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The persistence and size of the Greenland Ice Sheet (GrIS) through the Pleistocene is uncertain. This is important because reconstructing changes in the GrIS determines its contribution to sea level rise during prior warm climate periods and informs future projections. To understand better the history of Greenland's ice, we analyzed glacial till collected in 1993 from below 3 km of ice at Summit, Greenland. The till contains plant fragments, wood, insect parts, fungi, and cosmogenic nuclides showing that the bed of the GrIS at Summit is a long-lived, stable land surface preserving a record of deposition, exposure, and interglacial ecosystems. Knowing that central Greenland was tundra-covered during the Pleistocene informs the understanding of Arctic biosphere response to deglaciation., Competing Interests: Competing interests statement:The authors declare no competing interest.

Past interglacial climates with smaller ice sheets offer analogs for ice sheet response to future warming and contributions to sea level rise; however, well-dated geologic records from formerly ice-free areas are rare. Here we report that subglacial sediment from the Camp Century ice core preserves direct evidence that northwestern Greenland was ice free during the Marine Isotope Stage (MIS) 11 interglacial. Luminescence dating shows that sediment just beneath the ice sheet was deposited by flowing water in an ice-free environment 416 ± 38 thousand years ago. Provenance analyses and cosmogenic nuclide data and calculations suggest the sediment was reworked from local materials and exposed at the surface <16 thousand years before deposition. Ice sheet modeling indicates that ice-free conditions at Camp Century require at least 1.4 meters of sea level equivalent contribution from the Greenland Ice Sheet.

Understanding the history of the Greenland Ice Sheet (GrIS) is critical for determining its sensitivity to warming and contribution to sea level; however, that history is poorly known before the last interglacial. Most knowledge comes from interpretation of marine sediment, an indirect record of past ice-sheet extent and behavior. Subglacial sediment and rock, retrieved at the base of ice cores, provide terrestrial evidence for GrIS behavior during the Pleistocene. Here, we use multiple methods to determine GrIS history from subglacial sediment at the base of the Camp Century ice core collected in 1966. This material contains a stratigraphic record of glaciation and vegetation in northwestern Greenland spanning the Pleistocene. Enriched stable isotopes of pore-ice suggest precipitation at lower elevations implying ice-sheet absence. Plant macrofossils and biomarkers in the sediment indicate that paleo-ecosystems from previous interglacial periods are preserved beneath the GrIS. Cosmogenic 26 Al/ 10 Be and luminescence data bracket the burial of the lower-most sediment between <3.2 ± 0.4 Ma and >0.7 to 1.4 Ma. In the upper-most sediment, cosmogenic 26 Al/ 10 Be data require exposure within the last 1.0 ± 0.1 My. The unique subglacial sedimentary record from Camp Century documents at least two episodes of ice-free, vegetated conditions, each followed by glaciation. The lower sediment derives from an Early Pleistocene GrIS advance. 26 Al/ 10 Be ratios in the upper-most sediment match those in subglacial bedrock from central Greenland, suggesting similar ice-cover histories across the GrIS. We conclude that the GrIS persisted through much of the Pleistocene but melted and reformed at least once since 1.1 Ma., Competing Interests: The authors declare no competing interest.

Simulations of 21st century climate change for Great Britain predict increased seasonal precipitation that may lead to widespread soil loss by increasing surface runoff. Land use and different vegetation cover can respond differently to this scenario, mitigating or enhancing soil erosion. Here, by means of a sensitivity analysis of the PESERA soil erosion model, we test the potential for climate and vegetation to impact soil loss by surface-runoff to three differentiated British catchments. First, to understand general behaviours, we modelled soil erosion adopting regular increments for rainfall and temperature from the baseline values (1961-1990). Then, we tested future climate scenarios adopting projections from UKCP09 (UK Climate Projections) under the IPCC (Intergovernmental Panel on Climate Change) on a defined medium CO 2 emissions scenario, SRES-A1B (Nakicenovic et al., 2000), at the horizons 2010-39, 2040-69 and 2070-99. Our results indicate that the model reacts to the changes of the climatic parameters and the three catchments respond differently depending on their land use arrangement. Increases in rainfall produce a rise in soil erosion while higher temperatures tend to lower the process because of the mitigating action of the vegetation. Even under a significantly wetter climate, warmer air temperatures can limit soil erosion by enhancing primary productivity and in turn improving leaf interception, infiltration-capacity, and reducing soil erodibility. Consequently, for specific land uses, the increase in air temperature associated with climate change can modify the rainfall thresholds to generate soil loss, and soil erosion rates could decline by up to about 33% from 2070 to 2099. We deduce that enhanced primary productivity due to climate change can introduce a negative-feedback mechanism limiting soil loss by surface runoff as vegetation-induced impacts on soil hydrology and erodibility offset the effects of increased precipitation. The expansion of permanent vegetation cover could provide an adaptation strategy to reduce climate-driven soil loss., Competing Interests: Declaration of competing interest The authors declare that they have no known competing financial interests or personal relationships that could have appeared to influence the work reported in this paper., (Crown Copyright © 2020. Published by Elsevier B.V. All rights reserved.)

