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The Landrevus species group includes four Velarifictorus species that are related to Landrevinae crickets (Velarifictorus elephas Gorochov, 1992, Velarifictorus bubalus Gorochov, 1992, Velarifictorus gradifrons Ingrisch, 1998, and Velarifictorus landrevus Ma, Qiao & Zhang, 2019). A new species of the group is discovered in the Yunnan Province of China, and it is described and illustrated here. Velarifictorus yunnanensis Liu & Yin, 1993 is recognized as a junior synonym of Velarifictorus flavifrons Chopard, 1966. The morphological variety of V. flavifrons ectoparamere is documented and studied.

Genus Duolandrevus Kirby, 1906 Duolandrevus – Kirby, 1906: 50; Chopard, 1969: 147; Otte, 1988: 283; Gorochov, 1996: 35; Gorochov, 2016: 25 (key to subgenera) Type species. Gryllus brachypterus Haan, by original designation, Published as part of Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. & Robillard, Tony, 2022, New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah, pp. 201-228 in Zootaxa 5178 (3) on page 203, DOI: 10.11646/zootaxa.5178.3.1, http://zenodo.org/record/7026150, {"references":["Kirby W. F. (1906) A Synonymic Catalogue of Orthoptera. Vol. 2. Orthoptera Saltatoria. Part I. (Achetidae et Phasgonuridae). Trustees of the british Museum, London, 502 pp.","Chopard, L. (1969) Family Gryllidae: Subfamilies Mogoplistinae, Myrecophilinae, Scleropterinae, Cachoplistinae, Pteroplistinae, Pentacentrinae, Phalangopsinae, Trigonidiinae, Eneopterinae; Family Oecanthidae, Gryllotalpidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Vol. 12. Uitgeverij Dr. W. Junk N. V. ' s, Gravenhage, pp. 215 - 500.","Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334.","Gorochov, A. V. (1996) New and little known crickets from the collection of the Humboldt University and some other collections (Orthopera: Grylloidea). Part 2. Zoosystematica Rossica, 5 (1), 29 - 90","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97"]}

Duolandrevus (Duolandrevus) nobilis sp. nov. (Figs 3–6) Material examined. Holotype: EAST MALAYSIA • 1♂; Sabah State, Sabah State, Mount Silam, near Lahad Datu; N4.97667, E118.19135, 428.3± 7.9 m.a.s.l.; 13 May 2022, 21h36; on the ground along the trail; coll. M.K. Tan, T. Robillard & R. Japir; SBH.22.62 (FRC) Paratypes: • 1♂; Sabah State, Tabin Wildlife Reserve; N5.19438, E118.50294, 93.7±11.0 m.a.s.l.; 15 May 2022, 19h35; calling from a burrow on the clayey bank of a forest stream; coll. M.K. Tan, T. Robillard & R. Japir; SBH.22.84 (ZRC) • 1♂, 1♀; Sabah State, Mount Silam, near Lahad Datu, lowland forest; N4.97589, E118.19060, 341 m.a.s.l.; 13 May 2022, 21h10, video of male calling song (TR00022); male at the base of a large tree under the bark; female on the ground, near the entrance of burrow; coll. T. Robillard, M.K. Tan & R. Japir; TR22-12; (MNHNEO-ENSIF11147, MNHN-EO-ENSIF11148) Other material examined: EAST MALAYSIA • 1♀ juvenile; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N5.87395, E117.93871 (SAB14), 54 m. a.s.l.; 19 May 2022; coll. T. Robillard & M.K. Tan; TR22-39 (MNHN) Diagnosis. This new species differs from other species of the nominal subgenus Duolandrevus by the shape of the pseudepiphallus [epiphallus], metanotal gland and FW venation. Its male genitalia are most similar to Duolandrevus (Duolandrevus) kalimantan Gorochov, 2016 from Kalimantan; but the new species differs by FW with a larger harp area and more strongly curved oblique veins (instead of mostly straight), the metanotal gland more transverse; dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe], when viewed laterally, with the apex rounded (instead of subacute). It is also similar to Duolandrevus (Duolandrevus) brachypterus (Haan, 1844) from Java; but differs by the FW without a mirror area, more strongly curved oblique veins and the shapes of pseudepiphallic posteromedial lophi and pseudepiphallic lophi [posteromedial and posterolateral epiphallic lobes, respectively]. From other Bornean species of the subgenus: this new species differs from Duolandrevus (Duolandrevus) kubah Gorochov, 2016 and Duolandrevus (Duolandrevus) sabah Gorochov, 2016 by the absence of mirror on the FW and the dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe] broadly rounded (instead of slender or humped, respectively); from Duolandrevus (Duolandrevus) balikpapan Gorochov, 2016 and Duolandrevus (Duolandrevus) spinicauda Gorochov, 2016 by the pseudepiphallic lophi [posterolateral epiphallic lobe] not elongated and slender; from Duolandrevus (Duolandrevus) matang Gorochov, 2017 by the dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe] broadly rounded (instead of hooked). Remarks. This new species is placed under the nominal subgenus Duolandrevus according to these characters: the hindwings well surpassing metanotal gland; the male anal plate without a bundle of strong setae directed dorsad; the male genitalia with dorsal denticle at or near base of each pseudepiphallic lophi [posterolateral epiphallic lobe]. Etymology. The species name refers to the rarity of finding landrevine males forming burrows on the clayey bank of a forest stream. Often, landrevines call among cavity and crevices of tree trunk or branches. Description. Typical of the genus, body dorso-ventrally compressed and not pubescent, dark red-brown in colouration. Head rostrum not pubescent, about 1.7 times as wide as scape, with apex broadly rounded (in dorsal view) (Fig. 4A). Scapes brown (Fig. 4A). Eyes medium-sized, rather rounded (in profile view). Maxillary palpi yellow brown; with apical (fifth) segment yellow, longer than third and subapical (fourth) segments, slightly enlarged apically, with apex obtusely rounded; subapical segment, widens slightly apically, slightly longer than third segment (Fig. 4B). Head in anterior view slightly wider than tall (Fig. 4C). Lateral ocelli posterior of scapes oval; median ocellus between scapes small circular (Fig. 4C). Pronotum transverse, 1.7 times as broad as long; anterior margin as wide as posterior margin; dorsal disc with anterior margin lined with a row of setae and concave, and posterior margin feebly substraight (Fig. 4A). Pronotal lateral lobe about 1.3 times as long as wide; anterior half about 1.6 times as tall as posterior half, with ventral margin straight (Fig. 4B). TI with both tympana open and having oval tympanal membrane, inner one larger than outer one. TIII inner and outer margins with 4–5 stout articulated spurs (also known as movable spines) on each dorsal side; and 3 inner and ca. 9 outer basal spines, much smaller. TaIII with about 4 inner and 5 outer denticles. Male. FW glossy red-brown, harp area transparent (Figs 4D, 4E). Dorsal field not distinctly longer than lateral field, apical field stout (Fig. 4E). Venation (Figs. 4D, 4E): no distinct mirror present; harp area very large with 6 oblique veins (3 strongly curved long posterior veins and 3 substraight small anterior veins). Lateral field wide, R and M closely-spaced. Sc parallel to R and M, with four longitudinal and parallel branches; third vein (from dorsal) bifurcate in middle. Hind wings well surpassing metanotal gland (Fig. 4F). Metanotal gland with anterior margin with a row of strong setae, anterior lateral ends slightly swollen; two shallowly sunken divided from each other by a narrow longitudinal ridge; posterior margin broadly and faintly concave (Fig. 4F). Anal plate triangular with truncated apex, with setae along posterior margin but without bundle of strong setae directed dorsad (Fig. 5A). Male pseudepiphallus [epiphallus] (Figs 5B–D) notched in the middle, anterior half not raised. Pseudepiphallic posteromedial lophi [posteromedial epiphallic lobules] triangular, not elongated with obtuse apex; these lobules separated from each other by very shallow and narrow notch, appears fused together with a groove. Pseudepiphallic lophi [posterolateral epiphallic lobes] forming narrow plate-like projection, pointing dorsad at the apex (when viewed laterally); with a shorter and smaller projection pointing posteriorly (when viewed laterally); ventrally folded internally (when viewed dorsally or ventrally), also with a small projection pointing interno-posteriorly. Dorsal and posterior projections with apices bearing a few setae; apex of dorsal projection more broadly rounded, apices of posterior projections narrowly obtuse. Ectophallic fold [rachis] when viewed ventrally faintly surpassing pseudepiphallic posteromedial lophi, but not reaching the posterior end of pseudepiphallic lophi; very slender with acute apex. Each ectophallic apodeme [endparamere] forked posteriorly, and forming a Y-shape structure. Endophallic sclerite [formula] when viewed ventrally fairly stout, narrow anteriorly, widened and truncated at the posterior end. Rami not fused together by the anterior ends. Measurements (2♂, in mm). PronL = 2.7–2.9 (mean = 2.8); PronW = 4.6–5.3 (5.0); FWL = 7.1; FIIIL = 12.7; TIIIL = 8.9. Ecology. One of the males was found calling from a burrow on the clayey bank of a forest stream (Fig. 3). Another male was found on the ground along the trail. These suggest that this species is a more ground-dwelling landrevine. Distribution. Borneo, Sabah State: Mount Silam near Lahad Datu, Tabin Wildlife Reserve. Type locality. EAST MALAYSIA, Sabah State, Mount Silam near Lahad Datu. Calling song (1 ♂, in the field, 23.3°C) (Fig. 6). The calling song consists of an echeme made up of 32–48 syllables. The first few syllables are of lower amplitude but the amplitude of subsequent syllables increases to a maximum and remains relatively consistent. Each echeme has an average duration of 0.60±0.07 s (0.46–0.70 s). The interval between consecutive echemes is 5.01±1.84 s (3.19–8.47 s). Each syllable has an average duration of 9.1±0.5 ms (8.5–10.0 ms) and the interval between consecutive syllables is 5.6±0.6 ms (4.4–6.1 ms). The dominant frequency is 3.90±0.03 kHz (3.86–3.95 kHz)., Published as part of Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. & Robillard, Tony, 2022, New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah, pp. 201-228 in Zootaxa 5178 (3) on pages 206-210, DOI: 10.11646/zootaxa.5178.3.1, http://zenodo.org/record/7026150, {"references":["Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97","Gorochov, A. V. (2017) New data on Asiatic and Papuan crickets of the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 26 (2), 223 - 240."]}

