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International audience; Bilebullephlebia legendrei n. gen., n. sp. is described, illustrated, and placed into the small epiproctophoran family Burmaphlebiidae. This family was previously known from the mid-Cretaceous Kachin amber by two monotypic genera, but the new fossil shows that its diversity is underestimated. At least one additional genus with an enigmatic wing venation remains to be described from the same deposit. Nothing is known about the genitalia configuration of the Burmaphlebiidae but new fossils are expected to clarify the position of the family which is currently only discussed in light of wing venation characters.

The representatives of the superfamily Evanioidea are easily distinguishable from all other extant and extinct Hymenoptera because of their metasoma (sometimes rounded) articulated high on the propodeum and well above the metacoxae (e.g., Goulet & Huber, 1993). Recent phylogenetic studies on the superfamily Evanioidea have shown strong support for its monophyly and for the monophyly of the Evaniidae even when fossil taxa are included (e.g., Li et al., 2018; Parslow et al., 2020; Jouault et al., 2022). According to the most recent analysis, the superfamily arose during either the Upper Triassic or the Lower Jurassic. Still, its earliest species are recorded in the younger Middle Jurassic, and its crown-group representatives during the Lower Cretaceous (Jouault et al., 2022: fig. 8). The stem Evaniidae are estimated to arise during the Upper Jurassic while their crown group has a more recent origin, likely around the Cretaceous-Paleogene boundary. Extant evaniid wasps are common, nearly cosmopolitan, and moderately diversified (about 580 extant species in 21 genera) even if this diversity is underestimated (Deans, 2005; Mullins et al., 2012). Little is known about the biology of extant evaniids, but their larvae are considered predators of cockroach eggs in oothecae (Huben, 1995). The Evaniidae have a good fossil record with the oldest species known from the late Hauterivian, numerous species documented up to the Miocene, and an important diversity in Burmese amber (e.g., Rasnitsyn et al., 1998; Nel et al., 2002; Deans et al., 2004; Jennings et al., 2012; Shih et al., 2020). Nevertheless, their past diversity is still underestimated.

International audience; A new genus and species of the archaeorthopteran order Caloneurodea are described and illustrated from the Salagou Formation (France) as Lodevogramma pumilia gen. et sp. nov. The particular wing venation of this species precludes its placement within one of the already described families of Caloneurodea. Consequently, the family Lodevogrammatidae fam. nov. is created to accommodate this specimen. A detailed comparison of the new species with other Caloneurodea is provided. This new species differs from all other Caloneurodea because it has: petiolate wings; the area between C and R/RA is very narrow; vein RA ends near the wing apex; vein RP has two branches; the fork of CuP basad the base of M; and two very short anal veins are present. This new species increases the diversity of Caloneurodea in the Salagou Formation and provides additional information on the diversity of the order around the late Capitanian extinction. We compared the diversity of Caloneurodea with that of Megasecoptera, another order with a similar history between the Carboniferous and the Permian, and hypothesized that the decrease in the size of both groups could be an indicator of their declines, possibly related to floral changes, following a pattern similar to that of Permian tetrapods. The decline of Caloneurodea could also be related to the diversification of the Orthoptera during the middle–late Permian.

The Palaeocene–Eocene Thermal Maximum (PETM) is one of the most important Cenozoic climatic events shaping modern biodiversity, yet reconstructions of its palaeobiomes remain controversial. Here we describe Gesomyrmex gallicus sp. nov., a new, extinct species of the ant genus Gesomyrmex Mayr, 1868, based on minor and major workers preserved in the early Eocene amber of Oise, France. Although there are only seven known extant species in the genus, all confined to warm and humid forests of SE Asia, the fossil record of Gesomyrmex indicates that the genus once radiated across Eurasia. The new species therefore provides direct evidence that this radiation likely co-occurred with the PETM. This observation constrains palaeoclimatic reconstructions of the Early Eocene by requiring the presence of an extensive, homogeneous and interconnected rainforest-like biome across palaeo-Eurasia, a scenario otherwise corroborated by pollen assemblages. The phases of regression of warm and humid forests and the widening of dry biomes probably occurred later during the Eocene, in between hyperthermals, and were probably less extensive than some computer models predicted.

The first vespid species from the Lower Cretaceous Crato formation (Brazil) is described and figured as Curiosivespa sennlaubi Jouault, Rasnitsyn & Nel, n. sp. This new species can be differentiate...

