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TRIBE PHYSOCEPHALINI SMITH & PETERSON, 1987 Type genus Physocephala Schiner, 1861: 137. Dacops Speiser, 1923: 99; replacement name for Pseudodacus Kröber, 1915e (preoccupied); type – Pseudodacus abdominalis Kröber, 1915e, designated by Smith, 1980: 513. Pseudodacus Kröber, 1915e: 81; type Pseudodacus abdominalis Kröber, 1915e, designated by Smith, 1980: 513. Physocephala Schiner, 1861: 137; type – Conops rufipes Fabricius, 1781, original designation. Archiphysocephala Kröber, 1939a: 374, 392; type Physocephala gigantea Kröber, 1936 = Physocephala bimarginipennis Karsch, 1887, monotypy [teste Smith & Cunningham-Van Someren, 1970: 439]. Pseudophysocephala Kröber, 1940b: 221; type – Conops platycephala Loew, 1853, original designation.

SUBFAMILY ZODIONINAE RONDANI, 1856 Type genus Zodion Latreille, 1797: 162. Parazodion Kröber, 1927: 135; type – Parazodion schmidti Kröber 1927, monotypy. Robertsonomyia Malloch, 1919: 205; type – Zodion palpalis Robertson, 1901, original designation. Zodiomyia Camras, 1957b: 163; type – Zodiomyia sumbaensis Camras, 1957b, original designation. Zodion Latreille, 1797: 162; type – Myopa cinerea Fabricius, 1794, subsequent monotypy by Latreille, 1802: 444. Neozodion Szilady, 1926: 590; type Neozodion pruinosum Szilady, 1926, monotypy, syn. nov., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 81, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208

MYOPINAE MACQUART, 1834 Type genus Myopa Fabricius, 1775: 798. Included tribes: Myopini, Thecophorini trib. nov. Uncontroverted morphological autapomorphies: basisternum short, narrow, single sclerite (47); veins Sc and R 1 fused before reaching costa (73). A double row of black spines is present on the ventral surface of all femora (52) in all members of Myopinae; this character state is also observed in Baruerizodion and Robertsonomyia + Zodiomyia. Gibson et al. (2012) recover Myopinae as monophyletic with the same set of apomorphic character states, but only include representatives of Myopa, Thecophora, and Pseudoconops. They add the following additional apomorphic character state for those taxa: two, separate, anterior arms on the hypandrium. Macquart (1834) originally described Myopariae, including Myopa, Stachynia [now Dalmannia], and Zodion, with the following diagnostic characters: veins R 4+5 + M not petiolate; alula small; prementum elongate; labellum sometimes elongate and folded back along prementum. In the present analysis, both petiolate and nonpetiolate states of vein R 4+5 + M are observed in genera of Myopinae. The prementum in Myopinae is in the state plesiomorphic to Conopidae. The elongate and ventrally directed labellum is an autapomorphy of Conopidae. In addition to the present members of Myopini and Thecophorini, previous classifications also included genera presently placed within Dalmanniinae, Sicinae, and Zodioninae within Myopinae. See discussions of those subfamilies for details in distinguishing the members of each respective subfamily., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 67, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208, {"references":["Macquart J. 1834. Insectes dipteres du nord de la France. Athericeres: creophiles, oestrides, myopaires, conopsaires, scenopiniens, cephalopsides. Memoires de la Societe (Royale) des sciences, de l'agriculture et des arts a Lille 1833: 137 - 368.","Fabricius JC. 1775. Systema Entomologiae Sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Flensburgi et Lipsiae [= Flensburg and Liepzig]: Kortii.","Gibson JF, Skevington JH, Kelso S. 2012. A phylogenetic analysis of relationships between genera of Conopidae (Diptera) based on molecular and morphological data. Cladistics. doi: 10.1111 / j. 1096 - 0031.2012.00422. x."]}

SUBFAMILY DALMANNIINAE HENDEL, 1916 Type genus Dalmannia Robineau-Desvoidy, 1830. Baruerizodion Papavero, 1970: 121; type – Baruerizodion steyskali Papavero, 1970, original designation. Dalmannia Robineau-Desvoidy, 1830: 248; type – Myopa gemina Wiedemann, 1824 [= Conops aculeatus Linnaeus, 1767], automatic [by designation of the same species (by subsequent designation of Desmarest in d’Orbigny, 1848: 798) for Stachynia Macquart, 1835] (see note in Evenhuis et al., 2010). Stachynia Macquart, 1835; type – Myopa gemina Wiedemann, 1824 [= Conops aculeatus Linnaeus, 1767], designated by Desmarest in d’Orbigny, 1848: 798 [teste Coquillett, 1910: 608]. Dalmania Robineau-Desvoidy, 1853: 92, 146 (unjustified emendation) Robineau-Desvoidy, 1830., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 80, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208

