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Key to species of the genus Niladmirara gen. n. (by males) 1. Gonostyles with claw round apically (Figs 1, 23). Basal margin of genitalia with medial excision (Fig. 1). Parameres much longer than gonocoxites. Aedeagus with large basal swollen, prolonging far behind genitalia. Neck F5 1.7 times as long as burrel-shaped basal enlargement. Antennae 2+16-17 segments. Empodium as long as tarsal claw (Fig. 6, 7) or longer one. Body length 1.43 mm; length of wing 2.18 mm ……….......……………...……………................................... N. metula sp. n. – Gonostylus with very thin small pointed claw (Figs 10, 20, 24). Basal margin of genitalia is round (Figs 10, 24). Parameres shorter than gonocoxites. Rod of aedeagus reaches the base of genitaia (Fig. 24) or slightly longer one (Figs 10). Neck F5 1.2 or 1.8 times as long as basal enlargement (Fig. 11).......................... 2 2. Projections of parameres almost equal length (Fig. 24). Antennae 2+22-23 segments, apical flagellomere cone shaped. Neck of middle flagellomeres 1.2 times as long as subcylindric basal enlargement. Empodium longer than tarsal claw. Length of wing 3.2 mm.................................................................................. …………............................... N. mucronata (Mamaev et Zaitzev, 1998), comb. n. – One from projections of parameres much longer than other (Fig. 10). Antennae 2+15-16 segments, terminal flagellomere with round apex (Fig. 18). Neck F5 1.8 times as long as barrel-shaped basal enlargement (Fig. 17). Empodium shorter than tarsal claw (Fig. 15). Body length 1.58 mm; length of wing 2.46 mm ………. ……………………………………..…….............................. N. devergentia sp. n., Published as part of Fedotova, Z. A., 2018, New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East, pp. 1-12 in Far Eastern Entomologist 362 on pages 9-10, DOI: 10.25221/fee.362.1, http://zenodo.org/record/7164187

Ganseriella pankowskiorum Fedotova et Perkovsky, sp. n. urn:lsid:zoobank.org:act: 671FBEB1-9925-4D64-AEC6-53D035AB8D44 M a t e r i a l. Holotype. SIZK Bu–1, well preserved inclusion of male with tarsi missing, Burmese amber, earliest Cenomanian (fig. 3, 1 –10). D e s c r i p t i o n. Male (fig. 3, 1 –10). Body (with genitalia) is 1.3 times shorter than antennae. Antenna 1.5 times as long as wing. Head with large eyes that laterally cover all width of head. Scape 1.8 times as long as round pedicel. 1st flagellomere 6.0 times as long as wide. 2nd flagellomere as long as 1st. 5th flagellomere 3.8 times as long as wide, slightly shorter than 1st flagellomere; basal enlargement 1.8 times shorter than neck. 13th flagellomere 1.2 times as long as 14th and 0.56 times as long as 5th; 14th flagellomere with long projection and rounded apically. Palpi very narrow, parallel-sided laterally. 2nd, 3rd and 4th segments strongly prolonged, more than 2.0–2.5 times as long as 1st. Wing is 2.7 times as long as wide. Vein R 1+2 0.63 times as long as wing length. Hind femur 1.2 times as long as profemur. Ratio of hind tarsi 1:7.8:5.8:2.3:1.3. Gonostylus wide and almost parallel-sided distally, 1.7 times as long as gonocoxites. Tegmen wide, narrowed near apex, reaching short of gonostylus. M e a s u r e m e n t s (mm): body length, 1.60; antennal length, 2.13; head length, 0.26; width of lateral side of head, 0.15; palpi length, 0.23; thorax length, 0.38; wing length, 1.43; wing width, 0.53; vein R 1+2 , 0.90; halter length, 0.21; length of fore: coxa, 0.20; trochanter, 0.05, femur, 0.77, tibia, 0.71; metatarsus, 0.11; length of middle: femur, 0.69, tibia, 0.74, length of hind: femur, 0.93; tibia, 0.85; abdomen length, 0.98; genitalia length, 0.09. E t y m o l o g y. The species is named after Mark A. Pankowski and his father Mark S. Pankowski (Rockville, Maryland, USA) for donating the holotype to the amber collection of the Schmalhausen Institute of Zoology, the National Academy of Sciences of Ukraine. We are grateful to A. P. Rasnitsyn (Borissiak Paleontological Institute, Russian Academy of Sciences) for the constant help., Published as part of Perkovsky, E. E. & Fedotova, Z. A., 2017, The Second Cretaceous Gall Midge Genus Of The Tribe Diallactiini (Diptera, Cecidomyiidae) From The Late Cretaceous Burmese Amber, pp. 117-124 in Vestnik Zoologii 51 (2) on page 123, DOI: 10.1515/vzoo-2017-0017, http://zenodo.org/record/6404136

Genus Ganseriella Fedotova et Perkovsky, gen. n. urn:lsid:zoobank.org:act: 5093FEDA-A88E-485D-ABCD-9DA8145C85A8 T y p e s p e c i e s: Ganseriella pankowskiorum Fedotova et Perkovsky, sp. n. D e s c r i p t i o n (figs 1, 1–3; 2, 1 –8; 3, 1–10). Male. Body slightly longer than wings. Head elongate; eyes fused, positioned around head, without narrow bridge, occiput wide, rounded. Antennae 2+14-segmented, scape very long, slightly enlarged distally, basally narrower than round pedicel. All flagellomeres with distinct very narrow and long necks, but basal enlargements have different forms. Basal enlargement on proximal flagellomeres almost parallel-sided, on medial flagellomeres prolonged swollen, on distal flagellomeres slightly swollen or almost round. Basal enlargement of 1st and 2nd flagellomeres slightly narrowed medially. Proximal flagellomeres much wider and longer than distal ones. Basal enlargement of flagellomeres with basal whorls of very short dense setae and medial crenulate whorls of very long setae strongly curved basally. Distal part of basal enlargement with apical whorl of very long setae reaching of next flagellomere. Necks of flagellomeres always longer than basal enlargement, especially in middle flagellomeres. Mouthparts strongly elongate and wide. Palpi much longer than height of head, 4-segmented, the 3rd and 4th ones longest. Wing evenly and very strongly widened medially, vein m+rm slightly curved medially, R 5 slightly curved apically, joining margin of wing behind wing tip; rs very strong; Cu simple, positioned far from margin of wing, reaching wing edge, forming large lobe. Anal vein clear. Cell between veins C and R 1+2 distinctly sclerotized. Costal vein with break beyond its joining point with R 5 . Legs completely densely covered with short setae and sparse long setae. Fore and hind femora longer than their tibia, middle femur and tibia. Middle femora shorter than tibia. All tibia subequal in length. Abdomen slightly swollen near middle (2nd–4th segments) and narrowed distally. Abdominal segments covered by very long setae. Genitalia transverse. Gonocoxite wide, slightly rounded on outside margin, with dense long setae and almost rectangular on inner margin. Gonostylus long and strongly enlarged basally, with long apical dent. Tegmen very long, pointed apically, much longer than gonocoxites (lateral view, fig. 3, 9). C o m p a r i s o n. The shape of the body, 2+14-segmented antennae, wing venation, long legs and forms of gonocoxites and gonostyles of the new genus imply its close relation to Johnsonomyia Felt, 1908 (Felt, 1908; Mamaev, 1964, 1966; Spungis, 1985; Jaschhof, Jaschhof, 2013; Jaschhof, 2016), but the new genus differs from it in the elongated segments of palpi, very large eyes, absence of narrow eye bridge, different shape of flagellomeres, very long necks of flagellomeres, presence of very dense long setae of apical whorl of the basal enlargement of flagellomeres, wider and shorter wing, vein R 5 joining the wing margin far behind the wing tip, presence of sclerotisation of the wing cell between veins C and R 1+2 ; very long pointed tegmen, and smaller body size. The new genus is closely related to Chastomera Skuse, 1889, which is currently considered a synonym of Haplusia Karsch, 1877, but differs from it in a longer rs and part of R 1+2 from rs to fusing with C, in the very long palpi, in absence of dorso-apical projection on medial surface of gonocoxite, in the presence of long pointed tegmen, and in the smaller body size. Chastomera Skuse, 1888 stat. resurr. is obviously a separate genus, distinct from Haplusia, as the redescription of the holotype of type species Ch. bella Skuse shows (Skuse, 1888; Kolesik, Gagné, 2016) and does not meet the original diagnosis of Haplusia (Karsch, 1877) and the numerous diagnoses of the genus Haplusia given earlier (Mamaev, 1964, 1966; Gagné, 1978; Jaschhof, 2016; Kolesik, Gagné, 2016). The new genus differs from Haplusia in very long pointed tegmen, very wide cell of wing between veins C and R 4+5 , and also CuA and wing margin, very long and short dense setae in apical whorl of basal enlargement of flagellomeres (only short in Haplusia) and very long neck of middle flagellomeres, additionally.in the absence of a dark spot around vein rs and on vein CuA and in the not banded legs. E t y m o l o g y. The generic name is an arbitrary combination of letters. Gender feminine., Published as part of Perkovsky, E. E. & Fedotova, Z. A., 2017, The Second Cretaceous Gall Midge Genus Of The Tribe Diallactiini (Diptera, Cecidomyiidae) From The Late Cretaceous Burmese Amber, pp. 117-124 in Vestnik Zoologii 51 (2) on pages 119-123, DOI: 10.1515/vzoo-2017-0017, http://zenodo.org/record/6404136, {"references":["Felt, E. P. 1908. Appendix D. Pp. 286 - 422, 489 - 510, pls. 33 - 34. In his 23 d report of the State Entomologist on injurious and other insects of the State of New York 1907. New York State Museum Bulletin, 124, 5 - 541, 44 pl.","Mamaev, B. M. 1964. Gall midges of the USSR. 6. New species of the tribe Porricondylini (Diptera, Cecidomyiidae). Entomologicheskoe Obozrenie, 43, 894 - 913 [In Russian].","Mamaev, B. M. 1966. New and little known Palaearctic gall midges of the tribe Porricondylini (Diptera, Cecidomyiidae). Acta Entomologica Bohemoslovaca, 63, 213 - 239 [In Russian].","Spungis, V. 1985. Gall midges of the subtribe Diallactina (Diptera, Cecidomyiidae) in Latvia. Latvijas Entomologs, 28, 38 - 53.","Jaschhof, M., Jaschhof, C. 2013. The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392.","Jaschhof, M. 2016. A review of world Diallactiini (Diptera, Cecidomyiidae, Winnertziinae), with the description of six new genera and seventeen new species. Zootaxa, 4127 (2), 201 - 244.","Skuse, F. A. A. 1889. Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, (1988) 3 (1 - 2), 7 - 144, 2 - 3 pl.","Karsch, F. 1877. Revision der Gallmucken. E. C. Brunn, Munster, 1 - 58, 1 pl.","Kolesik, P., Gagne, R. J., 2016. P. Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae). Zootaxa, 4205 (4), 301 - 338.","Gagne, R. J. 1978. New synonymy and a review of Haplusia (Diptera: Cecidomyiidae). Proceedings of the Entomological Society of Washington, 80, 517 - 519."]}

