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Subgenus Dimetrotina Casey, 1911 of the genus Atheta Dimetrotina Casey, 1911: 143 (as subgenus in the genus Dimetrota Mulsant & Rey, 1873; type species of Dimetrotina: Dimetrota (Dimetrotina) vaniuscula Casey, 1911, by monotypy) Mycetota ��d��m, 1987: 13 (as genus; type species Aleochara laticollis Stephens, 1832, by original designation), syn. nov. Dimetrotina: Smetana, 2004 b: 380 (as junior synonym of the Atheta subgenus Dimetrota). Mycetota: Smetana, 2004 b: 389 (as valid subgenus of the genus Atheta). Dimetrotina: Sch��lke & Smetana, 2015: 529 (as junior synonym of the Atheta subgenus Dimetrota). Mycetota: Sch��lke & Smetana, 2015: 545 (as valid subgenus of the genus Atheta). (other references are omitted) The type species of Dimetrotina, Dimetrota vaniuscula, is a junior synonym of Atheta pasadenae, a species very similar to the type species of Mycetota (Aleochara laticollis). See above for a list of characters shared by the two species. Due to similarity of the two type species, the name Mycetota is placed in synonymy with Dimetrotina. This results in the following new subgeneric combinations in the genus Atheta: Atheta (Dimetrotina) laticollis (Stephens, 1832), new subgeneric assignment Atheta (Dimetrotina) mucronata (Kraatz, 1859), new subgeneric assignment Atheta (Dimetrotina) pasadenae Bernhauer, 1906, new subgeneric assignment, Published as part of Gusarov, Vladimir I., 2016, The importance of tracking introduced species: nine synonyms of Atheta (Dimetrotina) pasadenae Bernhauer, 1906 (Coleoptera, Staphylinidae, Aleocharinae), pp. 390-398 in Zootaxa 4066 (4) on page 395, DOI: 10.11646/zootaxa.4066.4.2, http://zenodo.org/record/270555, {"references":["Casey, T. L. (1911 a) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 1 - 245.","Mulsant, E. & Rey, C. (1873) Description de divers coleopteres brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189.","Adam, L. (1987) Oligotinae and Aleocharinae (Coleoptera: Staphylinidae) from the Hortobagy National Park. Folia Entomologica Hungarica, 48, 9 - 15.","Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, time of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V. Baldwin and Cradock, London, 240 pp. http: // dx. doi. org / 10.5962 / bhl. title. 8133","Smetana, A. (2004 b) Subfamily Aleocharinae Fleming, 1821. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophyloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 353 - 494.","Schulke, M. & Smetana, A. (2015) Family Staphylinidae Latreille, 1802. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Revised and updated edition. Hydrophyloidea - Staphylinoidea. Brill, Leiden, pp. 304 - 900.","Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25, 1 - 196. http: // dx. doi. org / 10.5962 / bhl. title. 66002","Bernhauer, M. (1906) Neue Aleocharinen aus Nordamerika (II. Teil). Deutsche Entomologische Zeitschrift, 1906 (2), 337 - 348."]}

Lypoglossa Fenyes, 1918 (Figs. 1 ���17, 25���70, 72��� 78) Lypoglossa Fenyes, 1918: 23 (type species: Dasyglossa fenyesi Bernhauer, 1907, by original designation). Lypoglossa: Fenyes, 1920: 239 (as valid genus). Lypoglossa: Bernhauer & Scheerpeltz, 1926: 597 (as valid genus in subtribe Athetina Casey, 1910). Megacrotona Scheerpeltz, 1968: 159 (unavailable name; as subgenus of Atheta Thomson, 1858; type species: Oxypoda lateralis Mannerheim, 1830, by original designation). Megacrotona Benick & Lohse, 1974: 103 (available name; as valid genus in tribe Callicerini Lohse, 1969). Lypoglossa: Moore & Legner, 1975: 448 (as valid genus). Lypoglossa: Seevers, 1978: 134 (as valid genus in tribe Athetini). Lypoglossa: Lohse & Smetana, 1985: 294 (as valid genus). Megacrotona: Lohse & Smetana, 1985: 294 (as synonym of Lypoglossa). Lypoglossa: Lohse, 1989: 207 (as valid genus in tribe Athetini). Megacrotona: Lohse, 1989: 207 (as synonym of Lypoglossa). Lypoglossa: Hoebeke, 1992: 382 (as valid genus). Megacrotona: Hoebeke, 1992: 382 (as synonym of Lypoglossa). Lypoglossa: Ashe in Newton, Thayer, Ashe & Chandler, 2000: 371 (as valid genus in tribe Athetini; unassigned to subtribe). (Other references for Palaearctic Lypoglossa are omitted) Diagnosis. Lypoglossa is distinguished from other athetine genera by the combination of the following characters: body parallel��sided; sensilla a of epipharynx long (Fig. 2); antennal article 2 as long as article 3 (Figs. 12���13); ligula with broad base, split in two lobes (Fig. 6); labial palpus with setae,, and present (Fig. 6); pronotum transverse, 1.2���1.4 times as wide as long, with microsetae directed anteriorly along midline, and laterally and posteriorly in lateral portions of the disc (Type I, Benick & Lohse 1974) (Fig. 16); pronotal macrosetae short and thin; pronotal hypomera invisible in lateral view; medial macroseta of mesotibia thin, much shorter than tibial width; mesothoracic process narrow (Fig. 17); posterior margin of elytra near posterolateral angle straight or slightly concave; tarsal formula 4��5 �� 5; metatarsal segment 1 much longer than segment 2; single empodial seta shorter than claws (Fig. 15); abdominal terga 3���5 with weak transverse basal impression; medial lamellae of internal sac present (Figs. 49���50); copulatory piece with pointed apex (Fig. 49) and without sclerotized suspensoria; spermatheca S��shaped, with large umbilicus (Figs. 32, 78). Description. Length 3.2���4.6 mm, pronotal width 0.81���1.09 mm. Body parallelsided; uniformly brown to dark brown with lighter appendages or with reddish brown pronotum and elytra. Head transverse; eye length to temple length ratio 0.9���1.5; infraorbital carina complete. Antennal article 2 as long as article 3, article 4 elongate, 5 elongate to subquadrate, 6���10 elongate to transverse, apical article without coeloconic sensilla, as long as articles 9 and 10 combined (Figs. 12���13). Labrum (Fig. 1) transverse, with slightly emarginate anterior margin. Epipharynx (Fig. 2) with long sensilla a, with three pairs of small marginal setae, medial field with 29 pores, lateral rows with 3 pores each, anterolateral groups with 4���5 pores each, transverse row with six pores, posterolateral groups with 4���5 pores each; two medial proximal pores on each side and one lateral proximal pore. Mandibles (Figs. 3���5) broad, right mandible with a blunt medial tooth; velvety patch of dorsal molar area visible at 400 X. Maxilla (Figs. 8���11) with galea projecting slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; internal margin of galea with long subapical setae (Fig. 9); distal comb of lacinia is divided into a group of 6 closely placed spines and two isolated spines (Figs. 10���11), middle portion produced medially and covered with numerous fine setae (Figs. 10���11), ventral surface of lacinia with a marginal group of 5 strong setae (Fig. 10), dorsal surface of lacinia with numerous weak setae (Fig. 11). Labium as in Figs. 6 ���7, 14; ligula with broad base, almost entirely split into two slightly divergent lobes (Fig. 6); medial area of prementum with 2 pores and 19 pseudopores, lateral areas each with two asetose pores, single setose pore and 8���11 pseudopores (Fig. 6). Hypopharyngeal lobes as in Fig. 7. Labial palpus with setae,, and present (Fig. 6). Mentum (Fig. 14) with concave anterior margin. Pronotum (Fig. 16) transverse, 1.2���1.4 times as wide as long, with microsetae directed anteriorly along midline, in lateral portions of the disc microsetae directed laterally and posteriorly (Type I, Benick & Lohse 1974); macrosetae short; hypomera invisible in lateral view. Meso�� and metasternum as in Fig. 17, mesosternal process narrow, extending about 2 / 3 length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 9: 4: 1; mesocoxal cavities margined posteriorly; mesocoxae contiguous. Medial macroseta of mesotibia thin, much shorter than tibial width. Tarsal segmentation 4��5 �� 5, metatarsal segment 1 much longer than segment 2 (Fig. 15). One empodial seta, shorter than claws (Fig. 15). Posterior margin of elytra near posterolateral angle straight or slightly concave. Wings fully developed. Abdominal terga 3���5 with weak basal impression. Tergum 7 as long as tergum 6. Terga 3���5 matte due to dense punctation, punctation on terga 6���7 sparser. Tergum 7 with wide white palisade fringe. Posterior margin of tergum 8 in both sexes slightly convex, without modifications (Figs. 51, 53). Posterior margin of sternum 8 in both sexes strongly convex (Figs. 52, 54), in male protruding posteriorly beyond the apex of tergum 8 (Figs. 51���52). Copulatory piece of the aedeagus with pointed apical process (Fig. 49); medial lamellae present (Figs. 49���50). Sclerite of paramere (Sklerit of Brundin (1940), velar sac sclerite of Seevers (1978)) triangular (Figs. 31, 69). Spermatheca S��shaped, with large umbilicus (Figs. 32, 70, 78). Type species. Dasyglossa fenyesi Bernhauer, 1907, by original designation. Discussion. When Scheerpeltz (1968) proposed the name Megacrotona he fixed the type species of this subgenus (Oxypoda lateralis Mannerheim, 1830) but did not mention any character of Megacrotona. Therefore, Scheerpeltz did not satisfy the requirements of Article 13.1 of the Code (ICZN 1999) and had not made the name Megacrotona available. As far as I know, the earliest work where any characters of Megacrotona were mentioned is the paper by Benick and Lohse (1974). Since Benick and Lohse were the first authors to make the name Megacrotona available the name should be attributed to them and not Scheerpeltz., Published as part of Gusarov, Vladimir I., 2004, A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-36 in Zootaxa 747 on pages 2-4, DOI: 10.5281/zenodo.158194, {"references":["Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110.","Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, Pennsylvania, pp. 1 - 183.","Scheerpeltz, O. (1968) Coleoptera-Staphylinidae. Catalogus Faunae Austriae, 15 fa. Springer- Verlag, Wien, 279 pp.","Mannerheim, C. G. (1830) Precis d'un nouvel arrangement de la famille des Brachelytres, de l'ordre des insectes Coleopteres. St. Petersbourg, 87 pp.","Benick, G. & Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 72 - 220.","Lohse, G. A. (1969) Vorschlage zur Anderung der Aleocharinensystematik (Coleoptera: Staphylinidae). In: Bericht uber die 10. Wanderversammlung deutscher Entomologen, 15. bis 19. September 1965 in Dresden. Deutsche Akademie der Landwirtschaftswissenschaften zu Berlin, Berlin, pp. 169 - 175.","Moore, I. & Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp.","Lohse, G. A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Maklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39 (3), 281 - 300.","Lohse, G. A. (1989) Erganzungen und Berichtigungen zu Freude - Harde - Lohse \" Die Kafer Mitteleuropas \" Band 5 (1974) In: Lohse, G. A. & Lucht, W. H. (Eds.), Die Kafer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke & Evers Verlag, Krefeld, pp. 185 - 243.","Hoebeke, E. R. (1992) Taxonomy and distribution of the athetine genus Lypoglossa Fenyes (Coleoptera: Staphylinidae: Aleocharinae) in North America, with description of a new species. Journal of the New York Entomological Society, 100 (2), 381 - 398.","Newton, A. F., Thayer, M. K., Ashe, J. S. & Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418.","Brundin, L. (1940) Studien uber die Atheta - Untergattung Oreostiba Ganglbauer (Col. Staphylinidae). Entomologisk Tidskrift, 61 (1 - 4), 56 - 130, Taf. I - XVIII.","ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp."]}

Key to Nearctic species of Halobrecta 1 Males: spatuliform apex of median lobe (in parameral view) smaller, 1.1 times as wide as median lobe measured subapically at its narrowest point (Figs. 22 –23, 26– 27). Body length 3.0– 3.9 mm, pronotal length 0.49–0.57 mm. Distributed along the Pacific coast of North America (California) (Fig. 66). Also known from Europe, Chile, New Zealand and Inaccessible Island of the Tristan da Cunha archipelago.......................................................................................................... 1. H. algophila (Fenyes) – Males: spatuliform apex of median lobe (in parameral view) larger, 1.4–1.6 times as wide as median lobe measured subapically at its narrowest point (Figs. 41–44). Body length 2.9–3.2 mm, pronotal length 0.43–0.51 mm. Distributed along the Atlantic coast of North America (Virginia, New York) (Fig. 66). Also known from Europe and Chile......................................................................................... 2. H. flavipes Thomson

2. Lypoglossa manitobae Gusarov, sp. n. (Figs. 51���63). Lypoglossa angularis: Hoebeke, 1992: 390, ex parte. Type material. Holotype: CANADA: Manitoba: ��, Riding Mtn. Nat. Park, Stathclair Trail, 4.5 km S Kinnis Creek, 12.ix. 1979 (A.Smetana) (CNCI). Paratypes: CANADA: Manitoba: 2 ����, 3 ����, same data as the holotype (CNCI (2), CUIC (1), KSEM (1), ZMUN (1)); ��, Riding Mtn. Nat. Park, 1 km SE Long Lake, 16.ix. 1979 (A.Smetana) (CNCI). Diagnosis. Lypoglossa manitobae differs from L. angularis in having a smaller body size, and shorter and blunter dentiform projections at the base of the apex of the median lobe (Figs. 55���59; 25���29). Lypoglossa manitobae can be distinguished from the Palaearctic L. lateralis by the following characters: pronotum and elytra lighter than head and abdomen; punctation of head and pronotum not asperate; and a distinct shape of the aedeagus and spermatheca (Figs. 55���63; 72���78). L. manitobae differs from L. franclemonti in having a smaller body size; the apex of the median lobe (in parameral view) constricted basally (Figs. 55���57; 64���65); with a dentiform projection on each side of the base of the apex (Figs. 55���59; 64���67); and a smaller spermatheca (Figs. 62���63; 70). Description. Length 3.2���3.5 mm. Head and abdomen brown; pronotum, elytra, legs, antennae and mouthparts light brown. Head surface glossy, with weak isodiametric microsculpture, with fine punctation, distance between punctures equals 1���2 times their diameter. Eyes 1.0��� 1.3 times as long as temples. Antennal articles 4���6 elongate, 7���10 subquadrate. Pronotum transverse, 1.4 times as wide as head, width 0.81���0.87 mm, length 0.60��� 0.66 mm, width to length ratio 1.3���1.4; surface glossy, with weak isodiametric microsculpture; punctation finer than on head, distance between punctures equals 2 times their diameter. Elytra wider (0.89���0.91 mm) and longer (0.66���0.74 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.1), 1.3 times as wide as long, surface glossy, with poorly visible microsculpture; punctation stronger and denser than on pronotum, distance between punctures equals ����� 1 times their diameter. Abdominal terga with very fine (poorly visible at 70 x) microsculpture consisting of transverse waves; terga 3���5 matte due to fine and dense punctation, distance between punctures equals 1���3 times their diameter, terga 6���7 glossy, with sparser punctation, distance between punctures equals 2���6 times their diameter. In males, head with weak medial impression and pronotum with broad impression along midline. Aedeagus as in Figs. 55���61. Spermatheca as in Figs. 62���63. Distribution. Lypoglossa manitobae is known only from Riding Mountains National Park in Manitoba (Fig. 18) where it is sympatric with L. franclemonti. Natural History. No information is available., Published as part of Gusarov, Vladimir I., 2004, A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-36 in Zootaxa 747 on pages 25-28, DOI: 10.5281/zenodo.158194, {"references":["Hoebeke, E. R. (1992) Taxonomy and distribution of the athetine genus Lypoglossa Fenyes (Coleoptera: Staphylinidae: Aleocharinae) in North America, with description of a new species. Journal of the New York Entomological Society, 100 (2), 381 - 398."]}

Key to the species of Lypoglossa 1 Punctation of head and pronotum not asperate. Pronotum and elytra usually much lighter than head and abdomen. Apex of median lobe wide, less than 2 times as long as wide (Figs. 25 ���27, 55���57, 64��� 65).Nearctic species................................................. 2 ��� Punctation of head and pronotum asperate. Body dark brown, pronotum and elytra almost as dark as head and abdomen. Apex of median lobe long and narrow, 2.5 times as long as wide (Figs. 72���73). Palaearctic species. Body length 3.6 ���4.0 mm........................................................................................................ 4. L. lateralis (Mannerheim) 2. Pronotal width to length ratio 1.2���1.3. Apex of median lobe (in parameral view) broadest at the base, not constricted (Figs. 64���65). No projections at the base of the apex (Figs. 64���67). Spermatheca larger, its proximal loop wider, almost circular (Fig. 70). Body length 3.8���4.6 mm, pronotal length 0.71���0.89 mm, pronotal width 0.90��� 1.09 mm. Distributed in interior Canada and eastern North America (Fig. 71).......................................................................................................... 3. L. franclemonti Hoebeke ��� Pronotal width to length ratio 1.3���1.4. Apex of median lobe (in parameral view) constricted basally (Figs. 25 ���27, 33���35, 55��� 57), with dentiform projection on each side of the apex base (Figs. 25 ���29, 55��� 59). Proximal loop of spermatheca narrower, with parallel sides (Figs. 32, 40, 48) or spermatheca smaller (Figs. 62���63).............................. 3 3. Dentiform projections at the base of the apex of median lobe longer and sharper (Figs. 25���29). Body larger (body length 3.9���4.4 mm, pronotal length 0.64���0.74 mm, pronotal width 0.91���1.04 mm). Three subspecies known from Alaska, British Columbia, Yukon Territory, and mountains of the western and north��eastern North America (Fig. 18).................................................................................................. 1. L. angularis (M��klin) ��� Dentiform projections at the base of the apex of median lobe shorter and blunter (Figs. 55���59). Body smaller (body length 3.2���3.5 mm, pronotal length 0.60���0.66 mm, pronotal width 0.81���0.87 mm). Known only from the Riding Mountains National Park, Manitoba (Fig. 18)............................................... 2. L. manitobae Gusarov, sp. n., Published as part of Gusarov, Vladimir I., 2004, A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-36 in Zootaxa 747 on pages 5-6, DOI: 10.5281/zenodo.158194

