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Three new species of the Clubiona corticalis group in China are described: Clubiona bidactylinasp. nov., C. camelasp. nov., and C. subhuimingsp. nov.

Clubiona flammaforma sp. nov. Figs 1, 2, 7 Type material. Holotype: ♁, CHINA: Hubei Province, Enshi City, Badong county, Yanduhe Town, Songziyuan Village, 31.35279°N, 110.39937°E, 1836 m, 27.IV.2016, W. Liu, T. Tian & C. Zeng leg. (HNU-HB-IV-1607). Paratypes: 1♁, same data as holotype (HNU-HB-IV-1607); 2♀, same place as holotype, 31.35067°N, 110.42625°E, 1340 m, 28.IV.2016, W. Liu et al. leg. (HNU-HB-IV-1608). Etymology. The specific epithet is the combination of the Latin flamma (fire) and formis (form), referring to the flame-shaped RTA, adjective. Diagnosis. Clubiona flammaforma sp. nov. resembles C. rostrata Paik, 1985 (see Zhu & Zhang 2011: fig. 266A–E) in having similar course of both embolus in male and copulatory ducts in female, but can be distinguished by: (1) the RTA trifurcate (Fig 1F) (vs. uniramous, with a ventral protuberance in C. rostrata); (2) the embolic base with a stronger dentiform process extended upwards distally in ventral view (Fig 1D) (vs. with a weak tooth extended horizontally in C. rostrata); (3) the posterior margin of epigyne recurve, without median excavation (Fig 2C) (vs. margine procurve, with oblong excavation in C. rostrata); (4) the proximal portion of copulatory ducts separated from each other and arc-shaped (Fig 2D) (vs. close to each other and straight in C. rostrata). Description. Male (holotype) (Fig. 1A, B). Total length 3.91; carapace 1.98 long, 1.35 wide; abdomen 2.15 long, 1.12 wide. Carapace dark yellow; fovea reddish and longitudinal, cervical groove inconspicuous, radial grooves visible. Eyes: AER slightly recurved, PER slightly precurved in dorsal view. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.12, PLE 0.10, AME–AME 0.04, AME–ALE 0.03, PME–PME 0.17, PME–PLE 0.10, MOQL 0.20, MOQA 0.26, MOQP 0.42. Chelicerae yellowish, with five promarginal teeth and seven retromarginal teeth. Endites and labium yellow. Sternum yellow, almost oval. Legs yellow; tibiae I and II with two pairs of ventral spines, metatarsus I and II with a pair of ventral spines. Leg measurements: I 4.04 (1.13, 1.76, 0.77, 0.45), II 4.22 (1.19, 1.77, 0.81, 0.45), III 3.59 (1.14, 1.25, 0.78, 0.42), IV 5.84 (1.91, 1.88, 1.59, 0.46). Abdomen elongate oval, dorsum yellowish white, with brown pinnate pattern posteriorly and two pairs of muscular depressions; venter yellowish white. Spinneret yellowish white. Palp (Fig. 1C–G). Femur and patella without apophysis. Tibia as long as patella and about 1/3 length of cymbium, RTA longer than tibia, sclerotized and trifurcated, flame-shaped, ventral branch tuberculate with blunt tip, lateral branch strongest with pointed tip, dorsal branch spiniform with pointed tip. Cymbium longer than wide. Tegulum relatively flattened, longer than wide. Conductor groovelike, membranous, located distally on retrolateral side of tegulum. Embolus slender, angled across the distal end of tegulum and extended to about 2/3 length of tegulum; embolic base broad with a stronger dentiform process extended upwards distally. Sperm duct slender and twisted in ventral view. Female (one of HNU-HB-IV-1608) (Fig. 2A–B). Total length 4.82. Carapace 2.09 long, 1.50 wide; abdomen 2.71 long, 1.58 wide. Eye sizes and interdistances: AME 0.13, ALE 0.14, PME 0.14, PLE 0.12, AME–AME 0.08, AME–ALE 0.06, PME–PME 0.22, PME–PLE 0.13, ALE–PLE 0.07, MOQL 0.27, MOQA 0.32, MOQP 0.47. Leg measurements: I 3.76 (1.14, 1.54, 0.70, 0.38), II 4.06 (1.26, 1.66, 0.72, 0.42), III 3.73 (1.14, 1.30, 0.90, 0.39), IV 5.77 (1.88, 1.77, 1.63, 0.49). Darker than male, patterns same as in male. Epigyne (Fig.2C, D).With trapezoid epigynal plate.Copulatory openings united at postmedian potion of epigyne. Copulatory ducts longitudinally arc-shaped and extended upwards, then twisted into oval loops. Spermathecae with two oval chambers respectively, interior chambers smaller than lateral chambers. Fertilization ducts located anteriorly inserted on spermathecal interior chambers. Distribution. Known only from the type locality (Fig. 7)., Published as part of Li, Li-Fen, Liu, Ping, Li, Bing & Peng, Xian-Jin, 2023, Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China, pp. 520-530 in Zootaxa 5263 (4) on pages 521-523, DOI: 10.11646/zootaxa.5263.4.3, http://zenodo.org/record/7835794, {"references":["Zhu M. S. & Zhang B. S. (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, 558 pp."]}

Two new species of the genus Clubiona Latreille, 1804 are described: C. tianpingshan sp. nov. (♂♀) and C. flammaforma sp. nov. (♂♀) from central and south China. The female of C. subcylindrata Wang et al., 2018 is described for the first time. Detailed descriptions, photographs of somatic features and copulatory organs, as well as a distribution map of these three species, are provided.