TribeHoloneurini Enderlein, 1936 The tribe is characterized by synapomorphies characteristic of the valid tribe rank in Porricondylinae (Mamaev, 2001; Fedotova, Sidorenko, 2007): antennae with 10–11 flagellomeres; reduction of wing venation; absent or reduced parameres; tarsal claws narrow and curved near midlength; male and female flagellomeres with dense sensorial reticulation, and female flagellomeres with a long neck. Their wing venation is variable, with M 3 absent or slightly visible, Cu never forked, CuA 2 slightly visible distally or completely disappearing, m+rm usually slightly curved distally, and rs situated at an angle to R 5. The ovipositor is 2-segmented. The larvae of some extant species of the Holoneurini have been associated with leaf litter and decaying wood (Mamaev, Krivosheina, 1965). Adult insects are usually captured in Malaise traps (Fedotova, Sidorenko, 2005, 2007). The tribe includes 12 genera and 43 species: 9 extant genera with 38 species (number of species in parentheses): Cassidoides Mamaev, 1960 (8); Coccopsis Meijere, 1901 (8); Cognitepidosis Fedotova et Sidorenko, 2005 (1); Cryptoneurus Mamaev, 1964 (3); Dalaiella Mamaev, 1996 (1); Holoneurus Kieffer, 1894 (14), Melicepidosis Fedotova et Sidorenko, 2007 (1); Putepidosis Fedotova et Sidorenko, 2007 (1), Pyxicoccopsis Fedotova et Sidorenko, 2007 (1) and 3 extinct genera with 5 species: Gratomyia (1), Volnococcopsis (1) and Rovnoholoneurus gen. n. (3)., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2017, New Genus And Species Of Gall Midges (Diptera, Cecidomyiidae: Porricondyinae, Holoneurini) From The Late Eocene Amber Of Olevsk (Zhitomir Region, Ukraine), pp. 23-30 in Vestnik Zoologii (Vestn. Zool.) (Vestn. Zool.) 51 (1) on page 24, DOI: 10.1515/vzoo-2017-0004, http://zenodo.org/record/6454042, {"references":["Mamaev, B. M. 2001. The tribe Holoneurini in the Palearctic (Diptera, Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, Pushkino, 17, 1 - 11 [In Russian].","Fedotova, Z. A., Sidorenko, V. S. 2007. New taxa of gall midges (Diptera, Cecidomyiidae, Porricondylinae) with reduced wing venation and flagellomeres from Russia. An International Journal of Dipterological Research, 18 (4), 235 - 277.","Mamaev, B. M., Krivosheina, N. P. 1965. Larvae of gall midges (Diptera, Cecidomyiidae). Comparative morphology, Biology and keys. Nauka, Moskow, 1 - 278 [In Russian].","Fedotova, Z. A., Sidorenko, V. S. 2005. New species of gall midges of the subfamily Porricondylinae from the Russian Far East (Diptera, Cecidomyiidae). An International Journal of Dipterological Research, 16 (2), 89 - 127."]}