1. Lypoglossa angularis (M��klin in Mannerheim, 1853) (Figs. 1 ���12, 15, 17, 25��� 50) Myrmedonia angularis M��klin in Mannerheim, 1853: 181. Euryusa obtusa LeConte, 1866: 373, syn. nov. Dasyglossa angularis: Bernhauer, 1906: 348 (as synonym of Dasyglossa prospera (Erichson, 1839)). Dasyglossa fenyesi Bernhauer, 1907: 402. Lypoglossa fenyesi: Fenyes, 1920: 239 (as valid species). Dasyglossa angularis: Fenyes, 1920: 359 (as synonym of Dasyglossa prospera (Erichson, 1839)). Lypoglossa fenyesi: Bernhauer & Scheerpeltz, 1926: 597 (as valid species). Dasyglossa angularis: Bernhauer & Scheerpeltz, 1926: 766 (as synonym of Dasyglossa prospera (Erichson, 1839)). Lypoglossa fenyesi: Moore & Legner, 1975: 448 (as valid species). Devia angularis: Moore & Legner, 1975: 396 (as synonym of Devia prospera (Erichson, 1839)). Lypoglossa fenyesi: Seevers, 1978: 271 (as valid species). Lypoglossa angularis: Lohse & Smetana, 1985: 293 (as valid species). Lypoglossa angularis: Hoebeke, 1992: 390, ex parte (as valid species). Lypoglossa fenyesi: Hoebeke, 1992: 391 (as synonym of L. angularis) Examined material. Listed separately for each subspecies except for the following female specimens unassigned to subspecies: UNITED STATES: Oregon: Deschutes Co.: ��, 12 mi. SW Sisters, FSR 1551, 4400 ', 23.vii. 1979 (J.M.Campbell & B.A.Smetana) (CNCI); California: El Dorado Co.: ��, Lake Tahoe, Cascade Lake, 6300 ', 11.viii. 1969 (A.Smetana) (CNCI). Diagnosis. Lypoglossa angularis differs from the Palaearctic L. lateralis in the following characters: pronotum and elytra lighter than head and abdomen; punctation of head and pronotum not asperate, and a distinct shape of the aedeagus (Figs. 25���30; 72���77) and spermatheca (Figs. 32; 78). Lypoglossa angularis differs from L. franclemonti in having a more transverse pronotum, less elongate antennal articles; the apex of the median lobe (in parameral view) constricted basally (Figs. 25 ���27, 33��� 35; 64���65), with a dentiform projection on each side of the apex base (Figs. 25���29; 64���67); the proximal loop of the spermatheca narrower, with parallel sides (Figs. 32, 40; 70). Lypoglossa angularis can be distinguished from L. manitobae in having a larger body size, and longer and sharper dentiform projections at the base of the apex of median lobe (Figs. 25���29; 55���59). Description. Length 3.9���4.4 mm. Head and abdomen dark brown; pronotum lighter, brownish red to brown with lighter margins; elytra brownish yellow to brown, with darker scutellar area and posterolateral angle; legs, mouthparts and 3 basal antennal articles brownish yellow, antennal articles 4���11 darker, reddish brown. Head surface glossy, with weak isodiametric microsculpture, with strong punctation, distance between punctures equals 1���2 times their diameter. Eyes 1.1���1.4 times as long as temples. Antennal articles 4���5 elongate, 6 slightly elongate or subquadrate, 7���10 subquadrate or slightly transverse (Fig. 12). Pronotum transverse, 1.5 times as wide as head, width 0.91���1.04 mm, length 0.64��� 0.74 mm, width to length ratio 1.4; surface glossy, with weak isodiametric microsculpture; punctation finer than on head, distance between punctures equal to their diameter. Elytra wider (1.00��� 1.17 mm) and longer (0.71���0.86 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.1), 1.3���1.4 times as wide as long, surface glossy, with weak isodiametric microsculpture; punctation slightly asperate and stronger than on pronotum, distance between punctures equals ����� 1 times their diameter. Abdominal terga with very fine (poorly visible at 70 x) microsculpture consisting of transverse waves; terga 3���5 matte due to fine and dense punctation, distance between punctures equals 1���2 times their diameter, terga 6���7 glossy, with sparser punctation, distance between punctures equals 2���5 times their diameter. In males, head with weak medial impression and pronotum with broad impression along midline. Aedeagus as in Figs. 25 ���31, 33���39, 41��� 47. Spermatheca as in Figs. 32, 40, 48. Distribution. Lypoglossa angularis is widely distributed in the western North America, from Alaska, Yukon Territory and British Columbia in the north to the high mountains of California, Colorado, New Mexico and Arizona in the south. There is an isolated population of this species in the northeastern states and provinces (Newfoundland, Quebec, Maine and New Hampshire) (Fig. 18). In Alaska L. angularis is continuously distributed and along the coast occurs at sea level. In interior British Columbia and Alberta and further south Lypoglossa angularis is restricted to boreal forests in high mountains and does not occur in valleys. Further south the localities are at higher elevations in the mountains. Despite the fact that in the interior United States the recorded distributions of L. lateralis consists of isolated populations, I could find no gaps in any morphological characters between the specimens from different isolates. Apparently the isolation between these populations is recent and L. angularis probably had a continuous distribution in the interior United States during colder and/or wetter periods of the Pleistocene. I recognize three valid subspecies within L. angularis (Fig. 18). The main reason for interpreting the three geographic varieties as subspecies is that there is no distinct hiatus between them and intermediate forms are occasionally found. The subspecies are para�� or allopatric in their distribution. The differences between these subspecies are discussed below. Natural History. Lypoglossa angularis is common in forest litter and moss in boreal forests. It was also collected in dung, gopher burrows and by pitfall traps., Published as part of Gusarov, Vladimir I., 2004, A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-36 in Zootaxa 747 on pages 6-10, DOI: 10.5281/zenodo.158194, {"references":["Mannerheim, C. G. (1853) Dritter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 26 (3), 95 - 273.","LeConte, J. L. (1866) Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Sciences of Philadelphia, 18, 361 - 394.","Bernhauer, M. (1906) Neue Aleocharinen aus Nordamerika. (II. Teil). Deutsche Entomologische Zeitschrift, 1906 (2), 337 - 348.","Erichson, W. F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. 1 - 400.","Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Moore, I. & Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp.","Lohse, G. A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Maklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39 (3), 281 - 300.","Hoebeke, E. R. (1992) Taxonomy and distribution of the athetine genus Lypoglossa Fenyes (Coleoptera: Staphylinidae: Aleocharinae) in North America, with description of a new species. Journal of the New York Entomological Society, 100 (2), 381 - 398."]}

2. Halobrecta flavipes Thomson, 1861 (Figs. 37–51) Halobrectha flavipes Thomson, 1861: 50. Homalota maritima Waterhouse, 1863: 138 (nec Mannerheim, 1843: 224) (replacement name for Homalota flavipes (Thomson, 1861), nec Homalota flavipes (Gravenhorst, 1806)). Homalota halobrectha Sharp, 1869: 139 (replacement name for Homalota flavipes (Thomson, 1861), nec Homalota flavipes (Gravenhorst, 1806)). Halobrectha flavipes: Mulsant & Rey, 1875: 45 (as synonym of Ha. anthracina (Fairmaire, 1853)). Atheta (s. str.) pocahontas Casey, 1910: 19, syn. nov. Atheta (s. str.) vaticina Casey, 1910: 19, syn. nov. Aloconota (s. str.) incertula Casey, 1910: 84, syn. nov. Atheta (Halobrecta) flavipes: Fenyes, 1920: 185 (as valid species). Atheta (Halobrecta) flavipes: Bernhauer & Scheerpeltz, 1926: 621 (as valid species). Atheta (Halobrecta) maritima (Waterhouse): Bernhauer & Scheerpeltz, 1926: 621 (as synonym of At. flavipes). Atheta (Halobrecta) halobrectha: Bernhauer & Scheerpeltz, 1926: 621 (as synonym of At. flavipes). Atheta (s. str.) pocahontas: Bernhauer & Scheerpeltz, 1926: 648 (as valid species). Atheta (s. str.) vaticina: Bernhauer & Scheerpeltz, 1926: 651 (as valid species). Atheta (s. str.) incertula: Bernhauer & Scheerpeltz, 1926: 644 (as valid species). Atheta (Halobrecta) flavipes: Scheerpeltz, 1934: 1600 (as valid species). Halobrecta flavipes: Benick & Lohse, 1974: 219 (as valid species). Halobrecta flavipes: Hammond, 2000: 275 (as valid species). Halobrecta flavipes: Pace, 2000: 376 (ex parte; as valid species). Type material. Lectotype of Halobrecta flavipes (here designated): ɗ, "L­a [Lomma, Sweden]" (MZLU (Thomson collection)). Lectotype of Atheta pocahontas (here designated): ɗ, "Va. [with two black dots under "a", Virginia: Fort Monroe (according to Casey locality code (FitzGerald 1962))]", " pocahontas ­ 3 PARATYPE USNM 39310 " (red label), " CASEY bequest 1925 ". Paralectotypes: Ψ, "Va. [with one black dot under "a", Virginia: Norfolk (FirzGerald 1962)]", " Atheta pocahontas Csy. ", " TYPE USNM 39310 " (red label), " CASEY bequest 1925 "; ɗ, with missing head, prothorax and elytra, ɗ, "Va. [with two black dots under "a", Virginia: Fort Monroe]", " pocahontas ­ 2 PARATYPE USNM 39310 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta vaticina (here designated): Ψ, "L.I. [Long Island, New York]", " Atheta vaticina Csy. ", " TYPE USNM 39311 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Aloconota incertula (here designated): Ψ, "Va. [with two black dots under "a", Virginia: Fort Monroe]", " incertula Csy. ", " TYPE USNM 39076 " (red label), " CASEY bequest 1925 " (NMNH). Additional material. CHILE: Llanquihue Prov.: 2 ɗɗ, 2 ΨΨ, 6 specimens (sex undetermined), Puerto Montt, Pilluco, sifting beach drift, 24.xii. 1984 (S. & J.Peck); ɗ, Ψ, 1 specimen (sex undetermined), Lenca, 34 km SE Puerto Montt, beach drift, 25.xii. 1984 (S. & J.Peck) (FMNH); GERMANY: Ψ, N Friesen Islands, Spekeroog Island, 20.v– 16.vi. 1992 (B.W.Johanning) (KSEM); SWEDEN: ɗ, Ψ, Oskarshamn, 10.vi. 1941 (H.Bachlund) (KSEM); UNITED KINGDOM: ɗ, Aldeburgh, 15.viii. 1925 (C.J.Saunders) (BMNH). Diagnosis. Halobrecta flavipes is closely related to Ha. algophila, but differs in having the body size smaller on average; and the spatuliform apex of the median lobe larger (in parameral view), the apex is 1.4–1.5 times as wide as the median lobe measured subapically at its narrowest point (1.1 times in Ha. algophila) (Figs. 41–44; 22 –23, 26– 27). Halobrecta flavipes differs from the Mediterranean species tentatively identified as Ha. halensis (see Discussion for Halobrecta) in having the median lobe of the aedeagus narrower (in parameral view) (Figs. 41–44; 64). Halobrecta flavipes differs from the Palaearctic Ha. algae and Ha. princeps by having the posterior margin of the male tergum 8 rounded and not crenulate (Fig. 37) (in Ha. algae and Ha. princeps the posterior margin is straight and may be slightly crenulate (Fig. 52)), and by the apex of the median lobe broader (Figs. 41–44; 56–57). Halobrecta flavipes differs from the Oriental Ha. cingulata in having a different shape of the median lobe (in lateral view) (Figs. 45–48 in this paper; Fig. 9, G in Sawada 1985). Halobrecta flavipes differs from the Neotropical Ha. discipula in having a different shape of the median lobe (in parameral view) (Figs. 41–44: this paper; Fig. 3: Pace 1999). Description. Length 2.9–3.2 mm. Pronotal width 0.53–0.61 mm, length 0.43–0.51 mm. Elytra 1.3–1.5 times as long as pronotum. In all character states very similar to Ha. algophila, but differs in the shape of the median lobe as described in the diagnosis. Aedeagus as in Figs. 41–50. Spermatheca as in Fig. 51. Discussion. The male types of At. pocahontas are similar in external characters to the lectotype of Ha. flavipes, as well as in the shape of the aedeagus. These names are deemed to be synonyms. The only existing type of Al. incertula is a female but it was collected in the same locality as the male lectotype of At. pocahontas. Since the type of Al. incertula is similar to examined specimens of Ha. flavipes, the first name is placed in synonymy with the second. The single existing type of At. vaticina is a female but it is similar in external characters to the examined specimens of Ha. flavipes. Considering that all examined males from the East Coast belonged to Ha. flavipes, I synonymize At. vaticina with that species. It seems appropriate to discuss the status of the names listed as synonyms of Ha. flavipes by Fenyes (1920) and Bernhauer and Scheerpeltz (1926). Unfortunately, these authors sometimes did not distinguish between newly proposed names and subsequent usage of these names. This practice, followed by some recent authors (e. g., Klimaszewski et al. 2002), is very confusing. The entries for " elongatula Stephens, 1832 ", " atricilla Scriba, 1866 " and " puncticeps Mulsant & Rey, 1875 " (see Fenyes 1920, p. 185; Bernhauer & Scheerpeltz 1926, p. 621; Klimaszewski et al. 2002, p. 483) in fact refer to subsequent citations of the names proposed by Gravenhorst (1802, as Aleochara elongatula); Erichson, (1839, as Homalota atricilla) and Thomson (1852, as Homalota puncticeps). Stephens (1832, p. 127), Scriba (1866, p. 290) and Mulsant and Rey (1875, p. 38) clearly did not intend to describe new species and referred to original papers of Gravenhorst, Erichson and Thomson. Even if Stephens, Scriba and Mulsant and Rey applied the same species names in a different sense, their usage did not create new nominal taxa but instead represents misidentifications. Despite the fact that Thomson described Ha. flavipes as a member of the genus Halobrecta, this taxon was not accepted as a genus separate from Atheta by some of his colleagues (e.g., Waterhouse 1863; Sharp 1869; Bernhauer & Scheerpeltz 1926). Since Waterhouse (1863) placed Halobrecta flavipes Thomson in Homalota, together with Aleochara flavipes Gravenhorst, 1806 (now considered a member of Notothecta Thomson, 1858), he had to introduce (Waterhouse 1863, p. 138) a new name (Homalota maritima) to replace the resulting secondary junior homonym Homalota flavipes (Thomson). Unfortunately, the name proposed by Waterhouse is preoccupied and Sharp (1869) had to propose a different replacement name for Homalota flavipes (Thomson). Now that Halobrecta flavipes Thomson, 1861 is a member of the genus Halobrecta and Aleochara flavipes Gravenhorst, 1806 is a member of Notothecta, the replacement names Homalota maritima Waterhouse and Ho. halobrectha Sharp are unnecessary and become objective synonyms of Ha. flavipes. Homalota atricilla Erichson, 1839, described from Sardinia, is usually considered a member of Halobrecta (e.g., Bernhauer & Scheerpeltz 1926). If this is true then Ho. atricilla is the oldest name in the genus Halobrecta. However the description of this species, particularly the shape of the antennal segments 4–10 does not fit any of the five species of Halobrecta examined by me. According to Kraatz (1857) and Scriba (1866) the single type of Ho. atricilla (deposited in the museum of Turin) is an immature poorly pigmented specimen. Scriba (1866) considered Ho. atricilla to be conspecific with Ha. flavipes but Sharp (1869) did not agree with this opinion because Erichson's description did not fit Sharp's specimens of Ha. flavipes (which he referred to as Ho. halobrectha). In my opinion, Ho. atricilla may not even belong to Halobrecta. Reexamination of the type of this species is necessary to clarify the status of this name. Two species of Halobrecta were described from the Channel coasts of northern France: Ha. pubes (Mulsant & Rey, 1873) from Normandy and Ha. anthracina Fairmaire, 1853) from the embouchure of the River Somme. One species, Halobrecta halensis Mulsant & Rey, 1873, was described from the Mediterranean coasts of France (Languedoc) and may be conspecific with the specimen from Italy illustrated in Figs. 64–65 (See Discussion in Halobrecta). Examination of types is necessary to clarify the status of these three species. The examined specimens from Chaiten, Chile, listed by Pace (2000) as Ha. flavipes belong in fact to Ha. algophila. Distribution. Halobrecta flavipes is known from the Atlantic coast of North America (Virginia and New York), from Chile and Europe. Natural History. Halobrecta flavipes is a marine littoral species., Published as part of Gusarov, Vladimir I., 2004, A revision of the Nearctic species of the genus Halobrecta Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae) with notes on some Palaearctic species of the genus, pp. 1-25 in Zootaxa 746 on pages 13-19, DOI: 10.5281/zenodo.158213, {"references":["Thomson, C. G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom III. Berlingska Boktryckeriet, Lund, 278 pp.","Waterhouse, G. R. (1863) [Untitled notes on the genus Homalota] Proceedings of the Entomological Society of London, 1863, 136 - 140.","Mannerheim, C. G. (1843) Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16 (1 - 2), 175 - 314.","Gravenhorst, J. L. C. (1806) Monographia coleopterorum micropterorum. Henricus Dieterich, Gottingen, xvi + 248 pp.","Sharp, D. (1869) A revision of the British species of Homalota. Transactions of the Entomological Society of London, 1869 (2 - 3), 91 - 272.","Muslant, E. & Rey, C. (1875) Tribu des Brevipennes. Famille des Aleochariens (suite). Septieme branche: Myrmedoniaires. 2 e partie. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4, 7 [1874], 27 - 496, Pl. vi - ix.","Fairmaire, L. (1853). Excursion entomologique dans la Baie de la Somme. Lettre a M. le Dr. Aube. Annales de la Societe Entomologique de France, (2) 10 [1852], 663 - 691.","Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Benick, G. & Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 72 - 220.","Hammond, P. M. (2000) 16. Coastal Staphylinidae (rove beetles) in the British Isles, with special reference to saltmarshes. In: Sherwood, B. R., Gardiner, B. G. & Harris T. (Eds.), British Saltmarshes. Joint Symposium on British Saltmarshes organized between the Linnean Society of London, the Royal Society for the Protection of Birds and English Nature (2000: London). Forrest Text, Cardingan, pp. 247 - 302.","Pace, R. (2000) Aleocharinae del Cile, nuove o poco note (Coleoptera, Staphylinidae). Bollettino del Museo regionale di Scienze naturali, Torino, 17 (2), 355 - 510.","FitzGerald, W. (1962) Casey locality code. Manuscript kept with Casey collection at the National Museum of National History, Washington, D. C.","Sawada, K. (1985) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). IV. Ten oriental species described by M. Bernhauer and M. Cameron. Contributions from the Biological Laboratory, Kyoto University, 27 (1), 91 - 110.","Pace, R. (1999) Aleocharinae del Cile (Coleoptera, Staphylinidae). Bollettino del Museo civico di Storia naturale di Verona, 23, 119 - 210.","Klimaszewski, J., Maus, C. & Gardiner, A. (2002) The importance of tracking introduced species: New records of athetine rove beetles from South Atlantic Inaccessible Island (Coleoptera, Staphylinidae, Aleocharinae). Coleopterists Bulletin, 56 (4), 481 - 490.","Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Baldwin and Cradock, London, 448 pp.","Scriba, W. (1866) Ueber Staphylinen. Berliner Entomologische Zeitschrift, 10 (1 - 3), 289 - 290.","Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non Exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium. C. Reichard, Brunswick, lxvi + 207 pp.","Erichson, W. F. (1839) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. 1 - 400.","Thomson, C. G. (1852) Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 9 (6), 131 - 146.","Thomson, C G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40.","Kraatz, G. (1857) Ueber Coleoptera. Berliner Entomologische Zeitschrift, 1, 175 - 178.","Muslant, E. & Rey, C. (1873) Description de divers Coleopteres Brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189."]}

4. Lypoglossa lateralis (Mannerheim, 1830) (Figs. 72���78) Oxypoda lateralis Mannerheim, 1830: 70. Oxypoda (s. str.) lateralis: Fenyes, 1920: 366 (as valid species). Oxypoda (s. str.) lateralis: Bernhauer & Scheerpeltz, 1926: 748 (as valid species). Atheta (Acrotona) lateralis: Brundin, 1952: 101 (as valid species). Atheta (Megacrotona) lateralis: Scheerpeltz, 1968: 159 (as valid species). Megacrotona lateralis: Benick and Lohse, 1974: 103 (as valid species). Lypoglossa lateralis: Lohse & Smetana, 1985: 294 (as valid species). Lypoglossa lateralis: Hoebeke, 1992: 396 (as valid species). (Other references for Lypoglossa lateralis are omitted) Type material. Lectotype of Oxypoda lateralis (designated by Hoebeke (1992)): FIN�� LAND: 30 km NW Turku, Villnas (MZHF). Additional material. RUSSIA: Murmansk Reg.: 5 specimens, 20 km SSE Kirovsk, pifall traps, 15.vi��� 15.viii. 1994 (M.Kozlov) (ZMUN); Tyumen��� Reg.: ��, Agap River, 22.vi. 1988 (Ye.Zinov���yev) (ZMUN); Magadan Reg.: ��, ��, 25 km N Magadan, in grass, 3.vii. 1985 (Yu.M.Marusik); 6 specimens, 30 km N Magadan, Snezhnaya Dolina, viii��� ix. 1996 (Yu.M.Marusik); 2 ����, ��, ditto but 12���14.ix. 1996; ��, upper reaches of the Kolyma River, Jack London Lake, Studyonyy Creek, 13.vii. 1986 (Yu.M.Marusik); 3 specimens, upper Kolyma River, Bol���shoy Annachag Mt. Range, env. of Sibit��Tyellakh, Olen��� River, 800m, Alnus, Pinus pumila, in forest litter, 15.vii. 1985 (Yu.M.Marusik) (ZMUN); Chukotkskiy Aut. Distr.: 8 specimens, E Chukotka, upper reaches of the Bol���shaya Osinovaya River (a tributary of the Belaya River), Chosenia, in forest litter, 13.vii. 1989 (Yu.M.Marusik); 11 specimens, 10 km E Amguema, 22.vii. 1988 (Yu.M.Marusik) (ZMUN). Diagnosis. Lypoglossa lateralis differs from all Nearctic species of Lypoglossa in having more uniform and darker body color; asperate punctation of head and pronotum; slightly transverse antennal segments 6���10; and a different shape of the aedeagus and spermatheca (Figs. 72���78). Description. Length 3.6 ���4.0 mm. Head and abdomen dark brown; pronotum dark brown or brown, in most specimens with light margins; elytra dark brown to brown, with lighter oblique area stretching from humeral angle to sutural angle; legs brown, mouthparts and antennae dark brown. Head surface glossy, with weak isodiametric microsculpture, with strong and asperate punctation, distance between punctures equals 1���1.5 times their diameter. Eyes 1.2���1.5 times as long as temples. Antennal article 4 elongate, 5 subquadrate, 6���10 slightly transverse (Fig. 13). Pronotum transverse, 1.5 times as wide as head, width 0.81���0.94 mm, length 0.59��� 0.67 mm, width to length ratio 1.4; surface glossy, with weak isodiametric microsculpture, punctation asperate and denser than on head, distance between punctures equals their diameter. Elytra wider (0.97���1.11 mm) and longer (0.73���0.86 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.3), 1.3 times as wide as long, surface glossy, with poorly visible microsculpture; punctation asperate, denser and stronger than on pronotum, distance between punctures equal to ����� 1 times their diameter. Abdominal terga with very fine (poorly visible at 70 x) microsculpture consisting of transverse waves; terga 3���5 matte due to fine and dense punctation, distance between punctures equals 1���3 times their diameter, terga 6���7 glossy, with sparser punctation, distance between punctures equals 2���6 times their diameter. Aedeagus as in Figs. 72���77. Spermatheca as in Fig. 78. Distribution. Lypoglossa lateralis is a Palaearctic species restricted to subarctic, northern boreal and alpine forests. Examined specimens from Murmansk and Magadan Regions, and Chukotka Peninsula were identical in all external characters and in the shape of the male and female genitalia. Natural History. Lypoglossa lateralis occurs in forest litter., Published as part of Gusarov, Vladimir I., 2004, A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-36 in Zootaxa 747 on pages 31-34, DOI: 10.5281/zenodo.158194, {"references":["Mannerheim, C. G. (1830) Precis d'un nouvel arrangement de la famille des Brachelytres, de l'ordre des insectes Coleopteres. St. Petersbourg, 87 pp.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Brundin, L. (1952) Acrotona - Studien. (Gattung Atheta, Col., Staphylinidae). Entomologisk Tidskrift, 73 (1 - 4), 93 - 145.","Scheerpeltz, O. (1968) Coleoptera-Staphylinidae. Catalogus Faunae Austriae, 15 fa. Springer- Verlag, Wien, 279 pp.","Benick, G. & Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 72 - 220.","Lohse, G. A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Maklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39 (3), 281 - 300.","Hoebeke, E. R. (1992) Taxonomy and distribution of the athetine genus Lypoglossa Fenyes (Coleoptera: Staphylinidae: Aleocharinae) in North America, with description of a new species. Journal of the New York Entomological Society, 100 (2), 381 - 398."]}

