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Key to species of Platyparadonus 1. Dark brown, legs and antennae lighter, contrasting with body; generally larger, 3mm or longer; known from the Greater Antilles......................................................... P. marleyi Etzler and Ivie, new species. 2. Amber (yellow-brown), legs and antennae not contrasting with body; generally smaller, less than 3mm; known from a single locality in Colombia...................................................... P. beckeri Etzler and Ivie, new species.

Platyparadonus marleyi Etzler and Ivie, new species urn:lsid:zoobank.org:act: D0E65DC2- 40F9-4201-A4A2-6FCC094564F7 (Figs. 1���3, 7���22, 24���27) Diagnosis.���A generally larger size, ��� 3mm, a dark brown body with con- trasting legs and antennae, and an aedeagus that is as long as the abdomen and curved basally, will readily separate this species from the only other known species, P. beckeri. Description.���Dark brown with legs and antennae amber, contrasting body. Length 3.0��� 3.6mm. Antennomeres 3���10 about 2X longer than wide, apical end lighter than basal two-thirds. Aedeagus (Figs. 25���27) relatively long, around 1mm, nearly as long as abdomen, with basal curve, basal struts of median lobe not reach base of parameres, not passing basal curve. Etymology.��� The specific name is named to honor Robert ���Bob��� Nesta Marley, to recognize his importance in spreading Reggae music throughout the world, and to recognize his connection to Jamaica, where he was born. Specimens examined.��� 11. Holotype: JAMAICA: Port Royal, 21���II��� 1908, M. Cameron/ W. I. 150/ M. Cameron Coll., B. M. 1936���555, (male, BMNH). Paratypes: CUBA: Maisi, Oriente, July 17, 1936 / Darlington Collector (1, MCZC). JAMAICA: Port Royal, 21���II���1908, M. Cameron/ W. I. 150/ M. Cameron Coll., B. M. 1936-555. (5, BMNH; 2, WIBF). U. S. VIRGIN IS: St. John, Europa Bay, 22 MAY���27 JUN 1979 / sifted from under wrack and under rocks on beach/ Colr. W. B. Muchmore (1, WIBF 017010); St. John, Lameshur Bay, V.I.E.R.S., 15 AUG 1980, at UV light/ M. A. Ivie Colr. (2, WIBF 017008, WIBF 017009). Discussion and field notes.��� To date, this species has only been collected on Cuba, Jamaica, and St. John, in the Greater Antilles. As a geographical note, the Virgin Islands belong to the geologic and biogeographic Greater Antilles (Donnelly 1988), the Anegada Passage being the divider between the Greater and Lesser Antilles. The Jamaican series of specimens was collected by Malcolm Cameron (1873��� 1954) a specialist on Staphylinidae. He collected in the West Indies while a naval surgeon with the British Royal Navy. During this period, he was particularly interested in coastal Aleocharinae, and produced three major descriptive works on West Indian staphylinids (Cameron 1913, 1922, 1923). Some hint of the habitat of Cameron���s specimens of P. marleyi can be gained from other species Cameron collected at Port Royal. Blackwelder���s (1943) revision of the West Indian Staphylinidae has two references to Port Royal, Jamaica, both associated with Cameron. First, Cameron described Cafius subtilis Cameron, 1922, from Jamaica, with no further mention of locality or date. Blackwelder (1943) examined Cameron���s types, and narrowed the type locality to Port Royal, apparently from label data on the types, but without date information. The habitat of C. subtilis is mentioned by Blackwelder (1943: 437) as ���I have found it under seaweed and drift on sandy beaches.��� A second Port Royal record from Cameron is Cafius bistriatus (Erichson, 1840) (Blackwelder 1943). Blackwelder (1943: 438) states about C. bistriatus, ���I have found it under seaweed and drift on sandy beaches.��� As a last bit of evidence that Cameron���s P. marleyi was found on sand, Port Royal itself is located on a sand spit, so sandy that it is subject to liquification during the periodic earthquakes that have devastated Port Royal as early as 1692 and as recently as 1907. The Lameshur and Europa Bay collecting localities are on the south���east coast of St. John in the U.S. Virgin Islands. "Lameshur Bay" by itself is used to refer to the Estate of Lameshur Bay, while the bays themselves are Europa, Great Lameshur and Little Lameshur Bays, and all occur next to one another across narrow peninsulas. The acronym "V.I.E.R.S.," refers to a research station (18.322235 N, 64.722873 W) built by the U.S. Navy, in cooperation with other agencies, in 1968 to support Project Tektite, an undersea habitat experiment platform from 1969���1970. Originally called the Virgin Islands Ecological Research Station (including at the time these specimens were collected), in 1992 it was renamed Virgin Islands Environmental Resource Station. Both William Breuleux Muchmore (1920���2017) (pers. comm.) and MAI used VIERS as their base of operations when these paratypes were collected. The station is built on an alluvial sand area about 1/ 4 km from the actual beach. MAI���s blacklight was placed to the back of one of the cabins, facing into