Duolandrevus (Duolandrevus) nobilis sp. nov. (Figs 3–6) Material examined. Holotype: EAST MALAYSIA • 1♂; Sabah State, Sabah State, Mount Silam, near Lahad Datu; N4.97667, E118.19135, 428.3± 7.9 m.a.s.l.; 13 May 2022, 21h36; on the ground along the trail; coll. M.K. Tan, T. Robillard & R. Japir; SBH.22.62 (FRC) Paratypes: • 1♂; Sabah State, Tabin Wildlife Reserve; N5.19438, E118.50294, 93.7±11.0 m.a.s.l.; 15 May 2022, 19h35; calling from a burrow on the clayey bank of a forest stream; coll. M.K. Tan, T. Robillard & R. Japir; SBH.22.84 (ZRC) • 1♂, 1♀; Sabah State, Mount Silam, near Lahad Datu, lowland forest; N4.97589, E118.19060, 341 m.a.s.l.; 13 May 2022, 21h10, video of male calling song (TR00022); male at the base of a large tree under the bark; female on the ground, near the entrance of burrow; coll. T. Robillard, M.K. Tan & R. Japir; TR22-12; (MNHNEO-ENSIF11147, MNHN-EO-ENSIF11148) Other material examined: EAST MALAYSIA • 1♀ juvenile; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N5.87395, E117.93871 (SAB14), 54 m. a.s.l.; 19 May 2022; coll. T. Robillard & M.K. Tan; TR22-39 (MNHN) Diagnosis. This new species differs from other species of the nominal subgenus Duolandrevus by the shape of the pseudepiphallus [epiphallus], metanotal gland and FW venation. Its male genitalia are most similar to Duolandrevus (Duolandrevus) kalimantan Gorochov, 2016 from Kalimantan; but the new species differs by FW with a larger harp area and more strongly curved oblique veins (instead of mostly straight), the metanotal gland more transverse; dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe], when viewed laterally, with the apex rounded (instead of subacute). It is also similar to Duolandrevus (Duolandrevus) brachypterus (Haan, 1844) from Java; but differs by the FW without a mirror area, more strongly curved oblique veins and the shapes of pseudepiphallic posteromedial lophi and pseudepiphallic lophi [posteromedial and posterolateral epiphallic lobes, respectively]. From other Bornean species of the subgenus: this new species differs from Duolandrevus (Duolandrevus) kubah Gorochov, 2016 and Duolandrevus (Duolandrevus) sabah Gorochov, 2016 by the absence of mirror on the FW and the dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe] broadly rounded (instead of slender or humped, respectively); from Duolandrevus (Duolandrevus) balikpapan Gorochov, 2016 and Duolandrevus (Duolandrevus) spinicauda Gorochov, 2016 by the pseudepiphallic lophi [posterolateral epiphallic lobe] not elongated and slender; from Duolandrevus (Duolandrevus) matang Gorochov, 2017 by the dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe] broadly rounded (instead of hooked). Remarks. This new species is placed under the nominal subgenus Duolandrevus according to these characters: the hindwings well surpassing metanotal gland; the male anal plate without a bundle of strong setae directed dorsad; the male genitalia with dorsal denticle at or near base of each pseudepiphallic lophi [posterolateral epiphallic lobe]. Etymology. The species name refers to the rarity of finding landrevine males forming burrows on the clayey bank of a forest stream. Often, landrevines call among cavity and crevices of tree trunk or branches. Description. Typical of the genus, body dorso-ventrally compressed and not pubescent, dark red-brown in colouration. Head rostrum not pubescent, about 1.7 times as wide as scape, with apex broadly rounded (in dorsal view) (Fig. 4A). Scapes brown (Fig. 4A). Eyes medium-sized, rather rounded (in profile view). Maxillary palpi yellow brown; with apical (fifth) segment yellow, longer than third and subapical (fourth) segments, slightly enlarged apically, with apex obtusely rounded; subapical segment, widens slightly apically, slightly longer than third segment (Fig. 4B). Head in anterior view slightly wider than tall (Fig. 4C). Lateral ocelli posterior of scapes oval; median ocellus between scapes small circular (Fig. 4C). Pronotum transverse, 1.7 times as broad as long; anterior margin as wide as posterior margin; dorsal disc with anterior margin lined with a row of setae and concave, and posterior margin feebly substraight (Fig. 4A). Pronotal lateral lobe about 1.3 times as long as wide; anterior half about 1.6 times as tall as posterior half, with ventral margin straight (Fig. 4B). TI with both tympana open and having oval tympanal membrane, inner one larger than outer one. TIII inner and outer margins with 4–5 stout articulated spurs (also known as movable spines) on each dorsal side; and 3 inner and ca. 9 outer basal spines, much smaller. TaIII with about 4 inner and 5 outer denticles. Male. FW glossy red-brown, harp area transparent (Figs 4D, 4E). Dorsal field not distinctly longer than lateral field, apical field stout (Fig. 4E). Venation (Figs. 4D, 4E): no distinct mirror present; harp area very large with 6 oblique veins (3 strongly curved long posterior veins and 3 substraight small anterior veins). Lateral field wide, R and M closely-spaced. Sc parallel to R and M, with four longitudinal and parallel branches; third vein (from dorsal) bifurcate in middle. Hind wings well surpassing metanotal gland (Fig. 4F). Metanotal gland with anterior margin with a row of strong setae, anterior lateral ends slightly swollen; two shallowly sunken divided from each other by a narrow longitudinal ridge; posterior margin broadly and faintly concave (Fig. 4F). Anal plate triangular with truncated apex, with setae along posterior margin but without bundle of strong setae directed dorsad (Fig. 5A). Male pseudepiphallus [epiphallus] (Figs 5B–D) notched in the middle, anterior half not raised. Pseudepiphallic posteromedial lophi [posteromedial epiphallic lobules] triangular, not elongated with obtuse apex; these lobules separated from each other by very shallow and narrow notch, appears fused together with a groove. Pseudepiphallic lophi [posterolateral epiphallic lobes] forming narrow plate-like projection, pointing dorsad at the apex (when viewed laterally); with a shorter and smaller projection pointing posteriorly (when viewed laterally); ventrally folded internally (when viewed dorsally or ventrally), also with a small projection pointing interno-posteriorly. Dorsal and posterior projections with apices bearing a few setae; apex of dorsal projection more broadly rounded, apices of posterior projections narrowly obtuse. Ectophallic fold [rachis] when viewed ventrally faintly surpassing pseudepiphallic posteromedial lophi, but not reaching the posterior end of pseudepiphallic lophi; very slender with acute apex. Each ectophallic apodeme [endparamere] forked posteriorly, and forming a Y-shape structure. Endophallic sclerite [formula] when viewed ventrally fairly stout, narrow anteriorly, widened and truncated at the posterior end. Rami not fused together by the anterior ends. Measurements (2♂, in mm). PronL = 2.7–2.9 (mean = 2.8); PronW = 4.6–5.3 (5.0); FWL = 7.1; FIIIL = 12.7; TIIIL = 8.9. Ecology. One of the males was found calling from a burrow on the clayey bank of a forest stream (Fig. 3). Another male was found on the ground along the trail. These suggest that this species is a more ground-dwelling landrevine. Distribution. Borneo, Sabah State: Mount Silam near Lahad Datu, Tabin Wildlife Reserve. Type locality. EAST MALAYSIA, Sabah State, Mount Silam near Lahad Datu. Calling song (1 ♂, in the field, 23.3°C) (Fig. 6). The calling song consists of an echeme made up of 32–48 syllables. The first few syllables are of lower amplitude but the amplitude of subsequent syllables increases to a maximum and remains relatively consistent. Each echeme has an average duration of 0.60±0.07 s (0.46–0.70 s). The interval between consecutive echemes is 5.01±1.84 s (3.19–8.47 s). Each syllable has an average duration of 9.1±0.5 ms (8.5–10.0 ms) and the interval between consecutive syllables is 5.6±0.6 ms (4.4–6.1 ms). The dominant frequency is 3.90±0.03 kHz (3.86–3.95 kHz).

Duolandrevus (Bejorama) lambir Gorochov, 2017 (Figs 1, 2) Duolandrevus (Bejorama) lambir – Gorochov, 2017: 230; Tan & Wahab, 2018: 128 Duolandrevus (Bejorama) nr. luzonensis – Tan & Wahab, 2017: 442 [tentative identification prior to species described by Gorochov (2017)] Material examined. EAST MALAYSIA • 2♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N5.87511, E117.94206, 41.5± 7.7 m.a.s.l.; 17 May 2022, 19h22; calling underneath the bark of a large tree; coll. M.K. Tan & T. Robillard; SBH.22.121, 122 (FRC) Remarks. This species was previously known from Lambir Hill in Sarawak and Kuala Belalong in Brunei Darussalam, both to the west of Borneo. Our specimens (Fig. 1) appear to resemble the descriptions from Lambir Hill by Gorochov (2017) and the figures in Tan & Wahab (2017). It should be noted that the pseudepiphallic lophi [posterolateral epiphallic lobes] appear narrower in lateral view in the Bruneian specimen (Tan & Wahab, 2017: Fig. 2C). Here, we add a new locality record for this species in the east of Borneo and describe its calling song for the first time. Ecology. This species seems to be relatively common. Multiples males were heard singing at night underneath barks of large trees. Distribution. Borneo, Sarawak State (Lambir Hill) and Sabah State (Sepilok), Brunei Darussalam (Kuala Belalong). Type locality. EAST MALAYSIA, Sarawak State, Lambir Hill Calling song (2 ♂, in the field, 26.5°C) (Fig. 2). The calling song consists of an echeme made up of 40–47 syllables. The first few syllables are of lower amplitude but the amplitude of subsequent syllables increases to a maximum and remains consistent. Each echeme has an average duration of 0.66±0.03 s (0.61–0.71 s). The interval between consecutive echemes is 0.87±0.14 s (0.70–1.15 s). Each syllable has an average duration of 11.6±0.7 ms (10.9–12.9 ms) and the interval between consecutive syllables is 3.5±0.8 ms (2.1–4.3 ms). The dominant frequency is 5.89±0.06 kHz (5.72–5.91 kHz)., Published as part of Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. & Robillard, Tony, 2022, New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah, pp. 201-228 in Zootaxa 5178 (3) on page 203, DOI: 10.11646/zootaxa.5178.3.1, http://zenodo.org/record/7026150, {"references":["Gorochov, A. V. (2017) New data on Asiatic and Papuan crickets of the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 26 (2), 223 - 240.","Tan, M. K. & Wahab, R. A. (2018) Preliminary study on the diversity of Orthoptera from Kuala Belalong Field Studies Centre, Brunei Darussalam, Borneo. Journal of Orthoptera Research, 27 (2), 119 - 142. https: // doi. org / 10.3897 / jor. 27.24152","Tan, M. K. & Wahab, R. A. (2017) New taxa and notes on crickets of the subfamily Landrevinae (Orthoptera: Gryllidae) from Brunei Darussalam, Borneo. Zootaxa, 4365 (4), 440 - 454. https: // doi. org / 10.11646 / zootaxa. 4365.4.4"]}

Duolandrevus (Eulandrevus) gorochovi Zhang, Liu & Shi, 2017 (Fig 6–7) Duolandrevus (Eulandrevus) gorochovi Zhang, Liu & Shi, 2017: 316 Examined materials. 5 males. China: Guangdong, Nanling, Sep. 22, 2020, coll. He, Zhixin (SNNU); 2 males. China: Guangxi, Daming Mountain, Oct. 1, 2020, coll. He, Zhixin (SNNU); 1 male. China: Guangxi, Longzhou, Nongang, Aug. 22, 2020, coll. He, Zhixin (SNNU); 1 male. China: Guangdong, Nanling, Pingding Mountain, Aug. 6, 2019, coll. He, Zhixin and Zhang, Tao (SNNU). Measurements (mm). Male: BL 20.14±1.62; HL 3.25±0.20; HW 5.29±0.40; PL 3.25±0.16; PW 5.20±0.28; FWL 7.70±0.39; HFL 9.60±0.44; HTL 6.66±0.08; Female: BL 19.90; HL 3.84; HW 5.12; PL 3.44; PW 5.53; FWL 2.62; HFL 10.75; HTL 6.88; OL 13.97. Description. Male: Head smooth and light, conspicuously wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly wide in dorsal view. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between the two areas ridged. Antennal socket and lower side of compound eye depressed rough or wrinkled; genae shiny and plump. Median ocellus small and transversely ovoid; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion is short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third. End section of labial palpi longer than remainder sections, depressed and widened. Pronotum widened and depressed; anterior margin concave and arch-like, posterior margin straight. Middle of both lateral edges slightly convex. Hindwings aplasia, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching the fifth abdominal tergite. The basal region of forewing elevated and slightly higher than the rest of the forewing surface. Oblique veins five; diagonal vein forked, not connected to CuA vein. Chord veins three, they proximally connected together and converging on CuA veins; the third chord vein distally forked, connected to 3A veins. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord vein and wing cells connected by several veins (numbered from two to four). Apical field armed with reticulated veins. Lateral field bearing seven branches of Sc vein. Both inner and outer tympana ovoid, inner slightly larger than outer one. Hind tibiae equipped with spines on the half of proximal portion and spurs on distal part (numbered 4:4), and armed with five apical spurs, of them, the inner ones numbering three, and the ventral one shortest with remainder ones nearly equal to each other; and outer apical spurs numbering two and dorsal one slightly longer than ventral one; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with six spines respectively on sides. Super-anal plate simple, trapezoidal, about 2.3 times longer than width. Subgenital plate pike-liked. Genitalia (Fig. 7): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus short. Lateral lobes of epiphallus about three times longer than the middle lobe, its tip not bifurcate, but armed with a small hook on lateral side. Lateral lobes of epiphallus apex obtuse and hairy. Ectoparameres acute, about two times longer than epiphallic middle lobe. Coloration: Head, thorax and abdomen colored dark brown. Tegmina brown. Femur colored light brown, tibial and claw brownish. Cercus yellowish-brown proximally and apically brownish. Female: Similar to male. Forewing reaching anterior to the second abdominal tergite, with longitudinal veins, which shorter inside and gradually lengthening outward. And only the sixth longitudinal vein yellowish-brown remainders colored brown. Ovipositor smooth, brown, arrow-shaped and slightly longer than posterior femur. Remark. This species is related to Duolandrevus (Duolandrevus) Kalimantan (Gorochov, A.V., 2016), but the shape of the lateral lobes of the epiphallus and the shape of tegmina are different. Especially, ectoparamere of this species is obviously shorter than epiphallic lateral lobes. Size of body and spines on hind tarsus as well as number and size of wing cells which replace mirror varied among individuals of this species. Most literature records that this species does not possess hindwings (Zhang et al., 2017a). We discovered that it does but they are undeveloped and rather small (almost reaching posterior margin of metanotal gland), and entirely covered by forewings.