Clubiona subcylindrata Wang, Chen & Zhang, 2018 Figs 3, 4, 7 Clubiona subcylindrata Wang et al. 2018: 328, figs. 16A–C, 17A–D (♁). Holotype from Fanjing Mountain Reserve, Heiwanhe (27°50.778′N, 108°46.362′E; elev. 533 m), 26.IX.2013, L.Y. Wang, D. Wang & X.K. Jiang leg. (Museum of Hebei University, Baoding, China, examined by Luyu Wang). New Records. 4♁ 5♀, CHINA: Hunan Province, Zhangjiajie City, Tianpingshan Nature Reserve, 29.76101°N, 110.04879°E, 551 m, 22.X.2014, J.H. Gan et al. leg. (HNU-HN-X-1417). Diagnosis. The males of Clubiona s ubcylindrata resemble those of C. cylindrata Liu et al., 2007 (see Liu et al. 2007: figs 13–15) in having a bulged tegulum and a slender embolus, but can be distinguished by: (1) the RTA finger-shaped with abruptly sharpened tip in retrolateral view (Fig. 3E) (vs. spinous in C. cylindrata); (2) the sperm duct invisible in ventral view (Fig. 3D) (vs. visible in C. cylindrata). The females of this species resemble those of C. cylindrata Liu et al., 2007 (see Liu et al. 2007: figs 11, 12) and C. lyriformis Song & Zhu, 1991 (see Song et al. 1991: fig. 4A, B) in having twisted tubular spermathecae, but can be distinguished by: (1) the anterior portion of atrium slightly wider than posterior portion (Fig. 4C) (vs. the anterior portion of atrium about half width of posterior portion in C. cylindrata); (2) the posterior margin of atrium opened (Fig. 4C) (vs. enclosed in C. lyriformis). Description. Male (one of HNU-HN-X-1417) (Fig. 3A, B). Total length 7.34. Carapace 3.19 long, 2.33 wide; abdomen 4.24 long, 2.03 wide. Carapace yellow, fovea reddish and longitudinal, cervical and radial grooves conspicuous. Eyes: AER slightly recurved, PER almost straight in dorsal view. Eye sizes and interdistances: AME 0.16, ALE 0.18, PME 0.14, PLE 0.15, AME–AME 0.12, AME–ALE 0.06, PME–PME 0.28, PME–PLE 0.15, ALE–PLE 0.08. MOQL 0.46, MOQA 0.52, MOQP 0.72. Chelicerae yellow with five promarginal teeth and two retromarginal teeth. Endites and labium yellowish brown. Sternum yellowish, almost oval. Legs yellow; tibiae I and II with two pairs of ventral spines, metatarsus I and II with one pair of ventral spines. Leg measurements: I 8.74 (2.46, 3.54, 1.80, 0.94), II 10.29 (2.87, 4.18, 2.23, 1.01), III 7.55 (2.28, 2.61, 1.96, 0.70), IV 10.65 (2.93, 3.59, 3.26, 0.87). Abdomen elongate oval, dorsum yellow with conspicuous tufts of hairs anteriorly and two pairs of muscular depressions, venter light yellow with four rows of dots medianly. Spinneret yellow. Palp (Figs. 3C–G) Femur and patella without apophysis. Tibia and patella longer than wide, about 1/2 length of cymbium. Tibia with two apophyses; VTA shorter with blunt tip, RTA longer than VTA, finger-shaped with abruptly sharpened tip. Cymbium longer than wide. Tegulum strongly bulged, posterior margin extended to patella. Conductor membranous and leaf-shaped, arising from the top of tegulum. Embolus slender and flagelliform, arising from the middle and prolateral side of tegulum. Sperm duct invisible in ventral view. Female (one of HNU-HN-X-1417) (Figs. 4A, B). Total length 8.51. Carapace 3.67 long, 2.57 wide; abdomen 4.70 long, 2.83 wide. Eye sizes and interdistances: AME 0.17, ALE 0.19, PME 0.15, PLE 0.17, AME–AME 0.16, AME–ALE 0.08, PME–PME 0.40, PME–PLE 0.13, ALE–PLE 0.10. MOQL 0.46, MOQA 0.53, MOQP 0.73. Leg measurements: I 8.20 (2.46, 3.35, 1.52, 0.87), II 8.42 (2.46, 3.39, 1.68, 0.89), III 7.08 (2.13, 2.40, 1.90, 0.65), IV 10.06 (2.92, 3.54, 2.78, 0.82). Colors and patterns same as in male. Epigyne (Fig. 4C, D). Epigynal plate almost rounded. Atrium almost calabash-shaped, longer than wide, and located anteriorly; posterior margin opened. Copulatory openings located at the posterolateral margin of atrium. Copulatory ducts semicircular. Spermathecae slender, tubular. Bursae oval. Fertilization ducts short and curved. Distribution. Guizhou and Hunan Provinces of China (Fig. 7)., Published as part of Li, Li-Fen, Liu, Ping, Li, Bing & Peng, Xian-Jin, 2023, Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China, pp. 520-530 in Zootaxa 5263 (4) on page 524, DOI: 10.11646/zootaxa.5263.4.3, http://zenodo.org/record/7835794, {"references":["Wang, L. Y., Chen, H. M., Wu, P. L., Zhang, F. & Zhang, Z. S. (2018) Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae). Zoological Systematics, 43 (4), 317 - 333. https: // doi. org / 10.11865 / zs. 201827","Liu, P., Yan, H. M., Griswold, C. & Ubick, D. (2007) Three new species of the genus Clubiona from China (Araneae: Clubionidae). Zootaxa, 1456, 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1456.1.3","Song, D. X., Zhu, M. S., Gao, S. S. & Guan, J. D. (1991 a) Six species of clubionid spiders (Araneae: Clubionidae) from China. Journal of Xinjiang University, 8, 66 - 72."]}

Clubiona tianpingshan sp. nov. Figs 5–7 Type material. Holotype: ♁, CHINA: Hunan Province, Zhangjiajie City, Tianpingshan Nature Reserve, 29.79004°N, 110.09161°E, 1442 m, 23.X.2014, J.H. Gan, Y.H. Gong, C. Wang & B. Zhou leg. (HNU-HN-X-1418). Paratypes: 1♀, same data as holotype (HNU-HN-X-1418); 2 ♁ 2♀, Tianpingshan Nature Reserve, 29.77190°N, 110.06751°E, 1348 m, 24.X.2014, J.H. Gan et al. leg. (HNU-HN-X-1419). Etymology. The specific epithet comes from the type locality, noun. Diagnosis. The males of Clubiona tianpingshan sp. nov. resemble those of C. huiming Wang et al., 2018 (see Wang et al. 2018: figs 8, 9) in having helical embolus and simple RTA and VTA, but can be distinguished by: (1) the embolic base about 1/3 width of the tegulum in ventral view (Fig. 5D) (vs. about 1/6 width of the tegulum in C. huiming); (2) the conductor arising from the anterior 1/3 of the tegulum and the terminal portion above the top of tegulum in retrolateral view (Fig. 5E) (vs. arising from the median portion of the tegulum and its terminal portion below the top of tegulum in C. huiming). The females of the new species resemble those of C. applanata Liu et al., 2007 (see Liu et al. 2007: figs 1, 2) and C. subapplanata Wang, et al., 2018 (see Wang et al. 2018: figs 14C, D, 15E, F) in having an oval atrium located anteriorly, a pair of large bursae and a pair of spermathecae with two chambers, but can be distinguished from the two species by (1) the margin of atrium thinner and posterior margin W-shaped in ventral view (Fig. 6C) (vs. thicker and rounded posteriorly in C. applanate and C. subapplanata); (2) the copulatory duct longer and twisted into two circular loops in dorsal view (Fig. 6D) (vs. shorter in C. applanate and C. subapplanata); (3) the posterior tubular chambers of spermathecae extended vertically in dorsal view (Fig. 6D) (vs. extended transversely in C. applanate and C. subapplanata). Description. Male (holotype) (Fig. 5A, B). Total length 3.85; carapace 1.80 long, 1.24 wide; abdomen 2.07 long, 1.29 wide. Carapace yellow; fovea reddish and longitudinal, cervical and radial grooves inconspicuous. Eyes: AER and PER slightly recurved in dorsal view. Eye sizes and interdistances: AME 0.08, ALE 0.11, PME 0.09, PLE 0.10, AME–AME 0.04, AME–ALE 0.03, PME–PME 0.11, PME–PLE 0.06, ALE–PLE 0.04, MOQL 0.27, MOQA 0.22, MOQP 0.37. Chelicerae yellow, each margin with five teeth. Endites and Labium yellowish brown. Sternum yellowish, almost oval. Legs yellowish; tibiae I and II with three pairs of ventral spines, metatarsus I and II with two pairs of ventral spines. Leg measurements: I 4.20 (1.28, 1.65, 0.75, 0.52), II 4.30 (1.34, 1.60, 0.89, 0.47), III 3.84 (1.17, 1.32, 0.93, 0.41), IV 5.46 (1.68, 1.84, 1.44, 0.50). Abdomen oval, dorsum light yellow, with conspicuous tufts of hairs anteriorly, and two pairs of muscular depressions, venter light yellow. Spinneret yellow. Palp (Fig. 5C–F). Femur without apophysis. Patella wider than long in dorsal view, with a thumb-shaped retroapical apophysis. Tibia shorter and narrower than patella, about 1/3 length of cymbium, with three apophyses; RTA sclerotized and flat, almost triangular with pointed tip in retrolateral view; VTA smaller than RTA, triangular with blunt tip; LTA smallest and with blunt tip. Cymbium longer than wide. Tegulum enlarged and protruded, slightly longer than wide. Conductor beak-shaped with sharp apices, located retrolaterally and arising distally from the 1/3 portion of tegulum in retrolateral view. Embolus arising from the top of tegulum, helical and extended anticlockwise, basal portion about 1/3 wide of tegulum, distal portion filiform. Sperm duct U-shaped in ventral view. Female (HNU-HN-1418) (Fig. 6A, B). Total length 4.23; carapace 1.88 long, 1.37 wide; abdomen 2.37 long, 1.40 wide. Eye sizes and interdistances: AME 0.08, ALE 0.12, PME 0.11, PLE 0.11, AME–AME 0.06, AME–ALE 0.03, PME–PME 0.19, PME–PLE 0.10, ALE–PLE 0.08, MOQL 0.31, MOQA 0.23, MOQP 0.40. Leg measurements: I 3.96 (1.21, 1.58, 0.75, 0.42), II 4.07 (1.29, 1.67, 0.73, 0.38), III 3.65 (1.06, 1.33, 0.89, 0.37), IV 5.05 (1.52, 1.70, 1.36, 0.47). Colors and patterns same as in male. Epigyne (Fig. 6C, D). Epigynal plate almost rounded. Atrium almost oval, wider than long, about 3/5 width of epigyne, located anteriorly, posterior margin thickened and W-shaped. Copulatory openings located in the posterior margin of atrium. Copulatory ducts twisted into two circular loops. Spermathecae meso-laterally located with two chambers respectively; anterior chamber globular, posterior chamber tubular and extended vertically. Bursae almost globular, located posteriorly. Fertilization ducts short and curved. Distribution. Known only from the type locality (Fig. 7)., Published as part of Li, Li-Fen, Liu, Ping, Li, Bing & Peng, Xian-Jin, 2023, Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China, pp. 520-530 in Zootaxa 5263 (4) on pages 524-528, DOI: 10.11646/zootaxa.5263.4.3, http://zenodo.org/record/7835794, {"references":["Wang, L. Y., Chen, H. M., Wu, P. L., Zhang, F. & Zhang, Z. S. (2018) Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae). Zoological Systematics, 43 (4), 317 - 333. https: // doi. org / 10.11865 / zs. 201827","Liu, P., Yan, H. M., Griswold, C. & Ubick, D. (2007) Three new species of the genus Clubiona from China (Araneae: Clubionidae). Zootaxa, 1456, 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1456.1.3"]}

