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The genusCicadulaZetterstedtis reported for the first time from Pakistan with a newly recordedCicadulasimlaensisViraktamath & Yeshwanth. In addition,Platymetopius fidelis(Distant) is also rediscovered from Pakistan and studied here. A brief diagnosis along with the digital photographs of the habitus and male genitalia of bothspecies are provided. General distribution of these species in both Palaearctic and Oriental regions is also briefly discussed.

The leafhopper Tambocerus viraktamathi Rao is a newly recorded species in Pakistan. So far, only two species have been identified from the genus Tambocerus Zhang & Webb in Pakistan. Their distinguishing features, along with illustrations, are provided below.

Stirellus pakistanicus Shah & Duan sp. n. (Fig. 1A–J) Measurement. Male: 3.9mm. Description. Coloration. General color chocolate brown (Fig. 1A–D). Crown anterior portion black with two conspicuous black spots normally present in middle adjacent to eyes (Fig. 1A, 1C). Eye reddish brown, surrounded with yellow band (Fig. 1A–D). Face black with yellowish patch on frontoclypeus (Fig. 1D). Pronotum with black band along the anterior margin, whitish across the middle while yellowish brown at the posterior margin (Fig. 1A, 1C). Mesonotum and scutellum yellowish brown (Fig. 1A, 1C). Thorax and abdomen black in ventral and lateral view (Fig. 1B). Abdominal segments separated by yellowish bands (Fig. 1B). Forewing dark brownish with prominent venation (Fig. 1A–B). Morphology. Crown narrower than pronotum, with anterior margin slightly angulate and smoothly curved to face (Fig. 1A–D). Crown almost equal to or a little longer than breadth between eyes (Fig. 1A, 1C). Ocellus next to eye slightly below anterior margin of crown (Fig. 1B, 1D). Antennae long and thin. Face slightly longer than broad (Fig. 1D). Pronotum median length a little smaller than median length of crown with posterior margin slightly concave (Fig. 1A, 1C). Mesonotum and scutellum generally shorter than pronotum. Scutellum with a transverse impression in middle (Fig. 1C). Forewing macropterous (Fig. 1A–B). Male genitalia. Pygofer lobe broad basally with apex truncate (Fig. 1E). Pygofer side with few dorsal macrosetae (Fig. 1E). Subgenital plate large, triangular, with a row of uniseriate macrosetae (Fig. 1F). Valve triangular. Style narrow with digitate apophysis, laterally curved (Fig. 1G). Connective H-shaped with stem slightly shorter than arms (Fig. 1H). Aedeagal shaft long, bent at near base, nearly parallel to aedeagal base, with a slight dorsal curve at mid-length in lateral view, apex pointed and slightly curved dorsad, gonopore apical (Fig. 1I–J). Material examined. Holotype: Pakistan: Punjab: ♂, Attock, 33°51′4.2192′′ N, 72°15′59.292′′ E, sweeping hand net, 16 June 2021, coll. Waqar Khan and Bismillah Shah. Distribution. Pakistan [Attock (Fig. 2)]. Remarks. The new species described in this paper appears to key out to S. indrus (couplet 8) in our previous key (See Shah et al., 2020b). However it can readily be distinguished from S. indrus by the crown with anterior portion black (Fig. 1A, 1C); two conspicuous black spots normally present in middle adjacent to eyes (Fig. 1A, 1C); and face black with yellowish patch on frontoclypeus (Fig. 1D). Also, the new species has coloration similar to some species described from the Americas (e.g., Stirellus bicolor, Stirellus picinus) and Australia (Stirellus petfordensis). Etymology. This species is named for the country Pakistan, where the type specimen has been collected., Published as part of Shah, Bismillah, Khan, Waqar, Naveed, Hassan, Shakeel, Muhammad & Duan, Yani, 2022, A new species in the grassland leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan, pp. 344-350 in Zootaxa 5125 (3) on pages 346-347, DOI: 10.11646/zootaxa.5125.3.6, http://zenodo.org/record/6443920, {"references":["Shah, B., Naveed, H. & Duan, Y. N. (2020 b) A new species and four new records of Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species. Zootaxa, 4895 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 4895.3.5"]}

Stirellus rotundus (Singh-Pruthi, 1930) Distribution. Pakistan [Muzaffarabad (Fig. 2)], India., Published as part of Shah, Bismillah, Khan, Waqar, Naveed, Hassan, Shakeel, Muhammad & Duan, Yani, 2022, A new species in the grassland leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan, pp. 344-350 in Zootaxa 5125 (3) on page 347, DOI: 10.11646/zootaxa.5125.3.6, http://zenodo.org/record/6443920, {"references":["Singh-Pruthi, H. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68."]}

Shah, Bismillah, Khan, Waqar, Naveed, Hassan, Shakeel, Muhammad, Duan, Yani (2022): A new species in the grassland leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan. Zootaxa 5125 (3): 344-350, DOI: https://doi.org/10.11646/zootaxa.5125.3.6

