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Aspidistra triradiata Vislobokov (2015: 269) (Fig. 3) Literature: — Wei (2018: 622), Wei et al. (2022: 112), Tillich (2023: 6). TYPE: — VIETNAM. Vinh Phuc province: Tam Dao district, Tam Dao National Park, about 6 km NW of Tam Dao town, 21°29.942’ N 105°36.973’ E, elev. 1208 m, rainforest, 19 November 2014, N. A. Vislobokov 14098 (holotype: MW: MW0595636!; flowering). Image of holotype available at: https://depo.msu.ru/open/public/en/item/MW0595636 Additional specimens examined: — CHINA. Guangxi Zhuang Autonomous Region: Baise City, Napo County, Baishen town, Nonglong village, slope of limestone hill, 23°14.377’ N 105°33.764’ E, elev. 1188 m, 22 November 2016, N. A. Vislobokov, M. S. Nuraliev, B. Pan, F. Wen, Z. B. Xin G30 (IBK: IBK00407784; MW: MW0753785, MW0753786; flowering); living plant from the same gathering cultivated in the Main Botanical Garden of the Russian Academy of Sciences (Moscow), garden number: 2016.12342; Guangxi Zhuang Autonomous Region: Laibin City, Jinxiu Yao Autonomous County, Dayao Mountain Park, Shengtang mountain, mountain slope, forest with bamboo, 23°58.493’ N 110°06.741’ E, elev. 1229 m, 11 November 2017, N. A. Vislobokov, M. S. Nuraliev, J. Dong G168 (IBK; MW: MW0754786, MW0754787, MW0754788; fruiting); living plant from the same gathering cultivated in the Main Botanical Garden of the Russian Academy of Sciences (Moscow), garden number: 2017.13475 (flowered in March 2018). Notes:— Aspidistra triradiata belongs to the so-called A. hainanensis species complex. This complex is characterized by tufted leaves, i.e., by presence of several foliage leaves per elementary shoot (Tillich & Averyanov 2012, Tillich 2014, 2023, Averyanov et al. 2018, Ding et al. 2021, Vislobokov et al. 2021), and distinguished from other species with tufted leaves by perigone lobes with 4–6 keels and mushroom-shaped pistils (Tillich 2023). Specimen identification within this complex is known to be complicated by incompletely understood morphological boundaries and geographical distribution of its species. A specimen Le Tuan Anh s.n. collected in Quang Tri province of Vietnam and assigned to A. triradiata was reported as a source for investigation of chemical compounds by Le et al. (2022, 2023a, 2023b). After investigation of analytical field photographs of this specimen kindly provided by Le Tuan Anh (see also LE: LE 01123061), a collector of this specimen, we concluded that it shows considerable differences from A. triradiata, including campanulate (vs urceolate) perigone tube and stigma entirely white above (vs with three purple radial lines). We here re-identify this specimen as A. hainanensis W.Y.Chun & F.C.How in Guangdong Institute of Botany (1977: 533) s.s. Wei (2018) included A. triradiata in a list of native plants of Guangxi and indicated this species to be found in Napo (where our collection Vislobokov et al. G 30 was also made). However, Wei (2018) did not cite any collections or references for this occurrence, which makes it impossible to be verified, especially given the taxonomic difficulties in A. hainanensis complex described above. Here, we report A. triradiata from China confirmed by specimen investigation and photographs of living flowers for the first time. Distribution:— China (Guangxi: counties Napo and Jinxiu), Vietnam (Vinh Phuc: Tam Dao district). Ecology and phenology:— Aspidistra triradiata inhabits diverse types of evergreen forests, including forests on limestone karst, at elevations around 1200 m a.s.l. Flowering and fruiting in nature in November., Published as part of Nuraliev, Maxim S., Romanov, Mikhail S., Drokina, Victoria G., Fu, Long-Fei & Vislobokov, Nikolay A., 2023, Two newly recorded species of Aspidistra (Asparagaceae) for the flora of China: A. obliqua and A. triradiata, pp. 248-254 in Phytotaxa 600 (4) on page 251, DOI: 10.11646/phytotaxa.600.4.4, http://zenodo.org/record/8093997, {"references":["Vislobokov, N. A. (2015) Two new species of Aspidistra (Asparagaceae, Nolinoideae) from northern Vietnam: A. clausa and A. triradiata. Phytotaxa 207 (3): 265 - 272. http: // dx. doi. org / 10.11646 / phytotaxa. 