International audience; Two new peculiar mid-Cretaceous braconid genera, Megalyrhyssalus gen. nov. (type species Megalyrhyssalus clavicornis sp. nov.) and Stephanorhyssalus gen. nov. (type species Stephanorhyssalus longiscapus sp. nov.), are described and illustrated from Burmese amber. According to the particular morphology and wing venation of Megalyrhyssalus gen. nov. a new subfamily Megalyrhyssalinae subfam. nov. is created. The subfamily Seneciobraconinae is synonymised under Protorhyssalinae (syn. nov.). Additionally, a key to the genera of Braconidae having second recurrent vein (2m-cu) in forewing is provided.

Using a fossilized birth-death model, a new phylogeny of the superfamily Evanioidea (including ensign wasps, nightshade wasps or hatchet wasps) is proposed with estimates of divergence times for its constitutive families, also corroborating the monophyly of Evanioidea. Additionally, our Bayesian analyses demonstrate the monophyly of †Anomopterellidae, †Othniodellithidae, †An-dreneliidae, Aulacidae, Gasteruptiidae, and Evaniidae, while the †Praeaulacidae and †Baissidae ap-pear to be paraphyletic lineages. Vectevaniavetula and Hyptiogastriteselectrinus are transferred in the Aulacidae. We estimate the divergence of Evanioidea in the Late Triassic (~203 Ma). Addition-ally, three new othniodellithid wasps are described and figured from mid-Cretaceous Burmese am-ber as Keradellithabasilici Jouault, Marechal, Wang & Perrichot, gen. et sp. nov., Keradellithaanubis Jouault, Marechal, Wang & Perrichot, sp. nov., and Keradellithakirina Marechal, Jouault & Perrichot sp. nov. We also document a temporal shift, in relative specific richness, between Ich-neumonoidea and Evanioidea.

Deciphering the timing and tempo of lineage diversification of organisms has greatly benefited from advances in Bayesian phylogenetic analyses using morphological data. Those advances, however, have not been used for termites despite a rich fossil record. Here, we estimate divergence times for living and fossil termites using the fossilized birth–death (FBD) process on a previously published morphological matrix expanded with two new fossils that we describe (see Appendices 1 & 2). Those fossils, based on soldier specimens, are the ‘mid’-Cretaceous mastotermitid Milesitermesengeligen. et sp. nov., and the Middle Eocene Reticulitermesgrimaldiisp. nov. The latter is the oldest occurrence of a Rhinotermitidae soldier and the first termite soldier described from Baltic amber. Our dating estimates provide new stem- and crown-ages for termites, suggesting older ages than previously thought for several lineages. Importantly, crown-Isoptera—and therefore eusociality—may have arisen ~200 Ma. We conclude with further directions to keep improving our understanding of the timing of differentiation in termites.Page 2 of 77Systematic Entomology

Wing coloration is a very ancient feature among insects. Even the wings of the oldest known Pterygota showed transverse colored bands involved in a putative disruptive function. However, no evidence of wing coloration in the representatives of the superorder Odonatoptera is recorded before the latest Triassic. These were the only insect flying-predators until the pterosaurs began their diversification. Here we argue that the situation dramatically changed in the Early Jurassic, with the simultaneous appearance of Odonata with patterns of coloration in phylogenetically distant clades. It is especially the case in the Heterophlebiidae, a small family closely related to the Anisoptera, in which we could record no less than five different patterns of coloration in the same rather small area of North-Western Europe. At the same time and in the same area, small potentially insectivorous pterosaurs greatly diversified. The increase of the predation pressure on the Odonata is the most probable cause of the appearance of patterns of colored spots and bands on the dragonfly wings at that time. In the period between the Middle Jurassic to Early Cretaceous, the number of Odonata with spots and bands of color on wings dramatically increased, we assume in relation to the predation pressure due to an increasing diversification of insectivorous pterosaurs, but also small feathered dinosaurs and birds.