Carteronius myene Bonaldo & Labarque sp. nov. Figs 13A, B; 14A, B; Map 1 Type. ♀ holotype from Parc National de Moukalaba-Doudou, Département de Ndougou, Province de Ogooué Maritime (02°35’S, 10°14’E), GABON, III–IV.2003, O.S.G. Olivier & M. Burger leg. (forêt marecaguese) (MRAC 220.827). Etymology. The specific name is a noun in apposition referring to the Myene people, which settled fishing communities along the coast of Gabon. Diagnosis. Females of Carteronius myene sp. nov. resemble those of C. gentilis comb. nov. by the slightly curved epigynal transversal ridge (Figs 14A, 19C), but can be recognized by the posterior sector relatively larger; spermathecae barely visible by transparency in anterior sector (Fig. 14A). Description. Male. unknown. Female. (MRAC 220.827). Measurements: Total length 11.29, CL 5.29, CW 4.85, AL 6.33, AW 4.59, SL 2.64, SW 2.30. Eye diameters and interdistances: AME 0.42, ALE 0.34, PLE 0.29, PME 0.32, AME-AME 0.32, AME-ALE 0.37, ALE-ALE 2.32, PME-PME 0.39, PME-PLE 0.62, PLE-PLE 2.71. Length of leg segments: I 4.68+2.28+4.48+3.46+1.34=16.24; II 4.96+2.29+4.27+3.58+1.41=16.58; III 3.59+1.71+2.83+2. 81+1.00=11.29; IV 4.03+1.69+3.43+3.44+1.15=13.74. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, proximal tooth largest; Leg spination: femora: II do 0-1-0, III do 0-1-0 ve 0-1-0, IV do 1-0-0; tibiae: I ve 2-2-2-2-2-2, II ve 2-2-2-2-2, III ve 2-2, IV ve 1-0-1; metatarsi: I ve 2-2-2-2, II ve 1 p -1 p -0-1 r -2, III pl 0-0-0-1 rl 0-0-0-1 ve 2-2-1, IV pl 2-0-1-0 ve 0-1-0 -0. Coloration: Carapace and chelicerae dark reddish-brown. Endites, labium and sternum dark reddish-brown. Legs reddish-brown, with femora I and II darker. Abdomen dark gray, with several small white spots, two pairs of small white spots in middle dorsally, and three faint chevrons posteriorly. Venter gray, with irregular white spots forming two transversal lines (Fig. 13A). Epigynum: CDv long, gently arched, ST2 tapering, anteriorly located, gland ducts present, approximately same size as ST1; CDd folded ventrally (Fig. 14B). Other material examined. None. Distribution. Only known from Gabon (Map 1).

Carteronius simoni Bonaldo & Shimano sp. nov. Fig. 17; Map. 1 Holotype. ♀ from GABON: leg. Mocquery (No further data) (MNHN-17.436). Etymology. The specific name is a patronym honoring French arachnologist Eugène Simon, who described Carteronius. Diagnosis. Females of C. simoni sp. nov. are similar to those of C. lumumba sp. nov. by the inconspicuous lateral plates of the posterior epigynal sector (Figs 16D, 17C) but can be recognized by the wider, divergent copulatory ducts (Fig. 17C). Description. Male. unknown. Female. (MNHN-17.436) Measurements: Total length 9.15, CL 4.14, CW 3.55, AL 4.99, AW 4.03, SL 1.68, SW 1.82. Eye diameters and interdistances:AME 0.35, ALE 0.24, PLE 0.25, PME 0.30, AME-AME 0.30, AME-ALE 0.40, ALE-ALE 2.15, PME-PME 0.56, PME-PLE 0.61, PLE-PLE 2.60. Length of leg segments: I 2.80+1.47+2.58+2.10+1.09=10.04; II 2.95+1.40+2.48+2.12+0.96=9.91; III 2.35+1.09+1.77+1.76+0.78 =7.75; IV 2.80+1.14+2.43+2.27+0.87=9.51. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two subequal teeth. Leg spination: tibiae: I ve 2-2-2-2-2, II ve 2-2-2-2, IV ve 0-0-1 p; metatarsi: I ve 2-2, II ve 2-2, III ve 2-1 p -2, IV pl 0-0-0-1 rl 0-1-0-1 ve 1 p -1 r - 1 p. Coloration: carapace, chelicerae, endites, labium and sternum reddish. All leg segments yellowish-red. Abdomen pale gray dorsally, with indistinct white spots (Fig. 17A). Epigynum: CDv long, straight, ST2 a small globe, anteriorly located, gland ducts inconspicuous, smaller than ST1; CDd large, S-shaped (Fig. 17D). Other material examined. None. Distribution. Only known from Gabon (Map 1)., Published as part of Eb. Bonaldo, Ramírez, Martín J., Om. Labarque, Shimano, Yulie, Silva-Junior, Cláudio J. & Haddad, Charles R., 2022, Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group, pp. 343-373 in Zootaxa 5205 (4) on page 363, DOI: 10.11646/zootaxa.5205.4.3, http://zenodo.org/record/7307035

