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The new genus Pumatiracia gen. nov. is described to accommodate a new species, P. venosa gen. et sp. nov. from Pu Mat National Park in Vietnam. The new genus is placed in the subtribe Parahiraciina of the Parahiraciini. It is compared with the genera Laohiracia Constant, 2021, Macrodarumoides Che, Zhang et Wang, 2012, Pseudochoutagus Che, Zhang et Wang, 2011, and Rostrolatum Che, Zhang et Wang, 2020 with which it shares possessing an elongate head. Illustrations of habitus, details, and male genitalia are given as well as a distribution map and photographs of the habitat. Laohiracia acuta Constant, 2021 is recorded for the first time from Vietnam, Pu Luong National Park; living specimens and habitat are illustrated, and the distribution map updated. The Parahiraciini fauna of Vietnam now comprises 14 species belonging to 11 genera.

The genus Hystatoderes Lameere, 1917 is reviewed and the species present in Vietnam are treated. The genus remained monotypic for a long time with H. weissi (Lameere, 1915) as the only species. However, further studies of recent material have differentiated three species within the genus, which are described and illustrated here. An identification key for male and female is provided. Hystatoderes vitalisi Lameere, 1917 is relieved from synonymy with H. weissi and considered a good species. The genus Laokaya Pic, 1928 is proposed as a junior synonym of Hystatoderes and its only species Laokaya vitalisi Pic, 1928 as a junior synonym of H. vitalisi. Hystatoderes antonkozlovi sp. nov. from Central Vietnam is described. The male of H. weissi is described for the first time. An updated list of species composing the genus Hystatoderes is given., Le genre Hystatoderes Lameere, 1917 est revu et les espèces présentes au Vietnam sont traitées. Le genre est resté longtemps monotypique avec comme seule espèce H. weissi (Lameere, 1915). De nouvelles études sur du matériel récent ont permis de différencier trois espèces dans le genre, qui sont décrites et illustrées ici. Une clé d’identification pour les mâles et femelles est fournie. Hystatoderes vitalisi Lameere, 1917 est relevé de la synonymie avec H. weissi et considéré comme une bonne espèce. Le genre Laokaya Pic, 1928 est proposé comme synonyme junior de Hystatoderes et son unique espèce Laokaya vitalisi Pic, 1928 comme synonyme junior d’Hystatoderes vitalisi. Une espèce nouvelle, H. antonkozlovi sp. nov., est décrite du centre du Vietnam. Le mâle de H. weissi est décrit pour la première fois. Une liste actualisée des espèces composant le genre Hystatoderes est donnée.

Cleotyche (Griseotyche) blanda Emeljanov, 2011 Figs 6, 16 Cleotyche blanda Emeljanov, 2011: 317 [described, compared with C. mariae]. Material examined Holotype (examined from photographs – Fig. 16) AUSTRALIA • ♀; [Western Australia]; White Gum Flat, Stirling Range N.P.;[34°23′47″S, 117°51′00″ E]; [25 Jan. 1979 – 6 Mar. 1979]; pit trap; “W.A., White Gum Flat, Stirling Range N.P., N° 1514, 25.I.1979 – 6.III.1979, pit trap ”; HMNH. Diagnosis Only species of the subgenus. It can be separated from all species of Cleotyche by the characters given to recognize the subgenus Griseotyche. Biology Unknown, however, White Gum Flat in the Stirling Range National Park is a picnic area in open forest, dominated by Eucalyptus wandoo Blakely (Myrtaceae). Distribution Australia, southwest Western Australia, Stirling Range National Park (Fig. 6)., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on page 91, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 2011. Improved tribal delimitation of the subfamily Dictyopharinae and description of new genera and new species (Homoptera, Fulgoroidea, Dictyopharidae). Entomological Review 91 (9): 1122 - 1145. https: // doi. org / 10.1134 / S 0013873811090053"]}

