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Abstract We present data on the distribution of seven wasp families, Ceraphronidae, Diapriidae, Ismaridae, Megaspilidae, Platygastridae, Proctotrupidae and Scelionidae, collected in Georgia (Sakartvelo). Thirty-three genera are recorded for the first time from Georgia and 25 of them for the Caucasus. From these, 16 species level taxa are newly recorded in Georgia, and 14 identified species are new records for the Caucasus. This brings the number of known insect species in the Lagodekhi Protected Area (LPA) to 1682.

Three species of the Acanopsilus Kieffer, 1908, six species of Acanosema Kieffer, 1908, two species of Polypeza Förster, 1856, and four species of Psilomma Förster, 1856 of the Russian fauna are reviewed, keyed and illustrated. Six of these species are described: Acanopsilus comadensis sp. n. (Russia: Primorsky krai), A. minimus sp. n. (Russia: Primorsky krai), Acanosema (Cardiopsilus) dentigastrum sp. n. (Russia: Primorsky krai), A. (C.) epicnemium sp. n. (Russia: Primorsky krai) A. (C.) setigerum sp. n. (Russia: East Siberia, Yakutia; Sweden), Psilomma calaris sp. n. (Russia: Kunashir Island; Japan). The generic status of Cardiopsilus Kieffer, 1908 is reduced to subgeneric one under Acanosema Kieffer, 1908 and a new combination is proposed for A. productum (Kieffer, 1908), comb. n. from Cardiopsilus Kieffer, 1908.

The taxonomic status of the genera Acropiesta Förster, 1856, Anommatium Förster, 1856, Erasikea Szabó, 1961 and Pantolyta Förster, 1856 is discussed. The following generic synonymies are proposed: Acropiesta, syn. nov., is synonymised with Pantolyta, and Erasikea, syn. nov., is synonymised with Anommatium. Erasikea nutrix Szabó, 1961, syn. nov., is a junior synonym of Anommatium ashmeadi Mayr, 1904. Three species are described as new to science: Pantolyta dichromia sp. nov. (Russia: European part, Primorskiy Territory), P. melniki sp. nov. (Russia: Kunashir Island; Republic of Korea) and P. simplicior sp. nov. (Russia: Kunashir Island). Eleven previously known Palaearctic species of the former genus Acropiesta and one species of Anommatium recorded in the fauna of Russia are reviewed and illustrated. The following new combinations are proposed: Pantolyta flavicauda (Ashmead, 1893), comb. nov.; P. flaviventris (Thomson, 1858), comb. nov.; P. flexinervis (Macek, 1998), comb. nov.; P. macrocera (Thomson, 1858), comb. nov.; P. micans (Macek, 1998), comb. nov.; P. nigrocincta (Kieffer, 1909), comb. nov.; P. nitida (Thomson, 1858), comb. nov.; P. pseudosciarivora (Macek, 1998), comb. nov.; P. pulchella (Whittaker, 1930), comb. nov.; P. radialis (Hellén, 1964), comb. nov.; P. rufiventris (Kieffer, 1909), comb. nov.; P. sciarivora (Kieffer, 1859), comb. nov.; P. semirufa (Fouts, 1924), comb. nov.; P. seticornis (Kieffer, 1910), comb. nov.; P. subaptera (Ashmead, 1893), comb. nov.; Pantoclis radiatula (Thomson, 1858), comb. nov. An identification key to all Palaearctic species of Pantolyta and Anommatium is given.

A total of 43 insect parasitoid species belonging to fourteen families (Aphelinidae, Braconidae, Ceraphronidae, Diapriidae, Encyrtidae, Eulophidae, Eurytomidae, Ichneumonidae, Megaspilidae, Mymaridae, Platygasteridae, Proctotrupidae, Pteromalidae, Trichogrammatidae) has been documented in the rice ecosystem using yellow pan trap. The observations were made at four important stages of rice crop like early tillering, active tillering, booting and panicle development. The parasitoids were also compared with the occurrence of sixteen insect pests that were recorded simultaneously in each stage of the crop. The result revealed that, there is a significant difference in the occurrence of parasitoids according to the stage of the crop and insect host availability. This understanding help in the introduction of specific parasitoids at respective stages for effective biocontrol.

A review of 17 European species of the genus Spilomicrus Westwood is provided with a key to and illustrations of all known European species except for the formosus group. Two species, S. diversus sp. n. (Europe, Siberia, Far East of Russia) and S. latus sp. n. (Europe), are described as new to science. The following new synonymies are established (valid name given first): S. antennatus Jurine, 1807 = S. quinquepunctatus Szabo, 1961, syn. n.; S. flavipes Thomson, 1859 = S. szelenyii Szabo, 1977, syn. n.; S. hemipterus Marshall, 1868 = S. inaequalis Tomsik, 1947, syn. n., = S. thomsoni Kieffer, 1911, syn. n.; S. integer Thomson, 1859 = S. major Vollenhoven, 1879, syn. n.; S. lusitanicus (Kieffer, 1910) = S. gracilicornis Kieffer, 1911, syn. n., = S. noctiger Szabo, 1977, syn. n.; S. nigriclavis Marshall, 1868 = S. punctatus Kozlov, 1978, syn. n., = S. nigriclavis var. subarmatus Kieffer, 1912, syn. n.; S. rufitarsis Kieffer, 1911 = S. pseudocursor Szabo, 1974, syn. n.; S. stigmaticalis Westwood, 1832 = S. basalyformis Marshall, 1868, syn. n., = S. barbatus Szabo, 1983, syn. n., = S. mediofurcatus Szabo, 1983, syn. n., = S. pilicornis Szabo, 1977, syn. n. Lectotypes are designated for 11 species: S. basalyformis Marshall, 1868, S. flavipes Thomson, 1859, S. hemipterus Marshall, 1868, S. inaequalis Tomsik, 1947, S. integer Thomson, 1859, S. modestus Tomsik, 1947, S. subarmatus Kieffer, 1912, S. thomsoni Kieffer, 1911, S. simplex Tomsik, 1947, S. stigmaticalis Westwood, 1832, and Trichopria lusitanica Kieffer, 1910.

The occurrence of the genera and morphospecies of Diapriidae (Insecta, Hymenoptera) in four habitats (stands) of eastern Mozambique, was studied. The genus Miota is recorded from the Afrotropical region for the first time. Overall, diapriids were represented by 52 morphospecies from 10 genera of the subfamilies Belytinae and Diapriinae. The highest occurrence (140 specimens) and the largest taxonomic diversity (42 morphospecies from 9 genera) were found for the diapriids in the native miombo stand (Brachystegia sp.), followed by the eucalypt, Eucalyptus spp., (92 specimens, 24 morphospecies from 8 genera), the loblolly pine, Pinus taeda, (62 specimens, 19 morphospecies from 7 genera), and the wild loquat (Uapaca kirkiana) stands (40 specimens, 14 morphospecies from 7 genera). The most morphospecies-rich genera were Basalys, Coptera and Trichopria (15, 14 and 7 morphospecies respectively), with Coptera being the most specimen-rich in all samples. The same diapriid morphospecies prevailed in all studied habitats. The abundance estimators ACE and Chao1 predicted 61 (60 in Chao1) species for miombo stand, that is nearly 3.0 times richer than the 22 species estimated for wild loquat, 2.1 (2.6) times richer than the 29 (23) species estimated for loblolly pine and 1.5 times richer than the 40 (39) species estimated for eucalypt. The diapriids of the exotic eucalypt and native miombo stands were most similar, and clustered together with diapriids of the exotic loblolly pine stand.