Maiestas maculata (Singh-Pruthi) n. rec. (Fig. 2) Cicadula maculata Singh-Pruthi, 1930: 58���59, figs 80���81, plate V, fig. 2. Thamnotettix prabha Singh-Pruthi, 1930: 62, figs 85, 86, plate V, figs 6, 6a. Synonymised by Webb & Viraktamath, 2009: 41. Recilia prabha, Ghauri, 1980: 166���169, figs 1, 3���11. Deltocephalus (Recilia) maculata, Dash & Viraktamath, 1998: 32, figs 260���269. Maiestas maculata, Webb & Viraktamath, 2009: 22, comb. nov. Maiestas maculata, Zhang & Duan, 2011: 37���39, figs 33���35, plate IV: E, plate V: P, plate VI: P. Length. Male: 3.4mm Description. Coloration. Pale to yellowish brown (Fig. 2A���D). Crown with variable black spots (Fig. 2A, 2C). Face stramineous to yellowish brown, with black patches adjacent to antennal bases (Fig. 2D). Pronotum occasionally with black patches. Visible mesonotum with basal angles black (Fig. 2A, 2C). Legs with or without blackish spots (Fig. 2B). Morphology. Head as broad as pronotum (Fig. 2A, 2C). Forewing macropterous or submacropterous; inner anteapical cell open at base, sometimes outer anteapical cell absent (Fig. 2A���B). Male genitalia. Subgenital plate moderately long, subtriangular, lateral margin somewhat convex (Fig. 2F). Style with preapical lobe short, apophysis digitate, long and faintly sinuate (Fig. 2F). Connective almost as long as aedeagus (Fig. 2G). Aedeagus boat-shaped dorsally (Fig. 2H), straight in lateral aspect and narrow at base, broader in middle and thereafter gradually narrowed to acute upturned apex (Fig. 2I). Material examined. Pakistan: Azad Kashmir: 1♂, Banjosa, 33��48���36.9432 N, 73��49���14.304 E, sweep net, 26 July 2018, coll. Bismillah Shah (AAU). Distribution. Pakistan (new record), China, India. Remarks. This species, originally described from India (Calcutta and Sikkim) is recorded here for the first time from Pakistan. It can readily be distinguished from other species of the genus by its external appearance, e.g., with variable black patches on head and thorax, and shape of the aedeagus and style., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on pages 404-405, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Singh-Pruthi, H. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and Description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Ghauri, M. S. K. (1980) Illustrated redescription of two of Pruthi's species of Cicadelloidea from India. Reichenbachia Staatliches Museum fur Tierkunde in Dresden, 18, 165 - 171.","Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Zhang, Y. L. & Duan, Y. N. (2011) Review of the Deltocephalus group of leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae). Zootaxa, 2870 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2870.1.1"]}

Maiestas setosa (Ahmed, Murtaza & Malik) Recilia setosa Ahmed, Murtaza & Malik, 1988: 412, fig. 2. Maiestas setosa, Webb & Viraktamath, 2009: 20, comb. nov.; Naveed et al., 2019: 287. Material examined. No material examined. Distribution. Pakistan. Remarks. The identity of this species remains uncertain due to the poor original description and figures and because the type series from Karachi could not be found in the repository indicated in the original description, Zoological Museum, University of Karachi (Khatri & Webb, 2010: 11)., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 406, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Ahmed, M., Murtaza, B. & Malik, K. F. (1988) Some new aphrodine leafhoppers from grasslands of Karachi, Pakistan. Pakistan Journal of Zoology, 20, 409 - 421.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16","Khatri I., & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1"]}

Maiestas sp. 1 (Fig. 7) M. samuelsoni, Naveed et al., 2019, fig. 3L���M, misidentification. Material examined. Pakistan: 2♂♂, Khyber Pakhtunkhwa: Pirsabak, 34��0���37.1196 N, 72��3���3.6648 E, sweep net, 12 July 2018; 3♂♂, 4♀♀, same data, light trap; 2♂♂, Balakot, 34��34���18.1668 N, 73��21���46.6668 E, sweep net, 12 August 2019; Punjab: 1♂, Hazro, 33��54���44.208 N, 72��29���12.084 E, sweep net, 31 July 2018. All collected by Bismillah Shah (AAU). Remarks. The identity of this species is uncertain. It is similar to the following un-named species in having the forewings with additional cross-veins and similar male genitalia but its colour pattern is different. Its male genitalia are also similar to those of some other Maiestas species, e.g., M. samuelsoni (Knight). Although resembling the latter species (originally described from the Pacific region) in shape of aedeagus and style, it differs by its longer head with more distinct markings and forewing with extra cross-veins. It is likely that the material studied here is the same species recorded from Pakistan as M. samuelsoni by Naveed et al. (2019). These authors recorded three males (NWAFU) from the same region (Khyber Pakhtunkhwa) as the specimens examined here, but with the figures of Zhang & Duan (2011) from Chinese specimens, the latter also possibly misidentified. The Pakistan specimens deposited in NWAFU need to be re-examined to confirm their identity., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 412, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16","Zhang, Y. L. & Duan, Y. N. (2011) Review of the Deltocephalus group of leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae). Zootaxa, 2870 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2870.1.1"]}

Maiestas trispinosa (Dash & Viraktamath) n. rec. (Fig. 6) Deltocephalus (Recilia) trispinosus Dash & Viraktamath, 1998: 35, figs 296���304 Maiestas trispinosa, Webb & Viraktamath, 2009: 38, comb. nov. Length. Male: 4.1mm. Description. Coloration. Yellowish brown (Fig. 6A���D). Head and thorax pale yellow (Fig. 6A, 6C). Crown with six small, dark brownish marks at anterior margin and some yellowish brown patches on each side of mid-line (Fig. 6A, 6C). Pronotum with ochraceous shades and six yellow brown longitudinal stripes (Fig. 6A, 6C). Mesonotum and scutellum pale with yellowish brown spots (Fig. 6A, 6C). Eye tinged with yellowish brown (Fig. 6A���D). Frontoclypeus chocolate brown with pale transverse stripes (Fig. 6D). Leg marked with dark brown (Fig. 6B). Forewing light brown, with prominent pale venation (Fig. 6A���B). Morphology. Head slightly wider than pronotum, almost equal in length to width between eyes (Fig. 6A, 6C). Ocellus next to eye on anterior margin. Pronotum median length almost equal to the median length of crown (Fig. 6A, 6C). Forewing macropterous, almost same length as hind wings (Fig. 6A���B). Male genitalia. Pygofer quadrate (Fig. 6E). Subgenital plate moderately long, triangular, lateral margins almost straight, with a row of uniseriate macrosetae, apically rounded and curved dorsad (Fig. 6F). Valve triangular (Fig. 6F). Style narrow, with prominent preapical shoulder, apex digitate and sinuate (Fig. 6F). Connective almost equal in length to aedeagus (Fig. 6G���I). Aedeagal shaft extended far beyond pygofer, curved ventrally (Fig. 6E); shaft compressed dorsoventrally, in lateral view broader in basal two thirds then abruptly narrowed to acute apex, in dorsoventral view with needle-like apex (Fig. 6G���H); pair of subapical spine-like lateral processes adpressed to shaft (Fig. 6G���H). Material examined. Pakistan: Punjab: 2♂♂, Bansra Gali, 33��54���17.532 N, 73��22���13.9656 E, sweep net, 1 August 2019, coll. Bismillah Shah (AAU). Distribution. Pakistan (new record), India. Remarks. This species can be distinguished by the lateral spine-like processes of the aedeagus. The Pakistan specimens show very slight differences compared to the original Indian material with respect to the shape of the subgenital plates and style apical process, as seen in the original figures given by Dash & Viraktamath (1998) and images of type material sent by C. Viraktamath., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on pages 408-412, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Khatri I., & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Zhang, Y. L. & Duan, Y. N. (2011) Review of the Deltocephalus group of leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae). Zootaxa, 2870 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2870.1.1","Dash, P. C. & Viraktamath, C. A. (1995) Two new species of grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) from Orissa, India. Hexapoda, 7, 71 - 78."]}