Sableta infulata Casey, 1910 (Figs. 1���26) Sableta (s. str.) infulata Casey, 1910: 107. Atheta (Sableta) infulata: Fenyes, 1920: 221 (as valid species). Atheta (Sableta) infulata: Bernhauer & Scheerpeltz, 1926: 666 (as valid species). Atheta (Sableta) infulata: Moore & Legner, 1975: 363 (as valid species). Sableta infulata: Seevers, 1978: 261 (as valid species). Sableta infulata: Ashe, 2000: 370 (as valid species). Holotype: Male, with labels as follows: ���Miss.[issippi, Vicksburg]���; hand written label ��� Sableta infulata Csy. ���; red label ��� TYPE USNM 39134 ���; ���Casey bequest 1925 ���; ��� Holotypus, Sableta infulata Casey, V. I. Gusarov rev. 2000 ��� (NMNH). The apical abdominal segments of the holotype are dissected; the permanent preparation of the dissected segments and genitalia are preserved in dimethylhydantoin formaldehyde resin on a slide pinned beneath the beetle. Casey (1910, p. 108) states specifically that he only has a single specimen of this species; that specimen is therefore the holotype. Additional material. UNITED STATES: Illinois: Jackson Co: 4.0 mi E Grand Tower, 24 October 1982, J. S. Ashe, ex Bjerkendera sp. (2 KSEM, in alcohol); Indiana: ���Smith Sta.[tion]���, 22 August 1942, C. H. Seevers, ex fungus (�� FMNH); Kansas: Douglas Co.: Bridenthal Ecological Preserve, 3.2 mi N. Baldwin City, 270 m, 12 October 1996, ex fungus��covered logs (4 KSEM); same locality and collector, 18 October 1995, ex Pleurotus sp. (3 KSEM); same locality, date and collector, ex Grifola frondosa (22 KSEM, in alcohol); same locality, 18 September 1999, V. I. Gusarov (3 ����, ZMUN); Michigan: Berrien Co.: Warren Woods, 3.2 km E Lakeview, 6 October 1982, J. S. Ashe, ex Ischnoderma fulginosum (4 KSEM); North Carolina: Montgomery Co.: Uwharrie Nat���l. Forest, 7.7 km W. Uwharrie Forest Rd 576, near Badin Lake, 17 August 1989, J. S. and A. K. Ashe, ex Pleurotus ostreatus (10, 2 on microslides, KSEM); same locality, date and collectors, ex Laetiporus sulfureus (1 KSEM); Rockingham Co.: Reidsville, 19 August 1989, J. S. Ashe, ex misc. mushrooms (1 KSEM); South Carolina: undetermined locality and date, 1990, G. Moulton (1 KSEM); Tennessee: Sevier Co.: Great Smoky Mtns. Natl. Pk., Metcalf Bottoms Picnic Area, 546 m, 35.68089��N, 83.64945��W, 18 July 2003, J. S. Ashe, # GSMNP 1 A03 0 21, ex Daedalea sp. (2 KSEM, in alcohol); Great Smoky Mtns. Natl. Pk., Laurel Falls Old Growth Forest, 1006 m, 35.68070��N, 83.860455��W, 19 July 2003, J. S. Ashe, # GSMNP 1 A03 0 31, ex Ganoderma tsugae (4 KSEM, in alcohol); Great Smoky Mtns. Natl. Pk., Greenbrier Area, Porter���s Creek Trail, 762 m, 35.67864��N, 83.89842��W, 20 July 2003, J. S. Ashe, # GSMNP 1 A03 0 32, ex Pleurotus ostreatus (6, KSEM, in alcohol). Diagnosis. Sableta infulata is the only described species remaining in the genus Sableta. It is similar to examined undescribed species of Sableta in the body proportions and color but differs in the shape of the genitalia. Description. Length 2.0���2.2 mm. Color pattern distinctive (Fig. 1); head light brown to brown, slightly darker than pronotum in most specimens; antenna light brown to brown; pronotum light brown to brown; elytra dark brown to blackish with outer basal angles flavate to light brown, dark brown to blackish color extended along suture to scutellum in most, extent of light color on external basal angles varying among specimens, from limited to external basal angles in some, up to �� of elytra light in others (most with about 1 / 3 of elytra with flavate/light brown color); mesosternum light brown to flavate; metasternum dark brown, some specimens with extreme antero��lateral angles and apical margin light brown to flavate; abdomen flavate, tergum V with middle of apical margin clouded with brown in some specimens, segment VI black to dark brown with middle of sternum VI lighter in color than tergum VI in most specimens, tergum VIII black to dark brown (medium brown in a few specimens) with base and baso��lateral areas flavate; sternum VIII flavate to light brown with apical margin darker brown in some specimens; lateral processes of tergum IX brown in some specimens. Integument moderately glossy. Antenna (Fig. 7) with article 4 subquadrate, articles 5���6 subquadrate or slightly transverse, 7 slightly transverse, 8���10 transverse, article 11 longer than articles 9 and 10 combined. Head, pronotum and elytra with inconspicuous, weakly reticulate to obsolete reticulate microsculpture; with fine punctation, distance between punctures equal to 1���2 times their diameter. Abdominal terga III���VIII with fine transverse microsculpture; distance between punctures equal to 2���4 times their diameter on terga III���V and 3���6 times on terga VI���VII. Secondary sexual characteristics: Male with very small tubercle in middle of tergum VII near posterior margin, tubercle reduced or absent in a few; posterior margin of tergum VIII with a strong spine on each side and a broad, subtruncate lobe medially, apex of medial lobe broadly and shallowly emarginate (Fig. 14). Posterior margin of male sternum VIII slightly produced into broad triangular lobe (Fig. 13). Aedeagus: Paramere as in Fig. 16. Median lobe as in Figs. 17���23, 25��� 26, long and tubular, athetine bridge broad and completely fused (Fig. 19, AB) so that apical lobe forms a long sclerotized tube. Copulatory piece (Figs. 22, 25���26) elongate and weakly sclerotized. Medial lamellae as in Figs. 21���23, 25��� 26. Spermatheca: As in Fig. 24; distal end of spermatheca bent towards spermathecal gland. Habitat. Adults are found on lignicolous gilled and polypore mushrooms. We have seen specimens collected from Pleurotus ostreatus, Bjerkendera sp., Ischnoderma fulginosum, Laetiporus sulfureus, Grifola frondosa, Ganoderma tsugae, Daedalea sp., undetermined rotting polypore mushrooms and fungus��covered logs. Fungal names were taken from specimen labels. The larval habitat is not known. Distribution. Sableta infulata is widely distributed in the eastern United States. We have seen specimens from Michigan, Illinois, Kansas, Indiana, North Carolina, South Carolina, Tennessee and Mississippi., Published as part of Ashe, James S. & Gusarov, Vladimir I., 2004, Review of the genus Sableta Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Athetini), pp. 1-15 in Zootaxa 745 on pages 8-11, DOI: 10.5281/zenodo.158570, {"references":["Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, Pennsylvania, pp. 1 - 183.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Brussels, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, 499 - 988.","Moore, I. & Legner, E. F. (1975) A catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, i - vi, 1 - 289.","Ashe, J. S. (2000) XII. Subfamily Aleocharinae Fleming 1821. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 358 - 374."]}

1. Halobrecta algophila (Fenyes, 1909) (Figs. 18–36) Atheta (Halobrectha) algophila Fenyes, 1909: 419. Atheta (s. str.) barbarae Casey, 1910: 18, syn. nov. Atheta (s. str.) importuna Casey, 1911: 111, syn. nov. Atheta (Halobrecta) algophila: Fenyes, 1920: 185 (as valid species). Atheta (Halobrecta) algophila: Bernhauer & Scheerpeltz, 1926: 621 (as synonym of At. flavipes). Atheta (s. str.) barbarae: Bernhauer & Scheerpeltz, 1926: 638 (as valid species). Atheta (s. str.) importuna: Bernhauer & Scheerpeltz, 1926: 644 (as valid species). Halobrecta algophila: Moore & Legner, 1975: 434 (as valid species). Halobrecta algophila: Seevers, 1978: 263 (as valid species). Halobrecta algophila: Ashe in Newton, Thayer, Ashe & Chandler, 2000: 369 (as valid species). Halobrecta algophila: Hammond, 2000: 275 (as valid species). Halobrecta flavipes: Pace, 2000: 376 (ex parte; as valid species; misidentification). Halobrecta flavipes: Klimaszewski, Maus & Gardiner, 2002: 483 (as valid species; misidentification). Halobrecta algophila: Klimaszewski, Maus & Gardiner, 2002: 483 (as synonym of Ha. flavipes). Type material. Lectotype of Atheta algophila (here designated): ɗ, "San Diego, Cal. [ifornia] Dr. A.Fenyes", " 2590 ", " algophila Feny. ", " algophila Fen. Cotypus Fenyes " (yellow label), "Chicago NHMus. M.Bernhauer Collection", " flavipes [pencil] not = [black pen; by Lohse] (= [pencil; crossed with a black pen; by Lohse] algophila) [pencil] no! Lohse [with a black pen; by Lohse]" (FMNH); Paralectotypes: Ψ, "San Diego, Cal. [ifornia] Dr. A.Fenyes", " 2597 ", " algophila Fen. Cotypus Fenyes " (yellow label), "Chicago NHMus. M.Bernhauer Collection" (FMNH); Ψ, "San Diego, Cal. [ifornia]", " 2315 ", " Type!", golden circle, "A.Fenyes Collection" (yellow label), " Atheta algophila Feny. ", "California Academy of Sciences. Type No. 4861 " (CASC). Holotype of Atheta barbarae: Ψ, "S. [anta] Barbara Cal. [ifornia] 2.7. 91 [7.ii. 1891]", " Atheta barbarae Csy. ", " TYPE USNM 39309 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta importuna (here designated): Ψ, "Cal. [with black dot inside C; California: San Francisco]", " importuna Csy. ", " TYPE USNM 39298 " (red label), " CASEY bequest 1925 " (NMNH). Additional material. CHILE: Palena Prov.: 2 ɗɗ, 8 specimens (sex undetermined), 4 km NW Chaiten, rocky beach drift, 31.i. 1985 (S. & J.Peck) (FMNH); FRANCE: ɗ, Ψ, “ Corsica ” (FMNH); NEW ZEALAND: ɗ, Auckland Wattle Bay, shore debris, 11.i. 1984 (P.M.Hammond) (BMNH); ɗ, 2 ΨΨ, 1 specimen (sex undetermined), Timaru, under littoral algae, 11.i. 1978 (S.Peck) (FMNH); UNITED KINGDOM: ɗ, " 41267 " (C.E.Tottenham); 2 specimens (sex undetermined), South sea, Sharp (iv. 1914); ɗ, Kent, Chatham (G.C.Champion) (BMNH); UNITED STATES: California: ɗ, without locality data (BMNH); Humboldt Co.: ɗ, Samoa, 21.vi. 1916 (F.E. Blaisdell) (CASC); Marin Co.: ɗ, Ψ, Chimney Rock, eelgrass drift, 24–30.iv. 2002 (P.M.Hammond) (BMNH, ZMUN); San Diego Co.: 7 ɗɗ, 2 ΨΨ, San Diego (A.Fenyes) (BMNH (2), CASC (5), KSEM (1), ZMUN (1)). Diagnosis. Halobrecta algophila is closely related to Ha. flavipes, but differs in having the body size smaller on average; and the spatuliform apex of the median lobe smaller (in parameral view), the apex is 1.1 times as wide as the median lobe measured subapically at its narrowest point (1.4–1.5 times in Ha. flavipes) (Figs. 22 –23, 26– 27; 41–44). Halobrecta algophila differs from the Mediterranean species tentatively identified as Ha. halensis (see Discussion for Halobrecta) in having a narrower median lobe of the aedeagus (in parameral view) (Figs. 22 –23, 26– 27; 64). Halobrecta algophila differs from the Palaearctic Ha. algae and Ha. princeps by having the posterior margin of the male tergum 8 rounded and not crenulate (Fig. 18) (in Ha. algae and Ha. princeps the posterior margin is straight and may be slightly crenulate (Fig. 52)); and by the apex of the median lobe broader (Figs. 23, 26; 56). Halobrecta algophila differs from the Oriental Ha. cingulata in having a different shape of the median lobe (in lateral view) (Figs. 24 –25, 28– 29: this paper; Fig. 9, G: Sawada 1985). Halobrecta algophila differs from the Neotropical Ha. discipula in having a different shape of the median lobe (in parameral view) (Figs. 22 –23, 26– 27: this paper; Fig. 3: Pace 1999). Description. Length 3.0– 3.9 mm. Body dark brown, legs, mouthparts and 3–4 basal antennal articles yellow. Head surface glossy, with weak isodiametric microsculpture, with strong punctation, distance between punctures equal to their diameter. Eyes 0.7–0.9 times as long as temples. Antennal article 2 slightly longer than article 3, article 4 elongate, 5 slightly elongate or subquadrate, 6 subquadrate or slightly transverse, 7–10 transverse, article 11 as long as articles 9 and 10 combined. Pronotum slightly transverse, 1.2 times as wide as head, width 0.57–0.66 mm, length 0.49–0.57 mm, width to length ratio 1.1–1.2; less glossy than head, isodiametric microsculpture stronger than on disk of head; punctation finer than on head, poorly visible on microsculptured background, distance between punctures equal to 1–2 times their diameter. Elytra much wider (0.57–0.93 mm) and longer (0.61–0.80 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.3–1.4), 0.9–1.2 times as wide as long, surface glossy, with weak isodiametric microsculpture; punctation finer than on disk of head but much stronger than on pronotum, distance between punctures equal to their diameter. Abdominal terga glossy, with poorly visible (at 70 x) microsculpture consisting of transverse waves; with fine punctation, distance between punctures equals 2–4 times their diameter on terga 3–5 and 2–6 times on terga 6–7. Posterior margin of male tergum 8 rounded and not crenulate (Fig. 18). Aedeagus as in Figs. 22 –31, 33– 36. Spermatheca as in Fig. 32. Discussion. The types of At. barbarae and At. importuna (both females) are similar in external characters to the examined types of Ha. algophila as well as in the shape of the spermatheca. Although Ha. algophila and Ha. flavipes cannot be reliably distinguished without examination of the male genitalia, I consider At. barbarae and At. importuna to be synonyms of Ha. algophila because all examined males from the West Coast belong to that species. The specimens from Inaccessible Island of the Tristan da Cunha archipelago identified by Klimaszewski et al. (2002) as Ha. flavipes seem to be misidentified. The (ab)parameral view of the median lobe (Fig. 1: Klimaszewski et al., 2002) does not agree with that of Ha. flavipes (Figs. 41–44: this paper;) but fits Ha. algophila (Figs. 22 –23, 26– 27: this paper). The examined specimens from Chaiten, Chile, listed by Pace (2000) as Ha. flavipes belong in fact to Ha. algophila. Distribution. Halobrecta algophila is known from the coasts of California, Europe, Chile, New Zealand and Inaccessible Island of the Tristan da Cunha archipelago (Klimaszewski et al., 2002). Natural History. Halobrecta algophila is a marine littoral species., Published as part of Gusarov, Vladimir I., 2004, A revision of the Nearctic species of the genus Halobrecta Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae) with notes on some Palaearctic species of the genus, pp. 1-25 in Zootaxa 746 on pages 8-13, DOI: 10.5281/zenodo.158213, {"references":["Fenyes, A. (1909) New Aleocharinae (Staphylinidae, Col.) of the U. S. Entomological News, 20, 1909, 418 - 425.","Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183.","Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 1 - 245.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Moore, I. & Legner, E. F. (1975) A catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp.","Newton, A. F., Thayer, M. K., Ashe, J. S. & Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418.","Hammond, P. M. (2000) 16. Coastal Staphylinidae (rove beetles) in the British Isles, with special reference to saltmarshes. In: Sherwood, B. R., Gardiner, B. G. & Harris T. (Eds.), British Saltmarshes. Joint Symposium on British Saltmarshes organized between the Linnean Society of London, the Royal Society for the Protection of Birds and English Nature (2000: London). Forrest Text, Cardingan, pp. 247 - 302.","Pace, R. (2000) Aleocharinae del Cile, nuove o poco note (Coleoptera, Staphylinidae). Bollettino del Museo regionale di Scienze naturali, Torino, 17 (2), 355 - 510.","Klimaszewski, J., Maus, C. & Gardiner, A. (2002) The importance of tracking introduced species: New records of athetine rove beetles from South Atlantic Inaccessible Island (Coleoptera, Staphylinidae, Aleocharinae). Coleopterists Bulletin, 56 (4), 481 - 490.","Sawada, K. (1985) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). IV. Ten oriental species described by M. Bernhauer and M. Cameron. Contributions from the Biological Laboratory, Kyoto University, 27 (1), 91 - 110.","Pace, R. (1999) Aleocharinae del Cile (Coleoptera, Staphylinidae). Bollettino del Museo civico di Storia naturale di Verona, 23, 119 - 210."]}