The biology of a koinobiont parasitoid of leaf-cutting ant larvae, Szelenyiopria talitae (Hymenoptera: Diapriidae), was studied from naturally infested Acromyrmex subterraneus (Hymenoptera: Formicidae) nests. Nests were collected in the field from the Atlantic rainforest biome in the state of Rio de Janeiro. A total of fifty-three nests were collected from 2015 to 2018. Parasitized nests were only found during the months of September and October. Approximately 22% of the nests collected over a four-year period were found to have been parasitized by S. talitae. The mean within-nest parasitism rate was 66.3%. This diapriid displayed both solitary (14%) and gregarious parasitism (86%), with up to a maximum of 12 parasitoids developing within a single host. Gregarious parasitism with two (29%) or three (21%) S. talitae per host was most frequently observed. There was a positive correlation between the number of parasitoids per host and host size (dry weight), indicating that S. talitae females oviposited a higher number of eggs in larger hosts. There was also a negative correlation between S. talitae pharate adult size and the number of parasitoids per host, which could have been caused by sibling competition for limited host resources. The high levels of parasitism seen here had a debilitating effect on the colonies. Acromyrmex subterraneus is a serious pest in Brazil, and these studies lay the foundation for understanding the impact of S. talitae on ant populations.

Spilomicrus succinalis sp. nov. urn:lsid:zoobank.org:act: C6DACEAE-5CBB-4BC5-A414-FC2458978B3D Figs 1H, 6E–H, Table 1 Diagnosis Head higher than long, minutely punctuate on vertex; scape with distinct punctuations; pedicel pearshaped, narrower at base; F1–F5 as long as wide; F6–F11 forming non-abrupt clava; F11 conical, longer than F10 (Fig. 6E, G–H); occipital flange foveate (Fig. 6F); pronotum laterally with small foveae along posterior margin (Fig. 6E); mesoscutum flat and smooth, with long hairs; notauli convergent, interrupting before posterior margin of mesoscutum; anterior scutellar pit bifoveate; posterior scutellar pits present (Fig. 6F); propodeum carinate; Sc+R pigmented, marginal vein wider than long, r-rs as long as marginal vein (Fig. 1H); first fore tarsomere bearing row of setae along inner margin; first hind tarsomere not longer than following two combined; petiole 1.5 times as long as wide; T2 with anterior margin wrinkled (Fig. 6F); two ring-like segments visible after T2. Etymology The species name derives from ‘Succinite’, a name given to Baltic amber due to its chemical composition. The specific epithet is to be treated as an adjective. Type material Holotype NHMD-607131, a complete female; paratype NHMD-608344, a complete female; paratype NHMD-608354, a complete female. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 1.96–2.65 mm. Head higher than long (HeL = 0.29–0.50 mm), with several hairs dorsally; head minutely punctuate on vertex; eye oval, higher than long; antennae inserted on shelf at level with lower half of eye; toruli distinct and slightly protruding; scape with distinct punctuations, 4.2 times as long as wide; pedicel pear-shaped, narrower at base; F1–F5 compact, as long as wide; following flagellomeres widening to form non-abrupt club; F11 wider and longer than F10, conical (antennomeres length of holotype, in mm: Sc-0.25; P-0.09; F1-0.05; F2-0.04; F3-0.04; F4-0.04; F5-0.05; F6-0.06; F7-0.06; F8-0.06; F9-0.08; F10-0.08; F11-0.10); mandibles simple, not forming beak; occipital flange foveate. MESOSOMA. Shorter than metasoma (MsL = 0.67–0.83 mm); pronotum not elongate, not visible in dorsal view, laterally with small foveae on posterior margin; mesoscutum wide, flat and smooth, with several long hairs; notauli present, deep, convergent posteriorly but not reaching posterior margin of mesoscutum; anterior scutellar pit bifoveate; scutellum with posterior scutellar pits present; propodeum carinate, with prominent median keel. Fore wing extending beyond gaster (FwL = 1.27–1.81 mm), hyaline and covered with micropubescence, rounded, bordered with long setae shortening toward wing apex; Sc+R pigmented and distinctly separated from wing margin; marginal vein thickening at apex of Sc+R and almost two times as wide as long; r-rs pigmented and elongate, almost as long as marginal vein. Hind wing half as long as fore wing; only C pigmented. Legs slender, with numerous long setae; first fore tarsomere bearing row of erect setae on inner margin; fore tibial spur long, curved; tibial spur formula 1-2-2; first hind tarsomere not longer than following two combined. METASOMA. Petiole longer than wide (PtL = 0.14–0.21 mm; PtW = 0.10–0.13 mm), covered with longitudinal carinae, glabrous; gaster beyond petiole broadly oval and narrowed at apex (GL = 0.87– 1.11 mm; GH = 0.23–0.43 mm), smooth and with few setae; T2 longest, covering two thirds of gaster, with anterior margin slightly wrinkled, overlapping petiole; S2 longest, as long as T2; segments 3 and 4 ring-like; segment 5 longer than 3 and 4 combined and sharply pointed at apex; ovipositor projecting as short stub. Male Unknown. Comments Using Nixon’s (1980) and Masner & García’s (2002) keys to the genera of Diapriinae, Spilomicrus succinalis sp. nov. clearly keys out to Spilomicrus because of the following characters: antenna with 13 segments, flagellum thickened towards apex, frons unarmed, ocellar triangle in the ocular zone, notauli present, scutellum with an anterior bifoveate pit, wings fully developed, fore wing with pigmented Sc+R, marginal vein not more than one and a half times as long as wide, anterior margin of syntergite straight, without a furrow or a cleft at base. Following the key to the British species of Nixon (1980) and to Eastern Palaearctic species of Chemyreva (2018), S. succinalis sp. nov. keys out near S. comatus Chemyreva, 2015. It differs from this species by having F11 longer than F10, the petiole less than two times as long as wide and lacking the long hairs on the postgena (Chemyreva 2015).