Maiestas indica (Singh-Pruthi) (Fig. 1E) Allophleps indica Singh-Pruthi, 1936: 120, fig. 132; plate 9, fig. 3. Allophleps delhiensis Rao & Ramakrishnan, 1990: 111, figs 1���9. Synonymised by Dash & Viraktamath, 1998: 35. Deltocephalus (Recilia) indicus, Dash & Viraktamath, 1998: 35���36, fig. 305. Maiestas indica, Webb & Viraktamath, 2009: 21, comb. nov.; Naveed et al. (2019: 287). Material examined. No material examined. Distribution. Pakistan, India. Remarks. This species was originally described based on both male and female specimens but a female from Pakistan ���Layallpur [Faisalabad], Punjab, at light; 10.X.29 (Coll. A. Rahman)��� (ZSI), was listed as the ��� holotype ��� together with ���four other specimens from the type locality and numerous specimens from several other localities in the Indian Museum collection���. The specimens other than the female holotype may be considered paratypes but it is not clear from which specimen the male genitalia were figured and the male genitalia slide is now missing (pers. com. C. Viraktamath). Therefore, Pruthi���s figure of the male could be from a specimen either from Pakistan or India. No further material of this species from Pakistan has been studied since the original description and although the original material was examined by Dash & Viraktamath (1998), including a male paratype, their figures are from their Indian specimens (pers. com. C. Viraktamath). Despite the above uncertainties, assuming that the female holotype and the male specimen illustrated by Pruthi are conspecific, the identity of the species is clear based on the aedeagus with a distinctive large subapical ventral process (Fig. 1E). The species can also be recognized by its accessary forewing cross-veins, similar to M. pruthii and Maiestas sp. 1 and Maiestas sp. 2, but from these it also differs by its shorter head as shown in Pruthi���s original figure., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 403, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Singh-Pruthi, H. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11, 101 - 131.","Rao, V. R. S. & Ramakrishnan, U. (1990) New species of Aconurella and Paramesodes (Homoptera: Cicadellidae) from India. Journal of the Bombay Natural History Society, 87, 268 - 271.","Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16"]}

Maiestas sinuata Shah & Duan sp. n. (Fig. 3) Length. Male: 3.3mm. Description. Coloration. Yellowish brown (Fig. 3A–B). Head and thorax creamy white with yellow markings (Fig. 3A, 3C). Crown with eight small, dark brownish marks, with ochraceous patches on each side of median longitudinal suture (Fig. 3A, 3C). Pronotum with four yellow longitudinal stripes (Fig. 3A, 3C). Mesonotum and scutellum pale with yellowish brown spots (Fig. 3A, 3C). Eye color tinged with yellowish grey. Ocellus white (Fig. 3A–C). Frontoclypeus dark brown with pale transverse stripes (Fig. 3D). Legs marked with dark brown (Fig. 3B). Forewing brown, with prominent pale venation (Fig. 3A–B). Morphology. Head slightly wider than pronotum and slightly longer than width between eyes (Fig. 3A, 3C). Ocellus next to eye on anterior margin (Fig. 3A–C). Pronotum median length almost equal to the median length of crown (Fig. 3A, 3C). Forewing macropterous, exceeding abdomen when at rest (Fig. 3A–B). Male genitalia. Pygofer side longer than height in lateral aspect with rounded apical margin (Fig. 3E). Subgenital plate moderately long, triangular, with row of marginal uniseriate macrosetae (Fig. 3F). Style narrow, with prominent preapical shoulder; apex digitate, slightly curved laterally (Fig. 3F). Connective almost equal in length to aedeagus (Fig. 3G–H). Aedeagal shaft in dorsal view broad at base (Fig. 3G), gradually tapered towards pointed apex; in lateral view, slightly sinuate with basal half concave ventrally and apical half concave dorsally, finely tapered from base to apex (Fig. 3H). Material examined. Holotype ♂: Pakistan: Punjab, Kahuta, 33°35′28.3776 N, 73°22′12.234 E, sweep net, 25 July 2018, coll. Bismillah Shah (AAU). Distribution. Pakistan. Etymology. The species named for the slightly sinuate aedeagal shaft in lateral view. Remarks. This species has a similar sinuate aedeagal shaft to M. chandrai Fletcher & Dai (2019) (replacement name for M. viraktamathi Fletcher & Dai, 2018), from Australia, but differs from the latter species in external appearance and other genitalia structures, i.e., the subgenital plate with apical fine setae, style with apical process thicker and aedeagus lacking a ventrobasal “heel”, the latter feature placing M. chandrai in the M. albomaculatus group of Dash & Viraktamath (1998).