International audience; Deciphering the timing of lineage diversification and extinction has greatly benefited in the last decade from methodological developments in fossil-based analyses. If these advances are increasingly used to study the past dynamics of vertebrates, other taxa such as insects remain relatively neglected. Our understanding of how insect clades waxed and waned or of the impact of major paleoenvironmental changes during their periods of diversification and extinction (mass extinction) are rarely investigated. Here, we compile and analyze the fossil record of Plecoptera (1,742 vetted occurrences) to investigate their genus-level diversification and diversity dynamics using a Bayesian process-based model that incorporates temporal preservation biases. We found that the Permian-Triassic mass extinction has drastically impacted Plecoptera, while the Cretaceous Terrestrial Revolution corresponds with a turnover of plecopteran fauna. We also unveiled three major gaps in the plecopteran fossil record: the Carboniferous-Permian transition, the late Early Cretaceous, and the late Cenomanian to Bartonian, which will need to be further investigated. Based on the life history of extant Plecoptera, we investigate the correlations between their past dynamic and a series of biotic (Red Queen hypothesis) and abiotic (Court Jester hypothesis) factors. These analyses highlight the major role of continental fragmentation in the evolutionary history of stoneflies, which is in line with phylogeny-based biogeographic analyses showing how vicariance drove their diversification. Our study advocates analyzing the fossil record with caution, while attempting to unveil the diversification and extinction periods plus the likely triggers of these past dynamics of diversification.Deciphering the timing of lineage diversification and extinction has greatly benefited in the last decade from methodological developments in fossil-based analyses. If these advances are increasingly used to study the past dynamics of vertebrates, other taxa such as insects remain relatively neglected. Our understanding of how insect clades waxed and waned or of the impact of major paleoenvironmental changes during their periods of diversification and extinction (mass extinction) are rarely investigated. Here, we compile and analyze the fossil record of Plecoptera (1,742 vetted occurrences) to investigate their genus-level diversification and diversity dynamics using a Bayesian process-based model that incorporates temporal preservation biases. We found that the Permian-Triassic mass extinction has drastically impacted Plecoptera, while the Cretaceous Terrestrial Revolution corresponds with a turnover of plecopteran fauna. We also unveiled three major gaps in the plecopteran fossil record: the Carboniferous-Permian transition, the late Early Cretaceous, and the late Cenomanian to Bartonian, which will need to be further investigated. Based on the life history of extant Plecoptera, we investigate the correlations between their past dynamic and a series of biotic (Red Queen hypothesis) and abiotic (Court Jester hypothesis) factors. These analyses highlight the major role of continental fragmentation in the evolutionary history of stoneflies, which is in line with phylogeny-based biogeographic analyses showing how vicariance drove their diversification. Our study advocates analyzing the fossil record with caution, while attempting to unveil the diversification and extinction periods plus the likely triggers of these past dynamics of diversification.

Pristaulacus jarzembowskii sp. nov., oldest fossil of the crown group Aulacidae, is described from the Paleocene of Menat (France). The previously oldest fossil crown-aulacid was known from the Lowermost Eocene amber of Oise (France). The inferences of parasitoidism on Cerambycidae or Buprestidae together with the preferences of warm evergreen forest with wildfires are in accordance with the known data on the Menat paleobiota.