Duolandrevus (Eulandrevus) axinus Zhang, Liu & Shi, 2017 (Fig 3–4) Duolandrevus (Eulandrevus) axinus Zhang, et al., 2017: 316; Chen, et al., 2019: 553 Examined materials. 2 males. China: Hannan, Yinggeling, Aug. 22, 2019, coll. He, Zhixin (SNNU); 2 males. China: Hannan, Wuzhishan, Aug. 16, 2019, coll. He, Zhixin (SNNU). Measurements (mm). Male: BL 20.11±2.12; HL 3.48±0.53; HW 5.10±0.57; PL 2.82±0.24; PW 5.00±0.33; FWL 6.78±0.45; HFL 9.18±0.02; HTL 6.66±0.08. Description. Male: Head smooth and light, conspicuously wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly wider in dorsal view, almost equal to the width of antennal scape. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between two areas ridged; cheek shiny and plump. Median ocellus small and transversely ovoid; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion is short. Eyes ovoid, situated at middle of head side, about 1/3 length of head. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi as long as the third, depressed and widened, with rounded apex; end section of labial palpi longer than remainder sections, depressed and widened. Pronotum widened and depressed; anterior margin concave and arch-like, posterior margin straight, about 1.7 times as long as wide; middle of both lateral edges slightly convex. Hind wings aplasia, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching the fifth abdominal tergite. The basal region of forewing elevated and slightly higher than the rest of the forewing surface. Oblique veins five; diagonal vein forked, not connected to CuA veins. Chord veins three, second and third chord veins basally connected, converging on CuA veins; second or third chord vein distally forked, connected to CuA veins. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord vein and wing cells are connected by veins numbered from two to four. Apical field armed with reticulated veins. Lateral field bearing five branches Sc vein. Both inner and outer tympanum ovoid and the inner slightly larger than outer one. Hind tibiae equipped with spines on the one half of proximal portion and spurs on distal part (numbered 4:4); and with five apical spurs, of them, the inner numbering three, and the ventral one shorter than the dorsal two; and the outer apical spur numbering two, the ventral one shorter than the dorsal, but longer than 1/2 its length; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with six spines respectively on sides. Supper-anal plate trapezoid-like. Subgenital plate pike-like. Genitalia (Fig. 4): Genitalia tapering distallyin dorsal view. Middle lobes of epiphallus short and thick. Lateral lobes of epiphallus about two times longer than the middle lobe; its tip apically bifurcate and included angle between branches formed as flat angle, and each branch armed with acute apex, and the lower branch slightly longer than the upper one. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic middle lobe, and the posterior one about two times longer than the anterior. Ectoparameres acute, about two times longer than epiphallic middle lobe. Coloration: Body colored brown. Femora light brown. Tegmina surface yellowish brown, lateral field brown. Cercus yellow-brown proximally and apically brown. Female unknown. Remark. This species is similar to Duolandrevus (Eulandrevus) hainanensis Liu, He & Ma, 2015, but the lateral lobes of the epiphallus are different; in Duolandrevus (Eulandrevus) axinus, both anterior and posterior edge of epiphallus lateral lobe sharp, but in Duolandrevus (Eulandrevus) hainanensis, anterior edge of epiphallus sharp, posterior one obtuse. Body size as well as number and size of wing cells which replacing mirror are varied among individuals of this species. And most literature note that this species does not possess hind wings (Chen et al., 2019; Zhang et al., 2017a). After observing some specimens, we discovered it bears hind wings. These wings are undeveloped, rather small (slightly longer than posterior margin of metanotal gland) and entirely covered by forewings., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on pages 256-257, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564, {"references":["Liu, Y., He, Z. Q. & Ma, L. B. (2015) A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 4013 (4), 594 - 599. https: // doi. org / 10.11646 / zootaxa. 4013.4.9","Chen, L., Shen, C. Z., Tian, D., Wu, X., Li, K. & He, Z. - Q. (2019) A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae). Zootaxa, 4701 (6), 553 - 562. https: // doi. org / 10.11646 / zootaxa. 4701.6.4","Zhang, D. X., Liu, H. Y. & Shi, F. M. (2017 a) Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species. Zootaxa, 4317 (2), 310 - 320. https: // doi. org / 10.11646 / zootaxa. 4317.2.7"]}

Duolandrevus (Duolandrevus) bannanus Zhang, Liu & Shi, 2017 Duolandrevus (Duolandrevus) bannanus Zhang, et al., 2017: 590 Holotype information. Type locality: China, Yunnan, Xishuangbanna. Deposited at Museum of Hebei University (MHBU), Baoding, China (not examined). Remark. After observing some specimens of the six species of Eulandrevus, we found they have hind wings. But these are small and undeveloped (almost reaching posterior margin of metanotal gland), and entirely covered by the forewings. Therefore, we suggest other species of Eulandrevus might possess this feature. Regarding species of the subgenus Duolandrevus, their hind wings are conspicuously longer than metanotal gland, but the species D. (D.) bannanus which was originally placed in this subgenus does not possess this feature (whose hind wings not longer than metanotal gland).

Duolandrevus (Eulandrevus) gorochovi Zhang, Liu & Shi, 2017 (Fig 6–7) Duolandrevus (Eulandrevus) gorochovi Zhang, Liu & Shi, 2017: 316 Examined materials. 5 males. China: Guangdong, Nanling, Sep. 22, 2020, coll. He, Zhixin (SNNU); 2 males. China: Guangxi, Daming Mountain, Oct. 1, 2020, coll. He, Zhixin (SNNU); 1 male. China: Guangxi, Longzhou, Nongang, Aug. 22, 2020, coll. He, Zhixin (SNNU); 1 male. China: Guangdong, Nanling, Pingding Mountain, Aug. 6, 2019, coll. He, Zhixin and Zhang, Tao (SNNU). Measurements (mm). Male: BL 20.14±1.62; HL 3.25±0.20; HW 5.29±0.40; PL 3.25±0.16; PW 5.20±0.28; FWL 7.70±0.39; HFL 9.60±0.44; HTL 6.66±0.08; Female: BL 19.90; HL 3.84; HW 5.12; PL 3.44; PW 5.53; FWL 2.62; HFL 10.75; HTL 6.88; OL 13.97. Description. Male: Head smooth and light, conspicuously wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly wide in dorsal view. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between the two areas ridged. Antennal socket and lower side of compound eye depressed rough or wrinkled; genae shiny and plump. Median ocellus small and transversely ovoid; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion is short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third. End section of labial palpi longer than remainder sections, depressed and widened. Pronotum widened and depressed; anterior margin concave and arch-like, posterior margin straight. Middle of both lateral edges slightly convex. Hindwings aplasia, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching the fifth abdominal tergite. The basal region of forewing elevated and slightly higher than the rest of the forewing surface. Oblique veins five; diagonal vein forked, not connected to CuA vein. Chord veins three, they proximally connected together and converging on CuA veins; the third chord vein distally forked, connected to 3A veins. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord vein and wing cells connected by several veins (numbered from two to four). Apical field armed with reticulated veins. Lateral field bearing seven branches of Sc vein. Both inner and outer tympana ovoid, inner slightly larger than outer one. Hind tibiae equipped with spines on the half of proximal portion and spurs on distal part (numbered 4:4), and armed with five apical spurs, of them, the inner ones numbering three, and the ventral one shortest with remainder ones nearly equal to each other; and outer apical spurs numbering two and dorsal one slightly longer than ventral one; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with six spines respectively on sides. Super-anal plate simple, trapezoidal, about 2.3 times longer than width. Subgenital plate pike-liked. Genitalia (Fig. 7): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus short. Lateral lobes of epiphallus about three times longer than the middle lobe, its tip not bifurcate, but armed with a small hook on lateral side. Lateral lobes of epiphallus apex obtuse and hairy. Ectoparameres acute, about two times longer than epiphallic middle lobe. Coloration: Head, thorax and abdomen colored dark brown. Tegmina brown. Femur colored light brown, tibial and claw brownish. Cercus yellowish-brown proximally and apically brownish. Female: Similar to male. Forewing reaching anterior to the second abdominal tergite, with longitudinal veins, which shorter inside and gradually lengthening outward. And only the sixth longitudinal vein yellowish-brown remainders colored brown. Ovipositor smooth, brown, arrow-shaped and slightly longer than posterior femur. Remark. This species is related to Duolandrevus (Duolandrevus) Kalimantan (Gorochov, A.V., 2016), but the shape of the lateral lobes of the epiphallus and the shape of tegmina are different. Especially, ectoparamere of this species is obviously shorter than epiphallic lateral lobes. Size of body and spines on hind tarsus as well as number and size of wing cells which replace mirror varied among individuals of this species. Most literature records that this species does not possess hindwings (Zhang et al., 2017a). We discovered that it does but they are undeveloped and rather small (almost reaching posterior margin of metanotal gland), and entirely covered by forewings., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on pages 259-260, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564, {"references":["Zhang, D. X., Liu, H. Y. & Shi, F. M. (2017 a) Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species. Zootaxa, 4317 (2), 310 - 320. https: // doi. org / 10.11646 / zootaxa. 4317.2.7"]}

Duolandrevus (Eulandrevus) hainanensis Liu, He & Ma, 2015 (Fig 8–9) Duolandrevus (Eulandrevus) hainanensis Liu, et al., 2015: 596; Gorochov, 2016: 27; Zhang, et al., 314; Chen, et al., 2019: 553–562 Examined materials. 4 males. China: Hainan, Ledong, Jianfengling, Aug. 19, 2019, coll. He, Zhixin and Zhang, Tao (SNNU); 2 males. China: Hainan, Wuzhishan, Aug. 17, 2019 coll. He, Zhixin and Zhang, Tao (SNNU). Measurements (mm). Male: BL 20.39±2.55; HL 3.78±1.12; HW 5.47±0.20; PL 2.70±0.17; PW 5.25±0.15; FWL6.74±0.42; HFL 9.63±0.02; HTL 5.10±0.04; Female: BL 16.90±0.20; HL 3.18±0.12; HW 4.74±0.10; PL 2.95±0.10; PW 4.76±0.11; FWL 3.63±0.18; HFL 10.21±0.23; OL 9.62±0.18. Description. Male: Body sized large for this genus. Head smooth and shining, conspicuously wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, somewhat wide in dorsal view and wider than antennal scape. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between two areas ridged; cheek shiny and plump. Median ocellus small and transversely oval-like; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion is short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third; end section of labial palpi longer than remainder sections, depressed and widened. Pronotum widened and depressed; anterior margin concave and arch-like, posterior margin straight. Middle of both lateral edges slightly convex. Hindwings aplasia, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching middle of the fifth abdominal tergite. The basal region of forewing elevated and slightly higher than the rest of wing surface, with 1A and 2A veins converging apically. Oblique veins six. Chord veins three, proximally connected together, converging on CuA veins. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord and wing cells connected by several veins (from two to four). Apical field armed with reticulated veins. Lateral field bearing six branches of Sc vein. Both inner and outer tympanum oval-like and inner slightly larger than outer one. Hind tibiae equipped with spines on the half of proximal portion and spurs on distal part (numbered 4:5), and with six apical spurs, of them, the inner apical numbering three, the ventral one shortest and remainder ones nearly equal to each other, and the ventral one shorter than 1/2 of remainder ones; the outer apical spur numbering two, the ventral one shorter than the dorsal, but longer than 1/2 of them; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with six spines respectively on sides. Super-anal plate simple, trapezoidal, about two times longer than width. Subgenital plate pike-liked. Genitalia (Fig. 9): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus short. Lateral lobes of epiphallus about three times longer than the middle lobe, its tip not bifurcate, but in lateral view with a small hook on the dorsal side. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic lateral lobe, and the posterior one about 1.8 times longer than the former. Ectoparameres acute, about two times longer than epiphallic middle lobe. Coloration: Body color dark brown. Upper half portion of clypeus dark brown and lower half portion yellowish. Labrum yellowish-brown. Tegmina brown. Femora yellowish-brown, with tibial and claw brown. Cercus yellowishbrown proximally and apically dark brown. Female: Similar to male. Forewing reaching posterior to the second abdominal segment, with longitudinal veins, which short inside and gradually lengthening outward. Ovipositor brown, smooth and arrow-shaped, slightly longer than posterior femur. Remark. This species is related to Duolandrevus (Eulandrevus) axinus (Gorochov, 1988), but the lateral lobes of the epiphallus are different. In Duolandrevus (Eulandrevus) hainanensis, anterior edge of epiphallus sharp, posterior one obtuse; but in Duolandrevus (Eulandrevus) axinus, both anterior and posterior edge of epiphallus lateral lobe sharp. Size of body as well as number and size of wing cells replacing mirror are varied among individuals of species. The chord veins are different in this species. They proximally connect together in most individuals, and in some individuals, the third chord converges with wing cells that replacing mirror. Most literature record this species does not possess hindwings (Chen et al., 2019; Gorochov, 2016; Liu et al., 2015; Zhang et al., 2017a). We found it bears hindwings. These wings are undeveloped, rather small (slighter longer than posterior margin of metanotal gland) and entirely covered by forewings., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on pages 261-262, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564, {"references":["Liu, Y., He, Z. Q. & Ma, L. B. (2015) A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 4013 (4), 594 - 599. https: // doi. org / 10.11646 / zootaxa. 4013.4.9","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 25 (21), 23 - 97.","Gorochov, A. V. (1988) New and little-known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and certain other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), Fauna i ekologiya nasekomykh Vetnama [The fauna and ecology of insects of Vietnam]. Nauka Publishing House, Moscow, pp. 5 - 21.","Chen, L., Shen, C. Z., Tian, D., Wu, X., Li, K. & He, Z. - Q. (2019) A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae). Zootaxa, 4701 (6), 553 - 562. https: // doi. org / 10.11646 / zootaxa. 4701.6.4","Zhang, D. X., Liu, H. Y. & Shi, F. M. (2017 a) Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species. Zootaxa, 4317 (2), 310 - 320. https: // doi. org / 10.11646 / zootaxa. 4317.2.7"]}

