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The family Hahniidae is a small family known from 23 genera. Only one genus Hahnia C.L. Koch, 1841 is known from South Africa represented by eight species of which seven are endemic to South Africa. Only one species Hahnia schubotzi Strand, 1913 was not previously listed from South Africa The African fauna was last revised by Bosmans (1981). Six of the species are listed as Least Concern and two species are data deficient.

Subtribe Thiratoscirtina Bodner & Maddison, 2012 Remarks. The Thiratoscirtina is a major species-rich group in African tropical forests ��� e.g., more than 52 species were found in 17 days of field work in Gabon by the second author ��� and yet its genera received little attention before Wanda Wesołowska began studying them. Of those 52 Gabonese species, we are able to confidently place fewer than half into described genera. Uncertainty in the group���s generic limits is heightened by their unusually variable genitalia, by the lack of taxonomic synthesis between vouchers of molecular data (Bodner & Maddison 2012), and by the growing number of named taxa. Faced with this, we could decide to stop describing species until genera are settled, to describe a new genus for each species of unclear affinity, or to place new species tentatively into existing genera. Describing species is so important for conservation and comparative biology that we do not want to hold back descriptions. We have thus chosen to describe two species here tentatively in Ajaraneola and Nimbarus, even though we must admit that their generic placements may change., Published as part of Sz��ts, Tam��s & Maddison, Wayne Paul, 2021, New species of the monotypic thiratoscirtine genera Ajaraneola and Nimbarus (Araneae: Salticidae: Aelurillini: Thiratoscirtina), pp. 119-126 in Zootaxa 4915 (1) on page 120, DOI: 10.11646/zootaxa.4915.1.8, http://zenodo.org/record/4452548, {"references":["Bodner, M. R. & Maddison, W. P. (2012) The biogeography and age of salticid spider radiations (Araneae: Salticidae). Molecular Phylogenetics and Evolution, 65 (1), 213 - 240. https: // doi. org / 10.1016 / j. ympev. 2012.06.005"]}

Hahnia ngai sp. nov. urn:lsid:zoobank.org:act: F9A7FE4B-77C4-4D9B-AFDB-A2FC6F5E0739 Figs 4, 6 a���c Diagnosis Hahnia ngai sp. nov. can be easily separated from other members of this genus by the simplified female genitalia. Copulatory ducts show only slightly swollen areas with glandular insertions (Figs 4g, 6 b���c) but do not form a receptacle or secondary spermathecae (as seen in Figs 5g, 6 e���f). Etymology The species epithet is a derivation of the Thai ngai (simple), in reference to the relatively simple vulva without the well-formed secondary spermathecae commonly seen in other species of Hahnia. Type material Holotype THAILAND ��� ♀; Chiang Mai Province, Doi Suthep National Park; 18��48.502��� N, 98��53.528��� E; 1409 m a.s.l.; 24���28 Jul. 2018; Booppa Petcharad, Jeremy Miller and F. Andr��s Rivera-Quiroz leg.; montane evergreen forest with pine; Winkler extractor; RMNH.ARA.18415 (four legs used for DNA extraction). Paratypes THAILAND ��� 1 ♀; Chiang Mai Province, Doi Inthanon National Park; 18��35.268��� N, 98��29.240��� E; 2572 m a.s.l.; 24���28 Jul. 2018; Booppa Petcharad, Jeremy Miller and F. Andr��s Rivera-Quiroz leg.; cloud forest; Winkler extractor; RMNH.ARA.18414 ��� 1 ♀; same collection data as for preceding; hand coll.; RMNH.ARA.18413. Description Female Carapace pear-shaped, reddish-brown, slightly darker in cephalic region; texture smooth (Fig. 4c). AME 0.04 mm, ALE 0.06 mm, PME 0.07 mm, PLE 0.04 mm, AME���AME 0.03 mm, AMEALE 0.02 mm, PME���PME 0.05 mm, PME���PLE 0.03 mm (Fig. 4d). Chelicerae with three promarginal and three retromarginal teeth (Fig. 4e). Legs pale brown, slightly darker on distal segments. Abdomen dark grey with light patches forming five to six chevron bands, oval, longer than wide (Fig. 4c). Tracheal spiracle near middle of abdomen (Fig. 4a). VULVA. Epigynal plate semitransparent, spermathecae well visible due to transparency. Copulatory openings close together, forming small semi-circular atrium (Figs 4 f���g, 6c). Spermatheca sub-spherical with brownish-red coloration (Fig. 4f). Copulatory ducts very simple, slightly swollen centrally (Figs 4f, 6 b���c). MEASUREMENTS (in mm). Total length 2.8, carapace 1.25 long, 0.91 wide; clypeus 0.09; chelicera 0.45 long, 0.25 wide; leg I: femur 0.95, patella 0.37, tibia 0.71, metatarsus 0.72, tarsus 0.55; leg II: femur 0.94, patella 0.34, tibia 0.72, metatarsus 0.68, tarsus 0.55; leg III: femur 0.89, patella 0.33, tibia 0.63, metatarsus 0.71, tarsus 0.51; leg IV: femur 1.12, patella 0.34, tibia 0.93, metatarsus 1.01, tarsus 0.62; leg formula IV-I-II-III; abdomen 1.65 long, 1.23 wide. Distribution Known from two localities in Chiang Mai, Thailand (Fig. 8): Doi Suthep National Park (type locality), and the neighboring Doi Inthanon National Park., Published as part of Rivera-Quiroz, Francisco Andr��s, Petcharad, Booppa & Miller, Jeremy A., 2020, First records and a new genus of comb-tailed spiders (Araneae: Hahniidae) from Thailand with comments on the six-eyed species of this family, pp. 51-69 in European Journal of Taxonomy 724 on pages 59-61, DOI: 10.5852/ejt.2020.724.1157, http://zenodo.org/record/4298074

Hahnia ngai sp. nov. urn:lsid:zoobank.org:act: F9A7FE4B-77C4-4D9B-AFDB-A2FC6F5E0739 Figs 4, 6 a–c Diagnosis Hahnia ngai sp. nov. can be easily separated from other members of this genus by the simplified female genitalia. Copulatory ducts show only slightly swollen areas with glandular insertions (Figs 4g, 6 b–c) but do not form a receptacle or secondary spermathecae (as seen in Figs 5g, 6 e–f). Etymology The species epithet is a derivation of the Thai ngai (simple), in reference to the relatively simple vulva without the well-formed secondary spermathecae commonly seen in other species of Hahnia. Type material Holotype THAILAND • ♀; Chiang Mai Province, Doi Suthep National Park; 18º48.502′ N, 98º53.528′ E; 1409 m a.s.l.; 24–28 Jul. 2018; Booppa Petcharad, Jeremy Miller and F. Andrés Rivera-Quiroz leg.; montane evergreen forest with pine; Winkler extractor; RMNH.ARA.18415 (four legs used for DNA extraction). Paratypes THAILAND • 1 ♀; Chiang Mai Province, Doi Inthanon National Park; 18º35.268′ N, 98º29.240′ E; 2572 m a.s.l.; 24–28 Jul. 2018; Booppa Petcharad, Jeremy Miller and F. Andrés Rivera-Quiroz leg.; cloud forest; Winkler extractor; RMNH.ARA.18414 • 1 ♀; same collection data as for preceding; hand coll.; RMNH.ARA.18413. Description Female Carapace pear-shaped, reddish-brown, slightly darker in cephalic region; texture smooth (Fig. 4c). AME 0.04 mm, ALE 0.06 mm, PME 0.07 mm, PLE 0.04 mm, AME–AME 0.03 mm, AMEALE 0.02 mm, PME–PME 0.05 mm, PME–PLE 0.03 mm (Fig. 4d). Chelicerae with three promarginal and three retromarginal teeth (Fig. 4e). Legs pale brown, slightly darker on distal segments. Abdomen dark grey with light patches forming five to six chevron bands, oval, longer than wide (Fig. 4c). Tracheal spiracle near middle of abdomen (Fig. 4a). VULVA. Epigynal plate semitransparent, spermathecae well visible due to transparency. Copulatory openings close together, forming small semi-circular atrium (Figs 4 f–g, 6c). Spermatheca sub-spherical with brownish-red coloration (Fig. 4f). Copulatory ducts very simple, slightly swollen centrally (Figs 4f, 6 b–c). MEASUREMENTS (in mm). Total length 2.8, carapace 1.25 long, 0.91 wide; clypeus 0.09; chelicera 0.45 long, 0.25 wide; leg I: femur 0.95, patella 0.37, tibia 0.71, metatarsus 0.72, tarsus 0.55; leg II: femur 0.94, patella 0.34, tibia 0.72, metatarsus 0.68, tarsus 0.55; leg III: femur 0.89, patella 0.33, tibia 0.63, metatarsus 0.71, tarsus 0.51; leg IV: femur 1.12, patella 0.34, tibia 0.93, metatarsus 1.01, tarsus 0.62; leg formula IV-I-II-III; abdomen 1.65 long, 1.23 wide. Distribution Known from two localities in Chiang Mai, Thailand (Fig. 8): Doi Suthep National Park (type locality), and the neighboring Doi Inthanon National Park.

Hahnia saccata Zhang, Li & Zheng, 2011 Figs 5, 6 d���f Hahnia saccata Zhang, Li & Zheng, 2011: 16, figs 14a���e, 15a���h, 16a���g. Material examined THAILAND ��� 2 ♀♀; Chiang Mai Province, Doi Suthep National Park; 18��48.780��� N, 98��55.928��� E; 643 m a.s.l.; 25���28 Jul. 2018; Booppa Petcharad, Jeremy Miller and F. Andr��s Rivera-Quiroz leg.; Dipterocarpus forest; hand coll.; RMNH.ARA.18412 (four legs of one specimen used for DNA extraction). Description Female Carapace pear-shaped, reddish-brown, slightly darker in cephalic region; texture smooth (Fig. 5c). AME 0.06 mm, ALE 0.11 mm, PME 0.08 mm, PLE 0.05 mm, AME���AME 0.02 mm, AMEALE 0.01 mm, PME���PME 0.06 mm, PME���PLE 0.04 mm (Fig. 5d). Chelicerae with three promarginal and seven retromarginal teeth (Fig. 5e). Legs same color as carapace, darker on proximal and distal part of each segment. Abdomen dark grey with light patches forming five to six chevron bands, oval, longer than wide (Fig. 5c). Tracheal spiracle near middle of abdomen (Fig. 5a). VULVA. Epigynal plate dark. Copulatory openings close together but not forming an atrium (Figs 5g, 6f). Spermatheca sub-speherical with brown coloration (Fig. 5f). Copulatory ducts forming secondary spermatheca (Figs 5f, 6 e���f). MEASUREMENTS (in mm). Total length 3.20, carapace 1.45 long, 1.04 wide; clypeus 0.10; chelicera 0.70 long, 0.33 wide; leg I: femur 1.22, patella 0.46, tibia 1.13, metatarsus 0.92, tarsus 0.63; leg II: femur 1.12, patella 0.45, tibia 0.90, metatarsus 0.81, tarsus 0.61; leg III: femur 0.98, patella 0.41, tibia 0.75, metatarsus 0.80, tarsus 0.49; leg IV: femur 1.31, patella 0.45, tibia 1.12, metatarsus 1.03, tarsus 0.65; leg formula IV-I-II-III; abdomen 1.73 long, 1.20 wide. Distribution Known from the Menglun Nature Reserve, Yunnan, China (type locality), and Doi Suthep National Park, Chiang Mai, Thailand (present work) (Fig. 8)., Published as part of Rivera-Quiroz, Francisco Andr��s, Petcharad, Booppa & Miller, Jeremy A., 2020, First records and a new genus of comb-tailed spiders (Araneae: Hahniidae) from Thailand with comments on the six-eyed species of this family, pp. 51-69 in European Journal of Taxonomy 724 on pages 61-62, DOI: 10.5852/ejt.2020.724.1157, http://zenodo.org/record/4298074, {"references":["Zhang Z. S., Li S. & Zheng G. 2011. Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa 2912 (1): 1 - 27. https: // doi. org / 10.11646 / zootaxa. 2912.1.1"]}

Genus Hahnia C.L. Koch, 1841 Hahnia C.L. Koch, 1841: 61. Type species Hahnia pusilla C.L. Koch, 1841., Published as part of Rivera-Quiroz, Francisco Andr��s, Petcharad, Booppa & Miller, Jeremy A., 2020, First records and a new genus of comb-tailed spiders (Araneae: Hahniidae) from Thailand with comments on the six-eyed species of this family, pp. 51-69 in European Journal of Taxonomy 724 on page 59, DOI: 10.5852/ejt.2020.724.1157, http://zenodo.org/record/4298074, {"references":["Koch C. L. 1841. Die Arachniden. Getreu nach der Natur abgebildet und beschrieben. Achter Band. C. H. Zeh'schen Buchhandlung, Nuremberg. https: // doi. org / 10.5962 / bhl. title. 43744"]}

The family Hahniidaei is a small family known from 23 genera. Only one genus Hahnia C.L. Koch, 1841 is known from South Africa represented by eight species of which seven are endemic to South Africa. Only one species Hahnia schubotzi Strand, 1913 was not previously listed from South Africa The African fauna was last revised by Bosmans (1981). Six of the species are listed as Least Concern and two species are data deficient.

