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The Florida State Collection of Arthropods (FSCA) is one of the top insect collections in the United States with in excess of eight million curated specimens and significant amounts of materials in bulk collections and other unprocessed samples. The subphylum Myriapoda currently comprises approximately 17,500 species in the four classes Diplopoda, Chilopoda, Symphyla, and Pauropoda. The FSCA houses type material in the Diplopoda orders Callipodida, Chordeumatida, Glomeridesmida, Julida, Polydesmida, Polyzoniida, Siphonophorida, Spirobolida, and Spirostreptida. This catalog documents the FSCA primary type material for 74 species in 45 genera in 25 families and nine orders within Diplopoda.

Petra sierwaldae, n. gen., n. sp. is described from males and females collected at four localities in Kootenai County, Idaho, USA. The new genus is defined by its unique gonopod and ninth leg anatomy, a notched collum and single ommatidium on each side of the head, and lacking third coxal flasks as well as having modified fifth coxae.

Calityla humboldtensis Shear, Richart & Wong, 2020: 20 Holotype: 1 ♂. CONOTYLIDAE / Calityla humboldtensis Shear et al/ MALE HOLOTYPE // CALIFORNIA: Humboldt Co., 2.75/ mi NNE Orleans, 800′ asl, 21/ December 1976, R. K. Johnson // [barcode label] FSCA 00070773. Conservation status: Good condition; in 75% EtOH., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 232, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A., Richart, C. H. & Wong, V. L. (2020) The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea). Zootaxa, 4753 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 4753.1.1"]}

Ralampia complexa Shear & Marek, 2022: 524 Holotype: 1 ♂. STRIARIIDAE / Ralampia complexa Shear & Marek / MALE HOLOTYPE // CALIFORNIA: Del Norte Co./ Patrick Creek Campground on Rt. 199, 7/ mi by road ENE of Gasquet, 41.8718°, -/ 123.8456°. 990′ asl/ 22 December 1977 A. R. Johnson/ Parts on SEM stub WS35-9 // FSCA // [hw label] Calif: Del Norte Co: Patrick Cr./ Campground on Hwy 199 at 7 mi. / (by rd.) ENE Gasquet, 900′./ 22 Dec 1977. A. K. Johnson. // [barcode label] FSCA 00051180. Conservation status: Good condition; in 75% EtOH, specimen in microvial.

Dybasia interamericana Loomis, 1974: 184 Holotype: 1 ♂. HOLOTYPE / 1♂/ DIPLOPODA / CLEIDOGONIDAE / Dybasia interamericana / Loomis/ det. H. F. Loomis/ COSTA RICA: Prov. of/ San Jose, San Isidro,/ Cerro de la Muerte,/ Iter-American Hwy./ 16-IV-1972 / E. M. & H. F. Loomis/ altitude 9,600 ft. // [barcode label] FSCA 00070696. Conservation status: Fair condition; in 75% EtOH., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 232, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Loomis, H. F. (1974) Millipeds from southern Costa Rican highlands. Florida Entomologist, 57 (2), 169 - 187. https: // doi. org / 10.2307 / 3493479"]}

Plaramia arcata Shear & Marek, 2022: 520 Holotype: 1 ♂. STRIARIIDAE / Plaramia arcata Shear & Marek / MALE HOLOTYPE // FSCA // [hw label] Calif: Humboldt Co: North Side Jolly/ Giant Canyon, 400–600′, Arcata/ Community Forest. 21 Feb 1978./ A. K. Johnson // [barcode label] FSCA 00051178. Conservation status: Good condition; in 75% EtOH, specimen in microvial., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 233, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. & Marek, P. E. (2022) The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatidae, Striarioidea). Zootaxa, 5205 (6), 501 - 531. https: // doi. org / 10.11646 / zootaxa. 5205.6.1"]}

Scoterpes jackdanieli Shear, 2010: 58 Holotype: 1 ♂. Motlow Cave, Moore Co. Tenn. / 17.i.1960 T C Barr Jr. // Scoterpes rudis Causey / det. Causey 1964 // HOLOTYPE/ Scoterpes jackdanieli Shear // PARATYPES / Scoterpes rudis Causey // [barcode label] FSCA 00051059. Conservation status: Fair condition; in 75% EtOH; specimen in microvial. Additional notes: Three female paratypes included: FSCA 00051153–00051155; for notes on the manuscript name Scoterpes rudis see Shear (2010)., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 234, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2010) The milliped family Trichopetalidae, Part 2: The genera Trichopetalum, Zygonopus and Scoterpes (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 2385, 1 - 62."]}

Causeyella causeyae Shear, 2003: 30 Holotype: 1 ♂. TRICHOPETALIDAE / Causeyella causeyae Shear / MALE HOLOTYPE // ARKANSAS: Independence Co./ Foushee Cave, 6 mi W of Locust/ Grove/ May 1978 Collector Unknown // [barcode label] FSCA 00051052. Conservation status: Good condition; in 75% EtOH., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 233, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 321, 1 - 36."]}

Nannopetalum fontis Shear, 2003: 17 Holotype: 1 ♂. TRICHOPETALIDAE / Nannopetalum fontis Shear / MALE HOLOTYPE // Springville Cave, Springville,/ St. Clair Co., ALA. [hw] D2 31. V.1962 [/hw] // [hw label] III-27-63 // FSCA // [barcode label] FSCA 00070684. Conservation status: Fair condition; in 75% EtOH., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 233, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 321, 1 - 36."]}

Scoterpes stewartpecki Shear, 2010: 51 Holotype: 1 ♂. ALABAMA: Javkson Co., Jess / Elliott C. #323, 3 mi E./ Mytop ix.3.65 S. Peck // [label hw] S.3 07 // FSCA // [barcode label] FSCA 00051156. Conservation status: Fair condition; in 75% EtOH; specimen in microvial. Additional notes: Three female paratypes included: FSCA 00051157–00051159., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 234, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2010) The milliped family Trichopetalidae, Part 2: The genera Trichopetalum, Zygonopus and Scoterpes (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 2385, 1 - 62."]}

Scoterpes tombarri Shear, 2010: 50 Holotype: 1 ♂. KY: Warren Co/ Plano Saltpeter Cave / 10 mi S Bowling Green / 18. vi. 73, S Peck // [hw pencil] S—16 // [barcode label] FSCA 00051056. Conservation status: Good condition; in 75% EtOH, specimen in microvial. Additional notes: Sixteen female paratypes included: FSCA 00051060–00051075., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 234, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2010) The milliped family Trichopetalidae, Part 2: The genera Trichopetalum, Zygonopus and Scoterpes (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 2385, 1 - 62."]}

Cleidogona bifurcata Loomis, 1974: 184 Holotype: 1 ♂. HOLOTYPE / 1♂/ DIPLOPODA / CLEIDOGONIDAE / Cleidogona bifurcata / Loomis/ det. H. F. Loomis/ COSTA RICA: Prov. of/ Cartago, Dulce Nombre/ 20-IV-1972 / E. M. & H. F. Loomis // [in microvial][hw label] HOLOTYPE ♂ // [barcode label] FSCA 00070697. Conservation status: Fair condition; in 75% EtOH; specimen in microvial., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 232, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Loomis, H. F. (1974) Millipeds from southern Costa Rican highlands. Florida Entomologist, 57 (2), 169 - 187. https: // doi. org / 10.2307 / 3493479"]}

Trigenotyla seminole Shear, 2003: 23 Holotype: 1 ♂. TRICHOPETALIDAE / Trigenotyla seminole Shear / MALE HOLOTYPE // Whiskey Cave, 9 mi SE Bowlegs/ Seminole Co., Oklahoma / 14.j.1974 H H Black // [barcode label] FSCA 00051057. Conservation status: Good condition; in 75% EtOH, specimen in microvial., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 234, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 321, 1 - 36."]}

Causeyella youngsteadtorum Shear, 2003: 31 Holotype: 1♂.TRICHOPETALIDA/ Causeyella youngsteadtorum /Shear/ MALE HOLOTYPE // ARKANSAS: Boone Co. / Brewer Cave, T17 N, R18 W, S8 (SE)/ July / August 1974 // [barcode label] FSCA 00051152. Conservation status: Fair condition; in 75% EtOH. Additional notes: Label data states “ T 17N, R 18W, S8 (SE), while the original description (Shear 2003) states “ T 16N, R 18W, S8”., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 233, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 321, 1 - 36."]}

Trigenotyla blacki Shear, 2003: 22 Holotype: 1 ♂. TRICHOPETALIDAE / Trigenotyla blacki Shear / MALE HOLOTYPE // Stansberry—January Cave/ 4 mi. N. of Colcord/ Delware Co., Okla/ 1/13/71/ J. H. Black // [barcode label] FSCA 00051058. Conservation status: Good condition; in 75% EtOH, specimen in microvial., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 234, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 321, 1 - 36."]}

Martenseya minutocaeca Shear, 2021: 111 Holotype: 1 ♂. CASEYIDAE / Martenseya minutocaeca Shear / MALE HOLOTYPE // CALIFORNIA: Marin Co./ Phoenix Lake, 0.5 mi W of Kentfield,/ 37.9552°, -122.5763°, 200′ asl/ 30 December 1977 A. K. Johnson // [hw label] Calif: Marin Co: Phoenix Lake,/ 200′, 1 ½ mi W Kentfield./ 30 Dec 1977. A. K. Johnson. // FSCA // [barcode label] FSCA 00070667. Conservation status: Good condition; in 75% EtOH., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 232, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. (2021) Additions to the millipede family Caseyidae. II. Martenseya, a new genus of miniature, blind millipedes from California (Diplopoda, Chordeumatida, Striariidea, Striarioidea, Caseyidae). Zootaxa, 4984 (1), 108 - 113. https: // doi. org / 10.11646 / zootaxa. 4984.1.10"]}

