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Epitettix parallelus (Podgornaya, 1986) (Fig. 12) Vaotettix parallelus Podgornaya, 1986: 17. Vaotettix parallelus — Blackith, 1992: 196; Kim & Pham, 2014: 59. Epitettix parallelus — Storozhenko, 2021: 182. Material examined. VIETNAM, Th ừa Thiên Hu ễ province, Bạch Mã National Park: •1Ô; Rhododendron Trail, heading to Four Lakes, N16.18950 E107.85360, 1160.0± 7.7 m.a.s.l.; 9 March 2023, 11h02; among leaf litter; coll. M.K. Tan & V. T. Trung; VIET.23.63 (ZRC) (Fig. 12). Remarks. This species is widely distributed in Vietnam. The specimen from Bạch Mã National Park, nevertheless, differs by the emarginated apex of posterior process of the pronotum (in typical form apex of process rounded) (Fig. 12). This is likely intraspecific variability. Ecology. The leaf litter area at which the specimen was collected was not too far from a forest stream, but the area is clearly elevated and probably not part of the water table. Distribution. So far, known only from central Vietnam in the provinces of Gia Lai and Thừa Thiên Huễ [new]. Type locality. Vietnam, Gia Lai Province, Buon Luoi.

3. Epitettix guangxiensis (Zheng & Jiang, 1994) (Figs. 4, 24C) Pseudepitettix guangxiensis Zheng & Jiang, 1994a: 36 [description] (holotype —♁, China: Guangxi prov., Shangsi County, Hongqi, in IZSNU, examined); Jiang & Zheng, 1998: 274; Liang & Zheng, 1998: 35; Liang & Jiang, 2004: 117; Zheng, 2005: 29; Deng, Zheng & Wei, 2007b: 29; Deng et al., 2015: 154; Deng, 2016: 28. Epitettix guangxiensis (Zheng & Jiang, 1994): Deng, 2020: 416. Formosatettix guangxiensis Zheng & Jiang, 1998 [description] (holotype —♁, China: Guangxi prov., Shangsi County, Hongqi, in IZSNU, examined); Jiang & Zheng, 1998: 344; Zheng, 2005: 352; Deng, Zheng & Wei, 2007b: 317; Deng, 2016: 272; Wei, Deng & Lu, 2019: 1003; syn. nov. Type material examined. Holotype Pseudepitettix guangxiensis, ♁, China: Guangxi prov., Shangsi County, Hongqi, 9. July 1991, in IZSNU. Holotype Formosatettix guangxiensis, ♁, China: Guangxi prov., Shangsi County, Hongqi, 9. July 1991, in IZSNU. Other material examined. 11♁ 10♀, China: Guangxi prov., Shangsi County, Wangle, 23 July 2013, EMHU; 7♁ 15♀, China: Guangxi prov., Fangchen County, Fulong, 20 July 2012, EMHU. Notes. We examined the type specimens of Pseudepitettix guangxiensis and Formosatettix guangxiensis, and found that the structures, type locality and coloration of the body are the same in both taxa. Therefore, a new synomimy is here established. The comparison of the holotype of Pseudepitettix guangxiensis and Formosatettix guangxiensis and their additional specimens show that these taxa are conspecific and characterized by the vertex is distinctly surpassing anterior margin of eyes as an evenly broad, rounded process; the vertex is much wider than one of the eyes; the apex of the hind pronotal process is concave., Published as part of Deng, Wei-An, Luo, Jie-Ling, Huang, Chao-Mei & Lin, Li-Liang, 2023, Review of the genus Formosatettix Tinkham (Orthoptera: Tetrigidae) with description of new taxa, new synonymy and an updated key to species, pp. 201-243 in Zootaxa 5228 (3) on page 208, DOI: 10.11646/zootaxa.5228.3.1, http://zenodo.org/record/7532348, {"references":["Zheng, Z. M. & Jiang, G. F. (1994 a) Five new species of Tetrigoidea from Guangxi (Orthoptera). Guangxi Sciences, 1 (2), 32 - 36.","Jiang, G. F. & Zheng, Z. M. (1998) Grasshoppers and Locusts from Guangxi. Guangxi. Normal University Press, Guilin, 390 pp.","Liang, G. Q. & Zheng, Z. M. (1998) Fauna Sinica, Insecta. Volume 12. Orthoptera, Tetrigoidea. Science Press, Beijing, 278 pp.","Liang, G. Q. & Jiang, G. F. (2004) Four new species of Tetrigoidea from Tianlin County, Guangxi, South China (Orthoptera). Acta Zootaxonomica Sinica, 29 (1), 115 - 120.","Deng, W. A., Zheng, Z. M. & Wei, S. Z. (2007 b) Fauna of Tetrigoidea from Yunnan and Guangxi. Science and Technology Press, Nanning, 