Central Iran
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Gholamalian, Hossein, primary, Hairapetian, Vachik, additional, and Poosti, Mohammad, additional
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- 2022
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18. Xenodiscidae Frech 1902
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Xenodiscidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Family Xenodiscidae Frech, 1902 Diagnosis Representatives of the superfamily Xenodiscoidea with a small to moderately large conch, in which the ontogeny displays up to three stages, beginning with an unsculptured initial stage followed by a juvenile stage with transverse ribs, and an adult stage with a weakening of the sculpture. Suture line with unserrated or weakly serrated, short but rather wide external lobe; adventive, lateral and umbilical lobe often multidentate; some species with a simplified suture line without serrations. Included Permian genera Iranites Teichert & Kummel, 1973; Laibinoceras Yang, 1987; Multisulcites Liang, 1983; Paraceltitoides Zheng & Chen, 1979; Penglaites Yang, 1987; Phisonites Shevyrev, 1965; Shevyrevites Teichert & Kummel in Teichert et al., 1973; Xenodiscus Waagen, 1879., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 18, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Teichert C., Kummel B. & Sweet W. C. 1973. Permian - Triassic strata, Kuh-e-Ali Bashi, Northwestern Iran. Bulletin of the Museum of Comparative Zoology, Harvard University 145 (8): 359 - 472.","Yang F. 1987. Late Late Permian strata and their ammonoid zones in Southwest China. Earth Science - Journal of Wuhan College of Geology 1985 (10): 129 - 144.","Liang X. 1983. New material of Permian ammonoids with discussion on the origin, migration of Araxoceratidae and the horizon of the Paratirolites. Acta Palaeontologica Sinica 22 (6): 606 - 615.","Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182."]}
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- 2021
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19. Pseudogastrioceras relicuum Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Pseudogastrioceras ,Animalia ,Pseudogastrioceras relicuum ,Biodiversity ,Paragastrioceratidae ,Taxonomy - Abstract
Pseudogastrioceras relicuum Korn & Ghaderi in Korn et al., 2016 Fig. 16; Table 1 Pseudogastrioceras relicuum Korn & Ghaderi in Korn et al., 2016: 852, text-fig. 12. Diagnosis Pseudogastrioceras with moderately large conch; diameter attaining 70 mm. Conch shape thinly discoidal (ww/dm=0.40–0.45), involute (uw/dm = 0.05–0.10) with converging flanks and broadly rounded venter. Ornament with about 15 faint spiral lines on the venter. Material examined Holotype IRAN • East Azerbaijan, Ali Bashi N section; Paratirolites Limestone; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 12); MB.C.25173. Additional material IRAN • 2 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29659 to MB.C.29660 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29661 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29662 to MB.C.29663. Description All specimens are rather poorly preserved, but MB.C.29661 is the best of them, though somewhat deformed (Fig. 16). It is a corroded specimen with a 56 mm conch diameter, which does not allow accurate measurements of the conch. Stratigraphic range Paratirolites Limestone; 6.80 to 3.30 m below the extinction horizon (Dzhulfites nodosus Zone to Paratirolites lanceolobatus Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 16-17, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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- 2021
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20. Lutites plicatus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. et sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Lutites plicatus ,Lutites ,Taxonomy - Abstract
Lutites plicatus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 1713B70A-3C3C-4FCD-9827-BBBA6F989797 Fig. 55; Table 29 Diagnosis Species of Lutites gen. nov. with conch reaching 120 mm dm. Subadult stage with oval, compressed whorl profile (ww/wh=0.75) and rounded venter; about 12 radial plications per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.72), weakly concave flanks, flattened venter and subangular ventrolateral shoulder; numerous weak and densely arranged plications and small ventrolateral nodes. Prongs of external lobe multiply serrated; altogether 20 notches of E, A and L lobes. Etymology From the Latin ‘ plicatus ’, meaning ‘corrugated’, referring to the dense ribbing. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 55A; MB.C.30018. Description Holotype MB.C.30018 is a specimen with 84 mm diameter, of which the last 120 degrees belong to the body chamber (Fig. 55A). Nearly one volution of the phragmocone can be studied; at this stage the whorl profile is oval and compressed with a rounded venter. The ornament consists of nearly 20 faint radial plications, which are coarsest near the umbilicus and in the ventrolateral area, where they form weak nodes. The ornament is weaker on the body chamber, where the plications are more numerous. The whorl profile is subtrapezoidal and compressed (ww/wh=0.72) in the adult stage with a rounded umbilical margin, weakly concave flanks and a flattened venter. The suture line of the holotype has a narrow external lobe with asymmetric, multiply serrated prongs (four and six small notches), an asymmetric and inflated ventrolateral saddle and multiply serrated adventive and lateral lobes (Fig. 55B). Both lobes show notches of various size. Remarks Lutites plicatus gen. et sp. nov. differs in the shape of the flattened, but convex venter from most of the other species of the genus (L. lyriformis gen. et sp. nov., L. profundus gen. et sp. nov., L. alius gen. et sp. nov.), which possess a completely flat or even concave venter. Stratigraphic range Upper part of the Hambast Formation; horizon unknown., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 85-87, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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- 2021
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21. Esfahanites armatus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. et sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Esfahanites ,Cephalopoda ,Mollusca ,Ammonoidea ,Esfahanites armatus ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Esfahanites armatus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: DACEC5B2-4BE5-40D0-AA2B-578741DC349C Figs 45–46; Table 22 ? Paratirolites sp. Ghaedi et al., 2009: pl. 2 figs 1, 4. ? Paratirolites waageni – Zakharov & Mussavi Abnavi 2013: text-fig. 12c. Diagnosis Species of Esfahanites gen. nov. with conch reaching 130 mm dm. Subadult stage with strongly trapezoidal, moderately depressed whorl profile (ww/wh =1.50–1.70) and rounded to tectiform venter; about 12 coarse ventrolateral nodes per volution. Adult stage with trapezoidal and moderately depressed whorl profile (ww/wh =1.50–1.70), flattened tectiform venter and angular ventrolateral shoulder; coarse rursiradiate ribs. Prongs of external lobe trifid or multiply serrated; altogether 15–21 notches of E, A and L lobes. Etymology From the Latin ‘ armatus ’, meaning ‘armed’, because of the coarse spiny sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 45; MB.C.29964. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29963 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29965 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29966 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29967 to MB.C.29968. Description Holotype MB.C.29964 is the best-preserved specimen; it has 106 mm conch diameter and allows the study of two whorls (Fig. 45A). The diameter of the phragmocone is 68 mm and the body chamber has a length of about 270 degrees. Throughout the last two preserved volutions, the whorl profile is trapezoidal, but with the flanks less strongly diverging at the end of growth. The venter is broadly rounded in the penultimate whorl and becomes more flattened on the terminal body chamber. The sculpture on the phragmocone has 14 coarse conical ventrolateral nodes on the last volution; these nodes show a rib-like elongation towards the umbilicus. The sculpture on the body chamber shows variably coarse, rounded ribs on the flank; they turn backward near the umbilicus and are strengthened in the ventrolateral area. The suture line of holotype MB.C.29964 is remarkable because of its very small external lobe, which is wider at the base and is constricted in the upper part (Fig. 45B). The prongs are inclined ventrally and possess five little notches each. The ventrolateral saddle is small and inflated. The asymmetric adventive lobe is strongly serrated with little notches being developed on the ventral side up to half of the lobe depth. The lateral lobe is slightly less strongly serrated than the adventive lobe. Paratype MB.C.29966 is an incomplete but otherwise rather well-preserved individual of 101 mm conch diameter; it allows the study of two and a half whorls (Fig. 46A). The body chamber begins at 60 mm conch diameter, meaning that the preserved part of the body chamber has a length of 270 degrees and is nearly complete. Conch shape changes can be observed during the last whorl of the specimen. While the whorl profile is circular with rounded venter at the binning of the last whorl, it changes to tectiform and has, at the end of growth, an almost flat venter, separated from the flat and diverging flanks by an angular ventrolateral shoulder. The sculpture of the phragmocone part of the specimen shows coarse conical nodes in the ventrolateral area, 12 on the last volution and 10 on the penultimate volution. At the beginning of the terminal body chamber, these nodes are transformed into elongate nodes and finally rounded ribs, which turn from the umbilicus in backward direction. Finally, at the end of the body chamber these ribs become weaker and only shallow rounded ribs and weak ventrolateral nodes can be seen. The suture line of paratype MB.C.29966 shows a very small external lobe that has not half the depth of the adventive lobe (Fig. 46B). It is parallel-sided with parallel-sided prongs that possess three small notches. The ventrolateral saddle is inverted U-shaped with a much longer dorsal flank. The adventive lobe has seven small notches and the lateral lobe five, both lobes are asymmetric and separated by a low semi-circular lateral saddle. Remarks Esfahanites armatus gen. et sp. nov. differs in the very small external lobe from all the species of Paratirolites. It can therefore not be confused with any other species. Stratigraphic range Upper part of the Hambast Formation; the only in situ collected specimen comes from 2.00 m below the extinction horizon (Alibashites ferdowsii Zone).