Maraplia schusteri Shear & Marek, 2022: 509 Holotype: 1 ♂. STRIARIIDAE / Maraplia schusteri Shear & Marek / MALE HOLOTYPE // FSCA / CALIFORNIA: Butte Col/ near Stringtown Hill, NE of/ Oroville/ 9Feb 1956 R. O. Schuster // [barcode label] FSCA 00051181. Conservation status: Good condition; in 75% EtOH, specimen in microvial., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 232, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Shear, W. A. & Marek, P. E. (2022) The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatidae, Striarioidea). Zootaxa, 5205 (6), 501 - 531. https: // doi. org / 10.11646 / zootaxa. 5205.6.1"]}

Trigenotyla parca Causey, 1951: 118 Holotype: 1 ♂. [hw] TRIGENTALUM GLESUM Causey/ ♂ holotype — ♀ paratype. X–29–49./ Blue Spg. Carroll Co.,Ark. Coll. N. Causey [/hw] // Chordeumatida: Trichopetalidae /?Trigenotyle sp., M, F/ Det. Shelley, 1999 // TRICHOPETALIDAE / T rigenotyle parca Causey / det. Shear 2003 // FSCA // [barcode label] FSCA 00051054. Conservation status: Good condition; in 75% EtOH, specimen and paratype in microvial. Additional notes: holotype gonopods missing. ♀ paratype in vial, same location data; FSCA 00051055., Published as part of Keller, Oliver, Dunn, Ann & Soto-Adames, Felipe, 2023, An annotated catalog of the primary type material of Myriapoda deposited in the Florida State Collection of Arthropods in Gainesville, FL, USA, pp. 230-250 in Zootaxa 5293 (2) on page 234, DOI: 10.11646/zootaxa.5293.2.2, http://zenodo.org/record/7960066, {"references":["Causey, N. B. (1951) On two new colobognath millipeds and records of some established species from east of the Rock Mountains. Proceedings of the Biological Society of Washington, 64, 137 - 140.","Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea). Zootaxa, 321, 1 - 36."]}

Two new genera and species, Stegostriaria dulcidormus (Trisariinae) and Kentrostriaria ohara (Striariinae), are described from Oregon and Idaho, respectively. The new species are distinct from any other striariids, and indeed from any other chordeumatidan millipedes, in having the second dorsal crests greatly enlarged, giving them a fanciful resemblance to stegosaurid dinosaurs. In spite of this similarity, the two species are so distinct they cannot be accommodated in the same genus or the same subfamily. The exaggerated metazonital crests are therefore attributed to convergent evolution. We present a key to the genera of the family Striariidae.

Kingaria Shear & Marek, new genus Type species: Kingaria prattensis Shear & Marek, n. sp. Diagnosis. This is the only genus of Striariidae known to have an internal, glandular apophysis on the mandibular stipes of males. Etymology. The name uses the county of the type locality (King Co., Washington) with the combining stem - aria, used in other striariid generic names. The gender is feminine. Description. As for the only included species, Kingaria prattensis Shear & Marek, n. sp. Distribution. Known only from the type locality of the single species in King Co., Washington., Published as part of Shear, William A. & Marek, Paul E., 2023, The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), pp. 323-340 in Zootaxa 5264 (3) on page 334, DOI: 10.11646/zootaxa.5264.3.2, http://zenodo.org/record/7836674

Kingaria prattensis Shear & Marek, n. sp. Figs 35–38, 60, 61. Types: Male holotype from Pratt Lake trailhead, north of I-90, 47.39715°N, - 121.48515°W, collected 25 October 2003 by W. Leonard. The holotype is mounted on SEM stub WS36-6. Deposited in CAS. Diagnosis. As for the genus, above. Etymology. The species name refers to the Pratt Lake Trail. This is not the same as “ pratensis, ” a Latin word meaning “of the meadow.” Description. Male holotype from Pratt Lake trailhead. Length 4.0 mm, width 0.45 mm. Twenty-eight postcephalic rings. Color white, with single black ommatidium on each side of head. Antennae robust, clavate; fifth antennomere longest, widest. Head (Fig. 35) except for labrum covered with fine setae and small tubercles. Labrum (lab, Fig. 35) not flattened, tapering, corners rounded. Collum ornamented by closely set small tubercles, becoming indistinct crests only mesally on most posterior part. Metazonites with twelve subequal crests (Fig. 37); metazonital setae long, prominent, with obscure longitudinal grooves and feathered tips occupying about 1/6 length of seta. Telson with indentations between lobes nearly obsolete, median lobe short; spinnerets directed posteriorly. Legs with specialized flattened setae with median ridges and long, filamentous extensions. The following secondary sexual characters occur in males. Head evenly rounded; labrum (lab, Fig. 35) with few setae, distal corners rounded. Mandibular stipes (m, Fig. 35) with serrate anteriodistal margin, angular blunt distal tooth, inner suface of stipes with strongly projecting, rounded apophysis bearing on its posterior surface a small pore (Figs 35, 36). First legpair enlarged, femora curved, lacking needle setae, distal podomeres without flattened setae ventrally but with tarsal comb. Second legpair shorter than first, openings of vas deferentia separate, subtended by long, flattened, ribbon-like setae. Third legpair less robust than first two, coxal flasks short (cf, Fig. 38), without subapical branch, strongly procurved and densely set with curled setae. Fourth legpair the largest; prefemora and femora of usual proportions. Fifth, sixth and seventh legpairs enlarged; without modified coxae. Tenth coxae with glands, not modified or enlarged. Gonopods (Fig. 60) separate from prominent, transverse sternum, coxae large, with two distinct groups of setae. Anterior angiocoxites (aac, Fig. 60) short, posterior margin membranous-fimbriate, without subdistal branch. Posterior angiocoxites (pac, Fig. 60) longer than anterior ones, posteriorly fimbriate, apparently not sheathing curved single flagellum or flagellocoxite (f, Fig. 60). Colpocoxites (cc, Fig. 60) as single, short, curved process. Ninth legs (Fig. 61) free from sternum, coxae and telopodites fused, coxae with strongly curved single process (cxp, Fig. 61) pore not observed; telopodites strongly flattened, setose, with ornament of pointed tubercles (as on other legs). Tenth coxae with pores, not modified. Females not collected. Distribution. Known only from the type locality. Notes. The glandular apophysis on the mandibular stipes of the male (Figs 35, 36) is unique among chordeumatid millipedes, so far as we are aware. The gonopods of this species are difficult to interpret, not closely resembling those of any other striariids. What we are calling the posterior angiocoxite could well be the subapical branch of the anterior angiocoxite, but that would mean that the posterior angiocoxite is missing. The unusual position of the flagellocoxite (or flagellum) shown in Fig. 60 could be a result of mounting the gonopod on a microscope slide with the pressure of the coverglass. Unfortunately a lack of material prevented us from making further observations., Published as part of Shear, William A. & Marek, Paul E., 2023, The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), pp. 323-340 in Zootaxa 5264 (3) on pages 334-339, DOI: 10.11646/zootaxa.5264.3.2, http://zenodo.org/record/7836674

Tigraria Shear & Marek, new genus Type species: Tigraria oregonensis Shear & Marek, new species Diagnosis. Distinct from other small (Petra Shear et al., 2022 in lacking flasks on the third coxae (Fig. 31) and having densely dentate processes on the fifth coxae (Fig. 32) of males. Distinct from Petra sierwaldae Shear et al., 2022 in that the gonopod tips are not enveloped in the telopodites of the ninth legpair but instead the ninth legpair (Figs 34, 59) resembles that of species of Miniaria n. gen. Etymology. The genus name refers to the type locality (Tiger Saddle) with the combining stem - aria commonly used in this family; gender is feminine. Description. As for the only included species, see below. Included species. Only Tigraria oregonensis Shear & Marek, n. sp. Distribution. A single locality in Umatilla Co., Oregon, USA., Published as part of Shear, William A. & Marek, Paul E., 2023, The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), pp. 323-340 in Zootaxa 5264 (3) on pages 331-332, DOI: 10.11646/zootaxa.5264.3.2, http://zenodo.org/record/7836674

Miniaria ramifera Shear & Marek, n. sp. Figs 1–11, 39–45 Types: All specimens deposited in CAS. Male holotype, 6 male paratypes and 6 female paratypes from Mashel River at State Route 7, 46.86695°N, - 122.33645°W, 600′asl, Pierce Co., Washington, collected 19 March 2004 by W. Leonard and C. Richart. Parts of the holotype are on SEM stub WS36-8. Diagnosis. Distinct from M. richarti n. sp. in having the gonopod anterior angiocoxite with a prominent subterminal posterior branch (aac, Figs 9, 11, 39–44). The gonopods are widely separated mesally, while those of M. richarti n. sp. are appressed to each other in the midline. The anterior angiocoxites are evenly curved, not sharply elbowed as in M. richarti n. sp.. Etymology. The species name is a Latin adjective, meaning “branch-bearing,” and refers to the subterminal posterior branch of the gonopod anterior angiocoxite. Description. Male paratype from Mashel River. Length 3.5 mm, greatest width 0.31 mm. Nonsexual characters and secondary sexual modifications as described for genus.Gonopods (Figs 9, 11, 39–44) small; anterior angiocoxites (aac, Fig. 9) curved posteriad, slightly knobbed at tips, distally with prominent fringed margin, long, flattened subterminal process. Posterior angiocoxites (pac, Fig. 9) less than half length of anterior ones, fimbriate especially along posterior margin, sheathing single curved flagellocoxite or flagellum. Colpocoxite reduced to single branch with membranous base. (cc, Figs 39–44). Ninth legs (Fig. 10) as described for genus. Females similar to males in all nonsexual respects. Distribution. OREGON: Multnomah Co.: Ainsworth State Park, 45.596°N, - 122.051°W, 210′ asl, 15 November 1967, E. M. Benedict, 2 mm. Gobbert Butte, Gresham, 45.4731°N, - 122.4378°W, 30 October 2016, P. Nosler, m. Washington Co.: 1.7 mi W of Timber, T3N, R5W, sec.?, 300′ asl, 27 November 1971, E. M. Benedict, m. WASHINGTON: Pierce Co.: Ohop Valley Road at State Route 7, 600′ asl, 46.86695°N, - 122.33645°W, 12 January 2004, W. Leonard, m. Skamania Co.: Cook-Underwood Road, 1.4 mi NE of Cook, 450′ asl, 45.7239°N, - 121.64765°W, 30 November 2003, W. Leonard, 4 mm. Notes. Gonopod variation in this species focuses on the degree to which the anterior angiocoxite is terminally knobbed and the extensiveness of the fimbriated posterior distal margin. In the Ainsworth State Park male (Fig. 43), the fimbriate part appears as a coalesced process. In the Timber (Fig. 44) and Mashel River (Figs 39, 40) males, it is more extensive and does not protrude, and while in the male from Cook-Underwood Road (Figs 41, 42) there is a broad fimbriate region. We do not have enough localities to determine if this is geographic variation or simply due to positioning on a microscope slide., Published as part of Shear, William A. & Marek, Paul E., 2023, The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), pp. 323-340 in Zootaxa 5264 (3) on pages 326-328, DOI: 10.11646/zootaxa.5264.3.2, http://zenodo.org/record/7836674