Pantolyta augustinusii sp. nov. urn:lsid:zoobank.org:act: DD4A2A42-E41A-4731-B91D-AD9EFC205E50 Figs 1E, 4, Table 1 Diagnosis Eye not pubescent; toruli protruding; antenna distinctly pubescent; F1 with indistinct emargination; F1 half of scape length (Fig. 4A–B); mesoscutum bearing long hairs; median propodeal keel simple; fore wing radial cell closed; postmarginal vein extending little beyond radial cell; Rs nebulous proximally to r-rs, marginal vein 0.7 times as long as its distance from basal vein; r-rs distinct (Fig. 1E); petiole 2.4 times as long as wide (Fig. 4D). Etymology We dedicate this species to the Augustinus Foundation, who generously contributed to the funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in genitive case. Type material Holotype NHMD-300829, a complete male; paratypes NHMD-608391, a complete male; NHMD- 608406, a complete male but partially hidden by a milky coat; NHMD-608412, a complete male; NHMD-608404, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. BL = 2.51–3.47 mm. Head subtriangular in frontal view (HeL = 0.39–0.48 mm); eye large, not pubescent; several hairs on frons; antennae inserted on distinct but not prominent shelf; toruli well developed, as long as one sixth of scape length from base to apex, separated by distinct cleft; antennae bearing numerous short setae; scape almost six times as long as wide; pedicel slightly longer than wide; flagellomeres elongate, cylindrical; F1 half of scape length, with an indistinct emargination basally; F2–F11 subequal in length; F12 longer and tapering at apex (antennomeres length of holotype, in mm: Sc-0.38; P-0.08; F1-0.21; F2-0.19; F3-0.19; F4-0.19; F5-0.19; F6-0.20; F7-0.19; F8-0.20; F9-0.21; F10- 0.21; F11-0.21; F12-0.23); mandibles short, not forming a beak. MESOSOMA. Shorter than metasoma (MsL = 0.90–1.29 mm); pronotum visible in dorsal view; epomia present as a simple keel; pronotal shoulders rounded; mesoscutum smooth, bearing long hairs; notauli deeply impressed, anteriorly converging towards scutellum, posteriorly parallel just before transscutal sulcus; anterior scutellar pit large, deep; scutellum without posterior scutellar pits; mesopleuron bare, mesopleural pit present; propodeum with median keel simple. Fore wing hyaline, extending far beyond metasoma (FwL = 2.60–3.54 mm), covered with micropubescence; C, Sc+R, M+Cu, basal vein and Cu pigmented; basal and costal cells closed; marginal vein slightly widened, about two thirds as long as its distance from basal vein and twice as long as r-rs; postmarginal vein pigmented, extending slightly beyond radial cell; radial cell closed; Rs tubular distally to r-rs, nebulous proximally; M inconspicuous towards apex. Legs slender, hind femur slightly thicker; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, 2.4 times as long as its middle width (PtL = 0.27–0.37 mm; PtW = 0.11– 0.15 mm), with longitudinal ribs, long hairs laterally and ventrally; gaster fusiform (GL = 0.90–1.33 mm; GH = 0.42–0.58 mm), smooth; T2 longest, narrowing toward petiole; other segments much shorter. Female Unknown. Comments Using Nixon’s (1957) key to the genera of Belytinae, Pantolyta augustinusii sp. nov. keys out near Acropiesta Förster, 1856 because of the following characters: notauli present, second flagellomere not modified; scutellum without foveae; mandibles not forming a beak; marginal vein shorter than both radial cell and its distance from basal vein; pronotum with reduced epomia; large tergite markedly tapering towards petiole. Pantolyta augustinusii especially share with Acropiesta the antennal shelf prominent with toruli separated by a furrow, the 14-segmented antennae with F1 modified, the pronotal shoulders rounded, the notauli complete, the anterior scutellar pit large and quadrate, the propodeum with median keel simple, the fore wing venation with C, Sc+R, M+Cu, Rs, basal and marginal vein tubular, the petiole cylindrical (Macek 1998). However, due to their close similarities, Acropiesta has recently been synonymized under Pantolyta by Chemyreva & Kolyada (2021). Four other Pantolyta species are present in Baltic amber, two having been described under Acropiesta and not duly transferred by Chemyreva & Kolyada (2021): Pantolyta antiqua Buhl, 1999, Pantolyta janzeni (Buhl, 2002) comb. nov., Pantolyta perrichoti (Jouault & Nel, 2020) comb. nov. and Pantolyta somnulenta Maneval, 1938. Pantolyta antiqua has the marginal vein as long as its distance from basal vein whereas it is distinctly shorter in Pantolyta augustinusii (Buhl 1999); P. janzeni has a distinct emargination of F 1 in the male antenna, the marginal vein is shorter and the postmarginal vein is longer than in Pantolyta augustinusii (Buhl 2002); P. perrichoti has the median propodeal keel bifid (Jouault & Nel 2020) and P. somnulenta has the radial cell shorter than the marginal vein (Maneval 1938).

Basalys villumi sp. nov. urn:lsid:zoobank.org:act: 4204B17E-4659-432C-9266-1C723B434E35 Figs 1F, 5F–H, 6A, Table 1 Diagnosis Body smooth; scape seven times as long as wide; pedicel conical; flagellomeres cylindrical and longer than wide; F1 longest; F2–F11 similar in length; F2 conspicuously emarginate (Fig. 5D–E); scutellum subquadrate without posterior scutellar pits (Fig. 5F); fore wing bordered with small hairs, r-rs short but distinct, M+Cu nebulous (Fig. 1F); hind wing narrow, two thirds of fore wing length, with long hairs along posterior margin; petiole three times as long as wide (Figs 5D, 6A). Etymology We dedicate this species to Villum Fonden, who generously contributed to funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in the genitive case. Type material Holotype NHMD-608360, a complete male; paratype NHMD-608369, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. BL = 1.82 mm. Body highly glabrous. Head globular, as long as high (HeL = 0.33 mm), with sparse short hairs; eye round; toruli situated at middle of eye height; antenna 14-segmented; scape seven times as long as wide; pedicel conical; flagellomeres cylindrical, elongate, distinctly longer than wide, with short hairs; F1 longest, F2–F11 subequal in length; F2 emarginate on anterior third; F12 as long as F1; tapering at apex (antennomeres length of holotype, in mm: Sc- 0.22 mm; P-0.09; F1-0.15; F2-0.09; F3-0.11; F4-0.10; F5-0.11; F6-0.10; F7-0.12; F8-0.10; F9-0.11; F10-0.12; F11-0.11; F12-0.14); mandibles weakly crossing at tip, not sickle or beak-shaped; occipital flange foveate, rather concave. MESOSOMA. Shorter than metasoma (MsL = 0.63 mm); pronotum not elongate, anterior part setose, epomia absent; mesoscutum slightly convex, smooth, notauli absent; anterior scutellar pit present, round; scutellum subquadrate, without posterior scutellar pits. Fore wing extending beyond metasoma (FwL = 1.78 mm), bordered with small hairs; Sc+R distinctly separated from anterior margin; marginal vein thickened along wing margin, longer than wide; r-rs short; basal vein curved, perpendicular to Sc+R, almost reaching M+Cu; M+Cu nebulous. Hind wing narrow, two thirds of fore wing length; bordered with long setae along posterior margin. Legs slender, covered with scattered setae; femur clavate; tibia weakly broadened apically; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, narrow, three times as long as wide (PtL = 0.21 mm; PtW = 0.07 mm), longitudinally striated; gaster ellipsoidal (GL = 0.61 mm; GH = 0.30 mm), glabrous; T2 and S2 longest, covering two thirds of gaster, anterior margin T2 straight; segments 3 and 4 ring-like, segment 5 longer than 3 and 4 combined. Female Unknown. Comments Using Nixon’s (1980) and Masner & García’s (2002) keys, Basalys villumi sp. nov. keys out to Basalys Westwood, 1833 because of the following characters: antenna with 14 segments, notauli absent, scutellum with anterior scutellar pit, wings fully developed, Sc+R separated from fore margin of wing, fore wing with distinct basal vein, basal margin of large tergite straight. The description of the species is also consistent with the diagnosis of Hou & Xu (2016). Basalys villumi sp. nov. differs from extant Basalys by having F1 longer than F2 (Nixon 1980; Hou & Xu 2016)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 76-77, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1980. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Diapriinae. Handbooks for the Identification of British Insects 8 (3 di): 1 - 55.","Masner L. & Garcia J. L. 2002. The genera of Diapriinae (Hymenoptera: Diapriidae) in the New World. Bulletin of the American Museum of Natural History 268: 1 - 138. https: // doi. org / c 7 zvsh","Hou Z. & Xu Z. 2016. First record of the genus Basalys Westwood, 1833 (Hymenoptera: Diapriidae) from China, with descriptions of two new species. Zoological Systematics 41 (3): 337 - 341. https: // doi. org / 10.11865 / zs. 201639"]}