458 pp.","Deng, W. A., Zheng, Z. M., Li, X. D., Lin, M. P., Wei, S. Z., Yuan, B. D. & Lin, L. L. (2015) The groundhopper fauna (Orthoptera: Tetrigidae) of Shiwanshan (Guangxi, China) with description of three new species. Zootaxa, 3925 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 3925.2.1","Deng, W. A. (2016) Taxonomic study of Tetrigoidea from China. Ph. D. Dissertation, Huazhong Agricultural University, Wuhan, 341 pp.","Deng, W. A. (2020) Taxonomic revision of the subfamily Cladonotinae (Orthoptera: Tetrigidae) from China with description of three new species. Zootaxa, 4789 (2), 403 - 440. https: // doi. org / 10.11646 / zootaxa. 4789.2.5","Wei, S. Z., Deng, W. A. & Lu, X. Y. (2019) Pygmy grasshoppers of the genus Formosatettix Tinkham, 1937 (Orthoptera: Tetrigidae: Tetriginae). Journal of Natural History, 53 (17 - 18), 1001 - 1022. https: // doi. org / 10.1080 / 00222933.2019.1609113"]}

Epitettix yunnanensis (Zheng, 1995), comb. nov. (Figure 5 G���H) Pseudepitettix yunnanensis Zheng, 1995a: 343 [description] (holotype��� ♂, China: Yunnan prov., Simao County, in IZSNU, examined); Liang & Zheng,1998: 34; Liang & Jiang, 2004:117; Zheng, 2005: 31; Deng et al., 2007: 31, 402. Type material examined. Holotype, ♂, China: Yunnan prov., Simao County, 12 IV 1974, IZSNU. Notes. The species is not a Pseudepitettix species because the humeral angles of pronotum are absent; the fastigium of the vertex distinctly surpasses the anterior margin of eyes and the anterior margin of the fastigium of vertex is distinctly arcuate. It belongs to Epitettix Hancock, 1907. Epitettix yunnanensis is easy to identify: the body is smooth; the vertex is much wider than one of the eyes; the median carina of the pronotum arch-like in profile, apex of hind pronotal process truncate. Distribution. China (Yunnan)., Published as part of Deng, Wei-An, 2020, Taxonomic revision of the subfamily Cladonotinae (Orthoptera: Tetrigidae) from China with description of three new species, pp. 403-440 in Zootaxa 4789 (2) on pages 416-417, DOI: 10.11646/zootaxa.4789.2.4, http://zenodo.org/record/3887709, {"references":["Zheng, Z. M. (1995 a) New genera and new species of Cladonotinae from China (Orthoptera: Tetrigidae). Acta Zootaxonomica Sinica, 20 (3), 342 - 347.","Liang, G. Q. & Zheng, Z. M. (1998) Fauna Sinica, Insecta. Vol. 12. Orthoptera, Tetrigoidea. Science Press, Beijing, 278 pp.","Liang, G. Q. & Jiang, G. F. (2004) Four new species of Tetrigoidea from Tianlin County, Guangxi, south China (Orthoptera). Acta Zootaxonomica Sinica, 29 (1), 115 - 120","Deng, W. A., Zheng, Z. M. & Wei, S. Z. (2007) Fauna of Tetrigoidea from Yunnan and Guangxi. Science and Technology Press, Nanning, 458 pp.","Hancock, J. L. (1907) Studies of the Tetriginae (Orthoptera) in the Oxford University Museum. Transactions of the Entomological Society of London, 1907, 213 - 244, pl. 21"]}

Key to Chinese species of Epitettix 1. Width of vertex 1.9 times width of an eye, frontal costa not concave between eyes; pronotum without interhumeral carinae, apex of hind pronotal process with weak excision.................. E. guangxiensis (Zheng & Jiang, 1994), comb. nov. - Width of vertex 2.8 times width of an eye, frontal costa distinctly concave between eyes; pronotum with interhumeral carinae, apex of hind pronotal process truncate................................... E. yunnanensis (Zheng, 1995), comb. nov., Published as part of Deng, Wei-An, 2020, Taxonomic revision of the subfamily Cladonotinae (Orthoptera: Tetrigidae) from China with description of three new species, pp. 403-440 in Zootaxa 4789 (2) on page 416, DOI: 10.11646/zootaxa.4789.2.4, http://zenodo.org/record/3887709, {"references":["Zheng, Z. M. & Jiang, G. F. (1994) Five new species of Tetrigoidea from Guangxi (Orthoptera). Guangxi Sciences, 1 (2), 32 - 36."]}