Shear, William A., Marek, Paul E. (2023): The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea). Zootaxa 5264 (3): 323-340, DOI: 10.11646/zootaxa.5264.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5264.3.2

Miniaria Shear & Marek, new genus Type species: Miniaria ramifera Shear & Marek, new species Diagnosis. A genus of Striariinae consisting of two species, distinct from other small striariids in the form of the male labrum, which is convex distally and with sharp lateral corners, but lacking curved processes (lab, Figs 1, 12). Additional distinguishing characters of the males: first legpair (Fig. 2) lacks needle setae; flasks of coxae 3 very short, with subterminal processes; fourth legpair largest but with short, wide podomeres (L4, Fig. 4); gonopod colpocoxites strongly reduced (Figs 9, 19); ninth legpair with coxal pores and curved coxal processes (cxp, Figs 10, 25). Etymology. The genus name is feminine in gender and is an arbitrary combination of letters. Description. Small species of Striariinae with 28 postcephalic rings. Length 3–4 mm, width about 0.25–0.30 mm. Color white, with single black ommatidium on each side of head. Antennae relatively short, robust, clavate. Head, except for labrum, covered with fine setae and small tubercles (Fig. 1). Collum ornamented by closely set small tubercles, coalescing into vague crests in posterior fifth of collum length. Metazonites with twelve subequal crests (Figs 13, 14); metazonital setae long, prominent, with longitudinal grooves and feathered tips (Figs 15). Telson with three lobes only shallowly separated, not prominent; spinnerets directed posteriad (sp, Fig. 16). Legs with specialized flattened setae with median ridges and long, filamentous extensions (Fig. 6). The following secondary sexual characters occur in males. Head frontally flattened with slight swellings below antennal sockets. Labrum flat and smooth, slightly concave, distal corners distinctly angular, not rounded (lab, Fig. 1). Mandibular stipes with serrate anteriodistal margin, prominent distal tooth (m, Fig 1). First legpair (Fig. 2) enlarged, femora curved, lacking needle setae, distal podomeres with few long, flattened setae ventrally. Second legpair larger than first; openings of vas deferentia separate, subtended by long, flattened, ribbon-like setae (vd, Figs 3, 5). Third legpair robust, coxal flasks short, with subterminal process (Fig. 5); tarsi with special sensory array including comb setae and recessed pore plate (Fig. 6; see Discussion section below for further details). Fourth legpair the largest; prefemora and femora nearly as wide as long (L4, Figs 4, 8). Fifth, sixth and seventh legpairs enlarged; seventh coxae unmodified. Tenth coxae with glands, not modified or enlarged (cx10, Fig. 10). Gonopods (Figs 9, 11, 19–24, 39–44, 46–53), separate from prominent, transverse sternum, coxae large, with two or three setae. Anterior angiocoxites curved, with fimbriate posterior margin distally. Posterior angiocoxites smaller than anterior ones, fimbriate, sheathing single flagellum or flagellocoxite. Colpocoxites membranous-fimbriate, much reduced. Ninth legs (Figs 10, 25, 45), variably free or partially fused to sternum; coxae and telopodites fused, coxae with prominent anterior lobe bearing gland pore, also with strongly curved single process; telopodites strongly flattened, setose, with ornament of pointed tubercles (as on other legs)., Published as part of Shear, William A. & Marek, Paul E., 2023, The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), pp. 323-340 in Zootaxa 5264 (3) on page 325, DOI: 10.11646/zootaxa.5264.3.2, http://zenodo.org/record/7836674

Tigraria oregonensis Shear & Marek, n. sp. Figs 26–34, 57–59 Types: All specimens deposited in CAS. Male holotype, two male and four female paratypes from Umatilla National Forest, 0.2 mi E of Tiger Saddle, 45.944417°N, - 118.009433°W, 4766′ asl, collected 24 October 2003 by W. Leonard. Diagnosis. As for the genus, see above. Etymology. The species name, an adjective, refers to the occurrence of the species in the state of Oregon. Description. Male paratype from Tiger Saddle. Length 4.3 mm, width 0.5mm. Twenty-eight postcephalic rings. Color white, with single black ommatidium on each side of head. Antennae robust, clavate; fifth antennomere longest, widest. Head (Fig. 26) except for labrum covered with fine setae and small tubercles. Labrum not flattened, tapering, corners rounded (lab, Fig. 26). Mandibular stipes (m, Fig. 26) with serrate edge suppressed, distally flattened and extended. Collum (col, Fig. 27) ornamented by closely set small tubercles, becoming indistinct crests only mesally on most posterior part. Metazonites with twelve subequal crests (Fig. 28); metazonital setae (Fig. 29) long, prominent, with longitudinal grooves and feathered tips occupying about 1/3 length of seta. Telson with lateral lobes practically obsolete, median lobe slightly projecting; spinnerets directed posteriorly. Legs with specialized flattened setae with median ridges and long, filamentous extensions. The following secondary sexual characters occur in males. Head (Fig. 26) frontally flattened with slight swellings below antennal sockets; labrum with few setae, distal corners rounded. Mandibular stipes (Fig. 26) with weakly serrate anteriodistal margin, prominent blunt distal tooth. First legpair (L1, Fig. 26) enlarged, femora curved, lacking needle setae, distal podomeres with few long, flattened setae ventrally. Second legpair (L2, Fig. 26; Fig. 30) shorter than first, openings of vas deferentia separate (vd, Fig. 30), subtended by long, flattened, ribbon-like setae, femur distally swollen. Third legpair (L3, Fig. 26; Fig. 31) less robust than first two, coxal flasks absent but coxae complexly excavate; tarsi with special sensory array including comb setae and recessed pore plate. Fourth legpair the largest; prefemora and femora of usual proportions. Fifth coxae (cx5, Fig. 32) with prominent projecting knobs densely set with cuticular teeth, sixth and seventh legpairs enlarged; seventh coxae unmodified. Tenth coxae with glands, not modified or enlarged. Gonopods (Figs 33, 57, 58) separate from prominent, transverse sternum (s, Fig. 33), coxae large (cx, Fig. 33), with five or six setae. Anterior angiocoxites curved, distally attenuated (aac, Figs 33, 57, 58), posterior margin not fimbriate, and with long, robust, curved branch arising near base and appearing almost to be articulated. Posterior angiocoxites (pac, Figs 33, 57, 58) smaller than anterior ones, fimbriate, divided into basal and distal parts, sheathing short, curved single flagellum or flagellocoxite (f, Figs 57, 58). Colpocoxites (cc, Figs 57, 58) membranous-fimbriate, not much reduced. Ninth legs (Fig. 34, 59) free from sternum, coxae and telopodites fused, coxae with prominent anterior poorly sclerotized lobe lacking obvious gland pore—also with strongly curved single process (cxp, Fig. 34) broad at base, distally hooked, with deep recess (pore?) at base; telopodites strongly flattened, setose, with ornament of pointed tubercles (as on other legs). Tenth coxae with pores, not modified. Females similar in all nonsexual characters. Notes. Despite the great distance separating their type localities, this species shares some significant characters with Petra sierwaldae, from Idaho, specifically the lack of flasks on the male third coxae and the modified coxae of legpair 5. The gonopods and second and ninth legpairs, however, are very different, justifying a new genus for this Oregon species. Tigraria oregonensis gonopods, however, are more conforming to the plan found in species of Miniaria, with a branched anterior angiocoxite and a two-part posterior angiocoxite. The male ninth legs also suggest a close relationship with Miniaria. The mandibular stipes is unique in that instead of a tooth-like process distally, the stipes is flattened into an extended lamella. Parts of a specimen from near Tiger Saddle are on SEM stub WS35-17., Published as part of Shear, William A. & Marek, Paul E., 2023, The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), pp. 323-340 in Zootaxa 5264 (3) on page 332, DOI: 10.11646/zootaxa.5264.3.2, http://zenodo.org/record/7836674

Vieteuma hubeiensis Mauriès & Nguyen Duy-Jacquemin, 1997 Figs 11, 12 Materials examined: 20 males and 38 females, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Guanshan, Yanzi Cave, 31°28’52.67” N, 109°43’59.70” E, alt. 2230 m, 12 August 2022, X.K. Jiang & H.M. Chen leg; 1 male and 4 females, Yintiaoling National Nature Reserve, Guanshan, Gangkou Cave, 31°30’32.22” N, 109°41’33.65” E, alt. 2070 m, 13 August 2022, X.K. Jiang & H.M. Chen leg. Distribution. Shennongjia (Hubei) and Yintiaoling (Chongqing). Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 11–12. The type locality of this species is in Shennongjia National Natural Reserve, Hubei Province (Mauriès & Nguyen Duy-Jacquemin, 1997), bordering on Yintiaoling. It is not surprising to collect this species in the region.