the forest over sand. Muchmore���s Europa Bay site (18.318483 N, 64.731362 W) is a km south-west of the Station, the wrack taken from the rockstrewn sand beach. Notebooks for the Darlington specimen collected from Cuba have not been located, but the maps of the area suggest a similar habitat to the above. All of these factors point towards P. marleyi as occurring on or in sandy substrates., Published as part of Etzler, Frank E. & Ivie, Michael A., 2019, Platyparadonus, A New Circum-Caribbean Genus Of Sand-Associated Negastriinae (Coleoptera: Elateridae), pp. 15-25 in Proceedings of the Entomological Society of Washington 121 (1) on pages 22-23, DOI: 10.4289/0013-8797.121.1.15, http://zenodo.org/record/3733353, {"references":["Donnelly, T. W. 1988. Geologic constraints on Caribbean biogeography, pp. 15 - 37. In Liebherr, J. K. (ed.). Zoogeography of Caribbean insects. Cornell Press, Ithaca, NY.","Cameron, M., 1913. Descriptions of new species of Staphylinidae from the West Indies. Part I. The Annals and Magazine of Natural History series 8, 12: 321 - 351.","Cameron, M. 1922. Descriptions of new species of Staphylinidae from the West Indies. Part II. The Annals and Magazine of Natural History 9: 633 - 652.","Cameron, M. 1923. Descriptions of new species of Staphylinidae from the West Indies. Part II (cont.). The Annals and Magazine of Natural History 11: 363 - 400.","Blackwelder, R. E. 1943. Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the U. S. National Museum 182: iviii, 1 - 658."]}

Platyparadonus Etzler and Ivie, new genus urn:lsid:zoobank.org:act: 766D28FF- 0F8F-4FFA-8769-2117263028E8 Type species: Platyparadonus marleyi Etzler and Ivie, new species. Diagnosis.��� Platyparadonus appears to be closely related to other Negastriinae genera that lack visible elytral striae, such as Paradonus Stibick (Fig. 5) and genera placed in the tribe Quasimusini Schimmel and Tarnawski 2009 (Fig. 6). Both Paradonus and Platyparadonus can be separated from the genera of Quasimusini by lacking hind angles that embrace the humeral angles of the elytra (see key above). Platyparadonus can be distinguished from Paradonus by the following combination of characters: lateral carina of pronotum limited to pronotal hind angle (Fig. 14), antennae passing posterior edge of pronotum by 3.5-4 antennomeres, antennomere 2 onethird longer than antennomere 3 (Fig. 9), and possessing laterally lobed tarsomeres (bilobed or cordiform, Fig. 16). Species of Paradonus possess a complete lateral carina on the pronotum; antennae that do not pass the posterior edge of the pronotum; antennomere 2 slightly longer than, but almost equal in length to, antennomere 3; and tarsi simple, with filiform tarsomeres. Description.���Body length 2.3���3.6 mm. Body uniform, yellow-brown to dark brown. Body dorso-ventrally flattened (Fig. 2). Head (Figs 10���12): punctures indistinct, small, simple, deep, separated by one to two diameters of punctures, interspaces rough and densely microreticulate, setae short, dense, yellow to grey; eyes semicircular, truncate dorsally, not bulging; supra-antennal carina short, merging with clypeal margin so frons not separate from clypeal margin; mouthparts prognathous, mandibles tridentate (Fig. 8), maxilla as in Fig. 7. Antennae (Fig. 9) feebly serrate, sometimes almost filiform; passing posterior edge of pronotum by 3.5���4 antennomeres; antennomere 1 (pedicel) expanded, curved and smooth posteriorly, densely setose anteriorly; antennomere 2 cylindrical, as long as 1; antennomere 3 feebly serrate, about two-thirds length of 2; antennomeres 4���10 feebly serrate, all subequal in length to one another; antennomere 11 oval, with single dorsal impression; antennomeres 4���11 with apical third often subtly lighter in color, each bearing a black spot near ventral side near midlength or apical two-thirds; setae erect; punctation on antennomeres 1���3 similar, subtly less than punctation on antennomeres 4���11. Thorax: pronotum (Figs. 13���15) wider than long, widest at mid-length, sides broadly arcuate, lateral carina limited to pronotal hind angle only, posterior edge with ���m���-shaped notch just anterior of scutellum, sublateral basal incisures not present; hind angles short, sharply pointed, not extending past posterior edge of pronotum, not embracing humeral angle of elytra, hind angle lacking dorsal carina; punctation large, lacking patterned placement but uniform (no difference between disc and lateral area), subumbilicate, adjacent to separated by 2X diameter of a puncture, interspaces dull, densely microreticulate. Hypomeron not separated from pronotum by carina, dull, appearing rough, interspaces densely microreticulate, punctures only visible along prosternal suture, separated by a diameter at most, hind angle visible, but not strongly projecting posteriorly, small medial projection behind procoxal cavity present. Prosternum broad, widest near anterior third, punctures large, lacking patterned placement but uniform throughout, subumbilicate, some nearly touching to separated