Carteronius arboreus Bonaldo & Haddad sp. nov. Figs 7, 8; Map 1 Types. ♂ holotype from Bas Congo, Mayombe, Luki Forest Reserve (05°37’S, 13°05’E), 28.IX.2007, DEMOCRATIC REPUBLIC OF THE CONGO, D. de Bakker & J. P. Michiels leg. (sieving along trail in primary rainforest) (MRAC 223.475). ♀ paratype, same locality and collectors, 10.XI.2006 (Fog 3, primary rainforest) (MRAC 220.925). Etymology. The specific name is a Latin adjective meaning arboreal, in reference to the fact that most of the known specimens were collected by canopy fogging. Diagnosis. Males of Carteronius arboreus sp. nov. are similar to those of C. ashanti sp. nov. by the presence of a sub-apical embolar process (Figs 8A, 10A), but differ by the dorsal lobe of the RTA, which is broad and retrolaterally oriented, and the rounded medial lobe, which is small in relation to the dorsal lobe, and share the same base (Fig. 8B). In C. ashanti sp. nov., the dorsal lobe is spoon-shaped and the medial lobe longer and fang-shaped, with its own base. Females resemble those of C. sudanus comb. nov. in the strongly recurved epigynal transversal ridge (Figs 4C, 8C), but differ by the lateral plates of the epigynal posterior sector being strongly sclerotized (Fig. 8C). Description. Male. (MRAC 223475). Measurements: Total length 6.03, CL 3.16, CW 2.71, AL 2.95, AW 2.24, SL 1.39, SW 1.49. Eye diameters and interdistances: AME 0.28, ALE 0.17, PLE 0.18, PME 0.16, AME-AME 0.65, AME-ALE 0.16, ALE-ALE 1.34, PME-PME 0.33, PME-PLE 0.37, PLE-PLE 1.57. Length of leg segments: I 3.08+1.17+2.67+2.29+1.22=10.43; II 3.10+1.24+3.10+2.24+1.27=10.95; III 2.07+0.93+1.61+1.65+0.96=7.22; IV 2.55+0.94+2.11+2.25+1.05=8.90. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: I do 0-1-0 pl 0-0-1-0, II do 0-1-0, III do 0-1-0, IV do 0-1-0; tibiae: I ve 2-2-2-2-2, II ve 1 p -1 r -1 p -2-2-2, III ve 1 r -1 p -2, IV rl 0-1-0-1 ve 1 p -0-1 p; metatarsi: I ve 2-2, II ve 2-2, III pl 0-1-0-1 rl 0-1-0-1 ve 2-2, IV pl 0-1-0-1 rl 0-1-0-1 ve 1 r -1 p -1 r -0. Coloration: carapace and chelicerae reddish-brown. Endites, labium and sternum reddish-brown. Legs I and II: coxae and trochanters reddish-brown; femora reddish-brown, yellowish distally; tibiae, metatarsi and tarsi yellowish. Legs III and IV yellowish. Abdomen dark gray dorsally, with two well defined white bands; white ventrally (Fig. 7A). Palp: RTA with apical spur short, curved and pointed, dorsal lobe with apical edges bent ventrally, ventral lobe rounded and excavated. Tegulum with short retrolateral apical tegular process, spermatic duct with long loop (Fig. 8A, B). Female. (MRAC 220925). Measurements: Total length 10.58, CL 4.10, CW 3.86, AL 6.37, AW 4.91, SL 1.97, SW 1.93. Eye diameters and interdistances: AME 0.31, ALE 0.21, PLE 0.21, PME 0.18, AME-AME 0.25, AME-ALE 0.29, ALE-ALE 1.75, PME-PME 0.45, PME-PLE 0.49, PLE-PLE 2.13. Length of leg segments: I 3.87+1.81+3.70+2.88+1.28=13.54; II 3.70+1.73+3.66+2.99+1.31=13.39; III 2.75+1.32+2.17+2.08+0.91=9.23; IV 3.25+1.31+2.76+2.84+0.98=11.14. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: I do 0-1-0 pl 0-1-0, II do 0-1-0, III do 0-1-0-1 p, IV do 0-1-0-1 p; tibiae: I ve 2-2-2-2-2-2-2, II ve 2-2-2-1 r -1 p -1 r -1 p -1 r, III ve 1 p -1 r -2, IV rl 0-1-0-1 ve 1p-0-1 p; metatarsi: I ve 2-0-2-0, II ve 2-2-2, III pl 0-1-0-1 rl 0-0-0-1 ve 2-2-1, IV pl 0-1-0-1 rl 1-0-1 ve 1 p -1 r -1 p - 0-0. Coloration: Carapace and chelicerae dark reddish-brown. Endites, labium and sternum reddish-brown. Legs reddish-brown, with femora I and II darker. Abdomen gray, dorsum with scattered small white spots, denser in middle, forming irregular white longitudinal band; posteriorly with large triangular white spot. Ventrally gray with two irregular lateroventral white bands (Fig. 7C). Epigynum: CDv slight folded posteriorly, ST2 globose, anteriorly located, larger than ST1, CDd almost straight (Fig. 8D). Other material examined. DEMOCRATIC REPUBLIC OF THE CONGO: Bas Congo, Mayombe, Luki Forest Reserve (05°37’S, 13°05’E), 18.IX.2007, D. de Bakker & J.P. Michiels leg. (Fog 5, old secondary forest), 1♀ (MRAC); same data but 22.IX.2007, 1♀ (MRAC); same data but 30.IX.2007, 1♂ (MRAC). Distribution. Only known from the Democratic Republic of the Congo (Map 1).

Eb.Bonaldo, Ramírez, Martín J., Om.Labarque, Shimano, Yulie, Silva-Junior, Cláudio J., Haddad, Charles R. (2022): Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group. Zootaxa 5205 (4): 343-373, DOI: https://doi.org/10.11646/zootaxa.5205.4.3

Carteronius teke Bonaldo & Bosselaers sp. nov. Figs 13C, D, 14C, D; Map 1 Type. ♀ holotype from Kivu, Rwankwi [01°19’S, 29°22’E], DEMOCRATIC REPUBLIC OF THE CONGO, VI.1946, J. Leroy leg. (MRAC 168.643). Etymology. The specific name is a noun in apposition referring to the Teke people, one of the three kingdoms that ruled Congo before the arrival of Europeans. Diagnosis. Females of Carteronius teke sp. nov. differ from all other species of Carteronius by the presence of completely straight epigynal transversal ridge (Fig. 14C). Description. Male. Unknown. Female. (MRAC 168.643) Measurements:Total length 13.45, CL 4.90, CW 4.39, AL 7.99, AW 5.24, SL 2.44, SW 2.15. Eye diameters and interdistances:AME 0.38, ALE 0.25, PLE 0.28, PME 0.25, AME-AME 0.36, AME-ALE 0.40, ALE-ALE 2.30, PME-PME 0.57, PME-PLE 0.67, PLE-PLE 2.81. Length of leg segments (sequence from femur to tarsus, and total): I 4.21+2.04+3.57+2.92+1.25=13.99; II 4.15+1.86+3.89+2.79 +1.19=13.88; III 2.83+1.43+2.52+2.20+1.04=10.02; IV 3.42+1.42+3.03+3.14+1.28=12.29. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: IV do 1-0-0; tibiae: I ve 1 p -2-2-2-2-2, II ve 0-0-2-2-2-2, III ve 0-0-1 p -0, IV ve 1 p -0-1 p -0; metatarsi: I ve 2-2-2-2, II ve 1 r -1 p -2-2, III ve 0-2-2-0, IV ve 0-2-2-0. Coloration: carapace and chelicerae dark reddish-brown. Endites, labium and sternum reddish-brown. Legs I and II reddish-brown, III and IV dark yellow. Abdomen dorsally pale gray, ventrally white with darker band converging at spinnerets (Fig. 13C). Epigynum: CDv long, arched, ST2 tapering, medially located, gland ducts inconspicuous, much smaller than ST1; CDd folded ventrally (Fig. 14D). Other material examined. None. Distribution. Only known from the Democratic Republic of the Congo (Map 1)., Published as part of Eb. Bonaldo, Ramírez, Martín J., Om. Labarque, Shimano, Yulie, Silva-Junior, Cláudio J. & Haddad, Charles R., 2022, Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group, pp. 343-373 in Zootaxa 5205 (4) on pages 359-361, DOI: 10.11646/zootaxa.5205.4.3, http://zenodo.org/record/7307035

A new genus of Clubionidae Wagner, 1887, Femorbiona Yu & Li, gen. nov., is described, with Clubiona brachyptera Zhu & Chen, 2012 (♂♀; Hainan, China) as the type species. Three species are included in Femorbionagen. nov.: F. brachypteracomb. nov., F. phami Yu & Li, sp. nov. (♂♀; Hai Phong, Vietnam), and F. shenzhen Yu & Li, sp. nov. (♂♀; Guangdong, China).