The millipedes of Yintiaoling National Natural Reserve, Southwest China, were investigated. In total, thirteen species from five orders (Sphaerotheriida, Spirobolida, Callipodida, Chordeumatida and Polydesmida) and eight families were discovered. Among them, six species are new to science: Zephronia linkouzi sp. nov., Paracortina inflata sp. nov., Orthomorpha laminata sp. nov., Polylobosoma corollifera sp. nov., Epanerchodus wuxi sp. nov. and Riukiaria spina sp. nov. As usual, the polydesmidan family Paradoxosomatidae is the most diverse group, containing five species. All new species are described and illustrated, and photographs of the habitus and the gonopods of all species are also provided.

Maurieseuma nontronense (Mauriès & Kime, 1999) gen. nov., comb. nov. Figs 1, 2 Hylebainosoma (?) nontronensis (sic!) Mauriès & Kime, 1999: 371–373, fig. 2, table 2. Hylebainosoma nontronensis (sic!) — Tajovský et al. (2014: 502, 516–521, fig. 39, table 1); Telfer et al. (2015: 22–29, figs 5–9, table 2). Hylebainosoma nontronense — Kime & Enghoff (2021: 111, 221, map 385). Diagnosis. As for the monospecific genus. Name. Mauriès & Kime (1999) named the species after Nontron, the region where it was discovered. Material examined. 1 ♂ (IZB, JSP200701-001), 3 ♀♀ (IZB, JSP200701-002), 6 ♂♂ (JSC, JSP130413-059 - 060 -061 -062, JSP141223-001), 8 ♀♀ (JSC, JSP130413-063 - 064 -065); FRANCE, Corrèze, Brive-la-Gaillarde, N Donzenac, 236 m a.l.s., 45.235°N, 1.5454°W; 22.10.2009; leg. A. Schönhofer. 3 ♂♂ (IZB, JSP150101-004; JSC, -002 -003); UK, Wales, Cefn Forest, N Blackwood, 51.697°N, 3.2044°W; 02.10.2014; leg. C. Owen. 3 ♂♂ (ZSM, GBOL11814, GBOL12287; JSC, JSP141209-036), 1 ♀ (JSC, JSP141209-033); UK, Wales, Groesfaen Woods, 51.701°N, 3.2559°W; 25.11.2014; leg. C. Owen. 2 ♂♂ (ZSM, GBOL11812; JSC, JSP141209-024), 2 ♀ (ZSM, GBOL12284; JSC, JSP141209-026); UK, Wales, Cwm Kendon Woods, 51.682°N, 3.1527°W; 25.11.2014; leg. C. Owen. 2 ♂♂ (ZSM, GBOL11813, GBOL12285); UK, Wales, Coed-y-moeth Comnon, 51.704°N, 3.2187°W; 26.11.2014; leg. C. Owen. 2 ♀♀ (ZSM, GBOL12286, GBOL12344); UK, Wales, Newbridge, 51.661°N, 3.1571°W; 27.11.2014; leg. C. Owen. Description. Medium-sized haaseid, with body length ca 8–10 mm, and with 30 rings (including telson) in adults. Colour yellowish-brown patterns, ommatidia black (Fig. 1A). Head with frontal side slightly concave in males (Fig. 1B), convex in females (Fig. 1C), with 12–14 ommatidia arranged in triangle. Antennae elongated, slender, with dorsomedial and ventromedial lines of small papillae-like outgrowths. Body rings with dorsolateral humps, bearing stout, long and trichoid macrochaetae (Haasea type sensu Spelda 2015). Leg-pairs 1 and 2 (typically reduced in size) in both sexes with tarsal combs, femora with several long and robust setae; postfemora and tibiae each with several long and robust setae arranged in a group (Fig. 1D). Males. Pregonopodal legs 3–6 enlarged, leg 7 only slightly enlarged (Fig. 1A, B). Tarsal papillae present only in distal halves of legs 3–7. Postgonopodal legs (including legs 10 and 11) without tarsal papillae. Leg-pair 10 reduced in size compared to other legs, without incrassate podomeres of telopodite; coxae enlarged, each with coxal sac and a prominent distal process directed mesad, with a single seta and nipples (Fig. 1B, F). Leg-pair 11 slightly reduced in size compared to other walking legs; with coxal sacs; no other peculiarities. Anterior gonopods (Figs 1G, H, 2A, B). Sternum (s) wide, with medial sternal process (msp) and three groups of setae on each half: anterior, lateral and anterolateral (similar to Tajovský et al. 2014: cf. figs 22, 27). Medial sternal process laterally flattened, rather sigmoid, directed anteriad, half the size of angiocoxites. Angiocoxites (a) robust, oriented rather anteriad, with several structures. In mesal view each angiocoxite with posteroproximal papillated cone (pc) partially covered with papillae-like outgrowths. Similar field with papillae-like outgrowths present anteriorly to pc. Mesal process (mp) long, acuminate, sabre-like or slightly sigmoid, similar in height as the other parts. Posterior lateral process (plp) rises from the posteroproximal shoulder completely straight up to half its length, and then abruptly bends anteriad at an angle of ca. 110°, distally bifurcated. Anterodistal part of angiocoxite (dp) massive, long, spoon-shaped, distally enclosed with fimbriae- and finger-like outgrowths, at the base of them slightly thickened. At base of the dp a complex structure with “seminal” opening (so) and numerous fimbriaelike outgrowths present. A “seminal” canal (sc) and “seminal” chamber (sch) visible. Unclear if posterior lamella present. Colpocoxites [= Opuntia -like membranous lobe sensu Antić & Akkari (2020)] with two lobes, smaller anterior one (cc1) and larger posterior one (cc2) digitiform in mesal and rather subtriangular in posterior view, both covered by spicule-like outgrowths. Posterior gonopods (Fig. 1I). Sternum reduced. Telopodite (t) 2-segmented, podomere 1 strongly developed with long setae distally, podomere 2 short, almost as long as its radius. Coxal process (cp) strongly developed, but apparently lower than telopodite, proximally with mesal bulge (b), distally wide with a posterior flange with teeth of variable appearance. Females. Leg-pair 3 reduced to sternum and one podomere (Fig. 1C–E); possibly with protective function. Vulvae merged posterobasally (Figs 1D, 2C); operculum (o) with lateral and mesal groups of setae (mainly 3+3) which are smaller in size than the setae of the bursa; lateral valve (lv) of bursa well-developed, with a bulge at the base, covered with numerous spicule-like outgrowths, nipples and long setae; mesal valve (mv) posteriorly with two ridges, of which mesal one covered with spicule-like outgrowths, nipples and long setae. Inside the bursa several ampullae (am). Without postgenital plate. Distribution. Known from the Massif Central in France and South Wales in UK (Kime & Enghoff 2021; Telfer et al. 2015)., Published as part of Antić, Dragan & Spelda, Jörg, 2022, Maurieseuma, a new genus to accommodate Hylebainosoma nontronense Mauriès & Kime, 1999, and the revalidation of the genus Romanosoma Mauriès, 2015 (Diplopoda: Chordeumatida: Haaseidae), pp. 193-200 in Zootaxa 5219 (2) on pages 195-198, DOI: 10.11646/zootaxa.5219.2.8, http://zenodo.org/record/7413491, {"references":["Mauries, J. - P. & Kime, R. D. (1999) Description, ecologie et chorologie de trois especes nouvelles de diplopodes (Myriapoda, Diplopoda) des zones peripheriques du Massif central et du centre de la France. Zoosystema, 21 (2), 367 - 378.","Tajovsky, K., Mock, A. & Papac, V. (2014) The genus Hylebainosoma Verhoeff, 1899 (Diplopoda, Chordeumatida, Haaseidae): Redescription of Hylebainosoma tatranum, description of a new troglobiont species and notes to the Hylebainosoma - Romanosoma species group. Zootaxa, 3764 (5), 501 - 523. https: // doi. org / 10.11646 / zootaxa. 3764.5.1","Telfer, M. G., Gregory, S. J., Kime, R. D., Owen, C. & Spelda, J. (2015) Ceratosphys amoena Ribaut, 1920 and Hylebainosoma nontronensis Mauries & Kime, 1999 new to Britain (Diplopoda: Chordeumatida). Bulletin of the British Myriapod and Isopod Group, 28, 15 - 30.","Kime, R. D. & Enghoff, H. (2021) Atlas of European millipedes 3: Order Chordeumatida (Class Diplopoda). European Journal of Taxonomy, 769, 1 - 244. https: // doi. org / 10.5852 / ejt. 2021.769.1497","Spelda, J. (2015) Class Diplopoda Order Chordeumatida. Revista IDE @ - SEA, 26 B, 1 - 15.","Antic, D. & Akkari, N. (2020) Haasea Verhoeff, 1895 - a genus of tumultuous history and chaotic records - redefinition, revision of taxonomy and geographic distributions, with descriptions of two new species from Austria and Serbia (Diplopoda, Chordeumatida, Haaseidae). Zootaxa, 4798 (1), 1 - 77. https: // doi. org / 10.11646 / zootaxa. 4798.1.1"]}