Gibson, Joel F., Skevington, Jeffrey H. (2013): Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data. Zoological Journal of the Linnean Society 167 (1): 43-81, DOI: 10.1111/j.1096-3642.2012.00873.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2012.00873.x

THECOPHORINI GIBSON TRIB. NOV. Type genus Thecophora Rondani, 1845: 15. Included genera: Pseudoconops Camras, 1962a: 183, Scatoccemyia Camras, 1957a: 13, Thecophora. Uncontroverted morphological autapomorphies: costa thickened at insertion of Sc + R 1 (74). Gibson et al. (2012) recover a monophyletic Thecophorini using molecular and morphological analysis, including representatives of Pseudoconops and Thecophora. Additional uncontroverted morphological autapomorphies provided are: dorsal half of proepisternum with patch of setae; hypandrium with broad, setose, anterior plate; and a repeated segment of DNA sequence in the D2 expansion section of the 28S rDNA region. The first character state is found by the present analysis to be an autapomorphy of Pseudoconops + Thecophora. Species examined – Pseudoconops antennatus; Scatoccemyia plaumanni; Thecophora africana; Thecophora atra; Thecophora australiana; Thecophora metallica; Thecophora modesta; Thecophora occidensis., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on pages 67-68, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208, {"references":["Rondani C. 1845. Sulle differenze sessuali delle Conopinae e Myopinae negli insetti ditteri. Memoria undecima per servire alla ditterologia italiana. Nuovi Annali delle Scienze Naturali. Bologna 3: 5 - 16.","Camras S. 1962 a. The Conopidae of Madagascar (Diptera). Memoires de l'Institut Scientifique de Madagascar (E) 13: 179 - 187.","Camras S. 1957 a. Descriptions and records of Neotropical Conopidae (Diptera). Psyche 64: 9 - 16.","Gibson JF, Skevington JH, Kelso S. 2012. A phylogenetic analysis of relationships between genera of Conopidae (Diptera) based on molecular and morphological data. Cladistics. doi: 10.1111 / j. 1096 - 0031.2012.00422. x."]}

SUBFAMILY CONOPINAE LATREILLE, 1802 Type genus Conops Linnaeus, 1758: 604., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 78, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208

SICINAE ZIMINA, 1960 Type genus Sicus Scopoli, 1763: 1004. Included genera: Carbonosicus Zimina, 1958: 933, Sicus. Uncontroverted morphological autapomorphies: male sternite 8 with dense, long, black setae (112). A secondary reversal to reduced posterolateral extensions of the basisternum is present in members of Sicinae (49); a parallel reversal has occurred in members of Conopinae. Gibson et al. (2012) did not test the monophyly of Sicinae, because only one species of Sicus was included in the analysis. Zimina (1958) first suggests the tribe Sicini, including the genera Carbonosicus and Sicus, but does not give diagnostic characters for the group. According to Sabrosky (1999), the lack of a description or diagnosis renders the genus-group name invalid. In Zimina (1960), however, a diagnosis of the tribe is provided with the following key characteristics: basisternum broad; veins Sc and R 1 parallel; cell cup elongate; long red bristles present on sides of mesonotum. According to the present analysis, the first three character states are plesiomorphic to Conopidae. Species examined – Carbonosicus carbonarius; Sicus abdominalis; Sicus ferrugineus., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 68, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208, {"references":["Zimina LV. 1960. New data on Conopidae system (Diptera) exemplified by the representatives of the USSR fauna. Zoologicheskii Zhurnal 39: 723 - 733.","Scopoli JA. 1763. Entomologia Carniolica Exhibens Insecta Carnioliae Indigena et Distributa in Ordines, Genera, Species, Varietates. Methodo Linnaeana. Vindobonae (= Vienna): I. T. Trattner.","Zimina LV. 1958. Carbonosicus, a new Palearctic genus of Conopidae (Diptera). Entomological Review 37: 807 - 809.","Gibson JF, Skevington JH, Kelso S. 2012. A phylogenetic analysis of relationships between genera of Conopidae (Diptera) based on molecular and morphological data. Cladistics. doi: 10.1111 / j. 1096 - 0031.2012.00422. x.","Sabrosky CW. 1999. Family-group names in Diptera. Myia 10: 1 - 371."]}