Genus Rovnoholoneurus Fedotova et Perkovsky, gen. n. urn:lsid:zoobank.org:act: CDD03952-9506-4EC9-841A-C28C89AF6F56 T y p e s p e c i e s: Rovnoholoneurus davidi Fedotova et Perkovsky, sp. n. D e s c r i p t i o n (figs 1–4). Body slightly shorter than wing. Head elongate; eyes fused, wide bridge positioned on 2/3 part of head, occiput wide, rounded. Head basally with very long setae. Male antenna 2+11-segmented; female antenna 2+10–11-segmented; scape slightly enlarged apically; flagellomeres narrow, basal enlargement with constriction near base or middle, neck prolonged (in male and female), necks of distal flagellomeres much shorter than those of medial ones. Basal enlargement with whorls of long setae of different sizes, especially with long setae in basal, medial whorls. 10th flagellomere sometimes fused, consisting of two segments. Proximal flagellomeres slightly longer than distal ones. Last flagellomere ovoid or conical, without apical projection. All female flagellomeres bearing two rings of sensorial filae with apical, basal round loops. Mouthparts strongly elongated, wide. Palpus 4-segmented, longer than head, with segments slender, almost parallel-sided, 4th segment longest. Scutellum strongly swollen. Wing longer than body, evenly and very strongly widened distally; vein R 5 almost straight, slightly curved near apex, joining margin almost at wing tip; rs strong, short; vein m+rm slightly curved; Cu 2 strongly developed, positioned far from margin of wing, but fork indistinct, not fused, with two poorly visible folds near CuA 2 ; vein CuP well developed. Legs with long setae ventrally. Abdomen slightly swollen near base, narrowed apically. Abdominal segments covered by very long setae. Ovipositor consisting of one pair of dorso-apical plates. C o m p a r i s o n. The new genus appears closely related to Holoneurus Kieffer, 1894 (Mamaev, 2001; Fedotova, Sidorenko, 2007), as evidenced by the body shape, 2+10–11-segmented antennae, long legs, and male and female flagellomere morphology. It differs from Holoneurus in the following characters: wide eye bridge, wing venation, elongated and swollen mouthparts, palpi very thin and much longer than head, different shape of flagellomeres, vein R 5 joining wing margin at apex, m+rm curved, vein rs very short, M 3 absent, CuA 2 well developed and reaching near wing margin, male hind femur longer than tibia, tarsi very long (almost as long as or longer than femur+tibia), long setae on ventral side of legs, and smaller body. S p e c i e s i n c l u d e d. Rovnoholoneurus davidi sp. n., R. miyae sp. n., R. laqueatus (Fedotova, 2005), comb. n. (for Bryocrypta laqueata Fedotova in: Fedotova, Perkovsky, 2005). All species are only known from Rovno amber. E t y m o l o g y. The generic name is formed from a combination of Rovno and the gall midge generic name Holoneurus. Gender masculine., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2017, New Genus And Species Of Gall Midges (Diptera, Cecidomyiidae: Porricondyinae, Holoneurini) From The Late Eocene Amber Of Olevsk (Zhitomir Region, Ukraine), pp. 23-30 in Vestnik Zoologii (Vestn. Zool.) (Vestn. Zool.) 51 (1) on pages 24-25, DOI: 10.1515/vzoo-2017-0004, http://zenodo.org/record/6454042, {"references":["Mamaev, B. M. 2001. The tribe Holoneurini in the Palearctic (Diptera, Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, Pushkino, 17, 1 - 11 [In Russian].","Fedotova, Z. A., Sidorenko, V. S. 2007. New taxa of gall midges (Diptera, Cecidomyiidae, Porricondylinae) with reduced wing venation and flagellomeres from Russia. An International Journal of Dipterological Research, 18 (4), 235 - 277.","Fedotova, Z. A., Perkovsky, E. E. 2005. New gall midges (Diptera, Cecidomyiidae) from the Rovno Amber: Subfamily Porricondylinae (Tribes Bryocryptini and Winnertziini) and subfamily Lasiopterinae (Tribes Brachineurini and Oligotrophini). Paleontologicheskii Zhurnal, 1, 42 - 53 [In Russian, English translation: Paleontological Journal, 39 (1), 41 - 51]."]}

Rovnoholoneurus davidi Fedotova et Perkovsky, sp. n. urn:lsid:zoobank.org:act: 70FAF245-2E6C-4403-81C2-CEB16E6512EF M a t e r i a l. Holotype. SIZK OL-8, a well-preserved male with partly missing tarsi from Olevsk, Rovno amber; Late Eocene. Syninclusions: SIZK OL-7, Carabidae, Psychodidae; SIZK OL-8, Chironomidae, Tipuloidea, Acari; SIZK OL-9, 2 Sciaridae, Psychodidae, Tipuloidea, Chironomidae (Orthocladiinae), Cecidomyiidae, Collembola (Symphypleona), Acari; SIZK OL-10, 2 Trichoptera, 18 Diptera (4 Sciaridae, 11 Chironomidae (Orthocladiinae), 2 Cecidomyiidae, Nematocera indet.). D e s c r i p t i o n. Male (fig. 1, 1 –6, 2, 1–11). Body 1.2 times as long as antenna, wing 1.3 times as long as body. Wing 1.5 times as long as antenna. 1st flagellomere 1.3 times as long as 2nd; proximal flagellomeres with long neck, almost as long as basal enlargement; 2nd flagellomere slightly shorter than 1st; distal flagellomeres with short neck; 5th flagellomere 3.8 times as long as wide; 11th flagellomere conical, 1.6 times as long as 10th. Palpi slightly longer than head, very narrow, parallel sided. Wing 3.5 times as long as wide. Vein R 1+2 is 2.2 times shorter than wing. Profemora and metafemora shorter than corresponding tibia; mesofemur longer than mesotibia. Genitalia transverse; gonostylus short, narrow. M e a s u r e m e n t s (mm): body length, 1.43; antennal length, 1.23; head length with mouth part, 0.31; head length without mouth part, 0.26; head height, 0.21; thorax length, 0.54; thorax width, 0.38; abdomen length, 0.93; wing length, 1.84; wing width, 0.52; vein R 1+2 , 0.85; length of fore leg 1.88: coxa, 0.10; trochanter, 0.07; femur, 0.69; tibia, 0.79; fore tarsi, 1.44; length of middle leg: coxa, 0.11; trochanter, 0.07; femur, 0.61; tibia, 0.57; 1st tarsal segment 0.08; 2nd 0.79; 3rd 0.38; 4th 0.16; 5th 0.11; middle tarsi, 1.52 and leg, 2.88; length of hind leg: coxa, 0.10; trochanter, 0.07; femur, 0.52; tibia, 0.66; 1st tarsal segment, 0.11; 2nd 0.67; 3rd 0.33; 4th 0.15; 5th 0.12; hind tarsi, 1.41 and leg, 3.14. E t y m o l o g y. The species is named after David, the son of the junior author., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2017, New Genus And Species Of Gall Midges (Diptera, Cecidomyiidae: Porricondyinae, Holoneurini) From The Late Eocene Amber Of Olevsk (Zhitomir Region, Ukraine), pp. 23-30 in Vestnik Zoologii (Vestn. Zool.) (Vestn. Zool.) 51 (1) on page 27, DOI: 10.1515/vzoo-2017-0004, http://zenodo.org/record/6454042

Rovnoholoneurus miyae Fedotova et Perkovsky, sp. n. urn:lsid:zoobank.org:act: C9136BCD-1536-4214-B583-C21CCF5F4DA9 M a t e r i a l. Holotype. SIZK OL-2, well preserved inclusion of a female with partly missing tarsi; Olevsk, Rovno amber; Late Eocene. D e s c r i p t i o n. Female (fig. 3, 1 –7, 4, 1–8). Body (with ovipositor) 1.7 times as long as antenna; wing 1.1 times as long as body. Wing 1.8 times as long as antenna. 1st flagellomere 5.0 times as long as wide, 1.2 times as long as 2nd; 5th flagellomere 3.9 times as long as wide, slightly shorter than 1st flagellomere; basal enlargement 2.7 times as long as wide, 0.42 times as long as neck; 10th flagellomere of left antenna 1.2 times as long as 9th, 0.83 times as long as 5th; 10th slightly narrowed to apex, rounded apically; 10th flagellomere of right antenna consisting of two fused segments. Palpi much longer than length or width of head, very narrow, parallel sided; 2nd, 3rd, 4th segments strongly prolonged, their ratio: 1: 2.5: 2.7: 4.3. Wing 2.9 times as long as wide. Vein R 1+2 is 0.48 times as long than wing length. All femora longer than corresponding tibiae. Ovipositor with large round apical plate, directed dorso-caudally, соvered with short setae. M e a s u r e m e n t s (mm): body length, 1.81; antennal length, 1.08; head length with mouth parts, 0.33; head length without mouth parts, 0.20; head width, 0.15; palpus length, 0.38; thorax length, 0.36; thorax width, 0.45; abdomen length, 1.16; wing length, 1.93; wing width, 0.66; vein R 1+2 , 0.88; length of fore leg, 3.53: coxa, 0.12; trochanter, 0.06; femur, 0.85; tibia, 0.82; 1st tarsal segment, 0.07; 2nd, , 0.79; 3rd, 0.44; 4th, 0.28; 5th, 0.10; length of middle: femur, 0.80; tibia; 0.57; 1st tarsal segment, 0.1; length of hind: coxa, 0.1; trochanter, 0.07; femur, 0.92, tibia, 0.47. E t y m o l o g y. The species is named after Mirabel Khelemskyy, daughter of Gennadiy Khelemskyy for his continuous help with the amber collection of the Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine. A key to the species of the genus Rovnoholoneurus gen. n. 1. Male. Wing 3.5 times as long as wide (fig. 1, 1). 5th flagellomere 3.8 times as long as wide (fig. 1, 4). Fore and hind femora slightly shorter than tibia. Hind femora 0.78 times as long as tibia. Body length 1.43 mm............................................................................................................................. B. davidi sp. n. — Female. Wing 2.6 or 2.9 times as long as wide (fig. 3, 1). 5th flagellomere 3.9–4.9 times as long as wide, fore femora longer than tibia................................................................................................................... 2 2. Wing 2.6 times as long as wide. 5th flagellomere 4.9 times as long as wide. Hind femora 1.4 times as long as tibia. Body length 1.74 mm..................................................... B. laqueatus (Fedotova) comb. n. — Wing 2.9 times as long as wide (fig. 3, 1). 5th flagellomere 3.9 times as long as wide (fig. 3, 4). Hind femora almost 2.0 times as long as tibia. Body length 1.81 mm...................................... B. miyae sp. n. We are grateful to A. P. Vlaskin (SIZK) for cutting and polishing the amber pieces, V. Yu. Nazarenko (SIZK) for photography, A. I. Zaitsev (Moscow Sity Pedagogical University) for valuable comments, and S. Bruce Archibald (Simon Fraser University, Burnaby, Canada), and Pyotr N. Petrov (Lomonosov Moscow State University, Moscow, Russia) for editing the English of the manuscript., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2017, New Genus And Species Of Gall Midges (Diptera, Cecidomyiidae: Porricondyinae, Holoneurini) From The Late Eocene Amber Of Olevsk (Zhitomir Region, Ukraine), pp. 23-30 in Vestnik Zoologii (Vestn. Zool.) (Vestn. Zool.) 51 (1) on page 29, DOI: 10.1515/vzoo-2017-0004, http://zenodo.org/record/6454042