Liaoninosoma marusiki sp. nov. Figs 1–27 Material examined. Holotype: 1 male (FSCB), China, Liaoning Prov., Fengcheng City, Mt. Phoenix, ca 40˚23′ N, 124˚05′ E, 280–550 m, 16– 18.10.2017, leg. Yu.M. Marusik; Paratypes: 7 males, 5 females (FSCB), 4 males, 4 females (ZMUM), same data as for holotype. Diagnosis. Same as for the genus. Description. Male. Length 7.0–8.0 mm, width about 1.0 mm with paraterga. Coloration in alcohol light beige. Antennae light beige. Legs beige-white. Eye patches black. Body with 30 rings (including telson). Head anterior part setose. Antennae long and slender (Fig. 1). Antennae in one male paratypes deformed, short, antennomeres almost globular. Eye patches triangular, with about 23 ommatidia. Collum semicircular. Both collum and ring 2 narrower than head with genae. Ring 2 somewhat wider than collum. Body width gradually increasing until ring 7, body parallel-sided on rings 7–21(22), thereafter gradually tapering. Paraterga beginning on ring 3, well developed on rings 6(7)–23, poorly developed on rings 24–26, onward missing. Paraterga of the ring 7 larger and broader. Metazonital macrochaetae in a transverse row on rings 27–29, like an extended (to different degrees) triangle on preceding rings. Anterolateral (median after Spelda 2001) macrochaetae shortest, caudolateral (exterior after Spelda 2001) and medial (interior after Spelda 2001) ones subequal in length. All macrochaetae pointed, but not very sharply so. Axial suture poorly-developed. Legs long and slender. Leg pairs 1 and 2 typically reduced in size, with usual tarsal brushes. Legs 3 and 4 not enlarged. Legs 5–7 enlarged, coxae with low subconical outgrowths, femora swollen and curved, tarsi very long, sabre-shaped (Figs 3 and 4). Sternites 3–7 frontally with a longitudinal roller (Figs 2–4). Legs 3–7 with tarsal papillae occupying about 1/3rd of the tarsus length distally (near claw) on leg pairs 3 and 4 (Fig. 2), while legs 5–7 with several (3–5) tarsal papillae near claw only (Fig. 14). Postgonopodal legs (including legs 10 and 11) without tarsal papillae. Claws of medium length. Claws of legs 1–4 at base with two small additional claws dorsally and a long setoid outgrowth ventrally. Claws of legs 5–7 at base with an outgrowth ventrally only (Fig. 14). Claws of postgonopodal legs (including legs 10 and 11) at base with a long setoid outgrowth ventrally and two small additional claws dorsally. Two additional claws gradually disappear toward the end of the body. Claws of hindmost legs at base without additional claws dorsally. Legs 10 and 11 with coxal glands (Figs 7, 8, 10, 11). Coxa 10 with a subconical process (c) setose apically (Figs 5–8). Trochanter 10 with low knob (t) setose apically (Fig. 6). Trochanter 11 with a caudoventral setose fingershaped process (d) rounded apically (Figs 10–13). Sternum 11 frontally with a longitudinal roller (Figs 10 and 11). Anterior gonopod coxosternum in anterior view with a central, small, conical papillate outgrowth and several setae (Fig. 25). Anterior gonopod telopodites (at) 1-segmented, relatively short (reaching middle of colpocoxite), flagelliform (ribbon-shaped in distal part), each positioned on posterior surface of posterior gonopod colpocoxite inside a shallow, partly open-edged sheath groove positioned on a broad, longitudinal structure (colpocoxite “pad’’ (cp) - a possible homologue of the lateral and mesal processes of the sheath groove) covered throughout with dense, forked and undivided spikes (Figs 15, 16, 19, 23) (the posterior angiocoxal process on the left is removed in Figs 15 and 16). Colpocoxite “pads” at base with the spear-shaped processes forming single structure (ms) (homologue of fused mesal processes of sheath groove). Posterior gonopod colpocoxites fused medially in basal half; their distal parts rectangular curved laterad and slightly caudad, posterior surface covered with tilted spikes. Colpocoxite apex (ax) as thin hook (Fig. 17). Subapically colpocoxite with a conical process (p). Each colpocoxite entire, undivided. Posterior angiocoxal process (pp) large, wide, curved c-shaped, with two basal blades: blade ba curved anteriad and blade b curved mesad, tightly clasping anterior gonopod (Figs 15, 18, 20) (the posterior angiocoxal process on the left is removed in Fig. 15). Angiocoxite (a) in posterior face with subglobule (sg) strongly protruding caudad and the proximal part covered with papillae and spikes (Figs 15, 18, 20). In anterior view the colpocoxite with an apically convex blade (bb) along its length (Figs 21, 22, 24). Two bb can be closed (Fig. 21) or located at some distance from each other (Figs 22, 24). Angiocoxite (a) in anterior face strongly elongated ventrally, convex centrally along its length (Fig. 21), however, it does not form an independent separate process (here I adhere to Shear’s (1990) opinion, see Discussion). Posterior gonopod telopodite 2-segmented, setose, located almost on the posterior surface of the gonopods; telopoditomere 1 with a very thin stem, telopoditomere 2 middle length. Female. Length 7.0–8.0 mm, width about 1.0 mm with paraterga. Claws of hindmost legs at base with two small additional claw-shaped knobs very closely pressed to main claw. Vulva as in Figs 26, 27. Etymology. The species is named in honor of the collector Dr. Yu. M. Marusik, a well-known Russian arachnologist. A noun in genitive., Published as part of Mikhaljova, Elena V., 2022, The millipede family Diplomaragnidae Attems, 1907 confirmed for mainland China, with descriptions of a new genus and new species (Diplopoda, Chordeumatida), pp. 185-192 in Zootaxa 5219 (2) on pages 186-191, DOI: 10.11646/zootaxa.5219.2.7, http://zenodo.org/record/7413471, {"references":["Spelda, J. (2001) Review of the millipede genus Pterygophorosoma Verhoeff, 1897 (Diplopoda, Chordeumatida, Craspedosomatidae). Andrias, 15, 29 - 48.","Shear, W. A. (1990) On the Central and East Asian milliped family Diplomaragnidae (Diplopoda, Chordeumatida, Diplomaragnoidea). American Museum Novitates, 2977, 1 - 40."]}

Key to the genera of the family Haaseidae 1. Coxal processes of the posterior gonopods surpassing telopodites in height............................. Romanosoma - Coxal processes of the posterior gonopods lower than the telopodites............................................ 2 2. Coxal processes of the posterior gonopods with two or three branches....................................... Haasea - Coxal processes of the posterior gonopods unipartite......................................................... 3 3. Medial sternal process of the anterior gonopods significantly exceeds half the height of the angiocoxites..... Hylebainosoma - Medial sternal process of the anterior gonopods shorter than half the height of angiocoxties.......................... 4 4. Coxal processes of the posterior gonopods robust and wide. Leg-pair 3 in females reduced to one podomere............................................................................................... Maurieseuma gen. nov. - Coxal processes of the posterior gonopods slender distally. Leg-pair 3 in females normal.................. Xylophageuma, Published as part of Antić, Dragan & Spelda, Jörg, 2022, Maurieseuma, a new genus to accommodate Hylebainosoma nontronense Mauriès & Kime, 1999, and the revalidation of the genus Romanosoma Mauriès, 2015 (Diplopoda: Chordeumatida: Haaseidae), pp. 193-200 in Zootaxa 5219 (2) on page 198, DOI: 10.11646/zootaxa.5219.2.8, http://zenodo.org/record/7413491

Liaoninosoma gen. nov. Type and only species: Liaoninosoma marusiki sp. nov., by present designation. Diagnosis. The genus differs from other genera of the family mainly by the posterior gonopod colpocoxite, which has an independent (separate), broad, longitudinal structure (“pad”) (cp), covered with dense spikes (Fig. 16), with a shallow, partly open-edged sheath groove for the anterior gonopod telopodite together with a rectangular laterally and slightly caudally curved distal part, in combination with a large posterior angiocoxal process. A structure similar to the above-mentioned “pad” is not observed among all known genera of the family. Description. Body with 30 rings (including telson). Ommatidia present, fields triangular. Medium-sized paraterga well-developed, horizontal. Male legs 3 and 4 not enlarged. Male legs 5–7 enlarged, coxae with low subconical outgrowths, femora swollen and curved, tarsi very long, sabre-shaped. Male legs 10 and 11 with coxal glands. Telopodites of anterior gonopods flagelliform, 1-segmented, sheathed. Posterior gonopod colpocoxite with an independent (separate), broad, longitudinal spiky structure (“pad”) (cp) with a shallow, partly open-edged sheath groove for the anterior gonopod telopodite (Figs 15, 16). Colpocoxites fused medially in its basal half; their distal parts curved laterally. Each colpocoxite entire, undivided. Angiocoxites with a subglobule (sg) strongly protruding caudad in posterior view (Figs 15, 18). Angiocoxites (a) in anterior view strongly elongated ventrally, centrally convex along their length, however, they do not form independent, separate processes, but are fused with to colpocoxites (Fig. 21). Anterior angiocoxal processes absent. Posterior angiocoxal processes present. Posterior gonopod telopodites 2-segmented, setose, telopoditomere 1 with a thin stem, telopoditomere 2 of medium length. Etymology. The generic name refers to the type locality, Liaoning Province. The name is a neuter noun. Distribution. China: Liaoning Province., Published as part of Mikhaljova, Elena V., 2022, The millipede family Diplomaragnidae Attems, 1907 confirmed for mainland China, with descriptions of a new genus and new species (Diplopoda, Chordeumatida), pp. 185-192 in Zootaxa 5219 (2) on page 186, DOI: 10.11646/zootaxa.5219.2.7, http://zenodo.org/record/7413471

Antić, Dragan, Spelda, Jörg (2022): Maurieseuma, a new genus to accommodate Hylebainosoma nontronense Mauriès & Kime, 1999, and the revalidation of the genus Romanosoma Mauriès, 2015 (Diplopoda: Chordeumatida: Haaseidae). Zootaxa 5219 (2): 193-200, DOI: https://doi.org/10.11646/zootaxa.5219.2.8

The family Diplomaragnidae Attems, 1907 (Chordeumatida) is confirmed for mainland China due to the discovery of the new genus and species: Liaoninosoma gen. nov. and Liaoninosoma marusiki sp. nov. The descriptions and taxonomic notes are given for the new taxa.