Maiestas Distant, 1917 Maiestas Distant, 1917: 312 ( Type species: Maiestas illustris Distant, 1917). For full details of synonymy, see Webb & Viraktamath, 2009: 14���15. Diagnosis. Generally pale brown with c rown often with line of submarginal angular spots and pronotum often with longitudinal stramineous bands (Fig 1A), sometimes without these markings (Fig. 2A). Crown triangularly produced, ocellus usually close to eye on anterior margin. Forewing with three anteapical cells, inner cell open. Pygofer without appendage. Subgenital plate short and ovate with outer margin convex to slightly concave; with uniseriate row of macrosetae laterally, sometimes with apical tuft of fine setae. Connective linear with arms longer than stem. Aedeagus fused to connective; shaft elongate, weakly curved dorsally, usually without processes, gonopore apical on dorsal surface, indistinct. Distribution. Old World. Remarks. Maiestas was revised by Webb & Viraktamath (2009). The genus is similar externally to Deltocephalus Burmeister and Recilia but differs by the shape of the aedeagus; in Maiestas the shaft is boat-shaped, at most weekly curved dorsally, sometimes with an apical ventral spine-like process and gonopore apical on dorsal surface. To the previous treatment of the genus from Pakistan (Naveed et al., 2019), a new species and two new country records are added in the present work, together with two species of uncertain identity (see Maiestas spp)., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 402, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Distant, W. L. (1917) The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr J. Stanley Gardiner, M. A. Vol. VI, No. VII-Rhynchota, Part II: Suborder Homoptera. Transactions of the Linnean Society of London, 17, 273 - 322. https: // doi. org / 10.1111 / j. 1096 - 3642.1917. tb 00469. x","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16"]}

Maiestas pruthii (Metcalf) (Fig. 4F���L) Deltocephlaus notatus Singh-Pruthi, 1936: 128���129, fig. 139, plate IX, fig. 10 (Prim.hom.: Deltocephalus notatus Melichar, 1896). Deltocephalus pruthii Metcalf, 1967: 1173 (nom. nov. pro Deltocephalus notatus Singh-Pruthi, 1936). Deltocephalus (Recilia) pruthii, Dash & Viraktamath, 1998: 22���23, figs 150���158. Maiestas pruthii, Webb & Viraktamath, 2009: 20, comb. nov.; Khatri & Webb, 2010: 11, plate 2a, fig. 13, misidentification? (see Remarks); Naveed et al., 2019: 286, figs 2A���C (incorrectly cited as M. subviridis, pers. com. Naveed), Fig. 3i, misidentification? (see Remarks). Material examined. No material examined. Distribution. Pakistan, India. (see Remarks) Remarks. The identity of this species is somewhat uncertain. It was described from the holotype male and allotype female from Pakistan, Layallpur [Faisalabad], but the male genitalia slide of the holotype (ZSI) is now missing (pers. com. Chandra Viraktamath). Although the types were examined by Dash & Viraktamath (1998) their figures are from Indian (Dehli) specimens (pers. com. Chandra Viraktamath) and although showing a very slight difference in the style apex to that of Singh-Pruthi���s original they show a similar relatively long apical extension of the aedeagus and rounded shaft apex (Figs 4J���L). Therefore, Dash & Viraktamath���s material is tentatively considered correctly identified but, to be certain, topotypical material of M. pruthii, i.e., from Faisalabad, should be obtained and examined. The Pakistan material identified as this species by Khatri & Webb (2010) with their figures reproduced by Naveed et al. (2019) show a more tapered aedeagus apex and shorter apical process and vertex with a small dot either side of the coronal suture apex, are here considered to be a different species (see Maiestas sp. 2 below). Despite the shortcomings of Singh-Pruthi���s original genitalia figures, his habitus figure of this species is very good and shows a species with a relatively long acute head with a pair of inverted U-shaped markings basally and forewings with extra cross veins. The extra forewing cross veins are also present in M. indica (described from the same type locality as M. pruthii) and Maiestas sp. 1 and 2 (see Remarks under these species). In conclusion, until the type genitalia or topotypical specimens can be found, the identity of this species is based on the figures of Dash & Viraktamath (1998) together with the images given here from the same specimen (Figs 4F���L)., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 406, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Singh-Pruthi, H. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11, 101 - 131.","Melichar, L. (1896) Cicadinen (Hemiptera-Homoptera) von Mittel-Europa. Dames, Berlin, 364 pp. https: // doi. org / 10.5962 / bhl. title. 8568","Metcalf, Z. P. (1967) s. n. In: General Catalogue of the Homoptera. Fascicle VI. Cicadelloidea. Part 10. Euscelidae Section II. Agricultural Research Service, U. S. Department of Agriculture, Washington, D. C., 1078 - 2074.","Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Khatri I., & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16"]}