by a diameter of a puncture, interspaces dull and microreticulate, carinate around procoxal cavities; anterior lobe short, rectangular, truncate to slightly convex medially, not covering ventral mouthparts, not deflexed ventrally, two black spots on either side of midline present basally (not visible in most noncleared specimens); prosternal sutures curved, marginate along hypomeral border, shining, not excavate or grooved anteriorly; prosternal process straight, slender, acutely pointed, very short past procoxae, without lateral expansions, flattened between procoxae. Mesosternal fossa shallow, parallel sided, not angled (Fig. 19). Mesepisternum and mesepimeron fused, not reaching mesocoxal cavity, mesocoxae surrounded by meso- and metaventrite, meso- and metaventrite fused medially. Metaventrite with deep, simple punctation, separated by 0.5X diameter medially, rapidly becoming sparser laterally, separated by 1.0 diameter or more, punctures not easily distinguished from interspaces, interspaces microreticulate and dull. Scutellar shield (Fig. 18) subpentagonal, weakly convex, about 2X longer than wide, anterior edge slightly curved, carinate and elevated above base, posterior edge broadly rounded, surface minutely punctured and weakly shining. Elytron (Fig. 17) with single stria near scutellar shield and 2X length of scutellar shield, otherwise lacking dorsal striae, punctures small, subumbilicate, separated by 1���2X diameters, apical fifth lacking punctures, interspaces microreticulate, dull, obscuring punctation, ventral striae (only visible ventrally or in cleared specimens) absent medio-basally. Legs (Figs. 16, 20) yellow-brown, femora enlarged and laterally flattened, tibiae moderately laterally flattened, metatibia more so, expanded distally, bearing a single apical spine, tarsomeres dorsoventrally flattened, bilobed (cordiform), all approximately equal in length, less pronounced bilobes on metatarsi, which are relatively longer compared to pro- and mesotarsi, claws simple, without basal setae, strongly curved; metacoxal plate (Fig. 20) with ventral expansion as wide as base until midlength, then rounded at 90-degree angle, extremely narrow at lateral margin. Hind wing (Fig. 24): well developed, 2.3X as long as wide; wing membrane notched in anal area. Radial cell reduced, 2.1X as long as wide, proximal, posterior angle obtuse; horizontal radial crossvein (r3) extending from radial cell 1.8X length of radius; radial vein thickened apically, with slight posterior projection at apex. Vein MP 4 without apparent crossvein to CuA 2. Wedge cell absent. Apex of wing membrane without venation, occupying 0.33X length of wing membrane, with two weakly sclerotized areas. Posterior membrane edge with setal fringe. Abdomen: punctures small, not separable from interspaces, which are microreticulate and dull. Fifth abdominal ventrite broadly rounded posteriorly. Terminal ventrite either wide, broadly rounded apically (females, Fig. 22), or narrow, hour-glass shaped, rounded apically (males, Fig. 23). Terminal tergite similar to female terminal ventrite in both sexes. Male genitalia (Figs. 25���29): Aedeagus equal to half length of abdomen to as long as abdomen, straight or possessing slight dorsal curve basally. Parameres with basal portions (basimere) fused to one another and medio-ventral strut of median lobe, forming rigid cylinder, dorsum covering ventrum side except near base; gradually attenuate basally, apex moderately rounded to subacute; apical portions (telomere) slender, separated dorsally as a movable condylite with digitate base and articulate to medio-ventral strut of median lobe, bearing one major subapical seta, apex narrowly rounded; full paramere complex subparallel throughout. Median lobe (centerpiece) broad, lateral struts slightly arcuate at base, otherwise straight, not reaching paramere base, apex usually moderately acuminate, exceeding apex of parameres. Basal piece small, weakly sclerotized, subquadrate. Female genitalia (Fig. 21): ovipositor weakly sclerotized, 0.8X length of abdomen. Coxites setose, lacking styli. Paraprocts elongate, 0.6X length of ovipositor. Uterus membraneous, without sclerotizations, bulbous; colleterial glands not visible, bursa copulatrix connected to uterus by narrow tube, divided into two membraneous arms [these arms may be the spermathecal gland]. Etymology.���The name Platyparadonus is the combination of Platy -, derived from the Greek word platus meaning broad or flat, referring to the dorso-ventrally flatted aspect, and Paradonus to denote a possible relationship with species in the genus Paradonus. The name is masculine in gender. Notes.