Petra sierwaldae Shear, Marek, Bond & Wesener, new species Figs 1–14 Types: Male holotype, two male paratypes and three female paratypes from Beauty Creek, Panhandle National Forest, 47.6018°N, - 116.6405°W, collected 4 November 2004 by W. Leonard, C. Richart and J. Baugh; male paratype from Beauty Creek Road (FSR438), 1.3 mi SE of SR97, 698' asl, 47.6015°N, - 116.6579°W collected 18 October 2009 by W. Leonard and C. Richart; Beauty Creek, 2.5 mi E of Lake Coeur d'Alene, 3202' asl, 47.6124°N, - 116.6355°W, 14 April 2004, W. Leonard, J. Baugh; male paratype from Bumblebee Creek, Panhandle National Forest, 2240' asl, 47.6325°N, - 116.2965°W, 6 November 2004, W. Leonard, C. Richart, J. Baugh. All localities are in Kootenai County, Idaho, USA. Parts of a male paratype are on SEM stub WS36-16. All specimens, including SEM stubs, deposited in Field Museum of Natural History, Chicago IL, USA (FMNH). Etymology. As for the genus, the specific epithet is a patronym in honor of Petra Sierwald. Diagnosis. As for the genus, see above. Description. Based on male holotype from Beauty Creek. Length, 3.8 mm, greatest width 0.56 mm. Twentyeight rings (collum + 26 leg-bearing rings + telson). Head (Figs 1, 2, 5) densely setose, with single ommatidium (om, Fig. 2) on each side. Lateral corners of labrum (lab, Fig. 5) rounded. Mandibular stipes (m, Fig. 5) with distal, short, acute process. Collum (col, Fig. 1) with distinct anterior rim, notched at position of lateralmost segmental seta (cn, Fig. 2), crests short, limited to posterior fourth of collum, sixth crest marginal. Subsequent rings (Figs 1, 3) with crests extending full length of metazonite, subequal in height. Metazonital setae long, with brush-like tips. Telson (tel, Fig. 4) short, lacking crests, lateral lobes suppressed but middle lobe obvious. Spinnerets directed posterioventrad. Paraprocts (pp, Fig. 4) with three setae, ornamented with elongate tubercles. Hypoproct (hp, Fig. 4) with two setae, similar tubercles. First leg pair (L1, Fig. 5) encrassate, without needle-like setae, few flattened, twisted setae on distal podomeres. Second leg pair (L2, Fig. 1) reduced, femora clavate. Leg pairs 3–7 strongly encrassate, third leg pair (L3, Fig. 6) lacking coxal flasks; fourth leg pair (L4, Fig. 6) the largest, coxae slightly excavated anteriorly; coxae of fifth leg pair (L5, Fig. 6) enlarged, with blunt anterior lobes set with acute tubercles (Fig. 7); sixth and seventh leg pair coxae unmodified. Proximal segments of all legs ornamented with typical flattened setae (Fig. 8). Gonopods (Figs 9, 10, 12, 13) with broad sternum (s, Fig. 9) bearing lobes extending laterally. Coxae (cx, Figs 9, 13) with 3–5 setae, long coxal process (cxp, Figs 10, 12, 13) parallels anterior angiocoxites (aac, 9, 10, 12, 13). Posterior angiocoxae (pac?, Fig. 12) sheaths single, large flagellocoxite (fc, Figs 12, 13). Colpocoxite (cc, Figs 10, 12, 13) as elongate, fimbriate, weakly sclerotized branch. Tips of gonopod elements insert into cavities in ninth leg pair telopodites (see Fig. 9). Ninth leg pair (Figs 11, 14) with weak, narrow sternum, coxae and telopodites appearing fused; coxal part with mesal group of acute setae, telopodite portion laterally heavily tuberculate, with setae somewhat ensiform, mesally a large, smooth lobe envelopes tips of gonopods. Tenth coxae unmodified, with gland openings. Female paratype similar to male but lacking secondary sexual modifications, 4.2 mm long., Published as part of Shear, William A., Marek, Paul E., Bond, Jason E. & Wesener, Thomas, 2022, The millipede family Striariidae Bollman, 1893. VII. Petra sierwaldae, n. gen., n. sp. a minute millipede from Idaho, USA (Diplopoda, Chordeumatida, Striariidae), pp. 287-293 in Zootaxa 5213 (3) on pages 289-290, DOI: 10.11646/zootaxa.5213.3.6, http://zenodo.org/record/7360003

Petra Shear, Marek, Bond & Wesener, new genus Type species: Petra sierwaldae Shear, Marek, Bond & Wesener, new species Diagnosis. A genus of Striariinae composed of a single species, distinct from other small striariids in having a single ommatidium on each side of the head (om, Fig. 2), a notch in the collum at the position of the outermost metazonital seta (cn, Fig. 2), absence of coxal flasks on the third legs (L3, Fig. 6), characteristically modified coxae of the fifth legs (Fig. 7) and the telopodites of the ninth legs enveloping the gonopods at rest (Fig. 9). None of these characters are found in previously described genera of striariids. Etymology. The genus name is feminine in gender and honors our respected colleague, mentor, and friend, Dr. Petra Sierwald, Curator of Arachnida and Myriapoda at the Field Museum of Natural History, Chicago, IL, USA, in recognition of her many contributions to Myriapodology, Arachnology, and to natural history collections and education. Description. As for the single species described below. Distribution. Currently known from only four localities in Kootenai County, Idaho, USA. Notes. This genus is part of a complex of genera yet to be described that includes very small species (et al. 2011; Richart & Hedin 2013; Shelley 1990)., Published as part of Shear, William A., Marek, Paul E., Bond, Jason E. & Wesener, Thomas, 2022, The millipede family Striariidae Bollman, 1893. VII. Petra sierwaldae, n. gen., n. sp. a minute millipede from Idaho, USA (Diplopoda, Chordeumatida, Striariidae), pp. 287-293 in Zootaxa 5213 (3) on pages 288-289, DOI: 10.11646/zootaxa.5213.3.6, http://zenodo.org/record/7360003, {"references":["Richart, C. H. & Hedin, M. (2013) Three new species in the harvestmen genus Acuclavella (Opiliones, Dyspnoi, Ischyropsalidoidea), including description of male Acuclavella quattuor Shear 1986. ZooKeys, 311, 19 - 68. https: // doi. org / 10.3897 / zookeys. 311.2920","Shelley, R. M. (1990) A new milliped of the genus Metaxycheir from the Pacific coast of Canada (Polydesmida: Xystodesmidae), with remarks on the tribe Chonaphini and the western Canadian and Alaskan diplopod fauna. Canadian Journal of Zoology, 68 (11), 2310 - 2322. https: // doi. org / 10.1139 / z 90 - 323"]}

Subfamily Trisariinae Shear, 2020 Note: The diagnosis of this subfamily, it now appears, was too narrowly based on the type genus, Trisaria Shear, 2020. However, we do believe that the genera grouped in it represent a monophyletic taxon. Presently we rely on gonopod and ninth leg characters for a diagnosis (see key above) but when more genera and species of Striariidae available have been studied and described, we expect to compose a more detailed diagnosis. The new genus Ralampia is particularly difficult to place; the two species may or may not have a flagellocoxite and though there is a branch of the gonopod that resembles one, it is not sheathed. Ralampia filamentosa n. gen., n. sp., has a lateral filamentous branch that appears similar to those seen in Trisaria. The ninth leg telopodites appear to be free. For the time being, however, we include this genus in Striariinae., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 503, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605, {"references":["Shear, W. A. (2020) The millipede family Striariidae Bollman, 1893. I. Introduction to the family, synonymy of Vaferaria Causey with Amplaria Chamberlin, the new subfamily Trisariinae, the new genus Trisaria, and three new species (Diplopoda, Chordeumatida, Striarioidea). Zootaxa, 4758 (2), 275 - 295. https: // doi. org / 10.11646 / zootaxa. 4758.2.4"]}

Maraplia chico, Shear & Marek, n. sp. Figs 12–15, 58–60 Types: Male holotype from West Branch of Feather River, Butte Co., California, 39.8290°N, 121.7734°W, 1600’ asl, sift in goak and grape litter(sample#81-291),collected 13April1981 byD. S.Chandler; Male paratype (SEMstubWS35-7) from same locality, but collected sifting laurel litter; two male paratypes from 4 mi SE of Chico, Butte Co., California, sifting elderberry and oak litter (sample 79-12), 39.6877°N, - 121.7734°W, 200’ asl, collected 24 February 1979 by D. S. Chandler. All specimens deposited in FMNH. Etymology: The species epithet is a noun in apposition, after one of the paratype localities near the town of Chico, California. Diagnosis: Differs from M. napa n. sp., in having males with 30 rings and from M. schusteri, n. sp., in the much shorter, sharply deflexed anterior angiocoxites of the gonopods (Figs 58, 59, aac). Description: Male paratype. Length, about 3.5 mm, width 0.35 mm. Thirty postcephalic rings including telson. Two black ommatidia on each side of head. Color white. Telson lobes obscure (Fig. 15, tel). Other characters as described for the genus. Third coxae flasks (Fig. 13, cf) short, reaching only to fourth coxae when reflexed posteriorly. Gonopods (Figs 14, 58, 59) small, compact. In lateral view, coxa (Fig. 58, cx) with about 6 or 7 setae. Coxal process broad, with deep, semicircular apical notch separating a curved, narrow anterior part from more broad, less curved posterior part. Anterior angiocoxite sharply deflexed (Figs 14, 58, 59, aac), with two branches. Posterior angiocoxite in lateral view (Fig. 58, pac) with two branches, one of which sheaths a thick, blunt, short, gently curved flagellocoxite (Fig 58, fc). Colpocoxite (Figs 58, 59, cc) shifted anteriomesally, with anterior branch densely fimbriate, posterior part swollen, with a few small, fine triangular fimbriae. Ninth legs (Fig. 60) with lateral coxosternal processes (Fig. 60, cp) broad at bases, with two branches, mesal branch hamate, lateral branch slightly curved, acute; sternal part with median knob, telopodite fungiform (Figs 14, 60, t 9), bearing mesal process set with 6 or 7 ensiform setae (these setae appear socketed in SEM view. Tenth leg coxae slightly enlarged, with glandular openings. Females not collected. Distribution: Presently known only from Butte County, California.