Genus Hahnia C.L. Koch, 1841 Type species. Hahnia pusilla C.L. Koch, 1841 Remarks. The type genus of Hahniidae, Hahnia, is the largest genus of this family. But obviously, it is not monophyletic, including nearly one hundred species from Africa, Europe, Asia, North and South America. Judged from the type species, H. pusilla from Europe, Hahnia s. str. seems to have the combined characters as follows: generally eight eyes (H. senaria Zhang, Li & Zheng, 2011 excluded), spiracle located in the middle between epigastric furrow and spinnerets in both sexes; male chelicerae with stridulatory files laterally, male palp with a hook-like patellar apophysis, almost semicircular and with a pointed RTA, filiform embolus originating retrolaterally, a membranous median apophysis retrolatero-distally, sometimes tegular setae (H. himalayaensis Hu & Zhang, 1990) and a deep cymbial furrow retrolaterally; epigyne with a pair of copulatory openings, located in a large atrium (H. thorntoni Brignoli, 1982) or not (H. pusilla), copulatory ducts long (H. himalayaensis) or short (H. pusilla), spherical subspermathecae, smaller than spermathecae and anteriorly located, a pair of hoods generally located behind the epigastric furrow., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655, {"references":["Zhang, Z. S, Li, S. & Zheng, G. (2011) Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa, 2912, 1 - 27.","Hu, J. L. & Zhang, F. P. (1990) A new species of the genus Hahnia from Xizang Autonomous Region, China (Araneae: Hahniidae). Acta Zootaxonomica Sinica, 15, 165 - 168."]}

Hahnia dongi Huang & Zhang, sp. nov. Figs 3���4, 16 Type material. CHINA: Yunnan Province: Holotype male , Baoshan City, Longyang District, Nankang, Gaoligong Mountains Nature Park, 24��49'40.195" N, 98��46'9.228" E, elev. 2203m, 8 November 2016, Z.W. Dong leg. (SWUC). Etymology. This new species is dedicated to Mr. Zhi-Wei Dong, collector of the new species and good friend of the first author. Diagnosis. Male of the new species can be distinguished from other Hahnia species by the relatively short RTA, the short and protuberant LTA retrolateralo-proximally and the presence of some strong tibial setae (Figs 3E��� G, 4A���C). Description. Male (holotype). Total length 2.20. Prosoma 1.07 long, 0.79 wide; Opisthosoma 1.26 long, 0.93 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.05, PLE 0.08; AME���AME 0.03, AME���ALE 0.02, PME���PME 0.08, PME���PLE 0.06, ALE���PLE 0.02. MOA 0.17 long, front width 0.15, back width 0.20. Clypeus height 0.14. Chelicerae with stridulatory files laterally (Figs 3C���D), 2 promarginal, 6 retromarginal teeth. Leg measurements: I 4.56 (1.30, 1.63, 1.06, 0.57); II 3.50 (0.99, 1.18, 0.82, 0.51); III 2.90 (0.88, 0.92, 0.69, 0.41); IV 3.35 (0.99, 1.09, 0.79, 0.48). Leg formula: 1243. Spiracle long and transverse, distance of spiracle to epigastric furrow 2.4 times as long as to spinnerets (Fig. 3B). Male palpal patella with a hook-like apophysis retrolatero-proximally, the patellar apophysis with a notch apically. Tibia covered with cluster of black setae at dorsal and retrolateral surface, with moderately short RTA extending dorsally and LTA proximally, tip of LTA covered with several black setae. Embolus originating retrolatero-proximally, tip of embolus lying near base of membranous median apophysis. Cymbial furrow deep and nearly as long as half of cymbium (Figs 3E���G, 4A���C). Female unknown. Distribution. Known only from the type locality in Yunnan, China (Fig. 16)., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655

Hahnia wangi Huang & Zhang sp. nov. Figs 13–16 Type material. CHINA: Yunnan Province: Holotype male , Nujiang Prefecture, between Gongshan County and Dulongjiang Township along Pula River, 27°47.468′ N, 98°35.132′ E, elev. 1955 m, 24 November 2011, Z.X Li & L.Y. Wang leg. (SWUC); Paratypes: 1 male, 3 females, same data as for holotype; 5 females, between Gongshan County and Dulongjiang Township along Heiwadi, 27°47.115′ N, 98°35.893′ E, elev. 1859 m, 24 November 2011, Z.X Li & L.Y. Wang leg.; 2 females, Nujiang Prefecture, Gongshan County, Bingzhongluo Township, 28°00.866′ N, E:98°35.953′ E, elev. 1840 m, 6 June 2011 Z.X. Li & G.C. Zhou leg.; 1 female, Baoshan City, Longyang District, Mangkuan Township, Baihualing, 25°17.756' N, 98°47.646' E, elev. 1575 m, 2 March 2011, Z.X Li & L.Y. Wang leg. (all SWUC). Etymology. This new species is dedicated to Mr. Luyu Wang (SWUC) who is one of the collectors of the type series. Diagnosis. Males of the new species are similar to those of H. submaginii Zhang et al., 2011 in having the male palpal patella with a bifurcate apophysis retrolaterally, one part hook-like proximad and the other horn-like distad, the female abdominal venter with a pair of curved hoods behind epigastric furrow, but can be distinguished from the latter by the male patellar apophysis with a constriction basally at proximad side of the horn-like part (red arrow in Figs 13E, 13G), the horn-like part not bifurcate apically, the hook-like part of patellar apophysis longer, RTA sharply curved, proximad (Figs 13E–G, 15A–C), the hoods oblique, epigyne without big atrium, spermathecae and subspermathecae bigger and closer together. Description. Male (holotype). Total length 2.70. Prosoma 1.28 long, 0.93 wide; Opisthosoma 1.47 long, 1.03 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.08, PLE 0.09; AME–AME 0.03, AME–ALE 0.02, PME–PME 0.05, PME–PLE 0.05, ALE–PLE 0.02. MOA 0.19 long, front width 0.12, back width 0.23. Clypeus height 0.11. Chelicerae with distinct stridulatory files laterally (Figs 16C–D), 3 promarginal, 7 retromarginal teeth. Leg measurements: I 3.58 (1.04, 1.18, 0.77, 0.59); II 3.36 (0.96, 1.09, 0.73, 0.58); III 3.04 (0.88, 0.95, 0.71, 0.51); IV 3.84 (1.09, 1.22, 0.92, 0.61). Leg formula: 4123. Spiracle long and transverse, the distance of spiracle to epigastric furrow as long as to spinnerets (Fig. 13B). Embolus originating retro-basally, tip of embolus lying near base of membranous median apophysis. Cymbial furrow deep and nearly as long as half of cymbium (Figs 13E–G, 15A–C). Female (paratypes). Total length 3.00–3.26. Prosoma 1.38–1.50 long, 1.00–1.10 wide; Opisthosoma 1.67–1.82 long, 1.19–1.30 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.08, PLE 0.09; AME–AME 0.04, AME–ALE 0.03, PME–PME 0.08, PME–PLE 0.05, ALE–PLE 0.01; MOA 0.19 long, front width 0.13, back width 0.27. Clypeus height 0.13. Leg measurements: I 3.81 (1.12, 1.31, 0.77, 0.60); II 3.53 (1.04, 1.17, 0.74, 0.58); III 3.21 (0.94, 1.01, 0.73, 0.54); IV 4.15 (1.20, 1.37, 0.93, 0.66). Other characters as in males. Epigyne with a pair of small copulatory openings centrally. Copulatory ducts bifurcated near copulatory openings, short branch connected with subspermathecae and with tuberculate expansion near base (green arrows and red arrows indicate in Figs 14D, 15E), long branch roundly coiled, finally connected with spermathecae, the latter sub-globular and located posterior-laterally. Subspermathecae oval, smaller than spermathecae and next to globular spermathecae. Fertilization ducts short, originating from posterior of spermathecae and pointing laterally (Figs 14B–D, 15D–E). Remarks. Both patellar apophyses of the male paratype are broken. Distribution. Known only from the type locality in Yunnan, China (Fig. 16). FIGURESA–E 15. Hahnia wangi sp. nov., holotype male (A–C) and paratype female (D–E). A–C Left palp (A ventral, B retrolateral, C dorsal). D Epigyne, ventral. E Vulva, dorsal. CO = copulatory openings; CD = copulatory duct; CF = cymbial furrow; Em = embolus; FD = fertilization duct; Ho = hood; MA = median apophysis; PA = patellar apophysis; RTA = retrolateral tibial apophysis; Sp = spermatheca; Ss = subspermatheca.

Hahnia wangi Huang & Zhang sp. nov. Figs 13���16 Type material. CHINA: Yunnan Province: Holotype male , Nujiang Prefecture, between Gongshan County and Dulongjiang Township along Pula River, 27��47.468��� N, 98��35.132��� E, elev. 1955 m, 24 November 2011, Z.X Li & L.Y. Wang leg. (SWUC); Paratypes: 1 male, 3 females, same data as for holotype; 5 females, between Gongshan County and Dulongjiang Township along Heiwadi, 27��47.115��� N, 98��35.893��� E, elev. 1859 m, 24 November 2011, Z.X Li & L.Y. Wang leg.; 2 females, Nujiang Prefecture, Gongshan County, Bingzhongluo Township, 28��00.866��� N, E:98��35.953��� E, elev. 1840 m, 6 June 2011 Z.X. Li & G.C. Zhou leg.; 1 female, Baoshan City, Longyang District, Mangkuan Township, Baihualing, 25��17.756' N, 98��47.646' E, elev. 1575 m, 2 March 2011, Z.X Li & L.Y. Wang leg. (all SWUC). Etymology. This new species is dedicated to Mr. Luyu Wang (SWUC) who is one of the collectors of the type series. Diagnosis. Males of the new species are similar to those of H. submaginii Zhang et al., 2011 in having the male palpal patella with a bifurcate apophysis retrolaterally, one part hook-like proximad and the other horn-like distad, the female abdominal venter with a pair of curved hoods behind epigastric furrow, but can be distinguished from the latter by the male patellar apophysis with a constriction basally at proximad side of the horn-like part (red arrow in Figs 13E, 13G), the horn-like part not bifurcate apically, the hook-like part of patellar apophysis longer, RTA sharply curved, proximad (Figs 13E���G, 15A���C), the hoods oblique, epigyne without big atrium, spermathecae and subspermathecae bigger and closer together. Description. Male (holotype). Total length 2.70. Prosoma 1.28 long, 0.93 wide; Opisthosoma 1.47 long, 1.03 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.08, PLE 0.09; AME���AME 0.03, AME���ALE 0.02, PME���PME 0.05, PME���PLE 0.05, ALE���PLE 0.02. MOA 0.19 long, front width 0.12, back width 0.23. Clypeus height 0.11. Chelicerae with distinct stridulatory files laterally (Figs 16C���D), 3 promarginal, 7 retromarginal teeth. Leg measurements: I 3.58 (1.04, 1.18, 0.77, 0.59); II 3.36 (0.96, 1.09, 0.73, 0.58); III 3.04 (0.88, 0.95, 0.71, 0.51); IV 3.84 (1.09, 1.22, 0.92, 0.61). Leg formula: 4123. Spiracle long and transverse, the distance of spiracle to epigastric furrow as long as to spinnerets (Fig. 13B). Embolus originating retro-basally, tip of embolus lying near base of membranous median apophysis. Cymbial furrow deep and nearly as long as half of cymbium (Figs 13E���G, 15A���C). Female (paratypes). Total length 3.00���3.26. Prosoma 1.38���1.50 long, 1.00���1.10 wide; Opisthosoma 1.67���1.82 long, 1.19���1.30 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.08, PLE 0.09; AME���AME 0.04, AME���ALE 0.03, PME���PME 0.08, PME���PLE 0.05, ALE���PLE 0.01; MOA 0.19 long, front width 0.13, back width 0.27. Clypeus height 0.13. Leg measurements: I 3.81 (1.12, 1.31, 0.77, 0.60); II 3.53 (1.04, 1.17, 0.74, 0.58); III 3.21 (0.94, 1.01, 0.73, 0.54); IV 4.15 (1.20, 1.37, 0.93, 0.66). Other characters as in males. Epigyne with a pair of small copulatory openings centrally. Copulatory ducts bifurcated near copulatory openings, short branch connected with subspermathecae and with tuberculate expansion near base (green arrows and red arrows indicate in Figs 14D, 15E), long branch roundly coiled, finally connected with spermathecae, the latter sub-globular and located posterior-laterally. Subspermathecae oval, smaller than spermathecae and next to globular spermathecae. Fertilization ducts short, originating from posterior of spermathecae and pointing laterally (Figs 14B���D, 15D���E). Remarks. Both patellar apophyses of the male paratype are broken. Distribution. Known only from the type locality in Yunnan, China (Fig. 16). FIGURESA���E 15. Hahnia wangi sp. nov., holotype male (A���C) and paratype female (D���E). A���C Left palp (A ventral, B retrolateral, C dorsal). D Epigyne, ventral. E Vulva, dorsal. CO = copulatory openings; CD = copulatory duct; CF = cymbial furrow; Em = embolus; FD = fertilization duct; Ho = hood; MA = median apophysis; PA = patellar apophysis; RTA = retrolateral tibial apophysis; Sp = spermatheca; Ss = subspermatheca., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655, {"references":["Zhang, Z. S, Li, S. & Zheng, G. (2011) Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa, 2912, 1 - 27."]}

Hahnia subsaccata Huang & Zhang sp. nov. Figs 10���12, 16 Type material. CHINA: Yunnan Province: Holotype male , Nujiang Prefecture, Lushui County, Yaojiaping Town, 25��58.711' N, 98��42.211' E, elev. 2784 m, 3 March 2011, Z.X. Li & L.Y. Wang leg. Paratypes: 3 males, 8 females, same data as for holotype; 7 males, 5 females, Lushui County, Pianma Town, between Pianma and Fengxue Yakou, 25��58.923'��� 26��00.494' N, 98��39.553' ��� 98��42.058' E, elev. 2295���2977 m, 3 March 2011, Z.X. Li & L.Y. Wang leg.; 15 males, 7 females, Lushui County, Pianma Town, between Pianma and Fengxue Yakou, 25��58.155��� ���25��59.305��� N, 98��39.413������98��41.050��� E, elev. 2643���3101 m, 28 November 2011, Z.X. Li & L.Y. Wang leg.; 7 males, 6 females, Lushui County, Pianma Town, 26��00.294��� N, 98��39.554��� E, elev. 2318 m, 27 November 2011, Z.X. Li & L.Y. Wang leg.; 5 males, 3 females, Lushui County, Pianma Town, 25��59.852��� N, 98��39.727��� E, elev. 2334 m, 28 November 2011, Z.X. Li & L.Y. Wang leg.; 1 male, Dali City, Mt. Cangshan, Baolinqing, 25 June 2002, Z.Z. Yang leg.; 1 female, Baoshan City, Bawan Town, Dasheyao Yakou, 24��55.665 N', 98��45.112' E, elev. 2363 m, 13 February 2011, Z.X. Li & L.Y. Wang leg.; 1 female, Baoshan City, Longyang District, Nankang, Yakou, 24��49.586' N, 98��45.959' E, elev. 2174 m, 28 February 2011, Z.X. Li & L.Y. Wang leg.; 1 male, Tengchong County, Wuhe Township, Xiaodifang Village, 24��53.112' N, 98��45.346' E, elev. 2210 m, 27 February 2011, Z.X. Li & L.Y. Wang leg.; 13 males, 13 females, Lushui County, Yaojiaping Town, 25��58.747��� N, 98��42.474��� E, elev. 2740 m, 28 November 2011, Z.X. Li & L.Y. Wang leg.; 2 males, 2 females, Lushui County, between Pianma Town and Yaojiaping Town, 23 May 2015, C. Jin & M. Wang leg.; 5 males, 6 females, Baoshan City, Longyang District, Mangkuan Township, Baihualing, Mts. Gaoligong, East Slope, 25��17.710��� N, 98��46.604��� E, elev. 2345 m, 13 April 2014, X.K. Jiang, T. Lu & Z.G. Yang leg. (all SWUC). Etymology. The specific epithet refers to its similarity with Hahnia saccata Zhang, Li & Zheng, 2011; adjective. Diagnosis. The new species is similar to H. saccata in having a similar palpal patellar apophysis, tegular setae, embolus, median apophysis and cymbial furrow of male palp and the same location of copulatory openings of epigyne, but can be distinguished from this species by the relatively thin and long RTA and the longer row of tegular setae approaching the prolateral margin of bulb in ventral view of male palp (Figs 10E���G, 12A���C); epigyne with a small oval atrium, copulatory opening hidden inside atrium; copulatory ducts coiled around subspermathecae, the latter smaller than spermathecae. Description. Male (holotype). Total length 2.71. Prosoma 1.43 long, 1.08 wide; Opisthosoma 1.44 long, 0.99 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.07, PLE 0.09; AME���AME 0.03, AME���ALE 0.03, PME���PME 0.07, PME���PLE 0.07, ALE���PLE 0.02. MOA 0.19 long, front width 0.13, back width 0.21. Clypeus height 0.14. Chelicerae with distinct stridulatory files (Figs 13C���D), 3 promarginal, 6 retromarginal teeth. Leg measurements: I 4.22 (1.24, 1.44, 0.90, 0.63); II 3.99 (1.18, 1.33, 0.87, 0.62); III 3.64 (1.03, 1.14, 0.88, 0.59); IV 4.62 (1.29, 1.49, 1.12, 0.72). Leg formula: 4123. Spiracle long and transverse, the distance of spiracle to epigastric furrow as long as to spinnerets (Fig. 10B). Male palpal patella with a hooked apophysis originating retro-laterally. RTA sharply curved. Tegulum with a series of setae marginally near prolateral and submarginally near proximal margin. Embolus originating retrolateroproximally, tip of embolus lying near the membranous median apophysis. Cymbial furrow deep and shorter than half cymbium (Figs 10E���G, 12A���C). Female (paratypes). Total length 2.76���3.22. Prosoma 1.38���1.50 long, 1.05���1.13 wide; Opisthosoma 1.52���1.66 long, 1.12���1.24 wide. One of them total length 3.22. Prosoma 1.50 long, 1.13 wide; Opisthosoma 1.66 long, 1.24 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.07, PLE 0.08; AME���AME 0.04, AME���ALE 0.02, PME���PME 0.07, PME���PLE 0.05, ALE���PLE 0.03; MOA 0.15 long, front width 0.19, back width 0.22. Clypeus height 0.13. Leg measurements: I 3.82 (1.12, 1.30, 0.80, 0.59); II 3.55 (1.05, 1.17, 0.77, 0.56); III 3.32 (0.94, 1.05, 0.78, 0.54); IV 4.30 (1.22, 1.38, 1.03, 0.67). Leg formula: 4123. Epigyne with a small and transversal oval atrium near epigastric furrow. Two copulatory openings hidden inside the anterio-lateral margin of atrium. Copulatory ducts firstly extending anterio-laterally, then coiled mesally and bifurcated, one branch short, connected with the subspermatheca, the other branch long, connected with the ball-like, posterior-laterally located and widely separated spermatheca. Fertilization ducts short, originating from posterior spermathecae, pointing laterally (Figs 11C���D, 12D���E). Behind epigastric furrow a pair of oblique curved hoods (Figs 11B���C, 12D). Distribution. Known only from the type locality in Yunnan, China (Fig. 16)., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655, {"references":["Zhang, Z. S, Li, S. & Zheng, G. (2011) Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa, 2912, 1 - 27."]}