Duolandrevus (Eulandrevus) longlouensis sp. n. (Figs. 1–2) Type material. Holotype. male. China: Hannan, Tongguling, Aug. 13, 2019, coll. He, Zhixin and Zhang, Tao (SNNU); Paratypes. 2 males and 2 females. China: Hannan, Tongguling, Aug. 13, 2019, coll. He, Zhixin and Zhang, Tao (SNNU). Measurements (mm). Male (n=3): BL 17.32±0.21; HL 2.71±0.07; HW 4.62±0.21; PL 2.53±0.02; PW 4.40±0.10; FWL 5.95±0.01; HFL 9.18±0.02; HTL 6.66±0.08; Female (n=2): BL 16.11±0.05; HL 2.77±0.01; HW 4.58±0.10; PL 2.91±0.05; PW 4.69±0.11; FWL 3.25±0.18; HFL 9.87±0.12; HTL 7.00±0.02; OL 9.62±0.18. Etymology. The specific epithet “ longlouensis ” refers to the town of collection locality of specimens. Diagnosis (Fig. 1). Body size small. Head slightly wider than pronotum. Tegmina reaching the fifth abdominal tergite. Basal region of forewing is elevated and slightly higher than the rest of forewing surface. Oblique veins four; the first oblique vein diverging laterally, one branch connected to basal region, the other linked to CuA vein. Mirror inconspicuous, replaced by multiple irregular cells; of them, most of proximal ones somewhat larger than the apical ones. The chord and nearby cells are connected by two veins. Apical field armed with reticulated veins (numerous cells). Middle lobes of epiphallus short and thick. Lateral lobes of epiphallus almost three times longer than the middle lobe; its tip apically bifurcate and included angle between branches formed as right angle; each branch armed with acute apex, and the lower branch about two times longer than the upper one. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic middle lobe, and the hind one about 1.5 times longer than the former. Ectoparameres acute apically, about two times longer than epiphallic middle lobe. Description. Male: Body sized small for this genus. Head smooth and light, slightly wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly wide dorsally viewed, almost equal to the width of antennal scape. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between two areas ridged; cheek shiny and plump. Median ocellus small and transversely ovoid; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion is short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third, depressed and widened, with rounded apex; end section of labial palpi longer than remainder sections, depressed and widened, with truncated tip. Pronotum flattened and widened; anterior margin concave and arch-like, posterior margin straight; middle of both lateral edges slightly convex. Hind wings undeveloped, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching the fourth abdominal tergite. The basal region of forewing elevated and slightly higher than the rest of the forewing surface. Oblique veins four, the first one diverging laterally, with one branch connected to the basal region and the other linked to CuA vein. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord vein and wing cells are connected by two transverse veins. Apical field armed with reticulated veins. Lateral field bearing five Sc branches. Legs densely hairy. Both inner and outer tympana oval-like and the inner slightly larger than outer one. Hind tibiae equipped with spines on proximally half portion and spurs on distal part (numbered 4:4); and with five apical spurs, of them, the inner numbering two and the ventral one shorter than the dorsal; and the outer apical spur numbering three, with the ventral one shortest and remainder ones nearly equal to each other; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with six spines respectively on sides; claw expanded proximally and possessing blunt teeth. Super-anal plate simple, trapezoid-like. Genitalia (Fig. 2): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus short and thick. Lateral lobes of epiphallus about three times longer than the middle lobe; its tip apically bifurcate and included angle between branches formed as right angle, and each branch armed with acute apex, and the lower branch about two times longer than the upper one. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic middle lobe, and the posterior one about 1.5 times longer than the former. Ectoparameres acute apically, about two times longer than epiphallic middle lobe. Coloration: Body color brown. Femora light brown. Tegmina surfaces yellowish brown, lateral field brown. Cercus yellow-brown proximally and apically brown. Female: Similar to male. Head vertex strongly inclined. Tegmina reaching the second abdominal tergite, with longitudinal veins, which shortened inside and gradually lengthening outward. Ovipositor brown, smooth and arrow-shaped, almost equal to the length of posterior femur. Remark. This species is very similar to D. (E.) obsidianu s (Chen et al., 2019) and D. (E.) dendrophilus in features of body and genitalia, but it has distinct epiphallus and ectoparameres. Shape of lateral lobes bifurcation and length of ectoparameres of new species is different from the others. Epiphallus lateral lobe branched as right angle, and posterior lateroabdominal projection of epiphallus short, apex not reaching lower branch of lateral lobe of epiphallus., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on pages 254-256, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564, {"references":["Chen, L., Shen, C. Z., Tian, D., Wu, X., Li, K. & He, Z. - Q. (2019) A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae). Zootaxa, 4701 (6), 553 - 562. https: // doi. org / 10.11646 / zootaxa. 4701.6.4"]}

Duolandrevus (Eulandrevus) longlouensis sp. n. (Figs. 1–2) Type material. Holotype. male. China: Hannan, Tongguling, Aug. 13, 2019, coll. He, Zhixin and Zhang, Tao (SNNU); Paratypes. 2 males and 2 females. China: Hannan, Tongguling, Aug. 13, 2019, coll. He, Zhixin and Zhang, Tao (SNNU). Measurements (mm). Male (n=3): BL 17.32±0.21; HL 2.71±0.07; HW 4.62±0.21; PL 2.53±0.02; PW 4.40±0.10; FWL 5.95±0.01; HFL 9.18±0.02; HTL 6.66±0.08; Female (n=2): BL 16.11±0.05; HL 2.77±0.01; HW 4.58±0.10; PL 2.91±0.05; PW 4.69±0.11; FWL 3.25±0.18; HFL 9.87±0.12; HTL 7.00±0.02; OL 9.62±0.18. Etymology. The specific epithet “ longlouensis ” refers to the town of collection locality of specimens. Diagnosis (Fig. 1). Body size small. Head slightly wider than pronotum. Tegmina reaching the fifth abdominal tergite. Basal region of forewing is elevated and slightly higher than the rest of forewing surface. Oblique veins four; the first oblique vein diverging laterally, one branch connected to basal region, the other linked to CuA vein. Mirror inconspicuous, replaced by multiple irregular cells; of them, most of proximal ones somewhat larger than the apical ones. The chord and nearby cells are connected by two veins. Apical field armed with reticulated veins (numerous cells). Middle lobes of epiphallus short and thick. Lateral lobes of epiphallus almost three times longer than the middle lobe; its tip apically bifurcate and included angle between branches formed as right angle; each branch armed with acute apex, and the lower branch about two times longer than the upper one. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic middle lobe, and the hind one about 1.5 times longer than the former. Ectoparameres acute apically, about two times longer than epiphallic middle lobe. Description. Male: Body sized small for this genus. Head smooth and light, slightly wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly wide dorsally viewed, almost equal to the width of antennal scape. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between two areas ridged; cheek shiny and plump. Median ocellus small and transversely ovoid; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion is short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third, depressed and widened, with rounded apex; end section of labial palpi longer than remainder sections, depressed and widened, with truncated tip. Pronotum flattened and widened; anterior margin concave and arch-like, posterior margin straight; middle of both lateral edges slightly convex. Hind wings undeveloped, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching the fourth abdominal tergite. The basal region of forewing elevated and slightly higher than the rest of the forewing surface. Oblique veins four, the first one diverging laterally, with one branch connected to the basal region and the other linked to CuA vein. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord vein and wing cells are connected by two transverse veins. Apical field armed with reticulated veins. Lateral field bearing five Sc branches. Legs densely hairy. Both inner and outer tympana oval-like and the inner slightly larger than outer one. Hind tibiae equipped with spines on proximally half portion and spurs on distal part (numbered 4:4); and with five apical spurs, of them, the inner numbering two and the ventral one shorter than the dorsal; and the outer apical spur numbering three, with the ventral one shortest and remainder ones nearly equal to each other; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with six spines respectively on sides; claw expanded proximally and possessing blunt teeth. Super-anal plate simple, trapezoid-like. Genitalia (Fig. 2): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus short and thick. Lateral lobes of epiphallus about three times longer than the middle lobe; its tip apically bifurcate and included angle between branches formed as right angle, and each branch armed with acute apex, and the lower branch about two times longer than the upper one. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic middle lobe, and the posterior one about 1.5 times longer than the former. Ectoparameres acute apically, about two times longer than epiphallic middle lobe. Coloration: Body color brown. Femora light brown. Tegmina surfaces yellowish brown, lateral field brown. Cercus yellow-brown proximally and apically brown. Female: Similar to male. Head vertex strongly inclined. Tegmina reaching the second abdominal tergite, with longitudinal veins, which shortened inside and gradually lengthening outward. Ovipositor brown, smooth and arrow-shaped, almost equal to the length of posterior femur. Remark. This species is very similar to D. (E.) obsidianu s (Chen et al., 2019) and D. (E.) dendrophilus in features of body and genitalia, but it has distinct epiphallus and ectoparameres. Shape of lateral lobes bifurcation and length of ectoparameres of new species is different from the others. Epiphallus lateral lobe branched as right angle, and posterior lateroabdominal projection of epiphallus short, apex not reaching lower branch of lateral lobe of epiphallus.

Duolandrevus (Eulandrevus) dendrophilus (Gorochov, 1988) (Fig 10–11) Eulandrevus dendrophilus Gorochov, 1988: 7 Duolandrevus (Eulandrevus) dendrophilus, Warchalowska-Sliwa, et al., 1997: 31; Kim & Pham, 2014: 62; Gorochov, 2016: 54; Zhang, et al., 2017: 313; Chen, et al., 2019: 553–562 Examined materials. 12 males. China: Sichuan, Chunzhou, Jiguanshan, Jul. 28, 2020, coll. He, Zhixin (SNNU); 11 males and 2 females. China: Sichuan, E’meishan, Aug. 4, 2020, coll. He, Zhixin (SNNU); 10 males and 8 females. China: Sichuan, Jinyunshan, Aug. 8, 2020, coll. He, Zhixin (SNNU); 1 male. China: Guangxi, Longzhou, Nongang, Sep. 22, 2020, coll. He, Zhixin and Zhang, Tao (SNNU); 5 males. China: Guangxi, Damingshan Jul. 28, 2019, coll. He, Zhixin (SNNU); 3 males. China: Hongkong, Dalanshan, May. 3, 2018, coll. Ma, Libin (SNNU); 10 males and 19 female female. China: Hongkong, Damaoshan, May. 9, 2018, coll. Ma, Libin (SNNU); 2 females. China: Hongkong, Ma’anshan, May. 14, 2018, coll. Ma, Libin (SNNU); 1 male. China: Shenzhen, Dapeng Peninsula National Geopark, Nov. 11, 2018, coll. Peng, Zhong (SNNU); 6 males. China: Guangdong, Nanling, Sep. 1, 2019, coll. He, Zhixin (SNNU); 2 males. China: Guangdong, Jiulong Lake, Oct. 31, 2014, coll. Zhang, Tao (SNNU). Measurements (mm). Male: BL 19.71±1.76; HL 3.50±0.58; HW 5.01±0.37; PL 3.50±0.84; PW 4.65±0.43; FWL 6.43±0.53; HFL 10.87±1.80; HTL 5.93±0.31; Female: BL 16.30±1.15; HL 2.88±0.05; HW 4.43±0.06; PL 2.71±0.02; PW 4.48±0.08; FWL 2.99±0.23; HFL 10.22±0.34; HTL 6.52±0.11; OL 8.58±0.51. Description. Male: Head smooth and shining, slightly wider than pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly broad in dorsal view and slightly wider than antennal scape. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between two areas ridged; cheek shiny and plump. Median ocellus small and transversely oval-like; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third; end section of labial palpi longer than remainder sections, depressed and widened. Pronotum widened and depressed; anterior margin concave and arch-like, posterior margin straight. Middle of both lateral edges slightly convex. Hind wings undeveloped, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching the middle of the sixth abdominal tergite. Oblique veins five. Chord veins three, basally connected and converging on CuA veins. Mirror inconspicuous, replaced by multiple irregular wing cells, of them, most of proximal ones somewhat larger than the apical ones. A chord vein and wing cells connected by two transvers veins. Apical field armed with reticulated veins. Lateral field bearing six branches of Sc vein. Both inner and outer tympana oval-like and the inner slightly larger than outer one. Hind tibiae equipped with spines on the half of proximal portion and spurs on distal part (numbered 4:5) and inner apical spurs longer than the outer ones. First tarsus of hind leg armed with spines respectively on sides. Super-anal plate simple, trapezoid-like. Subgenital plate pike-like. Genitalia (Fig. 11): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus short and thick. Lateral lobes of epiphallus about three times longer than the middle lobe; its tip apically bifurcate and included angle between branches formed as acute angle, and each branch armed with acute apex, and lower branches slightly shorter than upper one. Epiphallus ventrally possessing two lateral projections, the anterior one short and somewhat as long as epiphallic middle lobe, and the posterior one about 1.5 times longer than the anterior. Ectoparameres acute, about 1.5 times longer than epiphallic middle lobe. Coloration: Body dark brown. Lower part of fronto-clypeal suture proximally yellowish. Femur yellowishbrown. Cercus yellowish-brown proximally and apically brown. Female: Similar to male. Body colored brown. Forewing reaching posterior to the second abdominal segment, with longitudinal veins, which short inside and gradually lengthening outwards. Ovipositor brown, smooth and arrow-shaped, almost as long as posterior femur. Remark. This species is related to Duolandrevus (Eulandrevus) ivani (Gorochov, 1988), but is distinct in lateral lobes of the epiphallus which are bifurcate. In addition to the type locality of Vietnam, this species is widespread in the south of China. Size of body as well as size and number of wing cells that replacing mirror are varied among individuals in species. Most literature recorded this species do not possess hindwings (Chen et al., 2019; Gorochov, 1988, 2016; Kim & Pham, 2014; Warchalowska-Sliwa, Maryanska-Nadachowska, & Gorochov, 1997; Zhang et al., 2017a). However, we found it bears hind wings. These wings are undeveloped, rather small (almost reaching posterior margin of metanotal gland) and entirely covered by forewings., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on pages 263-264, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564, {"references":["Gorochov, A. V. (1988) New and little-known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and certain other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), Fauna i ekologiya nasekomykh Vetnama [The fauna and ecology of insects of Vietnam]. Nauka Publishing House, Moscow, pp. 5 - 21.","Kim, T. W. & Pham, H. T. (2014) Checklist of Vietnamese Orthoptera (Saltatoria). Zootaxa, 3811 (1), 53 - 82. https: // doi. org / 10.11646 / zootaxa. 3811.1.3","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 25 (21), 23 - 97.","Chen, L., Shen, C. Z., Tian, D., Wu, X., Li, K. & He, Z. - Q. (2019) A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae). Zootaxa, 4701 (6), 553 - 562. https: // doi. org / 10.11646 / zootaxa. 4701.6.4","Zhang, D. X., Liu, H. Y. & Shi, F. M. (2017 a) Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species. Zootaxa, 4317 (2), 310 - 320. https: // doi. org / 10.11646 / zootaxa. 4317.2.7"]}