Spiders of the genusClubionaLatreille, 1804 from Xishuangbanna, Yunnan Province, China are studied. A total of 47 species is reported and illustrated, including 14 new species and two new synonyms. Twelve of the new species belong to four species groups:C. dengpaoYu & Li,sp. nov.,C. subdidentataYu & Li,sp. nov.,C. tixingYu & Li,sp. nov.,C. xiaociYu & Li,sp. nov.,C. xiaokongYu & Li,sp. nov.,C. yejieiYu & Li,sp. nov.,C. zhaoiYu & Li,sp. nov.andC. zhigangiYu & Li,sp. nov.from theC. corticalisgroup;C. miiYu & Li,sp. nov.andC. subtongiYu & Li,sp. nov.from theC. ternatensisgroup;C. bannaYu & Li,sp. nov.from theC. filicatagroup; andC. menglunYu & Li,sp. nov.from theC. trivialisgroup. The remaining two new species,C. shuangsiYu & Li,sp. nov.andC. wangchengiYu & Li,sp. nov., are not readily assignable to any of the existing species groups. The female ofC. cochlearisYu & Li, 2019, the female ofC. tianeYu & Li, 2019, the female ofC. bicornisYu & Li, 2019, the male ofC. lalaJäger & Dankittipakul, 2010 and the true female ofC. suthepicaDankittipakul, 2008 are described for the first time. Two new synonyms are:C. vukomiJäger & Dankittipakul, 2010syn. nov. =C. circulataZhang & Yin, 1998;C. melanotheleThorell, 1895syn. nov. =Clubiona melanostictaThorell, 1890. A checklist ofClubionaspecies from Xishuangbanna is provided. The DNA barcodes of almost all of the species were obtained for species delimitation, matching of sexes and future use.

Clubiona qianlei J. Zhang, F. Zhang & H. Yu, sp. nov. Figs 1–4, 6 Type material. Holotype: ♂, CHINA: Hubei Province, Xianning City, Tongshan County, Jiugongshan Nature Reserve, Shilonggou (N29.41044598º, E114.64803038º, 480 m), 29.IV.2020, Q.L. Lu and Y. Zhong leg. Paratype: 1♀ (YHCLU0276), Jiugongshan Nature Reserve, Yunzhonghu (N29.40443038º, E114.67254442º, 1240 m), 1.VII.2020, Q.L. Lu et al. leg.; 1♂ (YHCLU0318), same data as holotype. Etymology. This species is a patronymic named after Mr. Qianle Lu (Shenzhen City, China), collector of the types, who has greatly helped us in our research. Diagnosis. The male of C. qianlei sp. nov. differs from those of all other group members by the absence of subapical barb on the RTA’s ventral process, and by the tegular apophysis width ca. 2/3 of bulb width (Figs 2B, C, 3B). In contrast, all other group members have subapical barb on the RTA’s ventral process, tegular apophysis narrower than half of bulb width, such as C. reclusa (Almquist 2006: 374, figs 323a, b) and C. interjecta (Figs 5A, C; Tang et al. 2005: 79, figs 3D, E). The female is similar to those of C. interjecta in the general appearance of the epigyne, but differs by: (1) copulatory openings slit-like, indistinct (Figs 4A–C) (vs. nearly circular, relatively conspicuous; Fig. 5D; Tang et al. 2005: 79, fig. 3B; Zhu & Zhang 2011: 358, fig. 257B); (2) dorsal part of spermatheca (SPd) with hyaline outermost surface (Figs 4D, E) (vs. surface sclerotized; Fig. 5E; Tang et al., 2005: 79, fig. 3C; Zhu & Zhang, 2011: 358, fig. 257B); (3) the two SPds separated by about one diameter (Figs 4D, E) (vs. closely spaced or separated by less than 0.5 diameter; Fig. 5E; Tang et al. 2005: 79, fig. 3C; Zhu & Zhang 2011: 358, fig. 257B) Description. Male (holotype): Total length 5.90. carapace 2.95 long, 2.05 wide; abdomen 2.95 long, 1.62 wide. Living holotype dark brown (Fig. 1A). Carapace yellowish-brown in ethanol (Figs 3D, F), slightly darker in front, without a distinct pattern; cephalic region distinctly narrowed, thoracic groove and radial grooves indistinct; tegument smooth, clothed with fine hairs. Eyes: AER slightly recurved, PER slightly wider than AER, almost straight in dorsal view. Eye sizes and interdistances: AME 0.15, ALE 0.14, PME 0.18, PLE 0.16, AME–AME 0.12, AME–ALE 0.07, PME–PME 0.33, PME–PLE 0.21, MOQL 0.34, MOQA 0.37, MOQP 0.60. Chelicerae protruding, robust, uniformly brownish red, with four teeth on promargin and three on retromargin. Sternum (Fig. 3E) yellowish white, 1.57 long, 1.02 wide. Labium coloured as chelicerae, endites brown. Legs light yellow, without distinct markings. Leg measurements: I 8.41 (2.37, 3.41, 1.66, 0.97), II 9.99 (2.55, 3.39, 3.11, 0.94), III 7.31 (2.21, 2.57, 1.90, 0.63), IV 10.36 (2.95, 3.44, 3.09, 0.88). Abdomen (Figs 3D–F) elongate, oval, dorsum centrally with a lengthwise heart mark, reaching half of abdomen length, with a pair of muscular depressions located on middle level of heart mark; laterally with lengthwise reticular pattern, posteriorly furnished with a fuzzy pattern; venter medially with two longitudinal dotted lines. Palp (Figs 2 A−D, 3A−C): Tibia short, ca. 1/3 length of cymbium. Retrolateral tibial apophysis (RTA) heavily sclerotized and bifurcated, slightly longer than tibia, broad at base; ventral process (VP) with sharp tip, shaped like a leaf in retrolateral view; dorsal process (DP) triangular, ca. 1/2 ventral process length. Genital bulb elongated and with a relatively flat tegulum, ca. 1.8 × longer than wide, with distinctive, sinuate sperm duct. Tegular apophysis (TA) represented by a large and flat sclerite, inverted triangular, anterior margin slightly concaved and rough, situated prolaterally on the tegulum, about 2/3 width of bulb. Embolar base (EB) inserted at approximately ten o’clock on tegulum, arising on the dorsal, hidden side of the tegular apophysis; free part of embolus (E) filamentous, as long as tegulum width, tip directed antero-mesally. Conductor (C) thick, finger-like, ca. 1/5 of tegulum length, originating from retrolateral, apical area of tegulum. Female. As in male, but slightly larger in size and lighter in colour. Living paratype light brown (Fig. 1B). Habitus in ethanol as Figs 4F, G. Total length 6.80; carapace 2.92 long, 2.09 wide; abdomen 3.88 long, 2.86 wide. Eye sizes and interdistances: AME 0.16, ALE 0.18, PME 0.19, PLE 0.11, AME–AME 0.05, AME–ALE 0.06, PME– PME 0.25, PME–PLE 0.20. MOQL 0.45, MOQA 0.38, MOQP 0.58. Sternum 1.56 long, 0.94 wide. Measurements of legs: I 6.49 (1.89, 2.68, 1.16, 0.76), II 6.68 (1.93, 2.72, 1.31, 0.72), III 5.97 (1.76, 2.13, 1.57, 0.51), IV 9.17 (2.63, 3.09, 2.76, 0.69). Epigyne (Figs 4 A−E): Epigyne plate a darkened disc, nearly triangular, anterior and lateral margins indistinct, posterior margin heavily sclerotized, middle part of posterior margin protruding; arrangement of the various parts of the vulva indistinctly visible through the tegument. Copulatory openings (CO) small, indistinct, represented by a short slit, situated at concave bends of epigynal plate posterior margin. Copulatory ducts (CD) firstly running along the posterior margin of epigyne, leading to large glands (GL), then ascending obliquely, finally connecting to anteriorly located spermathecae. Dorsal part of spermathecae (SPd) globular, innermost surface sclerotized, outermost surface hyaline, the two SPd separated by about one diameter. The ventral part of spermathecae (SPv) tubular or finger-like, curved medially, the two SPvs closely spaced. Fertilization ducts (FD) acicular, membranous, slightly shorter than diameter of SPv, located on dorsal surface of SPd. Natural History. Leaf-dwelling spiders. Photos of live specimens were taken after the spiders dropped on the ground due to beating. Distribution. Know only from the type locality, Mt. Jiugong, Hubei, China (Fig. 6). Clubiona interjecta L. Koch, 1879 Figs 5−6 Clubiona interjecta L. Koch 1879: 89, pl. 3, fig. 7 (♂ ♀); Zhang & Hu 1989: 57, figs 3, 18 (♂ ♀); Song et al. 1999: 416, figs 246F−G, 249E−F (♂ ♀); Tang et al. 2005: 79, figs 3A−E (♂ ♀); Zhu & Zhang 2011: 358, f. 257A-E (♂ ♀), World Spider Catalog 2022 (full list of taxonomic references). Material examined. CHINA: Hebei Province: Chengde City, Weichang County, 1♂ 5♀, Saihanba National Forest Park (N42.584537º, E117.837489º, 1500 m), 3.VIII.2018, X.B. Guo, H. Wang, J.X. Lai and Y.F. Li leg; Shijiazhuang City, Pingshan County, 1♂, Tuoliang Nature Reserve (N38.731813º, E113.824548º, 2000 m), 12. V.2018, X.B. Guo and Z.Y. Li leg; Zhangjiakou City, Zhuolu County, 27♀, Xiao Wutai Mountain Nature Reserve, Shanjiankou Village (N40.029531º, E115.063527º, 1300 m), 6.VII.2018, X.B. Guo, S. Qiao, J.X. Lai and Y.F. Li leg; Shanxi Province: Xinzhou City, Ningwu County, 1♂, Luya mountain (N38.837989º, E112.083789º, 1900 m), 6.VII.2011, C. Jin leg; Sichuan Province: Ganzi County, 1♂, Renguo village (N31.667885º, E99.803766º, 3420 m), 20. VI.2018, X.B. Guo Leg. Emended diagnosis. As the only two members of the reclusa -group in China, Clubiona interjecta and C. qianlei sp. nov. share almost all of the group characters, but they can be separated by the shape of the copulatory organs, and by some somatic characters. C. interjecta can be distinguished from C. qianlei sp. nov. by the following characters: for the males, RTA’s ventral process with a subapical barb, dorsal process finger-like in C. interjecta (Fig. 5C; Tang et al. 2005: 79, fig. 3E) (vs. subapical barb absent, dorsal process triangular; Fig. 2B); tegular apophysis about 1/3 width of bulb, distally narrowed and bearing a dentiform process (Fig. 5A; Tang et al. 2005: 79, fig. 3D) (vs. tegular apophysis about 2/3 width of bulb, distal margin wide and rough; Figs 2C, 3B); for the females, copulatory openings distinct, nearly circular in C. interjecta (Fig. 5D; Tang et al. 2005: 79, fig. 3B) (vs. copulatory openings indistinct, slit-like; Figs 4 A−C); SPd’ surface sclerotized, the two SPds nearly closely spaced (Fig.5E; Tang et al. 2005: 79 fig. 3C) (vs. SPd’ outermost surface hyaline, the two SPds separated by about one diameter; Figs 4D, E). In addition, the two species can be separated by the abdominal pattern: dorsum of abdomen anteriorly with distinct, longitudinal heart mark, reaching 1/2 of abdomen length, posteriorly with 4~6 transverse chevrons in C. interjecta (Figs 5F, G; Tang et al. 2005: 79, fig. 3A), but only with indistinct heart mark and muscular depressions in C. qianlei sp. nov. (Figs 3D, 4F). Description. See Tang et al. (2005). Male palp as in Figs 5 A−C, epigyne as in Figs 5D, E, habitus as in Figs 5F, G. Distribution. Russia (West Siberia to Far East), Mongolia, China (Inner Mongolia, Hebei, Shanxi, Henan, Sichuan, Jilin, Heiongjiang, as in Fig. 6).