���The known distribution of Platyparadonus is circum-Caribbean, known from the Greater Antillean islands delimiting the Caribbean to the north and the north coast of South America to the south. It would not be surprising if species were to be discovered along the Caribbean coast of Central America and in the Lesser Antilles, even the Lucayan Archipelago. Specimens were collected by sifting beach wrack or at an ultraviolet light. All localities seem to be on or near marine sand beaches. Their presence under beach wrack and the dense (hydrofuge?) setae on the body may suggest that they are able to float to the surface if their substrate is flooded. More detailed habitat data are presented under each species. Only two species are known in the genus. Species descriptions that follow are short, highlighting unique aspects of each species, otherwise both species share the characters given in the genus description., Published as part of Etzler, Frank E. & Ivie, Michael A., 2019, Platyparadonus, A New Circum-Caribbean Genus Of Sand-Associated Negastriinae (Coleoptera: Elateridae), pp. 15-25 in Proceedings of the Entomological Society of Washington 121 (1) on pages 17-22, DOI: 10.4289/0013-8797.121.1.15, http://zenodo.org/record/3733353, {"references":["Schimmel, R., and D. Tarnawski. 2009. Monograph of the tribe Quasimusini (Insecta: Coleoptera, Elateridae, Negastriinae). Polish Taxonomical Society, 169 pp."]}

Platyparadonus beckeri Etzler and Ivie, new species urn:lsid:zoobank.org:act: 7A979C50- E2D8-4775-9DE9-6A65F365DBA5 (Figs: 4, 23, 28���29) Diagnosis.���A generally smaller size, P. marleyi. Description.���Amber, head and pronotum darker amber, legs and antennae not contrasting with body. Length 2.3mm. Antennomeres 3���10 about 1.5X longer than wide, apical and basal portions concolorous. Aedeagus (Figs 28���29) ca. 0.5mm long, straight, basal struts of median lobe nearly reaching base of parameres. Etymology.��� The specific name is in honor of the late Dr. Edward Coulton Becker (1923���2008), to recognize his important contributions to the study of Elateridae and his early recognition of these species as belonging to a distinct genus. Specimens examined.��� 2. Holotype: Colombia: Magd. Parque, Tayrona, 21mi E. Sta. Marta, V���15���1973, Howden and Campbell (male, CNCI). Paratype: Same data as holotype (1, CNCI). Field notes.��� Both specimens were collected on a hot, overcast day, either in the afternoon or early evening at a blacklight (1:30���8PM) (Howden 1973). The collecting locality was a heavily cut-over area along the coast with coconut palms, so it appears that disturbed habitat does not necessarily affect this species. Satellite photographs of the locality show a prevalence of wrack on the beach immediately adjacent to the coconut groves. This suggests that this species may also be found under beach wrack, similar to P. marleyi., Published as part of Etzler, Frank E. & Ivie, Michael A., 2019, Platyparadonus, A New Circum-Caribbean Genus Of Sand-Associated Negastriinae (Coleoptera: Elateridae), pp. 15-25 in Proceedings of the Entomological Society of Washington 121 (1) on pages 23-24, DOI: 10.4289/0013-8797.121.1.15, http://zenodo.org/record/3733353, {"references":["Howden, H. A. 1973. Unpublished field notes dated 15 May [1973]. Deposited in the entomology department, Canadian Museum of Nature, Gatineau, Quebec, Canada."]}

Athoplastus acutus (Knull, 1946), new combination (Adult: Figs. 1, 5���7, 9���10, 12, 15���18) (Larva: Figs. 31���46) Aplastus acutus Knull 1946: 142 Ctenicera acutus Knull 1957: 202 Ctenicera idahoensis Knull 1958: 154. Invalid replacement name to prevent homonymy with Athous acutus Mulsant and Guillebeau, 1855 (= Metanomus infuscatus (Eschscholtz, 1829)). The Mulsant and Guillebeau name was considered a junior synonym of Metanomus infuscatus (Eschscholtz) since de Buysson (1887) but never appeared as a valid taxon name in combination with Ctenicera, so no homonymy exists (see ICZN 1999, Art. 60.2). Diagnosis. Males of A. acutus have the prosternum dorsoventrally flattened until it reaches the procoxae, where it is abruptly expanded ventrally (Fig. 9). Males of other species of the genus have the prosternum gradually expanding ventrally toward the procoxae (Fig. 8). Antennomeres 6 and 7 usually have a secondary expansion at approximately twothirds of total length from base, which is not present in the other species (Fig. 12). This character may be difficult to see or absent in some specimens, but it is usually visible on other antennomeres. Females have a slight impression on the prosternum anterad the procoxae and setae on the scutellum that are mostly projecting posteriorly. Description. Male. Length 15.0��� 17.5 mm, width 3.5���4.0 mm (measured across bases of elytra). Yellow-brown to dark brown (Figs. 1, 6���7). Head: Ocular index = 55.0���63.6 (mean = 58.1; mode = 60; n = 30). Antenna: Mean ratio of antennomeres [antennomere length relative to pedicle (antennomere 2)] 2���11: 1: 1.4: 3.5: 3.9: 3.9: 4.1: 4.1: 4.1: 3.9: 4.8 (n = 11). Antennomeres 6 and 7 often with secondary expansion at 2/3 total length (see Fig. 12). This character may also be on other antennomeres but is usually less pronounced. Pronotum: 1.2���1.7X wider than long. Lateral margin sinuate anterad divergent hind angles, widening just anterad midlength, lateral carina dorsoventrally flattened on anterior half. Punctures distinct, subumbilicate and dense on