TRIBE PLEUROCERINELLINI ZIMINA, 1974 Type genus Pleurocerinella Brunetti, 1923: 368. Pleurocerinella Brunetti, 1923: 368; type – Pleurocerinella dioctriaeformis Brunetti, 1923, original designation. Tammo Stuke, 2008: 50: type – Euconops rufus Camras, 1955a, original designation., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 79, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208

TRIBE TROPIDOMYIINI ZIMINA, 1960 Type genus: Tropidomyia Williston, 1888: 11. Physoconops Szilady, 1926: 588; type – Conops brachyrrhynchus Macquart, 1843 = Conops obscuripennis Williston, 1882 [misidentified, see note in Camras, 1955b], original designation. Physoconops (Aconops) Kröber, 1919: 142; type Conops antennatus Kröber, 1915f, designated by Malloch, 1933: 173 [see note in Camras, 1955b]. Physoconops (Aureoconops) Camras, 2004: 86; type – Physoconops aureolus Camras, 2004, original designation. Physoconops (Jelte) Stuke, 2008: 52; type – Pleurocerinella neotropica Camras, 2008, original designation, comb. nov. Physoconops (Kro e beroconops) Camras, 1955b: 175; type – Physoconops hermanni Kröber, 1915f, original designation. Physoconops (Pachyconops) Camras, 1955b: 161; type Conops bulbirostris Loew, 1853, original designation. Physoconops (Shannonoconops) Camras, 1955b: 172; type Physoconops apicalis Camras, 1955b, original designation. Schedophysoconops Gibson, gen. nov.; type Physoconops notatifrons Camras, 1962b: 219, present designation. Tropidomyia Williston, 1888: 11; type – Tropidomyia bimaculata Williston, 1888, monotypy., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 79, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208

BRACHYCERAEINI ZIMINA, 1960 Type genus Brachyceraea Röder, 1892: 366. Genera included: Brachyceraea, Microbrachyceraea Kröber, 1940b: 217, Neobrachyceraea Szilady, 1926: 587. Uncontroverted morphological autapomorphy: scape quadrate (27). The single species of Microbrachyceraea possesses a quadrate scape and petiolate male abdomen, the two key characteristics of Brachyceraeini, thereby warranting its inclusion. The presently proposed membership of Brachyceraeini matches the previously proposed definitions for the group. Zimina’s (1960) original description of Brachyceraeini includes the following diagnostic characters: antennae shortened; basisternum broad with very small posterolateral extensions; vein R 2+3 parallel to R 1 and meeting the costa midway between the end of R 1 and R 4+5. Hennig (1966) also suggests that the reduced antenna is probably a shared, derived apomorphy of the Brachyceraea. The present analysis confirms that the characteristic antennal shape is due to a reduction in the scape. The other characters included by Zimina are plesiomorphic to Conopinae. Species examined – Brachyceraea brevicornis; Microbrachyceraea pendleburyi Neobrachyceraea elongata; Neobrachyceraea obscuripennis., Published as part of Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, pp. 43-81 in Zoological Journal of the Linnean Society 167 (1) on page 63, DOI: 10.1111/j.1096-3642.2012.00873.x, http://zenodo.org/record/5292208, {"references":["Zimina LV. 1960. New data on Conopidae system (Diptera) exemplified by the representatives of the USSR fauna. Zoologicheskii Zhurnal 39: 723 - 733.","Roder VV. 1892. Brachyceraea nov. gen. novum genus Conopidarum (Diptera). Entomologische Nachrichten, Berlin 18: 366.","Krober O. 1940 b. Beitrage zur Kenntnis der Conopiden - VI. Annals and Magazine of Natural History Series 11, 5: 203 - 245.","Szilady Z. 1926. Dipterenstudien. Annales Musei Nationlis Hungarici 24: 586 - 593.","Hennig W. 1966. Conopidae im Baltischen Bernstein (Diptera; Cyclorrhapha). Stuttgarter Beitrage zur Naturkunde 154: 1 - 24."]}