Genus Cleotyche Emeljanov, 1997 Cleotyche Emeljanov, 1997: 78 [described, compared to Caliscelis De Laporte, 1833, Parorgerioides de Bergevin, 1928 and Anorgeriopus Kusnezov, 1930]. Type species Cleotyche mariae Emeljanov, 1997, by monotypy and original designation. Diagnosis Only genus in the tribe. Description BODY. Pyriform; anterior part of body slightly compressed laterally. HEAD. Trigons (triangular area delimited by carinae at anterior angles of vertex) well delimited. Frons subrectangular, narrow, parallel-sided, much longer than wide, with three longitudinal carinae and often two additional incomplete carinae on dorsal portion. Vertex subquadrate, longer than wide, with median carina. Clypeus elongate and narrow, subtriangular. Apical segment of the labium half as long as subapical segment. THORAX. Pronotum smooth, elongate (slightly shorter than vertex along mid-line), tricarinate, with a single lateral keel on each side and posterior margin nearly straight. Mesonotum about ⅔ as long as pronotum, smooth with carinae nearly obsolete. TEGMINA. Strongly reduced, without carina, truncate apically and slightly widening from base to apex; clavus not defined. Tegulae absent. LEGS. Procoxae with triangular, lobe-shaped carina. Profemora and protibiae flattened and dilated, often foliate; with internal margin more or less evenly curved. Mesofemora and mesotibiae elongate and slender. Metatibiae with two lateral spines, one near tibiofemoral joint and one in distal half of tibia. Apex of metatibiae with 6 teeth separated in two groups: 2 teeth on internal side and 4 on external side. First and second segments of metatarsus with platellae instead of spines, except large lateral common teeth without setae. Arolium with one pair of chetoid sensilla. Claws with 3 setae. ABDOMEN. Dorsoventrally flattened. Dorsum of abdomen without carinae. MALE TERMINALIA. Pygofer narrow in lateral view, suboval and about much wider than high in posterior view; anterior and posterior margins rounded in lateral view; posterior margin deeply notched in dorsal and ventral view. Gonostyli rather compact, longer than high in lateral view, very elongate in ventral view; dorsal margin smoothly sinuate in lateral view; posteroventral margin rounded in lateral view; strong basidorsal lateral hook. Aedeagus elongate and narrow in dorsal view, with 2 pairs of membranous processes, each bearing a sclerotized spine; phallobase with 2 elongate sclerotized dorsal processes; connective elongate and narrow. Anal tube more or less oval in dorsal view with rounded apical margin; ventral margin sinuate; 2 strong apicolateral teeth directed ventrad. Species included (type locality in parentheses) Cleotyche (Griseotyche) blanda Emeljanov, 2011 (Stirling Range National Park, Western Australia) Cleotyche (Cleotyche) christinae sp. nov. (Cania Gorge National Park, Queensland) Cleotyche (Cleotyche) francescoi sp. nov. (Eurimbula National Park, Queensland) Cleotyche (Cleotyche) mariae Emeljanov, 1997 (Pearl Beach, New South Wales) Cleotyche (Cleotyche) montana sp. nov. (Blackdown Tableland National Park, Queensland) Identification key to the subgenera and species of Cleotychini Emeljanov, 1997 1. Ground colour pale grey (Fig. 16A); transverse white stripe on frons continuing laterally along body, becoming less distinct on abdomen (Fig. 16B–C); no white band along apical margin of tegmina (Fig. 16A) (subgenus Griseotyche)................................. C. (Griseotyche) blanda Emeljanov, 2011 – Ground colour dark brown (Fig. 1); no transverse white stripe on frons nor lateral white stripe along side of body (Fig. 2C); white band along apical margin of tegmina (Fig. 1) (subgenus Cleotyche) 2 2. Profemora and protibiae narrower, more than 3 × as long as broad (Fig. 1A, D; Table 1).............. 3 – Profemora and protibiae broader, less than 3 × as long as broad (Fig. 1B–C; Table 1)................... 4 3. Profemora more slender, 3.78 × as long as broad; tibiae slightly broader than femora; vertex 1.56 × as long as broad (Fig. 1A)............................................................. C. (Cleotyche) christinae sp. nov. – Profemora broader, 3.16 × as long as broad; tibiae slightly narrower than femora; vertex 2.10 × as long as broad (Fig. 1D).................................................................. C. (Cleotyche) montana sp. nov. 4. Profemora and protibiae broader, 2.55 and 2.73 × as long as broad, respectively; vertex less elongate, 1.71 × as long as than broad (Fig. 1B)......................................... C. (Cleotyche) francescoi sp. nov. – Profemora and protibiae narrower, 2.91 and 2.93 × as long as broad, respectively; vertex less elongate, 2.04 × as long as broad (Fig. 1C)........................ C. (Cleotyche) mariae Emeljanov, 1997, Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on pages 69-70, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 1997. A new genus and species of the Dictyopharidae from Australia belonging to a new tribe (Homoptera, Cicadina). Zoosystematica Rossica 6 (1 - 2): 77 - 82.","Emeljanov A. F. 2011. Improved tribal delimitation of the subfamily Dictyopharinae and description of new genera and new species (Homoptera, Fulgoroidea, Dictyopharidae). Entomological Review 91 (9): 1122 - 1145. https: // doi. org / 10.1134 / S 0013873811090053"]}