Popovineura Fedotova et Perkovsky, gen. n. T y p e s p e c i e s: Popovineura nacta Fedotova et Perkovsky, sp. n. D e s c r i p t i o n. M a l e (fig. 3, 1–8; 8, 1, 2). Body is very wide and swollen, slightly longer than antennae and slightly shorter than wings. Head laterally is almost round. Eyes with wide bridge, occiput wide rounded. Antennae are 2+11-segmented, scape is enlarged distally, pedicel round. All flagellomeres with distinct necks, that are shorter than basal enlargements. Basal enlargement mostly parallelsided, only middle flagellomeres with almost round basal node. Distal flagellomeres with short necks. Eleventh flagellomere ovoid. Proximal flagellomeres are slightly wider and longer than distal ones. Flagellomeres with basal whorl of relatively short setae positioned in single row and medial whorl of very long setae densely covering medial part of basal enlargement. Sensorial filae of flagellomeres are invisible. Mouthparts are short and wide. Palpi two-segmented, second segment longer than first. Wing is widely enlarged and rounded distally, vein R 4+5 slightly curved, located near C, enters wing margin before its apex, R s absent, Cu located far from wing margin, not forming fork. Hind femur almost as long as middle, hind tibia is much longer than the middle one. Hind femur is shorter than tibia. Tarsal claw simple, hook-form. Abdomen is evenly swollen and wide rounded distally, strongly narrowed before genitalia. Genitalia are transversal. Gonocoxite is narrow, without basal lobe or enlargment, slightly curved distally. Gonostylus is narrow, with apical dent, slightly shorter than gonocoxites. Cerci are wide, cordiform, almost reaching apex of gonocoxites, with wide round lobe dissected by triangular excision. Parameres or basal enlargement of gonocoxites absent. Hypoproct is whole, rounded apically. Aedeagus is very narrow, almost parallelsided, much longer than gonocoxites. Рис. 3. Popovineura nacta, самец, голотип (инв. номер K–666 коллекции Института зоологии НАН Украины): 1 — общий вид; 2 — скапус, педицелл, 1-й и 2-й членики жгутика; 3 — 5–6-й членики жгутика; 4 — 8–11-й членики жгутика; 5 — щупик; 6 — передняя лапка; 7 — 5-й членик передней лапки; 8 — гениталии. Масштабные линейки 0.1 mm. C o m p a r i s o n. By the shape of the body, arrangement of wing veins, the new genus is closely related to Brachineura Rondani, 1840 (Mamaev, 1967; Gagné, Solinas, 1996), differing from it by the 2+11-segmented antennae, presence of much longer necks of flagellomeres not covered by dense scales, long legs and very narrow gonocoxites and gonostylus, very long aedeagus, short mouthparts, 2-segmented palpi, and smaller body size. E t y m o l o g y. The genus is named in honour of paleoentomologist Yu. A. Popov, the leading Russian expert in fossil Hemiptera. The name is combined of his second name and the second part of the name Brachineura. Gender feminine., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on pages 40-41, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Rondani, C. Sopra alcuni nuovi generi di insetti ditteri // Memoria seconda per servire alla ditterologia italiana. - Parma, 1840. - P. 1 - 27, Pl. 1.","Mamaev, B. M. Gall-midges of the USSR. 7. New species of non-gall-making gall midges of the tribe Oligotrophini (Diptera, Cecidomyiidae) // Entomologicheskoe Obozrenie. - 1967. - 46. - P. 873 - 883. - Russian: Мaмaiv B. М. Gaллitы СССР. 7. Нovыi vidы сvobodнozivuщik gaллit iz tрibы Oligotrophini (Diptera, Cecidomyiidae).","Gagne, R. J., Solinas, M. The Rondani Cecidomyiidae (Diptera) // Insecta Mundi. - 1996. - 10, N 1 - 4. - P. 69 - 75."]}

Rhizomyia Kieffer, 1898 Rhizomyia Kieffer, 1898: 56. T y p e s p e c i e s: Rhizomyia perplexa Kieffer, 1898: 57 (monotypy). D i a g n o s i s (fig. 5, 1–8, 7, 6, 7). Wing, body and legs are conspicuously covered by scales. Head almost completely covered by eyes or eye bridge wide. Antennae are 2+9–10-segmented. Male flagellomeres are subcylindrical with neck and swollen basal enlargement with numerous swollen peritremae of setae. Neck of male middle flagellomeres is longer, equal or slightly shorter than basal enlargement. Female flagellomeres are elongated, sessile, subcylindrical, without neck or with very short neck. Male fore tarsi usually with teeth, middle and hind tarsi with or without teeth. Palpi are 3–4-segmented, labrum is triangular. Wing is broad, R1+2 joins costa before its midlength. R4+5 is straight, joining C rather near wing apex or at wing apex; C broken at junction with R4+5; Cu forked. Claws with tooth. Genitalia are transversal. Gonocoxites are short, ovoid or parallel-sided. Gonostylus is long, narrowing apically, terminated by very little apical claw. Cerci with very deep excision. Hypoproct is slightly caved or excavated. Aedeagus is usually shorter than gonocoxites or equal in length. Basal outgrowths (or parameres) of gonocoxites absent. Ovipositor very short, not protrusible, apically with two terminal lamellae. S p e c i e s i n c l u d e d. Holarctic and Oriental genus with 29 species, 21 of them Palaearctic ones (Skuhravá, 1997; Mamaev, 1998; Mamaev, Zaitzev, 2002; Fedotova, 2004; Gagné, 2004, 2010; Fedotova, Sidorenko, 2005, 2006 b). Extinct species of Rhizomyia were unknown until now. B i o l o g y. Larvae of Rhizomyia perplexa Kieffer, 1898 and R. circumspinosa (Rübsaamen, 1899) develop on roots and in sheath of leaves of a sedge (Carex spp., Cyperaceae) (Mamaev, Krivosheina, 1965)., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on page 44, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Kieffer, J. J. Synopse des cecidomyies d'Europe et d'Algerie decrites jusqu'a ce jour // Bulletin de la Societe d'Histoire Naturelle de Metz. - 1898. - 2, N 8. - P. 1 - 64.","Skuhrava, M. Family Cecidomyiidae // Contributions to a Manual of Palaearctic Diptera (with Special Reference to Flies of Economic Importance): Nematocera and Lower Brachycera / Eds L. Papp, B. Darvas. - Budapest: Science Herald, 1997. - Vol. 2. - P. 71 - 204.","Mamaev, B. M., Zaitzev, A. I. Criteria and indicators for the separate position of gall midges belonging to the genus Rhizomyia (Diptera, Cecidomyiidae) // All-Russian Institute of Continuous Education in Forestry. - Pushkino, 2002. - 22. - P. 1 - 12. - Russian: Мaмaiv B. М., Zaйtiv A. I. Kрitiрii i iнdikatoры сaмoсtoitiльнoсti vidov gaллit рoda Rhizomyia (Diptera, Cecidomyiidae).","Gagne, R. J. A catalog of the Cecidomyiidae (Diptera) of the world // Memoirs of the Entomological Society of Washington. - 2004. - 25. - P. 1 - 408.","Gagne, R. J. Update for A catalog of the Cecidomyiidae (Diptera) of the world. - 2010. - http: // www. ars. usda. gov / SP 2 UserFiles / Place / 12754100 / Gagne _ 2010 _ World _ Catalog _ Cecidomyiidae. pdf","Mamaev, B. M., Krivosheina, N. P. Larvae of gall midges (Diptera, Cecidomyiidae). Comparative morphology, Biology and keys. - Mosсow: Nauka, 1965. - 278 p. - Russian: Мaмaiv B. М., Kрivosiiнa Н. P. Лiciнki gaллit (Diptera, Cecidomyiidae). Cрavнitiльнai мoрфoлogii, bioлogii."]}