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22. Pseudogastrioceras Spath 1930
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Pseudogastrioceras ,Animalia ,Biodiversity ,Paragastrioceratidae ,Taxonomy - Abstract
Genus Pseudogastrioceras Spath, 1930 For the composition of the genus, see Korn & Ghaderi in Korn et al. (2016)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 16, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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23. Stoyanowites aspinosus Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Stoyanowites ,Dzhulfitidae ,Animalia ,Biodiversity ,Stoyanowites aspinosus ,Taxonomy - Abstract
Stoyanowites aspinosus Korn & Ghaderi in Korn et al., 2016 Fig. 62; Table 36 Stoyanowites aspinosus Korn & Ghaderi in Korn et al., 2016: 885, text-fig. 43. Diagnosis Species of Stoyanowites with conch reaching 100 mm dm. Subadult stage with oval, weakly compressed whorl profile (ww/wh =0.70) and rounded venter; 15 narrow and rounded ribs on the flanks. Adult stage with oval and weakly compressed whorl profile (ww/wh = 0.60–0.70) and moderately wide to wide umbilicus (uw/dm =0.40–0.50), rounded venter and rounded ventrolateral shoulder; very weak ribs on the flanks, coarsest in ventrolateral area. Prongs of external lobe bifid; 8–9 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone, 0.95 m below the top; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 43a); MB.C.25458. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.300978 • 2 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.30098 to MB.C.30099 • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30100 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30101. Description Specimen MB.C.30100 is a fragmentary specimen with 72 mm diameter and shows the wide umbilicus (uw/dm =0.50), the nearly parallel-sided whorl profile and the rounded venter. The ornament of the last half volution consists of numerous (about 25) rather sharp ribles, which extend with a shallow sinus across the flanks and turn forward in the ventrolateral area, where they become coarser and for a hooklike rib (Fig. 62A). The venter is smooth. Remarks Stoyanowites aspinosus differs from S. dieneri in the absence of dorsolateral nodes and the presence of densely spaced curved riblets on the body chamber. Stoyanowites parallelus sp. nov. differs in the flat venter. Stratigraphic range Upper part of the Hambast Formation; 1.80 to 1.60 m below the extinction horizon (Alibashites ferdowsii Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 100-101, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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24. Lutites lyriformis Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. et sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Lutites lyriformis ,Animalia ,Biodiversity ,Lutites ,Taxonomy - Abstract
Lutites lyriformis Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 3FC3DB4D-AA14-43A6-B982-0137ED22D6B7 Figs 51–52; Table 26 Diagnosis Species of Lutites gen. nov. with conch reaching 140 mm dm. Subadult stage with parallel-sided, compressed whorl profile (ww/wh = 0.60–0.70) and rounded venter; about 14 ventrolateral nodes per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.50–0.60), concave flanks, concave venter and angular ventrolateral shoulder; numerous weak and densely arranged ribs. Prongs of external lobe bifid to multiply serrated; altogether 13–22 notches of E, A and L lobes. Etymology From the Latin ‘ lyra ’, referring to the shape of the whorl profile. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain A section; Hambast Formation; illustrated in Fig. 51A; MB.C.29995. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29994 • 2 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29996 to MB.C.29997 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29998 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29999 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30000 to MB.C.30001. Description Holotype MB.C.29995 is a rather well-preserved specimen with 96 mm conch diameter (Fig. 51A). It allows the study of three and a half volutions, of which only one third of a volution belongs to the body chamber. Adulthood is indicated by the weak crowding of the last septa at 73 mm diameter. The whorl profile is, at the largest preserved conch diameter, subtrapezoidal with a broadly rounded umbilical margin and concave and converging flanks. The weakly concave venter is separated from the flanks by an angular ventrolateral shoulder. Half a volution earlier, the venter is plane and another half a volution back, the venter is broadly rounded. The sculpture shows significant ontogenetic changes from the subadult stage with 13 coarse and blunt ventrolateral nodes, which originate from weak and rounded radial ribs, per volution. They become weaker and more numerous in the last quarter volution of the phragmocone (about 12 weak ventrolateral nodes within this interval). The body chamber possesses a very weak sculpture consisting only of weak and rounded radial riblets in the inner flank area. The suture line of the holotype possesses a parallel-sided external lobe with prongs being asymmetrically serrated by four and five small notches, respectively. The asymmetric ventrolateral saddle is nearly as wide as high; it is followed by an adventive lobe with oblique base serrated into seven small notches. The weakly asymmetric, inverted U-shaped lateral saddle is nearly as high as the ventrolateral saddle. Finally, the asymmetric lateral lobe has also seven small notches (Fig. 52A). The smaller paratype MB.C.29997 has 74 mm conch diameter, of which the phragmocone has a diameter of 64 mm and one quarter of a whorl belongs to the body chamber (Fig. 51B). The whorl profile is subtrapezoidal with concave, converging flanks and the venter is weakly concave. The sculpture generally resembles the holotype, but the ribs and nodes are weaker. The same is true for the additional paratypes MB.C.29996 (Fig. 51C) and MB.C.29999 (Fig. 51D), which is the conch geometry are nearly identical with the holotype. The suture line of paratype MB.C.29997 shows a parallel-sided external lobe that is deeper than the adventive lobe. The prongs are bifid and parallel-sided. All lobes and thus saddles are parallel-sided and all saddles have the same height. The adventive lobe and the lateral lobe possess six and four small notches (Fig. 52B). Remarks The new species differs in the coarse ventrolateral nodes of the subadult stage from most of the other species of Lutites gen. nov. with compressed whorl profile. Lutites lyriformis gen. et sp. nov. is similar to L. profundus gen. et sp. nov. but differs in the much weaker sculpture of the terminal body chamber, which shows only very weak ribs in L. lyriformis gen. et sp. nov. but ventrolateral nodes in L. profundus gen. et sp. nov. Stratigraphic range Upper part of the Hambast Formation; 1.45 to 1.00 m below the extinction horizon (Lutites profundus Zone to Abichites abichi Zone).
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25. Paratirolites coronatus Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Paratirolites coronatus ,Taxonomy - Abstract
Paratirolites coronatus Korn & Ghaderi in Korn et al., 2016 Fig. 31; Table 14 Paratirolites coronatus Korn & Ghaderi in Korn et al., 2016: 860, text-figs 17–18. Diagnosis Species of Paratirolites with conch reaching 125 mm dm. Subadult stage with trapezoidal, moderately depressed whorl profile (ww/wh =1.60–1.90) and flattened tectiform venter; 15 coarse conical ventrolateral nodes per volution. Adult stage with extremely trapezoidal and moderately depressed whorl profile (ww/wh =1.60–1.90), flattened tectiform venter and subangular ventrolateral shoulder; weak ventrolateral nodes. Prongs of external lobe trifid or multiply serrated; altogether 11–24 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone, 2.90 m below the top; illustrated by Korn et al. (2016: text-fig. 17A); MB.C.25095. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29842. Description Specimen MB.C.29842 is a fully chambered specimen with 55 mm conch diameter and allows the study of one and a half whorls (Fig. 31A). It has a strongly trapezoidal, depressed whorl profile (ww/wh =1.75), weakly sinuous flanks, an angular ventrolateral shoulder and a flattened tectiform venter. Half a volution earlier, the venter is weakly flattened but still rounded and another half volution back, the venter is broadly rounded. The sculpture changes during the last volution of the phragmocone; strong conical nodes at the beginning of this volution are transformed into smaller, longitudinally elongated nodes, which have a position at the ventrolateral shoulder. The suture line of specimen MB.C.29842 has a V-shaped external lobe with concave flanks and symmetric, narrowly V-shaped simple prongs. The ventrolateral saddle is asymmetric and dorsally inclined and the asymmetric, nearly parallel-sided adventive lobe has nine little notches. The lateral lobe is nearly flat at its base and subdivided in small equally sized notches (Fig. 31B). Remarks Paratirolites coronatus is the paratirolitid with the widest conch, resulting from the coronate ventrolateral shoulder. Therefore, it cannot be confused with any other species. Paratirolites coronatus differs from P. vediensis and the other species in the more strongly trapezoidal whorl profile of the adult stage. Stratigraphic range Upper part of the Hambast Formation; precise horizon at Baghuk Mountain not known.