Hahnia dongi Huang & Zhang, sp. nov. Figs 3–4, 16 Type material. CHINA: Yunnan Province: Holotype male , Baoshan City, Longyang District, Nankang, Gaoligong Mountains Nature Park, 24°49'40.195" N, 98°46'9.228" E, elev. 2203m, 8 November 2016, Z.W. Dong leg. (SWUC). Etymology. This new species is dedicated to Mr. Zhi-Wei Dong, collector of the new species and good friend of the first author. Diagnosis. Male of the new species can be distinguished from other Hahnia species by the relatively short RTA, the short and protuberant LTA retrolateralo-proximally and the presence of some strong tibial setae (Figs 3E– G, 4A–C). Description. Male (holotype). Total length 2.20. Prosoma 1.07 long, 0.79 wide; Opisthosoma 1.26 long, 0.93 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.05, PLE 0.08; AME–AME 0.03, AME–ALE 0.02, PME–PME 0.08, PME–PLE 0.06, ALE–PLE 0.02. MOA 0.17 long, front width 0.15, back width 0.20. Clypeus height 0.14. Chelicerae with stridulatory files laterally (Figs 3C–D), 2 promarginal, 6 retromarginal teeth. Leg measurements: I 4.56 (1.30, 1.63, 1.06, 0.57); II 3.50 (0.99, 1.18, 0.82, 0.51); III 2.90 (0.88, 0.92, 0.69, 0.41); IV 3.35 (0.99, 1.09, 0.79, 0.48). Leg formula: 1243. Spiracle long and transverse, distance of spiracle to epigastric furrow 2.4 times as long as to spinnerets (Fig. 3B). Male palpal patella with a hook-like apophysis retrolatero-proximally, the patellar apophysis with a notch apically. Tibia covered with cluster of black setae at dorsal and retrolateral surface, with moderately short RTA extending dorsally and LTA proximally, tip of LTA covered with several black setae. Embolus originating retrolatero-proximally, tip of embolus lying near base of membranous median apophysis. Cymbial furrow deep and nearly as long as half of cymbium (Figs 3E–G, 4A–C). Female unknown. Distribution. Known only from the type locality in Yunnan, China (Fig. 16).

Hahnia musica Brignoli, 1978 Figs 6���9, 16 Hahnia musica Brignoli, 1978: 50, figs 25, 27���28 (holotype male, 1 male, 1 female paratypes, NHMB, not examined; 1 male and 1 female paratypes, MSNV, examined; all from Bhutan, 87 km Phuntsholing-Thimphu, 22.5.1972). Material examined. CHINA: Yunnan Province: 5 males, 1 female, Nujiang Prefecture, Gongshan County, Bingzhongluo Township, 27 May 2015, C. Jin & M. Wang leg.; 1 male, Baoshan City, Tengchong County, Mt. Laifeng, 16 May 2015, C. Jin leg.; 1 female, Baoshan City, Longyang District, Nankang, Xiaodifang Village, Mts. Gaoligong, 24��51.347��� N, 98��45.514��� E, elev. 2114 m, 17 September 2013, X.W. Meng & Z.X. Li leg.; 4 males, 5 females, Nujiang Prefecture, Fugong County, Lumadeng Township, 24 May 2015, C. Jin & M. Wang leg.; 18 males, 2 females, Lushui County, Pianma Town, near Three-level Power Station, 22 May 2015, C. Jin & M. Wang leg.; 6 females, Lushui County, Pianma Town, 26��00.750' N, 98��37.305' E, elev. 1812 m, 3 March 2011, Z.X. Li & L.Y. Wang leg.; 1 female, Fugong County, Shiyueliang Township, Lishadi Village, 27��15.320��� N, 98��53.627��� E, elev. 1448 m, 8 June 2011, Z.X. Li & G.C. Zhou leg.; 2 males, 3 females, Gongshan County, Dandang Park, 27��44.776��� N, 98��39.826��� E, elev. 1621 m, 7 June 2011, Z.X. Li & G.C. Zhou leg.; 1 female, Gongshan County, Dulongjiang Township, along Shuanglawadi, 27��45.047��� N, 98��37.939��� E, elev. 1605 m, 24 November 2011, Z.X. Li & L.Y. Wang leg.; 2 females, Lushui County, Pianma Town, Jinsuolang, 26��00.555��� N, 98��36.959��� E, elev. 1802 m, 10 June 2011, Z.X. Li & G.C. Zhou leg.; 4 females, Gongshan County, Bingzhongluo Township, 28��00.866��� N, 98��35.953��� E, elev. 1840 m, 6 June 2011, Z.X. Li & G.C. Zhou leg.; 1 female, Lanping County, Hexi Township, 26��51.893��� N, 99��22.633��� E, elev. 2132 m, 17 June 2011, Z.X. Li & G.C. Zhou leg. (all SWUC). Diagnosis. The species is similar to H. himalayaensis in having quite similar appearance and pattern of genitalia. But it can be distinguished by less tegular setae not extending distally so far, short median apophysis not exceeding retrolateral margin of cymbium in ventral view (Figs 6C���E, 8E���G, 9A���C); epigyne with shorter distance between the copulatory openings and the epigastric furrow than the diameter of copulatory openings and less convoluted copulatory ducts than the latter (Figs 7C���E, 8C���D, 9D���E). Redescription. Male. Total length 2.44���3.02. Prosoma 1.11���1.51 long, 0.81���1.13 wide; Opisthosoma 1.30��� 1.53 long, 0.80���0.95 wide. Spiracle long and transverse, the distance of spiracle to epigastric furrow as long as to spinnerets. Male palpal patella with a hooked apophysis retrolatero-proximally. RTA long, sharply curved clockwise, tip pointing ventrally. Male palpal tegulum having a series of setae forming ���C��� around centre of bulb. Embolus originating retrolaterally, almost roundly coiled. Cymbial furrow deep and nearly as long as half cymbium (Figs 6C���E, 8E���G, 9A���C). Female. Total length 2.67���2.89. Prosoma 1.20���1.24 long, 0.79���0.84 wide; Opisthosoma 1.45���1.64 long, 1.02��� 1.33 wide. Spiracle long and transverse, the distance of spiracle to epigastric furrow as long as to spinnerets (Fig. 7B). Epigyne with a quite shallow and transversal oval atrium near epigastric furrow, two closely connected copulatory openings located inside the atrium. Copulatory ducts moderately long, strongly coiled and bifurcated, one branch short and connected with the subspermathecae, the other branch long and roundly coiled, finally connected with posterio-laterally located, elliptic spermathecae. Fertilization ducts short, arising posteriorly of spermathecae, pointing laterally (Figs 7C���E, 8C���D, 9D���E). Behind the epigastric furrow, a pair of oblique curved hoods located (Figs 7B���C, 8C, 9D). Distribution. China (Yunnan) (Fig. 16), Bhutan (Dechhi Paka, Phuntsholing). Remarks. We confirmed this is the first description of H. musica from China after the examination of male and female paratypes (Figs 6A���G, 7A���E) of H. musica., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655, {"references":["Brignoli, P. M. (1978) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Araneae: Fam. Oonopidae, Agelenidae, Hahniidae und Mimetidae. Entomologica Basiliensis, 3, 31 - 56."]}

Hahnia himalayaensis Hu & Zhang, 1990 Figs 5, 16 Hahnia himalayaensis Hu & Zhang, 1990: 165, figs 1���7 (male, female); Song et al. 1999: 361, figs 210K���L, 211L���M (male, female); Hu 2001: 112, figs 31. 1���5 (male, female); Zhang et al. 2011: 12, figs 10A���C, 11A���D, 12A���I, 13A���G (male, female); Zhang et al. 2013: 346, figs 3A���E (male, female). Material examined. CHINA: Yunnan Province: 2 females, Baoshan City, Tengchong County, Jietou Town, Datang Village, Daheling, Ganjiao, 25��44.269' N, 98��41.707' E, elev. 1979 m, 23 February 2011, Z.X. Li & L.Y. Wang leg.; 1 female, Longling County, Longxin Town, Xiaoheishan Nature Reserve, 24��31.218' N, 98��50.826' E, elev. 1968 m, 17 February 2011, Z.X. Li & L.Y. Wang leg.; 5 females, Tengchong County, Jietou Town, Datang Village, Daheling, 25��37.492' N, 98��40.290' E, elev. 1748 m, 21 February 2011, Z.X. Li & L.Y. Wang leg.; 2 females, Tengchong County, Jietou Town, Datang Village, 25��37.492' N, 98��40.290' E, elev. 1748 m, 24 February 2011, Z.X. Li & L.Y. Wang leg.; 1 male, 3 females, Nujiang Prefecture, Gongshan County, Bingzhongluo Township, 28��00.866��� N, 98��35.953��� E, elev. 1840 m, 6 June 2011, Z.X. Li & G.C. Zhou leg.; 3 females, Lushui County, Pianma Town, 26��00.750' N, 98��37.305' E, elev. 1812 m, 3 March 2011, Z.X. Li & L.Y. Wang leg.; 1 female, Gongshan County, Bingzhongluo Township, Shimenguan, Sijitong, 28��02.301��� N, 98��35.448��� E, elev. 1478 m, 23 November 2011, Z.X. Li & L.Y. Wang leg. (all SWUC). Description. See the redescription of both sexes in Zhang et al. (2011). Distribution. China (Tibet, Yunnan, Guizhou), Vietnam (Ninh Binh)., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655, {"references":["Hu, J. L. & Zhang, F. P. (1990) A new species of the genus Hahnia from Xizang Autonomous Region, China (Araneae: Hahniidae). Acta Zootaxonomica Sinica, 15, 165 - 168.","Song, D. X., Zhu, M. S. & Chen, J. (1999) The Spiders of China. Hebei University of Science and Techology Publishing House, Shijiazhuang, 640 pp.","Hu, J. L. (2001) Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, Zhengzhou, 658 pp.","Zhang, Z. S, Li, S. & Zheng, G. (2011) Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa, 2912, 1 - 27.","Zhang, Z. S., Li, S. Q. & Pham, D. S. (2013) First description of comb-tailed spiders (Araneae: Hahniidae) from Vietnam. Zootaxa, 3613 (4), 343 - 356."]}

Hahnia himalayaensis Hu & Zhang, 1990 Figs 5, 16 Hahnia himalayaensis Hu & Zhang, 1990: 165, figs 1–7 (male, female); Song et al. 1999: 361, figs 210K–L, 211L–M (male, female); Hu 2001: 112, figs 31. 1–5 (male, female); Zhang et al. 2011: 12, figs 10A–C, 11A–D, 12A–I, 13A–G (male, female); Zhang et al. 2013: 346, figs 3A–E (male, female). Material examined. CHINA: Yunnan Province: 2 females, Baoshan City, Tengchong County, Jietou Town, Datang Village, Daheling, Ganjiao, 25°44.269' N, 98°41.707' E, elev. 1979 m, 23 February 2011, Z.X. Li & L.Y. Wang leg.; 1 female, Longling County, Longxin Town, Xiaoheishan Nature Reserve, 24°31.218' N, 98°50.826' E, elev. 1968 m, 17 February 2011, Z.X. Li & L.Y. Wang leg.; 5 females, Tengchong County, Jietou Town, Datang Village, Daheling, 25°37.492' N, 98°40.290' E, elev. 1748 m, 21 February 2011, Z.X. Li & L.Y. Wang leg.; 2 females, Tengchong County, Jietou Town, Datang Village, 25°37.492' N, 98°40.290' E, elev. 1748 m, 24 February 2011, Z.X. Li & L.Y. Wang leg.; 1 male, 3 females, Nujiang Prefecture, Gongshan County, Bingzhongluo Township, 28°00.866′ N, 98°35.953′ E, elev. 1840 m, 6 June 2011, Z.X. Li & G.C. Zhou leg.; 3 females, Lushui County, Pianma Town, 26°00.750' N, 98°37.305' E, elev. 1812 m, 3 March 2011, Z.X. Li & L.Y. Wang leg.; 1 female, Gongshan County, Bingzhongluo Township, Shimenguan, Sijitong, 28°02.301′ N, 98°35.448′ E, elev. 1478 m, 23 November 2011, Z.X. Li & L.Y. Wang leg. (all SWUC). Description. See the redescription of both sexes in Zhang et al. (2011). Distribution. China (Tibet, Yunnan, Guizhou), Vietnam (Ninh Binh).