disc, separated by less than half their own diameter, although some specimens have impunctate areas just laterad midline; punctures becoming clearly umbilicate and denser along anterior and lateral margins. Prosternum: Dorsoventrally flattened with dense, shallow, umbilicate punctures; anterior lobe short, broadly rounded, shallowly deflexed. Posterior process short and arcuate, distinctly raised ventrally between procoxae, then sharply deflexed dorsally before ending in small hook; midline of prosternum often with raised section just anterad procoxal rise, sometimes nearly carinate, flanked by indentations in prosternum. Legs: Tarsomere 5 equal in length to tarsomere 1 on pro- and mesotarsi, tarsomere 5 equal in length to tarsomere 2 on metatarsi. Elytra: Elongate, subparallel, 5.2X pronotal length; striae shallowly impressed, deeper basally and apically, punctures evanescent; intervals shallowly convex, punctures rough, shallow, and irregularly distributed. Aedeagus (Figs. 16���18): Basal piece about 1/3 total length, with triangular membranous section on ventral surface (Fig. 17); median lobe 1.3X length of parameres, slightly deflexed ventrally at tip, median carina on ventral surface, basal struts coming off at an obtuse angle; parameres wide basally, edges sinuate and distinctly narrowing toward tip, tip with acute lateral expansion bearing 4���6 setae, either blunt or sharp (Fig. 18). Female. Length 21���26 mm, width 5���6 mm (measured across bases of elytra). Yellow-brown to red-brown (Fig. 5). Head: Ocular index = 70.8���74.1 (mean = 72.7, n = 3). Antenna: Mean ratio of antennomeres 2���11: 1: 1.2: 2.1: 1.9: 1.9: 1.8: 1.7: 1.6: 1.5: 1.7 (n = 3). Note: Values for antennomeres 8, 10, and 11 are each based only on two specimens. Pronotum: As wide as or 1.1X wider than long. Lateral edge sinuate anterad divergent hind angles, widening just anterad midlength. Scutellar shield: Setae directed posteriorly. Specimens Examined (45). Males. Holotype. USA: IDAHO: [Shoshone Co.], Wallace; 14.iv.[19]40, Otto Huellemann (FMNH). Paratypes. USA: IDAHO: [Shoshone Co.], Wallace, 14.iv.1940, [Otto Huellemann?] (2, FSCA; 1, FMNH); Wallace, Otto Huelleman (1, FMNH). Other Male Specimens Examined. USA: IDAHO: [Kootenai Co.] Ft. Sherman [Coeur d��� Alene], Mch. [March?] 1896, J. C. Merrill (1, USNM); 4 th of July Summit, 30.iv.1974, W. F. Barr (1, WFBM); Kidd Island Bay, 1987, E. Cardwell (1, WFBM); Latah Co., Harvard, Laird Pk, 03.iv.1962, S. D. Smith, attracted by U.V. light (1, USNM; 4, WFBM); Near Moscow, Spring 1932, P. L. Rice (1, USNM); Sect. @ Pond; Moscow, 01.v.[19]57, D. T. Allison, Collector (1, WFBM); [Shoshone Co.] Clarkia, 8.iv.1934, I. W. Bales (1, USNM); Wallace, 8.iv.1939, Huelleman (1, USNM); Wallace, 14. iv.1939, Huelleman (1, FMNH); Wallace, 26.iv.39, Huelleman (2, FMNH); Wallace, 1.v.1939, Huelleman (1, FMNH); Wallace, 9.v.39, Huelleman (1, FMNH); Wallace, 20.vii.1939, Huelleman (1, FMNH); Wallace, [no date], Huelleman (1, FMNH); Wallace, 12���20.iv.1940, O. Nudlemann [likely a mistype of Huelleman] (3, FMNH). MONTANA: Lincoln Co., Jnct. Kootenai & Yaak Rs., 48.5855��N, 115.9769��W, 12.iv.-03.vi.2014, [elev.] 1927ft, C.J. Hart, Lindfnl [with] PSB [lure] (1, MTEC, MTEC019710); Ravalli Co., Rombo Campgrd, 45.7644��N, 114.2815��W, 25. iv.2013, [elev.] 4505ft, F. E. Etzler, Handnet at Night (1, MTEC010381); Rombo Campgrd, 45.7680��N, 114.2810��W, 04.v.2013, [elev.] 4508ft, F. E. Etzler, Handnet at Night (1, MTEC); Rombo Campgrd, 45.7642��N, 114.2818��W, 19.iv.2014, [elev.] 4505ft, F. E. Etzler, Blacklight (8, MTEC). WASHINGTON: Stevens Co., Pleasant Valley, 4.v.1957, on maple, A. R. Gittins Collector (1, WFBM). Females. USA: IDAHO: [Latah Co.], Moscow Mt., iv.[19]12, Idaho (1, USNM); Moscow, 23.iv.1955, F. T. Cook (1, USNM; 1, WFBM). MONTANA: Mineral Co., Trout Creek Camp, 17.iii���22.vi.1995, D. L. Gustafson, pitfall trap (1, MTEC); Ravalli Co., Sand Crk - Blue Joint [Crk], 05.x.1996 ��� 06.vi.1997, D. L. Gustafson, pitfall trap (1, MTEC); Rombo Campgrd, 45.7644��N, 114.2815��W, 25.iv.2013, [elev.] 4505ft, F. E. Etzler, handnet at night (1, MTEC010382). Larvae. USA: IDAHO: Moscow Mtn, 15.v.1951, Lane-Downie (2, USNM). MONTANA: Mineral Co., Trout Creek Campground, 47.1174��N, 114.8673��W, 25.iv���19.vi.2013, elev. 2940ft, F. E. Etzler, pitfall (1, MTEC, MTEC008126); Ravalli Co., Blue Joint Creek, base of stump, 45.6985��N, 114.3586��W, 26. iv���5.v.2013, elev. 4960ft, F. E. Etzler, pitfall (1, MTEC, MTEC019709); Rombo Campground, 45.7680��N, 114.2810��W, 4.v.2013, elev. 4508ft, F. E. Etzler, by hand (1, MTEC, MTEC019713). Distribution. This species is known from westernmost Montana, northern Idaho (north of Idaho County), and the northeast corner of Washington State (Fig. 29). Taxonomic Remarks. This species is represented by the greatest number of specimens and has the broadest known species range of the four species in the genus.Due to this and the soft-bodiedness of the species, variation exists among specimens. However, no clear link to locality or date of collection