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26. Alibashites Korn & Ghaderi
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Alibashites ,Taxonomy - Abstract
Genus Alibashites Korn & Ghaderi in Korn et al., 2016 Type species Xenodiscus (Paratirolites ?) Mojsisovicsi Stoyanow, 1910, by original designation. Included species Alibashites ferdowsii Korn & Ghaderi in Korn et al., 2016; Xenodiscus (Paratirolites?) Mojsisovicsi Stoyanow, 1910; Alibashites uncinatus Korn & Ghaderi in Korn et al., 2016. Diagnosis Representatives of the family Dzhulfitidae with moderately large to large conch; maximum adult diameters 90 mm. Adult stage with subtrapezoidal or quadrate whorl profile. Subadult stage with large conical ventrolateral nodes, adult stage with weak ribs. Suture line with deep external lobe; depths of external lobe and adventive lobe nearly identical. Remarks Alibashites has an intermediate morphological position between Paratirolites and Abichites. It possesses the subadult sculpture with conical ventrolateral nodes of Paratirolites but lacks the trapezoidal whorl profile of that genus. Abichites has a similar adult conch morphology with a rectangular or subtrapezoidal whorl profile but does not possess the strong ventrolateral nodes developed in Alibashites., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 71, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135."]}
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27. Dzhulfites Shevyrev 1965
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Dzhulfites ,Animalia ,Biodiversity ,Taxonomy - Abstract
Genus Dzhulfites Shevyrev, 1965 Type species Dzhulfites spinosus Shevyrev, 1965, by original designation. Included species Dzhulfites hebes Korn & Ghaderi in Korn et al. (2016); Dzhulfites nodosus Shevyrev, 1965; Dzhulfites spinosus Shevyrev, 1965; Dzhulfites zalensis Korn & Ghaderi in Korn et al. (2016); Dzhulfites brevisellatus Korn & Hairapetian sp. nov. Diagnosis Representatives of the family Dzhulfitidae with moderately large to large conch; maximum adult diameters are between 80 and 160 mm. Subadult stage with trapezoidal whorl profile, adult stage variable. Subadult stage with small to large conical ventrolateral nodes, adult stage with weakening sculpture. Suture line with external lobe that does not reach the depth of the adventive lobe; prongs of external lobe simple to trifid. Remarks For a discussion of the genus, see Korn & Ghaderi in Korn et al. (2016)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 27, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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28. Paratirolites aduncus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Paratirolites aduncus ,Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Taxonomy - Abstract
Paratirolites aduncus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 86D671DC-3110-45AE-A379-2543EA8DEDD0 Figs 35–37; Table 17 Diagnosis Species of Paratirolites with conch reaching 125 mm dm. Subadult stage with circular or weakly depressed whorl profile (ww/wh = 1.00–1.30) and rounded venter; about 10 very coarse conical ventrolateral nodes per volution. Adult stage with weakly trapezoidal and weakly compressed whorl profile (ww/wh =0.75–0.90), flattened tectiform venter and subangular ventrolateral shoulder; about 10 hook-shaped ventrolateral nodes per half volution. Prongs of external lobe usually bifid but rarely trifid or multiply serrated; altogether 11–17 notches of E, A and L lobes. Etymology From the Latin ‘ aduncus ’, meaning ‘incurved’, because of the curved ventrolateral nodes. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 35A; MB.C.29894. Paratypes IRAN • 9 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29880 to MB.C.29888 • 5 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29889 to MB.C.29893 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29895 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29896 to MB.C.29900 • 17 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29901 to MB.C.29917 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29918 • 8 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29919 to MB.C.29926 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29927 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29928 to MB.C.29930. Description Holotype MB.C.29894 (Fig. 35A) is a fairly well-preserved specimen with 80 mm conch diameter. It shows, at a conch diameter of 60 mm, three closely approximated septa, after which the terminal body chamber begins with a sudden weakening of the ventrolateral nodes. While these nodes are coarse and conical on the phragmocone, they almost spontaneously become elongated and hook-shaped on the body chamber. The suture line of holotype MB.C.29894 shows a deep, parallel-sided external lobe with prongs multiply serrated asymmetrically (Fig. 35C). The ventrolateral saddle is nearly symmetric and inverted U-shaped, it is followed by an adventive lobe and a lateral lobe with six little notches each. Both lobes are semicircular at the base and slightly asymmetric. Paratype MB.C.29880 has 83 mm conch diameter; half of the last volution belongs to the body chamber (Fig. 35B). It largely agrees, in conch shape and sculpture, with the holotype. The specimen allows for the study of two whorls, on which transformations in conch shape and sculpture can be observed. The phragmocone has a nearly circular whorl profile (ww/wh about 1.15 at 60 mm conch diameter) with nine coarse conical ventrolateral nodes on its last volution, which sometimes are developed as sharp spines. On the body chamber, the transformation into a weakly trapezoidal whorl profile can be observed; here the slightly concave flanks diverge towards the subangular ventrolateral shoulder that separates the flank from the flattened, weakly tectiform venter. On the body chamber occurs also a change in the sculpture towards more numerous (about 10 per half whorl) short ribs, which are restricted to the outer flank and ventrally end in hook-shaped, sharp nodes. Paratype MB.C.29900 is a specimen of 104 mm conch diameter. It has a fairly well-preserved phragmocone but a weathered body chamber, of which a little more than half a volution is preserved (Fig. 36A). The shape of the body chamber whorl profile had to be reconstructed; it shows a trapezoidal shape with weakly diverging flanks, a rounded ventrolateral shoulder and a flattened tectiform venter. One volution earlier, the whorl profile is strongly trapezoidal with flat, diverging flanks and a flattened, broadly rounded venter. The sculpture is coarse on the phragmocone and weakens in strength significantly on the body chamber. The last volution of the phragmocone has 15 conical, spiny ventrolateral nodes, which are first very coarse and conical but later elongate, weaker and more closely spaced. On the body chamber, weak and rounded ribs with concave course are visible on the outer flank. The suture line of paratype MB.C.29900 shows a parallel-sided external lobe with parallel-sided prongs that are divided by four small notches. The very narrow ventrolateral saddle is weakly inclined towards the umbilicus. Following is a very asymmetric, weakly pouched adventive lobe with eight notches, which are already climbing up the ventral side. The weakly inflated, asymmetric lateral saddle is low and the asymmetric lateral lobe with steep dorsal flank possesses five notches (Fig. 36B). Remarks Paratirolites aduncus sp. nov. differs from the other species of the genus in the presence of the hookshaped ventrolateral nodes of the adult stage. Stratigraphic range Upper part of the Hambast Formation; 2.65 to 1.15 m below the extinction horizon (Paratirolites kittli Zone to Lutites profundus Zone).
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29. Arasella falcata Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Xenodiscidae ,Arasella ,Animalia ,Arasella falcata ,Biodiversity ,Taxonomy - Abstract
Arasella falcata Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: B2DA940A-842C-452B-8906-A29E07291FFE Fig. 21; Table 6 Diagnosis Species of Arasella with conch reaching 35 mm dm. Subadult and adult stage with compressed whorl profile (ww/wh=0.60–0.70) and rounded venter; 25 curved ribs which are coarsest on the inner flanks. Etymology From the Latin ‘ falx ’, meaning ‘sickle’, because of the rib course. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 0.05 m below top; illustrated in Fig. 21A; MB.C.29734. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29733 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29735 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29736 • 1 specimen; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29737. Description Holotype MB.C.29734 is a somewhat corroded specimen with a conch diameter of 24 mm (Fig. 21A). It is extremely discoidal (ww/dm =0.20) and subevolute (uw/dm =0.43) with a compressed whorl profile (ww/wh =0.63). The flanks are nearly parallel and the venter is rounded. The last volution shows about 25 ribs, which are sharpest and coarsest on the inner flank; they turn slightly forward and become weaker on the outer flank to diminish. Remarks Arasella minuta shows much coarser ribs, particularly on the outer flanks, while A. falcata sp. nov. possesses a much weaker sculpture with ribs strongest in in dorsolateral area. Stratigraphic range Topmost part of the Hambast Formation; immediately below the extinction horizon (Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 26-27, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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30. Shevyrevites corrugatus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Shevyrevites ,Ammonoidea ,Xenodiscidae ,Animalia ,Biodiversity ,Shevyrevites corrugatus ,Taxonomy - Abstract
Shevyrevites corrugatus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 89B3129F-6478-4C46-9CD6-DDCD7C735BF0 Fig. 19; Table 4 Diagnosis Species of Shevyrevites with conch reaching 60 mm dm. Subadult stage with oval, slightly compressed whorl profile (ww/wh = 0.60–0.75) and rounded venter; multiple fine ribs. Adult stage with parallelsided, slightly compressed whorl profile (ww/wh =0.60–0.75), rounded venter and rounded ventrolateral shoulder; multiple fine riblets on the flank. Prongs of external lobe bifid; altogether 11 notches of E, A and L lobes. Etymology From the Latin ‘ corrugatus ’, meaning ‘folded’, because of the corrugate sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain A section; Hambast Formation; illustrated in Fig. 19B; MB.C.29718. Paratype IRAN • 1 specimen; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29719; illustrated in Fig. 19A. Description Holotype MB.C.29718 has 37 mm conch diameter and shows half of a volution preserved that belongs to the terminal body chamber (Fig. 19B); the maximum phragmocone diameter is 27 mm. The phragmocone shows rather densely spaced septa (18 per volution), but the last four are then strikingly crowded and indicate adulthood. The conch is extremely discoidal (ww/dm=0.20) and subevolute (uw/dm =0.43) with applanate flanks and a rounded venter. The sculpture consists of numerous densely spaced and sharp radial riblets (about 30 on half a volution in the adult stage), which on the body chamber extend with a shallow concave sinus across the flanks. The suture line of the holotype has a nearly parallel-sided, weakly pouched external lobe that is almost as deep as the adventive lobe; it has asymmetric prongs that are subdivided by two notches. The slightly asymmetric ventrolateral saddle is inverted U-shaped and followed by the asymmetric adventive lobe with five small notches, an asymmetric lateral saddle and an asymmetric lateral lobe with four notches (Fig. 19C). Remarks The new species differs from the other species of Shevyrevites by the much weaker sculpture, which consists of feeble riblets in contrast to the coarser ribs in S. shevyrevi and the conical nodes in S. nodosus. Stratigraphic range Upper part of the Hambast Formation; float but probably 4.80 m below the extinction horizon (red nodular limestone in the Shevyrevites shevyrevi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 22-23, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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31. Shevyrevites Teichert & Kummel 1973