Cleotyche (Cleotyche) christinae sp. nov. urn:lsid:zoobank.org:act: 48303424-4988-454D-AA85-51B82488FEE5 Figs 2–6, Table 1 Diagnosis The species can be separated from the other species of Cleotyche (Cleotyche) by the combination of the following characters: 1. Profemora slender, 3.78 × as long as broad, and slightly narrower than tibiae (Figs 2A, 3A). 2. Vertex rather short, 1.56 × as long as broad (Figs 2F, 3F). 3. Vertex and anterior portion of pronotum dark brown (Figs 2F, 3F). Differential diagnosis This species differs from the three other species by the relatively slender profemora (profemora 3.78 × as long as broad vs max. 3.16 × in other species). The most similar species is C. (Cleotyche) montana sp. nov. which additionally differs from C. (Cleotyche) christinae sp. nov. by character 2 (vertex 2.10 × as long as broad vs 1.56 × in C. (Cleotyche) christinae sp. nov.). Etymology The species epithet refers to Dr Christine Lambkin (QM) in acknowledgement for all her proactive support of the authors’ research projects in Queensland. Type material Holotype AUSTRALIA • ♂; [Queensland]; Cania Gorge N.P.; 24°43′14″ S, 150°59′21″ E; 13 Dec. 2019; sweeping [grasses]; J. Constant, F. Martoni, M. Moir and L. Semeraro leg.; “ Australia Qld, Cania Gorge N.P., 24°43′14″S 150°59′21″E, 13.xii.2019, sweeping, leg. J. Constant, F. Martoni, M. Moir & L. Semeraro ”; QM. Paratype AUSTRALIA • 1 ♀; same collection data as for holotype; QM. Description MEASUREMENTS AND RATIOS. LT: ♂ (n = 1): 3.50 mm; ♀ (n = 1): 4.85. LTg/BTg = 1.39; LV/BV = 1.56; LF/BF = 3.12; LPf/BPf = 3.78; LPt/BPt = 3.34. HEAD (Figs 2F–H, 3F–H). Yellow-brown with vertex dark brown and clypeus black; labium yellow brown with distal part of terminal segment black. Vertex elongate, 1.56 × as long as broad, roundly pointed anteriorly, with complete median carina and with lateral margins slightly carinate and parallel; posterior margin roundly incurved. Frons elongate, straight in lateral view, with sides subparallel, 3.12 × as long as broad, anteriorly rounded in perpendicular view, with three carinae, one median and one along each lateral margin, all extending to apex of clypeus; two weak, short carinae between median and sublateral carinae extending from dorsal margin along 1 /5 of the way along the frons. Clypeus elongate and narrow, triangular. Eyes rather large, moderately protruding laterally. Antennae with scape short and cylindrical; pedicel short, inflated, barrel-shaped and with large sensory plates on ventral portion. Ocelli absent. Labium elongate and narrow, reaching posterior trochanters and with apical segment elongate, about half as long as penultimate one. THORAX (Figs 2E–H, 3E–H). Pronotum brown with posterior half whitish extending in a whitish band along posterior margin of paranotal lobes; mesonotum dark brown, darker than anterior portion of pronotum; thoracic sternites dark brown. Pronotum smooth with anterior margin strongly bisinuate, roundly projecting anteriorly behind vertex and roundly emarginate behind eyes, and posterior margin weakly incurved; median longitudinal carina and two lateral carinae on disc merging anteriorly along anterior margin; lateral carina behind eye; paranotal lobe broad, angularly rounded posteroventrally. Mesonotum very short, about ⅔ as long as pronotum, smooth with three very weakly marked obsolete carinae prolongating pronotal ones. Tegulae absent. TEGMINA (Figs 2A, C, F, H, 3A, C, F, H). Brown with rather broad white band along posterior margin, covering about 1 /5 of tegmina length; slightly elongate in dorsal view, 1.39 × as long as broad, slightly broadening from base to apex, truncate apically, convex, smooth; no trace of venation. LEGS (Figs 2A–D, I, 3A–D). Brown with pro- and mesocoxae and trochanters black; profemora paler than protibiae; apical pale yellowish interrupted ring on pro- and mesofemora; protibiae with basal pale yellow marking dorsally and ventrally; protarsi white; apex of metafemora and base of metatibiae darker; metatarsi brown with basal ⅔ of