Rudiaeschna jarzembowskii sp. nov. (Figs 2, 3) urn:lsid:zoobank.org:act: 6C6E7BA5-E55D-4A27-9C21- A217FF2BF300 Material. Holotype NIGP180347 (a complete imprint of a forewing), housed in the Nanjing Institute of Geology and Palaeontology, Nanjing, China. Etymology. Named in honor of our friend and colleague Edmund A. Jarzembowski. The specific epithet is to be treated as a noun in a genitive case. Diagnosis. Forewing characters only. Eleven postnodal crossveins; discoidal triangle divided into four cells; three cells in subdiscoidal triangle; four cells between RP3/4 and MAa along wing margin; four crossveins in space between base of IR2, RP1/2, RP2 and first oblique crossvein; five rows of cells between MAa and MP; presence of a distal fork of RP2, below distal end of pterostigma. Type locality and horizon. The locality near the Liutiaogou village, Dashuangmiao Township, Ningcheng County, Inner Mongolia, China; Lower Cretaceous Yixian Formation (Barremian to Aptian). Description. A complete forewing, hyaline and very well-preserved; pterostigma dark brown; wing length 46.7 mm, 12.2 mm wide; distance between base and arculus 6.0 mm, between arculus and nodus 17.2 mm; distance from nodus to pterostigma 13.5 mm, from pterostigma to wing apex 6.5 mm; Ax1 is well preserved but Ax2 is weakened and very similar to the secondary antenodal crossveins; distance between Ax1 and wing base 5.0 mm; 20 antenodal crossveins; arculus angular; pterostigma elongate and narrow, 4.5 mm long and 1.0 mm wide, covering four cells, not basally recessed; pterostigmal brace strong, well aligned with basal side of pterostigma, moderately oblique; 11 postnodal crossveins not aligned with 11 postsubnodal crossveins; median space free; submedian space only traversed by CuP-crossing and one distal cubito-anal crossvein above PsA; PsA distinctly sigmoidal; hypertriangle with two oblique and weak crossveins; discoidal triangle elongate, four-celled; MAb straight, 3.9 mm long; a well-defined three-celled subdiscoidal triangle; base of IR2 4.7 mm basal of nodus; base of RP3/4 6.5 mm basal of nodus; four crossveins between RP and IR2 basal of first oblique vein ‘O’; first oblique vein ‘ O’ 0.5 mm distad subnodus and second oblique vein three cells distally; pseudo-IR1 very short, originating from RP1 below distal side of pterostigma; area between RP1 and RP2 rather narrow with two-three rows of cells between them; base of RP2 well aligned with subnodus, RP2 smoothly undulated in its mid part and with a strong fork below distal end of pterostigma; IR2 only slightly undulated, area between it and RP2 strongly widened at their undulated parts, with three rows of cells in its broadest part; IR2 and RP2 basally parallel, with only one row of cells; a well-developed and strongly curved Rspl with three rows of cells between it and IR2; two short convex oblique and undulating secondary veins anastomosing between IR2 and RP3/4 directly basal of origin of Rspl; RP3/4 and MAa parallel and smoothly undulated, with one row of cells between them basally, two rows of cells more distally, and four rows along posterior margin of wing; MAa and MP more or less parallel, postdiscoidal area weakly widened with five rows of cells at level of nodus; no Mspl; MA and CuA basally parallel with one row of rectangular cells between them, distally strongly divergent with at least three intercalary veins originating from MP; CuA with six posterior branches, cubito-anal area with seven rows of cells between CuA and posterior wing margin; two rows of cells in anal area.

Genus Rudiaeschna Ren & Guo, 1996 Type species. Rudiaeschna limnobia Ren & Guo, 1996 Other species. Rudiaeschna jarzembowskii sp. nov., Published as part of HUANG, DI-YING, CORENTIN, JOUAULT & NEL, ANDRÉ, 2022, The second species of Rudiaeschna (Odonata, Rudiaeschnidae) discovered in the Lower Cretaceous of Inner Mongolia, Northeast China, pp. 240-245 in Palaeoentomology 5 (3) on page 241, DOI: 10.11646/palaeoentomology.5.3.6, http://zenodo.org/record/6820841, {"references":["Ren, D. & Guo, Z. G. (1996) Three new genera and three new species of dragonflies from the Late Jurassic of Northeast China (Anisoptera: Aeshnidae, Gomphidae, Corduliidae). Entomologia Sinica, 3, 95 - 105."]}

Examination of new fossil specimens of Prosyntexis from the Lower Cretaceous Crato Formation of Brazil reveled a new species namely Prosyntexis sennlaubi sp. nov. To ensure the validity of the new taxon we examined previously described and figured specimens but we also figured an additional specimen of Prosyntexis gouleti Sharkey, 1990. The new species can be differentiated from the other Prosyntexis species of the Crato formation by its larger size but also in having the cell 2R1 small, the cell 2M small and short, the cell 3R1 narrow and the vein a directed toward wing apex. We performed a Geometric Morphometric Analysis (GMA) to estimate the variation in the forewing venation of the two species from the Crato Formation and ensure our placement.

Korehelorus jinjuensis gen. et sp. nov., a new genus and species of the family Heloridae, is described and figured from a specimen preserved in the dark grey shales of the lower Albian Jinju Formation (Republic of Korea). This description further increases the diversity of the family Heloridae in the Mesozoic and provides additional evidence for a rapid decline of the latter in the Cretaceous. To date, no explanations nor hypotheses have been proposed to explain this particular pattern but we assume and hypothesize that the species and genus diversities of helorid wasps are linked with the diversity of their hypothetical hosts (i.e., Chrysopidae or closely related extinct lineages). We show that the Cretaceous chrysopoid diversity is dominated by the Mesochrysopidae and stem-chrysopid subfamily (Limaiinae), which declined and/or die out through the Cretaceous. We find a peak of diversity for the Heloridae, synchronous with that of the Mesochrysopidae and Limaiinae, leading us to suggest that the decline of these clades in the Cretaceous consequently triggered a decline in the Heloridae.