Genus Epitettix Hancock, 1907 Type species: Epitettix punctatus Hancock, 1907, by original designation. Composition. The genus includes eleven species distributed in Malaysia (Borneo), Indonesia (Java), Papua-New Guinea, Vietnam, India and Madagascar (Blackith, 1992; Eades et al., 2014). Originally, Epitettix included only E. punctatus Hancock, 1907 from North Borneo (Hancock, 1907). Acridium emarginatum Haan, 1843 was described from New Guinea (Haan, 1843), but later transferred to Epitettix by Willemse (1928). Four species (Epitettix fatigans G��nther, 1938, E. humilicolus G��nther, 1938, E. lativertex G��nther, 1938 and E. tumidus G��nther, 1938) were described from Papua-New Guinea (G��nther, 1938). Later the same author described E. elytratus G��nther, 1939 and E. tamilus G��nther, 1939 from India, E. dammermanni G��nther, 1939 from Java (G��nther, 1939), and E. spheniscus G��nther, 1974 from Madagascar (G��nther, 1974). Finally E. striganovae Storozhenko, 2012 was described from Vietnam (Storozhenko, 2012)., Published as part of Storozhenko, Sergey Yu. & Dawwrueng, Pattarawich, 2014, Three new species of the subfamily Cladonotinae (Orthoptera: Tetrigidae) from Thailand, pp. 325-337 in Zootaxa 3811 (3) on page 326, DOI: 10.11646/zootaxa.3811.3.3, http://zenodo.org/record/231482, {"references":["Hancock, J. L. (1907) Studies of the Tetriginae (Orthoptera) in the Oxford University Museum. Transactions of the Royal Entomological Society of London, 2, 213 - 244.","Blackith, R. E. (1992) Tetrigidae (Insecta: Orthoptera) of South-East Asia: Annotated catalogue with partial translated keys and bibliography. Ashford Co., APAGA, Rockbottom, 248 pp.","Eades, D. C., Otte, D., Cigliano, M. M. & Braun, H. (2014) Orthoptera Species File Online. Version 5.0 / 5.0 Available from: http: // Orthoptera. SpeciesFile. org (accessed 10 April 2014)","de Haan, W. (1843) Bijdragen tot de kennis der Orthoptera. In: Temminck (Ed.), Verhandelingen over de Natuurlijke Geschiedenis der Nederlansche Overzeesche Bezittingen, de Leden der Natuurkundige Commissie in Indie en andere Schrijvers, 19 - 20, 165 - 228.","Willemse, C. (1928) Revision des Acridoidea, descrites par de Haan, avec description de nouvelles especes. Zoologische Mededelingen, 11, 1 - 27.","Gunther, K. (1938) Acrydiinen (Orthoptera, Acrididae) von Neu Guinea, hauptsachlich aus den Ausbeuten von Professor Dr. Burgers (Deutsche Kaiserin Augusta Fluss-Expedition 1912 - 13), Dr. E. Mayr (1928), G. Stein (1931) und Miss L. Cheesman (1933 - 1934). Nova Guinea, New Series, 2, 1 - 46.","Gunther, K. (1939) Revision der Acrydiinae (Orthoptera), III Sectio Amorphopi (Metrodorae Bol. 1887, auct.). Abhandlungen und Berichte aus den Staatlichen Museen fur Tierkunde und Volkerkunde in Dresden. Bd. 20. Reihe A: Zoologie, Neue Folge, 1 (1), 16 - 335.","Gunther, K. (1974) Beitrag zur Kenntnis der Tetrigoidea (Orthoptera, Caelifera) von Madagascar und von Mauritius. Bulletin du Musee d'Histoire Naturelle de Paris. 3 e Serie. Zoologie, 236, 937 - 1031.","Storozhenko, S. Yu. (2012) A new species of the genus Epitettix Hancock, 1907 (Orthoptera: Tetrigidae, Cladonotinae) from Vietnam. Russian Entomological Journal, 21 (2), 185 - 187."]}

Epitettix obtusus Storozhenko et Dawwrueng, 2014 http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName: 465257 Material examined. Seven specimens (holotype and paratypes). Distribution. Endemic to Thailand (provinces Nakhon Ratchasima and Chiang Mai) (Storozhenko & Dawwrueng, 2014)., Published as part of Storozhenko, Sergey Yu. & Dawwrueng, Pattarawich, 2015, New and little-known pygmy grasshoppers (Orthoptera: Tetrigidae) from Thailand, pp. 527-554 in Zootaxa 4052 (5) on page 532, DOI: 10.11646/zootaxa.4052.5.2, http://zenodo.org/record/290102, {"references":["Dawwrueng, P. & Doodduem, Ch. (2014) A new species of the genus Zhengitettix (Orthoptera: Tetrigidae, Scelimeninae) from Thailand. Far Eastern Entomologist, 282, 1 - 6."]}