207.3.4","Wei, Y. G. (2018) The distribution and conservation status of native plants in Guangxi, China. China Forestry Publishing House, Beijing, 876 pp.","Wei, Y. G., Do, T. V. & Wen, F. (2022) A checklist to the plants of northern Vietnam. China Forestry Publishing House, Beijing, 606 pp.","Tillich, H. J. (2023) 200 years Aspidistra (Asparagaceae), and now more than 200 species: a new comprehensive determination key, and an annotated bibliography of the genus. Nordic Journal of Botany: e 03818. https: // doi. org / 10.1111 / njb. 03818","Tillich, H. - J. & Averyanov, L. V. (2012) Four new species of Aspidistra Ker Gawl. (Asparagaceae) from China and Vietnam with a comment on A. longifolia Hook. f. and A. hainanensis W. Y. Chun & F. C. How. Gardens' Bulletin Singapore 64 (1): 201 - 209.","Tillich, H. J. (2014) The genus Aspidistra Ker-Gawl. (Asparagaceae) in Vietnam. Taiwania 59 (1): 1 - 8. https: // doi. org / 10.6165 / tai. 2014.59.1","Averyanov, L. V., Tillich, H. J., Pham, V. T., Nguyen, S. K., Le, T. A., Nguyen, H. T., Maisak, T. V., Tuan, A. H. L., Nguyen, D. D., Truong, Q. C., Nguyen, T. L. T. & Vu, T. C. (2018) New taxa and taxonomic notes in Aspidistra (Convallariaceae s. s.) in China, Laos and Vietnam. Nordic Journal of Botany 36 (7): e 01833. https: // doi. org / 10.1111 / njb. 01833","Ding, H. B., Xiong, K. C., Yang, B., Yin, J. T., Bouamanivong, S. & Tan, Y. H. (2021) New species and taxonomic notes of Aspidistra (Asparagaceae) for the flora of China and Laos. Taiwania 66 (4): 439 - 449. https: // doi. org / 10.6165 / tai. 2021.66.439","Vislobokov, N. A., Fu, L. F., Wei, Y. G. & Nuraliev, M. S. (2021) Leaf epidermal micromorphology in Aspidistra (Asparagaceae): diversity and taxonomic significance. PhytoKeys 185: 65 - 86. https: // doi. org / 10.3897 / phytokeys. 185.72259","Le, H. B. T., Ho, D. V., Nguyen, H. M., Le, A. T., Tran, T. V. A. & Nguyen, H. T. (2022) A new spirostanol steroid and a new spirostanol steroidal saponin from Aspidistra triradiata and their cytotoxic activities. Natural Product Research: 1 - 10. https: // doi. org / 10.1080 / 14786419.2022.2095635","Le, H. B. T., Ho, D. V., Nguyen, H. M., Tran, T. V. A. & Nguyen, H. T. (2023 a) Aspidiatas C and D, two new spirostanol saponins from Aspidistra triradiata and their cytotoxic activities. Natural Product Research: 1 - 9. https: // doi. org / 10.1080 / 14786419.2023.2204432","Le, H. B. T., Tran, T. V. A., Nguyen, H. M., Nguyen, L. T. K., Ho, D. V. & Nguyen, H. T. (2023 b) Aspidiata E, a new spirostanol saponin from Aspidistra triradiata and its cytotoxic activity. Natural Product Communications 18 (6): 1 - 7. https: // doi. org / 10.1177 / 1934578 X 231179412","Guangdong Institute of Botany (1977) Liliaceae. In: Guangdong Institute of Botany (eds.) Flora Hainanica 4. Science Press, Beijing, pp. 107 - 119, 533 - 534. [in Chinese]"]}

Nuraliev, Maxim S., Romanov, Mikhail S., Drokina, Victoria G., Fu, Long-Fei, Vislobokov, Nikolay A. (2023): Two newly recorded species of Aspidistra (Asparagaceae) for the flora of China: A. obliqua and A. triradiata. Phytotaxa 600 (4): 248-254, DOI: 10.11646/phytotaxa.600.4.4, URL: http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN

Aspidistra obliqua K.S.Nguyen & Aver. in Averyanov et al. (2020a: 9) (Fig. 2) Literature: — Averyanov et al. (2020b: 155, Fig. 337, 338), Wei et al. (2022: 111), Tillich (2023: 11). TYPE: — VIETNAM. Ha Giang province: Quan Ba district, Can Ty commune, Sin Suoi Ho village, around point 23.09566° N 105.02142° E, very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800–1000 m a.s.l., remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest, terrestrial herb in shady place among rocks, 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak VR 1548 (holotype: LE: LE01067134 photo!; isotypes: HN, MW: MW0595840!; flowering and fruiting). Image of holotype available at: https://en.herbariumle.ru/?t=occ&id=12854 Photos of living plants used for preparation of type specimens available at: https://en.herbariumle.ru/?t=occ&id=13148 https://en.herbariumle.ru/?t=occ&id=99504 Additional specimens examined: — CHINA. Guangxi Zhuang Autonomous Region: Chongzuo City, Longzhou County, Zhubu town, Nongzai village, path to Sancunshan hill, limestone slope, 22°29.335’ N 106°56.212’ E, elev. 250 m, 25 November 2016, N. A. Vislobokov, M. S. Nuraliev, B. Pan, F. Wen G72 (IBK; MW: MW0753756, MW0753762, MW0753767, sterile); living plant from the same gathering cultivated in the Main Botanical Garden of the Russian Academy of Sciences (Moscow), garden number: 2016.12352 (and spirit material of flowers from this plant in collection of Moscow University, flowered in May 2022). Notes:—The Chinese specimen shows several significant morphological differences from the type specimen of A. obliqua, which, however, do not prevent us from assigning our specimen to this species. In particular, in the living material of Vislobokov et al. G 72, the leaf blade is lustrous (vs velvety); however, this difference is not evident when comparing dried specimens. Then, the stamens are inserted at level of 1/3 of perigone tube (vs at middle of perigone tube), and the pistil is evidently mushroom-shaped with convex stigma (vs obconoid). The pistil in the Chinese specimen is white in the basal half, whereas it is described as dark purple black to black in the protologue. At the same time, Figure 6 in Averyanov et al. (2020a) depicts a flower (right side of plate, second flower from above) with the same pistil coloration as in the Chinese specimen. In addition, in Vislobokov et al. G72, the stamens are free from the pistil (similarly to all the other known species of the genus), whereas Averyanov et al. (2020a) described the type material to uniformly have stamens densely appressed to the pistil right below stigma, and (1–)2(–3) of them fused with the pistil by filament apices. We suppose that such an unusual floral construction could be a result of a teratological nature of the type material. Distribution:— China (Guangxi: Longzhou County), Vietnam (Ha Giang: Quan Ba district). Ecology and phenology:— Aspidistra obliqua inhabits evergreen broad-leaved and coniferous forests on limestone karst at elevations of 250–1000 m a.s.l. Flowering and fruiting in nature in October–November., Published as part of Nuraliev, Maxim S., Romanov, Mikhail S., Drokina, Victoria G., Fu, Long-Fei & Vislobokov, Nikolay A., 2023, Two newly recorded species of Aspidistra (Asparagaceae) for the flora of China: A. obliqua and A. triradiata, pp. 248-254 in Phytotaxa 600 (4) on pages 249-251, DOI: 10.11646/phytotaxa.600.4.4, http://zenodo.org/record/8093997, {"references":["Averyanov, L. V., Nguyen, K. S., Son, H. T., Tillich, H. J., Wynn-Jones, B. & Maisak, T. V. (2020 a) New taxa and new records in Aspidistra (Convallariaceae s. s.) of Laos and Vietnam. Nordic Journal of Botany 38: e 02877. https: // doi. org / 10.1111 / njb. 02877","Averyanov, L. V., Nguyen, K. S., Tran, T. H., Averyanova, A. L., Maisak, T. V. & Nguyen, H. T. (2020 b) Plant diversity, flora and vegetation of Bat Dai Son Mountain area, northern Vietnam. Strata, Saint-Petersburg, 561 pp.","Wei, Y. G., Do, T. V. & Wen, F. (2022) A checklist to the plants of northern Vietnam. China Forestry Publishing House, Beijing, 606 pp.","Tillich, H. J. (2023) 200 years Aspidistra (Asparagaceae), and now more than 200 species: a new comprehensive determination key, and an annotated bibliography of the genus. Nordic Journal of Botany: e 03818. https: // doi. org / 10.1111 / njb. 03818"]}