was discovered.The presence of impunctate areas on either side of the midline is very rare, only occurring in a small number of northern Idaho specimens.Subtle variation also exists in male genitalia, but again there is no link to locality or collecting date. Mitochondrial DNA was amplified for Montana specimens. These results are discussed below. This is the only known species with associated larvae and ecological notes. Field Notes. The collecting localities of the Montana specimens of A. acutus were visited in early spring of 2013 to place pitfall traps in an attempt to capture fresh specimens and reliable DNA samples. While camped at Rombo Creek Campground near one of the localities of the MTEC females, the first author was examining trees around the campsite at dusk when a female was collected at the base of a ponderosa pine. A few minutes afterwards, a male was collected at the same spot, then additional males were observed flying and a few more collected. The collecting site was revisited in 2014, and specimens were collected at an ultra-violet (UV) light set up near a white sheet. Field observations were recorded and are discussed below. Larvae were collected at three Montana sites and tentatively associated with adults. Due to the apparent adult sexual dimorphism and lack of reared larvae, we confirmed associations using DNA barcoding., Published as part of Etzler, Frank E. & Johnson, Paul J., 2018, Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA, pp. 503-521 in The Coleopterists Bulletin 72 (3) on pages 513-516, DOI: 10.1649/0010-065X-72.3.503, http://zenodo.org/record/4789344, {"references":["Knull, J. F. 1946. A new species of Aplastus from Idaho (Coleoptera: Plastoceridae). The Ohio Journal of Science 46: 87 - 89.","Knull, J. F. 1957. Three new species of Aphricus with a note of Ctenicera (Coleoptera: Elateridae). The Ohio Journal of Science 57: 200 - 202.","Knull, J. F. 1958. Two new species of Ctenicera (Coleoptera: Elateridae). The Ohio Journal of Science 59: 87 - 89."]}

Athoplastus rainieri (Van Dyke, 1932), new combination (Adult: Figs. 4, 8, 22���24, 28) Ludius rainieri Van Dyke 1932: 397, 421. Ctenicera rainieri (Van Dyke, 1932); Lane 1971: 32. Diagnosis. This species is distinguished from all other species of the genus by the combination of close pronotal punctures (separated by less than one diameter of a puncture) (Fig. 10), sloped prosternum (Fig. 8), and dark markings at each puncture along the striae of the elytra (Fig. 4). This last character is likely not a good species-level character and may be related to specimen development. Athoplastus rainieri is quite similar to A. palouse, which can be distinguished as noted above. Description. Male. Length 16 mm, width 4 mm (measured across bases of elytra). Red-brown, carina and tips of mandibles darker; antennae, palps, coxae, and tarsi lighter, almost tawny yellow (Fig. 4). Head: Ocular index = 61.1. Antennae: Ratio of antennomeres (2���11): 1: 1.25: 3: 3.25: 3.5: 3.25: 3.75: 2(?): X: X. [Antennomere 9 appears to be broken, and 10 and 11 are missing on the specimen measured.] Pronotum: 1.1X wider than long. Lateral margin completely carinate, slightly sinuate, almost subparallel anterad hind angles; convergent and slightly dorsoventrally flattened along lateral edges on anterior fourth. Punctures distinct, subumbilicate and dense on disc, separated by less than 0.2X their own diameter, with 2 ���dimples��� on either side of midline just anterad midlength, with less close punctures; punctures becoming clearly umbilicate and denser along anterior and lateral margins. Prosternum: Slightly convex with dense, shallow, umbilicate punctures; anterior lobe short, broadly rounded, shallowly deflexed. Posterior process short and arcuate, gradually raised ventrally between procoxae, then sharply deflexed posteriorly; distinct ridge present immediately before procoxal cavity and along ventral surface of prosternal process. Legs: Tarsomere 5 equal to tarsomere 1 on pro- and mesotarsi, tarsomere 5 missing on both metatarsi. Elytra: Elongate, subparallel, 4X pronotal length; striae shallowly impressed, deeper apically, punctures marked by darkening of the integument; intervals shallowly convex, punctures fine, shallow, and irregularly distributed. Aedeagus (Figs. 22���24): Basal piece approximately 1/3 total length, with deep, narrowly U-shaped, membranous section on ventral surface (Fig. 23); median lobe 1.2X length of parameres, slightly deflexed ventrally at tip, median carina on ventral surface, basal struts following edges of median lobe; parameres wide basally, edges slightly sinuate and gradually narrowing toward tip, tip with subacute lateral expansion bearing 2 setae (Fig. 24). Female. Length 21 mm, width 5 mm (measured across bases of elytra). Yellow-brown. Head: Ocular index = 72.7. Antenna: Ratio of antennomeres 2���11: 1: 1.6: 2.2: 2: 1.8: 1.6: 1.8: 1.6: X: X: X. [Note: Antennae are broken on specimen examined, so the last three antennomeres of longest intact antenna are missing.] Pronotum: 1.25X wider than long, hind angles divergent, sides broadly sinuate. Legs: Tarsomere 5 intermediate in size between tarsomeres 1 and 2 on metatarsi; pro- and mesotarsi as in male. Scutellar shield: Setae radiating from midline. Specimens Examined (3). Male. Holotype. USA: WASHINGTON: [Pierce Co.]; Mt. Rainier [E.C. Van Dyke, date unknown] (CASC, 1, CASC type no. 3181). Female. USA: WASHINGTON: [Kittitas Co.] Lk. Cle Elum; 13.v.1936; M.C. Lane (1, USNM); Lk. Cle Elum [?], 26.v.1937, M.C. Lane (1, USNM). Distribution. This species is only known from the Cascade Range of mountains in Washington. Taxonomic Remarks. Van Dyke (1932) mentions a specimen seen ���in the collection of Prof. W. J. Chamberlin at Corvallis, Oregon.