Belyta knudhoejgaardi sp. nov. urn:lsid:zoobank.org:act: 618161BD-F876-422B-9C35-62BAD18B2457 Figs 1A, 2A–D, Table 1 Diagnosis Head nasiform; antennal shelf strongly projecting forward; scape as long as head; F1 longest; following flagellomere shorter but gradually lengthening (Fig. 2A); mesopleuron with one prominent longitudinal ridge (Fig. 2B); median propodeal keel simple (Fig. 2C); radial cell open, Rs fading before reaching fore margin; marginal vein hardly longer than r-rs (Fig. 1A); petiole 1.7 times as long as wide; gaster fusiform (Fig. 2B–C). Etymology We dedicate this species to the Knud Højgaard Foundation, who generously contributed to funding for purchasing the amber pieces studied. The specific epithet is to be treated as a noun in genitive case. Type material Holotype NHMD-608408, a complete female; paratype NHMD-608400, a slightly damaged female. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 4.8–5.3 mm. Body slightly pubescent, smooth and shiny. Head nasiform (HeL = 0.77– 0.89 mm), with sparse short hairs posteriorly; eye almond-shaped, glabrous; antennal shelf strongly prominent; antenna 15-segmented, with short heterogeneous setae; scape as long as head; pedicel thinner; F1 one and a half times as long as pedicel or other flagellomeres; F2–F13 gradually lengthening; F13 tapering at apex and slightly shorter than F1 (antennomere length of holotype, in mm: Sc-0.72; P-0.19; F1-0.29; F2-0.11; F3-0.12; F4-0.11; F5-0.12; F6-0.10; F7-0.11; F8-0.13; F9-0.13; F10-0.14; F11-0.15; F12-0.15; F13-0.28); mandibles simple. MESOSOMA. Almost as long as metasoma, flattened (MsL = 1.62–1.90 mm); pronotum with epomia extending from front coxa to posterodorsal pronotal corners; posterior margin of pronotum curved; mesoscutum with several hairs; notauli complete and deeply impressed; scutellum without posterior scutellar pits; mesopleuron with prominent longitudinal keel; propodeum with plicae, median propodeal keel simple. Fore wing hyaline, micro-pubescent, extending beyond metasoma (FwL = 2.74–3.78 mm); C, Sc+R, M+Cu, basal vein, marginal vein and r-rs pigmented; marginal vein shorter than its distance from basal vein; postmarginal vein barely longer than r-rs, pigmented on half length of radial cell; Rs nebulous proximally and pigmented distally from r-rs, projecting distally to postmarginal vein but fading before reaching wing margin; radial cell open. Hind wing hyaline, reduced and narrow (HwL =?– 2.39 mm). Legs slender, with hind femur and coxa enlarged, covered with scattered setae; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole glabrous, longitudinally striated and with transverse keels, 1.7 times longer (PtL = 0.50 mm) than wide (PtW = 0.30 mm); gaster fusiform, depressed (GL = 1.90–1.95 mm, GH = 0.48– 0.58 mm); S2 and T2 longest, covering two-thirds of gaster, T2 striated at junction with petiole; four ringlike segments distinguishable beyond large tergite, hypopygium bearing several long setae; ovipositor not exerted. Male Unknown. Comments Using Nixon’s (1957) key, Belyta knudhoejgaardi sp. nov. keys out to Belyta because of the following characters: macropterous, notauli present, mandibles of ordinary form, scutellum without a row of foveae along posterior margin, marginal vein shorter than its distance from basal vein, pronotum elongate, without a hollow on each side (= pronotal pits), epomia present, median keel of propodeum simple, petiole more than one and a half time as long as wide. The flattened body is generally accepted to diagnose Belyta spp. (Macek 1996) and is present in Belyta knudhoejgaardi, strengthening the placement of the species in Belyta. Following the key to European species of Belyta of Macek (1996), Belyta knudhoejgaardi sp. nov. keys out between B. subclausa Kieffer, 1916 and B. validicornis Thomson, 1858. It differs from the first by having a marked epomia and from the second by having the median propodeal keel simple (Macek 1996). The diagnostic character for Belyta knudhoejgaardi is the curiously marked longitudinal keel on the mesopleuron, which is absent in other species of Belyta. From the Eocene of Florissant is known B. mortuella Brues, 1910 (attributed to Belyta s. lat.), which differs from B. knudhoejgaardi by being shorter and having a less flattened body (Brues 1910)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 61-62, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Macek J. 1996. Revision of the European species of Belyta Jurine. Acta Musei Nationalis Prague, Series B, Historia Naturalis 51 (1 - 4): 29 - 39.","Brues C. T. 1910. The parasitic Hymenoptera of the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, Harvard College 54 (1): 3 - 125. Available from https: // www. biodiversitylibrary. org / page / 30208802 [accessed 2 Jan. 2021]."]}