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Shevyrevites ,Ammonoidea ,Xenodiscidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Genus Shevyrevites Teichert & Kummel in Teichert et al., 1973 Type species Shevyrevites shevyrevi Teichert & Kummel in Teichert et al., 1973, by original designation. Diagnosis Representatives of the family Xenodiscidae with a moderately large to large conch; maximum adult diameter between 60 and 80 mm. Subadult stage with a trapezoidal whorl profile, adult stage variable. Subadult stage with small to large conical ventrolateral nodes, adult stage with a weakening sculpture with predominant ribs. Suture line with external lobe that does not reach the depth of the adventive lobe; prongs of the external lobe simple or bifid. Included species Shevyrevites shevyrevi Teichert & Kummel in Teichert et al., 1973; Bernhardites nodosus Shevyrev, 1965; Shevyrevites corrugatus Korn & Hairapetian sp. nov. Remarks The species of Shevyrevites show a coarse ornament consisting of lateral ribs and sometimes also coarse conical nodes in the inner whorls, and are hereby clearly separated from the other genera of the family. Regarding the presence of ventrolateral nodes, there are some similarities to the genera of the family Dzhulfitidae, but these differ in the shape of the suture line: while the external lobe is wide in Shevyrevites, it is always narrow in the Dzhulfitidae. Teichert & Kummel (in Teichert et al. 1973) caused some nomenclatorial confusion with their opinion on the species of their new genus Shevyrevites. They made clear that “ Celtites radiosus ”, the Triassic type species of Bernhardites Shevyrev, 1965, differs from the Late Permian forms and that for the latter, a new genus name (Shevyrevites) had to be introduced. At the same time, they treated the two species “ Bernhardites radiosus (Frech in Noetling, 1905)” and “ Bernhardites nodosus Shevyrev, 1965 ”, both of which were separated by Shevyrev (1965), as synonyms; they described these under the new genus and species name Shevyrevites shevyrevi. This procedure is not accepTable because it suppresses the species name Shevyrevites nodosus, which in case of synonymy of the two species has priority over Shevyrevites shevyrevi. However, this problem is regarded here minor, as we accept both species as valid., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 18, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Teichert C., Kummel B. & Sweet W. C. 1973. Permian - Triassic strata, Kuh-e-Ali Bashi, Northwestern Iran. Bulletin of the Museum of Comparative Zoology, Harvard University 145 (8): 359 - 472.","Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182."]}
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32. Paratirolites trapezoidalis Shevyrev 1965
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Paratirolites trapezoidalis ,Taxonomy - Abstract
Paratirolites trapezoidalis Shevyrev, 1965 Fig. 32; Table 15 Paratirolites trapezoidalis Shevyrev, 1965: 177, pl. 24 fig. 1. Paratirolites trapezoidalis – Shevyrev 1968: 92, pl. 4 fig. 1. — Korn in Ghaderi et al. 2014: text-fig. 7e. — Korn & Ghaderi in Korn et al. 2016: 858, text-fig. 16. Diagnosis Species of Paratirolites with conch reaching 120 mm dm. Subadult stage with trapezoidal, depressed whorl profile (ww/wh= 1.40–1.60) and flattened, slightly tectiform venter; 10–15 coarse ribs, which end in coarse conical ventrolateral nodes, per volution. Adult stage with strongly trapezoidal and weakly depressed whorl profile (ww/wh =1.00–1.40), flattened venter and angular ventrolateral shoulder; weak straight riblets and numerous small ventrolateral nodes. Prongs of external lobe usually unsubdivided, rarely trifid; altogether 12–19 notches of E, A and L lobes. Type material Holotype AZERBAIJAN • Nakhichevan Province, Dorasham 2 section; Paratirolites Limestone; illustrated by Shevyrev (1965: pl. 24 fig. 1); PIN 1252/129. Material examined IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29843 • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29844 to MB.C.29845 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29846 to MB.C.29848. Description Specimen MB.C.29844 is a moderately well-preserved individual of 71 mm conch diameter and allows the study of one and a half whorls (Fig. 32B). The last segment of 210 degrees belongs to the body chamber, which has a weakly depressed trapezoidal whorl profile (ww/wh =1.05) with weakly concave flanks and a nearly flat venter and an angular ventrolateral shoulder. The sculpture of the subadult stage consists of conical and coarse, spiny ventrolateral nodes, 13 on the last phragmocone volution. The nodes become significantly weaker and more numerous on the body chamber, where about 16 of them are present on half a volution. Specimen MB.C.29844 allows only the study of the suture line on the venter. The external lobe is narrow and parallel-sided with non-serrated, narrow lanceolate prongs. The ventrolateral saddle is very asymmetric and dorsally inclined and the asymmetric adventive lobe has an oblique base that is serrated by six small notches (Fig. 32C). specimen MB.C.29845 (Fig. 32A) is similar in conch shape and sculpture but possesses a suture line with trifid prongs of the external lobe. Remarks Paratirolites vediensis has a similar conch shape, but differs from P. trapezoidalis in the stronger ventrolateral nodes in the adult stage. P. coronatus has an even wider whorl profile (ww/wh = 1.60–1.80 in P. coronatus but only 1.40 in P. trapezoidalis); the species has also less coarse but more numerous ventrolateral nodes in the subadult stage (15 in P. coronatus but only 10 in P. trapezoidalis). Stratigraphic range Upper part of the Hambast Formation; 3.05 to 2.80 m below the extinction horizon (Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 47-49, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Shevyrev A. A. 1968. Triasovye ammonoidei Yuga SSSR. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 119: 1 - 272.","Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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33. Abichites ovalis Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Abichites ,Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Abichites ovalis ,Taxonomy - Abstract
Abichites ovalis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 983F405A-033F-49EB-8D75-E5F917F23D86 Fig. 58; Table 32 Diagnosis Species of Abichites with conch reaching 85 mm dm. Subadult stage with subcircular, weakly compressed whorl profile (ww/wh =0.80–1.00) and rounded venter; about 12 shallow and rounded radial ribs. Adult stage with parallel sided or weakly trapezoidal, weakly compressed whorl profile (ww/wh =0.80–1.00), rounded venter and narrowly rounded ventrolateral shoulder; shallow wide ribs on the flank. Prongs of external lobe simple; altogether 7–11 notches of E, A and L lobes. Etymology From the Latin ‘ ovalis ’, meaning ‘oval’, because of the oval whorl profile in the subadult stage. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 58A; MB.C.30059. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30060 • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30061 to MB.C.30063 • 6 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30064 to MB.C.30069 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30070 to MB.C.30071 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30072. Description Holotype MB.C.30059 is a rather well-preserved specimen with 62 mm conch diameter; the phragmocone diameter is 44 mm and 190 degrees of the body chamber are preserved (Fig. 58A). The adult whorl profile is weakly trapezoidal with gently diverging, slightly concave flanks, a rounded ventrolateral shoulder and a slightly flattened but broadly rounded venter. One volution earlier, the whorl profile is subcircular and weakly compressed (ww/wh = 0.90). The sculpture on the phragmocone consists of 12 shallow and rounded radial ribs per volution. They become weaker on the body chamber but develop to elongate, rather sharp nodes on the outer flank. The suture line of holotype MB.C.30059 shows an external lobe with flanks standing parallel in the lower part and diverging in the upper part. The prongs are very narrow, lanceolate and not secondarily serrated. On the outer flank lies the symmetric adventive lobe with semicircular, strongly serrated base; it is followed by a rather high lateral saddle and a parallel-sided lateral lobe with five small notches (Fig. 58C). Paratype MB.C.30071 has 60 mm conch diameter and is, with its weakly trapezoidal whorl profile and the general conch shape very similar to the previous one but has a more slender conch (ww/wh =0.85). It shows, in the inner whorls, weak ventrolateral nodes connected to the shallow radial ribs (Fig. 58B). Remarks Abichites ovalis sp. nov. closely resembles A. paucinodum, but differs in the whorl profile (diverging flanks in A. ovalis sp. nov. but converging in A. paucinodum), the wider umbilicus in A. ovalis sp. nov. and in the rounded venter in A. ovalis sp. nov. (flat in A. paucinodum). Stratigraphic range Upper part of the Hambast Formation; 0.75 to 0.40 m below the extinction horizon (Abichites stoyanowi Zone to lowest part of the Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 92-93, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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34. Abichites stoyanowi
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Abichites ,Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy ,Abichites stoyanowi - Abstract
Abichites stoyanowi (Kiparisova in Voinova et al., 1947) Fig. 59; Table 33 Kashmirites ? stoyanowi Kiparisova in Voinova et al., 1947: 149, pl. 35 fig. 1. Xenodiscus radians – Stoyanow 1910: 86, pl. 9 fig. 5. Xenodiscus sp. indet. Stoyanow 1910: 87, pl. 9 fig. 6. Abichites stoyanowi – Shevyrev 1965: 179, pl. 24 figs 2–3; 1968: 94, pl. 3 fig. 5, pl. 4 fig. 2. — Korn in Ghaderi et al. 2014: text-fig. 7h. — Korn & Ghaderi in Korn et al. 2016: 881, text-fig. 38. Diagnosis Species of Abichites with conch reaching 70 mm dm. Subadult stage with circular, weakly compressed whorl profile (ww/wh =0.75–1.00) and rounded venter; 16–20 moderately weak ribs per volution. Adult stage with quadrate and weakly compressed whorl profile (ww/wh =0.90–1.00), flat venter and subangular to angular ventrolateral shoulder; altogether 12–14 sharp concavo-convex ribs per half volution on flanks. Prongs of external lobe simple or bifid; 6–11 notches of E, A and L lobes. Material examined IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30073 • 2 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30074 to MB.C.30075 • 3 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30076 to MB.C.30078 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.30079 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30080 to MB.C.30081 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30082. Description Specimen MB.C.30078 has 62 mm conch diameter and is rather well-preserved with the last half volution belonging to the body chamber (Fig. 59A). This has a subtrapezoidal, weakly compressed whorl profile (ww/wh =0.90) with a broadly rounded umbilical margin, weakly concave converging flanks, an angular ventrolateral shoulder and a weakly concave venter. Two stages of the ornament are visible; the subadult stage has coarse node-like ribs on the flank, which in the adult stage become increasingly weaker and more numerous (15 per half volution at the beginning of the body chamber). The suture line of specimen MB.C.30078 has a large external lobe with weakly diverging flanks and unsubdivided lanceolate prongs. The inverted U-shaped ventrolateral saddle is dorsally inclined and is followed by the pouched adventive lobe that has a very oblique base subdivided into three very shallow blunt notches. In contrast, the three notches of the parallel-sided lateral lobe are pointed denticles (Fig. 59E). The two smaller specimens MB.C.30074 (31 mm dm; Fig. 59D) and MB.C.30081 (34 mm dm; Fig. 59B) have similar conch shapes and proportions; they differ in the somewhat weaker sculpture. Remarks Abichites stoyanowi differs from A. abichi, which is another similar species with moderately coarse sculpture, in the much weaker ribs. Abichites subtrapezoidalis has a similar whorl profile, but only very weak riblets around the umbilicus on the adult body chamber. Stratigraphic range Upper part of the Hambast Formation; 0.80 to 0.05 m below the extinction horizon (Abichites stoyanowi Zone to lowest part of the Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 94, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Voinova E. V., Kiparisova L. D. & Robinson V. H. 1947. Klass Cephalopoda. Golovonogie., Atlas rukovodyashtchikh form iskopaemykh faun SSSR. VIII. Triasovaya sistema: 124 - 176, Moskva-Leningrad","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135.","Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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35. Paratirolites robustus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Paratirolites robustus ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Taxonomy - Abstract