Hahnia biapophysis Huang & Zhang, sp. nov. Figs 1���2, 16 Type material. CHINA: Yunnan Province: Holotype male , Nujiang Prefecture, Lushui County, Pianma Port, 26��00.745��� N, 98��37.792��� E, elev. 1987 m, 8 September 2013, X.W. Meng & Z.X. Li leg. (SWUC). Etymology. The specific name is combined by ��� bi -��� and ��� apophysis ���, referring the male palpal tibia with two apophyses (Figs 1F���G, 2B���C); term in apposition. Diagnosis. Males of the new species are similar to those of H. zhui Zhang & Chen, 2015 in having the spinelike LTA near dorsum of tibia and tegular setae proximally, but can be distinguished from the latter by the patellar apophysis much more strongly curved and pointed, RTA relatively short, not close to LTA in retrolateral view, tegulum unmodified, tegular setae located submarginally near prolatero-proximal margin (Figs 1E���G, 2A���C). Description. Male (holotype). Total length 2.78. Prosoma 1.28 long, 0.94 wide; Opisthosoma 1.42 long, 0.91 wide. Eye sizes and interdistances: AME 0.06, ALE 0.10, PME 0.08, PLE 0.09; AME���AME 0.02, AME���ALE 0.01, PME���PME 0.07, PME���PLE 0.03, ALE���PLE 0.01. MOA 0.18 long, front width 0.12, back width 0.25. Clypeus height 0.15. Chelicerae with stridulatory files laterally (Figs 1C���D), 3 promarginal, 5 retromarginal teeth. Leg measurements: I 4.13 (1.17, 1.39, 0.86, 0.71); II 3.89 (1.08, 1.29, 0.82, 0.70); III 3.51 (0.94, 1.11, 0.83, 0.63); IV 4.50 (1.22, 1.46, 1.05, 0.78). Leg formula: 4123. Spiracle long and transverse, distance of spiracle to epigastric furrow as long as to spinnerets (Fig. 1B). Male palpal patella with a hook-like apophysis. Tibia with a moderately short RTA extending dorsally and then proximally. Embolus originating retrolatero-proximally, tip of embolus situated near the base of membranous median apophysis, slightly extending beyond apex of cymbial furrow. Cymbial furrow deep and slightly longer than half of cymbium (Figs 1E���G, 2A���C). Female unknown. Distribution. Known only from the type locality in Yunnan, China (Fig. 16)., Published as part of Huang, Gui-Qiang, Zhang, Zhi-Sheng & Liu, Ying-Hong, 2017, Review of the comb-tailed spider genus Hahnia C. L. Koch 1841 (Hahniidae) from Gaoligong Mountains in Yunnan, China in Zootaxa 4344 (3), DOI: 10.11646/zootaxa.4344.3.2, http://zenodo.org/record/1043655, {"references":["Zhang, Z. S & Chen, H. M (2015) Hahnia zhui sp. nov. (Hahniidae), a special species with a unique shape of tegulum. Acta Arachnologica Sinica, 24 (2), 70 - 75."]}

Hahnia nava (Blackwall, 1841) Material examined. Site 13 (b: 1 ♀). Distribution. Europe, Russia, Japan, Kyrgyzstan, Tajikistan (Komnenov, 2014). Chorotype. Eurasian., Published as part of Marjan Komnenov, Eva Pitta, Konstantina Zografou & Maria Chatzaki, 2016, Discovering the still unexplored arachnofauna of the National Park of Dadia-Lefkimi-Soufli, NE Greece: a taxonomic review with description of new species, pp. 1-66 in Zootaxa 4096 (1) on page 35, DOI: 10.11646/zootaxa.4096.1.1, http://zenodo.org/record/263078, {"references":["Komnenov, M. (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana, 2, 1 - 267."]}

Hahnia nava (Blackwall, 1841) LITERATURE. — Isaia 2005; Negro et al. 2009; [Rossi & Bosio 2012]. CHOROTYPE. — SIE. MACROHABITAT. — Coniferous forests., Published as part of Paschetta, Mauro, Christille, Claretta, Marguerettaz, Fabio & Isaia, Marco, 2016, Regional catalogue of the spiders (Arachnida, Araneae) of Aosta Valley (NW Italy), pp. 49-125 in Zoosystema 38 (1) on page 72, DOI: 10.5252/z2016n1a3, http://zenodo.org/record/4578074, {"references":["ISAIA M. 2005. - Check list of spiders (Arachnids, Araneae) of Aosta Valley (Italy) with a new record for Italian fauna. Revue valdotaine d'Histoire naturelle 59: 25 - 43.","NEGRO M., ISAIA M., PALESTRINI C. & ROLANDO A. 2009. - The impact of forest ski-pistes on diversity of ground-dwelling arthropods and small mammals in the Alps. Biodiversity and Conservation 18 (11): 2799 - 2821. http: // dx. doi. org / 10.1007 / s 10531 - 009 - 9608 - 4","ROSSI G. & BOSIO F. 2012. - Elenco delle specie di ragni (Arachnida, Araneae) note per la Valle d ' Aosta. Revue valdotaine d'Histoire naturelle 66: 45 - 88."]}

Hahnia subcorticicola sp. nov. Figs 1 A���E, 2 A���I, 4 Type material. Holotype male, CHINA: Chongqing: Mt. Jinyun, 29 �� 50.1 ��� N, 106 �� 23.7 ��� E, Guankou, 5 April 2010, Z.S. Zhang, Z.X. Li, K.Y. Xu & L.Y. Wang leg. (SWUC). Paratypes: 12 males, 72 females (SWUC), with same data as holotype; 15 males, 69 females, with same location as holotype, 28 March 2010, Z.X. Li, K.Y. Xu & L.Y. Wang leg. (SWUC); 6 males, 93 females, with same location as holotype, 28 April 2010, Z.X. Li, K.Y. Xu & L.Y. Wang leg. (SWUC); 4 males, 9 females, with same location as holotype, 26 October 2011, Z.X. Li, Z. Li & L.Y. Wang leg. (SWUC); 7 males, 5 females, with same location as holotype, 12 November 2011, M.X. Liu & D. Wang leg. (SWUC); 11 males, 13 females, with same location as holotype, 29 November 2011, M.X. Liu & D. Wang leg. (SWUC); 15 males, 13 females, with same location as holotype, 27 November 2010, L.Y. Wang leg. (SWUC); 12 males, 19 females, Mt. Jinyun, Banzigou, 4 December 2010, Z. Li & Z.X. Li leg. (SWUC); 16 males, 23 females, Mt. Jinyun, Caijiagou, 12 December 2010, Z.S. Zhang leg. (SWUC); 71 males, 206 females, with same location as holotype, 30 March 2011, Z.X. Li, Z. Li & L.Y. Wang leg. (SWUC); 1 male, 6 females, with same location as holotype, 24 October 2011, L.Y. Wang, D. Wang & M.X. Liu leg. (SWUC); 3 females, with same location as holotype, 1 December 2010, Z.X. Li, Z. Li & L.Y. Wang leg. (SWUC); 3 females, with same location as holotype, 25 May 2011, Z. Li & L.Y. Wang leg. (SWUC); 1 female, Mt. Jinyun, Yunhe Village, 15 June 2010, L.Y. Wang leg. (SWUC); 1 female, Mt. Jinyun, Yunhe Village, 15 November 2010 (SWUC); 1 male, 15 females, Mt. Jinyun, Caijiagou, 10 April 2013, X.K. Jiang, X.W. Meng, T. Lu & J. Yang leg. (SWUC); 2 females, Mt. Jinyun, Caijiagou, 29 October 2011, L.Y. Wang, D. Wang & M.X. Liu leg. (SWUC); 1 male, 1 female, Mt. Jinyun, Fuxin Temple, 17 April, 2012, Z. Li & L.Y. Wang leg. (SWUC); 1 male, Mt. Jinyun, Beiwenquan, 9 December 2012, X.K. Jiang & M.X. Liu leg. (SWUC); Hubei: 9 females, Luotian County, Tiantangzhai, 30 �� 5.9 ��� N, 115 ��44.0��� E, 25���28 September 2009 (HUBU); 2 females, Tiantangzhai, 20���26 July 2009 (HUBU); 2 males, 5 males, Badong County, Yeshanguan Town, 30 �� 36.8 ��� N, 110 �� 19.8 ��� E, hillside behind the Sanxia wine factory, 24 March 2014, L.Y. Wang leg. (SWUC); Anhui: 123 males, 186 females, Xiuning County, Huangshan City, Mt. Qiyun, 29 �� 48.9 ��� N, 118 �� 2.7 ��� E, from Yuehua Street to the highest point within Qiyun, 19 April 2011, Z.X. Li & L.Y. Wang leg. (SWUC); 17 males, 49 females, Mt. Qiyun, Fanglazhai, 19 April 2011, Z.X. Li & L.Y. Wang leg. (SWUC); 1 female, Mt. Qiyun, 6 September 2011, L.Y. Wang & M.X. Liu leg. (SWUC); 10 males, 12 females, 18 April 2011, Z.X. Li & L.Y. Wang leg. (SWUC); 1 male, 2 females, Mt. Qiyun, 26 September 2011, L.Y. Wang & M.X. Liu leg. (SWUC); 3 males, 5 females, Mt. Qiyun, 27 September 2011, L.Y. Wang & M.X. Liu leg. (SWUC); 4 males, 14 females, Huangshan scenic area, 20 April 2011, Z.X. Li & L.Y. Wang leg. (SWUC). Material examined for comparison. Hahnia corticicola: 16 females, 17 males, CHINA: Sichuan: Jiuzhaigou County, Longkang Town (SWUC); 1 female, 1 male, Shaanxi: Zhouzhi County, Houzhenzi (SWUC). Etymology. The specific epithet refers to the species��� similarity to H. corticicola B��senberg & Strand 1906. Diagnosis. The new species is similar to H. corticicola (Figs 3 A���I) in having similarly shaped patellar apophysis, RTA, median apophysis, tegulum, and long and deep cymbial furrow as well as a similarly positioned copulatory opening, but it can be distinguished by the pointed apex of the male palpal cymbium, the equally long cymbium and cymbial furrow, the presence of a small tegular apophysis near the base of the embolus, the extremely slender embolus with its tip embedded in a pocket (Figs 1 A���C, 2 E���G), the protruding median part of the posterior margin of the copulatory opening, the long, multi-coiled copulatory ducts and the distinct subspermathecae of the epigyne (1 D���E, 2 H���I). Description. Male holotype (Figs 2 A, C���D) total length 1.77. Prosoma 0.78 long, 0.68 wide; opisthosoma 1.09 long, 0.81 wide. Carapace pear-shaped with dark markings in radial furrows and cervical groove. Fovea longitudinal, brown. Cephalic area elevated. Eight eyes with distinct black rims. Eye sizes and interdistances: AME 0.03, ALE 0.06, PME 0.05, PLE 0.05; AME���AME 0.03, AME���ALE 0.01, PME���PME 0.05, PME���PLE 0.02, ALE���PLE 0.01. MOA 0.12 long, front width 0.09, back width 0.14. Clypeus height 0.12. Chelicerae yellowish with three promarginal and three retromarginal teeth. Gnathocoxae of equal length and width. Labium wider than long. Sternum cordiform with dark marginal markings. Legs yellow with some spines. Leg measurements: I 2.25 (0.66, 0.75, 0.48, 0.36); II 2.11 (0.63, 0.68, 0.44, 0.36); III 1.95 (0.55, 0.62, 0.44, 0.34); IV 2.41 (0.67, 0.79, 0.56, 0.39). Leg formula: 4123. Opisthosoma oval, greyish black with a light quadrate and five light chevrons dorsally. Tracheal spiracle equidistant between the epigastric furrow and the base of spinnerets. Male palp (Figs 1 A���C, 2 E���G) with a hook-like, relatively thin and long patellar apophysis. Distal part of RTA filiform extending dorsally and then proximally. Cymbium with a very deep, long and semi-transparent retrolateral furrow. Embolus originating retrolaterally, clockwise curved along the cymbial margin, then anticlockwise curved inside the cymbial furrow. End of embolus embedded in a transparent pocket. Median apophysis membranous, located near the base of embolus. One female paratype (collected with holotype, Fig. 2 B) total length 2.03. Prosoma 0.81 long, 0.59 wide; opisthosoma 1.26 long, 0.95 wide. Eye sizes and interdistances: AME 0.04, ALE 0.05, PME 0.05, PLE 0.05; AME��� AME 0.02, AME���ALE 0.01, PME���PME 0.05, PME���PLE 0.03, ALE���PLE 0.01. MOA 0.13 long, front width 0.09, back width 0.13. Clypeus height 0.09. Leg measurements: I 1.96 (0.59, 0.66, 0.39, 0.32); II 1.83 (0.57, 0.59, 0.37, 0.30); III 1.72 (0.54, 0.54, 0.37, 0.27); IV 2.28 (0.66, 0.75, 0.50, 0.37). Leg formula: 4123. Epigyne (Figs 1 D���E, 2 H���I) with a pair of small copulatory openings medio-posteriorly with posterior margins visible. Copulatory ducts quite long and strongly convoluted as three coils. Subspermathecae located anteriorly, visible in the gap of copulatory duct loops. Spermathecae located centrally in the loops of copulatory ducts. Fertilization ducts extending posteriorly and curving medially. Distribution. China (Chongqing, Hubei, Anhui) (Fig. 4). Remark. The new species was collected in the leaf litter of an old-growth forest area, specifically, along the Yangtze River basin, close to the latitude 30 �� North., Published as part of Liu, Na, Huang, Gui-Qiang & Zhang, Zhi-Sheng, 2015, A new species of the genus Hahnia (Araneae: Hahniidae) from South China, pp. 295-300 in Zootaxa 3994 (2) on pages 295-299, DOI: 10.11646/zootaxa.3994.2.10, http://zenodo.org/record/240324

Genus Hahnia C.L. Koch, 1841, Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Pham, Dinh-Sac, 2013, First description of comb-tailed spiders (Araneae: Hahniidae) from Vietnam, pp. 343-356 in Zootaxa 3613 (4) on page 346, DOI: 10.11646/zootaxa.3613.4.2, http://zenodo.org/record/218582