Stirellus capitatus (Distant) n. rec. (Fig. 1) Viridomarus capitatus Distant, 1918: 70. Viridomarus brevialatus Xing, Dai & Li, 2009: 590. Viridomarus laticellus Xing, Dai & Li, 2009: 592. Stirellus capitatus, Li, Dai & Xing, 2011: 271. Stirellus capitatus, Duan, Webb & Zhang, 2016: 108. Measurement. Male: 3.0���4.0mm; female: 4.6mm. Description. Coloration. Stramineous. The vertex lateral margins generally pale stramineous (Fig. 1 A���C); vertex tip with small blackish spot anteriorly (Fig. 1A, 1C). Morphology. Crown longer than twice breadth between the eyes, elongately subtriangular, tapering to apex and widely subacute (Fig. 1 A���C). Ocellus on anterior margin far from eye. Eye comparatively small (Fig. 1 A���C). Anteclypeus narrowing to apex, not exceeding beyond normal curve of genae. Pronotum almost as long as 0.5 times crown median length (Fig. 1 A���C). Mesonotum and scutellum shorter than pronotum (Fig. 1 A���C). Forewing macropterous to submacropterous, generally exposing abdomen apex in female and occasionally in male (Fig. 1 A���B). Male genitalia. Pygofer side with a small number of macrosetae present dorsally, apex slightly angulate (Fig. 1F). Subgenital plate highly convex laterally, macrosetae almost uniseriate centrally (Fig. 1G). Style apophysis digitate, curved laterally (Fig. 1J). Connective with stem shorter than arms (Fig. 1I). Aedeagal shaft long, strongly bent dorsally near base, tip sharp, somewhat curved dorsad, gonopore apical (Fig. 1K). Material examined. Pakistan: Khyber Pakhtunkhwa: 1♀, Jalbai, 34��0���43.8444 N, 72��16���0.0228 E, sweeping hand net, 19 July 2019, coll. Bismillah Shah. Distribution. Pakistan, China, India, Thailand. Remarks. This species can be recognized by its comparatively long head, and by the tip of vertex with small dark brown spot. This is the first record of this species from Pakistan., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 401, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Xing, J. C., Dai, R. H. & Li, Z. Z. (2009) A taxonomic study on the genus Viridomarus Distant (Hemiptera: Cicadellidae: Euscelinae). Acta Zootaxonomica Sinica, 34 (3), 590 - 595.","Li, Z. Z., Dai, R. & Xing, J. (2011) Deltocephalinae from China (Hemiptera: Cicadellidae). Popular Science Press, Beijing, 336 pp.","Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus indrus (Distant) n. rec. (Fig. 2) Typhlocyba indra Distant, 1908: 415. Aconura bipunctella Matsumura, 1914: 232. Paivanana indra, Distant, 1918: 95. Paivanana centristriata Dai & Li, 2003 in Dai, Chen & Li, 2003: 31. Stirellus indrus, Li, Dai & Xing, 2011: 273; Duan, Webb & Zhang, 2016: 116. Measurement. Male: 2.9mm; female: 3.6mm. Description. Coloration. Normally pale greyish (Fig. 2 A���C). Crown with two conspicuous black spots normally between anterior margins of eyes, and to some extent adjacent to eyes than each other (Fig. 2A, 2C). Face having two blackish round marks on each side below the antennal sockets (Fig. 2D). Forewing with longitudinal brown bands, tinged with purple or green color (Fig. 2 A���B). Morphology. Head subconical, nearly subequal in width to pronotum (Fig. 2A, 2C), with anterior margin smoothly curved to face, acutely angled (Fig. 2D). Crown somewhat longer than breadth between eyes (Fig. 2A, 2C). Ocellus next to the eye at anterior margin. Eye comparatively large (Fig. 2 A���D). Anteclypeus tapering towards apex, exceeding beyond the normal curve of genae. Pronotum median length almost equal to the median length of crown (Fig. 2A, 2C). Mesonotum and scutellum smaller than pronotum. Forewing macropterous, normally exposing the apex of abdomen in females and occasionally in males (Fig. 2 A���B). Male genitalia. Pygofer side with few dorsal macrosetae, almost truncate apically (Fig. 2E). Subgenital plate a little concave laterally, macrosetae uniseriate laterally (Fig. 2F). Style narrow, with apophysis slender, digitate, to some extent curved laterally (Fig. 2F). Aedeagal shaft long, bent near base, parallel to base of aedeagus, with tip pointed, gonopore apical (Fig. 2I). Female hind margin of 7 th sternum almost straight. Material examined. Pakistan: Khyber Pakhtunkhwa: 1♀, Nowshera, 34��0���53.8056 N, 71��58���59.3184 E, sweeping hand net, 21 July 2019, coll. Bismillah Shah; 1♀, Faqir Baba, 33��54���53.4888 N, 72��9���56.2176 E, sweep-ing hand net, 20 July 2019, coll. Bismillah Shah; Punjab: 1��, Pir Sohawa, 33��45���37.674 N, 73��3���57.528 E, sweep-ing hand net, 6 August 2018, coll. Bismillah Shah. Distribution. Pakistan, China, India. Remarks. Though there is considerable variation in color pattern of this species, it can be recognized by the vertex with two conspicuous black spots between the anterior margins of the eyes, and by a brown longitudinal line on the forewing., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 401-402, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India, including Ceylon and Burma, 4, pp. 1 - 501.","Matsumura, S. (1914) Die Jassinen und einige neue Acocephalinen Japans. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, 5, 165 - 240.","Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Dai, R. H., Chen, X. X. & Li, Z. Z. (2003) A new species of the genus Paivanana (Homoptera: Cicadellidae: Euscelinae) from China. Entomotaxonomia, 25 (1), 31 - 33.","Li, Z. Z., Dai, R. & Xing, J. (2011) Deltocephalinae from China (Hemiptera: Cicadellidae). Popular Science Press, Beijing, 336 pp.","Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus rotundus (Pruthi) n. rec. (Fig. 4) Arya rotunda Pruthi, 1930: 40. Measurement. Male: 4.0mm. Description. Coloration. Pale to greyish (Fig. 4 A–D). Crown reddish-yellow (Fig. 4A, 4C). Eye and ocellus black (Fig. 4B, 4D). Pronotum with transverse darker greyish shade (Fig. 4A, 4C). Face light brown. Frons with reddish-brown transverse striae in lateral area (Fig. 4D). Forewing brown or stramineous, with brownish venation (Fig. 4 A–B). Legs brown. Abdomen black both dorsally and ventrally in males (Fig. 4 A–B). Morphology. Crown longer than width between eyes, rounded at anterior margin (Fig. 4A, 4C). Eye large, oblique, and projected over pronotal angles laterally (Fig. 4 A–D). Ocellus placed at anterior margin of vertex near eye, barely visible from above (Fig. 4D). Face as long as wide. Antennae long and thin. Pronotum a little longer than vertex with the posterior margin slightly concave (Fig. 4A, 4C). Scutellum with a transverse impression in middle. Male genitalia. Pygofer considerably produced in posterior direction (Fig. 4E). Subgenital plate large, triangular, with a row of uniseriate macrosetae (Fig. 4F). Valve triangular. Style narrow with digitate apophysis, laterally curved (Fig. 4F). Connective with arms well separated, stem slightly shorter than arms (Fig. 4F). Aedeagal shaft long, slender, parallel to aedeagal base, slightly sinuate, tip pointed, curved dorsad, gonopore apical (Fig. 4 G–H). Material examined. Pakistan: Azad Kashmir: 1³, Muzaffarabad, 34°21′44.1144 N, 73°27′10.62 E, sweeping hand net, 14 August 2019, coll. Bismillah Shah. Distribution. Pakistan, India. Remarks. Pruthi (1930) described this species for the first time from India. This is the first record of the species from Pakistan., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 404-406, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68."]}