Duolandrevus (Duolandrevus) bannanus Zhang, Liu & Shi, 2017 Duolandrevus (Duolandrevus) bannanus Zhang, et al., 2017: 590 Holotype information. Type locality: China, Yunnan, Xishuangbanna. Deposited at Museum of Hebei University (MHBU), Baoding, China (not examined). Remark. After observing some specimens of the six species of Eulandrevus, we found they have hind wings. But these are small and undeveloped (almost reaching posterior margin of metanotal gland), and entirely covered by the forewings. Therefore, we suggest other species of Eulandrevus might possess this feature. Regarding species of the subgenus Duolandrevus, their hind wings are conspicuously longer than metanotal gland, but the species D. (D.) bannanus which was originally placed in this subgenus does not possess this feature (whose hind wings not longer than metanotal gland)., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on page 265, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564

Duolandrevus (Eulandrevus) unguiculatus Ma, Gorochov & Zhang, 2015 (Fig 3, 5) Duolandrevus (Eulandrevus) unguiculatus Ma, et al., 2015: 446; Liu, et al., 2015: 598; Gorochov, 2016: 52; 2017: 228; Zhang, et al., 2017: 314; Chen, et al., 2019: 553–562 Examined materials. 1 male. China: Yunnan, Mengla, Wangtianshu, Aug. 6, 2017, coll. Ren, Guodong, Hou, Wenjun and Li, Yalin (MHBU). Measurements (mm). Male: BL 17.50; HL 3.64; HW 4.06; PL 2.53; PW 4.00; FWL 5.99; HFL 9.96; HTL 6.67. Description. Male: Body sized small for this genus. Head smooth and light, almost as wide as pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, slightly wider in dorsal view, and somewhat wider than antennal scape. Face transversely broad, area of antennal socket and area of lower side of eye distinctly depressed, between two areas ridged; cheek shiny and plump. Median ocellus small and transversely ovoid; lateral ocelli large and rounded. Clypeus extremely small, distal narrowed portion short. Labrum rhombus-like, with angle lateral margins and concave apically. End section of maxillary palpi longer than the third; end section of labial palpi longer than remainder sections, depressed and widened. Pronotum widened and depressed, about 1.7 times as long as wide; anterior margin concave and arch-like, posterior margin straight. Middle of both lateral edges slightly convex. Hind wings aplasia, small (reaching posterior margin of metanotal gland) and entirely covered by forewings. Tegmina reaching middle of the fifth abdominal tergite. Oblique veins five, the latter three veins basally connected, converging at the base of diagonal vein. Chord veins three, basally connected, converging on CuA veins. Mirror conspicuous, large, about 1.8 times wider than length and as long as base field. Apical field armed with reticulated veins. Lateral field bearing four branches of Sc vein. Both inner and outer tympanum oval-like and inner slightly larger than outer one. Hind tibiae equipped with spines on the half of proximal portion and spurs on distal part (numbered 4:4). And with five apical spurs, of them, the inner apical numbering three, and the ventral one shortest and remainder ones nearly equal to each other; as a whole, outer apical spurs longer than the inner ones. First tarsus of hind leg armed with two spines and an apical spur inside and outside respectively. Super-anal plate simple, trapezoidal. Subgenital plate acute. Genitalia (Fig. 5): Genitalia tapering distally in dorsal view. Middle lobes of epiphallus quite long, apical part bifurcate with claw-like tip. Lateral lobes of epiphallus about 2/3 length of middle lobe; its tip not bifurcate. Epiphallus ventrally possessing a lateral projection, about 1/3 length of middle lobe. Ectoparameres acute, about 2/3 length of epiphallic middle lobe. Coloration: Body brown. Tegmina surfaces yellowish-brown. Femur light brown. Female unknown. Remark. Male genitalia of this species are somewhat similar to those of Duolandrevus (Eulandrevus) sonorus, but in Duolandrevus (Eulandrevus) unguiculatus, middle lobes of epiphallus are much longer than in Duolandrevus (Eulandrevus) sonorus and in all the other known species of this subgenus. Most literature record this species as not having hind wings (Chen et al., 2019; Gorochov, 2016, 2017; Liu et al., 2015; Ma, Gorochov, & Zhang, 2015; Zhang et al., 2017a). After observing some specimens, we discovered it bears hind wings. However, these wings are very undeveloped, rather small (reaching posterior margin of metanotal gland.) and entirely covered by forewings., Published as part of Zheng, Yanna & Ma, Libin, 2021, Taxonomy of the genus Duolandrevus Kirby, 1906 (Orthoptera: Gryllidae; Landrevinae) from China with a new species of the subgenus Eulandrevus Gorochov, 1988, pp. 252-268 in Zootaxa 4942 (2) on pages 258-259, DOI: 10.11646/zootaxa.4942.2.7, http://zenodo.org/record/4600564, {"references":["Ma, L. B., Gorochov, A. V. & Zhang, Y. L. (2015) A new species of the genus Duolandrevus (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 3963 (3), 443 - 449. https: // doi. org / 10.11646 / zootaxa. 3963.3.8","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 25 (21), 23 - 97.","Gorochov, A. V. (2017) New data on Asiatic and Papuan crickets of the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 26 (22), 223 - 240.","Chen, L., Shen, C. Z., Tian, D., Wu, X., Li, K. & He, Z. - Q. (2019) A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae). Zootaxa, 4701 (6), 553 - 562. https: // doi. org / 10.11646 / zootaxa. 4701.6.4","Liu, Y., He, Z. Q. & Ma, L. B. (2015) A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 4013 (4), 594 - 599. https: // doi. org / 10.11646 / zootaxa. 4013.4.9","Zhang, D. X., Liu, H. Y. & Shi, F. M. (2017 a) Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species. Zootaxa, 4317 (2), 310 - 320. https: // doi. org / 10.11646 / zootaxa. 4317.2.7"]}

Duolandrevus (Eulandrevus) obsidianus He sp. nov. (Fig. 2E&F, 4 A–C, 5E) Holotype: male, CHINA, Guangxi, Baise, Napo, 20-iv-2019, coll. He Zhu-Qing. Paratypes: 3 males and 2 females, same data as holotype. Description. Fig. 2E. Head depressed dorsoventrally, slightly transverse, as wide as pronotum, rostrum almost equal to the width of antennal scape, eyes large but not convex, three ocelli nearly in one line with median one smaller, maxillary palps with distal three segments elongated and fifth segment little widened. Pronotum transverse and pubescent, almost 2 times as wide as long, lateral lobes narrow without lateral edge. Forewing short, reaching fifth abdominal tergite, five oblique veins curved with 3 longer ones jointed at base, mirror not well developed with reticulated veins instead, lateral field having 4 parallel longitudinal veins. Tympanum oval, on both inner and outer side of fore tibia with equal size, 5–9 proximal denticles with outer ones more than inner ones on hind tibia, and with 4–5 pairs of articulated spines, hind basitarsus with 5 pairs of dorsal denticles. Abdomen pubescent and cercus long, anal plate with widely rounded. Genitalia: Fig. 4, it is most similar to that of D. dendrophilus, but epiphallic lateral lobes are bifurcated apically in right angle. Moreover, the top of bifurcated lateral lobes is blunt and straight in the new species, while it is curved and acute in D. dendrophilus. Female. Similar to male. Forewing short, not as long as pronotum but touch each other in middle. Ovipositor straight (Fig. 2F). Coloration. Totally shinning black, with hind tibia reddish brown. Etymology. The name obsidianus is used to emphasize its color and texture similar to obsidian. Measurements (in mm). Male: body length (from head to anus) 21.04–22.09, pronotum length 3.11–3.15, forewing length 7.57–7.88, hind femur length 12.56–12.60; Female: body length 19.40–23.29, pronotum length 3.34–3.39, forewing length 2.49–2.67, hind femur length 13.06, ovipositor length 10.44–11.05. Distribution. China (Guangxi). Song. The song is continuous with 7.54± 0.61s cycle with 1.30±0.11 interval repeatedly. It includes 236±18.46 pulse in each chirp with pulse rate as 31.19±0.24 (Table 3., Fig. 5E).

Duolandrevus (Eulandrevus) obsidianus He sp. nov. (Fig. 2E&F, 4 A–C, 5E) Holotype: male, CHINA, Guangxi, Baise, Napo, 20-iv-2019, coll. He Zhu-Qing. Paratypes: 3 males and 2 females, same data as holotype. Description. Fig. 2E. Head depressed dorsoventrally, slightly transverse, as wide as pronotum, rostrum almost equal to the width of antennal scape, eyes large but not convex, three ocelli nearly in one line with median one smaller, maxillary palps with distal three segments elongated and fifth segment little widened. Pronotum transverse and pubescent, almost 2 times as wide as long, lateral lobes narrow without lateral edge. Forewing short, reaching fifth abdominal tergite, five oblique veins curved with 3 longer ones jointed at base, mirror not well developed with reticulated veins instead, lateral field having 4 parallel longitudinal veins. Tympanum oval, on both inner and outer side of fore tibia with equal size, 5–9 proximal denticles with outer ones more than inner ones on hind tibia, and with 4–5 pairs of articulated spines, hind basitarsus with 5 pairs of dorsal denticles. Abdomen pubescent and cercus long, anal plate with widely rounded. Genitalia: Fig. 4, it is most similar to that of D. dendrophilus, but epiphallic lateral lobes are bifurcated apically in right angle. Moreover, the top of bifurcated lateral lobes is blunt and straight in the new species, while it is curved and acute in D. dendrophilus. Female. Similar to male. Forewing short, not as long as pronotum but touch each other in middle. Ovipositor straight (Fig. 2F). Coloration. Totally shinning black, with hind tibia reddish brown. Etymology. The name obsidianus is used to emphasize its color and texture similar to obsidian. Measurements (in mm). Male: body length (from head to anus) 21.04–22.09, pronotum length 3.11–3.15, forewing length 7.57–7.88, hind femur length 12.56–12.60; Female: body length 19.40–23.29, pronotum length 3.34–3.39, forewing length 2.49–2.67, hind femur length 13.06, ovipositor length 10.44–11.05. Distribution. China (Guangxi). Song. The song is continuous with 7.54± 0.61s cycle with 1.30±0.11 interval repeatedly. It includes 236±18.46 pulse in each chirp with pulse rate as 31.19±0.24 (Table 3., Fig. 5E)., Published as part of Chen, Lin, Shen, Chu-Ze, Tian, Di, Wu, Xue, Li, Kai & He, Zhu-Qing, 2019, A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae), pp. 553-562 in Zootaxa 4701 (6) on pages 557-559, DOI: 10.11646/zootaxa.4701.6.4, http://zenodo.org/record/3562617

Subgenus Eulandrevus Gorochov, 1988 Type species: Eulandrevus sonorus Gorochov 1988, Published as part of Chen, Lin, Shen, Chu-Ze, Tian, Di, Wu, Xue, Li, Kai & He, Zhu-Qing, 2019, A new species of the subgenus Duolandrevus (Eulandrevus) Kirby, 1906 from Guangxi, China (Orthoptera: Gryllidae: Landrevinae), pp. 553-562 in Zootaxa 4701 (6) on page 557, DOI: 10.11646/zootaxa.4701.6.4, http://zenodo.org/record/3562617, {"references":["Gorochov, A. V. (1988) New and little known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and some other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), The Fauna and Ecology of Insects of Vietnam. Nauka Publishing House, Moscow, pp. 5 - 21."]}

Duolandrevus (Duolandrevus) bannanus sp. nov. (Figs. 2���7) http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:496078 Description. Male. Body medium, head short and slightly wider than pronotum (Fig. 2). Occiput somewhat convex, dorsal area of frontal rostrum flattened. Lateral ocelli rounded, located at side of frontal rostrum; median ocellus halfmoon-shaped, located at midpoint of frontal rostrum top. Frontal rostrum almost equal to the width of antennal scape. Eyes medium size, oval. Apical joint of maxillary palpi longer than the third, depressed and slightly widened. Apical joint of labial palpi longer than second, with rounded apex. Pronotum obviously transverse, middle of anterior margin slightly concave downward, almost equal to the width of posterior margin, posterior margin straight; midpoint of pronotum with a longitudinal concave, anterior margin of pronotum with more pubescent; lateral lobes trapezoidal, posterior margin slightly inclined, equipped with a ridge along margin; metanotal gland developed (Fig. 3), anterior margin of glandular fossa straight, obviously widened backward, with thick pubescent. Tegmen reaching 7th abdominal tergite, length about 1.4 times as long as width, with 5 oblique veins; mirror irregularly oblong, without dividing vein; apical area short, with irregular veins; lateral area with 6 oblique longitudinal veins. Wing short, almost reaching posterior margin of metanotal gland. Both inner and outer tympana present on fore tibia, oval-like, almost equal in size; hind tibia armed with small spines on the basal half portion and with 4 long spines respectively on both sides at the apical half portion. Supra anal plate, broad at base, posterior margin rounded (Fig. 4). Subgenital plate triangular; posterior narrowly, apical area arc-like. Genitalia (Figs. 5-7): epiphallus equipped widened, with pair of very small median lobes; lateral lobes developed, obviously narrowed to terminal, and slightly curved upward; epiphallic parameres obviously shorter than lateral lobes of epiphallus. Coloration. Body medium brown to dark brown, head and pronotum slightly darker. Tegmina pale brown, basal and apical portions respectively with a dark brown spot. Female. Unknown. Measurements (mm). Male: body 22.0, pronotum 2.9, tegmen 9.1, hind femur 11.9. Type material. Holotype. ♂, China, Yunnan, Xishuangbanna, 18. July, 2007, coll. Guo Zheng. Diagnosis. The new species is similar to D. (D.) thailandicus (Otte, 1988), but can be distinguished by anterior margin of metanotal gland narrower posterior margin; epiphallus widened, and apex of lateral lobes different shape; epiphallic parameres obviously short than the latter. Etymology. This species name is derived from the type locality on Banna (Xishuangbanna for short)., Published as part of Zhang, Dong-Xiao, Liu, Hao-Yu & Shi, Fu-Ming, 2017, First record of the subgenus Duolandrevus (Duolandrevus) (Orthoptera: Gryllidae: Landrevinae) from China, with description of a new species, pp. 589-592 in Zootaxa 4254 (5) on pages 590-591, DOI: 10.11646/zootaxa.4254.5.8, http://zenodo.org/record/556310, {"references":["Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334."]}