Hahnia zhejiangensis Song & Zheng, 1982 Figures 8 A���C, 9 A���I, 10 A���G, 11 A���E Hahnia zhejiangensis Song & Zheng, 1982: 81, figs 1���4 (female); Yin & Wang 1983: 141, figs 6���7 (male); Hu 1984: 210, figs 222.1 ��� 3 (female); Song 1987: 203, fig. 162 (male, female); Feng 1990: 144, figs 119.1 ��� 5 (male, female); Chen & Zhang 1991: 196, figs 196.1 ��� 5 (male, female); Song et al. 1999: 362, figs 211 D���E, 212 G���H (male, female); Chen et al. 2003: 26, figs 1 A���F, 2 A���B (male, female); Zhang et al. 2013: 350, figs 7 A���E (male, female). Hahnia thorntoni Brignoli, 1982: 346, figs 13���14 (male paratype only) (misspelled as H. thortoni); Song 1987: 201, fig. 160 (male) (misspelled as H. thortoni); Song et al. 1999: 361, figs 212 A���B (male). (misidentified) Hahnia yueluensis Yin & Wang, 1983: 143, figs 11���12 (male paratype only); Song 1987: 202, figs 161 C���D (male); Chen & Gao 1990: 121, figs 150 b���c (male); Song et al. 1999: 362, figs 212 E���F (male). (misidentified) Material examined. CHINA: ZHEJIANG: 1 female, Lin���an County, 19.X. 1974 (JLU); HUBEI: 1 male, Luotian County, Tiantangzhai, 25���28.IX. (HUBU); HUNAN: 1 male, 1 female, Changsha, Mt. Yuelu, 26.X. 2007, Z.Z. Yang (SWUC); CHONGQING: 1 male, Beibei District, Campus of Southwest University, 18.X. 2008, Y.F. Zhang (SWUC); 1 male, Shapingba District, Mt. Gele, 10.XI. 2008, Z.S. Zhang (SWUC); 2 females, Mt. Jinyun, Guankou, 28.III. 2010, Z.X. Li & L.Y. Wang (SWUC); 1 female, Guankou, 28.IV. 2010, Z.X. Li, L.Y. Wang & K.Y. Xu (SWUC); 1 female, front area of Mt. Jinyun, 20.III. 2010, Z.X. Li, L.Y. Wang & K.Y. Xu (SWUC); 1 male, 11 females, Mt. Jinyun, Banzigou, 17.IV. 2010, Z.X. Li, L.Y. Wang & K.Y. Xu (SWUC); 5 females, Banzigou, 24.IV. 2010, Z.X. Li, L.Y. Wang & K.Y. Xu (SWUC); 1 male, Banzigou, 1.XI. 2010, Z.X. Li & L.Y. Wang (SWUC); 1 female, Banzigou, 12.IV. 2008, Z.S. Zhang (BMNH); 1 female, Mt. Jinyun, Caijiagou, 29.IV. 2010, Z.X. Li & L.Y. Wang (SWUC); SICHUAN: 7 male, 1 female, Changning County, Zhuhai Town, Sansonghu Village, 28.��� 30.IX. 2008, Z.S. Zhang & L.Y. Wang (SWUC 6 males, 1 female, BMNH 1 male); 1 male, 1 female, Langzhong County, 8.IX. 2009, Z.S. Zhang (SWUC); JIANGXI: 1 male, 1 female, Wuzhou City, Donghua Park, 9.III. 2011, G.C. Zhou (SWUC); HONG KONG: Male paratype of H. thorntoni, 6.IV. 1965, I.W.B. Thornton (BMNH); VIETNAM (87 males, 72 females, see Zhang et al. 2013). Diagnosis. Males of this species can easily be distinguished from other Hahnia species by the relatively short RTA with an abruptly thin tip, the presence of a tibial process extending dorsally, and the long cymbial furrow as long as the cymbium (Figs 8 B���C, 9 H���I, 11 B���C). Females can be distinguished by the strongly sclerotized posterior margin of the copulatory openings and the strongly curved copulatory ducts (Figs 10 D���G, 11 D���E). Description. Male palp (Figs 8 A���C, 9 G���I, 11 A���C) with a small, thin and hooked patellar apophysis, tibia with a relatively short RTA, with tip abruptly thin, and a large process extending dorsally. Embolus originating proximally on retrolateral margin of tegulum, curved along the bulb, its tip staying in the cymbial furrow near the embolic base. Median apophysis membranous, almost transparent. Cymbial furrow as long as cymbium. Epigyne (Figs 10 D���G, 11 D���E) with a pair of copulatory openings anteriorly, with posterior margin strongly sclerotized, like two triangles. Copulatory ducts thin and long, strongly coiled. Subspermathecae small, anteriorly located. Spermathecae elliptic, more than twice as large as subspermathecae. For a more detailed description see Song & Zheng (1982) and Song (1987). Male habitus, eyes, cheliceral teeth and stridulatory files shown in Figures 9 A���F. Male palp shown in Figures 8 A���C, 9 G���I, 11 A���C. Female habitus shown in Figures 10 A���C. Epigyne and vulva shown in Figures 10 D���G, 11 D���E. Distribution. China (Zhejiang, Anhui, Jiangxi, Hunan, Hubei, Chongqing, Sichuan, Hong Kong, Taiwan); Vietnam. Remark. The type material of H. zhejiangensis (female holotype from Sanmen City, Zhejiang Province of China) is unavailable for this research, but the strongly sclerotized copulatory openings of epigyne and the particular tibial apophysis of male palp are enough to confirm this species and separated it from other Hahnia species., Published as part of Zhang, Zhi-Sheng & Zhang, Yao-Guang, 2013, Synonymy and misidentification of three Hahnia species (Araneae: Hahniidae) from China, pp. 521-533 in Zootaxa 3682 (4) on pages 529-532, DOI: 10.11646/zootaxa.3682.4.3, http://zenodo.org/record/249254, {"references":["Song, D. X. & Zheng, S. X. (1982) Two new species of Hahnia (Araneae: Hahniidae) from Zhejiang Province, China. Sinozoologica, 2, 81 - 84.","Yin, C. M. & Wang, J. F. (1983) A preliminary study on the Chinese spiders of the family Hahniidae (Arachnida: Araneida). Acta Zootaxonomica Sinica, 8, 141 - 145.","Hu, J. L. (1984) The Chinese Spiders Collected from the Fields and the Forests. Tianjin Press of Science and Techniques, Tianjin, 482 pp.","Song, D. X. (1987) Spiders from Agricultural Regions of China (Arachnida: Araneae). Agriculture Publishing House, Beijing, 376 pp.","Feng, Z. Q. (1990) Spiders of China in colour. Hunan Science and Technology Publishing House, Changsha, 256 pp.","Song, D. X., Zhu, M. S. & Chen, J. (1999) The Spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.","Chen, S. H., Wang, Y. & Chen, S. C. (2003) A newly recorded spider of the family Hahniidae (Arachnida, Araneae) from Taiwan. Journal of Taiwan normal University: Mathematics, Science & Technolgy, 48, 25 - 30.","Zhang, Z. S., Li, S. & Pham, D. S. (2013) First description of comb-tailed spiders (Araneae: Hahniidae) from Vietnam. Zootaxa, 3613 (4), 343 - 356. http: // dx. doi. org / 10.11646 / zootaxa. 3613.4.2","Brignoli, P. M. (1982) On a few spiders from China (Araneae). Bulletin of the British Arachnological Society, 5, 344 - 351.","Chen, Z. F. & Zhang, Z. H. (1990) Fauna of Zhejiang: Araneida. Zhejiang Science and Technology Publishing House, 356 pp."]}

Hahnia maginii Brignoli, 1977 Figures 1 A���F, 2 A���B Hahnia maginii Brignoli, 1977: 78, fig. 47 (female). Misidentifications: Hahnia maginii: Yu et al. 1984: 11 (female); Song 1987: 199, fig. 157 (female); Song et al. 1999: 361, figs 210 Q���R (female). Refers to H. thorntoni. Material examined. Female holotype from Toscana, Italy, 1.VII. 1973, F. Magini (MSNV). Diagnosis. The female of the species is similar to H. rossii Brignoli, 1977 from Italy (Brignoli 1977: 79, fig. 46), H. jocquei Bosmans, 1982 from Malawi (Bosmans 1982: 177, figs 2 ��� 3) and three Chinese species: H. ovata Song & Zheng, 1982 (Song & Zheng 1982: 81, figs 5���8), H. submaginii Zhang et al., 2011 (Zhang et al. 2011: 19, figs 18 A���F) and H. thorntoni Brignoli, 1982 (Figs 7 A���D) in having a distinctive epigynal atrium, but H. maginii can be distinguished from H. rossii, H. submaginii and H. thorntoni by the relatively small atrium (Figs 1 E, 2 A) and from H. jocquei and H. ovata by the short copulatory ducts (Figs 1 F, 2 B). Description. Female holotype (Figs 1 A���B) total length 2.70. AME smallest (Fig. 1 C). Both cheliceral margins with 4 teeth (Fig. 1 D). Leg formula: 4123. Epigyne (Figs 1 E���F, 2 A���B) with a distinct, shallow atrium, with anterior margin distinct. A pair of copulatory openings located in the atrium centrally and epigynal hoods visible behind the epigastric furrow. Copulatory ducts relatively short. Subspermathecae anteriorly located and their diameter the same length as half of that of the spermathecae. More details of the holotype can be found in Brignoli (1977). Male unknown. Distribution. Italy., Published as part of Zhang, Zhi-Sheng & Zhang, Yao-Guang, 2013, Synonymy and misidentification of three Hahnia species (Araneae: Hahniidae) from China, pp. 521-533 in Zootaxa 3682 (4) on pages 522-524, DOI: 10.11646/zootaxa.3682.4.3, http://zenodo.org/record/249254, {"references":["Brignoli, P. M. (1977) Ragni d'Italia XXVII. Nuovi dati su Agelenidae, Argyronetidae, Hahniidae, Oxyopidae e Pisauridae cavernicoli ed epigei (Araneae). Quaderni del Museo di Speleologia \" V. Rivera \", 4, 3 - 117.","Yu, S. Y., Shen, Q. Z. & Song, D. X. (1984) A new record of the genus Hahnia from China. Acta zootaxonomica sinica, 9 (1), 11.","Song, D. X. (1987) Spiders from Agricultural Regions of China (Arachnida: Araneae). Agriculture Publishing House, Beijing, 376 pp.","Song, D. X., Zhu, M. S. & Chen, J. (1999) The Spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.","Bosmans, R. (1982) Two new species of Hahnia from Malawi (Araneae: Hahniidae). Revue de zoologie africaine, 96, 174 - 178.","Song, D. X. & Zheng, S. X. (1982) Two new species of Hahnia (Araneae: Hahniidae) from Zhejiang Province, China. Sinozoologica, 2, 81 - 84.","Zhang, Z. S., Li, S. & Zheng, G. (2011) Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa, 2912, 1 - 27.","Brignoli, P. M. (1982) On a few spiders from China (Araneae). Bulletin of the British Arachnological Society, 5, 344 - 351."]}

Hahnia himalayaensis Hu & Zhang, 1990 Figs 3 A���E Hahnia himalayaensis Hu & Zhang, 1990: 165, figs 1���7; Song et al., 1999: 361, figs 210 K���L, 211 L���M; Hu, 2001: 112, figs 31.1 ��� 5; Zhang et al., 2011: 12, figs 10 A���C, 11 A���D, 12 A���I, 13 A���G. Material examined (58 males, 9 females). VIETNAM, NINH BINH PROVINCE, Cuc Phuong National Park: 10 males, 2 females, shrub-land, pitfall traps, 1���30.III. 2008; 4 males, shrub-land, pitfall traps, 1���30.V. 2007; 11 males, 1 female, natural forest, pitfall traps, 1���30. IV. 2007; 15 males, natural forest, pitfall traps, 1���30.VI. 2007; 7 males, 4 females, natural forest, pitfall traps, 1���30.VIII. 2007; 7 males, 1 female, natural forest, pitfall traps, 1���30.X. 2007; 4 males, 1 female, natural forest, pitfall traps, 1���30. XII. 2007. Description. See Zhang et al. (2011) for both sexes. Male palp shown in Figs 3 A���C. Epigyne in Figs 3 D���E. Distribution. Vietnam (Ninh Binh), China (Tibet and Yunnan)., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Pham, Dinh-Sac, 2013, First description of comb-tailed spiders (Araneae: Hahniidae) from Vietnam, pp. 343-356 in Zootaxa 3613 (4) on page 346, DOI: 10.11646/zootaxa.3613.4.2, http://zenodo.org/record/218582

Hahnia zhejiangensis Song & Zheng, 1982 Figs 7 A���E Hahnia zhejiangensis Song & Zheng, 1982: 81, figs 1���4; Yin & Wang, 1983: 141, figs 6���7; Hu, 1984: 210, figs 222.1 ��� 3; Song, 1987: 203, fig. 162; Feng, 1990: 144, figs 119.1 ��� 5; Chen & Zhang, 1991: 196, figs 196.1 ��� 5; Song et al., 1999: 362, figs 211 D���E, 212 G���H; Chen et al., 2003: 26, figs 1 A���F, 2 A���B. Material examined (87 males, 72 females). VIETNAM, NINH BINH PROVINCE: 1 male, Cuc Phuong National Park, disturbed forest, pitfall traps, 1���30.VI. 2007; 1 male, Cuc Phuong, disturbed forest, pitfall traps, 1���30.XII. 2007; 5 males, Cuc Phuong, natural forest, pitfall traps, 1���30. IV. 2007; 11 males, Cuc Phuong, natural forest, pitfall traps, 1���30.VI. 2007; 24 males, Cuc Phuong, natural forest, pitfall traps, 1���30.VII. 2007; 1 male and 26 females, Cuc Phuong, natural forest, pitfall traps, 1���30. XII. 2007; 2 males and 3 females, Cuc Phuong, natural forest, pitfall traps, 1���30.XII. 2007; 4 males, Cuc Phuong, shrub-land, pitfall traps, 1���30.III. 2008; 7 males and 1 female, Cuc Phuong, shrub-land, pitfall traps, 1���30.IV. 2007; 7 males and 2 females, Cuc Phuong, shrub-land, pitfall traps, 1���30.V. 2007; 1 female, Cuc Phuong, shrub-land, pitfall traps, 1���30.VI. 2007; 1 male and 13 females, Cuc Phuong, shrub-land, pitfall traps, 1���30.VII. 2007; 4 males, Cuc Phuong, shrub-land, pitfall traps, 1���30.VIII. 2007; 3 males and 1 female, Cuc Phuong, shrub-land, pitfall traps, 1���30.X. 2007; VINH PHUC PROVINCE: 1 male, Tam Dao national park, shrub-land, pitfall traps, 1���30.VI. 2007; HAI PHONG PROVINCE: 2 males and 2 females, Cat Ba National Park, disturbed forest, pitfall traps, 1���30.I. 2007; 1 male, Cat Ba, disturbed forest, pitfall traps, 1���29. II. 2008; 5 males and 3 females, Cat Ba, disturbed forest, pitfall traps, 1���30.V. 2007; 5 females, Cat Ba, disturbed forest, pitfall traps, 1���30.X. 2007; 1 female, Cat Ba, disturbed forest, 20 �� 47.277 ��N, 107 ��00.368��E, 117m, sieving, 19.III. 2008; 1 female, Cat Ba, disturbed forest, 20 �� 48.080 ��N, 107 ��00.212��E, 116m, sieving, 24.IV. 2007; 1 female, Cat Ba, disturbed forest, 20 �� 46.586 ��N, 107 ��01.029��E, 95m, sieving, 25.V. 2007; 1 female, same data as above, 24.VI. 2007; 1 female, same data as above, 20 �� 48.249 ��N, 107 ��00.016��E, 80m, sieving, 20.XII. 2007; 1 male, Cat Ba, shrub-land, pitfall traps, 1���29.II. 2008; 2 males and 1 female, Cat Ba, pitfall traps, 1���30.III. 2008; 1 male, same data as above, 1���30.VI. 2007; 1 female, same data as above, 1���30.VIII. 2007; 2 males and 2 females, same data as above, 1���30.XI. 2007; 1 male, Cat Ba, natural forest, pitfall traps, 1���30.IX. 2007; 1 female, same data as above, 20 �� 49.143 ��N, 107 ��02.193��E, 104m, sieving, 19.XII. 2007; 1 female, same data as above, 20 �� 48.486 ��N, 107 ��01.488��E, 137m, sieving, 19.XII. 2007; 2 males and 4 females, Cat Ba, Acacia plantation, pitfall traps, 1���30. III. 2008. Description. See Chen & Zhang (1991). Male palp shown in Figs 7 A���C, epigyne in Figs 7 D���E. Distribution. Vietnam (Ninh Binh, Hai Phong), China (Zhejiang, Hunan, Hong Kong, etc.), Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Pham, Dinh-Sac, 2013, First description of comb-tailed spiders (Araneae: Hahniidae) from Vietnam, pp. 343-356 in Zootaxa 3613 (4) on pages 350-351, DOI: 10.11646/zootaxa.3613.4.2, http://zenodo.org/record/218582