Pantoclis globosa sp. nov. urn:lsid:zoobank.org:act: 4BF21BF6-7925-4F2A-8A84-5D8C49BBD8D7 Figs 1D, 3D–H, Table 1 Diagnosis Eye glabrous; female antenna with: scape 5.5 times as long as wide, pedicel shorter than F1, F1 longest, F2–F13 as long as wide (Fig. 3D–E); male antenna with: scape 5.5 times as long as wide, pedicel rounded, F1 longest, emarginate on anterior third, F2–F12 subequal in length (Fig. 3G–H); mesosoma sparsely pubescent, shorter than metasoma; pronotum not elongate with epomia extending from pronotal shoulders to front coxa; notauli on mesoscutum complete and convergent posteriorly; propodeum with plicae; marginal vein shorter than radial cell and as long as three quarters of its distance to basal vein; radial cell closed; Rs inconspicuous proximally to r-rs (Fig. 1D); petiole as long as wide; gaster globous (Fig. 3F–G). Etymology The species name refers to the globular gaster of the species. The epithet is to be treated as an adjective. Type material Holotype NHMD-608414, a complete female but partially hidden by a milky coat; paratype NHMD- 608394, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 2.83 mm. Head oval, subtriangular in frontal view (HeL = 0.48 mm), bearing several short hairs, stouter on frons; eye oval, glabrous; antenna inserted on frontal shelf, at level with lower margin of eyes; toruli well separated; antenna bearing numerous short sensilla; scape shorter than head length, 5.5 times as long as wide; pedicel shorter than F1, longer than wide; 13 flagellomeres; F1 longest, two times as long as wide; F2–F13 as long as wide; F13 tapering at apex (antennomeres length of female holotype, in mm: Sc-0.44; P-0.14; F1-0.16; F2-0.10; F3-0.10; F4-0.09; F5-0.09; F6-0.09; F7-0.09; F8- 0.08; F9-0.09; F10-0.09; F11- 0.09; F12-0.09; F13-0.10); mandibles of ordinary form; occipital carina foveate. MESOSOMA. Smooth and shiny, shorter than metasoma (MsL = 0.97 mm), sparsely pubescent; pronotum not elongate with distinct and uninterrupted epomia; mesoscutum distinctly convex, notauli complete and convergent; anterior scutellar pit suboval, slightly narrower than apical distance between notauli; scutellum without posterior scutellar pit; propodeum with two prominent plicae; median propodeal keel simple. Fore wing hyaline, homogeneously micropubescent, extending beyond metasoma (FwL = 2.71 mm); C, Sc+R, M+Cu, basal vein, marginal vein, postmarginal vein and r-rs pigmented; Cu nebulous; marginal vein length around three quarters of its distance from basal vein; marginal vein shorter than radial cell; radial cell closed by Rs; Rs pigmented distally to r-rs, inconspicuous proximally. Legs slender, with short hairs; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole twice as long as wide (PtL = 0.27 mm; PtW = 0.12 mm), longitudinally striated, bearing long hairs dorsally; gaster globular, slightly longer than high (GL = 1.05 mm; GH = 0.67 mm), with sparse pubescence; S2 and T2 longest; four ring-like segments discernible beyond large tergite. Male BL = 3.50 mm; similar to female but antennae sexually dimorphic: scape shorter than head length (HeL = 0.53 mm), 5.5 times as long as wide, pedicel rounded, as long as wide, 12 flagellomeres elongate, cylindrical, longer than wide, F1 longest, with marked emargination on anterior third of segment, F2–F12 subequal in length, F12 tapering at apex (antennomeres length of male paratype, in mm: Sc- 0.38 mm; P-0.11; F1-0.25; F2-0.19; F3-0.19; F4-0.19; F5-0.18; F6-0.17; F7-0.16; F8-0.16; F9-0.16; F10-0.16; F11- 0.15; F12; 0.21). Remaining measurements: HeL = 0.53 mm; MsL = 1.24 mm; FwL = 2.84 mm; PtL = 0.32 mm; PtW = 0.15 mm; GL = 1.41 mm; GH = 0.82 mm. Comments Using Nixon’s (1957) key, Pantoclis globosa sp. nov. keys out to Pantoclis because of the following characters: antennae inserted on a frontal prominence, mandibles not forming a beak or sickle-shaped, notauli present, scutellum without foveae along posterior margin, marginal vein nearly as long as its distance from basal vein and presence of ring-like segments beyond the large tergite, marginal vein shorter than radial cell, epomia distinct, median propodeal keel simple, petiole at most one and a third times as long as its apical width. Pantoclis globosa also fits in Pantoclis according to the diagnosis of Hou et al. (2016). It differs from most extant representatives of Pantoclis by having the marginal vein much longer than the r-rs (Nixon 1957; Sharma 1980; Buhl 1998). Pantoclis longiscapa Chambers, 1974 has a similar fore wing configuration but differs by having F1 very long and F2 distinctly longer than the following flagellomeres (Chambers 1974). Pantoclis deperdita Brues, 1906, from the Eocene of Florissant has a shorter radial cell and marginal vein (Brues 1906) and is therefore considered as a different species from P. globosa., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 68-69, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Sharma S. K. 1980. On some new species of Belytinae and Diapriinae (Hymenoptera: Proctotrupoidea) from India. Oriental Insects 14 (1): 51 - 61. https: // doi. org / 10.1080 / 00305316.1980.10434583","Buhl P. N. 1998. New or little known Oriental and Australasian Belytinae (Hymenoptera: Diapriidae). Oriental Insects 32 (1): 41 - 58. https: // doi. org / 10.1080 / 00305316.1998.10433766","Chambers V. H. 1974. Taxonomic notes on the Belytinae, with a new species of Pantoclis Forster (Hym., Proctotrupoidea, Diapriidae). Journal of Entomology (B) 42 (2): 127 - 121. https: // doi. org / 10.1111 / j. 1365 - 3113.1974. tb 00064. x","Brues C. T. 1906. Fossil parasitic and phytophagous Hymenoptera from Florissant, Colorado. Bulletin of the American Museum of Natural History 22: 491 - 498. Available from http: // hdl. handle. net / 2246 / 1719 [accessed 2 Jan. 2021]."]}

Pantolyta chemyrevae sp. nov. urn:lsid:zoobank.org:act: 626E3F04-AB34-45C2-AA5A-82AD0A85CC8E Fig. 5A–C, Table 1 Diagnosis Eye round, extending at most a fourth of head length; scape as long as pedicel and first three flagellomeres combined; F1 two times as long as wide, narrower than pedicel; F2–F13 gradually widening towards apex (Fig. 5A–B); epomia present; anterior scutellar pit suboval, wider than shortest distance between notauli; plicae projecting posteriorly; fore wing not reaching middle of propodeum (brachypterous morph; Fig. 5A–B); petiole slightly longer than wide (Fig. 5C); gaster fusiform, elongate and tapering at apex; T2 striated dorsally at junction with petiole; ovipositor long, subequal in length to gaster (Fig. 5A). Etymology The species is dedicated to Vasilisa Chemyreva, Russian entomologist, in acknowledgment of her numerous publications on Diapriidae from the Palearctic region. The specific epithet is to be treated as a noun in the genitive case. Type material Holotype NHMD-608448, a complete female partially hidden by a milky coat. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 2.43 mm. Head smooth, bare (HeL = 0.37 mm); eye round, small, less than a third of head length (ED = 0.11 mm); antenna bearing numerous short setae; toruli separated by shallow cleft; scape as long as pedicel and first three flagellomeres combined, at least as long as head; pedicel three times as long as wide; 13 flagellomeres; F1 two times as long as wide, narrower than pedicel; following flagellomeres gradually widening towards apex; F13 largest and longest of all (antennomeres length of holotype, in mm: Sc-0.46; P-0.16; F1-0.12; F2-0.08; F3-0.08; F4-0.08; F5-0.09; F6-0.08; F7-0.08; F8- 0.08; F9-0.08; F10-0.10; F11- 0.10; F12; 0.11; F13-0.19); mandibles simple, just crossing at tip; labial palpi three-segmented, first long, second narrowing at apex and third wider. MESOSOMA. Shorter than metasoma (MsL = 0.75 mm); pronotum not elongate, epomia present; notauli deep and complete, convergent posteriorly; anterior scutellar pit suboval, wider than shortest distance between notauli; mesopleuron bare, with epicnemial pit and carina present; propodeum with plicae projecting posteriorly, forming teeth, median propodeal keel simple. Fore wing reduced, not reaching middle of propodeum (FwL = 0.32 mm). Hind wing shorter (HwL = 0.17 mm). Legs slender with few unorganized erect setae; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, slightly longer than wide (PtL = 0.20 mm; PtW = 0.18 mm), with distinct longitudinal sculpture, and short hairs dorsally; gaster subcylindrical, elongate and tapering at apex (GL = 1.11; GH = ca 0.40 mm); T2 and S2 longest; T2 slightly striated dorsally at junction with petiole; T6 and T7 not clearly separated and forming long triangle; ovipositor exerted, slightly shorter than gaster (OL = 0.89 mm). Male Unknown. Comments Using Nixon’s (1957) key, Pantolyta chemyrevae sp. nov. keys out to Pantolyta Förster, 1856 because of the following characters: brachypterous, toruli rims without a distinct cleft, antennae 15-segmented, epomia on pronotum defined at pronotal collar, notauli present on mesoscutum, lower side of gaster slightly down curved. With the keys to species of Pantolyta provided in Chemyreva & Kolyada (2019) and Nixon (1957), P. chemyrevae keys out near P. stylata Kieffer, 1908 (= P. vernalis sensu Nixon 1957) but has the antennal shelf less prominent and a longer scape (Chemyreva & Kolyada 2019). Among the Baltic amber species, it differs from other Pantolyta species as follow: P. antiqua is macropterous and has F1 more than two times as long as pedicel; Pantolyta augustinusii has the toruli separated by a distinct cleft, the mesoscutum bearing long hairs and the plicae not projecting posteriorly; P. janzeni and P. perrichoti has a longer petiole, more than two times as long as wide; P. somnulenta is very similar to P. chemyrevae sp. nov. but is macropterous, has anterior scutellar pit narrower than P. chemyrevae and the plicae less produced posteriorly., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 72-73, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Chemyreva V. G. & Kolyada V. A. 2019. Review of the Pantolyta genus (Hymenoptera: Diapriidae: Pantolytini) from Russia, with description of a new species. Zoosystematica Rossica 28: 163 - 176. https: // doi. org / 10.31610 / zsr / 2019.28.1.163"]}