Clubiona reclusa species-group Diagnosis. See Mikhailov (1995)., Published as part of Zhang, Jianshuang, Chen, Lulu, Ding, Yanmei, Zhang, Feng & Yu, Hao, 2022, On the Clubiona reclusa species-group in China, with the description of Clubiona qianlei sp. nov. (Araneae, Clubionidae), pp. 412-421 in Zootaxa 5129 (3) on page 413, DOI: 10.11646/zootaxa.5129.3.5, http://zenodo.org/record/6501213, {"references":["Mikhailov, K. G. (1995) Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta, 4 (2), 33 - 48."]}

The representation of the Clubiona reclusa species-group in China is here established in two species: Clubiona qianlei sp. nov. from Central China and C. interjecta L. Koch, 1879, which is mainly distributed in northern China. Detailed description, diagnosis, photographs of the new species are given. DNA barcodes (a partial fragment of the mitochondrial cytochrome oxidase subunit I gene, COI) of the new species were obtained to confirm matching of the sexes and for future use in molecular studies. Supplementary micrographs of C. interjecta are provided for the first time, alongside an emended diagnosis, to demonstrate the validity of C. qianlei sp. nov.. A distribution map of the reclusa group species in China is given.

The family Clubionidae is represented by 15 genera and 506 species and subspecies that occur worldwide (World Spider Catalog 2021). From South Africa one genus and 26 species are known of which 17 are South African endemics, six are African endemics and three are known throughout southern Africa. Fourteen species have a wide distribution and is of Least Concern but 12 are Data Deficient. Twenty of the species are known only from one sex. The genus Clubiona is presently being revised and several new species are expected.

The monotypic genus Bucliona Benoit, 1977 (type Clubiona dubia O. Pickard-Cambridge, 1870 from Saint Helena Island), previously considered a junior synonym of Clubiona Latreille, 1804, is resurrected. Three species are assigned to the genus Bucliona: B. dubia comb. reval., B. kirilli sp. n. (♂, Kenya), and B. jucunda (Karsch, 1879) comb. n., ex Clubiona (♂♀, Far East Asia).

Bucliona kirilli Yu & Li sp. n. Figs 2E, F, 5, 6, 7A, C Type material. Holotype ♂ (NMK), KENYA: Nyeri County, Naro Moru Town, Mount Kenya National Park, Metrological Station (S0.1702��, E37.2140��, 3000 m), 16.VIII.2018, Kioko G. leg. Other material examined. KENYA: Laikipia County, Nanyuki Town, Mount Kenya National Park, Sirimon Gate, Bamboo Forest (S0.2675��, E37.28276��, 3176 m), 1♂ (MGEU), 17.VIII.2018, Kioko G. leg. Etymology. The species is named in honour of Kirill G. Mikhailov from Moscow State University for his contributions to the systematics of clubionid spiders and on the occasion of his 60 th birthday; noun (name) in genitive case. Diagnosis. Males of B. kirilli sp. n. resemble those of B. dubia but differ by the parallel sides and a triangular tip of the RTA (vs. wide base and claw-like tip), as well as by the tip of the embolus which is directed antero-mesally (vs. anteriorly). Description. Male. Total length 7.66; carapace 3.65 long, 2.54 wide; abdomen 4.01 long, 2.14 wide. Carapace (Figs 6A���C) uniformly yellow-brown. Labium and endites dark reddish brown. Sternum coloured as carapace. Eyes: AER slightly recurved, PER wider than AER and straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.17, ALE 0.20, PME 0.17, PLE 0.18, AME���AME 0.14, AME���ALE 0.15, PME���PME 0.39, PME���PLE 0.28, MOQL 0.47, MOQA 0.44, MOQP 0.71. Legs yellowish, without distinct pattern. Leg measurements: I and II missing, III (2.50, ���, ���, ���), IV 11.45 (3.20, 3.95, 3.33, 0.97). Abdomen (Figs 6A���C) with wide dorsal scutum covering almost entire abdomen; dorsum anteriorly with triangular, dark, median stripe, reaching 1/3 of abdomen length, centrally without distinct patterns (faded in the preserved holotype but with 2 transverse chevrons in the second female), posteriorly with 3 transverse chevrons; venter anteriorly without any markings, posteriorly with 2 broken longitudinal stripes. Palp (Figs 5A���E, 7A, C): Tibia relatively long, over 1/2 cymbium length, ca. 1.9 times longer than wide, retrolateral tibial apophysis (RTA) about 1/2 of tibia length, about 5 times longer than wide, slightly wider at base, gradually tapering towards apex, tip triangular, sharply pointed. Embolus ��-shaped in ventral view, bent at almost a right angle, tip directed antero-mesally, base (EB) slightly longer than free part (E), embolus 6.6 times longer than wide. Tegular apophysis (TA) heavily sclerotized, arising retrolatero-distally from tegulum, finger shaped, directed proximally, distally curved retrolaterally. Female. Unknown. Distribution. Known only from the type locality., Published as part of Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao & Li, Shuqiang, 2021, Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae), pp. 195-207 in Zootaxa 5006 (1) on pages 203-206, DOI: 10.11646/zootaxa.5006.1.21, http://zenodo.org/record/5157237