Paratirolites robustus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 10C9CACF-9ED4-4E61-8140-FDBB70154F87 Figs 26–27; Table 11 Diagnosis Species of Paratirolites with conch reaching 120 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.30–1.50) and rounded venter; 8–12 coarse, radially elongated midflank nodes per volution. Adult stage with trapezoidal and weakly depressed whorl profile (ww/wh =1.00–1.10), flattened, tectiform venter and angular ventrolateral shoulder; numerous weak ribs on flank. Prongs of external lobe simple to trifid; altogether 12–17 notches of E, A and L lobes. Etymology From the Latin ‘ robustus ’, meaning ‘robust’, because of the coarse ribs. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation, 2.95 m below top; illustrated in Fig. 26A; MB.C.29778. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29776 to MB.C.29777 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29779 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29780 • 4 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29781 to MB.C.29784. Description Holotype MB.C.29778 has 80 mm conch diameter and consists of a rather well-preserved phragmocone and a more or less strongly weathered body chamber (Fig. 26A). Septal crowding indicates that the maximum phragmocone diameter is 48 mm. The conch shape changes during growth; there is a strongly trapezoidal whorl profile of the phragmocone and a weakly trapezoidal profile of the body chamber. The sculpture of the phragmocone shows 12 conical nodes, which are circular and symmetric in section in the penultimate whorl but become asymmetric and radially elongated towards the end of the phragmocone. The suture line of holotype MB.C.29778 has a Y-shaped external lobe, in which the prongs are weakly serrated including two notches. The adventive lobe is asymmetric and ventrally inclined with nearly parallel flanks and asymmetric notching. It is dorsolaterally followed by an asymmetric lateral saddle and a nearly symmetric lateral lobe with seven small notches (Fig. 27A). The slightly smaller paratype MB.C.29782 (73 mm dm) is very similar in conch shape and suture line. The penultimate whorl has eight coarse conical nodes, which transform, at the end of the phragmocone, into weakly curved elongated nodes (Fig. 26B). The suture line has an external lobe with weakly diverging flanks, an asymmetric ventrolateral saddle and a weakly asymmetric, strongly notched adventive lobe with nearly flat base and parallel flanks (Fig. 27B). Remarks Paratirolites robustus sp. nov. differs from the most similar of the Central Iranian species P. lanceolobatus sp. nov. in the stouter whorl profile (ww/wh = 1.20 in P. robustus sp. nov. at 70 mm conch diameter and only 0.85 in P. lanceolobatus). Furthermore, P. lanceolobatus sp. nov. has non-serrated prongs of the parallel-sided external lobe. Paratirolites robustus sp. nov. differs from P. vediensis in the rounded venter and from P. kittli in the radially elongated nodes of the subadult stage and the presence of weak radial ribs of the body chamber. Stratigraphic range Upper part of the Hambast Formation; 3.20 to 2.90 m below the extinction horizon (uppermost part of the Paratirolites lanceolobatus Zone to Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 36-40, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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36. Paratirolites quadratus Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Paratirolites quadratus ,Taxonomy - Abstract
Paratirolites quadratus Korn & Ghaderi in Korn et al., 2016 Figs 38–39; Table 18 Paratirolites quadratus Korn & Ghaderi in Korn et al., 2016: 866, text-fig. 25. Diagnosis Paratirolites with conch reaching 90 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.35–1.60) and broadly rounded venter; about 11–13 ribs ending in spiny ventrolateral nodes. Adult stage with weakly trapezoidal to nearly quadrate and weakly depressed whorl profile (ww/wh =1.00–1.20), weakly concave venter and angular ventrolateral shoulder; coarse and shallow ribs on flanks. Prongs of external lobe multiply serrated; altogether about 14–17 notches of E, A and L lobes. Material examined Holotype IRAN • 1 specimen; East Azerbaijan, Ali Bashi N section; Paratirolites Limestone, 1.70 m below the top; illustrated by Korn et al. (2016: text-fig. 25); MB.C.25296. Additional material IRAN • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29331 to MB.C.29333. Description Specimen MB.C.29932 is a nicely preserved specimen of 72 mm conch diameter (Fig. 38A). It shows weak septal crowding at 48 mm diameter and thus half of the last volution belongs to the body chamber. The whorl profile shape changes from trapezoidal with convex flanks of the subadult stage to parallelsided at the beginning of the body chamber; at the same time the shape of the venter changes from weakly convex to nearly totally flattened. The sculpture of the subadult stage possesses 13 conical ventrolateral nods per volution; they originate from rounded radial ribs. On the body chamber the nodes are weaker and become radially elongated, they are connected with low, rounded dorsolateral ribs. The suture line of specimen MB.C.29932 shows an asymmetric external lobe with weakly concave flanks, of which both prongs are trifid. Also on the venter follows the very narrow, weakly inflated ventrolateral saddle and the asymmetric adventive lobe (Fig. 38B). Specimen MB.C.29933 is a larger specimen with 101 mm conch diameter (Fig. 39A). It shows similar conch and sculpture characteristics but differs from the preceding specimen in the coarser ventrolateral nods and the larger phragmocone (62 mm dm). Its suture line has a parallel-sided external lobe with parallel-sided trifid prongs. The ventrolateral saddle is very narrow and the adventive lobe shows strong serration by small notches, which climb up the ventral flank to more than half of the lobe depth. The lateral saddle is weakly inflated and lower than the ventrolateral saddle and the lateral lobe is nearly parallel-sided and possesses five small notches (Fig. 39B). Remarks Paratirolites quadratus differs from the other species of Paratirolites in the slightly trapezoidal, nearly quadrate whorl profile in the adult conch and the very long, spiny ventrolateral nodes. Stratigraphic range At Baghuk Mountain only known from float from the upper part of the Hambast Formation; Stoyanowites dieneri Zone in the Julfa region.
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37. Clivotirolites Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Clivotirolites ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Genus Clivotirolites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 2F50EBFF-9384-40A5-84D9-573E68455DD2 Type species Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Representatives of the family Dzhulfitidae with large conch; maximum adult diameters are about 120 mm. Adult stage with trapezoidal whorl profile. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs. Suture line protracted on the flanks, with deep external lobe; depths of external lobe and adventive lobe nearly identical. Etymology From the Latin ‘ clivus ’, meaning ‘oblique’, because of the suture line and the similarity with Tirolites and Paratirolites. Included species Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov.; Clivotirolites petilus Korn & Hairapetian gen. et sp. nov. Remarks Clivotirolites gen. nov. differs from Paratirolites and related genera in the oblique protracting outline of the outer suture line., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 61-62, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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38. Abichites alibashiensis Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Abichites ,Cephalopoda ,Mollusca ,Ammonoidea ,Abichites alibashiensis ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Abichites alibashiensis Korn & Ghaderi in Korn et al., 2016 Fig. 57; Table 31 Abichites alibashiensis Korn & Ghaderi in Korn et al., 2016: 879, text-fig. 36. Diagnosis Species of Abichites with conch reaching 85 mm dm. Subadult stage with broadly oval, weakly depressed whorl profile (ww/wh =1.10–1.30) and rounded venter; 12–15 ventrolateral nodes per volution. Adult stage with quadrate of weakly subtrapezoidal, weakly compressed whorl profile (ww/wh =0.80–1.25), flattened venter and subangular to angular ventrolateral shoulder; sharp ribs. Prongs of external lobe simple or bifid; altogether 7–11 notches of E, A and L lobes. Material examined Holotype IRAN • East Azerbaijan, Ali Bashi N section; Paratirolites Limestone, 1.40 m below the top; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 36a); MB.C.25399. Additional material IRAN • 2 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30041 to MB.C.30042 • 3 specimens; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30043 to MB.C.30045 • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30046 to MB.C.30048 • 5 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30049 to MB.C.30053 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.30054 • 4 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30055 to MB.C.30058. Description Specimen MB.C.30051 has 62 mm conch diameter and is rather well preserved with about 190 degrees of the body chamber visible (Fig. 57A). The whorl profile of the terminal body chamber has a broadly rounded umbilical margin, weakly concave flanks that are gently converging towards the flat venter, which is deliminated by an angular ventrolateral shoulder. In the phragmocone, the whorl profile is oval with a broadly rounded venter. The change of the sculpture from the subadult stage with coarse ventrolateral nodes (12 per volution) towards the adult stage with only weak ribs on the flank happens already before the beginning of the terminal body chamber. Specimen MB.C.30046 is smaller (48 mm dm) and shows the transformation from the subadult stage with rounded ventrolateral margin to the adult stage with quadrate whorl profile, in which the venter is flat and separated from the flanks by an angular margin (Fig. 57B). The sculpture of the subadult stage possesses prominent ribs (about 13 per volution), which weaken out on the terminal body chamber. The suture line of specimen MB.C.30046 shows a simply subdivided external lobe with narrow, V-shaped prongs, a broadly rounded parabolic ventrolateral saddle and asymmetric, weakly serrated adventive and lateral lobes (Fig. 57D). Remarks Abichites alibashiensis differs from A. abichi, A. stoyanowi and A. subtrapezoidalis in the sharp ribs, which tend to end in ventrolateral nodes, in the subadult stage. Species of Alibashites possess subadult ribs but also ventrolateral nodes; these are A. mojsisovicsi (with very weak adult ribs) and A. ferdowsii (with a much more depressed whorl profile). Stratigraphic range Upper part of the Hambast Formation; 1.15 to 0.40 m below the extinction horizon (Abichites abichi Zone to lowest part of the Arasella minuta Zone).