first metatarsomere and most of third one, whitish. Profemora and protibiae moderately foliaceous, rather slender, 3.78 and 3.34 × as long as broad, respectively; protibiae 1.15 × as broad as profemora; profemora with anterior margin straight and posterior margin broadly rounded, with 12–14 small teeth; protibiae with margins broadly rounded and external margin roundly truncate apically; median and posterior legs elongate and slender; metatibiae broadening towards apex, with one ventrolateral spine at distal 3 /5 and 6 apical spines; first and second metatarsomeres with strong spine at each side and apical row of 12 platellae ventrally. Metatibiotarsal formula: (2) 6 (2-4)/2/2. ABDOMEN (Figs 2A–D, 3A–D). Black with median yellowish marking on dorsal surface of last segment in male (Fig. 2A), on two last segments in female (Fig. 3A); small yellowish marking on anal tube at base of anal column. Abdomen dorsoventrally flattened and smooth. MALE TERMINALIA (Fig. 4). Pygofer (Py) (Fig. 4A–E) narrow in lateral view, 2.5 × as high as long at mid-height, suboval and about 1.4 × as wide as high in posterior view; anterior and posterior margins rounded in lateral view; posterior margin deeply notched in dorsal and ventral view, with V-shaped notch dorsally and U-shaped notch ventrally. Gonostyli (G) (Fig. 4A–E) rather compact, 1.63 × as long as high in lateral view, 3.4 × as long as wide at base in ventral view, slightly surpassing anal tube; dorsal margin smoothly sinuate in lateral view; posteroventral margin strongly rounded in lateral view; strong basidorsal lateral hook (lh) curved lateroventrad, with dorsal margin rounded in caudal view. Aedeagus (ae) (Fig. 4F–J) elongate and narrow in dorsal view, with 2 pairs of membranous processes, each bearing an anteapical lateral sclerotized spine (sp); phallobase (phb) with 2 elongate, narrow, straight, sclerotized dorsal processes (dpp); connective (cv) elongate and narrow. Anal tube (An) (Fig. 4A–E) in dorsal view obovate, 1.45 × as long as wide, widest at basal 2 /5, with apical margin rounded and with anal opening at basal 36% of length; ventral margin sinuate with posterior half regularly roundly concave before 2 strong apicolateral teeth (alt); teeth with posterior margin slightly concave in lateral view, rather narrow and slightly diverging in caudal view; length of teeth along posterior margin in lateral view equals ⅓ of length of anal tube. Biology The specimens were collected by sweeping grass in an open area in the valley of Three Moon Creek (Fig. 5). Distribution Australia, SE Queensland, Cania Gorge National Park (Fig. 6)., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on pages 71-77, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 1997. A new genus and species of the Dictyopharidae from Australia belonging to a new tribe (Homoptera, Cicadina). Zoosystematica Rossica 6 (1 - 2): 77 - 82."]}

Subgenus Griseotyche subgen. nov. urn:lsid:zoobank.org:act: 24EC3313-F915-45FD-894B-B6091307CBDE Type species and locality Cleotyche blanda Emeljanov, 2011 (by present designation); Stirling Range National Park, Western Australia. Species included Cleotyche (Griseotyche) blanda Emeljanov, 2011. Diagnosis Ground colour pale grey (brown in Cleotyche (Cleotyche)). Transverse whitish stripe on frons continued on side of body, becoming less distinct and greyish on abdomen (no transverse white stripe on face nor lateral white stripe on body in Cleotyche (Cleotyche)). No white band along apical margin of tegmina (present in Cleotyche (Cleotyche)). Small dark spots on all visible segments of the abdomen dorsally where the sensory pits occur (no dark spots on dorsum of abdomen in Cleotyche (Cleotyche) due to the very dark ground colouration of species). Etymology The new subgenus name is formed from ‘ griseus ’ (adj., Latin) meaning ‘grey’, and ‘- tyche ’, the ending of ‘ Cleotyche ’. It refers to the mostly grey colouration of the member of the new subgenus., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on page 91, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 2011. Improved tribal delimitation of the subfamily Dictyopharinae and description of new genera and new species (Homoptera, Fulgoroidea, Dictyopharidae). Entomological Review 91 (9): 1122 - 1145. https: // doi. org / 10.1134 / S 0013873811090053"]}