Stirellus pakhtunensis Shah & Duan sp. n. (Fig. 3) Measurement. Male: 3.6mm. Description. Coloration. General color pale to greyish brown (Fig. 3 A–D). Crown with two spots near posterior margin (Fig. 3A, 3C). Pronotum and scutellum with two dark brown longitudinal stripes (Fig. 3A, 3C). Eye red, surrounded with yellow band (Fig. 3 A–D). Pronotum slightly darker (Fig. 3A, 3C). Scutellum yellowish brown. Forewing brownish with prominent white venation (Fig. 3 A–B). Morphology. Crown almost subequal to pronotum, almost as long as distance between eyes, anteriorly slightly angulate (Fig. 3A, 3C). Eye relatively large. Ocellus next to eye slightly below anterior margin of crown (Fig. 3B, 3D). Face as long as broad, with faded transverse markings near eyes (Fig. 3D). Pronotum median length nearly as long as median length of crown (Fig. 3A, 3C). Mesonotum and scutellum shorter than pronotum. Forewing macropterous; appendix distinctive, prolonged around wing apex; 4 apical and 3 anteapical cells, inner anteapical cell opened at base. (Fig. 3 A–B). Male genitalia. Pygofer lobe broad basally with apex truncate, without macrosetae (Fig. 3E). Subgenital plate slightly convex laterally, apically truncate, macrosetae not uniseriate laterally (Fig. 3F). Valve triangular. Styles narrow, apophysis digitate, slightly curved laterally (Fig. 3F). Connective with stem equal in length to arms (Fig. 3F). Aedeagal shaft long, geniculate basally, nearly parallel to aedeagal base, with preapical anterior expansion visible in lateral and dorsal views, apex pointed and slightly curved dorsad, gonopore apical (Fig. 3 G–H). Material examined. Holotype: Pakistan: Khyber Pakhtunkhwa: 1³, Manki, 34°0′34.5096 N, 73°19′44.4432 E, sweeping hand net, 20 August 2019, coll. Bismillah Shah. Distribution. Pakistan. Remarks. This species can be easily distinguished from other species in this genus by the distinctive external appearance. Etymology. This species is named for Pakhtun-khwa province in Pakistan where the type specimen has been collected.

Shah, Bismillah, Naveed, Hassan, Duan, Yani (2020): A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species. Zootaxa 4895 (3): 398-410, DOI: https://doi.org/10.11646/zootaxa.4895.3.5

Stirellus thattaensis Mahmood, Sultana & Waheed (See Shah et al., 2020: 195, Fig. 4) Stirellus thattaensis Mahmood, Sultana & Waheed, 1972: 82. Material examined. No material examined. Distribution. Pakistan. Remarks. Mahmood et al., (1972) has given detailed description of this species originally described from Pakistan. S. thattaensis is very similar to S. lahorensis in external appearance but differs in male genitalia., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 407, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Mahmood, S. H., Sultana, S. & Waheed, A. (1972) Two new species of Stirellus Osborn & Ball (Homoptera: Cicadellidae: Deltocephalinae) from West Pakistan. Pakistan Journal of Zoology, 4 (1), 79 - 84."]}