3. Lypoglossa franclemonti Hoebeke, 1992 (Figs. 64���70) Lypoglossa fenyesi: Lohse & Smetana, 1985: 294 (nec Bernhauer, 1907: 402) (Misidentification). Lypoglossa franclemonti Hoebeke, 1992: 386. Type material. Holotype: UNITED STATES: New York: Essex Co.: ��, Mt. Whiteface, 2000���4000 ', 22.viii. 1916 (CUIC). Paratypes: UNITED STATES: Maine: Washington Co.: 7 specimens, Carrying Place Cove, Lubec, 23.vii��� 24.viii. 1989 (H.Fraembs) (CUIC); New Hampshire: Coos Co.: ��, Bretton Woods (A.Fenyes) (CUIC); 1 specimen, White Mts., Glen Ellis Falls, 1900 ', 15.viii. 1976 (J.M. & B.A.Campbell) (CNCI); 1 specimen, White Mts., Glen Boulder Trail, 2500 ', 17.viii. 1976 (J.M. & B.A.Campbell) (CNCI); New York: Essex Co.: �� (allotype), Artist's Brook, 11.vi. 1933 (H.Dietrich) (CUIC); CANADA: Nova Scotia: 1 specimen, Cape Breton Hist. Nat. Park, 410 m, Benjies Lake area, PG 680786, sifting mosses, 18.ix. 1984 (J.M. Campbell & A.Davies) (CNCI); ��, ditto but 440 m, near Sammys Barren, PG 764861, sifting moss and Alnus litter, 24.ix. 1984 (CNCI); Quebec: 1 specimen, Mistassini Post, 13.vi. 1956 (J.R.Lonsway) (CNCI); 4 specimens, Parc Gaspesie, Mont Albert, 1000 ', 9.vii. 1972 (J.M. & B.A.Campbell); 2 specimens, Parc Gaspesie, Lac du Diable, 1500 ', 12.vii. 1972 (J.M. & B.A.Campbell) (CNCI); 1 specimen, ditto but Lac St. Anne (CNCI); 1 specimen, Parc Gaspesie, Lac Cascapedia, 1700 ', 14.vii. 1972 (J.M. & B.A.Campbell) (CNCI); 2 specimens, Parc des Laurentides, Mare��du��Sault, 2700 ', 15���17.viii. 1970 (J.M. & B.A.Campbell) (CNCI); Ontario: 9 specimens, Superior Prov. Park, Gargantua, 7.vi. 1973 (J.M.Campbell & R.Parry) (CNCI); 1 specimen, ditto but 9.vi. 1973 (CNCI); 3 specimens, ditto but Frater (CNCI); 1 specimen, ditto but 13.vi. 1973 (CNCI); 6 specimens, ditto but Agawa Bay (CNCI); 3 specimens, ditto but Old Woman Bay (CNCI); 1 specimen, ditto but Baldhead Lake, 8.vi. 1973 (J.M.Campbell & R.Parry) (CNCI); 5 specimens, 36 mi. S Pickle Lake, 22.vi. 1973 (Campbell & Parry) (CNCI); 1 specimen, 21 mi. N Pickle Lake, 19.vi. 1973 (Campbell & Parry) (CNCI); 1 specimen, ditto but 41 mi. N Pickle Lake (CNCI); 1 specimen, 49 mi. N Savant Lake, 23.vi. 1973 (Campbell & Parry) (CNCI); 2 specimens, 46 mi. N Hurkett, Black Sturgeon Lake, 29.vi. 1973 (R.Parry & J.M.Campbell) (CNCI); 5 specimens, Moosonee, 2.vii. 1973 (Parry & Campbell) (CNCI); 2 specimens, ditto but 30.vi. 1973 (CNCI); Manitoba: 6 specimens, Riding Mtn. Nat. Park, Moon Lake, 17.ix. 1979 (A.Smetana) (CNCI); 4 specimens, ditto but near Deep Lake, 7.ix. 1979; 1 specimen, ditto but Whirlpool River at Hwy. 19, 6.ix. 1979 (CNCI); Alberta: 8 specimens, George Lake, 53 �� 57 'N 114 ��06'W, moss and lichens, on rotten logs from black spruce bog, 1.vi. 1980 (J.S.Ashe); 1 specimen, ditto but berlese extract of sifted litter from birch��� Ledum bog, 12.vi. 1980; 2 specimens, ditto but berlese extract of sifted litter from spruce bog, 16.vi. 1980; 2 specimens, ditto but 2.viii. 1978; 2 specimens, ditto but berlese extract of sifted litter from birch bog, 10.viii. 1980; ��, ditto but pan trap, 5���22.v. 1970 (G.Gibson) (all ��� KSEM); 2 specimens, ditto but 1.x. 1966 (R.E.Leech) (CNCI); 2 specimens, ditto but 25.ix. 1966 (CNCI); 1 specimen, ditto but 10.ix. 1966 (CNCI); 3 specimens, Lusk Creek, Kananaskis F.E.S., 14.vii. 1971 (J.M. & B.A.Campbell) (CNCI); North West Territory: ��, 5 mi. SE Fort Simpson, Hwy. 3, 21.vi. 1972 (A. Smetana) (CNCI). Additional material. UNITED STATES: Vermont: Bennington Co.: 2 ����, 2 ����, Big Equinox Mtn., 3800 ', spruce���fir forest, berlese extract of litter, 25.ix. 1976 (A.Newton & M.Thayer) (FMNH); Franklin Co.: ��, E slope of Crocker Mt., 13 km SE Stratton, Appalachian Trail, 45 ��02.2'N 70 �� 21.9 W, 950 m, forest litter, Picea, Abies, Betula, 23.viii. 2001 (V.I.Gusarov) (ZMUN); New York: Essex Co.: ��, Mt. Marcy, 1.x (FMNH); Hamilton Co.: ��, Adirondack Park, 9 km SE Deerland, 2 km E Hwy. 28 N & 30, W slope of Blue Mountain, 43 �� 52.19 'N 74 �� 24.30 W, 1100 m, forest litter, Abies, Betula, Sorbus, Oxalis, 27.viii. 2001 (V.I.Gusarov) (ZMUN). Diagnosis. Lypoglossa franclemonti differs from the Palaearctic L. lateralis in having pronotum and elytra lighter than head and abdomen; punctation of head and pronotum not asperate; more elongate antennal articles; and in the shape of the aedeagus and spermatheca (Figs. 64���70; 72���78). Lypoglossa franclemonti can be distinguished from L. angularis by a less transverse pronotum, more elongate antennal articles; the apex of the median lobe (in parameral view) broadest at the base, not constricted (Figs. 64���65; 25���27); no projections at the base of the apex (Figs. 64���67; 25���29); a smaller sclerite of the parameres (Figs. 69; 31); and the proximal loop of the spermatheca wider, almost circular (Figs. 70; 32). Lypoglossa franclemonti differs from L. manitobae in having a larger body size; apex of the median lobe (in parameral view) broadest at the base, not constricted (Figs. 64���65; 55���57); no projections at the base of the apex (Figs. 64���67; 55���59); a smaller sclerite of parameres (Figs. 69; 61); and a larger spermatheca (Figs. 70; 62���63). Description. Length 3.8���4.6 mm. Head and abdomen brown, pronotum and elytra, legs, antennae and mouthparts reddish brown to brownish red, in some specimens elytra near scutellum and along suture darker. Head surface glossy, with weak isodiametric microsculpture, with weak punctation, distance between punctures equal to 1���1.5 times their diameter. Eyes 0.9���1.1 times as long as temples. All antennal articles elongate or articles 9���10 subquadrate. Pronotum transverse, 1.5 times as wide as head, width 0.90���1.09 mm, length 0.71��� 0.89 mm, width to length ratio 1.2���1.3; surface glossy, with weak isodiametric microsculpture; punctation finer and denser than on head, distance between punctures equals their diameter. Elytra wider (0.96���1.21 mm) than pronotum and as long (0.70���0.87 mm; measured from humeral angle) as pronotum, 1.3 times as wide as long, surface glossy, with poorly visible microsculpture; punctation slightly asperate and denser than on pronotum, distance between punctures equal to ����� 1 times their diameter. Abdominal terga with very fine (poorly visible at 70 x) microsculpture consisting of transverse waves; terga 3���5 matte due to fine and dense punctation, distance between punctures equals 1���2 times their diameter, terga 6���7 glossy, with sparser punctation, distance between punctures equals 2���4 times their diameter. In males, head with weak medial impression and pronotum with broad impression along midline. Aedeagus as in Figs. 64���69. Spermatheca as in Fig. 70. Distribution. Lypoglossa franclemonti is distributed in Canada and in the northeastern United States. It is known from the Northwest Territory, Alberta, Manitoba, Ontario, Quebec, Nova Scotia, New York, Vermont, New Hampshire and Maine (Fig. 71). In different parts of its range L. franclemonti is sympatric with L. manitobae (in Manitoba), L. angularis obtusa (in the North��East) and L. angularis fenyesi (in Alberta). Compared to L. angularis, L. franclemonti tends to be restricted to lower altitudes and more continental conditions. Natural History. Lypoglossa franclemonti is common in forest litter and in moss on forest floor., Published as part of Gusarov, Vladimir I., 2004, A revision of the genus Lypoglossa Fenyes, 1918 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-36 in Zootaxa 747 on pages 28-31, DOI: 10.5281/zenodo.158194, {"references":["Hoebeke, E. R. (1992) Taxonomy and distribution of the athetine genus Lypoglossa Fenyes (Coleoptera: Staphylinidae: Aleocharinae) in North America, with description of a new species. Journal of the New York Entomological Society, 100 (2), 381 - 398.","Lohse, G. A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Maklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39 (3), 281 - 300.","Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405."]}

Halobrecta Thomson, 1858 (Figs. 1–65) Halobrecta Thomson, 1858: 35 (type species Homalota puncticeps Thomson, 1852, by monotypy). Halobrectha: Thomson, 1861: 49 (as valid genus; incorrect subsequent spelling). Glaphya Mulsant & Rey, 1873: 172 (as subgenus of Dinaraea Thomson, 1858; type species Dinaraea pubes Mulsant & Rey, 1873, by monotypy). Halobrechta: Mulsant & Rey, 1875: 35 (as valid genus; incorrect subsequent spelling). Atheta (Halobrecta): Fenyes, 1920: 185 (as valid subgenus). Glaphya: Fenyes, 1920: 185 (as synonym of Atheta (Halobrecta)). Exatheta Cameron, 1920: 265 (type species Exatheta cingulata Cameron, 1920, by subsequent designation (Blackwelder 1952)). Atheta (Halobrecta): Bernhauer & Scheerpeltz, 1926: 621 (as valid subgenus in subtribe Athetina Casey, 1910). Glaphya: Bernhauer & Scheerpeltz, 1926: 621 (as synonym of Atheta (Halobrecta)). Exatheta: Bernhauer & Scheerpeltz, 1926: 681 (as valid genus in subtribe Schistogeniina Fenyes, 1918). Atheta (Halobrecta): Scheerpeltz, 1934: 1600 (as valid subgenus). Halobrecta: Blackwelder, 1952: 179 (as valid genus). Glaphya: Blackwelder, 1952: 170 (as synonym of Halobrecta). Exatheta: Blackwelder, 1952: 163 (as valid genus). Halobrecta: Benick & Lohse, 1974: 219 (as valid genus in tribe Callicerini Lohse, 1969). Halobrecta: Moore & Legner, 1975: 434 (as valid genus). Halobrecta: Seevers, 1978: 121 (as valid genus in subtribe Xenotae Seevers, 1978 (nomen nudum)). Halobrecta: Sawada, 1985: 108 (as valid genus in Coprothassa series). Exatheta: Sawada, 1985: 108 (as synonym of Halobrecta). Halobrecta: Lohse, 1989: 219 (as valid genus in tribe Athetini). Halobrecta: Ashe in Newton, Thayer, Ashe & Chandler, 2000: 369 (as valid genus in subtribe Athetina Casey, 1910). Diagnosis. Halobrecta is distinguished from other athetine genera by the combination of the following characters: body parallel­sided; anterior margin of labrum straight; sensilla a of epipharynx long; antennal article 2 longer than article 3, articles 7–10 transverse; ligula long, parallel­sided, with narrow base and slightly split apically (Fig. 6); labial palpus with setae and present (Fig. 6); pronotum slightly transverse, 1.1–1.2 times as wide as long, with microsetae directed anteriorly along anterior half of midline, and posteriorly along posterior half of midline; in lateral portions of the disc microsetae directed laterally (Type III, Benick & Lohse 1974) (Fig. 14); pronotal macrosetae long; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia thin, as long as tibial width; mesothoracic process narrow (Fig. 15); posterior margin of elytra emarginate; tarsal formula 4­5 ­ 5; metatarsal segment 1 longer than segment 2; single empodial seta longer than claws; abdominal terga 3–5 with transverse basal impression; posterior margin of female tergum 8 with comb of scattered tiny projections (Figs. 20, 39); median lobe of aedeagus without athetine bridge, apex of paramere long and narrow, medial lamellae of internal sac absent; copulatory piece with pointed apex (Figs. 30, 41, 62) and without sclerotized suspensoria; spermatheca short, not divided into distal and proximal portions, with large umbilicus (Figs. 32, 51). Halobrecta differs from other littoral aleocharines with the same tarsal formula (Adota Casey, 1910, Psammostiba Yosii & Sawada, 1976, Pontomalota Casey, 1885, Tarphiota Casey, 1894) in having a different type of pronotal pubescence (Fig. 14); the posterior margin of the elytra emarginate; the posterior margin of the female tergum 8 with a comb of scattered tiny projections (Figs. 20, 39); the median lobe without the athetine bridge (Figs. 25, 28); the paramere with a long apex (Figs. 36, 50) and a short spermatheca with a large umbilicus (Figs. 32, 51). Description. Length 2.9–3.9 mm, pronotal width 0.53–0.66 mm. Body parallelsided, dark brown, with legs, basal or all antennal articles, and mouthparts yellowish brown to yellow. Head transverse; eye length to temple length ratio 0.7–1.2; infraorbital carina complete. Antennal article 2 longer than article 3, article 4 elongate, 5 elongate or subquadrate, 6 subquadrate or transverse, 7–10 transverse, apical article without coeloconic sensilla, as long as articles 9 and 10 combined. Labrum (Fig. 1) transverse, with straight anterior margin. Epipharynx (Fig. 2) with long sensilla a, with three pairs of small marginal setae, medial field with 32 pores, lateral rows with two pores each, anterolateral groups with three pores each, transverse row with six pores, posterolateral groups with three–four pores each, with one medial proximal pore on each side and two lateral proximal pores. Mandibles (Figs. 3–5) broad, right mandible with a small medial tooth; velvety patch of dorsal molar area not visible at 400 x. Maxilla (Figs. 8–11) with galea projecting slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; internal margin of galea with long subapical setae (Fig. 9); distal comb of lacinia is divided into isolated groups of 5 and 2 closely placed spines (Figs. 10–11), middle portion produced medially and covered with numerous fine setae (Figs. 10–11), ventral surface of lacinia with a marginal group of 4 strong setae (Fig. 10), dorsal surface of lacinia with a row of 18 weak setae (Fig. 11). Labium as in Figs. 6 –7, 12; ligula long and parallel­sided, with narrow base, slightly split apically (Fig. 6); medial area of prementum with 2 pores and 12 pseudopores, lateral areas each with two asetose pores, single setose pore and 9–10 pseudopores (Fig. 6). Hypopharyngeal lobes as in Fig. 7. Labial palpus with setae and present (Fig. 6). Mentum (Fig. 12) with concave anterior margin. Pronotum (Fig. 14) slightly transverse, 1.1–1.2 times as wide as long, with microsetae directed anteriorly along anterior half of midline, posteriorly along posterior half of midline, and laterally in lateral portions of the disc (Type III, Benick & Lohse 1974) (Fig. 14); macrosetae long; hypomera fully visible in lateral view. Meso­ and metasternum as in Fig. 15, mesosternal process narrow, extending about 3 / 5 length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 3: 1: 1; mesocoxal cavities margined posteriorly; mesocoxae contiguous. Medial macroseta of mesotibia thin, as long as tibial width. Tarsal segmentation 4­5 ­ 5, metatarsal segment 1 longer than segment 2 (Fig. 17). One empodial seta, longer than claws (Fig. 16). Posterior margin of elytra emarginate. Wings fully developed. Abdominal terga 3–5 with moderate basal impressions. Tergum 7 is as long as tergum 6. Punctation on terga 6–7 finer and slightly sparser than on terga 3–5. Tergum 7 with wide white palisade fringe. Posterior margin of female sternum 8 with comb consisting of tiny projections (Fig. 21). Posterior margin of female tergum 8 with irregular comb of scattered projections (Fig. 20, 39) (in some males this comb is also present but may consist of just a few projections). Median lobe of aedeagus without athetine bridge (Figs. 25, 28), internal sac without medial lamellae (Figs. 62–65); copulatory piece with pointed apex (Figs. 30, 62, 64) and without sclerotized suspensoria; paramere with long and narrow apex (Figs. 33–36); spermatheca short, not divided into distal and proximal portions, with large umbilicus (Figs. 32, 51). Type species. Homalota puncticeps Thomson, 1852, by monotypy (Thomson 1858). Discussion. Halobrecta is usually placed in the tribe Athetini but it lacks the athetine bridge of the median lobe which is considered as an autapomorphy of Athetini (Seevers 1978; Muona 1987; Newton et al. 2000) although this character is also present in the tribes Lomechusini Fleming (1821) and Falagriini Mulsant & Rey (1874). The lack of the athetine bridge in Halobrecta may suggest that it belongs to Oxypodini Thomson (1859). Another character traditionally used to distinguish between the aleocharine tribes is the tarsal formula (e.g., Lohse 1974). In Halobrecta it is 4­5 ­ 5 and supports the placement of the genus in Athetini. However there are examples of changes in tarsal formula even within a genus (Gyronycha Casey, 1894 (Seevers 1978), Microlia Casey, 1910 (Gusarov 2002 a)). In addition to lacking the athetine bridge, Halobrecta is similar to some Oxypodini in having the apex of the paramere long; the spermatheca short with a large umbilicus; the posterior margin of the elytra emarginate; the ligula bisetose, long and narrow, split only at the very apex. The tribal placement of Halobrecta cannot be resolved until the phylogeny of Athetini and Oxypodini is analyzed. According to a recent review of the British intertidal beetles (Hammond 2000) Halobrecta includes at least four valid species, all of them recorded from the British Isles: Ha. algae (Hardy, 1851), Ha. flavipes Thomson, 1861, Ha. princeps (Sharp, 1869) and Ha. algophila (Fenyes, 1909). Two species from Singapore were described by Cameron (1920) in the genus Exatheta Cameron, 1920 (E. cingulata and E. consors) and subsequently transferred to Halobrecta and synonymized with each other by Sawada (1985, 1987). Pace (1999) described Halobrecta discipula from Chile. I am aware of an additional species of Halobrecta (AUSTRALIA: ɗ, Victoria (FMNH); GREECE: ɗ, 5 specimens (sex undetermined), Kérkira (J.Sahlberg); ɗ, Elevsís (J.Sahlberg) (FMNH); ITALY: 18 specimens (sex undetermined), Friuli, Lignano, 25.v. 1929 (A.Gagliardi); ɗ, 1 specimen (sex undetermined), Fiumicino, 7.vi. 1898 (FMNH); MACEDONIA: ɗ, Vardar (FMNH); YUGOSLAVIA: 5 specimens (sex undetermined), Sutorina (Paganetti); ɗ, Ψ, Sutorina near Castelnuovo; ɗ, 2 specimens (sex undetermined), Castelnuovo (Hummler) (FMNH)). This species differs from the other four species in the shape of the median lobe (Figs. 64– 65) and may be conspecific with Ha. halensis Mulsant & Rey, 1873 described from the coast of Languedoc (France)., Published as part of Gusarov, Vladimir I., 2004, A revision of the Nearctic species of the genus Halobrecta Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae) with notes on some Palaearctic species of the genus, pp. 1-25 in Zootaxa 746 on pages 2-7, DOI: 10.5281/zenodo.158213, {"references":["Thomson, C G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40.","Thomson, C. G. (1852) Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 9 (6), 131 - 146.","Thomson, C. G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom III. Berlingska Boktryckeriet, Lund, 278 pp.","Muslant, E. & Rey, C. (1873) Description de divers Coleopteres Brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189.","Muslant, E. & Rey, C. (1875) Tribu des Brevipennes. Famille des Aleochariens (suite). Septieme branche: Myrmedoniaires. 2 e partie. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4, 7 [1874], 27 - 496, Pl. vi - ix.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284.","Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae with an essay on genotypy. United States National Museum Bulletin, 200, i - iv + 483 pp.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183.","Fenyes, A. (1918) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110.","Benick, G. & Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 72 - 220.","Lohse, G. A. (1969) Vorschlage zur Anderung der Aleocharinensystematik (Coleoptera: Staphylinidae). In: Bericht uber die 10. Wanderversammlung deutscher Entomologen, 15. Bis 19. September 1965 in Dresden. Deutsche Akademie der Landwirtschaftswissenschaften zu Berlin, Berlin, pp. 169 - 175.","Moore, I. & Legner, E. F. (1975) A catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp.","Sawada, K. (1985) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). IV. Ten oriental species described by M. Bernhauer and M. Cameron. Contributions from the Biological Laboratory, Kyoto University, 27 (1), 91 - 110.","Lohse, G. A. (1989) Erganzungen und Berichtigungen zu Freude - Harde - Lohse \" Die Kafer Mitteleuropas \" Band 5 (1974) In: Lohse, G. A. & Lucht, W. H. (Eds.), Die Kafer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke & Evers Verlag, Krefeld, pp. 185 - 243.","Newton, A. F., Thayer, M. K., Ashe, J. S. & Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418.","Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140.","Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1 (4), 283 - 336.","Casey, T. L. (1894) Coleopterological notices. V. Annals of the New York Academy of Sciences, 7 [1893], 281 - 606.","Muona, J. (1987) Some aspects of Aleocharinae systematics - a response to Dr. G. A. Lohse. Entomologische Blatter, 83 (1), 19 - 24.","Fleming, J. (1821) Insecta. In: Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica, with preliminary dissertations on the history of the sciences. Vol. 5. Archibald Constable and Company, Edinburgh, pp. 41 - 56.","Muslant, E. & Rey, C. (1874) Tribu des Brevipennes. Famille des Aleochariens. Septieme branche: Myrmedoniaires. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4. 6 [1873], 33 - 738.","Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 1. Berlingska Boktryckeriet, Lund, v + 290 pp.","Gusarov, V. I. (2002 a) A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini). Zootaxa, 34, 1 - 24.","Hammond, P. M. (2000) 16. Coastal Staphylinidae (rove beetles) in the British Isles, with special reference to saltmarshes. In: Sherwood, B. R., Gardiner, B. G. & Harris T. (Eds.), British Saltmarshes. Joint Symposium on British Saltmarshes organized between the Linnean Society of London, the Royal Society for the Protection of Birds and English Nature (2000: London). Forrest Text, Cardingan, pp. 247 - 302.","Hardy, J. (1851) Family 1. Staphylinidae, Leach. In: Hardy, J. & Bold, T. J. A Catalogue of the Insects of Northumberland and Durham (Part ii). Transactions of the Tyneside Naturalists' Field Club, 2, 21 - 97.","Sharp, D. (1869) A revision of the British species of Homalota. Transactions of the Entomological Society of London, 1869 (2 - 3), 91 - 272.","Fenyes, A. (1909) New Aleocharinae (Staphylinidae, Col.) of the U. S. Entomological News, 20, 1909, 418 - 425.","Sawada, K. (1987) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). V. Singaporean species described in Cameron, 1920. Contributions from the Biological Laboratory, Kyoto University, 27 (2), 137 - 150.","Pace, R. (1999) Aleocharinae del Cile (Coleoptera, Staphylinidae). Bollettino del Museo civico di Storia naturale di Verona, 23, 119 - 210."]}