Cleotyche (Cleotyche) francescoi sp. nov. urn:lsid:zoobank.org:act: B3A245D8-CC75-487F-9D4B-1F1E825FFC5D Figs 6–10, Table 1 Diagnosis The species can be separated from the other Cleotyche (Cleotyche) species by the combination of the following characters: 1. Profemora very broad, only 2.55 × as long as broad, and slightly broader than protibiae (Figs 7A, 8A). 2. Vertex rather short, 1.71 × as long as broad (Figs 7F, 8F). 3. Vertex and anterior portion of pronotum pale brown (Figs 7F, 8F). Differential diagnosis This species differs from the three other species by its broad profemora (profemora 2.55 × as long as broad vs min. 2.91 × in the three other species). The most similar species is C. (Cleotyche) mariae Emeljanov, 1997 which additionally differs by character 2 (vertex more elongate, 2.04 × as long as broad vs 1.71 × in C. (Cleotyche) francescoi sp. nov.). Etymology The species epithet refers to Dr Francesco “ Bacon ” Martoni (VAIC) in acknowledgement for all his help and enthusiasm during the field work in Queensland in December 2019. Type material Holotype AUSTRALIA • ♂; [Queensland]; Eurimbula N.P., Ganoonga Noonga Lookout; 24°12′05″ S, 151°48′11″ E; 9 Dec. 2019; sweeping [grasses]; J. Constant, F. Martoni, M. Moir and L. Semeraro leg.; “ Australia Qld, Eurimbula N.P., Ganoonga Noonga Lookout, 24°12′05″S 151°48′11″E, 9.xii.2019, sweeping, leg. J. Constant, F. Martoni, M. Moir & L. Semeraro ”; QM. Paratypes AUSTRALIA • 4 ♂♂, 1 ♀; same collection data as for holotype; QM • 3 ♂♂, 1 ♀; same collection data as for holotype; DPIRD • 4 ♂♂; same collection data as for holotype; RBINS. Description MEASUREMENTS AND RATIOS. LT: ♂ (n = 2): 4.04 mm (3.94–4.14); ♀ (n = 1): 4.69. LTg/BTg = 1.12; LV/ BV = 1.71; LF/BF = 3.65; LPf/BPf = 2.55; LPt/BPt = 2.73. HEAD (Figs 7F–H, 8F–H). Yellow-brown with clypeus glossy black; labium brown with terminal segment black. Vertex elongate, 1.71 × as long as broad, projected anteriorly, rounded to a blunt point apically, with median carina not reaching anterior margin and with lateral margins carinate and incurved; posterior margin roundly concave. Frons elongate, with sides subparallel, weakly narrowing along eyes, 3.65 × as long as broad, straight in lateral view, anteriorly rounded, projected to a blunt point in perpendicular view, with three complete carinae, one median and one along each lateral margin, all extending to apex of clypeus; two weak, short carinae between median and sublateral carinae extending from dorsal margin along 1 /5 of the way along the frons. Clypeus elongate and narrow, triangular. Eyes rather large, moderately protruding laterally. Antennae with scape short and cylindrical; pedicel short, inflated, barrel-shaped and with large sensory plates on ventral portion. Ocelli absent. Labium elongate and narrow, reaching posterior trochanters and with apical segment elongate, about half as long as penultimate one. THORAX (Figs 7E–H, 8E–H). Pronotum yellow-brown with posterior half whitish extending in a whitish band along posterior margin of paranotal lobes; mesonotum dark brown, darker than anterior portion of pronotum, with apex of scutellum slightly paler; thoracic sternites glossy black. Pronotum smooth with anterior margin strongly bisinuate, roundly projecting anteriorly behind vertex and roundly emarginate behind eyes, and posterior margin weakly incurved; median longitudinal carina and two lateral carinae on disc merging anteriorly along anterior margin; lateral carina behind eye; paranotal lobe angularly rounded posteroventrally. Mesonotum very short, about ⅔ as long as pronotum, smooth with three weakly marked obsolete carinae prolongating pronotal ones. Tegulae absent. TEGMINA (Figs 7A, C, F, H, 8A, C, F, H). Brown as mesonotum with rather broad white band along posterior margin, covering about ¼ of tegmina length; slightly elongate in dorsal view, 1.12 × as long as broad, slightly broadening from base to apex, truncate apically, slightly rounded, particularly along lateral apical margin, convex, smooth; no venation apparent. LEGS (Figs 7A–D, I, 8A–D, I). Dark brown to black with pro- and mesocoxae and trochanters black; profemora slightly paler basally; apical pale yellowish interrupted ring on pro- and mesofemora; protibiae with basal pale yellow marking dorsally and ventrally; protarsi white; apex of metafemora and base of metatibiae darker; metatibiae turning yellowish from base to apex; metatarsomeres whitish except brown posterior portion of