Rovnobrachineura Fedotova et Perkovsky, gen. n. T y p e s p e c i e s: Rovnobrachineura kiryeyevi Fedotova et Perkovsky, sp. n. D e s c r i p t i o n (fig. 1, 1–6; 7, 1–3). Male. Body slightly longer than wings. Head is elongate; eyes fused, with narrow bridge, positioned on anterior part of head, occiput wide rounded. Antennae 2+10-segmented, scape narrower than round pedicel. All flagellomeres with distinct very narrow necks, but basal enlargements have different forms. Basal enlargement consists of basal node and narrower slightly swollen or parallelsided distal part: 1st and 2nd flagellomeres slightly narrowed medially; 3rd-7th with round basal node, wide distal part and little neck; 8th-9th with round basal node and narrow distal part; 10th flagellomere strongly narrowed between round basal node and wide oval distal part. Proximal flagellomeres much wider and shorter than distal ones. Flagellomeres with basal and medial whorls of very long setae. Distal part of flagellomeres bears two rings of sensorial filae. Mouthparts much elongate and wide. Palpi 2-segmented, the second one is longer. Wing evenly and very strongly widened distally, vein R4+5 straight, joining margin almost at wing tip, Rs absent, Cu positioned near margin of wing, not forming a fork. Fore and middle femora and tibiae shorter than hind femur and tibia. Length of hind femur and tibia almost equal. Abdomen slightly swollen near middle (2nd-4th segments) and narrowed distally. Genitalia transversal. Gonocoxite wide, almost straight on outside margin, with dense long setae and almost rectangular lobe on inner side. Gonostylus narrow, slightly curved basally. Cerci very short, far not reaching the apex of gonocoxites, with wide round lobes and triangular excision between its. Parameres or basal enlargement of gonocoxites absent. Hypoproct whole, rounded apically. Aedeagus very wide, much longer than gonocoxites. C o m p a r i s o n. By the shape of the body, 2+10-segmented antennae, wing venation, long legs and forms of gonocoxites and gonostylus the new genus is closely related to Brachyneurina Mamaev, 1967 (Mamaev, 1967; Harris, Evans, 1979), but differs from it by the elongated and swollen mouthparts, very narrow eye bridge, 2-segmented palpi, different shape of flagellomeres, vein R 4+5 joining with wing margin at the tip of the wing, very long aedeagus, presence of very dense long setae on the inner side of gonocoxites, and by smaller body size. E t y m o l o g y. The name is combined from the name of the amber Lagerstätte and name Brachineura. Gender feminine., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on page 37, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Mamaev, B. M. Gall-midges of the USSR. 7. New species of non-gall-making gall midges of the tribe Oligotrophini (Diptera, Cecidomyiidae) // Entomologicheskoe Obozrenie. - 1967. - 46. - P. 873 - 883. - Russian: Мaмaiv B. М. Gaллitы СССР. 7. Нovыi vidы сvobodнozivuщik gaллit iz tрibы Oligotrophini (Diptera, Cecidomyiidae).","Harris, K. M., Evans, R. E. Gall development in the fungus Peniophora cinerea (Fr.) Cooke induced by Brachyneurina peniophorae sp. n. (Diptera: Cecidomyiidae) // Entomologist's Gazette. - 1979. - 30. - P. 23 - 30."]}

Ledomyia Kieffer, 1895 Lepidomyia Kieffer, 1894: 201, preoccupied by Loew, 1864. T y p e s p e c i e s: L epidomyia lugens Kieffer, 1894: 211 (monotypy). Ledomyia Kieffer, 1895: cccxx, new name for Lepidomyia Kieffer. D i a g n o s i s (fig. 6, 1–7; 8, 3, 4). Wing, body and legs are conspicuously covered by scales. Antennae are 2+8–12-segmented. Male flagellomeres subcylindrical with neck and swollen basal enlargement. Female flagellomeres elongated, sessile, without neck. Tarsal claws toothed on the forelegs, toothed or simple on the mid- and hindlegs. Palpi are mostly 4-segmented, labrum triangular. Wing is broad, R 1+2 joins C before its midlength. R 4+5 is straight or curved, joins C rather far before wing apex; C broken at junction with R 4+5; Cu is forked. Genitalia are transversal. Gonocoxites are ovoid or parallel-sided. Gonostylus is short, excavated ventrally, terminated by very wide apical or dorso-apical claw, protruding from dorsal part of gonostylus. Cerci are cordiform with very deep excision. Hypoproct slightly caved. Aedeagus usually equal or longer than gonocoxites. Basal processes of gonocoxites are well developed, strongly narrowed apically. Ovipositor very long, extensible, apically with two terminal lamellae with two small, obtuse spines. Рис. 6. Ledomyia olgae, самец, голотип (инв. номер K–3697 коллекции Института зоологии НАН Украины): 1 — общий вид; 2, 3 — щупик (изменчивость формы); 4 — 9–11-й членики жгутика, 5 — 5-й членик жгутика, 6, 7 — гениталии (6 — снизу, 7 — сверху). Масштабные линейки 0.1 mm. Fig. 7. Rovnobrachineura kiryeyevi, самец, голотип (инв. номер K–3701 коллекции Института зоологии НАН Украины) (1–3); Brachineura polessica, самка, голотип (инв. номер K–1236) (4, 5); Rhizomyia parkalovi, самка, голотип (инв. номер K–7385) (6, 7): 1, 4, 6 — голова и антенна (1 — ×129; 4 — ×62; 6 — ×117); 2 — гениталии (×283); 3, 5, 7 — общий вид (3 — ×46; 5 — ×29; 7 — ×39). Fig. 8. Popovineura nacta, самец, голотип (инв. номер K–666 коллекции Института зоологии НАН Украины) (1, 2); Ledomyia olgae, самец, голотип (инв. номер K–3697) (3, 4); Nascensluna mellea, самка, голотип (инв. номер K–1601) (5–7): 1–3, 7 — общий вид (1 — ×45; 2 — ×38; 3 — ×48; 7 — ×64); 4 — антенна (4 — ×129); 5 — задняя нога (×82); 6 — голова и антеннa (×131). Species included. The genus is nearly cosmopolitan, two species were described from tropical regions (Fedotova, Sidorenko, 2007). Genus includes 26 species, 17 of them are Palaearctic (Mamaev, 1967; Skuhravá, 1997; Gagné, 2004, 2010; Fedotova, Sidorenko, 2007; Fedotova, Perkovsky, 2008). Some undescribed species are known from Middle Miocene Mexican amber (Gagné, 1973) and Late Eocene Baltic amber, part of species were mistakenly included in extinct genus Ledomyiella Meunier, 1904, known from Baltic amber. From Late Eocene Rovno amber was described Ledomyia dextra Fedotova et Perkovsky, 2008. Most of the included species have been reared from fresh-cut logs (Gagné, 1985, 2010; Rock, Jackson 1985). Ledomyia collarata Gagné, 1987 was reared from galls on a fungus Xylaria enterogena Mont. (Xylariaceae, Ascomycetes) (Larew et al., 1987); its adults are unknown and the species is only tentatively placed in this genus., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on pages 45-48, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Kieffer, J. J. Changement de nom // Bulletin de la Societe Entomologique de France. - 1895. - P. CCCXX.","Kieffer, J. J. Ueber die Heteropezinae // Wiener Entomologische Zeitung. - 1894. - 13. - S. 200 - 212, Pl. I.","Fedotova, Z. A., Sidorenko, V. S. Nine new species of genus Ledomyia Kieffer, 1895 (Diptera, Cecidomyiidae) from the Russian Far East // Far Eastern Entomologist. - 2007. - 169. - P. 1 - 19.","Mamaev, B. M. Gall-midges of the USSR. 7. New species of non-gall-making gall midges of the tribe Oligotrophini (Diptera, Cecidomyiidae) // Entomologicheskoe Obozrenie. - 1967. - 46. - P. 873 - 883. - Russian: Мaмaiv B. М. Gaллitы СССР. 7. Нovыi vidы сvobodнozivuщik gaллit iz tрibы Oligotrophini (Diptera, Cecidomyiidae).","Skuhrava, M. Family Cecidomyiidae // Contributions to a Manual of Palaearctic Diptera (with Special Reference to Flies of Economic Importance): Nematocera and Lower Brachycera / Eds L. Papp, B. Darvas. - Budapest: Science Herald, 1997. - Vol. 2. - P. 71 - 204.","Gagne, R. J. A catalog of the Cecidomyiidae (Diptera) of the world // Memoirs of the Entomological Society of Washington. - 2004. - 25. - P. 1 - 408.","Gagne, R. J. Update for A catalog of the Cecidomyiidae (Diptera) of the world. - 2010. - http: // www. ars. usda. gov / SP 2 UserFiles / Place / 12754100 / Gagne _ 2010 _ World _ Catalog _ Cecidomyiidae. pdf","Gagne, R. J. Cecidomyiidae from Mexican Tertiary amber (Diptera) // Proceedings of the Entomological Society of Washington. - 1973. - 75. - P. 169 - 171.","Meunier, F. Monographie des Cecidomyidae, des Sciaridae, des Mycetophilidae et des Chironomidae de l'ambre de la Baltique // Annales de la Societe Scientifique de Bruxelles. - 1904. - 28. - P. 12 - 275, pls. I - XVI.","Gagne, R. J. Descriptions of new Nearctic Cecidomyiidae (Diptera) that live in xylem vessels of fresh-cut wood, and a review of Ledomyia (s. str.) // Proceedings of the Entomological Society of Washington. - 1985. - 87. - P. 116 - 134.","Rock, E. A., Jackson D. The biology of xylophilic Cecidomyiidae (Diptera) // Proceedings of the Entomological Society of Washington. - 1985. - 87. - 135 - 141.","Larew, H. G, Gagne, R. J., Rossman, A. Y. Fungal gall caused by a new species of Ledomyia (Diptera: Cecidomyiidae) on Xylaria enterogena (Ascomycetes: Xylariaceae) // Annals of the Entomological Society of America. - 1987. - 80. - P. 502 - 507."]}