14. Seeversiella scabricollis Gusarov, sp. n. (Figs. 11, 15, 190�� 207) Type material. Holotype,, COSTA RICA: Cartago: Sendero a Cerro Chirrip��, 9 �� 26 ' 47 "N 83 �� 32 ' 12 "W, 2800 m, mixed oak forest litter (R.Anderson), 27.vi. 1999 (KSEM). Paratypes: COSTA RICA: Cartago: 42 specimens, same data as the holotype; 50 specimens, ditto but 9 �� 26 ' 37 "N 83 �� 32 ' 14 ", 2950 m, oak forest litter;, Cerro Chirrip��, Base Crestones, 9 �� 27 ' 17 "N 83 �� 30 ' 16 "W, 3350 m, elfin forest litter (R.Anderson), 25.vi. 1999; 17 specimens, 2 km S Villa Mills, 3000 m, ridgetop oak forest litter (R.S.Anderson), 10.ii. 1996;, 2.0 km E Villa Mills, 9 �� 34 '00"N 83 �� 41 ' 50 "W, 2750 m, oak forest litter (R.S.Anderson), 15.ii. 1998;, Cerro Buenavista, 9 �� 43 '02"N 83 �� 45 ' 14 "W, 3300 m, subparamo litter (R.Anderson); 10 specimens, Cerro de la Muerte, 4 km NE Ca��on, 2350 m, malaise trap (P.Hanson), i. 1995; 2 specimens, ditto but 1.i�� 30.iii. 1995; 3 specimens, ditto but 1��30.iv. 1995; 1 specimen, ditto but 1��30.v. 1995; 2 specimens, ditto but 1��30.vi. 1995; 6 specimens, ditto but 1��30.vii. 1995; 2 specimens, ditto but 1�� 30.viii. 1995; 1 specimen, ditto but 1��30.i. 1996; 1 specimen, ditto but 1.iv�� 30.v. 1996; 7 specimens, ditto but 1��30.vii. 1996; 3 specimens, ditto but 1��30.ix. 1996; 4 specimens, 2 km N P. N. Volc��n Iraz��, 10 ��00'00"N 83 �� 51 '00"W, 2900 m, berlese forest litter (R.Anderson), 18.vi. 1997; 6 specimens, 3 km N P. N. Volc��n Iraz��, 9 �� 59 ' 45 "N 83 �� 51 '00"W, 2750 m, berlese forest litter (R.Anderson), 18.vi. 1997;, Madreselva, Finca Los Lagos, A. C. Amistad, 2000��2600 m, 12��19.vi. 1993; 2, ditto but 28.vi�� 10.vii. 1993;, ditto but ix. 1993;, ditto but x. 1993; San Jose / Cartago: 11 specimens, km 71, Int. Amer. Hwy., nr. Tres de Junio, 9 �� 37 ' 44 "N 83 �� 50 ' 13 "W, 2900 m, wet cloud forest litter (R.Anderson), 23.vi. 1999; 3, km 69, Int. Amer. Hwy., nr. Tres de Junio, 9 �� 39 ' 30 "N 83 �� 51 ' 30 "W, 2600 m, flight intercept trap (S. & J.Peck), 7��19.vi. 1997; 11 specimens, km 78, Int. Amer. Hwy., nr. Ojo de Agua, 9 �� 36 ' 30 "N 83 �� 50 ' 20 "W, 2800 m, cloud forest litter (R.Anderson), 8.vii. 1999; 9 specimens, km 72, Int. Amer. Hwy., 3 km W Ojo de Agua, 9 �� 37 ' 30 "N 83 �� 50 ' 30 "W, 2950 m, berlese forest litter (R.Anderson), 7.vi. 1997;, km 87, Int. Amer. Hwy., nr. Cerro Buenavista, 9 �� 36 ' 30 "N 83 �� 46 '00"W, 3150 m, berlese forest litter (R.Anderson), 8.vi. 1997; 8 specimens, Cerro Buenavista, km 89, Int. Amer. Hwy., 9 �� 33 '00"N 83 �� 45 ' 30 "W, 3200 m, berlese leaf litter (R.Anderson), 18.vi. 1998;, 2, km 55, Int. Amer. Hwy., 3 km S El Empalme, 9 �� 42 ' 30 "N 83 �� 57 '00"W, 2350 m, berlese forest litter (R.Anderson), 8.vi. 1997; San Jose: 15 specimens, Estaci��n Cuerici, 4.6 km E Villa Mills, 9 �� 34 '00"N 83 �� 40 '00"W, 2600 m, berlese forest litter (R.Anderson), 26.vi. 1997; 3 specimens, ditto but 19.vi. 1997; , 2.0 km E Villa Mills, 9 �� 33 ' 30 "N 83 �� 42 '00"W, 2800 m, berlese forest litter (R.Anderson), 26.vi. 1997;, 2, km 97, Int. Amer. Hwy., Villa Mills, 3000 m, flight intercept traps (S. & J.Peck), 26.vi. 1997; 3,, Cerros de Escaz��, 2 km S San Antonio, 9 �� 53 ' 30 "N 84 ��09'00"W, 1650 m, berlese forest litter (R.Anderson), 13.vi. 1997; 8 specimens, Cerro de la Muerte, 6 km N San Gerardo, 2800 m (P.Hanson), viii��ix. 1992; 4 specimens, ditto but x��xii. 1992; 2 specimens, ditto but iv. 1992; 2, malaise trap, ditto but iv. 1993;, Cerro de la Muerte, 19 km S and 3 km W of Empalme, 2600 m, malaise trap (P.Hanson), ix��x. 1992; 7, 26 km N San Isidro, 2100 m (P.Hanson), xi. 1992 �� i. 1993;, ditto but vi��vii. 1990;, ditto but ix��x. 1992; 2 specimens, 2 km W Empalme, Cerro de la Muerte, 2300 m (P.Hanson), 1��30.vii. 1995, malaise trap; Heredia: 2, 2, 3 specimens, Porrosati, 6 km N San Jos�� de la Montana, 10 ��05' 30 "N 84 ��07'00"W, 1900 m, berlese forest litter (R.Anderson); 44 specimens, P. N. Braulio Carrillo Volc��n Barva, 10 ��07' 30 "N 84 ��07' 30 "W, 2600 m, berlese forest litter (R.Anderson), 11.vi. 1997; 21 specimens, ditto but, flight intercept trap (S. & J.Peck), 11��27.vi. 1997; 4 specimens, Vara Blanca, Finca Georgina, 2100 m (P.Hanson), vi��vii. 1990; Alajuela: 28 specimens, P. N. Volc��n Po��s, 10 �� 11 ' 30 "N 84 �� 14 '00"W, 2500 m, berlese forest litter (R.Anderson), 6.vi. 1997;,, ditto but flight intercept trap (S. & J.Peck), 6��28.vi. 1997; PANAMA: Chiriqu��: 31 specimens, 5.9 km NE Cerro Punta, Par. Nac. Volc��n Bar��, 8 �� 52 'N [the latitude on the label (8 �� 22 '0"N) is incorrect] 82 �� 34 'W, 2100 m, flight intercept traps (J.S.Ashe & R.Brooks), 14��16.vi. 1995;, ditto but 2400 m, bamboo forest litter (R.S.Anderson), 14.vi. 1995; 12 specimens, ditto but 2150 m, alder forest litter (R.S.Anderson), 14.vi. 1995; , 27.7 km W Volc��n Hartmann's Finca, 1450 m, 8 �� 51 ' 48 "N 82 �� 44 ' 36 "W, 1450 m, forest litter (R.S.Anderson), 14.vi. 1995; 22 specimens, NNE of Las Nubes @ continental divide, 2490 m (A.R.Gillogly), 14��15.vi. 1995; 5 specimens, N of Las Nubes, 2350��2450 m, continental divide (A.R.Gillogly), 15.vi. 1995; 2, Cerro Pando, 1875 m, 8 �� 54 ' 42 "N 82 �� 43 ' 18 "W, 1875 m, flight intercept trap (J.S.Ashe, R.Brooks), 17��18.vi. 1996;, ditto but 1850 m; 6 specimens, 11 km NW Boquete, Volc��n Bar��, 2150 m, 8 �� 48 '00"N 82 �� 29 '00"W, 2150 m, oak forest litter (R.S.Anderson), 18.vi. 1995 (all �� KSEM). Diagnosis. Seeversiella scabricollis can be distinguished from other species of Seeversiella by having dark brown body with brown elytra, legs and mouthparts; large eyes (temple length to eye length ratio 0.9��1.1); matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (Figs. 194��204, 206 �� 207) and the shape of spermatheca (Fig. 205). Seeversiella scabricollis differs from S. furcativentris in having parallel��sided apex of median lobe (in parameral view) (Figs. 194��196; 212��213). Seeversiella scabricollis differs from S. brunnea in having straight apex of median lobe (in lateral view) (Figs. 197��198; 241��242). Seeversiella scabricollis differs from S. brevipennis in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; apex of median lobe parallelsided (in parameral view) (Figs. 194��196; 226��227). Seeversiella scabricollis differs from S. curtipennis in having larger eyes; longer elytra; wings longer than elytra; posterior margin of male tergum 8 with medial emargination (Fig. 190); narrow apex of the median lobe (in parameral view) (Figs. 194��196; 251�� 252); short medial lamellae of the internal sac (Figs. 201; 258); lateral diverticula with sclerotized denticles (Figs. 200, 207; 259); and shorter spermatheca (Figs. 205, 255). CP ��� copulatory piece; LD ��� lateral diverticulum of internal sac. Description. Length 2.7��3.1 mm. Body dark brown, with brown elytra, legs and mouthparts. Head surface matte, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2 times their diameter. Temple length to eye length ratio 0.9��1.1. Antennal article 2 longer than article 3, article 4 subquadrate, 5 slightly transverse, 6��10 transverse (ratio 1.5 ��2.0). Pronotum slightly transverse, 1.2 times as wide as head, width 0.49��0.54 mm, length 0.41��0.46 mm, width to length ratio 1.2, surface matte, with fine isodiametric microsculpture; punctation fine and weak, distance between punctures equal to their diameter. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.3 times as wide as long, glossy, with fine and weak isodiametric microsculpture; with fine punctation, distance between punctures equal to ���� 1 time their diameter. Wings fully developed or partially reduced, longer than elytra. Abdominal terga glossy, with fine microsculpture consisting of strongly transverse meshes, with fine punctation, distance between punctures equals 2��4 times their diameter on terga 3��5 and 4��7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (Figs. 190, 192). In males posterior angles of tergum 3 projecting as spines up to 3 times as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to 2 / 3 as long as tergum length; posterior margin of tergum 8 with medial emargination delimited by short dentiform projections (Fig. 190). Aedeagus as in Figs. 194��204, 206 �� 207. Apex of paramere broad, proximal seta approximately as long as the other three setae (Fig. 202). Spermatheca as in Fig. 205. Distribution. Known from Costa Rica and Panama (Fig. 388). Natural History. Seeversiella scabricollis was collected in leaf litter and with flight intercept and malaise traps, at altitudes of 1450��3300 m., Published as part of Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-102 in Zootaxa 142 on pages 50-55, DOI: 10.5281/zenodo.156420