Epitettix obtusus Storozhenko et Dawwrueng, sp. nov. Figs. 1���13 Material examined. Holotype���female, Thailand: Nakhon Ratchasima Province, environs of National park Khao Yai, 500���1000 m, 26 October��� 4 November 2000, coll. A. Gorochov and L. Anisyutkin (Zoological Institute of the Russian Academy of Sciences, St.-Petersburg). Paratypes: 3 males, same data as in holotype (Zoological Institute of the Russian Academy of Sciences, St.-Petersburg); 3 females, Thailand: Chiang Mai Province, Chiang Mai, Huai Hongkhrai Royal Development Study Center, 18 �� 47 ��� 2 ��� N, 98 �� 59 ��� E, 500���700 m, 22���24 June 2013, coll. P. Dawwrueng (Department of Entomology, Kasetsart University, Bangkok). Description. Female (holotype). Body robust, medium sized for genus. Antennae filiform, 16 -segmented; antennal bases situated on the level of lower margin of eyes. Length of antennae 1.4 times longer than length of fore femur; mid segments of antennae 4.5���5 times as long as wide. Eyes not protruding above vertex in lateral view. Lateral ocelli placed near the middle of eyes. Fastigium of vertex with deep concavities behind lateral carinae, slightly produced in front of eyes from dorsal aspect, 2 times wider than width of one eye from above; anterior margin of fastigium broadly rounded, median carina of fastigium short; lateral carinae elevated. Frontal ridge in lateral view rounded. Width of frontal ridge near base of antennae 1.5 times more than width of 1 st antennal segment. Pronotum rugulose, not reaching apex of abdomen; in dorsal view anterior margin of pronotum acute angular in the middle; posterior process of pronotum in dorsal view with rounded apex. Median carina of pronotum almost straight in profile; in cross-section carina broadly triangular. Lateral carinae of pronotum in prozona reduced. Hind margin of lateral lobes of pronotum with lower sinus only; posterior angles of lateral lobes in dorsal view truncated. Tegmina and hind wings absent. Fore and mid femora with almost straight and finely serrate lower and upper carinae. Fore femur 3.2 times, mid femur 3.5 times as long as wide. Hind femur stout, 2.2 times as long as wide; upper and lower carinae smooth; lower outer keel of hind femora without projections. Upper side of hind tibia with 6���8 outer and 4 inner teeth; numerous denticles are situated in basal part of outer side and especially in apical and basal parts of inner side of tibia. First tarsal segment of hind legs 1.4 times longer than 3 rd segment (without claws). Subgenital plate as long as wide, with strongly angularly projecting posterior margin. Cerci conical, 1.5 times as long as wide near base, with pointed apex. Valves of ovipositor narrow, dentate. Upper valve of ovipositor 3.8 times longer than its maximum width. Lower valve of ovipositor 6.6 times longer than its maximum width. Body brown, with few blackish and light brown spots on pronotum and legs. Head dark brown, with light brown spot situated above lateral ocelli and light brown stripes below eyes. Antennae light brown with shiny dark brown apex. Dorsal side and lateral lobes of pronotun light brown. Fore and mid femora dark brown with indistinct light spots. Fore and mid tibiae blackish, with three light rings. First segment of fore and mid tarsi brown, second segment with whitish base and black apex; claws brown. Hind femora light brown, lower side with black pregenicular spot; upper and lower carinae of hind femora with numerous black marks. Hind tibiae brown, with blackish ring near the middle. First and second segment of hind tarsus brown, third segment with light brown base and blackish apex; claws brown. Abdomen light brown with two blackish longitudinal stripes on the lateral sides of tergites. Ovipositor light brown. Male (paratypes). Similar to female. Antennae 15 -segmented; length of antennae 1.4���1.6 times longer than length of fore femur; mid segments of antennae 3.8���4 times as long as wide. Fastigium of vertex 2.0��� 2.1 times wider than width of one eye from