Duolandrevus (Bejorama) nr. luzonensis Otte, 1988 (Figs. 1, 2) Material examined. 1 male (KB.17.30): Brunei Darussalam, Ulu Temburong, Kuala Belalong Field Studies Centre, along Ashton Trail, primary ridge dipterocarp forest, calling inside hanging dead leaves on tree, N4.54629, E115.15684, 122.5 �� 6.8 m, 7 January 2017, 1954 hours, coll. M. K. Tan (ZRC). Remarks. Our single male specimen (Fig. 1A) belongs to the subgenus Bejorama Otte, 1988 on the basis of the distinct stridulatory apparatus (Fig. 1B), presence of strong setae along posterior edge of male anal plate (Fig. 1C), and lack of distinct transverse fold and dorsal denticle on epiphallus (Fig. 2). It has similar male genitalia and tegminal venation to that of Duolandrevus luzonensis Otte, 1988 from Luzon Island, Philippines as illustrated in Otte (1988). While this may indicate a new locality record for the Filipino species, there are however still some slight differences: our male specimen has a reduced anal area of the tegmen and with mirror much broader than long (Fig. 1B vs. Fig. 1E); posterolateral epiphallic lobe more slender (when viewed dorsally and laterally) and with ventral process more elongated (Fig. 2). Owing to the lack of further information, we tentatively identify the specimen to be close to D. luzonensis., Published as part of Tan, Ming Kai & Wahab, Rodzay Bin Haji Abdul, 2017, New taxa and notes on crickets of the subfamily Landrevinae (Orthoptera: Gryllidae) from Brunei Darussalam, Borneo, pp. 440-454 in Zootaxa 4365 (4) on page 442, DOI: 10.11646/zootaxa.4365.4.4, http://zenodo.org/record/1119233, {"references":["Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334 .."]}

Duolandrevus (Eulandrevus) kawataredoki, new species (Figs. 3���6) http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:500654 Material examined. Holotype (male): Brunei Darussalam, Ulu Temburong, Kuala Belalong Field Studies Centre, near Sungei Mata Ikan, primary ridge dipterocarp forest, calling inside hanging dead leaves on tree, N4.54727, E115.15701, 81.7 �� 9.1 m, 5 January 2017, 2003 hours, coll. M. K. Tan (KB.17.2) (IBER). Paratypes: 1 female (KB.17.23), Brunei Darussalam, same locality as holotype, along Ashton Trail, primary ridge dipterocarp forest, feeding on dead leaves on tree, N4.54686, E115.15715, 116.1 �� 7.2 m, 6 January 2017, 2044 hours, coll. M. K. Tan; 1 male (KB.17.91), same locality as holotype, along Ashton Trail, primary ridge dipterocarp forest, calling inside hanging dead leaves on tree, N4.54614, E115.15677, 120.5 �� 6.1 m, 27 July 2017, 2053 hours, coll. M. K. Tan (all ZRC). Subgeneric status. We consider this species to belong to the subgenus Eulandrevus based on the following diagnostic characters: hind wing absent, male anal plate unspecialized without strong setae at apex, and lack of dorsal denticle at and near base of each posterolateral epiphallic lobe. Although the epiphallus lacks distinct transverse fold (barely visible on lateral parts), the species may still be included in this subgenus with the absence of a single diagnostic character (Gorochov, 2016). Diagnosis. The new species differs from all known species of Duolandrevus (Eulandrevus) by the combination of the following characters: relatively small habitus; developed tegminal venation with mirror; metanotal gland transverse with two small deep oval depressions; posterolateral epiphallic lobe (when viewed dorsally) elongated and straight, pointing posteriorly, and with apex subacute, with internal process before posterior third of the lobe; posterolateral epiphallic lobe (when viewed laterally) roundly tapering into a narrowly truncated apex, without roundly angular dorsal projection; formula very short and stout. Comparison with congeners. The new species resembles closely to Duolandrevus (Eulandrevus) borneo Gorochov, 2016 from Sabah by male genitalia; but differs by tegmen reaching sixth abdominal tergite (instead of fourth), pronounced mirror on the tegmen (highly reduced in D. (E.) borneo); metanotal gland with two oval depressions (absence in D. (E.) borneo); and anal plate more truncated (instead of rounded) apically. The male genitalia differs by posterolateral epiphallic lobe more slender and elongated rounded, not angulated and without roundly angular dorsal projection (instead of stouter and roundly angular in D. (E.) borneo); internal process of posterolateral epiphallic lobe more elongated but less wide (in profile); formula (f) distinctly shorter. The new species is also similar to Duolandrevus (Duolandrevus) rufus Chopard, 1931 from Peninsular Malaysia and Duolandrevus (Duolandrevus) kubah Gorochov, 2016 from Sarawak by the male genitalia but differs by tegminal venation; the lack of dorsal denticle at and near base of each posterolateral epiphallic lobe and shape of internal process of posterolateral epiphallic lobe; and length of formula. The new species is also similar to Duolandrevus (Eulandrevus) tawau Gorochov, 2016 from Sabah, Duolandrevus (Duolandrevus) sabah Gorochov, 2016 from Sabah, and Duolandrevus (Duolandrevus) curup Gorochov, 2016 from Sumatra by the two oval depressions of the metanotal gland but differs by tegminal venation and genitalia. Description. Fairly small and stout cricket: cylindrical, slightly dorso-ventrally compressed and fairly pubescent (Fig. 3). Head rostrum distinctly wider than scapus, with apex truncated (in dorsal view) (Fig. 4A). Maxillary palpi with apical (fifth) segment elongated triangular, distinctly larger, broader, and longer than third and fourth segments, with apex swollen and obtusely rounded; subapical (fourth) segment slightly shorter than third segment (Fig. 4B). Pronotum transverse; dorsal disc gently widens posteriorly, slightly wider at the posterior end than long; anterior and posterior margins with strong hairs; disc with anterior and posterior margins straight (Fig. 4A); lateral lobe longer than tall; ventral margin of lateral lobe slightly concave in the middle (Fig. 4B). Fore tibia with small and rounded inner and outer tympana. Hind tibia inner and outer margins with 4 stout articulated spurs (also known as movable spines) on each dorsal side; and 2���3 much smaller basal spines; inner margin with two long apical spurs, outer margin with one shorter apical spur (still longer than subapical spurs. Hind basitarsus with 4 inner and 4 outer denticles. Tegmen not pubescent, extending to middle of 6th abdominal tergite (Figs. 3, 4B); dorsal field with mirror oblong (longer than broad) and not separated by a vein, two cord veins, and eight harp veins (three anterior ones are small); anal area truncated; lateral field with five longitudinal veins (Fig. 4C). Hind wings absent. Metanotal gland transverse, with two small oval depressions nearly separated in the middle; anterior margin feebly concave with long and strong hairs; posterior margin deeply concave in the middle (Fig. 4D). Ninth abdominal tergite transverse, posterior margin slightly convex in the middle (Fig. 4E). Anal plate (fusion of tenth abdominal tergite and epiproct) broad at basal half (wider than entire length), tapering to a slightly narrower apical half (about as wide as entire length, and wider than length of apical half); apex truncated with a few setae (Fig. 4E). Subgenital plate slightly broader than long, tapering gently into a rounded apex (Fig. 4F). Male genitalia as shown in Figs. 4G���J. Male epiphallus deeply and fairly widely notched, not distinctly folded transversely (when viewed dorsally) (Fig. 4G). Posteromedial epiphallic lobules (when viewed dorsally) directed posteriorly, slightly curved internally, and with almost acute apex; narrowly notched between the lobules (Fig. 4J). Posterolateral epiphallic lobe (when viewed dorsally) elongated and straight, pointing posteriorly and slightly towards each other, apex subacute, with internal process before posterior third of the lobe (Fig. 4G); (when viewed laterally) roundly tapering into a narrowly truncated apex, without roundly angular dorsal projection (Fig. 4I). Internal process of posterolateral epiphallic lobe points anteriorly with obtuse apex; not touching, overlapping, or fusing with each other; but nearly touching the posteromedial epiphallic lobules (when viewed dorsally) (Fig. 4G); elongated rounded (when viewed laterally) (Fig. 4I). Principal apodeme widely forked posteriorly. Formula very short and stout, posterior apex broad (when viewed dorsally) (Fig. 4G). Ramus not fusing together at the anterior end, with flattened and slightly membranous exterior plate near posterior third (Fig. 4G). Coloration. Generally red brown or brown. Head red brown; antenna scapus and antennal segments lighter brown. Gena red brown; clypeus maxillary palpi yellow brown with apical end of each segment white. Pronotum red brown. Tegmen red brown, but ventral half of lateral field transparent. Legs generally red brown, tibiae more of yellow brown. Hind femur mostly red brown, except ventro-internal area white; hind tibia generally red brown, dorsally dark brown apical of knee; spines on hind tibia red brown with black tips; hind tarsus brown; spines on hind tibia and tarsus brown with black tips. Thoracic segments white. Abdominal tergites and sternites (including subgenital plate) red brown; anal plate dark brown. Female (Fig. 5). Cylindrical, similar to male (Fig. 5A). Wingless (Fig. 5B). Metanotal gland absent (Fig. 5B). Ninth abdominal tergite transverse (Fig. 5C). Anal plate trapezoid, about as long as broad, flattened, tapering into a truncated apex; posterior margin with hairs (Fig. 5C). Subgenital plate also trapezoid broader than long, apex fairly narrow and truncated (Fig. 5D). Ovipositor relatively long, surpassing middle of hind tibia and nearly reaching apical spurs; slender, feebly thicken towards apical valves; dorsal valves slightly longer than ventral valves; valves with margin smooth and with apices acute (especially so for the dorsal valves) (Fig. 5E). Measurements (in mm). Male BL = 11.0���12.2; HL = 2.2���2.3; PL = 2.7���2.8; PW = 3.1���3.5; TL = 6.7���5.8; HFL = 8.1���8.4; HTL = 5.2���5.3. Female BL = 12.4; HL = 2.4; PL = 3.0; PW = 3.6; HFL = 9.1; HTL = 5.7; OL = 6.5. Male calling song (Fig. 6). The male produces trilling calls of various length (between less than 1 sec to slightly more than 2 sec). The calling song has a mean frequency of 5.0 kHz; mean dominant frequency of 4.6 kHz, maximum dominant frequency of 5.9 kHz, minimum dominant frequency of 3.2. kHz. Each thrill gets louder before plateauing; dominant frequency also increased from 3.8 kHz to 4.9 kHz. Etymology. This species is named after kawataredoki (noun in apposition), a Romanized form of the antiquated Japanese word, translating to time of being ���who he is��� dusk or twilight. This reflects the interesting encounters of the strange cricket during dusk., Published as part of Tan, Ming Kai & Wahab, Rodzay Bin Haji Abdul, 2017, New taxa and notes on crickets of the subfamily Landrevinae (Orthoptera: Gryllidae) from Brunei Darussalam, Borneo, pp. 440-454 in Zootaxa 4365 (4) on pages 442-449, DOI: 10.11646/zootaxa.4365.4.4, http://zenodo.org/record/1119233, {"references":["Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97"]}

3. Duolandrevus (Eulandrevus) guntheri (Gorochov, 1988) Eulandrevus guntheri Gorochov, 1988: 10; Yin & Liu, 1995: 76; Ichikawa, Murai & Honda, 2000: 274. Duolandrevus (Eulandrevus) guntheri, Gorochov, 2016: 27. http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:35294 Material examined. NOT seen. Distribution. China (Taiwan)., Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on page 314, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343, {"references":["Gorochov, A. V. (1988) New and little known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and some other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), The fauna and ecology of insects of Uietnam. Nauka Publishing House, Moscow, pp. 5 - 21. [in Russian]","Yin, H. S. & Liu, X. W. (1995) Synopsis on the classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific and Technological Literature Publishing House, Shanghai, 237 pp. [in Chinese]","Ichikawa, A., Murai, T. & Honda, E. (2000) Monograph of Japanese crickets (Orthoptera; Grylloidea). Bulletin of the Hoshizaki Green Foundation, 4, 257 - 332. [in Japanese]","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 25 (1), 23 - 97."]}

5. Duolandrevus (Eulandrevus) unguiculatus Ma, Gorochov & Zhang, 2015 Duolandrevus (Eulandrevus) unguiculatus Ma, Gorochov & Zhang, 2015: 446; Liu, He & Ma, 2015: 589; Gorochov 2016: 52. http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:470139 Material examined. Holotype ♂, Wangtianshu, Mengla, Yunnan, China, 6��� 7 August, 2007 collected by Guodong Ren, Wenjun Hou and Yalin Li. Distribution. China (Yunnan), Laos., Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on page 314, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343, {"references":["Ma, L. B., Gorochov, A. V. & Zhang, Y. L. (2015) A new species of the genus Duolandrevus (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 3963 (3), 443 - 449. https: // doi. org / 10.11646 / zootaxa. 3963.3.8","Liu, Y., He, Z. Q. & Ma, L. B. (2015) A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 4013 (4), 594 - 599. https: // doi. org / 10.11646 / zootaxa. 4013.4.9","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 25 (1), 23 - 97."]}