Maraplia schusteri Shear & Marek, n. sp. Figs 10, 11, 55–57 Types: Male holotype and two male paratypes from near Stringtown Hill, NE of Oroville, Butte Co., California, 39.5219°N, - 121.3909°W, 1440’ asl, collected 9 February 1956 by R. O. Schuster. Types deposited in FSCA. Parts of holotype and one paratype on SEM stub WS35-4. Etymology: The species is named for entomologist Robert O. Schuster (1927–1989). Diagnosis: Distinct from the two other species of Maraplia n. gen. in the form of the gonopods. Maraplia schusteri n. sp., has long, thin, and evenly curved angiocoxites, rather than the sharply deflexed, relatively short anterior angiocoxites seen in the other species (Figs 10, 55, 56, aac). The coxal process is not flattened and platelike but narrower and distinctly bifid (Fig. 55, cp). Description: Male holotype. Length 3.5 mm, width about 0.32 mm. Two black ommatidia on each side of head. Color white. Sixth crests about twice height of C5, as short paranota. Telson lobes obscure. Other characters as for genus. Gonopods (Figs 10, 55, 56) small, compact. In lateral view, coxa (Fig. 55, cx) with about 6 or 7 setae, coxal process narrow, distally bifid, anterior branch longer, posterior branch slightly recurved (Fig. 55, cp). Anterior angiocoxite (Figs 10, 55, 56, aac) narrow, long, evenly curved, apically acute, with smaller basal triangular tooth. Posterior angiocoxite strongly reduced to single blunt rod, apparently not sheathing the very short, straight flagellocoxite. Colpocoxite (Figs 55, 56, cc) shifted anteriomesally, with anterior branch densely fimbriate, posterior part swollen, with a few small, fine triangular fimbriae. Ninth legs (Fig. 57) with mesal coxosternal process low, triangular (Fig. 57, cp), telopodite fungiform (Figs 10, 57, t 9), bearing mesal process set with 6 or 7 ensiform setae and second curved, roughly triangular posterior process. Tenth leg coxae slightly enlarged, with glandular openings. Females not collected. Distribution: Presently known only from Butte County, California., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 509, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Rampalia Shear & Marek, new genus Type species: Rampalia cheathamensis Shear & Marek , n. sp. Etymology: The genus name is an arbitrary combination of letters, an anagram of the related genus name Amplaria forming a Latin neologism to be treated as feminine in gender. Diagnosis: A genus of the subfamily Striariinae by virtue of having gonopods with a flagellocoxite and ninth legs with free telopodites. The gonopod flagellocoxite is inconspicuous. The anterior angiocoxites are erect and fused, a character unique in the family, while the posterior angiocoxites appear to be reduced to just a sheath for the single, small flagellocoxite. The ninth legs are distinctive in the bifurcate coxosternal process and the rather small, elongate telopodites bearing a subterminal lobe. Description: See the description of the single known species, below. Included species: Only the type species. Distribution: Humboldt County, California., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 517, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Rampalia cheathamensis Shear & Marek, n. sp. Figs 33–36, 72–75 Types: Male holotype and paratype from Owen R. Cheatham Redwood Grove, along Rt. 36 and Van Duzen River, 40.4829°N, - 123.9630°W, 270’ asl, Humboldt Co., California, collected 18 December 1977 by A. K. Johnson. Parts of male paratype on SEM stub WS35-5. Deposited in FSCA. Male paratype from Along Rt. 101, 1.5 mi by road south of Scotia, 40.4626°N, - 124.0871°W, 120’ asl, Humboldt Co., California, collected 3 January 1977 by A. K. Johnson, deposited in FSCA. Etymology: The name is derived from the type locality. Diagnosis: As for the genus, see above. Description: Male holotype. 30 rings. Length, 5.0 mm, width 0.45 mm. Three black ommatidia on each side of head (Fig. 33, om). Sixth crests well extended as broad paranota. Telson lobes clearly divided. Metazonital setae not seen, absent or concealed by heavy cerotegument. Color after long preservation pale whitish tan with slight purplish brown markings on crests. First legs enlarged, with needle-like setae. Second legpair smaller, when coxae apposite, appearing to have common seminal opening. Flasks of third coxae long (Fig. 34, cf), when extended posteriorly reaching to posterior margin of sixth coxae; telopodites with broad, flattened prefemora (Fig. 34, pf3). Fourth through sixth legs encrassate, podomeres flattened, with prominent characteristic modified setae. Seventh coxae with large, flattened lobes covering bases of gonopods (Fig. 35, cx7). Other characters as described for genus. Gonopods (Figs 36, 72, 73) moderately large. Sternum massive, short (Fig. 36, s). Coxae (Figs 72, 72, cx) with 3–4 setae on mesal side, 2 setae laterally, extended into long, twisted and apically flattened processes bearing short, recurved subterminal branch (Figs 72, 73, cp). Anterior angiocoxites projecting anteriorly between coxae, fused in midline, simple (Figs 36, 72, 73, aac). Posterior angiocoxites reduced to sheath for single flagellocoxite (Fig. 72, fc) Colpocoxites (Fig. 73, cc) pushed anteriorly, with many small finger-like extensions. Ninth legs with median, apically bifurcate coxosternal process (Figs 74, 75, cxp); telopodites free, small, narrow, with subapical lobe densely setose in posterior view, fitting into locking notch in pleurotergite 7 (Fig. 36, pt7). Tenth coxae not much swollen, gland openings anteriodorsal. Females not collected. Distribution: Known only from two localities in Humboldt Co., California. Note: The gonopods of this species presented difficulties in interpretation and we cannot be entirely sure that the homologies we have suggested are correct. Alternatively, the fused median structure we have called the anterior angiocoxites could be a process from the sternum, but Fig. 36 shows them quite separate from the sternum and basically as a double structure. The coxal processes in this interpretation might be the anterior angiocoxites, but they are continuous with the coxae and the coxal setae are set more distally than we have seen in other striariids. The collection of additional material and further study with SEM should resolve the question. The unique ninth legs seem to set this species well apart from the others described here.