��� Chamberlain��� s collection was deposited at the Oregon State Arthropod Collection (OSAC) at Oregon State University (Oman 1970). However, a search of the OSAC yielded no specimen matching this description., Published as part of Etzler, Frank E. & Johnson, Paul J., 2018, Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA, pp. 503-521 in The Coleopterists Bulletin 72 (3) on pages 517-518, DOI: 10.1649/0010-065X-72.3.503, http://zenodo.org/record/4789344, {"references":["Van Dyke, E. C. 1932. Miscellaneous studies in the Elateridae and related families of Coleoptera. Proceedings of the California Academy of Science 20: 291 - 465.","Lane, M. C. 1971. Family Elateridae [pp. 6 - 34, excluding Cardiophorinae]. In: The Beetles of the Pacific Northwest, Part V: Rhipiceroida, Sternoxi, Phytophaga, Rynchophora, and Lamellicornia (M. H. Hatch, editor). University of Washington Press, Seattle, WA.","Oman, P. 1970. The Oregon State University insect collection. Bulletin of the Entomological Society of America 16: 165."]}

Athoplastus palouse Etzler and Johnson, new species (Adult: Figs. 2, 14, 25���27) Zoobank.org/ urn:lsid:zoobank.org:act: 9601386F-4698-4376-8144-FB8FED38DAB2 Diagnosis. This species can be distinguished from other Athoplastus species by the combination of the non-compressed prosternum (Fig. 8), dense pronotal punctation (Fig. 10), and elytra without dark markings along the striae of the elytra. More subtle characters that help distinguish this species are the more parallel-sided pronotum (Fig. 2) and the slightly wider antennomeres (Fig. 14). These last two characters are best used when comparing specimens of different species and become quite clear in this way. However, when identifying just a single specimen, these characters are less useful, so the characters from the key should be used to eliminate other potential species. This species is quite similar to A. rainieri, but the paramere tips and V-shaped membranous area of the basal piece of the male genitalia (Figs. 25���27) and geographic distribution, in addition to the subtle characters above, will distinguish these two species. Description. Male. Length 15���17 mm in length, width 3���4 mm (measured across bases of elytra). Red-brown to brown, carina, palps, and tips of mandibles darker; posterior margin of hypomeron, coxae, and tarsi lighter, almost tawny yellow (Fig. 2). Head: Ocular index = 60.0���68.4 (mean = 65.0, n = 3). Antennae: Mean ratio of antennomeres 2- 11: 1: 1.25: 3.4: 3.6: 4; 4: 3.8: 4: 5.2 (n = 2). [Note: Antennomeres 8���11 are adjusted values based only on one specimen. The antennomeres, on average, are less narrowed than in other species. This is difficult to describe, so compare Fig. 14 to Figs. 12 and 13.] Pronotum: 1.4���1.6X wider than long. Lateral margin completely carinate, subparallel anterad divergent hind angles; subtly dorsoventrally flattened along lateral edges near anterior fourth. Punctures distinct, simple and dense on disc, separated by at most 1 diameter, often by about 0.5X diameter of puncture; punctures becoming denser along margins, nearly continuous laterally. Prosternum: Slightly convex with dense, shallow, subumbilicate punctures; anterior lobe short, broadly rounded, shallowly deflexed. Posterior process short and arcuate, gradually raised ventrally between procoxae, then deflexed posteriorly. Legs: Tarsomere 5 equal to tarsomere 1 on all tarsi. Elytra: Elongate, subparallel, 4.0���4.5X pronotal length; striae shallowly impressed, deeper apically; intervals flat, punctures fine, shallow, and irregularly distributed. Aedeagus (Figs. 25���27): Basal piece 2/5 total length, with broadly triangular membranous section on ventral surface (Fig. 26); median lobe 1.3X length of parameres, slightly deflexed ventrally at tip, median carina on ventral surface, basal struts following edges of median lobe; parameres wide basally, edges slightly sinuate and gradually narrowing toward tip, tip with very acute lateral expansion bearing 2 setae (Fig. 27). Female. Unknown. Specimens Examined (8). Holotype. USA: WASHINGTON: [Whitman Co.], Pullman, 7.vi. [18]98, C. V. Piper (1, WSUC). Paratypes. USA: IDAHO: Kootenai Co., Athol, 29.iv.1957, W. F. Barr Collector (1, WFBM). WASHINGTON: Washington T[erritory] (1, USNM); Pullman, Sept. [year unknown] (1, USNM); [Whitman Co.], Pullman, 8.vi.