second one. Profemora and protibiae foliaceous, broad, 2.55 and 2.73 × as long as broad, respectively; protibiae 0.96 × as broad as profemora; profemora with anterior margin weakly curved and posterior margin broadly rounded, with 14–16 small teeth; protibiae with margins broadly rounded and external margin roundly truncate apically; median and posterior legs elongate and slender; metatibiae broadening towards apex, with one ventrolateral spine at distal 3 /5 and 6 apical spines; first and second metatarsomeres with strong spine at each side and apical row of 12 platellae ventrally. Metatibiotarsal formula: (2) 6 (2-4)/2/2. ABDOMEN (Figs 7A–D, 8A–D). Glossy black with median yellowish marking on last segment in male, on two last segments in female; small yellowish marking on anal tube at base of anal column. Abdomen dorsoventrally flattened and smooth. MALE TERMINALIA (Fig. 9). Pygofer (Py) (Fig. 9A–E) narrow in lateral view, 2.9 × as high as long at midheight, suboval and about 1.25 × as wide as high in posterior view; anterior and posterior margins rounded in lateral view; posterior margin deeply notched in dorsal and ventral view, with roundly V-shaped notch dorsally and U-shaped notch ventrally. Gonostyli (G) (Fig. 9A–E) moderately elongate, 1.75 × as long as high in lateral view, 3.4 × as long as wide at base in ventral view, slightly surpassing anal tube; dorsal margin sinuate in lateral view; posteroventral margin regularly rounded in lateral view; strong basidorsal lateral hook (lh) curved lateroventrad, with dorsal margin strongly rounded in caudal view. Aedeagus (ae) (Fig. 9F–J) elongate and narrow in dorsal view, with 2 pairs of membranous processes, each bearing an anteapical lateral sclerotized spine (sp); phallobase (phb) with 2 elongate, narrow, straight, sclerotized dorsal processes (dpp); connective (cv) elongate and narrow. Anal tube (An) (Fig. 9A–E) in dorsal view obovate, 1.4 × as long as wide, widest at basal 2 /5, with apical margin rounded and with anal opening at basal ⅓ of length; ventral margin straight, abruptly rounded at straight angle before strong apicolateral teeth (alt); teeth with posterior margin weakly concave in lateral view, narrow and slightly diverging in caudal view; length of teeth along posterior margin in lateral view equals 0.47 × length of anal tube. Biology The specimens were collected by sweeping grass in and around the trail to Ganoonga Noonga Lookout, in relatively open Eucalyptus woodland (Fig. 10). Distribution Australia, SE Queensland, Eurimbula National Park (Fig. 6)., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on pages 77-83, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 1997. A new genus and species of the Dictyopharidae from Australia belonging to a new tribe (Homoptera, Cicadina). Zoosystematica Rossica 6 (1 - 2): 77 - 82."]}

Cleotyche (Cleotyche) mariae Emeljanov, 1997 Figs 6, 11–12, Table 1 Diagnosis The species can be separated from the other Cleotyche (Cleotyche) species by the combination of the following characters: 1. Profemora rather broad, only 2.91 × as long as broad, and slightly narrower than protibiae (Fig. 1C). 2. Vertex rather elongate, 2.04 × as long as broad (Figs 11B, 12B). 3. Vertex and anterior portion of pronotum pale brown (Figs 11B, 12B). Differential diagnosis This species differs from C. (Cleotyche) christinae sp. nov. and C. (Cleotyche) montana sp. nov., by the proportions of the profemora (profemora 2.91 × as long as broad vs min. 3.16 × in the two other species). The most similar species is C. (Cleotyche) francescoi sp. nov. which additionally differs by having broader profemora (2.55 × as long as broad vs 2.91 × in C. (Cleotyche) mariae) and less elongate vertex (1.71 × as long as broad vs 2.04 × in C. (Cleotyche) mariae). Material examined Holotype (examined from photographs – Fig. 11) AUSTRALIA • ♂; [New South Wales, Pearl Beach]; [33°32′59″ S, 151°17′57″ E]; [13 Mar. 1997]; Emeljanov leg.; “Australia, NSW, Pearl Beach, 13.III.1997, Emeljanov”; “Holotypus Cleotyche mariae Emeljanov 1997 ”; “ANIC Database No. 20 011325”; ANIC. Paratype (examined from photographs – Fig. 12) AUSTRALIA • ♀; [New South Wales, Pearl Beach]; [33°32′59″ S, 151°17′57″ E]; [9 Mar. 1997]; Emeljanov leg.; “Australia, NSW, Pearl Beach, 9.III.1997, Emeljanov”; “Paratypus Cleotyche mariae Emeljanov 1997 ”; “ANIC Database No. 20 011326”; ANIC. Supplementary description MEASUREMENTS AND RATIOS. LTg/BTg = 1.17; LV/BV = 2.04; LF/BF = 3.94; LPf/BPf = 2.91; LPt/ BPt = 2.93. Distribution Australia, New South Wales, Pearl Beach (Fig. 6)., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on pages 83-86, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 1997. A new genus and species of the Dictyopharidae from Australia belonging to a new tribe (Homoptera, Cicadina). Zoosystematica Rossica 6 (1 - 2): 77 - 82."]}