Cinetus elongatus sp. nov. urn:lsid:zoobank.org:act: A7DAC194-D7C5-41B4-B925-6BE4250CB42C Figs 1C, 3A–C, Table 1 Diagnosis Scape longer than head length, F1 longest flagellomere, deeply emarginated on anterior half, wider at apex of emargination (male; Fig. 3A–B); metapleural carina extending from hind coxa and forked anteriorly (Fig. 3C); marginal vein slightly shorter than its distance from basal vein, radial cell elongate, longer than marginal vein, postmarginal vein 1.5 times as long as radial cell (Fig. 1C); petiole 5.3 times as long as wide, with a few long setae (Fig. 3C). Etymology The species name derives from the general elongate aspect of the species. The specific epithet is to be treated as an adjective. Type material Holotype NHMD-608402, a complete male but partially hidden by a milky coat. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. 4.46 mm. Head higher than long, homogeneously pubescent (HeL = 0.54 mm); eye oval; antenna inserted on distinct shelf; toruli broadly separated; antenna with numerous short hairs; scape longer than head length; pedicel more than two times as short as shortest flagellomere, almost as wide as long; flagellomeres cylindrical, elongate, much longer than wide; F1 longest, with deep emargination basally, wider at apex of emargination; F2–F11 shorter, slightly decreasing in length; F12 longer than F7–F11, conical (antennomeres length of holotype, in mm: Sc-0.61; P-0.12; F1-0.41; F2-0.38; F3-0.36; F4-0.34; F5-0.35; F6-0.33; F7-0.31; F8-0.32; F9-0.29; F10-0.29; F11-0.26; F12-0.33); mandibles of ordinary form, bidentate, slightly crossing at tip. MESOSOMA. Pubescent, with short setae (MsL = 1.52 mm); pronotum reduced in length, epomia present; mesoscutum large, convex, notauli complete, convergent anteriorly, slightly diverging posteriorly but terminating in anterior scutellar pit; anterior scutellar pit deep, large, suboval with posterior margin convex; metapleural carina distinct, forked anteriorly; propodeum with plicae, median propodeal keel simple. Fore wing extending beyond metasoma, homogeneously micro-pubescent (FwL = 3.89 mm); C, Sc+R, basal vein, M+Cu, M, Cu, marginal vein, postmarginal vein, r-rs and Rs fully pigmented; marginal vein slightly shorter than its distance from basal vein; r-rs well developed; Rs closing radial cell in straight line; radial cell elongate, longer than marginal vein; postmarginal vein 1.5 times as long as radial cell. Hind wing shorter and narrower than fore wing; C pigmented; basal cell opened; three hamuli present. Legs slender, bearing numerous un-organized setae; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole 5.3 times as long as wide (PtL = 0.79 mm; PtW = 0.15 mm), longitudinally ribbed, with sparse very long setae; gaster ellipsoidal (GL = 1.61 mm; GH = 0.68 mm), pointed at apex, with sparse setae on first segments, more numerous at apex; T2 and S2 longest; four ring-like segments posterior to large tergite, genitalia externalized. Female Unknown. Comments Using Nixon’s (1957) key, Cinetus elongatus sp. nov. keys out near Leptorhaptus Förster, 1865 sensu Nixon (1957) because of the following characters: fourth antennal segment not modified, mandibles of ordinary form, scutellum without a row of foveae along posterior margin, marginal vein hardly shorter than the radial cell, petiole much longer than wide, notauli subparallel, terminates in scutellar hollow. Leptorhaptus is currently regarded as a synonym of Cinetus (Masner 1964). Cinetus elongatus displays all the diagnostic characters of Cinetus provided by Quadros & Brandão (2017), especially the large anterior scutellar pit, the posterior extremity of notauli directed toward a point within the anterior scutellar pit, the marginal vein shorter than radial cell and slightly shorter than its distance to basal vein. Among the Baltic amber species, it differs from other Cinetus species as follow: C. balticus has the scape as long as F1 (longer in Cinetus elongatus sp. nov.), the petiole shorter (ratio length/middle height: 4 vs 5.3) and the marginal vein longer than its distance from basal vein; Cinetus breviscapus sp. nov. has the scape shorter than F1, the petiole shorter (ratio length/middle height: 2.8), the marginal vein longer than its distance from basal vein and the postmarginal vein shorter than radial cell; C. inclusus has a shorter petiole (ratio length/middle height: 1.5) and a smaller body (1.9 mm)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 66-68, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Nixon G. E. J. 1957. Hymenoptera, Proctotrupoidea, Diapriidae, subfamily Belytinae. Handbooks for the Identification of British Insects 8 (dii): 1 - 107. https: // doi. org / 10.5281 / zenodo. 23920","Masner L. 1964. A comparison of some Nearctic and Palaearctic genera of Proctotrupoidea (Hymenoptera) with revisional notes. Casopis Ceskoslovenske Spolecnosti Entomologicke 61: 123 - 155.","Quadros A. L. & Brandao C. R. F. 2017. Genera of Belytinae (Hymenoptera: Diapriidae) recorded in the Atlantic dense ombrophilous forest from Paraiba to Santa Catarina, Brazil. Papeis Avulsos de Zoologia 57 (6): 57 - 91. https: // doi. org / 10.11606 / 0031 - 1049.2017.57.06"]}

