We have described a new, distinct camellia cultivar, Camellia + 'Daisy Eagleson' (Byrd ex Meyer), found on a rootstock of hexaploid C. sasanqua Thunb. 'Maiden's Blush' on which a scion of diploid C. japonica L. had been grafted. The new cultivar is a graft chimera with epidermal tissue from C. sasanqua and the internal tissue from C. japonica. Many characters expressed and measured at the cellular or tissue level showed complete independence in the chimera. These included chromsome number, anthocyanin synthesis, pH, cell size, and fragrance. Some characters expressed at the organ or whole plant level showed an interaction between the two components. These included floral morphology, leaf size, time of bloom, bud hardiness, and growth habit. One entirely new character we found was the presence of 6-8 styles in the chimeral flowers in contrast to the three styles typical of the component species. While the two components grew together in apparent harmonious and stable fashion, we have recovered each of the pure, homogeneous component cultivars a number of times from tissue replacement and displacement phenomena in the shoot and as adventitious shoots from roots. 'Daisy Eagleson' is only the third, fully-authenticated, woody graft chimera. PLANT CHIMERAS, which can persist because of the pattern of ontogenesis in higher plants (angiosperms and gymnosperms), have an important application in ontogenetic studies (Satina, Blakeslee, and Avery, 1940; Dermen, 1945, 1960). They are of practical importance in that they can produce a desirable change in color or various other characteristics with a minimum change in cultural requirements or other important horticultural qualities. The most useful, most common, and the only persistent forms are the periclinal chimeras which were first described and the term applied to geranium (Pelargonium) chimeras by Baur in 1909. His material was chimeral as the result of somatic, plastogene mutation confined to a single apical layer and to its derivatives in the primary body. In 1910 Winkler recognized that some of the offtype shoots he had obtained in 1907 from callus at graft unions of Solanum nigrumn and S. esculentum were periclinal chimeras. Similar periclinal graft chimeras have been produced between herbaceous species of Solanacae, but among woody plants in no more than two well-documented cases (+2 Laburnocytisus adamii and + Crataegomespilus asnieresii: Bergann, 1958; Dermen, 1969; Neilsen-Jones, 1969; Balkema, 1971). The majority of periclinal chimeras described in the literature have arisen as the result of somatic mutation within a single layer of the shoot apical meristem which was confined to derivatives of that layer throughout the plant body. The layers are 1 Received for publication 11 October 1971. 2 + designates the asexual association of two components in a graft-chimera. genetically identical except for the one mutant characteristic which may be due to a chromosomal gene (Stewart and Arisumi, 1966), a plastogene (Stewart, 1965), or chromosome number (Dermen and May, 1966). The graft chimeras are of particular interest because one apical layer differs from the others in many genetic characteristics. They offer unique opportunities for studying the interaction of genetically diverse tissues growing in intimate association and provide many morphological and physiological markers of the cell lineages for ontogenetic studies in contrast to the single marker in chimeras which arise as the result of mutation (Stewart and Dermen, 1970a, b). A newly discovered graft chimera of two commonly cultivated Camellia species has provided material for complete documentation of a third woody chimera and makes this material available for teaching and research purposes. There are many obvious cases of complete independence of phenotypic expression of specific characters as well as a number of instances of profound influence of tissue derived from one component upon the development of the other. MATERIALS AND METHODS-The plant described here in detail and formally named Camellia + 'Daisy Eagleson' (hereafter referred to as DE), was reported by L. R. Byrd, Jr. (1970). DE originated in 1954 after the grafting by Mr. Tom Eagleson of Port Arthur, Texas, of a C. japonica scion with dark-rose colored, white-blotched, completely double, tiered, and sterile flowers onto an