Bucliona dubia (O. Pickard-Cambridge, 1870) comb. reval. Fig. 1 Clubiona dubia O. Pickard-Cambridge, 1870: 532, pl. 42, fig. 1 (♀); O. Pickard-Cambridge, 1873: 213, pl. 24, fig. 3 (♂). Bucliona dubia: Benoit 1977: 68, figs 26a���c, 27a���d (♂ ♀). Clubiona (Bucliona) dubia: Mikhailov 1997: 96, figs 1���4 (♂ ♀). Type examined. Photograph of the holotype female from OUMNH, Mr. Melliss���s Saint Helena Island collection, May 1869. Diagnosis and description. See Benoit (1977) and Mikhailov (1997). Epigyne and habitus of the holotype female are as in Fig. 1A���D. Comments. The holotype female was thought to be lost (Benoit 1977), however Zo�� Simmons, the curator of OUMNH has managed to find the vial with the holotype female. Distribution. The species is known from Saint Helena Island only., Published as part of Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao & Li, Shuqiang, 2021, Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae), pp. 195-207 in Zootaxa 5006 (1) on page 198, DOI: 10.11646/zootaxa.5006.1.21, http://zenodo.org/record/5157237, {"references":["Pickard-Cambridge, O. (1870) Notes on some spiders and scorpions from St Helena, with descriptions of new species. Proceedings of the Zoological Society of London, 37 (3, 1869), 531 - 544, pl. 42.","Benoit, P. L. G. (1977) Fam. Clubionidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences Zoologiques (Zool. - Ser. 8 °), 220, 64 - 81.","Mikhailov, K. G. (1997) Redescription of Clubiona dubia O. Pickard-Cambridge, 1869 (Aranei Clubionidae) from Saint Helena Island. Arthropoda Selecta, 6 (1 / 2), 95 - 97."]}

Genus Bucliona Benoit, 1977, status revalidated Bucliona Benoit, 1977: 68. Bicluona Mikhailov, 1994: 52 (described as a subgenus of Clubiona, type Liocranum jucundum Karsch, 1879). Bucliona: Mikhailov 1997: 95 (downgraded to subgenus of Clubiona and synonymized with Bicluona). Type species. Clubiona dubia O. Pickard-Cambridge, 1870 from Saint Helena Island, by monotypy. Diagnosis. Bucliona differs from Clubiona sensu stricto (type species Araneus pallidulus Clerck, 1757) by: having a distinct mesal ridge on the male chelicerae (Fig. 2A, C) (vs. lacking in Clubiona); the presence of a dorsal abdominal scutum in males (Figs 4E, F, 6A, C) (vs. absent); the undivided, short tibial apophysis of the male palp (Figs 3B, 5B, 7B���D) (vs. divided in many species); the presence of a tegular (= median) apophysis and the lack of a meandering sperm duct (Figs 3B���E, 5A���E) (vs. lack of a tegular apophysis and presence of a meandering sperm duct); the subtegulum with a prolateral extension (Fig. 8A) (vs. without, Figs 8B���F); the tip of the embolus directed anteriorly or antero-mesally (Figs 3D, 5D, 7A, B) (vs. retrolaterally or posteriorly). Females of Bucliona differ from those of Clubiona sensu stricto in having an epigyne with a distinct, undivided epigynal fovea with an anterior hood and a pair of posterior hoods (Figs 1C, D, 4A, C) (vs. fovea and hoods lacking) and longer copulatory ducts (Fig. 4D). Both the males and females of Bucliona can be recognized by the anterior tibiae with a prolateral ���scopula���, which is reduced to a single row (Fig. 2D���F) (vs. dense scopula ventrally on the anterior tibiae). Description. Medium to large size, with the body length of males 4.90���7.75 and of females 6.50���9.50. Male. Body yellow-brown, legs uniformly coloured as carapace (Figs 4E, F, 6A���C). Carapace (Figs 4E, F, 6A, C): elongate-oval, widest at midpoint, pars cephalica slightly elevated above thorax, pars thoracica distinctly wider and slightly higher than pars cephalica; integument smooth; distinctly darker in ocular region, without distinct pattern; cervical groove and radial grooves distinct; fovea a short slit on posterior half of carapace, longitudinal and reddish. Clypeus height distinctly narrower than AME (Figs 2A, C). Chelicerae (Figs 2A, C, 4E, F, 6A���C): robust and red brownish, consisting of a coniform paturon and claw-shaped fang; cheliceral paturon with distinct mesal ridge, distally with several setae; fang furrow with 3 promarginal and 2 retromarginal teeth. Both endites and labium longer than wide; endites depressed posteriorly, slightly convergent anteriorly, with dense scopulae on inner margin; labium nearly trapezoidal and depressed laterally (Fig. 6B). Sternum (Fig. 6B): shield shaped and brown, longer than wide, anteriorly straight; posterior region protruding strongly between coxae IV. Eyes: in dorsal view, AER slightly recurved, PER almost straight (Figs 4E, 6A); in anterior view, AER almost straight, PER slightly recurved (Figs 2A, C); AME very slightly smaller than ALE, or equal in diameter, AME closer to ALE than to each other; in dorsal view, PME and PLE nearly equal in diameter, well-separated, PME located about 2���3 diameters apart, PME���PLE distance ca. 2/3 PME���PME distance. Legs: leg formula 4213; anterior legs with conspicuous scopulae disto-prolaterally on tibiae, and entire metatarsi and tarsi (Figs 2D, F); anterior femora with 3���5 dorsal spines, posterior femora with 5���7 dorsal spines; all patellae with 0���1 dorsal or retrolateral spine; tibiae I���II with 2 or 3 pairs of ventral spines, 1 retrolateral, and 1 prolateral spine; metatarsi I���II with 1 or 2 pair of spines, 1 retrolateral, and 1 prolateral spine; tibiae and metatarsi of posterior legs with more spines than anterior legs but spination varies among different individuals. Abdomen (Figs 4E, F 6A���C): elongate-oval, tapering posteriorly, dorsal scutum over 3/4 of length of abdomen; without distinct colour patterns in the type species and B. jucunda (with median band and chevrons in B. kirilli sp. n., see description below). Palp: femur and patella unmodified, tibia cylindrical with single, unbranched retrolateral apophysis (RTA), apophysis shorter than diameter of tibia; cymbium unmodified, lacking spines; bulb elongate-oval; subtegulum (ST) with characteristic prolateral extension (Figs 3A, C, 5A, C, 8A); sperm duct not meandering, U-shaped in ventral view of tegulum; tegulum with well-developed hook-shaped distal apophysis (TA); embolus helical, originating prolatero-distally, either filamentous and wrapping around dorsal side of tegulum (B. kirilli sp. n.) or shorter, tip directed either anteriorly (B. dubia) or antero-mesally (B. jucunda). Female. General characters as in males, from which the females differ by (1) the cheliceral base without a mesal ridge (Fig. 2B); (2) the abdomen without a scutum (Figs 1A, 4G); and (3) a denser ���scopula��� on anterior legs (Fig. 2E). Epigyne: epigynal plate with large fovea, located posteriorly, with 1 anterior (AH) and 2 posterior hoods (PH); copulatory openings (CO) small and indistinct, located in central portion of fovea; copulatory ducts (CD) hyaline, strongly convoluted; receptacles (R) small and located posteriorly; bursae (BS) situated anteriorly, balloon shaped, larger than receptacles. Composition. Bucliona dubia (Saint Helena Island), B. kirilli sp. n. (Kenya), and B. jucunda (Far East Russia, China, Korea, Japan)., Published as part of Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao & Li, Shuqiang, 2021, Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae), pp. 195-207 in Zootaxa 5006 (1) on pages 196-197, DOI: 10.11646/zootaxa.5006.1.21, http://zenodo.org/record/5157237, {"references":["Benoit, P. L. G. (1977) Fam. Clubionidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences Zoologiques (Zool. - Ser. 8 °), 220, 64 - 81.","Mikhailov, K. G. (1994) Bicluona Mikhailov, subgen. n., a new subgenus of spiders of the genus Clubiona (Aranei, Clubionidae) from eastern Asia. Zoologicheskii Zhurnal, 73 (11), 52 - 57.","Karsch, F. (1879) Baustoffe zu einer Spinnenfauna von Japan. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 36, 57 - 105.","Mikhailov, K. G. (1997) Redescription of Clubiona dubia O. Pickard-Cambridge, 1869 (Aranei Clubionidae) from Saint Helena Island. Arthropoda Selecta, 6 (1 / 2), 95 - 97.","Pickard-Cambridge, O. (1870) Notes on some spiders and scorpions from St Helena, with descriptions of new species. Proceedings of the Zoological Society of London, 37 (3, 1869), 531 - 544, pl. 42."]}