Hahnia rimiformis sp. nov. Figs 4 A���F, 5 A���G, 6 A���G Type material. Holotype: male, VIETNAM, NINH BINH PROVINCE, Cuc Phuong National Park, natural forest, pitfall traps, 1���30.X. 2007. Paratypes (57 males, 27 females): 26 males, 1 female, same data as holotype; 1 male, 2 females, same data as holotype but: 1���30.VI. 2007; 1 female, same data as holotype but: 1���30.VII. 2007; 28 males, same data as holotype but: 1���30.VIII. 2007; 2 males, 23 females, same data as holotype but: 1���30.XI. 2007. Etymology. The specific name is taken from the Latin word ��� rimiformis ���, meaning split-like and referring to the shape of the epigynal atrium, adjective. Diagnosis. The new species is similar to H. himalayaensis (Figs 3 A���E) in having tegular setae and long copulatory ducts, but differs from the latter by the smaller number of setae and the long cymbial furrow; and a splitlike epigynal atrium, the small subspermathecae far away from spermathecae (Figs 4 A���B, 4 D, 4 F). Description. Male holotype (Figs 5 A���D): Total length 2.25. Prosoma 1.13 long, 0.83 wide; opisthosoma 1.25 long, 0.82 wide. Eye sizes and interdistances: AME 0.05, ALE 0.09, PME 0.07, PLE 0.08; AME���AME 0.03, AME���ALE 0.01, PME���PME 0.05, PME���PLE 0.03, ALE���PLE 0.02. MOA 0.17 long, front width 0.12, back width 0.19. Clypeus height 0.12. Leg measurements: I 3.61 (1.02, 1.20, 0.80, 0.59); II 3.35 (0.94, 1.07, 0.75, 0.59); III 3.11 (0.85, 0.95, 0.78, 0.53); IV 3.78 (1.01, 1.24, 0.96, 0.67). Leg formula: 4123. Chelicerae with 3 promarginal and 5 retromarginal teeth. Male palp (Figs 4 A���C, 5 E���G) with a sheet-like, hooked patellar apophysis and a long and curved RTA. Another protuberance present on the dorsal margin of retrolateral side of tibia. Embolus long, retrolateralproximally originated and its tip lying in the cymbial furrow. Median apophysis membranous, originating retrolaterally. Cymbial furrow long and deep. One of female paratypes (Figs 6 A���B) total length 2.34. Prosoma 1.67 long, 0.81 wide; opisthosoma 1.29 long, 0.94 wide. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.07, PLE 0.09; AME���AME 0.04, AME���ALE 0.02, PME���PME 0.06, PME���PLE 0.05, ALE���PLE 0.03. MOA 0.18 long, front width 0.13, back width 0.21. Clypeus height 0.12. Leg measurements: I 3.15 (0.94, 1.11, 0.56, 0.54); II 2.99 (0.89, 0.97, 0.62, 0.51); III 2.77 (0.77, 0.86, 0.63, 0.51); IV 3.68 (1.05, 1.18, 0.86, 0.59). Chelicerae with 3 promarginal and 5���6 retromarginal teeth (some other female paratypes with 3���4 promarginal and 5���6 retromarginal ones). Other characters similar to the male holotype. Epigyne with an elevated, split-like atrium (Figs 4 D, 6 C���D) near the epigastric furrow. Atrium pit-like if degraded backwards (Figs 4 E, 6 E���F). Copulatory ducts slender and strongly curved. Subspermathece slightly inflated and located anteriorly. Spermathecae located posteriorly, widely separated from each other (Figs 4 F, 6 G). A pair of epigynal hoods can be located behind of the epigastric furrow, their distance wider than twice of the width of copulatory openings (Fig. 4 D). Distribution. Vietnam (Ninh Binh)., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Pham, Dinh-Sac, 2013, First description of comb-tailed spiders (Araneae: Hahniidae) from Vietnam, pp. 343-356 in Zootaxa 3613 (4) on pages 347-350, DOI: 10.11646/zootaxa.3613.4.2, http://zenodo.org/record/218582

Hahnia thorntoni Brignoli, 1982 Figures 3 A���E, 4 A���I, 5 A���C, 6 A���G, 7 A���D Hahnia thorntoni Brignoli, 1982: 346 (male holotype only; male paratype misidentified) (misspelled as H. thortoni). Hahnia maginii: Yu et al. 1984: 11 (female); Song 1987: 199, fig. 157 (female); Song et al. 1999: 361, figs 210 Q���R (female). (Misidentified) Hahnia flagellifera Zhu et al., 1989: 148, figs 1���6 (male, female); Chen & Gao 1990: 119, figs 149 a���f (male, female); Song et al. 1999: 361, figs 210 I���J, 211 J���K (male, female); Zhang et al. 2011: 9, figs 6 A���C, 7 A���D, 8 A���I, 9 A���G (male, female); J��ger & Praxaysombath 2011: 28. (New synonymy) Hahnia yueluensis Yin & Wang, 1983: 143, figs 9���10 (female); Song 1987: 202, figs 161 A���B (female); Chen & Gao 1990: 121, fig. 150 a (female); Song et al. 1999: 362, figs 211 B���C (female). (New synonymy) Material examined. CHINA: HONG KONG: Male holotype, 6.IV. 1965, I.W.B. Thornton (BMNH); SICHUAN: 1 male and 1 female (80 ��� 73, might be paratypes of H. flagellifera, not indicated), from cornfield, Guangyuan City, X.E. Chen (JLU); 6 males, Mt. Emei, 22.IX.��� 3.X. 1975 (JLU); 12 males, Changning County, Zhuhai Town, Sansonghu Village, 28.��� 30.IX. 2008, Z.S. Zhang & L.Y. Wang (SWUC 11 males; BMNH 1 male); 3 males, Changning County, Meidong Town, Gaojian Village, 29.IX. 2008, R.Y. Zuo & Y.F. Zhang (SWUC); CHONGQING: 1 female, Beibei District, Jindaoxia Scenic Spot, 23.XI. 2008, Z.S. Zhang (SWUC); 1 female, Mt. Jinyun, Banzigou, 12.IV. 2008, Z.S. Zhang (SWUC); 3 females, Baizigou, 17.IV. 2010, Z.X. Li, L.Y. Wang, Z. Li & K.Y. Xu (SWUC); 1 female, Jiangjin District, Simianshan Town, 6.III. 2009, Z.S. Zhang & Q.Y. Feng (SWUC); 1 female, Simian Mountain Natural Reserve, Dawuopu, 11.IX. 2011, Z.X. LI & L.Y. WANG (BMNH); YUNNAN: 3 males, 63 females, Mengla County, Menglun Town, X. 2006 ��� IV. 2007, G. Zheng (IZCAS); 1 female, Mengla County, Menglun Town, 17.X. 2009, G. Tang & Z.Y. Yao (IZCAS); 1 male, Menglun Town, 27.XI. 2009, G. Tang & Z.Y. Yao (IZCAS); 1 female, Menglun, 20.XI. 2009, G. Tang & Z.Y. Yao (IZCAS); JIANGXI: 1 male, Wuzhou, Campus of East China Institute of Technology, 30.X. 2010, G.C. Zhou (SWUC). Diagnosis. Females of the species are similar to H. jocquei (Bosmans 1982: 177, figs 2 ��� 3), H. maginii (Figs 1 A���F, 2 A���B), H. ovata (Song & Zheng 1982: 81, figs 5���8), H. rossii (Brignoli 1977: 79, fig. 46), and H. submaginii (Zhang et al. 2011: 19, figs 18 A���F) in having an epigynal atrium and small, anteriorly located subspermathecae, but differ from the others by the large, oval atrium with distinct margin (Figs 6 D, 6 E, 7 A, 7 C). Males of the species are similar to H. submaginii (Zhang et al. 2011: 19, figs 17 A���C), but are distinguished by the simple patellar apophysis (Figs 3 C���E, 4 G���I, 5 A���C), rather than a bifurcate one. Description. See Brignoli (1982: 346) and the description of H. flagellifera in Zhang et al. (2011: 19). Male holotype���s habitus, stridulatory files and palp are shown in Figures 3 A, 3 B, 3 C���E. Photos of specimens from Menglun in Yunnan are shown in Figures 4 A���I, 5 A���C, 6 A���G, 7 A���D. Distribution. China (Hong Kong, Yunan, Jiangxi, Hunan, Chongqing, Sichuan), Laos., Published as part of Zhang, Zhi-Sheng & Zhang, Yao-Guang, 2013, Synonymy and misidentification of three Hahnia species (Araneae: Hahniidae) from China, pp. 521-533 in Zootaxa 3682 (4) on pages 525-526, DOI: 10.11646/zootaxa.3682.4.3, http://zenodo.org/record/249254, {"references":["Brignoli, P. M. (1982) On a few spiders from China (Araneae). Bulletin of the British Arachnological Society, 5, 344 - 351.","Yu, S. Y., Shen, Q. Z. & Song, D. X. (1984) A new record of the genus Hahnia from China. Acta zootaxonomica sinica, 9 (1), 11.","Song, D. X. (1987) Spiders from Agricultural Regions of China (Arachnida: Araneae). Agriculture Publishing House, Beijing, 376 pp.","Song, D. X., Zhu, M. S. & Chen, J. (1999) The Spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.","Zhu, C. D., Chen, X. E. & Sha, Y. H. (1989) A new species of genus Hahnia from Sichuan Province, China (Araneae: Hahniidae). Journal of Norman Bethune University of medical science, 15, 148 - 150.","Chen, Z. F. & Zhang, Z. H. (1990) Fauna of Zhejiang: Araneida. Zhejiang Science and Technology Publishing House, 356 pp.","Zhang, Z. S., Li, S. & Zheng, G. (2011) Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae). Zootaxa, 2912, 1 - 27.","Jager, P. & Praxaysombath, B. (2011) Spiders from Laos with forty-three new records and first results from the provinces Bolikhamsay and Champasak (Arachnida: Araneae). Acta Arachnologica, 60 (1), 9 - 31. http: // dx. doi. org / 10.2476 / asjaa. 60.9","Yin, C. M. & Wang, J. F. (1983) A preliminary study on the Chinese spiders of the family Hahniidae (Arachnida: Araneida). Acta Zootaxonomica Sinica, 8, 141 - 145.","Bosmans, R. (1982) Two new species of Hahnia from Malawi (Araneae: Hahniidae). Revue de zoologie africaine, 96, 174 - 178.","Song, D. X. & Zheng, S. X. (1982) Two new species of Hahnia (Araneae: Hahniidae) from Zhejiang Province, China. Sinozoologica, 2, 81 - 84.","Brignoli, P. M. (1977) Ragni d'Italia XXVII. Nuovi dati su Agelenidae, Argyronetidae, Hahniidae, Oxyopidae e Pisauridae cavernicoli ed epigei (Araneae). Quaderni del Museo di Speleologia \" V. Rivera \", 4, 3 - 117."]}

Hahnia flagellifera Zhu, Chen & Sha, 1989 Figs 6���9 Hahnia flagellifera Zhu et al. 1989: 148, figs 1 ��� 6 (Holotype female, allotype male and paratype female from rice field, Emei, Sichuan Province, China, August 1980, deposited in Jilin Unversity, not examined); Chen & Gao 1990: 119, figs 149 a ��� f. Song et al. 1999: 361, figs 210 I ��� J, 211 J ��� K. Material examined. 2 males: China, Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Town (active searching), 5 ��� 12 October 2006, G. Zheng leg.; 1 male: Menglun Town (active searching), 5 ��� 12 November 2006, G. Zheng leg.; 63 females: Menglun Town (active searching), October 2006 ��� April 2007, G. Zheng leg. Diagnosis. Females of this species are similar to those of H. maginii Brignoli, 1977 (female holotype from Italy, deposited in Museo Civico di Storia Naturale, Verona, Italy, examined) in having a single atrium and small sub-spermathecae, but differing from the latter by the large epigynal atrium (Figs 7 A, 7 C, 9 D���E) and the widely separated sub-spermathecae (Figs 7 B, 7 D, 9 F���G). This species is also similar to H. submaginii sp. nov. (Figs 17��� 20) in having similar RTA, embolus, median apophysis and cymbial furrow, but males can be distinguished by the single palpal patellar apophysis (Figs 6 A���C, 8 G���I) (bifid in H. submaginii), females by the anterior margin of the epigynal atrium arc-shaped or somewhat protruding in dorsal view and the relatively small sub-spermathecae with wide distance from each other (Figs 7 B, 7 D, 9 F���G). Redescription. Male. Total length 2.40 ��� 2.60. One of them (Figs 8 A���F) total length 2.49. Prosoma 1.28 long, 0.95 wide; opisthosoma 1.32 long, 0.93 wide. Chelicerae with conspicuous stridulatory files on the lateral surface, 3 promarginal and 6 ��� 8 retromarginal teeth. Leg formula: 1423. Opisthosoma with some black spots and 4 pale chevron-like marings dorsally. Spiracle located centrally. Male palpal patellar apophysis recurved and hook-like. RTA long. Embolus originating retro-basally and spiraling clockwise along the margin of tegulum nearly 360 �� (left palp). Median apophysis membranous, originating retrolaterally. Cymbium furrow deep, which length is almost equal to half of cymbial length (Figs 6 A���C, 8 G���I). Female. Total length 2.80 ��� 3.20. One female (Figs 9 A���C): total length 2.87. Prosoma 1.27 long, 0.97 wide; opisthosoma 1.70 long, 1.37 wide. Leg formula: 4123. Chelicerae with 6 retromarginal teeth. Stridulatory file absent. Other characters as in male. Epigyne with large, posteriorly extended atrium. Copulatory ducts relatively short, bifurcate in front of atrium. Sub-spermathecae ball-like or tuberculate. Spermathecae located posteriorly, separated by large atrium. Fertilization ducts small, pointing laterally. A pair of hoods found posterior of epigastric furrow, distance of which is wider than width of spermathecae (Figs 7 A���D, 9 G���I). Distribution. China (Sichuan, Yunnan). Biology. All male and most of female specimens of H. flagellifera were collected from Rubber-Tea and Paramichelia baillonii -plantations near the Menglun Nature Reserve. Remark. There are two pairs of receptacula located on the vulva dorsally, which were named basal and second receptacula by Forster (1970) or primary and secondary receptacula by Bosmans and Thijs (1980). Here the basal or primary receptacula connecting with fertilization ducts are named spermathecae, which might be homologue to those of other spider groups. And the second or secondary receptacula are named sub-spermathecae, which are present only in a few other groups, such as Clubiona (Clubionidae)., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Zheng, Guo, 2011, Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae), pp. 1-27 in Zootaxa 2912 on pages 8-12, DOI: 10.5281/zenodo.277820, {"references":["Zhu, C. D., Chen, X. E. & Sha, Y. H. (1989) A new species of genus Hahnia from Sichuan Province, China (Araneae: Hahniidae). Journal of Norman Bethune University of Medical Science, 15, 148 - 150.","Chen, X. E. & Gao, J. C. (1990) The Sichuan Farmland Spiders in China. Sichuan Scientific and Technical Publishing House, Chengdu, 226 pp.","Song, D. X., Zhu, M. S. & Chen, J. (1999) The Spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.","Forster, R. R. (1970) The spiders of New Zealand. Part III. Otago Museum Bulletin, 3, 1 - 184.","Bosmans, R. & Thijs, E. (1980) Spiders of the family Hahniidae from Mount Kenya: Scientific report of the Belgian Mt. Kenya Bio-Expedition 1975, no. 20. Revue de Zoologie Africaine, 94, 559 - 569."]}