Rhizomyia parkalovi Fedotova et Perkovsky, sp. n. M a t e r i a l. Holotype. SIZK N K–7385, well preserved inclusion of female with tarsi missing, Klesov, Rovno amber, Late Eocene (fig. 5, 1–8, 7, 6, 7). Syninclusions: N K–7383: Collembola Entomobryomorpha, 2 ex.; Cecidomyiidae: Rhizomyia sp., K–7384: Sciaridae. D e s c r i p t i o n. F e m a l e (fig. 5, 1–8, 7, 6, 7). Body (without genitalia) is 1.8 times longer than antennae. Wings 2.2 times as long as antennae. Length of head 2.2 times as long as lateral side of head. Antennae are 2+10-segmented. Head completely covered by eyes laterally, eye bridge narrow. Scape enlarged distally, almost as long as round pedicel. Necks of flagellomeres very short. First flagellomere slightly longer than 2nd; 5th flagellomere 2.0 times as long as wide. Ninth flagellomere without neck; 10th flagellomere 2.9 times as long as wide; narrowed distally. Palpi are very narrow, 2-segmented, 2nd segment is 1.8 times longer than the first. Fore and middle femur have equal length, shorter than tibia ones. Lengths of fore and middle tibia also almost equal. First tarsomere with lateral long projection is covered by setae. Wing is 2.3 times as long as wide. Vein R4+5 strongly curved near apex, joining wing margin at smaller distance from tip of wing than Cu2. All tibiae subequal to, or longer than, respective femora. Fore tibia 1.1 times as long as femur. Ovipositor is not protrusible. 8th abdominal segment without ventral lobes or projections. M e a s u r e m e n t s, mm: body length, 1.05; wing length, 1.27; wing width, 0.65; antennal length, 0.59; head length, 0.24 (without mouth parts); width of lateral side of head, 0.11; thorax length, 0.34; abdomen length, 0.61; length of fore: coxa, 0.10; trochanter, 0.07; femur 0.45; tibia 0.48; 1st segment, 0.07; length of middle: coxa, 0.11; trochanter, 0.07; femur 0.43; tibia 0.45; length of hind: coxa, 0.14; trochanter, 0.06; femur, 0.45; tibia, 0.47; 1st segment, 0.07. C o m p a r i s o n. By the 2+10-segmented antennae, arrangement of wing veins, long flagellomeres new species is closely related to R. operculata Fedotova et Sidorenko, 2006, differing from it by the elongated segments of 2-segmented palpi; very short neck and wide basal enlargement of middle flagellomeres; narrowing distally 10th flagellomere; in wing, that is enlarged distally (not proximally); in R 4+5, that joining C before wing apex, and smaller body size. E t y m o l o g y. The species is named after Andrej Parkalov who permanently helps the amber collection of Schmalhausen Institute of Zoology, NAS of Ukraine., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on pages 44-45, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791

Nascensluna Fedotova et Perkovsky, gen. n. T y p e s p e c i e s: Nascensluna mellea Fedotova et Perkovsky, sp. n. D e s c r i p t i o n. Male (fig. 4, 1–4; 8, 5–7). Body very wide and swollen, much longer than antennae and slightly shorter than wings. Body and antennae densely covered by scales. Head laterally almost oval. Eye bridge wide, occiput wide rounded. Antennae are 2+10-segmented, scape is enlarged distally, pedicel is round. All flagellomeres without neck, and slightly rounded laterally. Flagellomeres evenly narrowed and shortened distally. Tenth flagellomere widely rounded apically. Flagellomeres with basal whorl of short setae located in single row and medial whorl of very long setae rarely covering medial part of flagellomere. Sensorial filae of flagellomere are invisible. Wing is wide enlarged and rounded distally, vein R 4+5 is short and straight, almost fused with R 1+2 and C, joins margin of the wing near its middle, Cu is positioned far from margin of wing, without fork. Fore femur and tibia are slightly longer than middle ones and shorter than hind femur and tibia. Hind femur and tibia have equal length. Hind tarsus is 1.6 times as long as femur. Tarsal claw is simple, hook-form. Hind leg is 1.7 times as long as body and 2.1 times as long as wing. Thorax and abdomen are strongly swollen and wide rounded distally. Ovipositor is not protrusible. Cerci are bilobed, very short and narrow. Рис. 4. Nascensluna mellea, самка, голотип (инв. номер K–1601 коллекции Института зоологии НАН Украины): 1 — общий вид; 2 — 8–10-й членики жгутика; 3 — 5-й членик жгутика; 4 — яйцеклад. Масштабные линейки 0.1 mm. C o m p a r i s o n. By the shape of the swollen body, 2+10-segmented antennae, covered by dense scales, the new genus is closely related to Brachineura Rondani, 1840 (Mamaev, 1967; Gagné, Solinas, 1996), differing from it by the wing venation (fused veins R 1+2 and R 4+5), much shorter flagellomeres, longer legs, narrow and short cerci of ovipositor and smaller body size. Wings of new genus are similar to Lasioptera Meigen, 1818, but new genus differs by short antennae and not protrusible ovipositor. E t y m o l o g y. Name is based on the Latin Nascens luna (new moon or moonrise). Gender feminine., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on pages 42-43, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Rondani, C. Sopra alcuni nuovi generi di insetti ditteri // Memoria seconda per servire alla ditterologia italiana. - Parma, 1840. - P. 1 - 27, Pl. 1.","Mamaev, B. M. Gall-midges of the USSR. 7. New species of non-gall-making gall midges of the tribe Oligotrophini (Diptera, Cecidomyiidae) // Entomologicheskoe Obozrenie. - 1967. - 46. - P. 873 - 883. - Russian: Мaмaiv B. М. Gaллitы СССР. 7. Нovыi vidы сvobodнozivuщik gaллit iz tрibы Oligotrophini (Diptera, Cecidomyiidae).","Gagne, R. J., Solinas, M. The Rondani Cecidomyiidae (Diptera) // Insecta Mundi. - 1996. - 10, N 1 - 4. - P. 69 - 75."]}