9. Duolandrevus (Eulandrevus) axinus sp. nov. (Figs 8, 11, 14, 18–21) http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:498561 Description. Male. Body medium (Fig. 8), occiput somewhat convex, frontal rostrum bluntly round, slightly wider than antennal scape, antennal scape shield-shaped. Lateral ocelli, round, located at the base of side rostrum; median ocellus like half moon, located at antennal fossa. Apical joint of maxillary palpi depressed, arc-shaped, slightly equal with the third joint; apical joint of labial palpi equal with second joint, depressed and flat, truncate. Eyes medium, oval, ventral margin with a lot of fine transverse striation, inside of the bottom corner slightly protruding. Pronotum obviously transverse, flat, with more pubescent. Anterior margin concave, posterior margin straight, about 1.7 times as long as wide. Metanotal gland developed, as in Fig. 14; central area of anterior margin and posterior margin concave; metanotal gland central area with few pubescent. Tegmina short, extending to base of 5th the abdominal tergite, present with 6 oblique veins, and without mirror, apical area with irregular crossvenation; with 6 longitudinal veins in lateral field; apical portions armed with reticulated veins. Fore tibiae with two tympana, oval, internal larger than external; hind tibiae armed with small spines on the basal half portion and with 4 long spines respectively on both sides at the apical half portion. Supra-anal plate short and trapezoid (Fig. 11), about 3.5 times as long as wide. Posterior margin straight, with bristle. Genitalia (Figs. 18–21): Epiphallus equipped with a pair of thick median lobes; lateral lobes wide and armed with two acute apex, axe-shaped; ectoparamere longer than lateral lobes. Female. Unknown. Coloration. Body brown. Head light, labrum and eye brown; ocelli light yellow. Tegmina copper brown. Measurements (mm). Body 16.8–21.9; pronotum 2.9–3.9; tegmen 6.2–8.0; hind femur 12.1–13.1. Type material. Holotype. ♂, Weng Village, Yuanmen, Baisha, Hainan, China, 400m, 20 May, 2008, collected by Yibin Ba and Juntong Lang. Paratypes. 1♂, Hongmao, Yuanmen, Baisha, Hainan, China, 29 May, 2007, collected by Yibin Ba and Juntong Lang; 1♂, Kaibo, Nankai, Baisha, Hainan, China, 12 June, 2007, collected by Yibin Ba and Juntong Lang; 1♂, Yinggezui, Baisha, Hainan, China, 23 May, 2014, collected by Junyan Su. Distribution. China (Hainan). Diagnosis. This new species is related to Duolandrevus (Eulandrevus) dendrophilus (Gorochov, 1988), but can be distinguished on the shape of the lateral lobes of the epiphallus, the lateral lobe of the epiphallus bifurcate; ectoparamere longer than lateral lobes. Etymology. This new specific name is derived from Greek axin, referring the lateral lobe of epiphallus with an axe-shaped apex, Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on pages 316-319, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343, {"references":["Gorochov, A. V. (1988) New and little known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and some other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), The fauna and ecology of insects of Uietnam. Nauka Publishing House, Moscow, pp. 5 - 21. [in Russian]"]}

4. Duolandrevus (Eulandrevus) hainanensis Liu, He & Ma, 2015 Duolandrevus (Eulandrevus) hainanensis Liu, He & Ma, 2015: 596; Gorochov, 2016: 27. http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:471458 Description. Female (Fig. 5). Body medium, Occiput somewhat protruding, as wide as pronotum. Lateral ocellus, oblong, located at base side of antennal scape; middle ocellus round, located at midpoint of frontal rostrum top. Frontal rostrum bluntly round, antennal scape semicircular, frontal rostrum wider than antennal scape, almost 1.2 times. Tegmina short, extending to base of the 3th abdominal tergite; with 7 parallel veins on dorsal surface, with 6 longitudinal veins in lateral field. Ovipositor long, sword-shaped, about 0.9 times as long as the length of hind femur. Measurements (mm). Body: 15.2; pronotum: 2.7; tegmen: 4.1; hind femur: 12.1; ovipositor: 11.0. Material examined. ♀, Jianfengling, Ledong, Hainan, China, 15 November, 2011, collected by Haoyu Liu. Remarks. This species was originally described from a male, and we identified a female specimen which is the same location as holotype. We identify it as Duolandrevus (Eulandrevus) hainanensis and describe it above. Distribution. China (Hainan)., Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on page 314, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343, {"references":["Liu, Y., He, Z. Q. & Ma, L. B. (2015) A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 4013 (4), 594 - 599. https: // doi. org / 10.11646 / zootaxa. 4013.4.9","Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 25 (1), 23 - 97."]}

6. Duolandrevus (Eulandrevus) infuscatus Liu & Bi, 2010 stat. nov. http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:498550 Duolandrevus (Duolandrevus) infuscatus Liu & Bi, 2010: 94. Material examined. NOT seen. Remark. According to the description and illustrations of this species, we referred it to Duolandrevus (Eulandrevus). Liu & Bi (2010) also regarded Duolandrevus (Duolandrevus) coriaceus is very similar to Duolandrevus (Duolandrevus) infuscatus, and apparently misplaced location. They conform to the characteristic of the subgenus Eulandrevus, so we put Duolandrevus (Duolandrevus) infuscatus into Duolandrevus (Eulandrevus). Distribution. China (Zhejiang)., Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on page 314, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343, {"references":["Liu, X. W. & Bi, W. X. (2010) Grylloidea (Gryllidae). In: Xu, H. C. & Ye, T. X. (Eds.), Insects of Fengyangshan National Reserve. China Forestry Publishing House, Beijing, pp. 92 - 97."]}

7. Duolandrevus (Eulandrevus) ziyunensis sp. nov. (Figs 6, 9, 12, 15) http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:498555 Description. Male. Body medium (Fig. 6), occiput somewhat convex, frontal rostrum smooth, as wide as antennal scape, antennal scape short, trapezoid, anterior margin somewhat convex. Lateral ocelli oval, located at the base of side frontal rostrum; median ocellus fissuriform, located at midpoint of frontal rostrum top. Eyes medium size, oval. Apical joint of maxillary palpi depressed, inclined, and longer than the third joint; apical joint of labial palpi longer than second joint, about 1.5 times as long as wide, depressed and truncate. Width of head subequal with anterior margin of pronotum; pronotum obviously transverse, flat, with more pubescent; anterior margin concave, posterior margin straight, about 1.5 times as long as wide. Metanotal gland developed, as in Fig.12; width of anterior margin of glandular fossa equal with posterior margin, central area with hair. Tegmina extending to base of 8th the abdominal tergite; with 5 oblique veins on dorsal surface; without mirror, apical area with irregular cross-venation; with 5 longitudinal veins in lateral field. Fore tibiae with two tympana, oblong, almost equal in size; hind tibiae armed with small spines on the basal half portion and with 4 long spines respectively on both sides at the apical half portion. Supra-anal plate simple (Fig. 9), trapezoid, about 2 times as long as wide, posterior margin arc-shaped, apex sunk. Genitalia (Fig. 15): Epiphallus equipped with a pair of small median lobes; lateral lobes wide and armed with somewhat squared apex, anterior margin of lateral lobes with a denticle curved upward; apical lobes of ectoparamere large and apically bluntly round in lateral view, lateral lobes with ectoparamere almost equal in length. Female. Unknown. Coloration: Head and pronotum black brown. Ocelli orange yellow; labrum and eye bright brown. Antenna, legs and tegmina copper yellow. Measurements (mm). Body:14.2; pronotum:3.1; tegmen: 7.5; hind femur 13.0. Type material. Holotype ♂, Ziyun, Fujian, China, 29 July, 2014, collected by Limei Li. Distribution. China (Fujian). Diagnosis. This new species is related to Duolandrevus (Eulandrevus) kalimantan, but it can be distinguished on the length of ectoparamere, the former lateral lobes with ectoparamere almost equal in length; anterior margin of lateral lobes with a denticle curved upward. Etymology. The specifc name is derived from the type locality Ziyun (China: Fujian)., Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on pages 315-316, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343

8. Duolandrevus (Eulandrevus) gorochovi sp. nov. (Figs 7, 10, 13, 16, 17) http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName:498559 Description. Male. Body slightly larger (Fig. 7), occiput flat, frontal rostrum bluntly round, slightly protruding, frontal rostrum wider than antennal scape, antennal scape almost trapezoid, anterior margin somewhat convex. Lateral ocellus, oval, located at the base of side rostrum; median ocellus, like half moon, located at midpoint of rostrum top. Eyes lager, ventral margin with a lot of fine transverse striation. Apical joint of maxillary palpi depressed, arc-shaped, and slightly longer than the third joint; apical joint of labial palpi 2 times as length as second joint, depressed and flat, truncate. Head slightly wider than pronotum, obviously transverse, pronotum with pubescent; anterior margin concave, posterior margin straight, about 2 times as long as wide. Tegmina short, extending to base of 5th the abdominal tergite; without mirror; with 5���7 oblique veins on dorsal surface and 5 longitudinal veins in lateral field; apical portions armed with reticulated veins. Metanotal gland developed, as in Fig.13; anterior margin of metanotal gland shorter than posterior margin (1: 1.3). Fore tibiae with two tympana, external tympanum oblong, interal tympanum oval; hind tibiae armed with small spines on the basal half portion and with 4 long spines respectively on both sides at the apical half portion. Supra-anal plate simple (Fig. 10), flat, trapezoid, about 2.3 times as long as wide. Posterior margin arc-shaped, with bristle. Genitalia (Figs. 16���17): Epiphallus equipped with a pair of long and thin median lobes; lateral lobes with a concavity, and divided into two small lobes; ectoparamere shorter than lateral lobes. Female. Unknown. Coloration. Head and pronotum dark brown. Antenna light brown; labrum and eye brown; ocellus light yellow. Tegmina and legs light brown. Measurements (mm). Body: 20.4���23.0; pronotum: 3.8���4.1; tegmen: 7.8���8.5; hind femur: 13.0���17.9. Type material. Holotype. ♂, Nanling, Ruyuan, Guangdong, China, 22 August, 2010, collected by Cuixi Du. Paratype: 1♂, the information same as holotype. Distribution. China (Guangdong). Diagnosis. This new species is related to Duolandrevus (Eulandrevu s) kalimantan, but can be distinguished on the shape of the median lobes of the epiphallus, the former median lobes of the epiphallus is long and thin; lateral lobes with a concavity, longer than ectoparamere. Etymology. This species is named after Dr. A. V. Gorochov, honoring this Russian entomologist., Published as part of Shi, Fu-Ming, 2017, Taxonomy of the subgenus Duolandrevus (Eulandrevus) Gorochov (Orthoptera: Gryllidae: Landrevinae: Landrevini) from China, with descriptions of three new species, pp. 310-320 in Zootaxa 4317 (2) on page 316, DOI: 10.11646/zootaxa.4317.2.7, http://zenodo.org/record/884343

Duolandrevus (Eulandrevus) unguiculatus Ma, Gorochov & Zhang Duolandrevus (Eulandrevus) unguiculatus Ma, Gorochov & Zhang, 2015: 446 Type specimen information. Type locality (holotype): Wangtianshu Park, Bubang, MenglaTai, Yunnan Province, China. Deposited at Museum of Hebei University, Baoding, Hebei Province, China (MHBU). Materials examined. China: Holotype: male, China, Yunnan Prov., Mengla, Wangtianshu (Bubang), coll. Ren Guodong, Hou Wenjun and Li Yalin, 6 ~ 7 August 2007 (MHBU). Distribution (Fig. 1). China (Yunnan). Measurements. BL 17.5, HW 4, PL 2.5, PW 4, FWL 6, HFL 10. Diagnosis. Body size small for most Landrevinae species, brown. Male tegmina with a small but distinct mirror in the stridulatory apparatus. Dorsal field of forewings somewhat square and armed with arc-like margin laterally. Both inner and outer fore tibiae tympanum present. Metanotal gland shiny and smooth for a quarter of apical portion and armed with long hairs for most of base portion. Ramiform branches of epiphallic median lobes armed with claw-like apex. Remarks. This species was reported recently, located in Yunnan Province, China. Its male genitalia are very similar to those of D. (E.) sonorus in the epiphallic median lobes, having long apical lobules. But, in D. (E.) unguiculatus, the latter lobules are much longer than in D. (E.) sonorus and in all the other species of this subgenus., Published as part of He, Zhuqing & Ma, Libin, 2015, A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China, pp. 594-599 in Zootaxa 4013 (4) on page 598, DOI: 10.11646/zootaxa.4013.4.9, http://zenodo.org/record/236809, {"references":["Ma, L. B., Gorochov, A. V. & Zhang, Y. L. (2015) A new species of the genus Duolandrevus (Orthoptera: Gryllidae: Landrevinae) from China. Zootaxa, 3963 (3), 443 - 449. http: // dx. doi. org / 10.11646 / zootaxa. 3963.3.8"]}