Doliopria baltica sp. nov. urn:lsid:zoobank.org:act: 36E9B60D-6971-454F-AD9F-DD69D430C16B Figs 1G, 6B–D, Table 1 Diagnosis Head globular, with sparse long hairs on vertex; apical segment of maxillary palpi bearing one long seta (Fig. 6C); antenna 11-segmented; scape as long as pedicel + F1–F5 combined, scape with apical rim slightly notched; pedicel cylindrical, elongate; F1 narrower; F2–F6 widening; F7–F9 forming distinct club; F7–F8 two times as wide as long; F9 as wide as long (Fig. 6B); anterior scutellar pit suboval, not divided; scutellum subquadrate; propodeum punctuate and carinate (Fig. 6D); hind wing bordered with long setae along posterior margin and distal part of fore margin; petiole cylindrical, slightly longer than wide; gaster ellipsoidal; tergite 2 with anterior margin excised (Fig. 6D). Etymology The species name refers to the origin of the amber piece containing the specimen. The specific epithet is to be treated as an adjective. Type material Holotype NHMD-608374, a complete female partially hidden by a milky coat. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 1.20 mm. Body smooth and shiny, sparsely pubescent, hairs very long. Head globular, slightly higher than long with hairs posteriorly (HeL = 0.23 mm); eye oval, glabrous; maxillary palpi 5-segmented with one long seta on apical segment; antenna 11-segmented inserted on transverse shelf of low elevation; toruli separated by shallow cleft; scape as long as pedicel + F1–F5 combined, as long as head, with apical circumference slightly notched; pedicel as wide as scape, cylindrical; F1 narrower than pedicel and longer than wide; F2–F5 slightly shortening and widening; F6 two times as wide as long; F7–F9 wider than long, longer than previous flagellomeres, forming clava; F9 ovoid (antennomeres length of holotype, in mm: Sc-0.19; P-0.05; F1-0.03; F2-0.02; F3-0.02; F4-0.03; F5-0.03; F6-0.03; F7- 0.07; F8-0.07; F9-0.10); mandibles short, only crossing at tips. MESOSOMA. Slightly shorter than metasoma (MsL = 0.43 mm); pronotum with lateral posterior margin straight; mesoscutum large and slightly convex, without notauli or any other sulci; anterior scutellar pit deep, not divided, suboval; scutellum subquadrate; propodeum minutely punctuate in anterior part and carinate. Fore wing extending beyond metasoma (FwL = 0.77 mm), micropubescent and bordered with short setae; only Sc+R present, slightly separate from anterior margin of wing, distally ending in slight thickening of marginal vein. Hind wing length two thirds of length of fore wing (HwL = 0.55 mm), micropubescent; with long setae along posterior margin and distal part of costal margin; three hamuli present; basal cell open. Legs slender, with only hind coxa and femur stouter; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole cylindrical, narrow, longer than wide (PtL = 0.10 mm; PtW = 0.06 mm), longitudinally striated, bearing several long hairs; gaster ellipsoidal (GL = 0.44 mm; GH = 0.21 mm), not sharply pointed at apex, smooth and glabrous; T2 and S2 longest; T2 covering at least two thirds of gaster, its anterior margin medially excised for distance subequal to petiole length; ovipositor slightly exerted. Male Unknown. Comments In Masner & García’s (2002) keys to the genera of Diapriinae, Doliopria baltica sp. nov. keys out to Doliopria Kieffer, 1910 because of the following characters: frons unarmed, antenna 11-segmented, notauli absent, wings fully developed, Sc+R separate from anterior margin of wing, basal vein absent. The description of the specimen is consistent with the original diagnosis of the genus by Kieffer (1910) and the revised diagnosis of Masner & García (2002). Doliopria baltica differs from extant representatives of this genus as follow: D. americana Fouts, 1926 has a two-segmented club (Fouts 1926); D. brachyptera Ogloblin, 1960 has F9 more lengthened and is brachypterous; D. foersteri Ogloblin, 1960 has short hairs on eyes, flagellomeres longer than wide and lacks the anterior scutellar pit; D. equatoriana Ogloblin, 1960 has F2–F5 longer than wide (Oglobin 1960); D. collegii Ferrière, 1929 has F9 longer (Loiácono et al. 2013); D. flavipes Kieffer, 1910 has F5–6 as long as wide and the petiole shorter (Kieffer 1910); D. myrmecobia Kieffer, 1921 has a two-segmented club and the apical flagellomere sharply pointed at the apex (Loiácono et al. 2013)., Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 77-79, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Masner L. & Garcia J. L. 2002. The genera of Diapriinae (Hymenoptera: Diapriidae) in the New World. Bulletin of the American Museum of Natural History 268: 1 - 138. https: // doi. org / c 7 zvsh","Kieffer J. J. 1910. Beschreibung neuer sudamerikanischer im Zoologischen Museum zu Berlin aufbewahrter Diapriiden. Entomologische Rundschau 1: 46 - 48. Available from https: // www. biodiversitylibrary. org / page / 43669771 [accessed 2 Jan. 2021].","Fouts R. M. 1926. Notes on Serphoidea with descriptions of new species (Hymenoptera). Proceedings of the Entomological Society of Washington 28 (8): 167 - 179. https: // doi. org / 10.5281 / zenodo. 24228","Ogloblin A. A. 1960. Tres especies nuevas del genero Doliopria del Ecuador (Diapriidae, Hymenoptera). Revista de la Sociedad Entomologica Argentina 22: 69 - 76.","Loiacono M. S., Margaria C. B. & Aquino D. A. 2013. Diapriinae wasps (Hymenoptera: Diaprioidea: Diapriidae) associated with ants (Hymenoptera: Formicidae) in Argentina. Psyche 2013: 320590. https: // doi. org / 10.1155 / 2013 / 320590"]}

Pantolyta similis sp. nov. urn:lsid:zoobank.org:act: 1E2F7650-238A-4AC8-9900-38A6B9DFB59F Fig. 5D–E, Table 1 Diagnosis Head as high as long; eye small but functional; antennal shelf projecting forward and upward; antenna with homogeneous short setae; pedicel conical, with base narrower; F1 second longest; F2–F13 gradually widening; F13 longest and widest (Fig. 5D); epomia absent; median propodeal keel simple (Fig. 5E); fore wing not extending beyond petiole (brachypterous morph); petiole 1.3 times as long as wide; gaster fusiform, elongate; T2 and S2 covering anterior half of gaster; T2 with numerous fine and long striations medially at junction with petiole (Fig. 5D). Etymology The specific epithet refers to its similarity with the extant species Pantolyta macrocera (Thomson, 1858) and is to be treated as an adjective. Type material Holotype NHMD-608468, a complete female. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Female BODY. BL = 3.52 mm. Head glabrous, smooth, as high as long (HeL = 0.50 mm); eye round, small, less than a third of head length (ED = 0.18 mm); frontal prominence elongate; toruli facing upwards; antenna slightly but homogeneously pubescent; scape longer than head length; pedicel conical, thinner at base; F1 second longest; F2–F12 of subequal length, gradually widening; F12 as long as wide; F13 widest and longest, conical (antennomeres length, in mm: Sc- 0.58 mm; P-0.11; F1-0.17; F2-0.10; F3-0.10; F4- 0.11; F5-0.11; F6-0.11; F7-0.12; F8-0.11; F9-0.11; F10-0.11; F11-0.11; F12-0.12; F13-0.21); mandibles of ordinary form. MESOSOMA. Shorter than metasoma (MsL = 1.10 mm); pronotum not elongate, with straight posterior margin, epomia absent; mesoscutum convex, with notauli deep and complete; anterior scutellar pit large, rounded; scutellum sub-quadrate, without posterior scutellar pits; propodeum dorsally with prominent plicae, median keel simple. Fore wing very reduced, hardly reaching posterior margin of petiole (FwL = 1.04 mm). Hind wing reduced, shorter than fore wing (HwL = 0.52 mm). Legs slender, with only hind coxa widened; erect setae along hind tibia and tarsomere; tibial spur formula 1-2-2; tarsal claws simple. METASOMA. Petiole 1.3 times as long as wide (PtL = 0.28 mm; PtW = 0.22 mm), longitudinally ribbed; gaster fusiform and elongate (GL = 1.64 mm; GH = 0.59 mm); S2 and T2 longest, covering anterior half of gaster; T2 striated medially at junction with petiole; three ring-like segments visible posteriorly to large tergite; apical sternites fused into long, smooth triangle; ovipositor exerted, shorter than gaster (OL = 0.74 mm). Male Unknown Comments The specimen was originally identified as the extant species Pantolyta macrocera (Thomson, 1858) by Buhl (2002) because of the brachypterous morph. However, we can observe some differences on the petiole and gaster. Pantolyta similis sp. nov. displays numerous fine longitudinal ribs dorsally on the petiole whereas there are fewer ribs in P. macrocera. Additionally, P. macrocera has shorter and stronger medial striations on T2 than Pantolyla similis (Chemyreva & Kolyada 2021: fig. 11g). Among the Baltic amber species, it differs from other Pantolyta species as follows: P. antiqua and P. somnulenta are macropterous and have a shorter petiole; Pantolyta augustinusii sp. nov. has the epomia present, whereas it is absent in Pantolyta similis sp. nov.; Pantolyta chemyrevae sp. nov. has the epomia present, the anterior scutellar pit more quadrate, the petiole more elongate and the striations on T2 shorter; P. janzeni and P. perrichoti has a longer petiole, more than two times as long as wide. Subfamily Diapriinae Haliday, 1833 Genus Basalys Westwood, 1833, Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 73-76, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167, {"references":["Buhl P. N. 2002. On a Baltic amber collection of Platygastridae and Diapriidae (Hymenoptera). Entomologiske Meddelelser 70: 57 - 61.","Chemyreva V. G. & Kolyada V. A. 2021. Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystemica Rossica 30: 137 - 162. https: // doi. org / 10.31610 / zsr / 2021.30.1.137"]}