Fedotova, Z. A., Perkovsky, E. E. (2014): New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber. Vestnik Zoologii 48 (1): 35-50, DOI: 10.2478/vzoo-2014-0004, URL: http://dx.doi.org/10.2478/vzoo-2014-0004

Brachineura Rondani, 1840 T y p e s p e c i e s: Brachineura fuscogrisea Rondani, 1840: 17 (monotypy). D i a g n o s i s (fig. 2, 1 –11; 7, 4, 5). Body and appendages are covered by very dense dark scales (flagellomeres and tarsi particularly so). Antennae 2+10-segmented: male flagellomeres subcylindrical with short neck; female ones cylindrical, rarely with swollen peritremae. Length of flagellomeres increases toward apex. Claws with denticle, empodium is developed. Palpi 1–3-segmented. Vein R 1+2 joins C before middle of wing, R 4+5 straight or curved, join C very far before wing apex; Cu simple, almost invisible distally. Cerci of male genitalia often cordiform, with very deep triangular excision. Hypoproct is almost as long as cerci and gonocoxites, deep excavated, with oval or semiround excision. Basal outgrouths of gonocoxites undeveloped. Ovipositor is not protrusible, with two dorsal lobes and one ventral. Рис. 2. Brachineura polessica, самка, голотип (инв. номер K–1236 коллекции Института зоологии НАН Украины): 1 — общий вид; 2 — скапус, педицелл, 1-й и 2-й членики жгутика; 3 — 9–10-й членики жгутика; 4 — 6–7-й членики жгутика; 5 — 3–5-й членики жгутика; 6 — щупик; 7 — 3–5-й членики средней лапки; 8 — 1–2-й членики средней лапки; 9 — 1-й членик средней лапки; 10 — 5-й членик средней лапки; 11 — яйцеклад сбоку. Масштабные линейки 0.1 mm. S p e c i e s i n c l u d e d. Cosmopolitan genus, includes 3 subgenera and 23 species, 19 of them Palaearctic ones (Mamaev, 1967, 1994; Fedotova, 2004; Fedotova, Sidorenko, 2006 a; Gagné, 2004, 2010), 13 species are recorded from Russian Far East, 2 subgenera and 12 species were recently described (Fedotova, 2004; Fedotova, Sidorenko, 2006 a). Rovno amber species is the first extinct representative of the genus. B i o l o g y. Larvae are associated with fungi and decaying leaves, small knots and bark fragments, other vegetable matter and fungi (Mamaev, Krivosheina, 1965; Gagné, Solinas, 1996). Known, that Brachineura fungicola (Mamaev, 1967) were reared from mycelium on thin alder snags and dead fruit bodies of Polyporus sulphureus (Mamaev, Krivosheina, 1965). Brachineura squamigera (Winnerts, 1853) was reared from mushrooms belonging to genera Russula and Rhizopogon (Bayram, Skuhravá, 2004)., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on pages 38-39, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Rondani, C. Sopra alcuni nuovi generi di insetti ditteri // Memoria seconda per servire alla ditterologia italiana. - Parma, 1840. - P. 1 - 27, Pl. 1.","Mamaev, B. M. Gall-midges of the USSR. 7. New species of non-gall-making gall midges of the tribe Oligotrophini (Diptera, Cecidomyiidae) // Entomologicheskoe Obozrenie. - 1967. - 46. - P. 873 - 883. - Russian: Мaмaiv B. М. Gaллitы СССР. 7. Нovыi vidы сvobodнozivuщik gaллit iz tрibы Oligotrophini (Diptera, Cecidomyiidae).","Mamaev, B. M. A contribution to the gall midge fauna (Diptera, Cecidomyiidae) of Kamchatka with descriptions of new species // Vestnik Zoologii. - 1994. - N 2. - P. 28 - 32. - Russian: Мaмaiv B. М. Мatiрiaлы po фauнi gaллit Kaмcatki (Diptera, Cecidomyiidae) с opiсaнiiм нovыk vidov.","Fedotova, Z. A., Sidorenko, V. S. New species of gall midges from the genus Brachineura Rondani, 1840 and new related genera (Diptera: Cecidomyiidae, Brachineurini) from the Russian Far East // International Journal of Dipterological Research. - 2006 a. - 17. - P. 77 - 97.","Gagne, R. J. A catalog of the Cecidomyiidae (Diptera) of the world // Memoirs of the Entomological Society of Washington. - 2004. - 25. - P. 1 - 408.","Gagne, R. J. Update for A catalog of the Cecidomyiidae (Diptera) of the world. - 2010. - http: // www. ars. usda. gov / SP 2 UserFiles / Place / 12754100 / Gagne _ 2010 _ World _ Catalog _ Cecidomyiidae. pdf","Mamaev, B. M., Krivosheina, N. P. Larvae of gall midges (Diptera, Cecidomyiidae). Comparative morphology, Biology and keys. - Mosсow: Nauka, 1965. - 278 p. - Russian: Мaмaiv B. М., Kрivosiiнa Н. P. Лiciнki gaллit (Diptera, Cecidomyiidae). Cрavнitiльнai мoрфoлogii, bioлogii.","Gagne, R. J., Solinas, M. The Rondani Cecidomyiidae (Diptera) // Insecta Mundi. - 1996. - 10, N 1 - 4. - P. 69 - 75.","Bayram, S., Skuhrava, M. New records of Cecidomyiidae (Diptera) from Turkey // J. Entomol. Res. Soc. - 2004. - 6, N 3. - P. 9 - 13."]}

Rovnobrachineura kiryeyevi Fedotova et Perkovsky, sp. n. M a t e r i a l. Holotype. SIZK N K–3701, well preserved inclusion of male with tarsi missing, Klesov, Rovno amber, Late Eocene (fig. 1, 1–6; 7, 1–3). Syninclusions: Chironomidae; Cecidomyiidae: Porricondyla sp., 1 °. D e s c r i p t i o n. M a l e (fig. 1, 1–6; 7, 1–3). Body (without genitalia) is 1.6 times longer than antennae. Wings 1.4 times as long as antennae. Head with narrow eyes, that laterally cover only 2/3 of width of head. Scape enlarged distally, shorter than pedicel. First flagellomere 2.2 times as long as wide; 2nd flagellomere strongly narrower than 1st and slightly shorter than one. Sixth flagellomere 2.2 times as long as wide, slightly shorter than 1st flagellomere; 9th and 10th flagellomeres are longer than middle ones, enlarged distally, rounded apically. The palpi are very narrow, slightly rounded or parallelsided laterally; 2nd palpomere is 1.3–1.4 times as long as 1st. Wing is 2.7 times as long as wide. Vein R 1+2 is less than 2 times shorter than wing length. Gonocoxites are slightly longer than gonostylus. Gonostylus is almost parallelsided distally, 1.2 times shorter than gonocoxites. Cerci are not reaching before middle of gonocoxites. Aedeagus is conical. Рис. 1. Rovnobrachineurina kiryeyevi, самец, голотип (инв. номер K–3701 коллекции Института зоологии НАН Украины): 1 — общий вид; 2 — скапус, педицелл, 1-й и 2-й членики жгутика; 3 — 8–10-й членики жгутика; 4 — 6–7-й членики жгутика; 5 — ротовые органы и щупики; 6 — гениталии сверху. Масштабные линейки 0.1 mm. M e a s u r e m e n t s, mm: body length, 0.78; antennal length, 0.49; head length, 0.19; width of lateral side of head, 0.11; palpus length, 0.07; thorax length, 0.23; wing length, 0.71; wing width 0.26; halter length, 0.12; length of fore: femur, 0.22; tibia, 0.27; metatarsus 0.04; length of middle: femur, 0.22; tibia; 0.21; length of hind: femur 0.25; tibia 0.24; abdomen length, 0.42; genitalia length, 0.06. E t y m o l o g y. The species is named after Denys Kiryeyev for the help to the amber collection of Schmalhausen Institute of Zoology, NAS of Ukraine., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on pages 37-38, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791