Genus Duolandrevus Kirby, 1906 Type species. Gryllus brachypterus Haan, 1842 (Java) Notes. This genus is a most usual genus for this tribe. It is widely distributed in Southeast Asia (excepting Indian Subcontinent and Sri Lanka I.) and the Papuan Region (New Guinea and adjacent islands). Duolandrevus is characterized by the following set of characteristics: body is slightly or significantly depressed dorsoventrally; head is wide but not high, with an angular or rounded rostral projection in the profile; pronotum is transverse, without ridges along lateral edges of the disc; male tegmina are shortened and with developed stridulatory apparatus, but sometimes this apparatus is slightly reduced or almost lost; female tegmina are from distinctly shortened to scale-like and always without stridulatory apparatus; hind wings are from strongly shortened to absent; male metanotal gland is developed; legs are strong but not long, with two (inner and outer) tympanal openings in the fore tibia (but sometimes these openings or one of them may be lost), with denticles or unarticulated spines situated proximad to the longer articulated spines of this tibia, and with laterally compressed (not widened) second tarsal segments; male genitalia are with a large but not very long epiphallus deeply bifurcated in the posterior part and having a pair of additional posteromedial lobules (sometimes almost fused with each other) between the larger or longer posterolateral lobes, with rather small and compact ectoparameres partly articulated or fused with the epiphallic posterolateral lobes, with normal endoparameres having a median bridge and rather long apodemes, with short and almost membranous or indistinct rachis (= guiding rod), with a small formula (= mold of spermatophore attachment plate), and with a distinctly sclerotized rami; ovipositor is more or less long, straight, and without drilling denticles on the apical part. Duolandrevus is divided into eight subgenera separated from each other morphologically and more or less geographically. These subgenera and some data on their distributions are given below, in a preliminary key to the Duolandrevus subgenera. However, this key may be significantly modified in the future because many new species must be discovered in the enormous unstudied or almost unstudied regions of Southeast Asia and on numerous islands between Asia and Australia, and some described species are in need of restudy, e.g. subgeneric position is quite uncertain in D. luzonensis Otte, 1988, D. balabacus Otte, 1988 and D. gingoogus Otte, 1988 from the Philippines as well as in D. palauensis Otte, 1988 from a small island situated between the Philippines and New Guinea (Otte, 1988)., Published as part of Ma, Libin, Gorochov, Andrej V. & Zhang, Yalin, 2015, A new species of the genus Duolandrevus (Orthoptera: Gryllidae: Landrevinae) from China, pp. 443-449 in Zootaxa 3963 (3) on page 445, DOI: 10.11646/zootaxa.3963.3.8, http://zenodo.org/record/234228, {"references":["Kirby, W. F. (1906) A Synonymic Catalogue of Orthoptera. Vol. II. Orthoptera Saltatoria Part I. (Achetidae et Phasgonuridae). Order of the Trustees of the British Museum, London, 502 pp.","Haan, W. D. (1842) Bijdragen tot de kennis der Orthoptera. In: Temminck, C. J. (Ed.), Verhandelingen over de natuurlijke geschiedenis der Nederlandsche overzeesche bezittingen. 16 Zoologie. Leiden, pp. 45 - 248.","Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334."]}

Duolandrevus (Eulandrevus) hainanensis sp. n. (Figs. 1, 2 A, 3 A, 3 C, 3 E) Type material. Holotype. male. China: Hainan, Jianfengling, Jul. 22, 2009, coll. He Zhuqing (SNNU). Measurements. BL 21.04, HW 5.43, PL 2.77, PW 4.91, FWL 6.78, MTL 4.58, HFL 12.59, HTL 8.75, CL> 6.35 (destroyed). Etymology. The specific epithet “ hainanensis ” refers to the location of specimen. Diagnosis. Large body size. Elytra shorter than uncovered abdomen portions. Six oblique veins (Fig. 2 A). Apical portions armed with reticulated veins. Both inner and outer tympanum present, oval shaped. Epiphallic lateral lobes somewhat squared apically, equipped with a denticle pointed backward (Fig. 3 C). Description. Body size large for this genus. Head large and transverse, conspicuously wider than pronotum. Occiput broad and somewhat flattened, slightly inclined to rostrum. Rostrum broad and flattened, almost equal to the width of antennal scape. Portions under antennal socket concave and flattened, and portions under eyes concave but wrinkled. Straight rostrum suture. Labrium rhombus-like, with angle lateral margins and concave apical margin. End section of maxillary palpi longer than the third, depressed and widened, with rounded apex. End section of labrial palpi longer than remainder basal sections, depressed and widened, with truncated tip. Median ocellus small and transverse oval-like; lateral ocelli large and rounded. Pronotum short, disk almost smooth, bearing few hairs; anterior margin concave and arc-like, posterior margin straight. Pronotal disk somewhat rounded and curved to lateral lobes without lateral edges. Lateral lobes vertically short, equipped with a ridge along margin and concave at hind corner beyond the ridge. Hind wings absent. Tegmina shorter than abdomen and with 6 oblique veins; apical portions bearing reticulated veins. Both inner and outer tympanum present on fore tibia, oval-like and inner slightly larger than the outer. Fore tibiae laterally depressed. Hind tibiae equipped with spines proximally and with spurs halfway on the proximal portion, numbered 4: 5 (inner: outer); inner apical spurs longer than the outer; ventral inner apical spur shortest, median outer apical spur longest. Abdomen armed with few hairs, almost smooth. Supra-anal plate simple, trapezoid-like with straight apex. Cercus robust, pubescent. Subgenital plate simple, fusiform, with acute apex. Genitalia: Epiphallus slightly widened and constricted apically, equipped with pair of small median lobes; lateral lobes longitudinally wide and armed with somewhat squared apex, with a denticle pointed backward. Top lobe of ectoparamere large and apically truncated in lateral view, with a rather small notch beyond the small bottom lobe. Coloration (Fig. 2 A). Body brown-colored. Antenna, legs, cercus and bottom of abdomen light-colored. FIGURE 3. Genitalia. A, C, E. D. (E.) hainanensis; B, D, F. D. (E.) hongkongae. A~B, dorsal view; C~D, lateral view; E~F, ventral view. Remarks. So far, all Eulandrevus species could be basically divided into two types from the shape of their epiphallic tips—simple rod-like and bifurcated. The new species is distinct from its relatives. Its epiphallic tip is simple but short, thick and somewhat square-like when laterally viewed, and also armed with a tooth pointed toward the head. Additionally, the new one is slightly larger than the previous, with longer elytra that reach the 5 th abdominal tergite (that of D. (E.) hongkongae is just extended to the apex of the 4 th tergite).

Duolandrevus (Eulandrevus) unguiculatus sp. nov. (Figs 2, A���C; Figs 3, A���D) http://lsid.speciesfile.org/urn:lsid: Orthoptera.speciesfile.org:TaxonName: 470139 Etymology. The specific epithet ��� unguiculatus ��� refers to the epiphallus having a pair of long and thin (spine-like) posteromedial lobules. Material studied. Holotype: male, China, Yunnan Prov., Mengla, Wangtianshu (Bubang), coll. Ren Guodong, Hou Wenjun and Li Yalin, 6���7 August 2007 (MHBU). Differential diagnosis. General appearance is as in representatives of the subgenus Eulandrevus having a rather small body, a more or less brown head without wrinkles on the face, male tegmina with a small but distinct mirror in the stridulatory apparatus, and fore tibia with both inner and outer tympana developed. Male genitalia of the new species are most similar to those of D. sonorus in the median epiphallic projection having long apical lubules, but in D. unguiculatus, the latter lobules are much longer than in D. sonorus and in all the other species of this subgenus. Description. Male. Anterior part of epicranium under rostral apex 0.93 times as high as antennal cavity and under eyes 0.5 times as high as this cavity; these parts of epicranium smooth, shine and without transverse wrinkles, but with small concave areas under antennal cavities; rostrum between antennal cavities almost as wide as scape. Pronotum distinctly transverse, 1.7 times as wide as long; its lateral lobes moderately narrow, 1.6 times as long as high. Metanotal gland as in Fig. 3 D. Tegmina reaching fifth abdominal tergite, with very short (narrow) and roundly truncate apical area of dorsal field, with rather short and somewhat oblique stridulatory vein, with moderately large and transverse mirror (approximately 1.8 times as wide as long) lacking dividing vein but having a few dividing veinlets in distal part, with wide harp area having seven oblique veins, with diagonal vein bifurcated as base, with strongly arcuate chords connecting with mirror by one crossvein and with diagonal vein by another crossvein, and with lateral field having 4���5 parallel longitudinal veins. Hind wings completely absent. Fore tibia with inner tympanum moderately large and ovoid, with outer tympanum slightly smaller and oval; hind tibiae dorsally with 4 inner and 4 outer articulated spines, and with 7 inner and 6 outer more proximal denticles (small unarticulated spines); hind basitarsus with 5 pairs of dorsal denticles. Supra-anal plate somewhat trapezoidal with straight posterior margin having several setae but lacking both distinct bundle of strong setae in dorsoapical part and dorsoapical denticle. Genitalia. Epiphallus: proximal epiphallic half wide and with a pair of distinct transverse folds on lateral parts of dorsal surface; distal epiphallic half distinctly narrower than proximal one, with a pair of rather large and angular (in profile) posterolateral lobes lacking denticles or strong setae, and with rather long median projection gradually narrowing backwards and having a pair of very long apical lobules slightly curved upwards but almost straight in distal half. Ectoparameres with rather large rounded lobe. Rachis (= guiding rod) rather large, semimembranous in median part but with a pair of lateral elongate (almost stick-like) sclerites; each of these sclerites with rather small rounded lateroproximal lobe and with comparatively thin but not acute apex. Formula small and rather narrow, with moderately long apodeme directed more or less forwards. Colouration. Head dark brown but with following pattern: dorsum of epicranium with 4 somewhat lighter longitudinal stripes; antennal scape, clypeus and labrum light brown with brown patches. Pronotum dark brown with anterior margin, middle part of disc, and lower part of lateral lobes with slightly lighter marks. Legs light brown, but hind femur with brown apical part. Tegmina: dorsal field light brown but having dark brown basal area and brown distal area; lateral field almost uniformly brown. Abdomen brown to dark brown, but its anterior part light brown, and third-seventh segments with small irregular light marks. Female unknown. Measurements. BL 17.5, HW 4, PL 2.5, PW 4, FWL 6, HFL 10., Published as part of Ma, Libin, Gorochov, Andrej V. & Zhang, Yalin, 2015, A new species of the genus Duolandrevus (Orthoptera: Gryllidae: Landrevinae) from China, pp. 443-449 in Zootaxa 3963 (3) on pages 446-448, DOI: 10.11646/zootaxa.3963.3.8, http://zenodo.org/record/234228

Duolandrevus (Eulandrevus) hainanensis sp. n. (Figs. 1, 2 A, 3 A, 3 C, 3 E) Type material. Holotype. male. China: Hainan, Jianfengling, Jul. 22, 2009, coll. He Zhuqing (SNNU). Measurements. BL 21.04, HW 5.43, PL 2.77, PW 4.91, FWL 6.78, MTL 4.58, HFL 12.59, HTL 8.75, CL> 6.35 (destroyed). Etymology. The specific epithet ��� hainanensis ��� refers to the location of specimen. Diagnosis. Large body size. Elytra shorter than uncovered abdomen portions. Six oblique veins (Fig. 2 A). Apical portions armed with reticulated veins. Both inner and outer tympanum present, oval shaped. Epiphallic lateral lobes somewhat squared apically, equipped with a denticle pointed backward (Fig. 3 C). Description. Body size large for this genus. Head large and transverse, conspicuously wider than pronotum. Occiput broad and somewhat flattened, slightly inclined to rostrum. Rostrum broad and flattened, almost equal to the width of antennal scape. Portions under antennal socket concave and flattened, and portions under eyes concave but wrinkled. Straight rostrum suture. Labrium rhombus-like, with angle lateral margins and concave apical margin. End section of maxillary palpi longer than the third, depressed and widened, with rounded apex. End section of labrial palpi longer than remainder basal sections, depressed and widened, with truncated tip. Median ocellus small and transverse oval-like; lateral ocelli large and rounded. Pronotum short, disk almost smooth, bearing few hairs; anterior margin concave and arc-like, posterior margin straight. Pronotal disk somewhat rounded and curved to lateral lobes without lateral edges. Lateral lobes vertically short, equipped with a ridge along margin and concave at hind corner beyond the ridge. Hind wings absent. Tegmina shorter than abdomen and with 6 oblique veins; apical portions bearing reticulated veins. Both inner and outer tympanum present on fore tibia, oval-like and inner slightly larger than the outer. Fore tibiae laterally depressed. Hind tibiae equipped with spines proximally and with spurs halfway on the proximal portion, numbered 4: 5 (inner: outer); inner apical spurs longer than the outer; ventral inner apical spur shortest, median outer apical spur longest. Abdomen armed with few hairs, almost smooth. Supra-anal plate simple, trapezoid-like with straight apex. Cercus robust, pubescent. Subgenital plate simple, fusiform, with acute apex. Genitalia: Epiphallus slightly widened and constricted apically, equipped with pair of small median lobes; lateral lobes longitudinally wide and armed with somewhat squared apex, with a denticle pointed backward. Top lobe of ectoparamere large and apically truncated in lateral view, with a rather small notch beyond the small bottom lobe. Coloration (Fig. 2 A). Body brown-colored. Antenna, legs, cercus and bottom of abdomen light-colored. FIGURE 3. Genitalia. A, C, E. D. (E.) hainanensis; B, D, F. D. (E.) hongkongae. A~B, dorsal view; C~D, lateral view; E~F, ventral view. Remarks. So far, all Eulandrevus species could be basically divided into two types from the shape of their epiphallic tips���simple rod-like and bifurcated. The new species is distinct from its relatives. Its epiphallic tip is simple but short, thick and somewhat square-like when laterally viewed, and also armed with a tooth pointed toward the head. Additionally, the new one is slightly larger than the previous, with longer elytra that reach the 5 th abdominal tergite (that of D. (E.) hongkongae is just extended to the apex of the 4 th tergite)., Published as part of He, Zhuqing & Ma, Libin, 2015, A new species of subgenus Eulandrevus Gorochov, 1988 (Orthoptera: Gryllidae: Landrevinae) from China, pp. 594-599 in Zootaxa 4013 (4) on pages 596-597, DOI: 10.11646/zootaxa.4013.4.9, http://zenodo.org/record/236809