Mitreola spathulifolia D.Fang & L.S.Zhou in Fang et al. (1995: 33, Fig. 9–12) (Fig. 4) Literature:— Huang (2007: 79), Chen (2011: 761). Type: — CHINA. Guangxi Zhuang Autonomous Region: Mashan, Guling, in cavern of limestone hill, 310–330 m, 07 June 1994, Fang D., Zhou L. S ., Xie Z. M. 3146 (lectotype, designated here: GXMI: GXMI050420 photo!; isolectotypes: GXMI: GXMI050418 photo!, GXMI050419 photo!, GXMI050421 photo!; GXNM). Images of lectotype and isolectotypes available at https://www.cvh.ac.cn/spms/detail.php?id=09174c08 https://www.cvh.ac.cn/spms/detail.php?id=09174aeb https://www.cvh.ac.cn/spms/detail.php?id=09174b79 https://www.cvh.ac.cn/spms/detail.php?id=09174c92 Nomenclatural notes: —In the original description of Mitreola spathulifolia, Fang et al. (1995) indicated the duplicate of the collection Fang et al. 3146 at GXMI as a holotype, and its duplicate at GXNM as an isotype. However, we have found four duplicates of Fang et al. 3146 in GXMI matching the detailed description and specimen label information included in the protologue. These duplicates should be treated as syntypes in accordance with Art. 9.6 of ICN (Turland et al. 2018). We selected the specimen with the barcode GXMI 050420 as the lectotype following Arts. 9.3 and 9.11 of ICN (Turland et al. 2018) as it represents best the species, having an intact well-developed stem and numerous inflorescences. The other specimens at GXMI (GXMI 050418, GXMI 050419, GXMI 050421), as well as the one at GXNM (not seen), are isolectotypes. Of them, the specimen GXMI 050418 has been labeled in shed. as a “ holotypus ” during or after preparation of the protologue; the plant in this specimen has only the distal portion of the stem., Published as part of Nuraliev, Maxim S., Lyskov, Dmitry F., Kuznetsov, Andrey N., Kuznetsova, Svetlana P. & Fu, Long-Fei, 2023, Novelties in Asian Mitreola (Loganiaceae): M. capitata, a new species from Vietnam, and lectotypification of M. spathulifolia, pp. 39-47 in Phytotaxa 585 (1) on page 44, DOI: 10.11646/phytotaxa.585.1.4, http://zenodo.org/record/7672924, {"references":["Fang, D., Qin, D. H., Zhou, L. S. & Lu, X. H. (1995) Four new species of Mitreola Linn. (Loganiaceae) from Guangxi. Journal of Tropical and Subtropical Botany 3 (3): 30 - 35. https: // dx. doi. org / 10.3969 / j. issn. 1005 - 3395.1995.3.004","Huang, S. X. (Ed.) (2007) Plants endemic to Guangxi province. Vol. 1. Guangxi Science & Technology Publishing House, Nanning, 160 pp. [in Chinese]","Chen, X. X. (2011) Loganiaceae. In: Guangxi Inst. of Botany (Org.) Flora of Guangxi. Vol. 3. Guangxi Science and Technology Press, Nanning, pp. 756 - 772. [In Chinese]","Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. & Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. [Regnum Vegetabile 159]. Koeltz Botanical Books, Glashutten. https: // doi. org / 10.12705 / Code. 2018"]}

Mitreola capitata Nuraliev (Fig. 1–3) Diagnosis: — Mitreola capitata is unique among the species of the genus Mitreola in having the following combination of characters: short stem with rosulate leaves, hairs on leaves and peduncle and a capitate inflorescence with a long peduncle. It is most close to M. spathulifolia, differing in leaves 5.9–12 cm long, a dense capitate inflorescence 0.8–2.3 cm in diameter when flattened, pedicel ca. 0.5 mm long, hairy calyx, and capsule 2 mm long and 2–2.5 mm wide. Type: — VIETNAM. Quang Binh Province: Bo Trach District, Tan Trach commune, Phong Nha-Ke Bang National Park, limestone rock, 17°24’41’’N 106°13’59’’E, elev. 650 m, 30 March 2022, Nuraliev M. S ., Lyskov D. F., Dac