Ledomyia olgae Fedotova et Perkovsky, sp. n. M a t e r i a l. Holotype. SIZK N K–3697, well preserved inclusion of male, Klesov, Rovno amber, Late Eocene (fig. 6, 1–7; 8, 3, 4). Syninclusion: Diptera. D e s c r i p t i o n. M a l e (fig. 6, 1–7; 8, 3, 4). Body (without genitalia) is 1.3 times longer than antennae. Wings are 1.3 times as long as antennae. Antennae are 2+12-segmented. Head with large eyes that laterally cover only dorsal part of head. Scape is enlarged distally, almost as long as wide round pedicel. First flagellomere slightly longer than 2nd, its neck is shorter than the neck of the 2nd one; 5th flagellomere is 1.9 times as long as wide, its neck is 3.0 times shorter than basal enlargement. Tenth and 11th flagellomeres have very short neck; 10th flagellomere is 1.5 times as long as wide; 11th flagellomere is slightly narrowed distally and shorter than 10th. 12th flagellomere is oviform. Palpi are very narrow, 2–3-segmented, slightly swollen laterally. Fore tarsus with 2nd tarsomere 5 times as long as 1st. Wing is 2.2 times as long as wide. Vein R 1+2 joins C before its midlength; R 4+5 is strongly curved near apex, joins the wing margin closer to the apex than Cu 2. All tibiae subequal to or longer than respective femora. Fore tibia 1.2 times as long as femur. Hind leg is longer than fore and middle ones. Gonocoxites are slightly rounded laterally, 2.6 times as long as wide. Gonostylus is slightly curved, almost parallelsided, with wide apical claw, 1.7 times shorter than gonocoxites. Cerci are wider than gonocoxites. Aedeagus almost triangular. M e a s u r e m e n t s, mm: body length, 0.76 (without genitalia); 0.83 (with genitalia); wing length, 0.77; wing width, 0.35; antennal length, 0.59; head length, 0.20 (without mouth parts); 0.25 (with mouth parts); width of lateral side of head, 0.11; palpi length, 0.12; thorax length, 0.28; abdomen length, 0.41; length of fore leg, 1.38; coxa and trochanter, 0.09; tarsus, 0.61; femur 0.32; tibia 0.36; 1st segment, 0.06, 2nd, 0.30; 3rd, 0.06; 4th, 0.09; 5th, 0.07; length of middle: leg, 1.31; femur 0.33; tibia 0.37; tarsus, 0.61; coxa and trochanter, 0.09; 1st segment, 0.06, 2nd, 0.28; 3rd, 0.12; 4th, 0.09; 5th, 0.06; length of hind: leg, 1.46; coxa and trochanter, 0.09; tarsus, 0.66; femur, 0.34; tibia, 0.37; 1st segment, 0.06, 2nd, 0.32; 3rd, 0.11; 4th, 0.10; 5th, 0.07. C o m p a r i s o n. By the shape of the body, wing venation, long legs and form of flagellomeres the new species is closely related to Ledomyia dextra Fedotova et Perkovsky, 2008 from Dubrovitsa, differing from it by the 2+12-segmented antennae (vs. 2+11 of L. dextra); wing ratio (2.2 vs. 2.6); very long 2nd tarsomere; wider and shorter middle flagellomeres with wide and short necks; all tibiae longer than respective femora (hind tibia of L. dextra shorter than femur); oviform apical flagellomere (vs. oval of L. dextra); larger body size. E t y m o l o g y. The species is named after Olga — daughter of E. E. Perkovsky. All in all, in Rovno amber Brachineurini and Ledomyiini account for 11.9 % of all species of the superfamily Cecidomyioidea (27 vs. 227), in Baltic amber — for 9.0 % (6 vs. 67) (Fedotova, Perkovsky, 2009)., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on page 48, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791, {"references":["Fedotova, Z. A., Perkovsky, E. E. New gall midges of the tribe Leptosynini (Diptera, Cecidomyiidae) from the Late Eocene Ambers and the classification of the supertribe Heteropezidi // Paleontol. J. - 2009. - 43, N 9. - P. 1101 - 1179."]}

Brachineura polessica Fedotova et Perkovsky, sp. n. M a t e r i a l. Holotype. SIZK N K–1236, poorly preserved inclusion of female with deformed wing and partly missing tarsi, Klesov, Rovno amber, Late Eocene (fig. 2, 1 –11; 7, 4, 5). Syninclusions: K–1233: Phoridae; K–1234: Dolichopodidae; K–1235: Dolichopodidae; K–1236: spider web; K–1237: Collembola Entomobryomorpha. D e s c r i p t i o n. F e m a l e (fig. 2, 1 –11; 7, 4, 5). Body is very wide, as long as wings, 1.7 times longer than antennae. Wings are 1.7 times as long as antennae. Head with wide eye bridge. Head length 1.7 times as long as width. Scape is enlarged distally, longer than transversal pedicel. First flagellomere is 1.8 times as long as wide; 2nd flagellomere is as long as 1st one, with very short neck. Middle flagellomeres are almost parallelsided, 5th flagellomere is 2.6 times as long as wide, almost as long as 1st flagellomere one. Lengths of 9th and 10th flagellomeres are equal, 9th is narrowed distally, apically rounded. Flagellomeres with basal and medial whorls of very long setae. Basal enlargement with rings of sensorial filae, sometimes with little wide loops around lateral sides of flagellomeres. Palpi are 2-segmented, very narrow, almost parallelsided laterally, 2nd 1.4 times as long as 1st. Wing is deformed. Vein Cu is simple, without fork, curved. Fore and middle femur and tibia are shorter than hind femur and tibia. All legs with femur that longer than tibia or have almost equal length with it. First tarsal segment with narrow apical projection laterally (fig. 2, 9). Second middle tarsomere 4.8 times as long as 1st. Tarsal claw is simple, hook-form. Ovipositor is directed dorsocaudally, apical lobes are wide-oval, covered by long setae. M e a s u r e m e n t s, mm: body length, 1.54; antennal length, 0.91; head length, 0.36; width of lateral side of head, 0.21; palpus length, 0.14;1st segment, 0.06; 2nd segment, 0.08 mm; wing length, 1.54; wing width 0.44 (unclear); length of fore: femur, 0.50; tibia, 0.52; 1st tarsal segment 0.10; length of middle: coxa, 0.18; trochanter, 0.06; femur, 0.55; tibia, 0.53; tarsus, 1.06: 1st segment, 0.10, 2nd, 0.48; 3rd, 0.19; 4th, 0.15; 5th, 0.13; length of hind: femur, 0.55; tibia, 0.58; metatarsus, 0.10. C o m p a r i s o n. By the shape and size of the body, 2+10-segmented antennae, simple vein Cu, short legs and form of large round plates of ovipositor the new species is closely related to Brachineura (s. str.) enigmatosa Fedotova et Sidorenko, 2006, differing from it by shorter 2-segmented palpi; short and wide 1st and 2nd flagellomeres; very little neck of middle flagellomeres and presence of numerous round loops of sensorial filae around middle flagellomeres; presence of 1–2 swollen peritremae on basal enlargement of flagellomeres. E t y m o l o g y. From the toponym Polesje (adj.)., Published as part of Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, pp. 35-50 in Vestnik Zoologii 48 (1) on page 40, DOI: 10.2478/vzoo-2014-0004, http://zenodo.org/record/6452791