Stirellus Osborn & Ball Athysanus (Stirellus) Osborn & Ball, 1902: 250 (Type: Athysanus bicolor Van Duzee, 1892). Stirellus, Kirkaldy, 1907: 58. Doratulina Melichar, 1903: 198 (Type: Doratulina jocosa Melichar, 1903). Phrynophyes Kirkaldy, 1906: 237 (Type: Phrynophyes phrynophyes Kirkaldy, 1906). Anemochrea Kirkaldy, 1906: 329 (Type: Anemochrea mitis Kirkaldy, 1906). Anemolua Kirkaldy, 1906: 329 (Type: Anemolua hanuala Kirkaldy, 1906). Giffardia Kirkaldy, 1906: 336 (Type: Giffardia dolichocephala Kirkaldy, 1906). Arya Distant, 1908: 338 (Type: Arya rubrolineatus Distant, 1908). Trebellius Distant, 1918: 52 (Type: Trebellius albifrons Distant, 1918). Campbellinella Distant, 1918: 69 (Type: Campbellinella illustrata Distant, 1918). Viridomarus Distant, 1918: 69 (Type: Viridomarus capitatus Distant, 1918). Bituitus Distant, 1918: 70 (Type: Bituitus projectus Distant, 1918). Paternus Distant, 1918: 71 (Type: Paternus pusana Distant, 1918). Volusenus Distant, 1918: 72 (Type: Volusenus lahorensis Distant, 1918). Nandidrug Distant, 1918: 74 (Type: Nandidrug speciosus Distant, 1918). Allectus Distant, 1918: 75 (Type: Allectus notata Distant, 1918). Galerius Distant, 1918: 78 (Type: Galerius indicatrix Distant, 1918). Paivanana Distant, 1918: 95 (Type: Typhlocyba indrus Distant, 1908). Gillettiella Osborn, 1930: 689 (Type: Deltocephalus labiatus Gillette, 1898). Bella Pruthi, 1930: 44 (Type: Bella apicalis Pruthi, 1930). Penestirellus Beamer & Tuthill, 1934: 21 (Type: Penestirellus catalinus Beamer & Tuthill, 1934). Sunda Pruthi, 1936: 112 (Type: Sunda ribeiroi Pruthi, 1936). Pseudaconura Linnavuori, 1952: 182 (Type: Pseudaconura luxorensis Linnavuori, 1952). Malagasiella Evans, 1954: 110 (Type: Malagasiella minima Evans, 1954). Umesaona Ishihara, 1961: 246 (Type: Umesaona asiatica Ishihara, 1961). Doratulina (Cymbopogonella) Viraktamath, 1976: 79 (Type: Doratulina longivertex Viraktamath, 1976). Anaconura Emeljanov, 1999: 559 (Type: Stirellus aristidae Emeljanov, 1968). Synonymized by Duan et al., 2019. Coloration and morphology. Yellowish brown to greenish in color. Head variably shaped, slightly to strongly produced. Ocellus present slightly below or on anterior margin. Frontoclypeus straight to slightly convex in profile. Clypeal sulcus prominent. Forewing variably developed, brachypterous to macropterous, usually with three anteapical cells, inner anteapical cell opened at base. Male genitalia. Pygofer triangular in lateral view, dorsal margin sloping posteriorly and mostly straight or concave distally with short tooth present on the apical half near ventral margins. Valve long, triangular. Style with preapical angle distinct. Connective Y-shaped or H-shaped. Aedeagal base triangular, shaft elongate and slender with no basal hinge and strongly curved anterodorsally. Gonopore apical or subapical. Distribution. Worldwide. Remarks. Duan et al. (2019) provided a detailed description of the tribe. Stirellus is heterogeneous and the largest genus of the tribe, distributed worldwide. Variation in external morphological characters including body proportions and coloration explain why many diverse forms were considered previously to represent separate genera. Previously Khatri & Webb (2010) reported 3 known Stirellus species from Pakistan. Recently, Naveed et al. (2020) added 2 additional new species to make a total of 5 known species from Pakistan. Checklist of the species of Stirellus Osborn & Ball from Pakistan Stirellus kumratensis Naveed & Zhang, 2020. Stirellus lahorensis Distant, 1918. Stirellus mankiensis Shah & Duan sp. n. Stirellus neoconvexus Naveed & Zhang, 2020. Stirellus thattaensis Mahmood, Sultana & Waheed, 1972. Stirellus tolla Pruthi, 1930 n. rec. Stirellus viridulus Pruthi, 1930. Key to species of Stirellus Osborn & Ball from Pakistan 1. Crown 1.5x longer than breadth between eyes (Fig. 2A).............................................. S. lahorensis ��� Crown less than 1.5x or equal to breadth between eyes (Figs 1 A���B, 3A���C, 4A, 5A, 6A, 8A, 9A)........................2 2. Species with yellowish green in color (Figs 6 A���C, 8A���C).................................................... 3 ��� Species with ochraceous to brownish in color (Figs 1 A���D, 3A���E, 9A���C).........................................5 3. Crown anterior margin very slightly angulate (Figs 6A, 6C, 8A, 8D)................................... S. tolla n. rec. ��� Crown anterior margin acutely angled (Figs 4A, 5A)......................................................... 4 4. Male pygofer long, with rounded apex (Fig. 5C)................................................... S. viridulus ��� Male pygofer short with pointed apex (Fig. 4E)................................................... S. thattaensis 5. Subgenital plate with macrosetae uniseriate laterally (Fig. 1H)........................................ S. kumratensis ��� Subgenital plate with macrosetae not uniseriate laterally (Fig. 3K, 10B).......................................... 6 6. Connective stem shorter than anterior arms (Fig. 3L), aedeagal shaft with blunt apex (Fig. 3 M���N)........... S. neoconvexus ��� Connective stem longer than anterior arms (Fig. 10C), aedeagal shaft with pointed apex (Fig. 10 D���E).... S. mankiensis sp. n., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species, pp. 189-202 in Zootaxa 4763 (2) on pages 190-191, DOI: 10.11646/zootaxa.4763.2.3, http://zenodo.org/record/3758208, {"references":["Osborn, H. & Ball, E. D. (1902) A review of the North American species of Athysanus (Jassidae). The Ohio Naturalist. A Journal Devoted More Especially to the Natural History of Ohio, 2, 231 - 256.","Van Duzee, E. P. (1892) New North American Homoptera. IV. The Canadian Entomologist, 24, 113 - 117. https: // doi. org / 10.4039 / Ent 24113 - 5","Kirkaldy, G. W. (1907) Leafhoppers supplement (Hemiptera). Report of Work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology Bulletin, 3, 1 - 186.","Melichar, L. (1903) Homopteren-fauna von Ceylon. Verlag von Felix L. Damer, Berlin, 248 pp.","Kirkaldy, G. W. (1906) Leaf-hoppers and their natural enemies. (Pt. IX Leaf-hoppers. Hemiptera). Report of Work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology Bulletin, 1 (9), 271 - 479.","Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India, including Ceylon and Burma, 4, pp. 1 - 501.","Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Osborn, H. (1930) North American leafhoppers of the Athysanella group (Homoptera: Cicadellidae). Annals of the Entomological Society of America, 23, 687 - 720. https: // doi. org / 10.1093 / aesa / 23.4.687","Gillette, C. P. (1898) A few new species of Deltocephalus and Athysanus from Colorado. Bulletin of the Colorado Agricultural Experiment Station, 43, 23 - 29.","Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Beamer, R. H. & Tuthill, L. D. (1934) Some new species and a new genus of deltocephaloid leafhoppers. (Homoptera: Cicadellidae). Journal of the Kansas Entomological Society, 7, 1 - 24.","Pruthi, H. S. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11 (3), 101 - 131.","Linnavuori, R. (1952) Studies on some Palearctic Hemiptera. Annales Entomologici Fennici, 18, 181 - 187.","Evans, J. W. (1954) Les Cicadellidae de Madagascar (Homopteres). Memoires de l'Institut Scientifique de Madagascar, 4, 87 - 137.","Ishihara, T. (1961) Homoptera of Southeast Asia collected by the Osaka city university biological expedition to Southeast Asia 1957 - 1958. Nature & Life in Southeast Asia, 1, 225 - 257.","Viraktamath, C. A. (1976) New species of Doratulina and Bumizana (Homoptera: Cicadellidae) from Karnataka. Oriental Insects, 10 (1), 79 - 86. https: // doi. org / 10.1080 / 00305316.1976.10432323","Emeljanov, A. F. (1999) A key to genera of the subfamily Deltocephalinae s. l. (Homoptera: Cicadellidae) from Kazakhstan, Middle Asia, and Mongolia with description of new genera and subgenera. Entomological Review, 79 (5), 547 - 562.","Emeljanov, A. F. (1968) Notes on the tribe Stirellini (Homoptera: Cicadellidae). Zoologicheskii Zhurnal, 47 (2), 249 - 253.","Duan, Y. N., Zhang, Y. T. & Dietrich, C. H. (2019) Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records. Zootaxa, 4603 (2), 201 - 241. https: // doi. org / 10.11646 / zootaxa. 4603.2.1","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H., Shah, B., Sohail, K. & Zhang, Y. L. (2020) Review of the leafhopper genus Stirellus Osborn & Ball, 1902 (Hemiptera: Cicadellidae: Deltocephalinae) with description of two new species from Pakistan. Zootaxa, 4722 (5), 479 - 485. https: // doi. org / 10.11646 / zootaxa. 4722.5.6","Mahmood, S. H., Sultana, S. & Waheed, A. (1972) Two new species of Stirellus Osborn & Ball (Homoptera: Cicadellidae: Deltocephalinae) from West Pakistan. Pakistan Journal of Zoology, 4 (1), 79 - 84."]}