1. Seeversiella globicollis (Bernhauer, 1907) (Figs. 1 ��10, 13, 17, 20�� 36) Atheta (Microdota) globicollis Bernhauer, 1907: 388. Atheta (Microdota) globicollis: Bernhauer & Scheerpeltz, 1926: 632. Sipalia cristata Fenyes, in litteris (manuscript name). Seeversiella bispinosa Ashe, 1986: 503, syn. nov. Type material. Lectotype of A. globicollis (here designated): CANADA: Ontario:, ���Nepigon 0 6.6. 18 Ont[ario]. Dr. A.Fenyes���, ��� 248 ���, ��� 95.��� (in red ink), ��� globicollis Brh. Cotypus ��� (yellow label) (FMNH); paralectotype: CANADA: Ontario:, ���Nepigon 0 6.6. 18 Ont[ario]. Dr. A.Fenyes���, ��� 249 ���, ��� globicollis Brh. Typus ��� (yellow label) (FMNH). The purpose of the lectotype designation is to assure correct and consistent application of the name in the future. The specimen designated as lectotype was supplied with the red lectotype label. Paratypes of S. bispinosa: UNITED STATES: Arizona: Coconino Co.:,, San Francisco Mts., Hart Prairie Rd. (418), woody debris, base of dead ponderosa pine (M.W.Sanderson), 11.ix. 1982 (FMNH). Additional material. CANADA: British Columbia:, 2 mi. S Salmo (Campbell & Smetana), 9.vi. 1968;, Mt. Garibaldi, 9 mi. N Squamish, 1500 ��� (Campbell & Smetana), 30.v. 1968; 1 specimen, 26.4 mi. N Quesnel, 2350 ���, soil and moss under log (B.D.Ainscough), 3.xi. 1978; 1 specimen, Birkenhead Pr. Pk., 1600 ���, humus under maple (B.D.Ainscough), 16.iii. 1977;, McBride, river debris (D.E.Bright), 15.vii. 1981;, 46 km W McBride, sifting Populus litter at edge of spruce��cedar forest (J.M.Campbell), 26.viii. 1983; Alberta: 4 specimens, Waterton Lks. N. P., Rowe Cr., 5400 ���, under ground squirrel carcass (J.M.Campbell), 19.vi. 1980; 6 specimens, Waterton River meadow, sod (I.M.Smith), 9.vi. 1980; 1 specimen, Waterton Lks. N. P., mi. 3 Chief Mt. Hwy., 4500 ��� (J.M.Campbell), 14��17.vi. 1980; 1 specimen, George Lake, 53 �� 57 'N 114 ��06'W (R.E.Leech), 10.ix. 1966;, Cypress Hills Prov. Pk. (Redner & Starr), 9��15.vi. 1973 (all �� CNCI); Quebec: Hull Co.:, Gatineau Park, King Mountain, pine stump buttress (W.S.Suter), 24.v. 1979 (FMNH);, ���Sapini��re M��tis, Fosse L��4, 23.viii. 1993 ���;, Lac des ��tang, B��tulaie jaune �� sapin, 4��11.viii. 1999;, Lac Poissonneux, B��tulaie jaune �� sapin, pitfall trap, 4��11.viii. 1999;, ditto but 13��20.vii. 1999;, ditto but 8��15.vi. 1999 (all �� SCFK);, ditto but 15��22.vi. 1999;, Lac Mitis, sapini��re m��tis (Lindgren), 28.vii. 1993; , ditto but 21.vi. 1993;, Anticosti (Lindgren), 16.vi. 1993 (all �� SPSU); UNITED STATES: Arizona: Apache Co.: 2, 30 km S Eagar, Apache National Forest, 33 �� 51.57 'N 109 �� 11.73 'W, 2500 m, in forest litter, Picea, Pinus (V.I.Gusarov), 6.viii. 1999 (KSEM); 1 specimen, Apache N. F., Escudilla Mt., 8 mi. NE Alpine, 2730 m (J.M.Campbell), 17.vii. 1976 (CNCI);, 22 mi. SW Eagar, 2700 m, squirrel midden (P.M.Hammond), 15.vii. 1976; 2, Chuska Mts., 2600 m, squirrel midden (P.M.Hammond), 11.vii. 1976 (all �� BMNH); Cochise Co.: 3 specimens, Huachuca Mts., Bear Saddle, 8100 ��� (A.Smetana), 5.viii. 1979 (CNCI); 7 specimens, Huachuca Mts., Carr Cny., 2150 m, leaf oak litter (P.M.Hammond), 23.vii. 1976 (BMNH); 2 specimens, Huachuca Mts., Miller Ch., 7200�� 7800 ��� (A.Smetana), 4.viii. 1979; 1 specimen, Huachuca Mts., Miller Cn., 5500 ��� (A.Smetana), 1.viii. 1969; 1 specimen, Chiric Mts. Rustler Park, 8400 ��� (A.Smetana), 24.vii. 1969; , Chiricahua Mts. Nr. Ash Spg., 6300 ��� (A.Smetana), 29.vii. 1979;, Chiricahua Mts., East Turkey Creek, 6400��6800 ��� (A.Smetana), 28.vii. 1979; 1 specimen, Chiricahua Mts., near Barfoot Park, 7500 ��� (A.Smetana), 27.vii. 1979; 1 specimen, Pinaleno Mts. Wet Canyon, 6000 ��� (A.Smetana), 29.vii. 1969 (all �� CNCI); Coconino Co.: 15 specimens, Hutch Mountain, Pseudotsuga menziesi duff (M.W.Sanderson), 14.x. 1984 (KSEM); Gila Co.: 7 specimens, 21 mi. NE Payson, FR 289 between Tonto Creek (6000 ft) and Horton Creek (6500 ft), maple litter near Horton Spring (L.Herman), 5.vi. 1986 (AMNH);, Pinal Mts., Pioneer Pass, 1660 m, leaf litter (P.M.Hammond), 25.vii. 1976 (BMNH); Graham Co.:, 29 mi. SSE Safford, Rt. 266 in Penaleno Mts., Alder Spring near Stockton Pass Campground, 5700 ft, oak litter (L.Herman), 13.v. 1986 (AMNH); 4 specimens, Pinaleno Mts., Turkey Flat, 7200 ��� (A.Smetana), 27.vii. 1969; (CNCI); Navajo Co.: 17 specimens, Black Lake, Sitgreaves National Forest, 2120��2240 m, leaf litter (P.M.Hammond), 13�� 14.vii. 1976 (BMNH); Santa Cruz Co.: 3 specimens, Santa Rita Mts., Madera Canyon, Bog Spring, 5800 ft (L.Herman), 6.vi. 1981 (AMNH); 3 specimens, Santa Rita Mts., Mt. Wrightson, 8000��8500 ��� (A.Smetana), 9.viii. 1979 (CNCI); Yavapai Co.:, 14 mi. SW Clarkdale, Forest Road 104, 7200 ft (L.Herman), 2.vi. 1986 (AMNH); New Mexico: Lincoln Co.:,, 15 mi. N Ruidoso, Sacramento Mts., Kraut Cyn., 7400 ft, litter at seep (L.Herman), 3.ix. 1985;, 14 mi. N Ruidoso, Sacramento Mts., Mills Cyn., 7000 ft, litter near spring runoff (L.Herman), 3.ix. 1985; Torrance Co.:,, 6 mi. W Manzano Mts., Red Canyon Campground, 7800 ft, oak, willow, pine litter (L.Herman), 12.v. 1988 (all �� AMNH); Bernalillo Co.: 3 specimens, Sandia Mts., Cibola N. F., Tree Spr. Trail, 8500 ��� (A.Smetana), 6.vii. 1969;, ditto but Las Huertas Crk., 8.vii. 1969; Otero Co.: 6 specimens, Lincoln N. F., 2 mi. SE Cloudcroft, 8500 ��� (A.Smetana), 13.vii. 1969;, ditto but 1 mi. SE Cloudcroft, 8750 ���, 14.vii. 1969; 2 specimens, ditto but 13��18.vii. 1969 (CNCI); Utah: San Juan Co.: 2 specimens, 5 mi. W Monticello, Dalton Spring, poplar, willow, oak litter, 8200 ft (L.Herman), 11.vi. 1987;, 8 mi. SW Monticello, South Creek Road, Cold Springs, 8600 ft (L.Herman), 10.vi. 1987; Washington Co.: 2 specimens, 20 mi. NE Veyo, via Pine Valley Road, 6800 ft (L.Herman), 10.vi. 1987; Colorado: Hinsdale Co.:, 20 mi. NW Pagosa Springs, near Bridge Campground, Turkey Peak Spring, 8200 ft (L.Herman), 22.viii. 1982;,, ditto but Forest Road 639 to Trail Creek, 8000��8400 ft, 21.viii. 1982 (all �� AMNH); Las Animas Co.:, 10 km SEE Cuchara, SEE of Cordova Pass, San Isabel National Forest, 37 �� 20.28 'N 104 �� 59.66 'W, 3150 m, in forest litter, Picea, Pinus, Populus (V.I.Gusarov), 3.viii. 1999; Archuleta Co.:, 30 km W Pagosa Springs, N of Hwy. 160, San Juan National Forest, 37 �� 13.88 'N 107 �� 21.04 'W, 2250 m, in forest litter, Quercus, Pinus, Fraxinus, Pseudotsuga (V.I.Gusarov), 4.viii. 1999; Rio Grande Co.:, 9.5 km S South Fork, env. of Beaver Creek Reservoir campground, Rio Grande National Forest, 37 �� 34.86 'N 106 �� 38.96 'W, 2800 m, in forest litter (V.I.Gusarov), 4.viii. 1999; Montana: Flathead Co.:, 47 km NE Kalispell, Fish Creek, 48 �� 32.71 'N 113 �� 59.08 'W, 1100 m, in forest litter, Pinus, Tuja (V.I.Gusarov), 26.viii. 2000; Idaho: Boundary Co.:,, 19 km NE Bonners Ferry, Meadow Creek, Kaniksu National Forest, 48 �� 49.28 'N 116 ��08.47'W, 2200 m, in forest litter (V.I.Gusarov), 28.viii. 2000 (all �� KSEM); South Dakota: Custer Co.:,, North Pole Spring, 5500 ft (L.Herman), 15.vi. 1981; Wisconsin: Ashland Co.: , 9 mi. SW Marengo, near Beaver Lake (L.& N.Herman), 7.viii. 1978; 2, ditto but (L.Herman), 4.ix. 1982 (all �� AMNH); Eau Claire Co.:, 2 mi. S Eau Claire (W.Suter), 6.v. 1976 (FMNH); Minnesota: Pine Co.: 2, 16 mi. E Hinckley, St. Croix State Park (L.Herman), 1.ix. 1982 (AMNH); New Hampshire: Coos Co.: 2,, Bretton Woods (CASC); MEXICO: Nuevo Leon:,, 31.5 km SW Linares, 750 m, under leaves (R.Brooks, R.Leschen), 24.iii. 1991;, 1 specimen, ditto but under leaves in streambed, 22.iii. 1991; 2, 1 specimen, 37 km SW Linares, 900 m, flight intercept trap (R.Brooks, R.Leschen), 17��24.iii. 1991; Veracruz:,, 2 specimens, 3.2 km SW Las Vigas, Hwy. 140, 2830 m, pine forest litter (J.S.Ashe), 11.vii. 1992; Puebla:, 10 km NE Zacatepec, Hwy. 140, 2500 m, leaf litter in barranca (J.S.Ashe), 10.vii. 1992; Michoacan:, 4.8 km W Mil Cumbres, 2820 m, oak and pine forest litter (R.S.Anderson), 27.vii. 1988; Guerrero: 2, 9 specimens, 10.3 km SW Filo de Caballo, 2700 m, oak, pine and fir forest litter (R.S.Anderson), 13.vii. 1992;, ditto but 15.vii. 1992; 2, ditto but 17.vii. 1992;, 3 specimens, ditto but 18.vii. 1992;, 3 specimens, 9.3 km SW Filo de Caballo, 2400 m, alder forest litter (R.S.Anderson), 15.vii. 1992;, 2 specimens, 15 km SW Filo de Caballo, 2500 m, oak forest litter (R.S.Anderson), 16.vii. 1992;,, 8 specimens, 5.6 km SW Filo de Caballo, 2310 m, alder forest litter (R.S.Anderson), 17.vii. 1992; Oaxaca:, 62.5 km SW Valle Nacional, km 115.5, 2650 m, oak and pine forest litter (R.S.Anderson), 28.vii. 1992;,, 5 specimens, 64.5 km SW Valle Nacional, km 117.5, 2600 m, oak forest litter (R.S.Anderson), 28.vii. 1992;, 89.5 km SW Valle Nacional, km 142.5, 2430 m, pine forest litter (R.S.Anderson), 28.vii. 1992;, 2 mi. S Cerro Pelon, 8000��9000 ' (M.A.Ivie), 3.vii. 1982;, 2 mi. S Cerro Pelon, 8000��9000 ' (R.S.Miller), 2.vii. 1982;, 3.2 km S San Jose de Pacifico, Hwy. 175, 2440 m, forest litter (J.S.Ashe), 22.vii. 1992; Chiapas: 2, 4 specimens, Cerro Huitepec, ca. 5 km W San Cristobal, 2700 m, oak forest litter (R.S.Anderson), 14.ix. 1992; 3,, 23 specimens, Volc��n Tacana, lower slopes, ca. 4 km N Union Juarez, 2000 m, cloud forest litter (R.S.Anderson), 19.ix. 1992; GUATEMALA: Quetzaltenango:, 8 km SE Zunil, 2480 m, flight intercept trap (J.S.Ashe, R.Brooks), 20.viii. 1994;, 12 km SW Zunil, NE Face Cerro Zunil, 2700��2760 m, hardwood forest litter (R.S.Anderson), 28.v. 1991; Guatemala:, Guatemala City, 1 km SE La Pueblito, 1880 m, oak forest litter (R.S.Anderson), 10.vi. 1991; HONDURAS: Comayagua:, 2, 18 km E Comayagua, 2000 m, liquidambar litter (R.S.Anderson), 20.viii. 1994; Ocotepeque:, 24 km E Ocotepeque, El Guisayote, 14 �� 25 'N 89 ��04'W, 2170 m, flight intercept trap (J.S.Ashe, R.Brooks), 14��16.vi. 1994;, ditto but 16.vi. 1994;, 12.7 km E and 10.6 km S Ocotepeque, lower slopes El Pital, 14 �� 25 'N 89 ��04'W, 2050 m, oak litter (R.S.Anderson), 15.vi. 1994; El Paraiso:, 6.9 km W Yuscar��n, Cerro Monserrat, 13 �� 55 'N 86 �� 24 'W, 1760 m, forest litter (R.S.Anderson), 7.vii. 1994 (all �� KSEM). Diagnosis. Seeversiella globicollis can be distinguished from other species of Seeversiella by having dark brown body; temples 0.8��2.5 times as long as eyes; glossy pronotum with weak microsculpture; elytra longer than pronotum; wings fully developed, 4 times as long as elytra; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (Figs. 24 ��31, 33�� 36), especially the hook��shaped distal sclerites of internal sac (Figs. 31, 35); and the shape of spermatheca (Fig. 32). CP ��� copulatory piece; DS ��� distal sclerite of internal sac; LD ��� lateral diverticulum of internal sac; ML ��� medial lamellae. Seeversiella globicollis differs from closely related S. texana in having darker body colour; less distinct microsculpture of head and pronotum; elytra longer than pronotum; fully developed wings, 4 times as long as elytra; straight apex of median lobe (in lateral view) (Figs. 26��27; 43��44); longer copulatory piece of internal sac (Figs. 29, 34; 46, 49�� 50) and spermatheca without umbilicus (Fig. 32). Seeversiella globicollis differs from the other similar species of Seeversiella (3, 5��8, 10�� 12) with long elytra and weak pronotal microsculpture in having strongly sclerotized hook��shaped distal sclerites of internal sac and L��shaped spermatheca without umbilicus. Description. Length 2.4��3.3 mm. Body dark brown, sometimes with lighter elytra and mouthparts. Head surface glossy, partially with weak isodiametric microsculpture, with fine and weak punctation, distance between punctures equals 2��3 times their diameter. Temples 0.8��2.5 times as long as eyes. Antennal article 2 longer than article 3, article 4 slightly transverse, 5��10 transverse or strongly transverse (ratio 1.5 ��2.0) (Fig. 9). Pronotum slightly transverse, 1.2 times as wide as head, width 0.44��0.54 mm, length 0.40��0.47 mm, width to length ratio 1.1, surface glossy, with weak and poorly visible (at 70 x) isodiametric microsculpture; punctation as on head. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.9), 1.2 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equal to 1��2 times their diameter. Wings fully developed, 4 times as long as elytra. Abdominal terga glossy, with fine microsculpture consisting of strongly transverse meshes, with fine punctation, distance between punctures equals 2��4 times their diameter on terga 3��5 and 4��6 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (Figs. 20, 22). In males posterior angles of tergum 3 projecting as spines up to 1 time as long as tergum 3 (measured medially) (Fig. 17), tergum 7 with medial carina along midline, the carina up to 2 / 3 as long as tergum length (Fig. 17). Posterior margin of male tergum 8 without emargination (Fig. 20). Aedeagus as in Figs. 24 ��31, 33�� 36. Distal sclerites of internal sac hook��shaped (Figs. 31, 35). Proximal seta of the apex of paramere is much longer than the other three setae (Fig. 33). Spermatheca as in Fig. 32, without umbilicus. Synonyms. The types of A. globicollis agree completely with the detailed description (Ashe 1986) and with two examined paratypes of S. bispinosa from the type locality, in external characters and in male and female genitalia. Three specimens in Fenyes collection (CASC) bear a label with the manuscript name ��� Sipalia cristata Fenyes ���. Fenyes never published a description of this species. Distribution. Widespread in North America, from Canada to Honduras (Figs. 385�� 387). Natural History. Seeversiella globicollis was collected in leaf litter, often near water. In the north of its range S. globicollis occurs at the sea level, in the southern United States and further south the species is restricted to mountainous forests, mostly above 2000 m., Published as part of Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-102 in Zootaxa 142 on pages 16-22, DOI: 10.5281/zenodo.156420, {"references":["Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907, 381 - 405.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. Berlin: W. Junk, pp. 499 - 988.","Ashe, J. S. (1986) Seeversiella bispinosa, a new genus and species of athetine Aleocharinae (Coleoptera: Staphylinidae) from North America. Journal of the New York Entomological Society, 94 (4), 500 - 511."]}

1. Adota maritima (Mannerheim, 1843) (Figs. 1 ��16, 18�� 30) Homalota maritima Mannerheim, 1843: 224. Atheta (Adota) massettensis Casey, 1910: 68. Atheta (Adota) subintima Casey, 1910: 68, syn. nov. Atheta (Panalota) setositarsis Casey, 1910: 71, syn. nov. Atheta (Adota) scortea Casey, 1911: 124, syn. nov. Atheta (Adota) scolopacina Casey, 1911: 124, syn. nov. Atheta (Adota) insons Casey, 1911: 125, syn. nov. Adota massettensis: Fenyes, 1920: 176 (misspelled as masettensis; as valid species). Adota subintima: Fenyes, 1920: 176 (as valid species). Adota scortea: Fenyes, 1920: 176 (as valid species). Adota scolopacina: Fenyes, 1920: 176 (as valid species). Adota insons: Fenyes, 1920: 176 (as valid species). Atheta (Metaxya) maritima: Fenyes, 1920: 199. Panalota setositarsis: Fenyes, 1920: 243 (as valid species). Atheta (Panalota) maritima: Bernhauer & Scheerpeltz, 1926: 611 (as valid species). Atheta (Panalota) setositarsis: Bernhauer & Scheerpeltz, 1926: 611 (as possible synonym of At. maritima). Atheta (Adota) insons: Bernhauer & Scheerpeltz, 1926: 659 (as valid species). Atheta (Adota) massettensis: Bernhauer & Scheerpeltz, 1926: 659 (misspelled as masettensis; as valid species). Atheta (Adota) scolopacina: Bernhauer & Scheerpeltz, 1926: 659 (as valid species). Atheta (Adota) scortea: Bernhauer & Scheerpeltz, 1926: 659 (as valid species). Atheta (Adota) subintima: Bernhauer & Scheerpeltz, 1926: 659 (as valid species). Atheta (Panalota) maritima: Brundin, 1943: 24. Atheta (Brundinia) maritima: Moore & Legner, 1975: 366 (as valid species). Atheta (Adota) massettensis: Moore & Legner, 1975: 366 (as valid species). Atheta (Adota) subintima: Moore & Legner, 1975: 366 (as synonym of Atheta massettensis). Atheta (Adota) insons: Moore & Legner, 1975: 364 (as valid species). Atheta (Panalota) setositarsis: Moore & Legner, 1975: 366 (as valid species). Atheta (Adota) scolopacina: Moore & Legner, 1975: 373 (as valid species). Atheta (Adota) scortea: Moore & Legner, 1975: 373 (as valid species). Panalota maritima: Seevers, 1978: 265 (as valid species). Panalota setositarsis: Seevers, 1978: 265 (as valid species). Xenota massettensis: Seevers, 1978: 270 (as valid species). Xenota subintima: Seevers, 1978: 271 (as valid species). Xenota scortea: Seevers, 1978: 271 (as valid species). Xenota scolopacina: Seevers, 1978: 271 (as valid species). Xenota insons: Seevers, 1978: 270 (as valid species). Adota maritima: Lohse & Smetana, 1985: 282 (as valid species). Adota massettensis: Lohse & Smetana, 1985: 282 (misspelled as massetensis; as synonym of Adota maritima). Adota setositarsis: Lohse & Smetana, 1985: 282 (as valid species). Type material. Lectotype of Homalota maritima (designated by Lohse and Smetana (1985)),, UNITED STATES: Alaska: Sitka (MZHF). Lectotype of Atheta massettensis (here designated):, "Q.[ueen] C.[harlotte] I.[slands] [Massett (J.H.Keen)]", " Adota massettensis Csy. ", " TYPE USNM 39364 " (red label), " CASEY bequest 1925 " (NMNH). Paralectotypes: 4, "Q.[ueen] C.[harlotte] I.[slands] [Massett (J.H.Keen)]", " massettensis PARATYPE USNM 39364 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta subintima (here designated):, "Metlakatla, B.[ritish] Col.[umbia] (Keen)", " subintima Csy. ", " TYPE USNM 39365 " (red label), " CASEY bequest 1925 " (NMNH). Paralectotypes: 14 specimens, "Metlakatla, B.[ritish] Col.[umbia] (Keen)", " subintima PARATYPE USNM 39365 " (red label), " CASEY bequest 1925 " (NMNH). Holotype of Atheta setositarsis:, "Cal.[ifornia, San Francisco]", " Panalota setositarsis Csy. ", " TYPE USNM 39358 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta scortea (here designated):, "Cal.[ifornia, Berkeley, Alameda Co.]", " scortea Csy. ", " TYPE USNM 39367 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta scolopacina (here designated):, "Cal.[ifornia, Berkeley, Alameda Co.]", " scolopacina Csy. ", " TYPE USNM 39369 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta insons (here designated):, "Cal.[ifornia, San Diego]", " insons Csy. ", " TYPE USNM 39370 " (red label), " CASEY bequest 1925 " (NMNH). Additional material. UNITED STATES: Alaska:, Kodiak (MZHF); Kenai Peninsula Co.: 8 specimens, Homer, S shore of Homer spit, 59 �� 36.33 'N 151 �� 25.71 'W, sandy seashore, in seaweed (V.I.Gusarov), 22.vii. 1998 (AMNH, SPSU); Haines Co.:, Haines, sifting algae on beach (Campbell & Smetana), 3.vii. 1968 (CNCI); California: Marin Co.:, Stinson State Beach, under seaweed on sand beach (K.J.Ahn & J.S.Ashe), 14.v. 1991 (KSEM); Los Angeles Co.: 5, Santa Catalina Isl., Descanso Beach, under seaweed on rocks with numerous Diptera larvae (K.J.Ahn), 31.v. 1991 (KSEM); 2 specimens, Sonoma Co.: 34.5 km SWW Santa Rosa, Bodega Head, University of California Marine Laboratory, 38 �� 19.013 'N 123 ��04.254'W �� 10m, seashore, in seaweed (V.I.Gusarov), 4.vi. 2002 (KSEM); Santa Cruz Co.:, 10.5 km NW Davenport, nr. Hwy. 1, nr. Greyhound Rock, 37 ��04.821'N 122 �� 16.062 'W �� 6m, seashore, in seaweed (V.I.Gusarov), 19.vi. 2002 (SPSU); Mendocino Co.: 2, Needle Rock, on beach (D.Giuliani), 6.x. 1974 (UCR); San Francisco Co.:, 2, without locality label (MCZ); CANADA: British Columbia: 2, 2, Miracle Beach, 17 mi. S Campbell River (J.M. & B.A.Campbell), 22.vii. 1975 (CNCI);,, Massett (BMNH). Diagnosis. Adota maritima is very similar to Ad. gnypetoides, but differs in having larger body size, longer antennal segments (Figs. 18; 17), wider apex of median lobe (Figs. 23��24; 35��36), posterior margin of male tergum 8 emarginate medially (Fig. 19) and larger spermatheca (Figs. 29��30; 40��41). Adota maritima differs from Ad. colpophila by having matte body, posterior margin of male tergum 8 with four blunt projections (Fig. 19), median lobe with blunt apex (Figs. 23�� 24; 46��47) and the distinct shape of spermatheca (Figs. 29��30). Description. Length 2.5��3.2 mm. Body black with dark brown legs and light brown tarsi. Head surface matte, with strong and dense isodiametric microsculpture, with fine punctation, distance between punctures equals 1��2 times their diameter. Frons with sparser punctation, in both sexes with longitudinal impression. Temple length to eye length ratio 1.0�� 1.2. Antennal article 2 is 1.1 times as long as article 3, articles 4��5 slightly elongate (length to width ratio 1.2��1.3), articles 6��7 as long as wide, articles 8��10 slightly transverse (length to width ratio 0.8) (Fig. 18). Pronotum subquadrate or slightly transverse, 1.1 times as wide as head, width 0.47�� 0.61 mm, length 0.41��0.51 mm, width to length ratio 1.2; matte, with strong and dense isodiametric microsculpture; punctation finer than on head, poorly visible on matte background. Elytra much wider (0.63��0.89 mm) and longer (0.57��0.79 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.5), 1.1 times wider than long, surface matte, with strong and dense isodiametric microsculpture; punctation poorly visible. Abdominal terga matte, with strong and dense isodiametric microsculpture; with fine punctation, distance between punctures equals 2��3 times their diameter on terga 3��5 and 3�� 5 times on terga 6��7. Apical margin of tergum 7 with white palisade fringe. Posterior margin of male tergum 8 with four blunt projections, emarginate medially (Fig. 19). Aedeagus as in Figs. 23��28, median lobe with wide apex (Figs. 23��24). Spermatheca as in Figs. 29��30. Discussion. The types of Homalota maritima, Atheta massettensis, At. subintima, At. setositarsis, At. scortea, At. scolopacina and At. insons fall within the range of variability of Adota maritima and have similar shape of genitalia and male abdominal tergum 8. The holotype of Atheta insons is smaller (pronotal width 0.47 mm) than most specimens of Adota maritima. However, it differs from Adota gnypetoides by the wide apex of median lobe and the shape of posterior margin of the male tergum 8, and I interpret it as a small specimen of Ad. maritima. Examined specimens of Ad. maritima from southern California are smaller than the specimens from British Columbia and Alaska and approach in size examined specimens of Ad. gnypetoides. Additional collecting is needed to study the variability of Ad. maritima. Distribution. Known from the Pacific coast of North America, from Alaska to California (Fig. 52). Natural History. Adota maritima is a littoral species, common in decomposing seaweed on a beach., Published as part of Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-35 in Zootaxa 185 on pages 9-14, DOI: 10.5281/zenodo.156388, {"references":["Mannerheim, C. G. (1843) Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16 (1 - 2), 175 - 314.","Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. Lancaster: The New Era Printing Company, pp. 1 - 183.","Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. Lancaster: The New Era Printing Company, pp. 1 - 245.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. Bruxelles: L. Desmet-Verteneuil, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. Berlin: W. Junk, pp. 499 - 988.","Brundin, L. (1943) Zur Kenntnis einiger in die Atheta - Untergattung Metaxya M. & R. gestellten Arten (Col. Staphylinidae). Lunds Universitets Arsskrift, N. F., Avd. 2, 39 (4), 3 - 37, Taf. 1 - 7.","Moore, I. & Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, 1 - 275.","Lohse, G. A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Maklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39 (3), 281 - 300."]}

Key to Nearctic species of Stethusa 1 Antennal article 11 with two subbasal impressions densely covered with microsetae (Figs. 15­16). Internal sac of aedeagus with two strong distal spines (Figs. 34 ­35, 37­ 38). Female with accessory sclerite (Fig. 40). Distal end of spermatheca bent towards spermathecal gland (Figs. 41­42). Body length 2.0­3.0 mm, pronotal length 0.41­0.53 mm. Widely distributed in the eastern United States (Fig. 110). Also known from Bolivia, Paraguay, the Galápagos Islands and the Antilles.. 1. S. dichroa (Gravenhorst) – Antennal article 11 without subbasal impressions, microsetae evenly distributed. Internal sac of aedeagus without strong distal spines (Figs. 53 ­54, 67­ 68). Female without accessory sclerite. Distal end of spermatheca bent away from spermathecal gland (Figs. 57, 70)....................................................................................................... 2 2 Body larger, length 3.2­3.5 mm, pronotal length 0.54­0.61 mm. Apex of median lobe in lateral view straight (Figs. 50­51). Spermatheca longer (Fig. 57). Known from Louisiana, Mississippi and southern Indiana (Fig. 111)........ 2. S. klimschi (Bernhauer) – Body smaller, length 2.1­2.5 mm, pronotal length 0.36­0.44 mm. Apex of median lobe in lateral view bent paramerally (Figs. 65­66). Spermatheca shorter (Fig. 70). Widely distributed in the eastern United States (Fig. 111).......... 3. S. spuriella (Casey)