Cinetus breviscapus sp. nov. urn:lsid:zoobank.org:act: CCD7DD88-2504-4697-9B70-80EC819B1C24 Figs 1B, 2E–F, Table 1 Diagnosis Head with sparse long hairs; antenna with uniform pilosity; scape less than five times as long as wide; pedicel rounded, as long as wide; flagellomeres cylindrical, elongate, longer than wide; F1 1.2 times as long as scape, deeply emarginated (Fig. 2F); anterior scutellar pits bean-shaped; propodeum with strong plicae; median propodeal keel simple; marginal vein longer than its distance from basal vein and as long as radial cell; postmarginal vein present on half radial cell; Rs directed toward base of wing; hind wing with basal cell open (Fig. 1B); hind tibia and tarsomeres with long recumbent hairs; petiole 2.8 times as long as high, with longitudinal ribs; gaster globose; T2 covering two thirds of gaster; S2 longer than T2 (Fig. 2E). Etymology The specific epithet is to be treated as an adjective and is composed of ‘ brevi- ’, meaning ‘short’, and ‘- scapus ’, a reference to the small size of the scape. Type material Holotype NHMD-300622, a complete male. Locality and horizon Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma. Description Male BODY. BL = 3.61 mm. Head with sparse long hairs (HeL = 0.47 mm); eye oval, glabrous; antennae inserted on transverse shelf; 14 antennomeres with uniform pilosity; scape elongate, less than five times as long as wide; pedicel rounded, as long as wide; flagellomeres elongate, cylindrical; F1 1.2 times as long as scape, with a deep emargination covering almost half of segment; F2–F6 slightly shorter, of subequal length; F6–F12 decreasing in size but still much more longer than wide; F12 tapering at apex (antennomeres length, in mm, when possible: Sc-0.32; P-0.06; F1-0.39; F2-0.34; F3-0.32); mandibles of ordinary form, not forming a beak. MESOSOMA. Shorter than metasoma (MsL = 1.21 mm); pronotum without epomia, only visible in lateral view; mesoscutum smooth; notauli deep, diverging at posterior terminations toward a point outside scutellum; anterior scutellar pit bean-shaped; scutellum without posterior scutellar pits; mesopleuron with epicnemial pit, epicnemial bridge and subalar bridge present; propodeum with prominent plicae; median propodeal keel simple. Fore wing hyaline, uniformly micropubescent (FwL= 3.58 mm); venation complete with C, Sc+R, basal vein, M+Cu, marginal vein, r-rs and Rs pigmented; marginal vein very long, longer than its distance from basal vein and as long as radial cell; postmarginal vein fading after middle of radial cell; Rs proximally directed toward base of wing; M inconspicuous towards apex; Cu short and nebulous; radial cell closed. Hind wing narrow, with C pigmented and basal cell opened (HwL = 2.23 mm). Legs slender, covered with setae; tibial spur formula 1-2-2; hind tibia and tarsomeres with long recumbent hairs. METASOMA. Petiole elongate, almost three times as long as its middle height (PtL = 0.56 mm; PtW = 0.19 mm), with longitudinal ribs; gaster globose, oval, smooth (GL = 1.37 mm; GH = 0.71 mm); six segments identifiable; T2 longest, covering two thirds of gaster; other tergites reduced, decreasing in length until they become indistinguishable; S2 longest, longer than T2; following sternites very short; pygidium and hypopygium at least as long as previous segments; genitalia internalized. Female Unknown. Comments Using Nixon’s (1957) key, Cinetus breviscapus sp. nov. keys out to Cinetus Jurine, 1807 because of the following characters: mandibles of ordinary form, scutellum without a row of foveae posteriorly, marginal vein very long, notauli slightly divergent posteriorly. Following Nixon’s (1957) key to male species of Cinetus, Cinetus breviscapus keys out near C. aletes Nixon, 1957 because of its scape that is shorter than F1 but differs in having the first flagellomere less distinctly longer than the scape (ratio scape length/F1 length: 1.2 vs 1.43; Nixon 1957), the petiole longer (three times as long as wide vs “about two and a half times as long as wide”) and being bigger (body length ca 3.6 mm vs ca 2.2 mm). In Baltic amber, Cinetus breviscapus sp. nov. is considered as separate from Cinetus inclusus Maneval, 1938 and Cinetus balticus Szabó & Oehlke, 1986 because it clearly does not fit with their descriptions. The petiole of C. balticus is longer than that of C. breviscapus (ratio length/middle height: 4 vs 2.8) and the fore wing venation is different (C. breviscapus has a longer marginal vein, a longer radial cell, the postmarginal vein does not extend beyond the radial cell and Rs is conspicuous for almost all its length). Cinetus inclusus has a shorter petiole (ratio length/middle height: 1.5) and the scape as long as pedicel, F1 and half of F2 combined, unlike C. breviscapus whose scape is shorter than flagellomere 1.

Acropiesta perrichoti sp. nov., a new belytine wasp is described based on a well preserved male specimen from Eocene Baltic amber. This is the second fossil species of this extant genus, known already from the same amber deposit by Acropiesta janzeni Buhl, 2002.

A new species of the parasitic wasp Coptera Say was previously distinguished from other species via correspondence between ecological (host) differences and DNA barcodes. A description and figures for Coptera tonicsp. nov., along with revisions to existing keys that allow it to be distinguished from other Nearctic species without the aid of molecular characters, is provided in this work.

Heraty et al. (2011) and Sharkey et al. (2012) defined their Proctotrupoidea s. str. as a monophyletic group, a clade, using analytical methods and including all relevant extant taxa. These analyses strongly supported the monophyly of their new concept of the superfamily Proctotrupoidea. It included seven families: Austroniidae, Heloridae, Pelecinidae, Peradeniidae, Proctotrupidae, Roproniidae and Vanhorniidae. Diapriidae and Monomachidae, formerly considered to be proctotrupoids, were placed (with other taxa) in the sister-group to Proctotrupoidea sensu stricto.

Trichopria anastrephae Lima and Pachycrepoideus vindemmiae (Rondani) naturally occur together with the pest Drosophila suzukii (Matsumura) in Brazil. Due to their co-occurrence, it is important to understand competitive interactions between these pupal parasitoids and how this may impact D. suzukii parasitism levels. Thus, the aim of the study was to evaluate the interspecific competition between T. anastrephae and P. vindemmiae. Bioassays were performed with and without time advantage, with different exposure (treatments) of D. suzukii pupae to parasitoid species. Verified that there was a higher parasitism and emergence rate of T. anastrephae adults. This was not observed for P. vindemmiae that decreases the parasitism rate in the presence of T. anastrephae. In strawberry fruits bioassay, T. anastrephae females showed capacity to parasitize pupae of D. suzukii inside strawberry fruits. However, P. vindemmiae had a lower parasitism rate and consequently, fewer insects emerged when used alone or associated with the presence of T. anastrephae. Therefore, we can conclude that there is interspecific competition between T. anastrephae and P. vindemmiae in D. suzukii pupae, with T. anastrephae having a higher competitive advantage.