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39. Arasella minuta
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Xenodiscidae ,Arasella ,Animalia ,Biodiversity ,Arasella minuta ,Taxonomy - Abstract
Arasella minuta (Zakharov, 1983) Fig. 20; Table 5 Sinoceltites ? minutus Zakharov in Kotlyar et al., 1983: 153, pl. 15 fig. 1, 2. Arasella minuta – Korn in Ghaderi et al. 2014: text-fig. 7I. — Korn & Ghaderi in Korn et al. 2016: 887, text-fig. 44. Diagnosis Species of Arasella with conch reaching 35 mm dm. Subadult and adult stage with circular whorl profile (ww/wh =0.85–1.05) and rounded venter; 20–22 sharp ribs on the flanks. External lobe with V-shaped prongs; adventive and lateral lobes broadly rounded. Type material Holotype AZERBAIJAN • Nakhichevan Province, Akhura; probably top of the Paratirolites Limestone; illustrated by Zakharov (in Kotlyar et al. 1983: pl. 15 fig. 1); BPI 4/813. Material examined IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29720 • 3 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29721 to MB.C.29723 • 8 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29724 to MB.C.29729 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29730 • 2 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29731 to MB.C.29732. Description Specimen MB.C.29720 has 25 mm conch diameter and is moderately well preserved (Fig. 20A). Less than half of the last volution belongs to the body chamber. The extremely and evolute conch (uw/ dm =0.48) shows a nearly circular whorl profile of the body chamber. The sculpture of the phragmocone and the terminal body chamber does not differ markedly; the phragmocone has slightly coarser but more rounded radial and sharp ribs on the flank, while the ribs on the body chamber are sharper and coarsest in the ventrolateral area. There are about 15 ribs per half volution. Remarks Species of Arasella minuta can easily be separated from all other contemporaneous ammonoids from the Iranian localities because of its very simple suture line with rounded adventive and lateral lobes and the short external lobe (Korn et al. 2016). Arasella falcata sp. nov. possesses a much weaker sculpture with ribs coarsest in in dorsolateral area. Stratigraphic range Topmost part of the Hambast Formation; immediately below the extinction horizon (Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 24-25, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Kotlyar G. V., Zakharov Y. D., Koczyrkevicz B. V., Kropatcheva G. S., Rostovcev L. O., Chedija I. O., Vuks G. P. & Guseva E. A. 1983. Posdnepermskiy etap evolyutsii organicheskogo mirA. Dzhulficheskiy i dorashamskiy yarusy SSSR. NAUKA, Leningrad.","Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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40. Arasella Korn
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Xenodiscidae ,Arasella ,Animalia ,Biodiversity ,Taxonomy - Abstract
Genus Arasella Korn in Ghaderi et al., 2014 Type species Sinoceltites? minutus Zakharov in Kotlyar et al., 1983, by original designation Included species Sinoceltites? minutus Zakharov in Kotlyar et al., 1983; Arasella falcata Korn & Hairapetian sp. nov. Remarks The genus Arasella is somewhat problematic because of its very simple suture line; an unambiguous attribution to a distinct family is therefore difficult. The shape of the external lobe rather speaks for placing it in the family Xenodiscidae rather than the Dzhulfitidae. The shape of the conch and the sculpture are similar to the early Changhsingian Shevyrevites, but this genus has multidentate adventive and lateral lobes., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 24, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Kotlyar G. V., Zakharov Y. D., Koczyrkevicz B. V., Kropatcheva G. S., Rostovcev L. O., Chedija I. O., Vuks G. P. & Guseva E. A. 1983. Posdnepermskiy etap evolyutsii organicheskogo mirA. Dzhulficheskiy i dorashamskiy yarusy SSSR. NAUKA, Leningrad."]}
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41. Esfahanites Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Esfahanites ,Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Genus Esfahanites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 61EDFE2C-1650-4303-BDC4-3E9F0434CDB8 Type species Esfahanites armatus Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Representative of the family Dzhulfitidae with large conch; maximum adult diameters about 120 mm. Adult stage with trapezoidal whorl profile. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs. Suture line with very small external lobe reaching only half the depth of adventive lobe. Etymology Named after the city of Esfahan. Included species Esfahanites armatus Korn & Hairapetian gen. et sp. nov. Remarks Esfahanites gen. nov. differs from Paratirolites and related genera in the very small external lobe.
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42. Paratirolites baghukensis Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Paratirolites baghukensis ,Taxonomy - Abstract
Paratirolites baghukensis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 27027DEB-9F15-4F61-8EA3-3F15E997A1D9 Figs 33–34; Table 16 Diagnosis Species of Paratirolites with conch reaching 130 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.00–1.20) and rounded venter; about 12 moderately coarse ribs ending in conical ventrolateral nodes per volution. Adult stage with trapezoidal and weakly compressed whorl profile (ww/wh =0.70–0.90), flattened tectiform venter and subangular ventrolateral shoulder; numerous sharp ribs ending in sharp radially elongated ventrolateral nodes. Prongs of external lobe usually trifid; altogether 13–15 notches of E, A and L lobes. Etymology Named after the occurrence at Baghuk Mountain. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain G section; Hambast Formation; illustrated in Fig. 33A; MB.C.29868. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29849 to MB.C.29850 • 4 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29851 to MB.C.29854 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29855 to MB.C.29859 • 8 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29860 to MB.C.29867 • 7 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29869 to MB.C.29875 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29876 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29877 to MB.C.29879. Description Holotype MB.C.29868 has 70 mm conch diameter and is fairly well-preserved but slightly corroded. The last third of the terminal volution belongs to the body chamber (which is thus only incompletely preserved); adulthood of the specimen is obvious because of the crowding of the last septa of the phragmocone (Fig. 33A). The penultimate of the preserved whorls shows a trapezoidal profile with a rounded venter. The body chamber finally shows a trapezoidal whorl profile with weakly diverging flanks, an angular ventrolateral shoulder and a nearly flat, tectiform venter. The sculpture changes with the beginning of the terminal body chamber, but this change is continuous without sudden breaks. One subadult volution has ten isolated spiny ventrolateral nodes.Already towards the end of the phragmocone, these nodes become radially elongate and form coarse ribs across the flank. At the beginning of the terminal body chamber, the ventrolateral nodes weaken and finally, only sharp ribs, about 15 per quarter of a volution, are present. The suture line of holotype MB.C.29868 shows a slightly pouched external lobe with trifid, asymmetric prongs. The ventrolateral saddle is weakly inflated, the adventive and lateral lobes are flattened at the base and subdivided in five and four notches, respectively (Fig. 33C). Paratype MB.C.29860 (Fig. 33B) is smaller with 61 mm conch diameter but confirms the description given for the holotype. It shows four whorls preserved and thus allows for the study of the sculpture of the inner volutions. The two larger paratypes MB.C.29874 and MB.C.29850 (Fig. 34) show then the transformation into the weakly ornamented adult stage, in which a tendency towards a parallel-sided whorl profile is visible. Remarks Paratirolites baghukensis sp. nov. belongs to the slender species of the genus. It differs from the other species of the genus in the sharp ribs on the adult body chamber. Paratirolites multiconus also has a slender conch, but differs from P. baghukensis sp. nov. in the much more numerous ventrolateral nodes. Stratigraphic range Upper part of the Hambast Formation; 3.10 to 2.35 m below the extinction horizon (Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 49-52, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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43. Dzhulfites brevisellatus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Dzhulfites ,Dzhulfites brevisellatus ,Animalia ,Biodiversity ,Taxonomy - Abstract
Dzhulfites brevisellatus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 1FDCA9D7-546A-48FA-94E1-0053EB7F7C23 Fig. 23; Table 8 Diagnosis Species of Dzhulfites with conch reaching 160 mm dm. Subadult stage with weakly trapezoidal, moderately depressed whorl profile (ww/wh =1.50–1.80) and broadly rounded venter; 10–13 coarse conical nodes on midflank per volution. Adult stage with numerous weak ribs and small ventrolateral nodes. Prongs of external lobe bifid or trifid; altogether 15–20 notches of E, A and L lobes. Etymology From the Latin ‘ brevis ’, meaning ‘low’ and ‘ sella ’, meaning ‘saddle’, after the low amplitude of lobes and saddles in the suture line. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain H section; Hambast Formation, 5.70 m below top; illustrated in Fig. 23B; MB.C.29748. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29746 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29747. Description The large paratype MB.C.29746 has 125 mm conch diameter, of which half of the last whorl belongs to the strongly weathered body chamber (Fig. 23A). The phragmocone is 90 mm in diameter and is partly crushed; it shows that the whorl profile is trapezoidal with a broadly rounded venter. The sculpture changes dramatically throughout ontogeny. At 40 mm dm there are 13 coarse, spiny ventrolateral nodes per volution; these nodes become weaker and more numerous so that the last 120° of the phragmocone have about 20 weak ventrolateral nodes, which are connected with weak and slightly sinuous radial ribs on the flank. The suture line of the specimen suffered from weathering, but, most probably, the prongs of the very short external lobe are bifid. Holotype MB.C.29748 is a fairly well-preserved, fully septate specimen with 51 mm conch diameter (Fig. 23B). It has a weakly trapezoidal whorl profile with broadly rounded flanks and venter. The sculpture shows about ten coarse conical nodes, which are positioned on the midflank, on the last preserved volution. The suture line of the holotype shows an external lobe with subparallel, weakly diverging flanks and weakly asymmetric, trifid prongs. All three saddles of the outer suture line are similar in shape, being inverted U-shaped and broadly rounded. The adventive lobe and the lateral lobe are similar in shape with parallel flanks and many small notches at the base. They differ in their depth; the adventive lobe is much deeper than the external lobe (Fig. 23C). Remarks Among the species of Dzhulfites, only D. nodosus is similar in conch shape and sculpture, but this species shows an applanate venter in the adult stage. Dzhulfites brevisellatus sp. nov. differs from D. spinosus and the other species of the genus in the suture line, which in D. brevisellatus sp. nov. shows much wider elements; particularly the saddles are wider than the lobes. In the shape of the suture line, D. brevisellatus sp. nov. has some resemblance to representatives of the family Xenodiscidae. Stratigraphic range Upper part of the Hambast Formation; 5.70 to 4.70 m below the extinction horizon (Dzhulfites spinosus Zone to Shevyrevites shevyrevi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 30-32, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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44. Alibashites ferdowsii Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Alibashites ferdowsii ,Biodiversity ,Alibashites ,Taxonomy - Abstract