above. Width of frontal ridge near the base of antennae 1.6���1.8 times more than width of 1 st antennal segment. Pronotum as in female. Tegmina and hind wings absent. Fore and mid femora as in female, but lower carina of femora slightly sinuate; fore femur 3.1���3.4 times, mid femur 3.1���3.2 times as long as wide. Hind femur 2.2���2.4 times as long as wide; upper and lower carinae smooth; lower outer keel of hind femora without projections. Upper side of hind tibia with 6���8 outer and 4���6 inner teeth and with numerous denticles situated as in female. First tarsal segment of hind legs 1.4���1.5 times longer than 3 rd segment (without claws). Epiproct narrow triangular, with pointed apex. Subgenital plate in lateral view short with pointed apex; in ventral view apex of plate excised. Cerci 1.5���1.7 times as long as wide, with pointed apex. There are two color forms. Body of two paratypes dark brown. Head blackish, without spots or stripes. Antennae colored as in female. Dorsal side and lateral lobes of pronotum dark brown. Fore and mid legs, hind tibiae and tarsi colored as in female. Hind femora blackish brown, with indistinct light marks; lower side completely black. Abdomen from above blackish with two light brown longitudinal stripes. Sternites brown, except blackish subgenital plate and apical sternite. Epiproct blackish brown, cerci brown. Third male differs from other paratypes in color of pronotum, hind femora and tibiae. In this specimen the lateral lobes of pronotum and anterior quarter of disk black, posterior 3 / 4 of pronotum light brown, with 5 small black spots situated around lateral carina; hind femora light brown like in female, but blackish pregenicular spot on lower side indistinct; hind tibiae light brown, without blackish ring near the middle. Measurements (in mm). Length of body (from frontal ridge to apex of subgenital plate) male 8.4���8.5, female 8.5���8.6; pronotum male 7.5���7.7, female 7.7���7.8; antenna male 2.3���2.5, female 2.5���2.6; fore femur male 1.6���1.7, female 1.7���1.8; mid femur male 1.9, female 2.1; hind femur male 5.1���5.5, female 5.5; ovipositor 1.4. Distribution. Thailand: provinces Nakhon Ratchasima and Chiang Mai. Diagnosis. The new species is similar to E. punctatus from Borneo and E. striganovae from Vietnam. It is distinguished from the former by acute angular anterior margin of pronotum (in E. punctatus anterior margin of pronotum truncate) and from the latter by median carina of fastigium reaching the anterior margin of eyes, the almost straight lower side of posterior process of pronotum, and the color of the hind tibiae (in E. striganovae median carina of fastigium reaching the mid of eyes, lower side of posterior process of pronotum sinuate, and hind tibiae black, with two light rings). The new species differs from E. tamilus, E. dammermanni, E. humilicolus and E. spheniscus by the absence of lateral carinae in the prozona (in all above mentioned species the prozona bears distinct lateral carinae), and similar in this respect to E. emarginatus, E. fatigans, E. lativertex and E. tumidus from New Guinea. However, these species are clearly distinguished from E. obtusus sp. nov. by their sinuate lower side of posterior process of pronotum and by the finely punctured pronotal integument (in E. obtusus sp. nov. the lower side of posterior process is almost straight and the pronotal integument is coarsely punctured). Habitats. New species was collected in the mountain mixed forests at elevation 500���1000 m in the moist place between fallen leaves. Etymology. The name of new species is originated from Latin adjective ��� obtusus��� (blunt)., Published as part of Storozhenko, Sergey Yu. & Dawwrueng, Pattarawich, 2014, Three new species of the subfamily Cladonotinae (Orthoptera: Tetrigidae) from Thailand, pp. 325-337 in Zootaxa 3811 (3) on pages 326-329, DOI: 10.11646/zootaxa.3811.3.3, http://zenodo.org/record/231482