Stethusa lurida (Erichson, 1839), new combination (Figs. 10, 37��38, 46, 53 in Klimaszewski & Peck, 1998) Homalota lurida Erichson, 1839: 108. Atheta (Metaxya) lurida: Fenyes, 1920: 199 (as valid species). Atheta (s. str.) lurida: Bernhauer & Scheerpeltz, 1926: 645 (as valid species). Atheta lurida: Blackwelder, 1944: 160 (as valid species). Atheta lurida: Pace, 1985: 454 (as valid species). Atheta lurida: Klimaszewski & Peck, 1998: 238 (as valid species). Material. No specimens of S. lurida have been examined. Diagnosis. See Klimaszewski & Peck, 1998. Discussion. Stethusa lurida is similar to the Nearctic species of the genus in the shape of posterior margin of the male tergum 8 (Fig. 53: Klimaszewski & Peck 1998); the pronotal pubescence of type I (Fig. 10: Klimaszewski & Peck 1998); the long pronotal macrosetae (Fig. 10: Klimaszewski & Peck 1998); the shape of the aedeagus (Figs. 37��38: Klimaszewski & Peck 1998), particularly the shape of the copulatory piece of the internal sac (indicated by denser punctation in both lateral and abparameral views of the median lobe (Figs. 37��38: Klimaszewski & Peck 1998)); and the shape of the spermatheca (Fig. 46: Klimaszewski & Peck 1998)). Since I have not examined any specimens of S. lurida, I rely on the published diagnosis and illustrations provided by Klimaszewski and Peck (1998) who noted that they had examined a syntype of Homalota lurida from Brasil (ZMNB). Distribution. Stethusa lurida is known from Brasil, Argentina and the Gal��pagos Islands., Published as part of Gusarov, Vladimir I., 2003, A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-43 in Zootaxa 239 on pages 29-30, DOI: 10.5281/zenodo.156493, {"references":["Erichson, W. F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. 1 - 400.","Klimaszewski, J. & Peck, S. B. (1998) A review of Aleocharine Rove Beetles from the Galapagos Islands, Ecuador (Coleoptera: Staphylinidae: Aleocharinae). Revue suisse de Zoologie, 105 (2), 221 - 260.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185 (1), i-xii, 1 - 188.","Pace, R. (1985) Aleocharinae delle isole Galapagos (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France (N. S.), 21 (4), 449 - 456."]}

Idahotyphlus alleni Gusarov, sp. n. (Figs. 1���22) Type material. Holotype: ��, UNITED STATES: Idaho: Boise Co.: Beaver Creek Summit, funneled from Ponderosa pine litter (A.D.Allen), 3.vii. 1978 (KSEM). Paratypes: UNITED STATES: Idaho: Boise Co.: 3 ����, 13 ����, same data as the holotype (AACB, KSEM, SPSU, WFBM); 3 ����, ca. 30 mi. NE Boise, Ten Mile Camp Area, funneled from Ponderosa pine litter (A.D.Allen), 29.v. 1978 (AACB, SPSU); ��, 2 ����, ditto but 20.v. 1982 (FMNH, SPSU); Valley Co.: 3 ����, Warm Lake, pine litter (A.D.Allen), 4.vi. 1982 (FMNH). Diagnosis. Idahotyphlus alleni is the only known species of the genus Idahotyphlus. If any additional species of Idahotyphlus are ever discovered, I. alleni could probably be distinguished by the shape of the aedeagus (Figs. 15���20) and the shape of the female accessory sclerites (Fig. 22). Description. Length 1.4���1.7 mm. Head with short setae arranged as in Figs. 1���2. Second antennal article slightly longer than wide, twice as long and 1.4 times as wide as article 3, articles 3���9 transverse, 1.2���1.5 times as wide as long, article 10 is 1.3 times as wide as long, last article elongate, 1.3 times as long as wide. Pronotum with short setae arranged as in Fig. 1, narrowed posteriad, 1.1 times as wide as long, as wide as head and elytra. Elytra with short setae, with lateral sides diverging posteriad (Fig. 1), 1.1 times as wide as long (length measured from humeral level). Male mesotrochanter without modification (Fig. 13). In males, setae on ventral side of protarsus thicker than in females (cf. Figs. 11 and 12), but in both sexes the setae lack apical adhesive disks. Male abdominal sternum 8 with broad and shallow apical emargination, without zone devoid of setae (Fig. 21). Aedeagus long and narrow, without denticle on parameral surface (Figs. 15���20). Internal sac with long needle��shaped sclerite which is as long as median lobe; when retracted the sclerite exposed apically. Female accessory sclerites as in Fig. 22, poorly sclerotized. Distribution. Known from three localities in Idaho (Fig. 23). The males from both localities in Boise County are identical in the shape of the aedeagus and its internal sclerite. No males are known from the locality in Valley County but the females are identical to those from the type locality in all external characters, including the shape of the terminal abdominal segments, and in the shape of the female accessory sclerites. Natural History. Idahotyphlus alleni was found in funnel extract of forest litter taken at the base of a big Ponderosa pine tree (Pinus ponderosa) (Allen, personal communication)., Published as part of Gusarov, Vladimir I., 2003, Idahotyphlus alleni Gusarov, gen. n., sp. n., a new leptotyphline staphylinid beetle from Idaho (Coleoptera: Staphylinidae: Leptotyphlinae), pp. 1-8 in Zootaxa 345 on pages 5-7, DOI: 10.5281/zenodo.156647

2. Psammostiba kenaii Gusarov, sp. n. (Figs. 78��87) Type material. Holotype:, UNITED STATES: Alaska: Haines Co.: Haines (Campbell & Smetana), 3.vii. 1968 (CNCI). Paratypes: UNITED STATES: Alaska: Haines Co.:, 4, Haines, sifting algae on beach (Campbell & Smetana), 3.vii. 1968 (CNCI, SPSU); Kenai Peninsula Co.:, Homer, S shore of Homer spit, 59 �� 36.33 'N 151 �� 25.71 'W, sandy seashore, in seaweed (V.I.Gusarov), 22.vii. 1998 (SPSU). Additional material. UNITED STATES: California: Mendocino Co.:, Needle Rock, on beach (D.Giuliani), 6.x. 1974 (UCR); CANADA: British Columbia:, Victoria (CASC). Diagnosis. Psammostiba kenaii can be distinguished from P. comparabilis by smaller body, smaller aedeagus with narrower apex (Figs. 82��83; 73��74) and smaller spermatheca (Figs. 87; 77). Psammostiba kenaii can be distinguished from the Palaearctic species of Psammostiba by the fine crenulation of the posterior margin of male tergum 8 (Fig. 78), and by the distinct shape of aedeagus (Figs. 82��86) and spermatheca (Fig. 87). Description. Length 3.0�� 3.1 mm. Body black, elytra dark brown to black, legs brown, tarsi yellowish brown. Head surface matte, with strong and dense isodiametric microsculpture, with fine and poorly visible punctation, distance between punctures equals their diameter; in both sexes with weak medial impression. Temple length to eye length ratio 0.9 ��1.0. Antennal article 3 longer than article 2, articles 4��7 elongate, 8��10 elongate or subquadrate, article 11 shorter than articles 9 and 10 combined. Pronotum slightly transverse, 1.2 times as wide as head, width 0.59��0.64 mm, length 0.47��0.53 mm, width to length ratio 1.2; surface matte, with strong and dense isodiametric microsculpture; punctation finer than on head, poorly visible on matte background, distance between punctures equal to ���� 1 times their diameter. Elytra much wider (0.76��0.86 mm) and longer (0.70��0.76 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.5), 1.1 times as wide as long, surface matte, with strong and dense isodiametric microsculpture; punctation as on pronotum. Abdominal terga matte, with strong and dense isodiametric microsculpture, with fine punctation, punctation on terga 6��7 almost as dense as on terga 3��5, distance between punctures equals 1��3 times their diameter. Apical margin of tergum 7 with white palisade fringe. Posterior margin of male tergum 8 with fine crenulation (Fig. 78). Posterior margin of male sternum 8 convex (Fig. 79). Aedeagus as in Figs. 82��86. Spermatheca as in Fig. 87. Distribution. Known from the Pacific coast of Alaska (Fig. 88). Records from British Columbia and California are based on female specimens and need to be reconfirmed when male specimens become available. Natural History. Psammostiba kenaii is a littoral species, inhabiting decomposing seaweed on a beach., Published as part of Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-35 in Zootaxa 185 on pages 28-32, DOI: 10.5281/zenodo.156388

Psammostiba Yosii & Sawada, 1976, stat. nov. (Figs. 53��88) Atheta (Panalota): Brundin, 1943: 19, ex parte. Atheta (Psammostiba Yosii & Sawada, 1976): 82 (type species: Homalota hilleri Weise, 1877, by original designation). Diagnosis. Psammostiba can be distinguished from the other athetine genera by the combination of the following characters: body parallel��sided; anterior margin of labrum concave; antennal articles 5��10 slightly elongate or subquadrate (Fig. 64); ligula long and split in apical half (Fig. 58); labial palpus with setae and present (Fig. 58); pronotum slightly transverse, 1.1 times as wide as long, with microsetae directed anteriorly along the midline; in lateral portions of the disc microsetae directed laterally (Type I, Benick & Lohse 1974) (Fig. 62); pronotal macrosetae short; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia inconspicuous and thin, as long as tibial width; tarsal formula 4��5 �� 5; metatarsal segment 1 as long as segment 2 (Fig. 65); no empodial setae (best observed in lateral view; Fig. 65); tarsal claws of different length (best observed in dorsal view; Fig. 66), external claw longer than internal; copulatory piece without pointed apex (Figs. 67��68); proximal portion of spermatheca with one coil (Figs. 77, 87). Psammostiba can be distinguished from Atheta by strong isodiametric microsculpture of the entire body; by the concave anterior margin of the labrum; by basal impression on the tergum 6; by lacking empodial seta and by tarsal claws of different length. Psammostiba differs from Adota in having antennal article 3 longer than article 2 (Fig. 64); larger body; tarsal claws of different length; and in lacking the empodial seta. Description. Length 3.0��5.0 mm, pronotal width 0.59��0.89 mm. Body black with brown legs and light brown tarsi. Head 1.1 times as wide as long; eyes large, temple length to eye length ratio 0.9��1.1; infraorbital carina incomplete, reaching middle of eye or very short. Antennal article 3 slightly longer than article 2, articles 4��10 slightly elongate or subquadrate, terminal article without coeloconic sensilla, shorter or as long as articles 9 and 10 combined (Fig. 64). Labrum (Fig. 58) transverse, with concave anterior margin. Adoral surface of labrum (epipharynx) as in Fig. 54. Mandibles (Figs. 55��57) broad, right mandible with a small medial tooth; dorsal molar area with velvety patch consisting of very small denticles (invisible at 400 x). Maxilla (Fig. 60) with galea projecting slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; internal margin of galea with long subapical setae; apical 1 / 3 of lacinia with row of closely spaced spines, middle portion produced medially and covered with numerous setae (Fig. 60). Labium as in Figs. 58 ��59, 61; ligula long and split in apical half; medial area of prementum with 2 pores and with 18��30 pseudopores, lateral areas each with two asetose pores, single setose pore and 18��33 pseudopores (Fig. 58). Hypopharyngeal lobes as in Fig. 59. Labial palpus with setae and present (Fig. 58). Mentum (Fig. 61) with concave anterior margin. Pronotum slightly transverse, 1.2 times as wide as long, with microsetae directed anteriorly in midline; in lateral portions of disc microsetae directed laterally (Type I, Benick & Lohse 1974) (Fig. 62); macrosetae short; hypomera fully visible in lateral view. Meso�� and metasternum as in Fig. 63, mesosternal process narrow, extending about 4 / 7 length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 4: 1: 2; mesocoxal cavities margined posteriorly; mesocoxae narrowly separated. Medial macroseta of mesotibia inconspicuous, shorter than tibial width. Tarsal segmentation 4��5 �� 5, metatarsal segment 1 as long as segment 2 (Fig. 65). No empodial setae (best observed in lateral view; Fig. 65); tarsal claws of different length (best observed in dorsal view; Fig. 66), external claw longer than internal. Posterior margin of elytra straight. Wings fully developed. Abdominal terga 3��6 with moderate basal impression. Tergum 7 as long as tergum 6. Punctation on terga 6��7 almost as dense as on terga 3��5. Tergum 7 with wide white palisade fringe. Internal sac of aedeagus with a medial pair of partially sclerotized structures (Figs. 67�� 68; SS) which may be homologous to medial lamellae present in many genera of Athetini. Copulatory piece without pointed apex (Figs. 67��68); proximal portion of spermatheca with a single coil (Figs. 77, 87). Type species. Homalota hilleri Weise, 1877, by original designation. Discussion. Although originally proposed as a subgenus of Atheta, Psammostiba differs from Atheta in many characters (see Diagnosis) and it is considered here as a distinct genus of the tribe Athetini. Yosii & Sawada (1976) state that in P. hilleri and P. jessoensis the "claws of all legs are variable in length and strength". In all twenty specimens of P. jessoensis examined by me the external claws were significantly longer than the internal ones. Psammostiba seems to be restricted to the coasts of the Northern Pacific. Psammostiba comparabilis (M��klin in Mannerheim, 1853) and P. kenaii Gusarov, sp. n. are the only known Nearctic species of this genus. Psammostiba includes three additional Palaearctic species distributed in the Far East: P. hilleri (Weise, 1877), P. jessoensis (Brundin, 1943) and P. kamtschatica (Brundin, 1943)., Published as part of Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-35 in Zootaxa 185 on pages 20-24, DOI: 10.5281/zenodo.156388, {"references":["Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of Genera and Subgenera with description of representative Species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140.","Brundin, L. (1943) Zur Kenntnis einiger in die Atheta - Untergattung Metaxya M. & R. gestellten Arten (Col. Staphylinidae). Lunds Universitets Arsskrift, N. F., Avd. 2, 39 (4), 3 - 37, Taf. 1 - 7.","Weise, J. (1877) Japanische Staphylinidae und Pselaphidae. In: Beitrage zur Kaferfauna von Japan, meist auf R. Hiller's Sammlungen basirt. Deutsche Entomologische Zeitschrift, 21, 88 - 100.","Benick, G. & Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Krefeld: Goecke & Evers Verlag, pp. 72 - 220.","Mannerheim, C. G. (1853) Dritter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 26 (3), 95 - 273."]}

Gusarov, Vladimir I. (2003): A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 185: 1-35, DOI: 10.5281/zenodo.156388

10. Seeversiella mexicana Gusarov, sp. n. (Figs. 135��149) Type material. Holotype,, MEXICO: Oaxaca: 4.6 km S Suchixtepec, 2150 m, leaf litter in riparian alder forest (R.S.Anderson), 23.vii. 1992 (KSEM). Paratypes: MEXICO: Oaxaca: 2, 5, same data as the holotype;, 2, ditto but 5.1 km S Suchixtepec, oak, pine and alder forest litter, 25.vii. 1992;, 4, ditto but 24.vii. 1992; Guerrero: 3, 4, 71 km NE Atoyac de Alvarez, 1700 m, forest litter (J.S.Ashe), 25.vii. 1992; 3,, 5.6 km SW Filo de Caballo, 2310 m, alder forest litter (R.S.Anderson), 13.vii. 1992 (all �� KSEM). Diagnosis. Seeversiella mexicana can be distinguished from other species of Seeversiella by having brown body; temples 1.2 ��2.0 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer or a little shorter than pronotum (elytral length to pronotal length ratio 1.2 �� 0.9); wings fully developed or reduced; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (Figs. 139��148) and spermatheca (Fig. 149). Seeversiella mexicana can be distinguished from S. globicollis and the other similar species (3, 5��8, 11�� 12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook��shaped and have strong spicules (Fig. 146); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (Figs. 141��142), in parameral view narrow and with weak basal constriction (Figs. 139��140); spermatheca Sshaped with umbilicus facing proximally (Fig. 149). Description. Length 2.2��2.8 mm, pronotal width 0.41��0.53 mm. Body brown to dark brown with lighter elytra, legs brown to yellowish brown. In all character states S. mexicana is very similar to S. globicollis, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis. In large males posterior angles of male tergum 3 projecting as spines up to twice as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2 / 3 as long as tergum length. Aedeagus as in Figs. 139��148. Spermatheca as in Fig. 149. DS ��� distal sclerite of internal sac. Distribution. Known from Mexico (Fig. 387). Natural History. Seeversiella mexicana was collected in forest litter at altitude of 1700��2310 m., Published as part of Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-102 in Zootaxa 142 on pages 40-42, DOI: 10.5281/zenodo.156420

16. Seeversiella brevipennis Gusarov, sp. n. (Figs. 222��234) Type material. Holotype,, PANAMA: Chiriqu��: 5.9 km N Cerro Punta, Par. Nac. Volc��n Bar��, 8 �� 52 'N [the latitude on the label (8 �� 22 '0"N) is incorrect] 82 �� 34 '0"W, 2150 m, alder forest litter (R.Anderson), 14.vi. 1995 (KSEM). Paratypes: PANAMA: Chiriqu��:,, same data as the holotype (KSEM). Diagnosis. Seeversiella brevipennis can be distinguished from other species of Seeversiella by having brown or reddish brown body with yellowish brown elytra, abdominal segments 3��4, legs and mouthparts; large eyes (temple length to eye length ratio 1.6 ��2.0); glossy pronotum with weak isodiametric microsculpture and asperate punctation; elytra as long as pronotum; wings partially reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (Figs. 226��233) and spermatheca (Fig. 234). Seeversiella brevipennis differs from S. scabricollis in having glossy pronotum with asperate punctation; elytra as long as pronotum; apex of median lobe in parameral view constricted basally (Figs. 226��227; 194��196). Seeversiella brevipennis differs from S. furcativentris in having glossy pronotum with asperate punctation; elytra as long as pronotum; larger median lobe with longer apex (Figs. 226��229; 212��215). Seeversiella brevipennis differs from S. brunnea in having glossy pronotum with asperate punctation; elytra as long as pronotum; straight apex of median lobe (in lateral view) (Figs. 228��229; 241��242). Description. Length 2.6 ��3.0 mm. Body brown or reddish brown, elytra, abdominal segments 3 and 4, legs and mouthparts yellowish brown. Head surface glossy, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2��3 times their diameter. Temple length to eye length ratio 1.6 ��2.0. Antennal article 2 as long as 3, articles 4��10 transverse (ratio 1.3�� 1.6). Pronotum slightly transverse, 1.2 times as wide as head, width 0.53��0.56 mm, length 0.47��0.5 mm, width to length ratio 1.1, surface glossy, with fine and weak isodiametric microsculpture; punctation fine and asperate, distance between punctures equal to 1��1.5 times their diameter. Elytra wider and as long (measured from humeral angle) as pronotum (pronotal length to elytral length ratio 1.0), 1.4 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to 1��2 times their diameter. Wings partially reduced. Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 3��6 times their diameter on terga 3��5 and 4��7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (Figs. 222, 224). In males posterior angles of tergum 3 projecting as spines up to 2 times as long as tergum 3 (measured medially); tergum 7 in front of posterior margin with medial tubercle; posterior margin of male tergum 8 with emargination delimited by short projections (Fig. 222). Aedeagus as in Figs. 226��233. Apex of paramere broad, proximal seta approximately as long as the other three setae (Fig. 233). Spermatheca as in Fig. 234. Distribution. Known from Cerro Punta, Panama (Fig. 388). In this locality S. brevipennis is sympatric S. scabricollis. Natural History. Seeversiella brevipennis was collected in forest litter at altitude of 2150 m., Published as part of Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-102 in Zootaxa 142 on pages 58-62, DOI: 10.5281/zenodo.156420