[18]99 (1, WSUC); Pullman, 31.iii. [19]11 (1, WSUC); Pullman, 01.iv.[19]17 (1, USNM); Pullman, Apr [il] [19]20, Chester Dixon (1, USNM); Kamiak Butte, 23.ii.[19]34 Wn, J.F. Clarke (1, USNM). Distribution. This species is only known from the areas of Pullman, Washington and Athol, Idaho (Fig. 29). Etymology. The specific name ��� palouse ��� is a noun in apposition derived from the geographic area of collection of the majority of specimens. Taxonomic Remarks. The distance between localities is quite a surprise, but the morphology of specimens from the two localities is consistent. Athoplastus palouse may co-occur with A. acutus., Published as part of Etzler, Frank E. & Johnson, Paul J., 2018, Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA, pp. 503-521 in The Coleopterists Bulletin 72 (3) on pages 516-517, DOI: 10.1649/0010-065X-72.3.503, http://zenodo.org/record/4789344

KEY TO THE SPECIES OF ADULT MALE ATHOPLASTUS 1. Pronotum with discal punctures separated by less than puncture diameter (Fig. 10), closer laterally; aedeagus with parameres not broadly rounded apically or membraneous........... 2 1 ��. Pronotum with discal punctures separated by more than puncture diameter (Fig. 11); aedeagus relatively narrower and paramere tips broadly rounded and membranous apically (Figs. 19���21).......... Athoplastus lochsa Etzler and Johnson, new species (Fig. 3), Published as part of Etzler, Frank E. & Johnson, Paul J., 2018, Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA, pp. 503-521 in The Coleopterists Bulletin 72 (3) on page 511, DOI: 10.1649/0010-065X-72.3.503, http://zenodo.org/record/4789344

Athoplastus lochsa Etzler and Johnson, new species (Adult: Figs. 3, 11, 13, 19���21) Zoobank.org/ urn:lsid:zoobank.org:act: 53CAEDD4-4FC0-4042-ABEF-8D7369F90FCC Diagnosis. This species can be separated from all others in the genus by the combination of relatively sparse pronotal punctation (separated by greater than the diameter of a single puncture) (Fig. 11), sloped, non-depressed prosternum (Fig. 8), and antennomeres without secondary expansions (Fig. 13). The male genitalia are also distinctive, with the parameres broadly rounded apically (Fig. 21). Description. Male. Length 16.0 mm in length, width 3.5 mm (measured across bases of elytra). Red-brown, carina, tips of mandibles, and palps darker; edges of hypomeron and abdomen lighter, almost tawny yellow (Fig. 3). Head: Ocular index = 63.1. Antenna: Ratio of antennomeres 2���11: 1: 1.25: 2.9: 3: 3: 2.9: 2.9: 2.9: 2.9: 3.25 (Fig. 13). Pronotum: 1.5X wider than long. Lateral margin completely carinate, sinuate anterad divergent hind angles, widening at anterior third; dorsoventrally flattened along lateral edges on anterior half. Punctures distinct, simple and moderately dense on disc, separated by at least diameter of a puncture, with 2 ���dimples��� on either side of midline just anterad midlength that are nearly impunctate; punctures remain uniform along lateral edges, becoming denser on hind angles. Prosternum: Slightly convex with dense, shallow, subumbilicate punctures; anterior lobe short, broadly rounded, shallowly deflexed. Posterior process short and arcuate, gradually raised ventrally between procoxae, then gradually deflexed posteriorly before becoming parallel to body plane. Legs: Tarsomere 5 longest on pro- and mesotarsi, tarsomere 5 equal in length to tarsomere 1 on metatarsi. Elytra: Elongate, subparallel, 4.8X length of pronotum; striae shallowly impressed, deeper apically; intervals flat, punctures rough, shallow, and irregularly distributed. Aedeagus (Figs. 19���21): Basal piece 1/3 total length, with roughly trapezoidal membranous section on ventral surface (Fig. 20); median lobe just slightly longer than parameres when measured from basal struts, slightly deflexed ventrally at tip, median carina on ventral surface, basal struts coming off at an obtuse angle; parameres subparallel, gradually narrowing toward tip, tip with expanded lateral expansion bearing 4 setae (Fig. 21), which is broadly rounded apically. Female. Unknown. Specimens Examined (1). Holotype. USA: IDAHO: Idaho Co.; Lochsa RNA [Research Natural Area]; 29.iv.1985; P.J. Johnson (1, PJJC to USNM). Distribution. This species is only known from the Lochsa River Valley in Idaho County, Idaho (Fig. 29). Etymology. The specific name ��� lochsa ��� is a noun in apposition representing the Nez Perce name for the Lochsa River, along which the only known specimen was collected. Taxonomic Remarks. While it is best to describe a species based on more than one specimen, the unique pronotal punctation and male genitalia support recognition as a new species. We also feel that this description will promote collecting trips to collect more specimens of this species and expand its known range., Published as part of Etzler, Frank E. & Johnson, Paul J., 2018, Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA, pp. 503-521 in The Coleopterists Bulletin 72 (3) on page 516, DOI: 10.1649/0010-065X-72.3.503, http://zenodo.org/record/4789344