L. X., Ha L. M., Thang D. Q. NUR 3614 (holotype: MW: MW0595774; isotype: MW: MW0595775). Description:—Herb lithophytic, perennial. Stem rhizome-like, ascending to vertical, to 4 cm long, ca. 5 mm in diameter (in old plants), branched in old plants, pubescent with dense spreading white hairs ca. 0.3 mm long when young; internodes usually vanishingly short, in young shoots up to ca. 4 mm long, rarely up to 12 mm long especially in lateral branches; plant with tap root up to ca. 1.5 mm in diameter at base, at basal portion of stem (with leaves withered) sometimes with thin adventitious roots ca. 0.4 mm in diameter. Stipules interpetiolar, flange-like to scalelike, in latter case broadly triangular to ovate, up to ca. 1 mm long and 2 mm wide, glabrous; nodes in axils of stipules and leaves bearing numerous narrowly clavate gland-like papillae (colleters) ca. 0.5–1 mm long with blackish apices. Leaves rosulate, opposite, equal in each pair, sessile; leaves persistent in several distal nodes of each shoot. Leaf blade chartaceous in sicco, obovate (to broadly oblanceolate), adaxially dark green with a distinct whitish green midvein in vivo, abaxially pale green in vivo, both sides brownish green in sicco, larger blades on each shoot (5.9)7.0–11.0(12.0) cm long and (2.6)2.9–4.4(5.2) cm wide, (1.6)1.8–3.7 times as long as wide; base cuneate to nearly attenuate; apex very broadly acute to obtuse or almost rounded; margin entire, ciliate with white hairs up to 0.5 mm long; both surfaces pubescent with dense erect white hairs ca. 0.5 mm long; midvein ca. 0.8–1 mm wide at base, gradually becoming thinner towards apex; secondary veins pinnately arranged, 6–7(8) on each side of midvein, ascending; midvein and secondary veins adaxially slightly impressed, abaxially prominently raised; tertiary veins conspicuous on both sides. Inflorescences evidently terminal, possibly also in axils of upper leaves; peduncle erect, green, 8.5–19.5 cm long, covered with dense spreading white hairs 0.5 mm long. Inflorescence an umbel-like thyrsoid with several cymes (cymes mostly dichasial proximally and monochasial distally), or umbel of such thyrsoids, dense, capitate (head-like), ca. 1.0–2.0 cm in diameter (0.8–2.3 cm when flattened), with numerous congested flowers; all axes similar to peduncle but thinner and with shorter hairs. Bracts appressed to axes, narrowly triangular, green, basal ones (1.5)2.0–2.5(3.0) mm long and 0.4–1.0 mm wide at base, distal ones gradually becoming smaller, adaxially glabrous, abaxially sparsely to densely hairy, with acute apex and slightly serrate margin; small hairy colleters usually present near bract bases. Flower actinomorphic, 5-merous (except for gynoecium). Pedicel ca. 0.5 mm long (rarely flower sessile), with spreading white hairs. Calyx cupuliform-campanulate, 1.5–1.8 mm long, externally (abaxially) covered with sparse spreading white hairs, internally (adaxially) glabrous; tube green, 0.6–0.7 mm long; lobes triangular, green with greenish white margins and apex, 0.7–1.2 mm long, 0.9–1.2 mm wide at base, with acute apex and minutely ciliate margin. Corolla urceolate, white, somewhat translucent in proximal half, ca. 2.8–3.2 mm long and ca. 3.5 mm in diameter when fully open; tube 2.0– 2.5 mm long, 2.0 mm in diameter; lobes horizontally spreading and somewhat curved outwards, imbricate in bud, broadly triangular, 1.0– 1.1 mm long, 0.7–0.9 mm wide at base, with obtuse apex and entire margin; corolla inside with a circular thickening at level of lobe bases bearing a horizontal row of uniseriate multicellular trichomes 0.6 mm long; trichomes pointing at ca. 45° upwards, forming a conical structure and completely obstructing the orifice of the corolla tube; corolla otherwise glabrous. Stamens included in corolla tube, glabrous; filaments inserted at middle