The genus Ampulicomorpha Ashmead, 1893 (Hymenoptera: Embolemidae) currently includes 24 extant described species (Perkovsky et al., 2020). They are known from all the zoogeographic regions, excluding Antarctica (see Chény et al., 2020: fig. 1). Information about their biology is extremely reduced and needs further investigations. The few species whose biology has been studied parasitize nymphs of Achilidae (Hemiptera: Auchenorrhyncha) feeding on fungі inside rotten logs (Guglielmino & Bückle, 2013; Olmi et al., 2014b; Perkovsky et al., 2020).

Acropiesta perrichoti sp. nov., a new belytine wasp is described based on a well preserved male specimen from Eocene Baltic amber. This is the second fossil species of this extant genus, known already from the same amber deposit by Acropiesta janzeni Buhl, 2002.

The extant representatives of the order Mecoptera (except Siphonaptera now considered to belong to the Mecoptera: Tihelka et al., 2020) are famous among insects because of their long beaklike rostrum but also, because the males of the family Panorpidae have enlarged genitals raised over the body, and are called scorpionflies. The order Mecoptera is thought to have arisen during the Carboniferous period (Misof et al., 2014), a dating corroborated by their abundant fossil record during the Permian period (https://paleobiodb.org), and the presence of the antliophoran sister group Amphiesmenoptera during the late Carboniferous (Nel et al., 2007). The origin of the main family of Mecoptera i.e., the Panorpidae, was estimated to date to the Early Cretaceous (ca. 122.5 Ma) (Miao et al., 2019). The phylogenetic relationships within the family Panorpidae were recently investigated and revised using morphological data (Wang & Hua, 2021) and new hypotheses were proposed for their past dispersal events. Wang & Hua (2021: fig. 17) assumed that the Panorpidae originated from Asia and that at least two dispersal events have shaped the west European fauna. Here we describe a new specimen of scorpionfly from the Miocene of Murat (Cantal, France).

A new genus and two new species of Eremoneura are described from mid-Cretaceous amber of the Charentes region, in south-western France. Chimeromyia vulloi sp. nov., from early Cenomanian amber of Fouras, is the first record of Chimeromyiidae from France. It extends the geographical distribution of this family, which was hitherto known only from Cretaceous ambers of Lebanon, Spain, and Myanmar. A key to all known species of Chimeromyia is provided. Additionally, a specimen from latest Albian–earliest Cenomanian amber from Archingeay, which was studied using synchrotron imaging, is described as Francomyina incomparabilis gen. et sp. nov., and is left unassigned to family within Eremoneura. The affinities of both new species are discussed.

Palaeobiogeographic hypotheses need to be based on accurate palaeomaps and phylogenies of the concerned clades with a fossil record as complete as possible. Here we re-analyse the palaeobiogeography of the small fly family Scenopinidae, based on its first record in the mid-Cretaceous Burmese amber that we describe herein as Burmaprorates alagracilis gen. et sp. nov. and we assign to the subfamily Proratinae. At that time, the corresponding palaeobiota was an isolated island in the Southern part of the Tethys Ocean far away from the Laurasian plate and with general wind circulation unfavorable for aerial transport from Laurasia to the “Burmese Terrane”. Our results challenge and question the hypothesis of a North-Central American origin for the Scenopinidae at the beginning of the Cretaceous, and support a new hypothesis of a South Gondwanan origin.

A new wood wasp, Cratosirex sennlaubi gen. et sp. nov., is described and figured from one specimen collected from the Lower Cretaceous Crato Formation in northeastern Brazil. This new genus is placed in the new siricid subfamily Cratosiricinae subfam. nov., based on a combination of plesiomorphic and autapomorphic characters. The presence of small and sub-equal forewing cells 1R1 and 2R1 is a synapomorphy with the extant subfamily Siricinae, absent in the other extinct subfamilies †Auliscinae and †Gigasiricinae, supporting a sister group relationships with the Siricinae. Our new discovery expands the distribution range of Siricidae fossil records, highlights the antiquity of the family, and emphasizes the need for more studies of this particular insect lineage in the Mesozoic deposits. Currently, all the representatives of the crown group of the extant Siricidae are Cenozoic.