9. Seeversiella grandis Gusarov, sp. n. (Figs. 126��134) Type material. Holotype,, GUATEMALA: Quetzaltenango: 8 km SE Zunil, 2620 m, flight intercept trap (J.S.Ashe & R.Brooks), 19��21.vi. 1993 (KSEM). Diagnosis. Seeversiella grandis can be distinguished from other species of Seeversiella by having large body (3.7 mm), brownish black body with brown elytra and legs; large eyes (temple length to eye length ratio 0.9); matte pronotum with strong isodiametric microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (Figs. 129��134). Description. Length 3.7 mm. Body brownish black, elytra and legs brown. Head surface matte, with fine isodiametric microsculpture, fine punctation, distance between punctures equals 1��2 times their diameter, anteromedial portion of the disc without punctation. Temple length to eye length ratio 0.9. Antennal article 2 as long as 3, article 4 elongate (ratio 1.2), 5 quadrate, 6��10 transverse (ratio 1.2��1.3). Pronotum slightly transverse, 1.3 times as wide as head, width 0.74 mm, length 0.66 mm, width to length ratio 1.1, surface matte, with fine isodiametric microsculpture and fine punctation, distance between punctures equals ���� 1 times their diameter. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.3 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to ���� 1 time their diameter. Wings fully developed. Abdominal terga glossy, terga 3��6 with fine microsculpture consisting of transverse waves and strongly transverse meshes, with fine punctation, distance between punctures equals 3��6 times their diameter. Tergum 7 with microsculpture consisting of transverse meshes and with fine and sparse punctation, distance between punctures equal to 4��7 times their diameter. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (Fig. 126). In the only known male specimen pronotum with wide medial impression, posterior margin of male tergum 3 with two short and obtuse projections (as in Fig. 19), tergum 7 with medial tubercle in front of posterior margin. Posterior margin of male tergum 8 without emargination (Fig. 126). Aedeagus as in Figs. 129��134. Proximal seta of the apex of paramere is much longer than the other three setae (Fig. 134). Female unknown. Distribution. Known from Guatemala (Fig. 387). Natural History. The only known specimen of S. grandis was collected with a flight intercept trap at altitude of 2620 m., Published as part of Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-102 in Zootaxa 142 on pages 38-40, DOI: 10.5281/zenodo.156420

Atheta texana Casey, 1910 (Figs. 84��87) Atheta (Stethusa) texana Casey, 1910: 6. Atheta (Stethusa) mendosa Casey, 1910: 8, syn. nov. Atheta (Hypatheta) texana: Fenyes, 1920: 207 (as valid species). Atheta (Hypatheta) mendosa: Fenyes, 1920: 208 (as valid species). Atheta (Hypatheta) texana: Bernhauer & Scheerpeltz, 1926: 654 (as valid species). Atheta (Hypatheta) mendosa: Bernhauer & Scheerpeltz, 1926: 654 (as valid species). Atheta (Stethusa) texana: Moore & Legner, 1975: 375 (as valid species). Atheta (Stethusa) mendosa: Moore & Legner, 1975: 366 (as valid species). Xenota texana: Seevers, 1978: 269 (as valid species). Xenota mendosa: Seevers, 1978: 268 (as valid species). Type material. Lectotype of Atheta texana (here designated): ��, "Tex. [Texas, Austin]", " texana Csy. ", " TYPE USNM 39444 " (red label), " CASEY bequest 1925 " (NMNH). Holotype of Atheta mendosa: ��, "Va. [Virginia, Fort Monroe]", " mendosa Csy. ", " TYPE USNM 39453 " (red label), " CASEY bequest 1925 " (NMNH). Diagnosis. Atheta texana can be recognized by the pronotal setation of type I, antennal articles 4��5 quadrate, articles 6��10 transverse, mesothoracic process narrow, tergum 7 with microsculpture consisting of transverse waves and very transverse cells; and the distinct shape of the spermatheca (Figs. 86��87). Body length 2.7 ��3.0 mm, pronotal width 0.69��0.70 mm. Discussion. The types of At. texana and At. mendosa are very similar in external characters and the shape of the spermatheca. Based on this similarity, the second name is placed in synonymy with the first. The systematic position of this species is not clear until the males are discovered. Distribution. Atheta mendosa is known from Virginia and Texas. Natural History. No information is available., Published as part of Gusarov, Vladimir I., 2003, A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-43 in Zootaxa 239 on pages 33-34, DOI: 10.5281/zenodo.156493, {"references":["Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Moore, I. & Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp."]}

3. Adota colpophila Gusarov, sp. n. (Figs. 42��51) Type material. Holotype:, MEXICO: Sonora: Punta Cirio, 29 �� 53 'N 112 �� 40 'W, wrack on sandy beach (V.Roth & W.Brown), 20.iii. 1974 (UCR). Paratypes: MEXICO: Sonora: 69 specimens, sama data as the holotype (UCR, KSEM, SPSU); 57 specimens, Punta Chueca, 29 ��00'N 112 ��05'W, light trap on beach (V.Roth), 18.i. 1974 (UCR, KSEM, SPSU). Diagnosis. Adota colpophila differs from Ad. maritima and Ad. gnypetoides in having glossy body, despite isodiametric microsculpture; posterior margin of male tergum 8 straight (Fig. 42); median lobe with pointed apex (Figs. 46��47; 23 ��24, 35�� 36) and the distinct shape of spermatheca (Fig. 51). Description. Length 2.4��2.6 mm. Body black, elytra dark brown to reddish brown, antennae black to brown, legs brown to yellowish brown. Head surface glossy, with weak and dense isodiametric microsculpture, with fine punctation, distance between punctures equals their diameter. Frons with sparser punctation, in both sexes with weak impression. Temples as long as eyes. Antennal article 2 longer than article 3, articles 4��6 slightly elongate, article 7 as long as wide, articles 8��10 slightly transverse. Pronotum slightly transverse, 1.1 times as wide as head, width 0.44��0.50 mm, length 0.37��0.43 mm, width to length ratio 1.1��1.3; glossy, with weak and dense isodiametric microsculpture; punctation poorly visible and finer than on head, distance between punctures equal to ���� 1 times their diameter. Elytra much wider (0.61��0.66 mm) and longer (0.56��0.61 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.5), 1.1 times as wide as long, surface glossy, with weak and dense isodiametric microsculpture; punctation as on pronotum. Abdominal terga glossy, with weak isodiametric microsculpture; with fine punctation, distance between punctures equals 1��3 times their diameter on terga 3��5 and 2��5 times on terga 6��7. Apical margin of tergum 7 with white palisade fringe. Posterior margin of male tergum 8 with straight posterior margin (Fig. 42). Aedeagus as in Figs. 46��50, median lobe with pointed apex (Figs. 46��47). Spermatheca as in Fig. 51. Distribution. Known from the Gulf of California (Fig. 52). Natural History. Adota colpophila is a littoral species, inhabiting decomposing seaweed on a beach., Published as part of Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-35 in Zootaxa 185 on pages 16-18, DOI: 10.5281/zenodo.156388

Adota magnipennis (Bernhauer, 1943), comb. nov. (Figs. 36, F��R in Yosii & Sawada 1976) Atheta (Anopleta) magnipennis Bernhauer, 1943: 184. Atheta (Anopleta) magnipennis: G.Benick, 1970: 94. Atheta (Halostiba) magnipennis: Yosii & Sawada, 1976: 88. Atheta (Halostiba) magnipennis: Sawada, 1977: 173. Diagnosis. Adota magnipennis can be recognized by the distinct shape of male tergum 8 (Fig. 36, K: Yosii & Sawada 1976), the presence of few bifurcate microsetae at the apex of male sternum 8 (Fig. 36, M) and by the distinct shape of the aedeagus and spermatheca (Figs. 36, N��R). Adota magnipennis can be distinguished from the Nearctic species of Adota by the concave and slightly crenulate posterior margin of the male tergum 8 and different shape of the aedeagus and spermatheca. Distribution. Adota magnipennis is known from Japan (Honshu and Kyushu)., Published as part of Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-35 in Zootaxa 185 on page 20, DOI: 10.5281/zenodo.156388, {"references":["Bernhauer, M. (1943) Neuheiten der palaearktischen Staphylinidenfauna. Mitteilungen der Munchner Entomologischen Gesellschaft, 33, 169 - 188.","Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of Genera and Subgenera with description of representative Species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140.","Benick, G. (1970) Revision der Untergattung Anopleta Muls. et Rey. Entomologische Blatter, 66, 83 - 110.","Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). III: Japanese Species described by the previous Authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222."]}

Key to Nearctic species of Psammostiba 1 Body larger, length 4.0­ 4.6 mm, pronotal width 0.80­0.89 mm. Aedeagus larger (Figs. 73­76) with wider apex (Figs. 73­74). Spermatheca larger (Fig. 77)......................................................................................................................... 1. P. comparabilis (Mäklin) – Body smaller, length 3.0­ 3.1 mm, pronotal width 0.59­0.64 mm. Aedeagus smaller (Figs. 82­86) with narrower apex (Figs. 82­83). Spermatheca smaller (Fig. 87)........................................................................................................ 2. P. kenaii Gusarov, sp. n.

8. Seeversiella nigriceps Gusarov, sp. n. (Figs. 117��125) Type material. Holotype,, MEXICO: Puebla: 1.6 km E Nicolas Bravo, 2410 m, forest litter (J.S.Ashe), 17.vii. 1992 (KSEM). Diagnosis. Seeversiella nigriceps can be distinguished from other species of Seeversiella by having brown body; temples twice as long as eyes; glossy pronotum with weak and partially missing microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (Figs. 119��125). Seeversiella nigriceps differs from closely related S. sonomotoides in having narrower distal sclerites of internal sac of aedeagus (Figs. 124; 63, 65). Seeversiella nigriceps can be distinguished from S. globicollis and the other similar species (5 ��7, 10�� 12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac sclerotized and dentiform (Fig. 124); apex of median lobe in lateral view not widened subapically (Figs. 121��122). Description. Length 2.4 mm, pronotal width 0.47 mm. Body dark brown, elytra and legs brown. In all character states S. nigriceps is very similar to S. sonomotoides, but differs in the shape of the distal sclerites of internal sac as described in the diagnosis. In the only known male specimen the posterior angles of male tergum 3 not projecting as spines, tergum 7 in front of posterior margin with small medial tubercle. Aedeagus as in Figs. 119��125. Female unknown. Distribution. Known from Mexico (Fig. 387). Natural History. The only known specimen of S. nigriceps was collected in forest litter at altitude of 2410 m., Published as part of Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-102 in Zootaxa 142 on pages 36-38, DOI: 10.5281/zenodo.156420

Mayatyphlus carltoni Gusarov, sp. n. (Figs. 1��18) Type material. Holotype:, BELIZE: Orange Walk Distr., Rio Bravo Conservation / Management Area, La Milpa Station, Chiche Trail, 17 �� 50 ' 22 "N 89 ��01' 12 "W, lot # 119, berlese (C.E.Carlton & D.Murray), 18.v. 1997 (KSEM). Paratypes: BELIZE:, 2, same data as the holotype (LSAC). Diagnosis. Mayatyphlus carltoni is the only known species of the genus Mayatyphlus. If any additional species of Mayatyphlus are ever discovered, M. carltoni could be distinguished by the shape of the aedeagus, especially the apex of the median lobe (Figs. 13��15), and by the shape of the female accessory sclerites (Fig. 17). Description. Length 0.7��0.8 mm. Head with short setae arranged as in Figs. 1��2. Second and third antennal articles as long as wide and twice as wide as article 3, articles 3��9 strongly transverse, 1.6 ��2.0 times as wide as long, article 10 1.5 times as wide as long, last article as long as wide (Fig. 9). Pronotum with short setae arranged as in Fig. 1, narrowed posteriad, 1.1 times as long as wide, as wide as head and elytra (Fig. 1). Elytra with short setae, with parallel lateral sides (Fig. 1), 1.1 times as long as wide (length measured from humeral level). Abdominal terga with long setae (Fig. 1). Male protarsus with adhesive setae (Fig. 10). Male mesotrochanter with denticle (Fig. 11). Male abdominal sternum 8 with broad apical emargination and narrow medial zone devoid of setae (Fig. 16). Aedeagus with short and narrow parameres, apex of median lobe in lateral view Sshaped, in parameral view asymmetrical, with two subapical processes pointed to the left (Figs. 13��15). Female abdominal sternum 8 as in Fig. 18. Female accessory sclerites as in Fig. 17. Distribution. Known from a single locality in Belize. Natural History. Mayatyphlus carltoni presumably inhabits soil and/or leaf litter, and was collected using Berlese funnels., Published as part of Gusarov, Vladimir I., 2003, Mayatyphlus carltoni Gusarov, a new genus and species of leptotyphline staphylinid beetle from Belize (Coleoptera: Staphylinidae: Leptotyphlinae), pp. 1-7 in Zootaxa 165 on pages 4-5, DOI: 10.5281/zenodo.156671

Psammostiba kamtschatica ( Brundin, 1943), comb. nov. (Figs. 20, 26 in Brundin 1943; Fig. 35 in Yosii & Sawada 1976) Atheta (Panalota) kamtschatica Brundin, 1943: 21 Atheta (Psammostiba) kamtschatica: Yosii & Sawada, 1976: 84. Examined material: Russia: Sakhalin Region:, Kuril Islands, Rasshua Island, SW part, W of Serp Mt., seashore, 47 �� 43.24 'N 152 �� 58.36 'E (Yu.M.Marusik), 12��13.viii. 1992; 3 , Kuril Islands, Urup Island, Negodnaya Bay, Vstrechnyy Ck., seashore, 45 �� 57.55 'N 150 �� 10.33 'E (Yu.M.Marusik), 29.viii. 1995 (all �� SPSU). Diagnosis. Psammostiba kamtschatica can be recognized by smooth posterior margin of male tergum 8, and by the distinct shape of the aedeagus (Figs. 35, H��K: Yosii & Sawada 1976) and spermatheca (Fig. 26: Brundin 1943) Psammostiba kamtschatica differs from P. comparabilis and P. k e n a i i by lack of crenulation of the posterior margin of male tergum 8. Distribution. Psammostiba kamtschatica is known from Russia (Kamchatka Peninsula, Kuril Islands) and northern Japan (Hokkaido)., Published as part of Gusarov, Vladimir I., 2003, A revision of Nearctic species of the genera Adota Casey, 1910 and Psammostiba Yosii & Sawada, 1976 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-35 in Zootaxa 185 on page 33, DOI: 10.5281/zenodo.156388, {"references":["Brundin, L. (1943) Zur Kenntnis einiger in die Atheta - Untergattung Metaxya M. & R. gestellten Arten (Col. Staphylinidae). Lunds Universitets Arsskrift, N. F., Avd. 2, 39 (4), 3 - 37, Taf. 1 - 7.","Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of Genera and Subgenera with description of representative Species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140."]}

Stethusa spuriella (Casey, 1910) (Figs. 58��70) Atheta (s. str.) unigranosa Bernhauer, 1909: 521 (nec Bernhauer, 1908: 363), syn. nov. Atheta (Stethusa) spuriella Casey, 1910: 8. Atheta (Stethusa) subdebilis Casey, 1910: 7, syn. nov. Atheta (s. str.) duplicata Fenyes, 1920: 213 (replacement name for At. unigranosa Bernhauer, 1909, nec 1908), syn. nov. Atheta (Hypatheta) spuriella: Fenyes, 1920: 207 (as valid species). Atheta (Hypatheta) subdebilis: Fenyes, 1920: 207 (as valid species). Atheta macrops Notman, 1920: 725, syn. nov. Atheta (s. str.) unigranosa: Bernhauer & Scheerpeltz, 1926: 642 (as synonym of At. duplicata). Atheta (Hypatheta) spuriella: Bernhauer & Scheerpeltz, 1926: 654 (as valid species). Atheta (Hypatheta) subdebilis: Bernhauer & Scheerpeltz, 1926: 654 (as valid species). Atheta (s. str.) duplicata: Bernhauer & Scheerpeltz, 1926: 642 (as valid species). Atheta (s. str.) macrops: Bernhauer & Scheerpeltz, 1926: 645 (as valid species). Atheta (s. str.) duplicata: Moore & Legner, 1975: 360 (as valid species). Atheta (s. str.) unigranosa: Moore & Legner, 1975: 360 (as synonym of At. duplicata). Atheta (Stethusa) spuriella: Moore & Legner, 1975: 374 (as valid species). Atheta (Stethusa) subdebilis: Moore & Legner, 1975: 374 (as valid species). Xenota unigranosa: Seevers, 1978: 269 (as valid species). Xenota duplicata: Seevers, 1978: 269 (as synonym of At. unigranosa). Xenota spuriella: Seevers, 1978: 269 (as valid species). Xenota subdebilis: Seevers, 1978: 269 (as valid species). Microdota macrops: Seevers, 1978: 265 (as valid species). Type material. Lectotype of Atheta unigranosa (here designated): ��, "Newark, Del. [aware], 5��22.1905 [22.v. 1905]", " 7803 ", "Houghton", " unigranosa Bernh. Typus. Fenyes" (yellow label), "Chicago NHMus. M.Bernhauer Collection (FMNH). Lectotype of Atheta spuriella (here designated): ��, "N. Y. [New York, Catskill Mountains]", " spuriella Csy. ", " TYPE USNM 39439 " (red label), " CASEY bequest 1925 " (NMNH). Lectotype of Atheta subdebilis (here designated): �� (with missing aedeagus), "MO: [Missouri, St. Louis]", " subdebilis Csy. ", " TYPE USNM 39446 " (red label), " CASEY bequest 1925 " (NMNH). Paralectotype: ��, "MO: [Missouri, St.Louis]", " subdebilis PARATYPE USNM 39446 " (red label), " CASEY bequest 1925 " (NMNH). Paratypes of Atheta macrops: UNITED STATES: Florida: 2 ����, "Fla." (no locality data) (AMNH, FMNH); Jefferson Co.: ��, Monticello, 4��8.x. 1914 (AMNH). Additional material. UNITED STATES: Pennsylvania: Westmoreland Co.: ��, Jeannette (H.G.Klages) (FMNH); Ohio: ��, without locality data (Frost) (CASC); Indiana: Lawrence Co.: 2 ���� (W.S.Blatchley), 29.vii. 1909 (CASC); Georgia: Rabun Co.: 2 ����, Clayton, 2000��3700 ' (Leng) (CASC). Diagnosis. Stethusa spuriella differs from S. dichroa in the following characters: a smaller body size; the lack of subbasal impressions of the terminal antennal article; the lack of the distal spines of the internal sac (Figs. 68; 38��39); the shape of the spermatheca (Figs. 70; 41��42); and the lack of a female accessory sclerite. Stethusa spuriella differs from S. klimschi in having a smaller body size; the bent apex of the median lobe (in lateral view; Figs. 65��66; 50��51); and a shorter spermatheca (Figs. 70; 57). Description. Length 2.1��2.5 mm. Head and abdomen dark brown, abdominal segments 3��4 and apical half of segment 7 lighter, pronotum brown, elytra brownish yellow, in some with darker area around scutellum, legs yellow, mouthparts and antennae brown, two basal antennal segments yellow. Head surface glossy, with weak isodiametric microsculpture, with fine punctation, distance between punctures equals 3 times their diameter. Eyes very large, 3.5��4.5 times as long as temples. Antennal article 2 is 1.1 times as long as article 3, articles 4��10 transverse, article 11 slightly longer than articles 9 and 10 combined. Pronotum transverse, 1.2 times as wide as head, width 0.46��0.57 mm, length 0.36��0.44 mm, width to length ratio 1.3; glossy, with very weak isodiametric microsculpture (hardly visible at 70 x); punctation as fine as on head but denser, distance between punctures equal to 1��2 times their diameter. Elytra much wider (0.59��0.73 mm) and longer (0.43��0.57 mm; measured from humeral angle) than pronotum (elytral length to pronotal length ratio 1.2), 1.3 times wider than long, surface glossy, with poorly visible (at 70 x) microsculpture; punctation as on pronotum, slightly asperate. Abdominal terga glossy, with weak microsculpture consisting of transverse waves; with fine punctation, distance between punctures equals 2��4 times their diameter on terga 3��5 and 3��6 times on terga 6��7. Apical margin of tergum 7 with white palisade fringe. Male tergum 7 with weak elongate medial tubercle in front of posterior margin. Posterior margin of male tergum 8 with two blunt medial projections and two lateral denticles (Figs. 58, 60). Aedeagus as in Figs. 63��69, internal sac without distal spines (Fig. 68). Female without accessory sclerite. Spermatheca as in Fig. 70. Discussion. The three species synonymized here with S. spuriella are identical with the types of that species in external characters and the shape of the aedeagus. Distribution. Apparently, S. spuriella is widely distributed in the eastern United States but its distribution is poorly documented (Fig. 111). Natural History. No information is available., Published as part of Gusarov, Vladimir I., 2003, A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-43 in Zootaxa 239 on pages 25-28, DOI: 10.5281/zenodo.156493, {"references":["Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp.","Bernhauer, M. (1909) Neue Aleocharini aus Nordamerika. (Col.) (4. Stuck.). Deutsche Entomologische Zeitschrift, 1909 (4), 515 - 528.","Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1 (2 - 3 )), 283 - 372.","Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.","Notman, H. (1920) Staphylinidae from Florida in the collection of the American Museum of Natural History, with descriptions of new genera and species. Bulletin of the American Museum of Natural History, 42, 693 - 732, plate 39.","Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.","Moore, I. & Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.","Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp."]}