Hahnia submaginii sp. nov. Figs 17���20 Type material. Holotype: Male, China, Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Nature Reserve (active searching), 5 ��� 12 November 2006, G. Zheng leg. Paratypes: 2 males, 24 females, same locality as for holotype (active searching), September 2006 ��� May 2007, G. Zheng leg. Etymology. The specific name is taken from the similarity to H. maginii Brignoli, 1977; noun in apposition. Diagnosis. Females of the new species similar to those of H. maginii Brignoli, 1977 in having an epigynal atrium, spherical spermathecae and sub-spermathecae and relatively short copulatory ducts, but differ from the latter by the large epigynal atrium (Figs 18 A, 18 C, 18 E, 20 E), the small distance between anterior atrial margin and the anterior margin of copulatory ducts (Figs 18 A, 18 C, 18 E, 20 E) and the relatively thin copulatory ducts (Figs 18 B, 18 D, 18 F, 20 F). The new species is also similar to H. flagellifera Zhu, Chen & Sha, 1989 (Figs 15���17), but can be distinguished by the bifurcate palpal patellar apophysis (Figs 17 A���C, 19 E���G), the distinct margin of epigynal copulatory openings (Figs 18 A, 18 C, 18 E, 20 E) and the procurved anterior margin of atrium in dorsal view (Figs 18 B, 18 D, 18 F, 20 F). Descripton. Male. Total length 1.90 ��� 2.50. Holotype (Figs 19 A���D): Total length 2.45. Prosoma 1.28 long, 0.99 wide; opisthosoma 1.29 long, 0.89 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.07, PLE 0.07; AME���AME 0.04, AME���ALE 0.03, PME���PME 0.07, PME���PLE 0.06, ALE���PLE 0.02. MOA 0.19 long, front width 0.15, back width 0.21. Clypeus height 0.13. Chelicerae with distinct lateral condyle, stridulatory files laterally, 3 or 4 promarginal and 5 or 6 retromarginal teeth. Leg measurements: I 4.19 (1.23, 1.44, 0.93, 0.59); II 3.82 (1.10, 1.30, 0.88, 0.54); III 3.16 (0.91, 1.00, 0.78, 0.47); IV 3.83 (1.09, 1.25, 0.94, 0.55). Leg formula: 1423. Opisthosoma greyish, with 3 chevron-like markings dorsally. Spiracle located centrally. Male palpal patella with a bifurcate apophysis retrolaterally. One part pointing proximally, hook-like and the other pointing distally. RTA long and curved dorsally. Embolus originating retro-basally and curved around tegulum. Median apophysis membranous. Cymbial furrow deep, which length about half of cymbial length (Figs 17 A��� C, 19 E���G). Female. Total length 2.30 ��� 2.80. One female paratype (Figs 20 A���D): Total length 2.50. Prosoma 1.05 long, 0.75 wide; opisthosoma 1.55 long, 1.25 wide. Eye sizes and interdistances: AME 0.05, ALE 0.08, PME 0.08, PLE 0.08; AME���AME 0.03, AME���ALE 0.02, PME���PME 0.05, PME���PLE 0.05, ALE���PLE 0.01. Clypeus height 0.08. Leg measurements: I 2.80 (0.85, 0.95, 0.60, 0.40); II 2.55 (0.75, 0.85, 0.55, 0.40); III 2.25 (0.70, 0.75, 0.55, 0.35); IV 3.00 (0.90, 1.00, 0.70, 0.40). Leg formula: 4123. Other characters as in male. Epigynal atrium large, with distinct anterior margin. Copulatory openings located in atrium, with visible anterior margin, close to the anterior margin of atrium. Sub-spermathecae of similar size as spermathecae. Spermathecae located posteriorly, near the big atrium laterally. Fertilization ducts located posterior to spermathecae (Figs 18 A���F, 20 E���F). Distribution. Known only from Xishuangbanna rainforest, Yunnan Province, China., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Zheng, Guo, 2011, Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae), pp. 1-27 in Zootaxa 2912 on pages 19-20, DOI: 10.5281/zenodo.277820, {"references":["Zhu, C. D., Chen, X. E. & Sha, Y. H. (1989) A new species of genus Hahnia from Sichuan Province, China (Araneae: Hahniidae). Journal of Norman Bethune University of Medical Science, 15, 148 - 150."]}

Hahnia saccata sp. nov. Figs 14���16 Type material. Holotype: Male, China, Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Nature Reserve, ground of secondary tropical montane evergreen forest (active searching), 5 ��� 12 September 2006, G. Zheng leg. Paratypes: 16 males, same locality as for holotype (active searching and pitfall traps), from July to September 2006, G. Zheng leg.; 66 females, same locality as for holotype (active searching and pitfall traps), September 2006 ��� May 2007, G. Zheng leg. Etymology. The specific name is taken from the Latin adjective ��� saccatus ���, meaning saccate, shaped like a pouch or sac and referring its saccate sub-spermathecae; adjective. Diagnosis. The new species is similar to H. himalayaensis Hu & Zhang, 1990 (Figs 10���13), H. cinerea Emerton, 1890 and H. arizonica Chamberlin & Ivie, 1942 in having tegular setea on male palp and two relative small copulatory openings of epigyne, but can be distinguished from these species by the stout RTA (Figs 14 B���C, 15 G���H), the fewer tegular setae (Figs 14 A, 15 F) of male palp and the posteriorly located copulatory openings (Figs 14 D, 16 E���F), the very short copulatory ducts between openings and bifurcation points and the spherical subspermathecae with relatively wide distance from each other (Figs 14 E, 16 G). Description. Male. Total length 2.30 ��� 2.60. Holotype (Figs 15 A���E): Total length 2.46. Prosoma 1.32 long, 1.02 wide; opisthosoma 1.20 long, 0.84 wide. Eye sizes and interdistances: AME 0.07, ALE 0.10, PME 0.09, PLE 0.11; AME���AME 0.03, AME���ALE 0.02, PME���PME 0.06, PME���PLE 0.04, ALE���PLE 0.02. MOA 0.22 long, front width 0.16, back width 0.24. Clypeus height 0.13. Chelicerae with distinct lateral condyle, stridulatory files laterally and 3 promarginal, 5 ��� 6 retromarginal teeth. Leg measurements: I 4.45 (1.27, 1.47, 1.01, 0.70); II 4.06 (1.19, 1.32, 0.91, 0.64); III 3.58 (1.02, 1.10, 0.87, 0.59); IV 4.46 (1.24, 1.40, 1.10, 0.72). Leg formula: 4123. Opisthosoma with 4 chevron-like markings dorsally. Spiracle located centrally. Male palpal patellar apophysis hooked, originating retro-basally. RTA sharply curved backward, tip pointed. Embolus originating retro-basally, curved clockwise around the tegulum (left palp) and ending near the membranous median apophysis. Set of setae located on tegulum prolaterally. Cymbial furrow deep, its length slightly shorter than half of cymbial length (Figs 14 A���C, 15 F���H). Female. Total length 2.40 ��� 3.00. One female paratype (Figs 16 A���D): Total length 2.55. Prosoma 1.35 long, 0.95 wide; opisthosoma 1.40 long, 1.15 wide. Eye sizes and interdistances: AME 0.05, ALE 0.10, PME 0.08, PLE 0.10; AME���AME 0.05, ALE���ALE 0.03, PME���PME 0.08, PME���PLE 0.05, ALE���PLE 0.02. Clypeus height 0.13. Chelicerae with 6 retromarginal teeth (7 or 8 teeh rarely found). Leg measurements: I 3.75 (1.10, 1.30, 0.80, 0.55); II 3.40 (1.00, 1.15, 0.75, 0.50); III 3.15 (0.90, 1.05, 0.75, 0.45); IV 4.20 (1.25, 1.35, 1.00, 0.60). Leg formula: 4123. Other characters as in male. Epigyne with two small copulatory openings, located in shallow, small atrium and close to epigastric furrow. Copulatory ducts bifurcate near the openings. One branch connecting with sub-spermathecae relatively thick and the other slender. Sub-spermathecae located anteriorly. Spermathecae elliptic, near the posterior margin of epigyne. Fertilization ducts originating from the spermathecae posteriorly and pointing laterally (Figs 14 D���E, 16 E���G). Distribution. Known only from Xishuangbanna rainforest, Yunnan Province, China., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Zheng, Guo, 2011, Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae), pp. 1-27 in Zootaxa 2912 on pages 16-19, DOI: 10.5281/zenodo.277820, {"references":["Hu, J. L. & Zhang, F. P. (1990) A new species of the genus Hahnia from Xizang Autonomous Region, China (Araneae: Hahniidae). Acta Zootaxonomica Sinica, 15, 165 - 168."]}

Hahnia himalayaensis Hu & Zhang, 1990 Figs 10–13 Hahnia himalayaensis Hu & Zhang 1990: 165, figs 1 – 7 (Holotype female, allotype male and 2 female paratypes, 3 male paratypes from Linzhi County, Tibet, China, 9 September 1987, Juping Zhang leg., deposited in Shandong University, China, not examined); Song et al. 1999: 361, figs 210 K – L, 211 L – M; Hu 2001: 112, figs 31.1 – 5. Material examined. 85 males, 140 females, China, Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Town (active searching and pitfall traps), August 2006 – June 2007, G. Zheng leg. Diagnosis. The species is similar to H. barbata Bosmans, 1992 from Sulawesi (Bosmans 1992: 84, figs 1–6) in having a row of long tegular setae, a pair of copulatory openings and coiled copulatory ducts, but can be distinguished by the relatively long and anteriorly curved row of tegular setae, the tiny RTA, the bigger patellar apophysis of the male palp and the relatively long copulatory ducts of the epigyne. It is also similar to H. saccata sp. nov. (Figs 14–16), H. cinerea Emerton, 1890 (Opell & Beatty 1976: 423, figs 78–83) and H. arizonica Chamberlin & Ivie, 1942 (Opell & Beatty 1976: 424, figs 84–88) in having a submarginal prolateral tegular row of setae on the male palp and a pair of copulatory openings, but distinguished from others by the conspicuously long tegular setae row of male palp (Figs 10 A–B, 12 G–H) and two copulatory openings located centrally, close to each other (Figs 11 A, 11 C, 13 D–E), copulatory ducts long and strongly twisted (Figs 11 B, 11 D, 13 F–G). Redescription. Male. Total length 2.35 – 2.60. One male (Figs 12 A–F): total length 2.50. Prosoma 1.30 long, 0.90 wide; opisthosoma 1.30 long, 0.90 wide. Chelicerae with 3 promarginal and 5 – 6 retromarginal teeth (4 or 7 retromarginal teeth can be rarely found). Stridulatory files present on the lateral surface of chelicerae. Leg formula: 4123. Opisthosoma with 4 chevron-like markings dorsally. Spiracle located centrally. Male palpal patella with a hooked apophysis retro-basally. RTA long, sharply curved and pointing ventrally. Embolus originating retro-basally, extending around the tegulum, ending at the base of membranous median apophysis. Length of deep cymbial furrow is about half of cymbial length (Figs 10 A–C, 12 G–I). Female. Total length 2.40 – 3.10. One female (Figs 13 A–C): total length 2.55. Prosoma 1.25 long, 0.85 wide; opisthosoma 1.45 long, 1.15 wide. Chelicerae without stridulatory files, with 6 retromarginal teeth. AME smallest. Leg formula: 4123. Epigyne with a pair of small copulatory openings centrally. Copulatory ducts long, bifurcated and strongly swirled. Sub-spermathecae is nearly as large as spermathecae and close to the latter anteriorly. Fertilization ducts small, posterior to spermathecae, pointing laterally (Figs 11 A–D, 13 D–G). Distribution. China (Tibet, Yunnan) Biology. This is a dominant species on the ground of Xishuangbanna, whether in tropical rainforest or in plantations. Nearly all mature male individuals were collected in November and December, while females collected almost all time of a year. Apparently, H. himalayaensis completes only one generation per year and the male die as soon as the copulation is finished. Female lives apparently longer after copulation.

Hahnia himalayaensis Hu & Zhang, 1990 Figs 10���13 Hahnia himalayaensis Hu & Zhang 1990: 165, figs 1 ��� 7 (Holotype female, allotype male and 2 female paratypes, 3 male paratypes from Linzhi County, Tibet, China, 9 September 1987, Juping Zhang leg., deposited in Shandong University, China, not examined); Song et al. 1999: 361, figs 210 K ��� L, 211 L ��� M; Hu 2001: 112, figs 31.1 ��� 5. Material examined. 85 males, 140 females, China, Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Town (active searching and pitfall traps), August 2006 ��� June 2007, G. Zheng leg. Diagnosis. The species is similar to H. barbata Bosmans, 1992 from Sulawesi (Bosmans 1992: 84, figs 1���6) in having a row of long tegular setae, a pair of copulatory openings and coiled copulatory ducts, but can be distinguished by the relatively long and anteriorly curved row of tegular setae, the tiny RTA, the bigger patellar apophysis of the male palp and the relatively long copulatory ducts of the epigyne. It is also similar to H. saccata sp. nov. (Figs 14���16), H. cinerea Emerton, 1890 (Opell & Beatty 1976: 423, figs 78���83) and H. arizonica Chamberlin & Ivie, 1942 (Opell & Beatty 1976: 424, figs 84���88) in having a submarginal prolateral tegular row of setae on the male palp and a pair of copulatory openings, but distinguished from others by the conspicuously long tegular setae row of male palp (Figs 10 A���B, 12 G���H) and two copulatory openings located centrally, close to each other (Figs 11 A, 11 C, 13 D���E), copulatory ducts long and strongly twisted (Figs 11 B, 11 D, 13 F���G). Redescription. Male. Total length 2.35 ��� 2.60. One male (Figs 12 A���F): total length 2.50. Prosoma 1.30 long, 0.90 wide; opisthosoma 1.30 long, 0.90 wide. Chelicerae with 3 promarginal and 5 ��� 6 retromarginal teeth (4 or 7 retromarginal teeth can be rarely found). Stridulatory files present on the lateral surface of chelicerae. Leg formula: 4123. Opisthosoma with 4 chevron-like markings dorsally. Spiracle located centrally. Male palpal patella with a hooked apophysis retro-basally. RTA long, sharply curved and pointing ventrally. Embolus originating retro-basally, extending around the tegulum, ending at the base of membranous median apophysis. Length of deep cymbial furrow is about half of cymbial length (Figs 10 A���C, 12 G���I). Female. Total length 2.40 ��� 3.10. One female (Figs 13 A���C): total length 2.55. Prosoma 1.25 long, 0.85 wide; opisthosoma 1.45 long, 1.15 wide. Chelicerae without stridulatory files, with 6 retromarginal teeth. AME smallest. Leg formula: 4123. Epigyne with a pair of small copulatory openings centrally. Copulatory ducts long, bifurcated and strongly swirled. Sub-spermathecae is nearly as large as spermathecae and close to the latter anteriorly. Fertilization ducts small, posterior to spermathecae, pointing laterally (Figs 11 A���D, 13 D���G). Distribution. China (Tibet, Yunnan) Biology. This is a dominant species on the ground of Xishuangbanna, whether in tropical rainforest or in plantations. Nearly all mature male individuals were collected in November and December, while females collected almost all time of a year. Apparently, H. himalayaensis completes only one generation per year and the male die as soon as the copulation is finished. Female lives apparently longer after copulation., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Zheng, Guo, 2011, Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae), pp. 1-27 in Zootaxa 2912 on pages 12-16, DOI: 10.5281/zenodo.277820, {"references":["Hu, J. L. & Zhang, F. P. (1990) A new species of the genus Hahnia from Xizang Autonomous Region, China (Araneae: Hahniidae). Acta Zootaxonomica Sinica, 15, 165 - 168.","Song, D. X., Zhu, M. S. & Chen, J. (1999) The Spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.","Hu, J. L. (2001) Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, Zhengzhou, 658 pp.","Opell, B. D. & Beatty, J. A. (1976) The Nearctic Hahniidae (Arachnida: Araneae). Bulletin of the Museum of Comparative Zoology at Harvard University, 147, 393 - 433."]}