Two new bethylid wasps from Cenomanian Burmese amber, one tentatively placed in †Protopristocerinae: Cretapristocera longiscapa gen. & sp. nov. and one in †Holopsenellinae: Megalopsenella pouilloni gen. & sp. nov., are described and illustrated here. They provide new data on the Cretaceous diversity of the family and extend the Cenomanian Burmese amber records to six species. We discuss their systematic placements and provide identification keys to species of †Holopsenellinae and †Protopristocerinae to assist future taxonomic studies. We phylogenetically analyse Bethylidae, adding several fossil taxa to a recent study, suggesting a new hypothesis for the relationships between the different subfamilies. We also address the distribution and underestimated diversity of the family during the Cretaceous and the evolution of the venation of their wings. Finally, we discuss the possible misplacement of the families Plumariidae and †Falsiformicidae.

Examination of mid-Cretaceous Burmese amber reveals a new species of Atelestidae: Alavesia myanmarensis sp. nov., and the female of the dolichopodid Microphorites pouilloni Ngô-Muller & Nel, 2020. Both are described and illustrated. Alavesia myanmarensis sp. nov. is the first species of Alavesia from the mid-Cretaceous amber of northern Myanmar. The oldest records of this genus of small Diptera are from the Early to Late Cretaceous ambers of Spain, while the Burmese amber was probably produced on an island during the mid-Cretaceous, which had separated from Gondwana during the Jurassic. It suggests a possible Late Jurassic origin of the genus Alavesia.

International audience; A new specimen of the species Litholingia rhora Ren, 2002, from the Middle-Upper Jurassic deposit of Daohugou, is used to redescribe the species and precise its diagnosis. We also propose a series of wing venation drawings highlighting the main characteristics of the wing venation all Litholingia species. Finally, we summarize the current diversity of the lacewing family Grammolingiidae.

A new species, namely Doratomantispa pouilloni sp. nov., of the mantidfly subfamily Doratomantispinae, is described and illustrated from a complete individual preserved in mid-Cretaceous amber from Tanai, northern Myanmar. Doratomantispa pouilloni sp. nov. preserves nearly all the diagnostic characters of this subfamily, but differs from the two other species of the genus Doratomantispa, inter alia, in its forelegs with stout trochanter, femora with nine spines on outer edge, two spines on inner edge, tibia with parallel sides, probasitarsomeres with around six pairs of black and thick spines; wings elongate and narrow, forewings with costal veinlets simple before ScP and RA fusion, coloration pattern present. Doratomantispa pouilloni sp. nov. highlights the diversity and the niche specialization of mid-Cretaceous Mantispidae from Tanai amber.

International audience; A new species, namely Astioberotha coutreti sp. nov., of the thorny lacewing subfamily Paraberothinae, is described and illustrated from a complete individual preserved in mid-Cretaceous amber from Tanai, northern Myanmar. Astioberotha coutreti sp. nov. preserves nearly all the diagnostic characters of this subfamily, and differs from the type species of the genus Astioberotha owing to the fore femur with one long basal spine and 34 additional smaller spines; protibia bearing five distal spines; probasitarsus with two spines on inner edge located near mid-length; forewing with numerous maculation spots; intra-RP crossvein between RP3 and RP4 present (4rp3-rp4). Astioberotha coutreti sp. nov. together with the recent description of several other Paraberothinae species from the mid-Cretaceous Burmese amber, are used to discuss the palaeobiogeography of the subfamily.

International audience; A new fossil kalotermitid species of the genus Cryptotermes is described and illustrated as Cryptotermes pouilloni n.sp. from the Miocene Dominican amber (Dominican Republic). This new species can be differentiated from the otherCryptotermes species, inter alia, by its small size (one of the smallest Cryptotermes species), anterior margin of anteclypeusconcave, 15 antennomeres, and forewing RP with five secondary branches reaching wing anterior margin and M joining RPafter wing midlength. The new species originates from La Toca mines while the exact origins of the other Dominican fossilCryptotermes species are unknown. We summarize the Isoptera diversity from Dominican amber and rapidly discuss theimpact of inaccurate dating of the Dominican amber on the time divergence estimate of the isopteran constitutive families.