Stirellus mankiensis Shah & Duan (See Shah et al., 2020: 200���201, Figs 9���10) Stirellus mankiensis Shah & Duan, 2020 in Shah et al., 2020: 200. Measurement. Male: 3.8���4.0mm; female: 5.0mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 11����, 2♀♀, Manki, 34��0���34.5096 N, 73��19���44.4432 E, sweeping hand net, 20 August 2019, coll. Bismillah Shah. Distribution. Pakistan. Remarks. The species can be distinguished from other species of Stirellus by its distinctive external appearance and crown with anterior margin slightly bulbous. This species is somewhat similar to Stirellus neospeciosus Duan, Webb & Zhang in the form of the aedeagus but can easily be separated by the subgenital plate with macrosetae not uniseriate and the connective stem longer than arms; it is also very different from in external appearance., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 402-403, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3"]}

Stirellus kumratensis Naveed & Zhang (See Shah et al., 2020: 192, Fig. 1) Stirellus kumratensis Naveed & Zhang, 2020 in Naveed et al., 2020: 481. Measurement. Male: 3.5–3.6mm; female: 3.9mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 2³³, 1♀, Abbottabad, 34°10′7.5036 N, 73°13′17.3892 E, sweeping hand net, 23 July 2018, coll. Bismillah Shah; 1♀, Barkot, 34°35′58.3944 N, 73°22′41.8764 E, sweep-ing hand net, 11 August 2019, coll. Bismillah Shah; 14³³, 11♀♀, Balakot, 34°34′18.1668 N, 73°21′46.6668 E, sweeping hand net, 12 August 2019, coll. Bismillah Shah; Punjab: 6³³, 3♀♀, Pathriata, 33°52′29.6292 N, 73°27′23.5044 E, sweeping hand net, 29 July 2019, coll. Bismillah Shah; 1³, 7♀♀, Bansra Gali, 33°54′17.532 N, 73°22′13.9656 E, sweeping hand net, 1 August 2019, coll. Bismillah Shah; 4♀♀, Bhurban, 33°56′27.3228 N, 73°26′48.7428 E, sweeping hand net, 1 August 2019, coll. Bismillah Shah; 4³³, 3♀♀, Sunny Bank, 33°55′1.1244 N, 73°23′39.8256 E, sweeping hand net, 15 August 2019, coll. Bismillah Shah; 5♀♀, Lower Topa, 33°53′51.198 N, 73°25′53.8932 E, sweeping hand net, 16 August 2019, coll. Bismillah Shah; 3♀♀, New Murree, 33°52′36.8148 N, 73°27′2.4912 E, sweeping hand net, 16 August 2019, coll. Bismillah Shah; Azad Kashmir: 1♀, Muzaffarabad, 34°21′44.1144 N, 73°27′10.62 E, sweeping hand net, 14 August 2019, coll. Bismillah Shah. Distribution. Pakistan. Remarks. This species looks similar to Stirellus laetus (Melichar) but can easily be separated by the crown with a pair of orange longitudinal stripes along the mesal margins of eyes, the male pygofer with macrosetae, the style apophysis small and the aedeagus with a spine-like apex., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 402, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Naveed, H., Shah, B., Sohail, K. & Zhang, Y. L. (2020) Review of the leafhopper genus Stirellus Osborn & Ball, 1902 (Hemiptera: Cicadellidae: Deltocephalinae) with description of two new species from Pakistan. Zootaxa, 4722 (5), 479 - 485. https: // doi. org / 10.11646 / zootaxa. 4722.5.6"]}

Stirellus viridulus (Pruthi) (See Shah et al., 2020: 196, Fig. 5) Paternus viridula Pruthi, 1930: 42. Paternus viridulus, Metcalf, 1967: 2350. Stirellus viridulus, Khatri & Webb, 2010: 1���47. Measurement. Male: 3.3���3.7mm; female: 3.9mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 1��, 4♀♀, Pirsabak, 34��0���37.1196 N, 72��3���3.6648 E, light trap, 12 July 2018, coll. Bismillah Shah; 1��, Kund Park, 33��55���50.0304 N, 72��14���5.0064 E, sweeping hand net, 30 July 2018, coll. Bismillah Shah; 2����, 1♀, Khairabad, 33��54���15.1596 N, 72��13���37.8264 E, sweeping hand net, 20 August 2018, coll. Bismillah Shah; 1��, 2♀♀, Faqir Baba, 33��54���53.4888 N, 72��9���56.2176 E, sweeping hand net, 20 July 2019, coll. Bismillah Shah; 2����, Manki, 34��0���34.5096 N, 73��19���44.4432 E, sweeping hand net, 20 August 2019, coll. Bismillah Shah; Punjab: 28����, 12♀♀, Kuldana, 33��55���15.8232 N, 73��23���58.0812 E, sweep-ing hand net, 1 August 2019, coll. Bismillah Shah. Distribution. Pakistan, India. Remarks. This species closely resemble S. lahorensis (Distant) in external appearance and male genitalia but varies in color and in the pygofer lobe. The caudal region of pygofer lobe in S. viridulus (Pruthi) is straight but curved in S. lahorensis (Distant)., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 409, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Metcalf, Z. P. (1967) General catalogue of the Homoptera. Fascicle VI. Cicadelloidea. Part 10. Section III. Euscelidae. U. S. Department of Agriculture, Agriculture Research Service, Washington, D. C., pp. 2075 - 2695.","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1"]}