of corolla tube, slender and slightly narrowing towards apex, white, ca. 0.4 mm long, pointing at ca. 45° upwards; anthers dorsifixed, ovate, pale yellow, 0.2 mm long, arranged at level of stigmas, introrse. Gynoecium green, glabrous; ovary superior, 2-locular with axile placentation, broadly conical, 0.2 mm long, 0.7–0.8 mm in diameter at base, each locule bearing about 20 ovules; styles 2, tightly connivent with each other into a common style; common style included in corolla tube, very narrowly conical, ca. 0.8 mm long; common stigma capitulate. Fruit (capsule) 2 mm long, 2–2.5 mm wide, glabrous, laterally slightly compressed, bilobed at distal 1/2, lobes divergent, dehiscent along ventral sutures of lobes, bearing persistent calyx ca. 1 mm long (when withered). Seeds several per fruit, black, ca. 0.5 mm in diameter. Etymology: —The specific epithet “ capitata ” refers to head-like inflorescences which distinguish the new species from all its congeners. Phenology: —On 30 March 2022, the plants were in full flower, and also bore completely withered fruiting inflorescences (with dehisced capsules) of the previous season. Distribution and ecology: —The new species is currently known from a single location in Phong Nha-Ke Bang National Park, Vietnam. The species inhabits a prominent outcrop of limestone karst at elevation of 650 m a.s.l., where it occurs in crevices and on vertical walls. Several dozens of individuals were observed. Mitreola capitata is associated with the dominant tree species Calocedrus rupestris Aver., T.H.Nguyên & L.K.Phan (Cupressaceae) and the herbs including Selaginella sp. (Selaginellaceae), Amorphophallus glossophyllus Hett. (Araceae), Ophiopogon sp. (Asparagaceae), Polygala umbonata Craib (Polygalaceae), Elatostema spp. (Urticaceae), Begonia sp. (Begoniaceae), Strobilanthes sp. (Acanthaceae) and various ferns. Taxonomic relationships: — Mitreola capitata is readily distinguishable from most of its congeners by short stem with rosulate leaves, presence of indumentum (especially the hairs on leaves and peduncle) and compact longpedunculate inflorescence. Among the previously described species of Mitreola, this combination of features is only found in M. spathulifolia endemic to Daxin and Mashan counties of Guangxi, China. Apart from the habit, the two species share confinement to limestone cliffs. The air distance between the type location of M. capitata and the nearest reported population of M. spathulifolia (its paratype from Daxin: Fang, Zeng 195) is about 590 km. The main morphological differences between M. capitata and M. spathulifolia are summarized in Table 1. Most important of them are the leaf length (5.9–12 cm vs. 7–28 cm), size and appearance of inflorescence (0.8–2.3 cm in diameter when flattened, dense and capitate vs. 2–5 cm in diameter when flattened, quite lax), pedicel length (ca. 0.5 mm vs. 1–2 mm), calyx indumentum (hairy vs. glabrous) and capsule size (2 mm long, 2–2.5 mm wide vs. 3 mm long, 5 mm wide). The inflorescence appearance, although not described in the protologue of M. spathulifolia, seems to be especially useful for field recognition of this species against M. capitata: the lax inflorescences are evident from the photos provided by Huang (2007) and presented in this study (Fig. 4B)., Published as part of Nuraliev, Maxim S., Lyskov, Dmitry F., Kuznetsov, Andrey N., Kuznetsova, Svetlana P. & Fu, Long-Fei, 2023, Novelties in Asian Mitreola (Loganiaceae): M. capitata, a new species from Vietnam, and lectotypification of M. spathulifolia, pp. 39-47 in Phytotaxa 585 (1) on pages 40-44, DOI: 10.11646/phytotaxa.585.1.4, http://zenodo.org/record/7672924, {"references":["Huang, S. X. (Ed.) (2007) Plants endemic to Guangxi province. Vol. 1. Guangxi Science & Technology Publishing House, Nanning, 160 pp. [in Chinese]"]}