Cleotyche (Cleotyche) montana sp. nov. urn:lsid:zoobank.org:act: 40E65B47-662A-42CA-80BC-239F869205EB Figs 6, 13–15, Table 1 Diagnosis The species can be separated from the other Cleotyche (Cleotyche) species by the combination of the following characters: 1. Profemora rather slender, 3.16 × as long as broad, and slightly broader than protibiae (Fig. 13A). 2. Vertex elongate, 2.10 × as long as broad (Fig. 13F). 3. Vertex and anterior portion of pronotum medium brown (Figs 13F). Differential diagnosis This species differs from C. (Cleotyche) francescoi sp. nov. and C. (Cleotyche) mariae Emeljanov, 1997 by its relatively more slender profemora (profemora 3.16 × as long as broad vs max. 2.91 × in the two other species). The most similar species is C. (Cleotyche) christinae sp. nov. which differs by its narrower profemora (3.78 × as long as broad vs 3.16 × in C. (Cleotyche) montana sp. nov.), and the proportions of the vertex (vertex 2.10 × as long as broad vs 1.56 × in C. (Cleotyche) christinae sp. nov.). Etymology The species epithet is a Latin adjective meaning ‘associated with mountain’ and refers to the mountainous habitat on top of Blackdown Tableland where the species was discovered. Type material Holotype AUSTRALIA • ♂; [Queensland, Blackdown Tableland N.P.]; [11–12 Mar. 2020]; 23°42′48″ S, 149°07′06″ E; J. Constant and L. Semeraro leg.; “Australia Qld, Blackdown Tableland N.P., 11–12 Mar 2020, 23°42′48″S 149°07′06″E, leg. J. Constant & L. Semeraro, Leopold III Funds Expedition”; QM. Description MEASUREMENTS AND RATIOS. LT: ♂ (n = 1): 3.9 mm. LTg/BTg = 1.32; LV/BV = 2.10; LF/BF = 3.39; LPf/ BPf = 3.16; LPt/BPt = 3.52. HEAD (Fig. 13F–H). Rufous-brown with clypeus black; labium brown with terminal segment black. Vertex elongate, 2.10 × as long as broad, projected anteriorly, rounded to a blunt point apically, with median carina not reaching anterior margin and with lateral margins carinate, subparallel along eye and incurved anteriorly; posterior margin roundly incurved. Frons elongate, very weakly concave in lateral view, with sides subparallel, weakly narrowing along eyes, 3.29 × as long as broad, anteriorly pointly rounded in perpendicular view, with three complete carinae, one median and one along each lateral margin, all extending to apex of clypeus; two weak, short carinae between median and sublateral carinae extending from dorsal margin along 1 /5 of the way along the frons. Clypeus elongate and narrow, triangular. Eyes rather large, moderately protruding laterally. Antennae with scape short and cylindrical; pedicel short, inflated, barrel-shaped and with large sensory plates on ventral portion. Ocelli absent. Labium elongate and narrow, reaching posterior trochanters and with apical segment elongate, about half as long as penultimate one. THORAX (Fig. 13E–H). Pronotum rufous-brown with posterior half whitish, paranotal lobes slightly darker towards posterior and ventral margins; mesonotum dark brown, darker than anterior portion of pronotum, with apex of scutellum slightly paler; thoracic sternites dark brown. Pronotum smooth with anterior margin strongly bisinuate, roundly projecting anteriorly behind vertex and roundly emarginate behind eyes, and posterior margin weakly incurved; median longitudinal carina and two lateral carinae on disc merging anteriorly along anterior margin; lateral carina behind eye; paranotal lobe angularly rounded posteroventrally. Mesonotum very short, about ⅔ as long as pronotum, smooth with three hardly visible obsolete carinae prolongating pronotal ones. Tegulae absent. TEGMINA (Fig. 13A, C, F, H). Rufous-brown as mesonotum with rather broad white band along posterior margin, covering slightly less than 1 /5 of tegmina length; slightly elongate in dorsal view, 1.32 × as long as broad, slightly broadening from base to apex, broadly truncate apically, slightly rounded along lateral apical margin, convex, smooth; no trace of venation. LEGS (Fig. 