Alibashites ferdowsii Korn & Ghaderi in Korn et al., 2016 Figs 48–49; Table 24 Alibashites ferdowsii Korn & Ghaderi in Korn et al., 2016: 871, text-fig. 30. Diagnosis Species of Alibashites with conch reaching 100 mm dm. Subadult stage with circular to subtrapezoidal, weakly depressed whorl profile (ww/wh =1.25–1.50) and broadly rounded venter; 10–12 very coarse ventrolateral nodes. Adult stage with subtrapezoidal and weakly depressed whorl profile (ww/wh =1.20– 1.50) with converging flanks, almost flat venter and angular ventrolateral shoulder; sharp ribs on flanks, forming short dorsolateral nodes and stronger ventrolateral nodes. Prongs of external lobe variable in shape, bifid to multiply serrated; altogether 15–18 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 30a); MB.C.25327. Additional material IRAN • 6 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29976 to MB.C.29978 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29979 • 7 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29980 to MB.C.29986 • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29987 to MB.C.29988 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29989 to MB.C.29990. Description Specimen MB.C.29974 is the largest available specimen with 59 mm conch diameter (Fig. 48A). Slightly less than half of a volution belongs to the body chamber. The profile of the aperture is characterised by a broadly rounded umbilical margin, weakly concave flanks, an angular ventrolateral shoulder and a broadly rounded but somehow flattened venter. The whorl profile in this stage is weakly depressed with a ww/wh ratio of 1.35. One volution earlier, the venter is more rounded and the flanks are widely curved, producing a depressed oval whorl profile. The sculpture changes from the subadult stage with coarse ventrolateral nodes per volution into the body chamber sculpture with sharp dorsolateral ribs, which are connected by low and rounded folds with weak ventrolateral nodes. The suture line of specimen MB.C.29974 is characterised by a Y-shaped external lobe, of which the prongs are subdivided by four small notches of varying depth. The ventrolateral saddle is strikingly asymmetric and dorsally inclined. It is followed by the asymmetric adventive lobe with seven small notches, the asymmetric parabolic lateral saddle and a lateral lobe with three notches (Fig. 49A). Specimen MB.C.29987 is a specimen with 51 mm conch diameter. Its profile is subtrapezoidal at the aperture; it shows a convex and oblique umbilical wall and is widest in the dorsolateral portion of the flank and weakly concave in the outer flank area (Fig. 48C). The flattened venter is separated from the flank by an angular ventrolateral shoulder. One whorl earlier is the whorl profile trapezoidal with a rounded ventrolateral shoulder and a broadly rounded venter. The sculpture on the phragmocone consists of twelve coarse conical nodes; on the body chamber these nodes become weaker and are connected with elongate rounded dorsolateral ribs by shallow, rounded folds. The suture line shows a slightly pouched external lobe with lanceolate, unsubdivided prongs. The ventrolateral saddle is turned U-shaped with nearly parallel flanks. Both flanks of the adventive lobe stand parallel, but its base is oblique and serrated into six little notches (Fig. 49D). Remarks The specimens from Baghuk Mountain are morphologically very similar to the material from the type area, and the sizes of the specimens are nearly identical. Differences occur in the degree of sutural subdivision; while the prongs of the external lobe in the material from Julfa are secondarily subdivided (two, three or four notches), many of the specimens from Baghuk Mountain show non-subdivided lanceolate prongs. Alibashites ferdowsii shows an intermediate morphological position between the genera Paratirolites and Abichites. Characteristic for Paratirolites is the intermediate growth stage with trapezoidal whorl profile with the prominent conical ventrolateral nodes; similar to Abichites is the flat venter in the adult stage. The intermediate position of A. ferdowsii is also visible in the suture line; on the one side is the more complex external lobe in Paratirolites (with serrated prongs of the external lobe), and on the other side is Abichites (with a trend towards unserrated prongs of the external lobe). Stratigraphic range Upper part of the Hambast Formation; 2.25 to 0.70 m below the extinction horizon (Alibashites ferdowsii Zone to lower part of the Abichites stoyanowi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 73-76, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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45. Stoyanowites parallelus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Stoyanowites ,Dzhulfitidae ,Animalia ,Biodiversity ,Stoyanowites parallelus ,Taxonomy - Abstract
Stoyanowites parallelus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: E6C0C2D8-02EF-4E72-9E1E-78816A50573E Fig. 63; Table 37 Diagnosis Species of Stoyanowites with conch reaching 100 mm dm. Subadult stage with rectangular, compressed whorl profile (ww/wh =0.70), subangular ventrolateral shoulder and flattened venter; 10 rather coarse ventrolateral spines per half volution. Adult stage with parallel-sided, compressed whorl profile (ww/ wh= 0.60), flat venter and angular ventrolateral shoulder; 15 ventrolateral spines per half volution. Prongs of external lobe simple; altogether 13 notches of E, A and L lobes. Etymology Named after the parallel flanks of the whorl profile. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 1.70 m below the top; illustrated in Fig. 63A; MB.C.30102. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30103 to MB.C.30104 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30105. Description Holotype MB.C.30102 is an incomplete specimen of 72 mm diameter; it allows the study of conch shape, ornament and suture line. The conch is extremely discoidal (ww/dm= 0.19) with a wide umbilicus (uw/ dm =0.47) and a compressed whorl profile (ww/wh =0.61). It has nearly parallel, slightly concave flanks, a prominent subangular ventrolateral shoulder and a weakly concave venter at the largest diameter (Fig. 63A). The sculpture is dominated by coarse ventrolateral spines, which become weaker towards the end of growth, but of which are still 15 are present on half a volution of the body chamber. They are not always connected with short ribs on the inner flank. The suture line of holotype MB.C.30102 shows the short external lobe, which has simple, weakly pouched prongs. The ventrolateral saddle is rather narrow; the adventive and lateral lobes possess six notches (Fig. 63B). Remarks Stoyanowites parallelus sp. nov. differs from S. dieneri and S. aspinosus in the flat venter and the numerous sharp ventrolateral nodes in the adult stage. Stratigraphic range Upper part of the Hambast Formation; 1.70 m below the extinction horizon (Alibashites ferdowsii Zone).
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46. Clivotirolites petilus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. et sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Clivotirolites ,Dzhulfitidae ,Animalia ,Biodiversity ,Clivotirolites petilus ,Taxonomy - Abstract
Clivotirolites petilus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 6608C3B9-8B98-4E05-91E6-DA463119748F Fig. 44; Table 21 Diagnosis Species of Clivotirolites gen. nov. with conch reaching 120 mm dm. Subadult stage with trapezoidal, weakly compressed whorl profile (ww/wh=0.90) and flattened tectiform; 15 coarse ventrolateral nodes per volution. Adult stage with trapezoidal, weakly compressed whorl profile (ww/wh =0.70), flat venter and angular ventrolateral shoulder; weak ribs and ventrolateral nodes. Prongs of external lobe bifid; altogether 16 notches of E, A and L lobes. Etymology From the Latin ‘ petilus ’, meaning ‘slim’, because of the slender conch. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 44A; MB.C.29962. Description Holotype MB.C.29962 is a comparatively well-preserved individual with 80 mm conch diameter and allows the study of two whorls (Fig. 44A). Nearly three quarters of the last volution belong to the body chamber. The adult whorl profile is compressed (ww/wh ~ 0.80) and weakly trapezoidal with gently diverging flanks, a subangular ventrolateral shoulder and a flattened but still weakly rounded venter. The sculpture of the last phragmocone volution has 15 coarse ventrolateral nodes. With beginning of the terminal body chamber, these nodes become weaker and more numerous; at the end of the body chamber, ten faint nodes occur in a quarter of a volution. The suture line of the holotype has an external lobe with weakly converging flanks. Its prongs are slightly pouched and possess three denticles. The ventrolateral saddle is inflated and followed by an asymmetric, pouched adventive lobe with oblique, weakly rounded base and seven small notches. The lateral lobe is generally V-shaped with weakly diverging flanks; it has six notches (Fig. 44B). Remarks Clivotirolites petilus gen. et sp. nov. differs in the lower number (about 15 per volution) of ventrolateral nodes from the species of Clivotirolites decorates gen. et sp. nov. Stratigraphic range The single specimen was collected from float of the upper part of the Hambast Formation.