Clubiona reclusa O. Pickard-Cambridge, 1863 Material. 1 ♂, 1 ♀ (NMNHS), Vitosha Mts, Bistritsa village, N 42.5927 ˚, E 23.3401 ˚, 1070 m, 03.07.1982, 1 ♀ (NMNHS), 17.10.1982, L. Penev leg.; 1 ♂ (NMNHS), Vitosha Mts, Tintyava hut, N 42.6178 ˚, E 23.2584 ˚, 1620 m, 27.06.1984, 1 ♀ (NMNHS), 19.07.1984, C. Deltshev leg. Remarks. Widespread in Europe and Turkey to South Siberia and Kazakhstan (Nentwig et al. 2021; World Spider Catalog 2021)., Published as part of Naumova, Maria, Blagoev, Gergin & Deltshev, Christo, 2021, Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae), pp. 228-257 in Zootaxa 4984 (1) on page 231, DOI: 10.11646/zootaxa.4984.1.18, http://zenodo.org/record/4927030, {"references":["World Spider Catalog (2021) World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 28 February 2021) https: // doi. org / 10.24436 / 2"]}

This study is a part of an ongoing comprehensive inventory of Bulgarian spiders. A total of fifty spider species belonging to thirteen families are reported for the first time from Bulgaria. Four species are rejected from the Bulgarian spider checklist due to misidentification: Callilepis concolor Simon, 1914, Centromerus capucinus (Simon, 1884), Hoplopholcus labyrinthi (Kulczyński, 1903) and Mansuphantes prope fragilis (Thorell, 1875). Another four species (Drassodes villosus (Thorell, 1856), Entelecara flavipes (Blackwall, 1834), Lepthyphantes notabilis Kulczyński, 1887 and Singa semiatra L. Koch, 1867) are rejected after a new interpretation of the locations. Six species were omitted from the list of Bulgarian spiders as obviously doubtful records (Dysdera nicaeensis Thorell, 1873, Haplodrassus rufipes (Lucas, 1846), Harpactea hispana (Simon, 1882), Macaroeris cata (Blackwall, 1867), Nomisia celerrima (Simon, 1914) and Tenuiphantes monachus (Simon, 1884)). A new synonymy is proposed for Cyclosa strandjae Drensky, 1915 (= Cyclosa sierrae Simon, 1870 syn. nov.). In addition, new images with essential taxonomic value are provided for twenty-five species to facilitate their identification or to illustrate their intraspecific variability. To ensure correct identification, DNA barcoding was additionally used in some species.

ClubionaLatreille, 1804, with more than 500 named species, is one of the largest genera of Araneae. The genus has 15 synonyms, most of which are not listed in the World Spider Catalog (2018) and unknown to many arachnologists. The most comprehensive survey ofClubionasensu lato by Wunderlich (2011) also lacked a few synonyms. In this paper all genus group names described inClubionaare listed with their type species. Most of these names correspond to the species groups recognised inClubionasensu lato. We agree thatPorrhoclubionaLohmander, 1944 (=Clubionagenevensis-group) deserves a status of a separate genus and provide the diagnosis of this taxon. Three species ofPorrhoclubionathat occur in Central Asia are surveyed, and two of them are described as new to science:P.laudata(O. Pickard-Cambridge, 1885),comb. n. (♂♀, Xinjiang, Tibet, China),P.bosmansisp. n.(♂♀, Tajikistan), andP.moradmandisp. n.(♂♀, Fars, Iran). It seems that all records ofP.genevensisL. Koch, 1866 from China refer toP.laudata. The records ofClubionavegetaSimon, 1918 from Tajikistan and Iran refer toP.bosmansisp. n. andP.moradmandisp. n., respectively. The following new combinations have been established:Porrhoclubionadecora(Blackwall, 1859),comb. n.,P.diniensis(Simon, 1878),comb. n.,P.leucaspis(Simon, 1932),comb. n.,P.minor(Wunderlich, 1987),comb. n.,P.pseudominor(Wunderlich, 1987),comb. n.,P.pteronetoides(Deeleman-Reinhold, 2001),comb. n.,P.vegeta(Simon, 1918),comb. n.,P.viridula(Ono, 1989),comb. n., andP.wunderlichi(Mikhailov 1992),comb. n.(all ex.Clubiona). SEM study of the structure considered earlier as scopula inClubionaandPorrhoclubionareveals that it is represented by several lateral rows of movable macrosetae (spines) with a locking mechanism.

The Clubiona zilla-group is a relatively small species group, distributed exclusively in East Asia, with only three species clearly documented so far. Clubiona hooda Dong & Zhang, 2016, which was previously placed in the C. trivialis-group, is assigned to the C. zilla-group in the present paper. A new spider of the C. zilla-group from Jiugong Mountain in China is described under the name of C. jiugong sp. nov. Detailed descriptions and photographs of the new species are provided.

En esta nota se ofrece una relción de nuevas localidades para diversas especies de las familias Clubionidae, Zoridae, Sparassidae, Thomisidae, y Philodromidae. Se amplian con ello considerablemente los datos faunísticos para la zona de estudio, añadiéndose algunas precisiones taxonómicas, y otros comentarios de interés entorno a las especies mencionadas. In this paper new records of several species of families Clubionidae, Zoridae, Sparassidae, Thomisidae, and Philodromidae, are ennumerated. New data extend considerably the faunistical image. Some remarks, and others comments of interest are introduced.

CLUBIONIDAE Unidentified sp.: Auplopus militaris., Published as part of Kurczewski, Frank E., West, Rick C., Waichert, Cecilia, Kissane, Kelly C., Ubick, Darrell & Pitts, James P., 2020, New and unusual host records for North American and South American spider wasps (Hymenoptera: Pompilidae), pp. 1-112 in Zootaxa 4891 (1) on page 111, DOI: 10.11646/zootaxa.4891.1.1, http://zenodo.org/record/4309249

Kurczewski, Frank E., West, Rick C., Waichert, Cecilia, Kissane, Kelly C., Ubick, Darrell, Pitts, James P. (2020): New and unusual host records for North American and South American spider wasps (Hymenoptera: Pompilidae). Zootaxa 4891 (1): 1-112, DOI: https://doi.org/10.11646/zootaxa.4891.1.1

Clubiona milingae Barrion-Dupo, Barrion & Heong, 2013 was described from a single male and no additional specimens have been recorded. The original description was brief and the illustrations were inadequate. Clubiona milingae is redescribed and illustrated based on new material from the type locality and the new distribution region (Jianfeng Mountains and Limu Mountains of Hainan Island, China). The female is reported for the first time.

Pristidia cervicornutaYu, Zhang & Chen, 2017 is redescribed based on new material from the type locality, Diaoluo Mountains of Hainan Island, China. The female is described and illustrated for the first time. In addition, this paper further illustrates the male, and provides a supplementary description.