13A–D, I). Brown with profemora along lateral margins, protibiae and metafemora dark brown; pro- and mesocoxae and trochanters black-brown; apical pale yellowish interrupted ring on pro- and mesofemora; protibiae with basal pale yellow marking dorsally and ventrally; protarsi white; base of metatibiae darker; metatibiae turning yellowish from base to apex; metatarsomeres brown with first one whitish on basal ⅔. Profemora and protibiae moderately foliaceous, rather slender, 3.16 and 3.52 × as long as broad, respectively; protibiae 0.92 × as broad as profemora; profemora with anterior margin weakly curved and posterior margin broadly rounded, with about 15 small teeth, teeth progressively smaller towards base of femur; protibiae with margins broadly rounded, external margin rather straight along median ⅔, and roundly truncate apically; median and posterior legs elongate and slender; metatibiae broadening towards apex, with one ventrolateral spine at distal 3 /5 and 6 apical spines; first and second metatarsomeres with strong spine at each side and apical row of 12 platellae ventrally. Metatibiotarsal formula: (2) 6 (2-4)/2/2. ABDOMEN (Fig. 13A–D). Black with median yellowish marking on last segment in male (female unknown); small yellowish marking on anal tube at base of anal column. Abdomen dorsoventrally flattened and smooth. MALE TERMINALIA (Fig. 14). Pygofer (Py) (Fig. 14A–E) narrow in lateral view, 2.7 × as high as long at mid-height, suboval and about 1.25 × as wide as high in posterior view; anterior and posterior margins rounded in lateral view; posterior margin deeply notched in dorsal and ventral view, with rounded notch dorsally and U-shaped notch ventrally. Gonostyli (G) (Fig. 14A–E) rather elongate, 1.9 × as long as high in lateral view, 3.4 × as long as wide at base in ventral view, slightly surpassing anal tube; dorsal margin sinuate in lateral view; posteroventral margin straight in basal ⅓ then rounded in lateral view; strong basidorsal lateral hook (lh) curved lateroventrad, with dorsal margin oblique in caudal view. Aedeagus (ae) (Fig. 14F–J) elongate and narrow in dorsal view, with 2 pairs of membranous processes, each bearing an anteapical lateral sclerotized spine (sp); phallobase (phb) with 2 elongate, narrow, straight, sclerotized dorsal processes (dpp); connective (cv) elongate and rather strong.Anal tube (An) (Fig. 14A– E) in dorsal view obovate, 1.45 × as long as broad in dorsal view, widest at basal ⅓, with apical margin rounded and with anal opening at basal ⅓ of length; ventral margin straight, strongly rounded before strong apicolateral teeth (alt); teeth with posterior margin weakly concave in lateral view, narrow and subparallel in caudal view; length of teeth along posterior margin in lateral view 0.47 × as long as anal tube. Biology The specimen was collected by sweeping grass in an open Eucalyptus forest up the plateau, not far from Yaddamen Dhina – Horseshoe Lookout (Fig. 15). Distribution Australia, SE Queensland, Blackdown Tableland National Park (Fig. 6)., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on pages 86-90, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 1997. A new genus and species of the Dictyopharidae from Australia belonging to a new tribe (Homoptera, Cicadina). Zoosystematica Rossica 6 (1 - 2): 77 - 82."]}

Cleotyche (Cleotyche) Emeljanov, 1997 Type species Cleotyche mariae Emeljanov, 1997, by present designation. Diagnosis Ground colour brown to dark brown (pale grey in Cleotyche (Griseotyche)). No transverse white stripe on face nor lateral white stripe on body (transverse whitish stripe on frons continues along side of body, becoming less distinct and greyish on abdomen in Cleotyche (Griseotyche)). White band along apical margin of tegmina (absent in Cleotyche (Griseotyche)). No visible dark spots on segments of the abdomen dorsally where the sensory pits occur (small dark spots on all visible segments of the abdomen dorsally where the sensory pits occur in Cleotyche (Griseotyche))., Published as part of Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), pp. 66-95 in European Journal of Taxonomy 836 on page 70, DOI: 10.5852/ejt.2022.836.1917, http://zenodo.org/record/7061621, {"references":["Emeljanov A. F. 1997. A new genus and species of the Dictyopharidae from Australia belonging to a new tribe (Homoptera, Cicadina). Zoosystematica Rossica 6 (1 - 2): 77 - 82."]}