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47. Abichites abichi Shevyrev 1965
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Abichites ,Cephalopoda ,Mollusca ,Abichites abichi ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Abichites abichi Shevyrev, 1965 Fig. 56; Table 30 Abichites abichi Shevyrev, 1965: 181, pl. 24 fig. 4. Xenodiscus aff. kapila – Stoyanow 1910: 87, pl. 9 fig. 3. Abichites abichi – Shevyrev 1968: 96, pl. 4 fig. 4. — Korn & Ghaderi in Korn et al. 2016: 878, textfig. 35. Diagnosis Species of Abichites with conch reaching 75 mm dm. Subadult stage with subquadrate, weakly depressed to compressed whorl profile (ww/wh =1.20) and slightly flattened venter; with 15 rounded straight ribs on the flanks. Adult stage with quadrate whorl profile (ww/wh =1.00), parallel flanks and flattened venter; with weak rounded ribs on the flanks. Prongs of external lobe usually simple; 7–12 notches of E, A and L lobes. Type material Holotype AZERBAIJAN • Nakhichevan Province, Dorasham 2 section; Paratirolites Limestone; illustrated by Shevyrev (1965: pl. 24 fig. 4); PIN 1252/137. Material examined IRAN • 4 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30019 to MB.C.30022 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30023 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30024 to MB.C.30028 • 6 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30029 to MB.C.30034 • 4 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30035 to MB.C.30038 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30039 to MB.C.30040. Description Specimen MB.C.30039 has 44 mm diameter and is a fragment of half a volution (Fig. 56A). However, it allows the study of the conch shape with a nearly quadrate whorl profile. On half of a volution there are twelve sharp radial ribs, which are coarsest in the outer flank. Specimen MB.C.30019 has 40 mm conch diameter and allows the study of only one volution (Fig. 56B). The whorl profile is subtrapezoidal with broadly rounded umbilical margin, weakly converging flanks, a subangular ventrolateral shoulder and a flat venter. The sculpture is weaker than in the first specimen and the ribs are coarsest on the inner flank. The suture line of specimen MB.C.30019 is characterised by a large external lobe, of which the flanks stand parallel in the middle part. Its prongs are narrowly V-shaped and non-serrated. After a ventrolateral saddle that is as wide as the external lobe follow a small, nearly parallel-sided adventive lobe with three notches, a small lateral saddle and a small parallel-sided lateral lobe with three notches (Fig. 56C). Remarks Abichites abichi differs from the otherwise similar species A. subtrapezoidalis in the quadrate adult whorl profile (subtrapezoidal in A. subtrapezoidalis). The body chamber of A. abichi has stronger ribs than A. subtrapezoidalis does. A. alibashiensis has a similar body chamber, but differs from A. abichi in the coarse ventrolateral nodes of the subadult stage. Stratigraphic range Upper part of the Hambast Formation; 1.20 to 0.05 m below the extinction horizon (Abichites abichi Zone to Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 87-89, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135.","Shevyrev A. A. 1968. Triasovye ammonoidei Yuga SSSR. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 119: 1 - 272.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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48. Paratirolites serus Korn & Ghaderi 2016
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Paratirolites ,Animalia ,Biodiversity ,Paratirolites serus ,Taxonomy - Abstract
Paratirolites serus Korn & Ghaderi in Korn et al., 2016 Fig. 40; Table 19 Paratirolites serus Korn & Ghaderi in Korn et al., 2016: 868, text-fig. 27. Diagnosis Paratirolites with conch reaching 80 mm dm. Subadult stage with weakly to strongly trapezoidal, depressed whorl profile (ww/wh =1.40–1.80) and broadly rounded venter; 12–14 coarse conical ventrolateral nodes. Adult stage with circular or slightly trapezoidal, weakly to moderately depressed whorl profile (ww/wh =1.20–1.60), flattened venter and subangular to angular ventrolateral shoulder; numerous ribs on the flank. Prongs of external lobe simple; altogether 6–9 notches of E, A and L lobes. Material examined Holotype IRAN • East Azerbaijan, Ali Bashi N section; Paratirolites Limestone, 0.65 m below the top; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 27a); MB.C.25300. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29934 • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29935 to MB.C.29936 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29937. Description Specimen MB.C.29936 has 62 mm conch diameter and is the largest available specimen (Fig. 40A). It is corroded but still rather well preserved and allows the study of three whorls. A little more than half a volution belongs to the body chamber. The weakly depressed whorl profile (ww/wh = 1.05) of the body chamber is trapezoidal with weakly diverging flanks and the flattened venter is separated from the flanks by a subangular ventrolateral shoulder. The sculpture changes, on the last portion of the phragmocone, from a subadult stage with 14 coarse ventrolateral nodes per volution to the adult stage with numerous weak radial ribs with weak ventrolateral nodes on the flank. Specimen MB.C.29934 is a smaller individual with 48 mm conch diameter and is better preserved than the preceding one with half a volution of the body chamber preserved (Fig. 40B). It shows a weakly trapezoidal, weakly depressed body chamber whorl profile (ww/wh=1.05) with rounded ventrolateral shoulder and a broadly rounded venter, while the phragmocone has a more depressed whorl profile (ww/ wh= 1.40). The sculpture in the subadult stage has twelve coarse ventrolateral nodes, which become weaker and radially elongated on the body chamber, where they finally disappear and instead shallow radial dorsolateral ribs take place. The suture line of specimen MB.C.29934 has an external lobe with nearly parallel flanks and unserrated, narrow lanceolate prongs. The ventrolateral saddle is weakly asymmetric and the adventive lobe is strongly asymmetric with a steep ventral flank, four little notches and an oblique dorsal side. The lateral lobe possesses only three small notches (Fig. 40D). Remarks Paratirolites serus is the stratigraphically youngest species of the genus in the Baghuk Mountain sections. It differs from the other by its small conch size and the simple suture line with unsubdivided prongs of the external lobe. Subadult specimens of P. serus resemble some stratigraphically older species such as P. kittli and particularly P. vediensis. Paratirolites serus differs from these in the smaller conch diameter (40 mm maximum phragmocone diameter in contrast to 70–80 mm in P. kittli and P. vediensis) and a simpler suture line with unsubdivided prongs of the external lobe. Stratigraphic range Upper part of the Hambast Formation; 0.75 to 0.60 m below the extinction horizon (Abichites stoyanowi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 59-61, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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49. Lutites Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Dzhulfitidae ,Animalia ,Biodiversity ,Lutites ,Taxonomy - Abstract
Genus Lutites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 5B2AB929-F903-46B0-B108-A1C57458A1DA Type species Lutites lyriformis Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Genus of the family Dzhulfitidae with large conch; maximum adult diameters about 140 mm. Adult stage with compressed, subtrapezoidal whorl profile; flanks concave, venter flat or concave. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs or delicate plications. Suture line with deep external lobe (deeper than adventive lobe), ventral prongs unsubdivided or notched. Etymology Named after the Lut Desert, which extends to the North-East of Baghuk Mountain. Included species Alibashites stepanovi Korn & Ghaderi in Korn et al., 2016; Lutites paucis Korn & Hairapetian gen. et sp. nov.; Lutites alius Korn & Hairapetian gen. et sp. nov.; Lutites profundus Korn & Hairapetian gen. et sp. nov.; Lutites lyriformis Korn & Hairapetian gen. et sp. nov.; Lutites plicatus Korn & Hairapetian gen. et sp. nov. Remarks Lutites gen. nov. differs from Paratirolites and related genera in the compressed subtrapezoidal, pearshaped whorl profile and in the very deep external lobe., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 76, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}
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50. Clivotirolites decoratus Korn & Hairapetian & Ghaderi & Leda & Schobben & Akbari 2021, gen. et sp. nov
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Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin, and Akbari, Amir
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Cephalopoda ,Mollusca ,Ammonoidea ,Clivotirolites ,Clivotirolites decoratus ,Dzhulfitidae ,Animalia ,Biodiversity ,Taxonomy - Abstract
Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 0A7C7606-6D92-4630-BAF3-1A8E30B6486D Figs 41–43; Table 20 Diagnosis Species of Clivotirolites gen. nov. with conch reaching 110 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh=1.10–1.20) and slightly flattened venter; about 20–24 weak ventrolateral spines per volution. Adult stage with parallel or weakly diverging flanks and weakly compressed whorl profile (ww/wh =0.70–0.80), flattened, slightly tectiform venter and angular ventrolateral shoulder; numerous rounded ribs on flank ending in sharp ventrolateral nodes. Prongs of external lobe variable, simple to digitate; altogether 10–17 notches of E, A and L lobes. Etymology From the Latin ‘ decoratus ’, meaning ‘decorated’, because of the sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 1.35 m below the top; illustrated in Fig. 41A; MB.C.29941. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29938 • 2 specimens; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29939 to MB.C.29940 • 14 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29942 to MB.C.29955 • 2 specimens; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29956 to MB.C.29957 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29958 to MB.C.29959 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29960 to MB.C.29961. Description Holotype MB.C.29941 is a rather well-preserved specimen of 64 mm conch diameter and shows all typical characters for the species (Fig. 41A). The specimen shows one and a half whorls preserved. It displays septal crowding and the end of the phragmocone appears at 38 mm diameter.About 210 degrees of the last volution belong to the body chamber, which shows a weakly trapezoidal whorl profile with weakly concave, weakly diverging flanks, a subangular ventrolateral shoulder and a flattened tectiform venter. The whorl belonging to the phragmocone has a more pronounced trapezoidal profile with convex, diverging flanks. The sculpture on the phragmocone part consists of rather small, pointed ventrolateral tubercles, 22 per volution. On the body chamber, these tubercles are connected with weak and rounded radial ribs, which are strongest on the inner flank. Paratype MB.C.29946 is a similar specimen with 58 mm conch diameter and the last septum at 33 mm dm (Fig. 41B). It possesses a nearly quadrate whorl cross section of the body chamber but also shows that the profile is trapezoidal in an earlier growth stage. On the last volution of the phragmocone, 20 sharp ventrolateral nodes can be counted. Larger paratypes such as MB.C.29947 (50 mm phragmocone diameter), MB.C.29944 (52 mm; Fig. 42B), MB.C.29938 (52 mm; Fig. 42A) and MB.C.29958 (53 mm; Fig. 43A) are generally similar but show stronger diverging flanks of the body chamber. Suture lines were drawn from four specimens, all of them show principally similar outlines with some common characters (Fig. 43B–E). The external lobe is not as deep as the adventive lobe, it is wider at its base, from where the flanks converge towards the ventrolateral saddle. All four specimens show asymmetric notching of its prongs; usually there are three notches but also more intense serration is possible. The ventrolateral saddle is usually slightly inflated and the adventive lobe is parallel-sided with five or six notches at its base. Remarks Clivotirolites decoratus gen. et sp. nov. differs in the large number (more than 20 per volution) of ventrolateral nodes from the species of Clivotirolites petilus gen. et sp. nov. (about 15 nodes per volution). Stratigraphic range Upper part of the Hambast Formation; 2.05 to 1.10 m below the extinction horizon (Alinashites ferdowsii Zone to Abichites abichi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 62-64, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715
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