Hahnia submaginii sp. nov. Figs 17–20 Type material. Holotype: Male, China, Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Nature Reserve (active searching), 5 – 12 November 2006, G. Zheng leg. Paratypes: 2 males, 24 females, same locality as for holotype (active searching), September 2006 – May 2007, G. Zheng leg. Etymology. The specific name is taken from the similarity to H. maginii Brignoli, 1977; noun in apposition. Diagnosis. Females of the new species similar to those of H. maginii Brignoli, 1977 in having an epigynal atrium, spherical spermathecae and sub-spermathecae and relatively short copulatory ducts, but differ from the latter by the large epigynal atrium (Figs 18 A, 18 C, 18 E, 20 E), the small distance between anterior atrial margin and the anterior margin of copulatory ducts (Figs 18 A, 18 C, 18 E, 20 E) and the relatively thin copulatory ducts (Figs 18 B, 18 D, 18 F, 20 F). The new species is also similar to H. flagellifera Zhu, Chen & Sha, 1989 (Figs 15–17), but can be distinguished by the bifurcate palpal patellar apophysis (Figs 17 A–C, 19 E–G), the distinct margin of epigynal copulatory openings (Figs 18 A, 18 C, 18 E, 20 E) and the procurved anterior margin of atrium in dorsal view (Figs 18 B, 18 D, 18 F, 20 F). Descripton. Male. Total length 1.90 – 2.50. Holotype (Figs 19 A–D): Total length 2.45. Prosoma 1.28 long, 0.99 wide; opisthosoma 1.29 long, 0.89 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.07, PLE 0.07; AME–AME 0.04, AME–ALE 0.03, PME–PME 0.07, PME–PLE 0.06, ALE–PLE 0.02. MOA 0.19 long, front width 0.15, back width 0.21. Clypeus height 0.13. Chelicerae with distinct lateral condyle, stridulatory files laterally, 3 or 4 promarginal and 5 or 6 retromarginal teeth. Leg measurements: I 4.19 (1.23, 1.44, 0.93, 0.59); II 3.82 (1.10, 1.30, 0.88, 0.54); III 3.16 (0.91, 1.00, 0.78, 0.47); IV 3.83 (1.09, 1.25, 0.94, 0.55). Leg formula: 1423. Opisthosoma greyish, with 3 chevron-like markings dorsally. Spiracle located centrally. Male palpal patella with a bifurcate apophysis retrolaterally. One part pointing proximally, hook-like and the other pointing distally. RTA long and curved dorsally. Embolus originating retro-basally and curved around tegulum. Median apophysis membranous. Cymbial furrow deep, which length about half of cymbial length (Figs 17 A– C, 19 E–G). Female. Total length 2.30 – 2.80. One female paratype (Figs 20 A–D): Total length 2.50. Prosoma 1.05 long, 0.75 wide; opisthosoma 1.55 long, 1.25 wide. Eye sizes and interdistances: AME 0.05, ALE 0.08, PME 0.08, PLE 0.08; AME–AME 0.03, AME–ALE 0.02, PME–PME 0.05, PME–PLE 0.05, ALE–PLE 0.01. Clypeus height 0.08. Leg measurements: I 2.80 (0.85, 0.95, 0.60, 0.40); II 2.55 (0.75, 0.85, 0.55, 0.40); III 2.25 (0.70, 0.75, 0.55, 0.35); IV 3.00 (0.90, 1.00, 0.70, 0.40). Leg formula: 4123. Other characters as in male. Epigynal atrium large, with distinct anterior margin. Copulatory openings located in atrium, with visible anterior margin, close to the anterior margin of atrium. Sub-spermathecae of similar size as spermathecae. Spermathecae located posteriorly, near the big atrium laterally. Fertilization ducts located posterior to spermathecae (Figs 18 A–F, 20 E–F). Distribution. Known only from Xishuangbanna rainforest, Yunnan Province, China.

Hahnia senaria sp. nov. Figs 21���23 Type material. Holotype: Male, China, Yunan Province, Xishuangbanna Dai Autonomous Prefecture, Mengla County, Menglun Nature Reserve (active searching), 4 ��� 11 April 2007, G. Zheng leg. Paratypes: 216 males, 298 females, same locality as for holotype (active searching and pitfall traps), August 2006 ��� June 2007, G. Zheng leg. Etymology. The specific name is taken from the Latin word ��� senarius ���, meaning consisting of six each, referring to the presence of six eyes of this new species; adjective. Diagnosis. The new species can be easily distinguished from other Hahnia species by the very small body with 6 eyes and shallow body color (Figs 22 A���C, 23 A���C), the midway located spiracle, the absence of stridulatory files on chelicerae, the short embolus and very short cymbial furrow of male palp (Figs 21 A���D, 22 D���F), the spherical sub-spermathecae of female epigyne (Figs 21 F, 23 E). Description. Male (holotype, Figs 22 A���C). Total length 0.95. Prosoma 0.46 long, 0.39 wide; opisthosoma 0.52 long, 0.40 wide. Dorsal prosoma yellowish, with dark cephalic area. Fovea longitudinal and brown. The highest point of prosoma located between the eye area and fovea. AME reduced. Posterior eye row distinctively procurved. ALE nearly black, others white. Eye pigments obvious. ALE equal to PLE (0.03), PME smallest (0.02); ALE���ALE 0.04, PME���PME 0.03, PME���PLE 0.01, ALE���PLE 0.01. ���Clypeus height��� (from the anterior margin of ALE to the margin of clypeus) 0.05. Chelicerae yellow, with indistinct lateral condyle and stridulatory file. Promarginal teeth absent, 2 small retromarginal teeth. Labium, gnathocoxas, sternum and legs yellowish. Leg measurements: I 1.32 (0.41, 0.46, 0.24, 0.21); II 1.23 (0.37, 0.43, 0.23, 0.20); III 1.13 (0.37, 0.34, 0.23, 0.19); IV 1.37 (0.40, 0.45, 0.29, 0.23). Leg formula: 4123. Opisthosoma oval and greyish, wish indistinct markings. Spiracle located centrally. Male palpal patella with a small hooked apophysis basally. RTA originating ventro- proximally, curved dorsally. Embolus originating proximally, curved around tegulum, semicircular. Conductor absent. Median apophysis membranous, originating distal-retrolaterally. Cymbial furrow short, its length about 1 / 4 of cymbial length (Figs 21 A���D, 22 D���F). Female. (one paratype, Figs 23 A���C): Total length 1.15. Prosoma 0.43 long, 0.37 wide; opisthosoma 0.74 long, 0.57 wide. ALE = PME = PLE = 0.03; ALE���ALE 0.03, PME���PME 0.04, PME���PLE = ALE���PLE = 0.01. ���Clypeus height��� 0.03. Leg measurements: I 1.12 (0.37, 0.38, 0.19, 0.18); II 0.99 (0.30, 0.33, 0.19, 0.17); III 0.87 (0.26, 0.28, 0.18, 0.15); IV 1.17 (0.36, 0.39, 0.23, 0.19). Leg formula as male. Opisthosoma dark greyish yellow, with some blackish spots and 4 bright thin chevron-like markings dorsally. Epigyne small, with 2 tiny copulatory crevices located posteriorly, close to each other. Copulatory ducts extending forward, bifurcate and reaching sub-spermathecae (small, close to each other) and spermathecae (spherical, separated by the copulatory ducts). Fertilization ducts small, originating posterior to spermathecae (Figs 21 E��� F, 23 D���E). Distribution. Known only from rainforest of Xishuangbanna, Yunnan, China. Biology. Hahnia senaria sp. nov. is a dominant species in Xishuangbanna, in natural habitats and in plantations. All specimens were collected from August 2006 to June 2007. The months with largest abundances are November 2006 (40 females and 42 males) and May 2007 (88 females and 49 males). Apparently, two generations occurs throughout a year., Published as part of Zhang, Zhi-Sheng, Li, Shuqiang & Zheng, Guo, 2011, Comb-tailed spiders from Xishuangbanna, Yunnan Province, China (Araneae, Hahniidae), pp. 1-27 in Zootaxa 2912 on pages 24-26, DOI: 10.5281/zenodo.277820

Hahnia carmelita n. sp. Figs. 40, 41 Holotype. adult female from Ramat HaNadiv, Mt. Carmel, Israel, leg. Iris Musli, May 7 2004 (HUJ 15484), pitfall trap; male unknown. Etymology. The specific name refers to the type locality, Mt. Carmel. Description. Female. Carapace dark brown anteriorly and light brown mottled with black streaks posteriorly. Eyes surrounded by deep black patch. Clypeus barely projecting anteriorly. Chelicerae with 2 retromarginal teeth. Labium wider than long. Legs brown. Opisthosoma dark coloured with a few very light chevrons dorsally. Measurements (holotype + 2 ����; holotype listed first): total length 1.8, 1.7 ���2.0; carapace length 0.70, 0.70���0.72, width 0.60, 0.54���0.57, index 1.16, 1.23���1.33; labium length/width 0.50, 0.69���0.75; clypeal index 2.5, 2.0��� 2.5; leg lengths: I 1.72, 1.60���1.77, II 1.60, 1.42���1.64, III 1.49, 1.32���1.47, IV 1.97, 1.85���2.16; patellatibia index 0.87, 0.76���0.88. Epigynum. Plate with median, almost U-shaped, narrow opening, and long coiled spermathecal ducts discerning in the background (Fig. 40). Spermathecae form one pair of seminal receptacles; complex of winding ducts only partly exposed (Fig. 41). FIGURES 38 ���41, 38, 39. Zodarion odem n. sp. ��, holotype. 38. Epigynum, ventral view. 39. Spermathecae, dorsal (inner) view. 40, 41. Hahnia carmelita n. sp. ��, holotype. 40. Epigynum, ventral view. 39. Spermathecae, dorsal (inner) view; detail. Diagnosis. Based on female. The shape of the central epigynal opening and the complicated trajectory of the spermathecal ducts distinguish Hahnia carmelita n. sp. distinctly from all other Hahnia species. Distribution. Israel. Known only from the type locality on Mt. Carmel. Comments. Adult females of Hahnia carmelita n. sp. were collected by using pitfall traps in April and May, unlike females of H. candida that were found in winter (January; Levy 2003). Generally, the biotopes of both species are rather similar and further collecting is needed to clarify the consistency in the different seasonal occurrence of each species., Published as part of Levy, Gershom, 2007, Calommata (Atypidae) and new spider species (Araneae) from Israel, pp. 1-30 in Zootaxa 1551 on pages 15-16, DOI: 10.5281/zenodo.178107, {"references":["Levy, G. (2003) Spiders of the families Anyphaenidae, Hahniidae, Ctenidae, Zoridae and Hersiliidae (Araneae) from Israel. Israel Journal of Zoology, 49, 1 - 31."]}

Hahnia corticicola (B��senberg & Strand, 1905) * Material examined: 3 ♂ ♀, [19] 43��25'N 133��28'E, short stop on creek, 9.08.1998, YM; 89 ♂ ♀, [12] 43��55'N 134��23'E, Pravava Izvilinka Riv, 1-2.08.1998, YM.; 42 ♂ ♀, [13] 43��52'N 134��17'E, Pravaya Izvilinka Riv, 2.08.1998, YM.; 2 ♀♀, [18] 43��17'N 134��12'E, short stop on creek, 5.08.1998, YM; 1 ♀, [09] env. of Anisimovka Vill, Kangaus River 43��06'N 132��47'E, 28.07- 9.08.1998, YM; 39 ♂ ♀, [17] Lazovski Reserve, Korpad��� Camp, 43��16'N 134��08'E, 6-9.08.1998, YM; 10 ♀♀, Lazovski Reserve, America Kordon, 43��16.5'N 134��03'E, 14- 17.05.1999, YS; 1 ♀, Lazovsky Reserve, Proselochny Kordon, 43��00.5'N 134��08'E, 4 ��� 6.10.1999, YS; 19 ♂ ♀, Lazovsky Distr, Gersimov River, 43��24'N 133��53.7'E, 13.09.1998, YS; 1 ♂ 1 ♀, Lazo Town, Lazovka River valley, 43��23'N 133��54'E, 17.04.1999, YS. COMMENTS. The first record in the province, earlier reported from Magadan and Sakhalin areas [Marusik & Logunov, 1991]. In addition to Russia, known also from Korea, Japan and China [Song et al., 1999]., Published as part of Marusik, Yuri M. & Koponen, Seppo, 2000, New data on spiders (Aranei) from the Maritime Province, Russian Far East, pp. 55-68 in Arthropoda Selecta 9 (1) on page 60, DOI: 10.5281/zenodo.822713, {"references":["Marusik Yu. M., Logunov D. V. 1991. Spiders of the superfamily Amaurobioidea (Aranei) from Sakhalin and Kurily Islands / / Ibid. Vol. 70. No. 9. P. 87 - 94 [in Russian, with English summary],","Song DX, Zhu, M. S, Chen, J. 1999. The spiders of China. Hebei Science & Technology Press. 640 pp."]}

H. pusilla C. L. Koch: Steiermark (422: 1 ♂, 5 2♀., Published as part of Thaler, K., 1978, Bodenspinnen aus der Steiermark und ihren Nachbarl��ndern, gesammelt von Prof. Dr. R. Schuster, pp. 173-183 in Mitt. Abt. Zool. Landesmus. Joanneum 7 (3) on page 178

Hahnia helveola Simon: Steiermark (422: 1 2)., Published as part of Thaler, K., 1978, Bodenspinnen aus der Steiermark und ihren Nachbarl��ndern, gesammelt von Prof. Dr. R. Schuster, pp. 173-183 in Mitt. Abt. Zool. Landesmus. Joanneum 7 (3) on page 178

H. ononidum Simon: Steiermark (422: 1 2, 615: 1 ♂, 617: 1 ♀., Published as part of Thaler, K., 1978, Bodenspinnen aus der Steiermark und ihren Nachbarl��ndern, gesammelt von Prof. Dr. R. Schuster, pp. 173-183 in Mitt. Abt. Zool. Landesmus. Joanneum 7 (3) on page 178