The Azores, Madeira, Selvagens, Canary Islands and Cabo Verde are commonly united under the term "Macaronesia". This study investigates the coherency and validity of Macaronesia as a biogeographic unit using six marine groups with very different dispersal abilities: coastal fishes, echinoderms, gastropod molluscs, brachyuran decapod crustaceans, polychaete annelids, and macroalgae. We found no support for the current concept of Macaronesia as a coherent marine biogeographic unit. All marine groups studied suggest the exclusion of Cabo Verde from the remaining Macaronesian archipelagos and thus, Cabo Verde should be given the status of a biogeographic subprovince within the West African Transition province. We propose to redefine the Lusitanian biogeographical province, in which we include four ecoregions: the South European Atlantic Shelf, the Saharan Upwelling, the Azores, and a new ecoregion herein named Webbnesia, which comprises the archipelagos of Madeira, Selvagens and the Canary Islands.

Coastal boulder deposits (CBD) are archives of extreme wave events. They are emplaced well above high tide, and may include megagravel clasts weighing tens or even hundreds of tonnes. But do they represent storms or tsunami? Many are interpreted as tsunami deposits based simply on clast size and inferences about transport, despite the fact that there are no direct observations documenting formation of these inbricated boulder clusters and ridges. In this study, we use force-balanced, dynamically scaled wave-tank experiments to model storm wave interactions with boulders, and show that storm waves can produce all the features of imbricated CBD. This means that CBD, even when containing megagravel, cannot be used as de facto tsunami indicators. On the contrary, CBD should be evaluated for inclusion in long-term storminess analysis.

A synthetic model is presented to enlarge the evolutionary framework of the General Dynamic Model (GDM) and the Glacial Sensitive Model (GSM) of oceanic island biogeography from the terrestrial to the marine realm. The proposed 'Sea-Level Sensitive' dynamic model (SLS) of marine island biogeography integrates historical and ecological biogeography with patterns of glacio-eustasy, merging concepts from areas as diverse as taxonomy, biogeography, marine biology, volcanology, sedimentology, stratigraphy, palaeontology, geochronology and geomorphology. Fundamental to the SLS model is the dynamic variation of the littoral area of volcanic oceanic islands (defined as the area between the intertidal and the 50-m isobath) in response to sea-level oscillations driven by glacial-interglacial cycles. The following questions are considered by means of this revision: (i) what was the impact of (global) glacio-eustatic sea-level oscillations, particularly those of the Pleistocene glacial-interglacial episodes, on the littoral marine fauna and flora of volcanic oceanic islands? (ii) What are the main factors that explain the present littoral marine biodiversity on volcanic oceanic islands? (iii) How can differences in historical and ecological biogeography be reconciled, from a marine point of view? These questions are addressed by compiling the bathymetry of 11 Atlantic archipelagos/islands to obtain quantitative data regarding changes in the littoral area based on Pleistocene sea-level oscillations, from 150 thousand years ago (ka) to the present. Within the framework of a model sensitive to changing sea levels, we discuss the principal factors affecting the geographical range of marine species; the relationships between modes of larval development, dispersal strategies and geographical range; the relationships between times of speciation, modes of larval development, ecological zonation and geographical range; the influence of sea-surface temperatures and latitude on littoral marine species diversity; the effect of eustatic sea-level changes and their impact on the littoral marine biota; island marine species-area relationships; and finally, the physical effects of island ontogeny and its associated submarine topography and marine substrate on littoral biota. Based on the SLS dynamic model, we offer a number of predictions for tropical, subtropical and temperate volcanic oceanic islands on how rates of immigration, colonization, in-situ speciation, local disappearance, and extinction interact and affect the marine biodiversity around islands during glacials and interglacials, thus allowing future testing of the theory., (© 2019 Cambridge Philosophical Society.)

Past climate changes provide important clues for advancement of studies on current global change biology. We have tested large-scale biogeographic patterns through four marine groups from twelve Atlantic Ocean archipelagos and searched for patterns between species richness/endemism and littoral area, age, isolation, latitude and mean annual sea-surface temperatures. Species richness is strongly correlated with littoral area. Two reinforcing effects take place during glacial episodes: i) species richness is expected to decrease (in comparison with interglacial periods) due to the local disappearance of sandy/muddy-associated species; ii) because littoral area is minimal during glacial episodes, area per se induces a decrease on species richness (by extirpation/extinction of marine species) as well as affecting speciation rates. Maximum speciation rates are expected to occur during the interglacial periods, whereas immigration rates are expected to be higher at the LGM. Finally, sea-level changes are a paramount factor influencing marine biodiversity of animals and plants living on oceanic islands., (Copyright © 2017 Elsevier Ltd. All rights reserved.)