Lamparia pratensis Shear & Marek, n. sp. Figs 26–29, 66–68 Types: Male holotype from Grassy Flat Campground, Rt. 199, 5 mi by road E of Gasquet, Del Norte Co., California, 41.8564°N, - 123.8890°W, 700’ asl, collected 25 March 1976 by A. K. Johnson. The specimen is mounted on SEM stub WS35-10, deposited in FMNH. Etymology: The species epithet is a Latin adjective, meaning “of the meadow,” and refers to Grassy Flats, the type locality. Diagnosis: Very similar to the L. millicoma n. sp., but distinct in the form of the ninth legs, which lack the expanded coxosternum, the median coxosternal knob, and the curved coxosternal process. Instead, there is a smaller triangular coxosternal process (compare Figs 68 and 71). Description: Male holotype. Length, 5.0 mm, width 0.45 mm. Two black ommatidia on each side of head. Sixth crests well extended as broad paranota (Figs 26, 27). Telson lobes nearly completely suppressed (Fig. 29, tel). Metazonital setae not seen, absent or concealed by heavy cerotegument. Color after long preservation medium brown. Flasks of third coxae short, when extended posteriorly reaching only to anterior margin of fifth coxae. Other characters as described for genus. Gonopods (Figs 66, 67) moderately large. Coxae with 3 setae, bulbous, with acute angle distally (Fig. 66, cx). Anterior angiocoxites narrow, erect, with slightly expanded, curved tip, posterior and anterior surface finely fimbriate from base to about half length of coxite (Figs 66, 67, aac). Posterior angiocoxite much reduced; flagellocoxite not sheathed. Flagellocoxite (Figs 66, 67, fc) single, thin, short. Colpocoxite bulbous, shifted anteriomesally, set with a few triangular, fine cuticular teeth (Fig. 67, cc). Ninth legs (Fig. 68) with coxosternum. Coxosternal processes flattened, long, bluntly triangular (Fig. 68, cp), arising near midline. Telopodites (Fig. 68, cp) free, rounded, with projecting posterior lobe. Tenth coxae not much swollen, gland openings anteriodorsal. Females not collected. Distribution: Known only from the type locality., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 514, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Key to species of Maraplia based on males 1a. Gonopod coxal process broad, plate-like, not divided, flagellocoxite distinctly hooked, Anterior part of posterior angiocoxite botrydial................................................................................. M. napa, n. sp. 1b. Not as above.........................................................................................2. 2a. Gonopod anterior angiocoxite long, evenly curved............................................ M. schusteri, n. sp. 2b. Gonopod anterior angiocoxite short, sharply reflexed............................................. M. chico, n. sp., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 507, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Lamparia curryensis Shear & Marek, n. sp. Figs 16–19, 49–51 Types: Male holotype from NE of Brookings Curry Co., Oregon, north bend of Chetco River Road, 0.6 mi NW of entrance to Alfred A. Loeb State Park, Redwood Nature Trail, 135’ asl, 42.1187°N, - 124.1959°W, collected 4 January 2011 by C. Richart et al; male and female paratypes from 13 mi W, 5 mi N of Brookings, T39S, R14W, S5 (42.1239°N, - 124.0141°W), 2000’ asl, collected 12 February 1972 by E. M. Benedict. All material deposited in CAS; male holotype is on SEM stub WS35-13. Etymology: The species epithet refers to Curry County, Oregon, which includes both known collection localities for this species. Diagnosis: Obviously close to L. bentonensis, n. sp., but distinct in details of the gonopods (compare Figs 61, 62 with Figs 63, 64) and in the much longer coxosternal process of the ninth legs (Fig. 19, cp). The sixth crests of this species are nearly twice as high as those of L. bentonensis n. sp. (compare Figs 17, 18 with Figs 20, 21). Description: Male holotype. Length, 4.8 mm, width 0.48 mm. Two black ommatidia on each side of head (Fig. 16, om). Sixth crests well extended as broad paranota (Figs 17, 18). Telson lobes nearly completely suppressed. Metazonital setae not seen, absent or concealed by heavy cerotegument. Color white to pale tan, faintly suffused pale purplish brown. Flasks of third coxae long, when extended posteriorly reaching to anterior margin of seventh coxae. Other characters as described for genus. Gonopods (Figs 19, 61, 62) moderately large. Coxae with 3 setae, bulbous (Fig. 61, cx). Anterior angiocoxites narrow, erect, with flattened, bulbous tip, posterior surface finely fimbriate from base to just below bulbous tip (Figs 19, 61, 62, aac). Posterior angiocoxite complex (Fig. 62, pac) with two branches. Flagellocoxite possibly not sheathed. Two flagellocoxites, each narrow, slightly curved, comparatively short (Figs 61, 62, fc). Colpocoxite large and bulbous, shifted anteriomesally, densely set with triangular, fine cuticular teeth (Fig. 62, cc). Ninth legs with coxosternum. Coxosternal processes (Fig. 19, cp) flattened, elongate, distally slightly expanded. Telopodites (Fig. 19, tp) free, apically flattened, mesally curved anteriorly, with sawtooth edges, interlocking with gonopods, laterally fitting into shallow indentations in ventrolateral edges of seventh ring. Tenth coxae not much swollen, gland openings anteriodorsal. Female paratypes similar to males in nonsexual characters. Distribution: Presently known from two nearby localities in Curry Co., Oregon. Note: Additional collecting is required to establish the distribution of this species and the following one. They are evidently closely related and may in fact be variants of a single species., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 512, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Plaramia Shear & Marek, new genus Type species: Plaramia arcata Shear & Marek , n. sp. Etymology: The genus name is an arbitrary combination of letters, an anagram of the related genus name Amplaria forming a Latin neologism to be treated as feminine in gender. Diagnosis: The flagellocoxites are long, thin and largely concealed in this genus and the ninth leg telopodites are free. Thus we place the genus in Striariinae. Three of the striariine genera described in this paper have their metazonital crests all roughly subequal and not paranota-like. The two species of Plaramia, n. gen., differ from those of Maraplia, n. gen., in that the gonopod anterior angiocoxites are erect and sinuously curved, rather than reflexed anteriorly and either short or if long, curved in an arc. In Ralampia, n. gen., the gonopods are quite compact, and the colpocoxites have long, erect, branched, stiff fimbriae distally. Description: Small striariines, about 3–4 mm long (Fig. 46). Thirty postcephalic rings, including telson. Two ommatidia on each side of head (Fig. 45, om). Head (Fig. 37) densely setose, with small tubercles. Labral hook lacking. Mandibular stipes with serrate margin, acute triangular process. Collum with crests limited to posterior region, studded with small tubercles. Metazonital crests 1–5 subequal (Fig. 41), crest 6 slightly elevated as paranota (though variably so; Fig. 46); metazonital setae long, with brush-like tips. Telson with lobes very shallowly demarcated or nearly indiscernable, sulci shallow. Legpair 1 (Fig. 37, L 1) larger than legpairs 2 or 3, femora and tibiae with long, needle-like setae, tarsus with comb. Legpair 2 with short telopodites, vas deferens openings in coxae facing mesally, coxae notched so that when appressed, a single pore is formed. Legpair 3 with relatively short flasks (Figs 38, 48, cf), reaching fifth coxae when extended posteriorly, heavily set with raised cuticular scales, few contorted setae. Legpairs 4–7 crassate, legpair 7 lacking coxal lobes. Gonopod sternum large, broader than long (Fig 43, s). Coxa with 4–6 setae, coxal process absent. Anterior angiocoxites long, not deflexed, sinuously curved or with hooked termination (Figs 42, 43, 72, 73). Posterior angiocoxites small, inconsipicuous, forming short sheath for flagellocoxites. Flagellocoxites single, long, thin, concealed by angiocoxites. Colpocoxites large, sac-like, seemingly well sclerotized, with fine longitudinal ridges but lacking fimbriae. Ninth legs (Figs 44, 78) with coxosternite and free telopodites. Coxosternite with two processes from coxal part, sternal part sometimes expanded, with central knob. Telopodite large, variable, with pebbled sculpture and setae, with deep notch to accommodate gonopods. Telopodite locks into deep notch in ventrolateral margin of pleurotergite 7. Tenth legs with coxal pores, coxae slightly enlarged. Distribution: Humboldt Co., California., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on pages 519-520, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Plaramia arcata Shear & Marek, n. sp. Figs 37–40, 76–78 Types: Male holotype, 2 male paratypes and female paratype from the north side of Jolly Giant Canyon, Arcata Community Forest, 40.8759°N, - 124.0681°W, 400–600’ asl, collected 21 February 1978 by A. K. Johnson; two male paratypes from Arcata Community Forest, 40.8699°N, - 124.0725°W, collected 31 March 2011 by C. Richart et al .; two male paratypes (parts of these are on SEM stub WS35-3) from along Jacoby Road and Kilpatrick Quarry Road, 100–200’ asl, T5N, R1E, Sec 14 (40.8279°N, - 124.0390°W) collected 24 November 1977 by A. K. Johnson; six male and two female paratypes from behind Humboldt State University campus, Arcata Community Forest, 40.8748°N, - 124.0506°W, 200–700’ asl, collected 1 April 1979 by A. K. Johnson. All material deposited in FSCA. Etymology: The name is derived from the type locality, Arcata Community Forest, and is a noun in apposition. Diagnosis: The two species of this genus are closely similar, but Plaramia arcata, n. sp., can be distinguished from P. johnsonae, n. sp., by details of the gonopods and ninth legs. The gonopods of the latter species have more curved anterior angiocoxites than the former, with a hooked termination, and the ninth leg telopodites are deeply notched. C6 of this species are very prominent, they are less so in P. johnsonae, but this difference can only really be seen in comparison. Description: Male holotype. Length 3.8 mm, width 0.4 mm; 30 rings. Two black ommatidia on each side of head (Fig. 37). C6 moderately extended as paranota. Telson lobes shallowly divided, nearly obscure. Metazonital setae obvious only on anterior rings, relatively short, acute. Color after long preservation pale yellowish tan. First legs enlarged (Fig. 37, L 1), with needle-like setae. Second legpair smaller, when coxae apposite, appearing to have common seminal opening. Flasks of third coxae short (Fig. 38, cf), when extended posteriorly reaching to anterior margin of fifth coxae; telopodites with broad, flattened prefemora. Fourth through sixth legs encrassate, podomeres flattened, with prominent characteristic modified setae. Seventh coxae lacking lobes. Gonopods Figs 39, 40, 76, 77) moderately large. Sternum massive, short. Coxae with 3 or 4 setae on lateral side, extended into short, triangular process (Fig. 76, cx). Anterior angiocoxites curved laterally, then mesally, without branches, slightly flattened at tip (Figs 40, 76, 77, aac). Posterior angiocoxites small, low, with two processes; reduced to sheath for single flagellocoxite (Fig. 77, fc). Single flagellocoxite long, thin, evenly curved, possibly movable (there may be a basal articulation). Ninth legs (Fig. 78) with broad, plate-like coxosternal process (Fig. 77, cxp); telopodites (Fig. 39, 77, t 9) free, large, flattened, notched on mesal surface to embrace gonopod anterior angiocoxites, laterally fitting tightly into notch in seventh pleurotergite (Fig. 39). Tenth coxae not much swollen, gland openings anteriodorsal. Females similar to males in nonsexual characters. Distribution: Known only from Arcata Community Forest, Humboldt Co. California., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 520, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Shear, William A., Marek, Paul E. (2022): The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea). Zootaxa 5205 (6): 501-531, DOI: https://doi.org/10.11646/zootaxa.5205.6.1

Nototrisaria ornata Shear & Marek, n. sp. Figs 1–6, 49–51 Types: Male holotype and male paratype from 9.6 miles up Elochoman Valley Road from Rt. 4, Wahakiakum Co., Washington, 46.289633°N, - 123.26485°W, 450’ asl, collected 28 March 2004 by C. Richart, W. Leonard; male paratype from 11.7 mi up Elochoman Valley Road from Rt. 4, 450’ asl, 46.316783°N, - 123.2617°W, 28 March 2004, collected by C. Richart; male and female paratypes from Trap Creek, 1.1 mi S of SR6, Pacific Co. Washington, 46.5468°N, - 123.6161°W, 180’ asl, collected 11 January 2003 and 3 May 2003 by W. Leonard. Parts of a male paratype are mounted on SEM stub WS3-8. Types deposited in the CAS. Etymology: The species name, a Latin adjective, refers to the ornate dorsum. Diagnosis: As for the genus; see above. Description: Male paratype from Elochoman River Road. Length, 5.5 mm, width 0.6 mm (includes paranota). Thirty postcephalic rings including telson. Head (Fig. 2) densely setose, with sculpture of tiny, acute tubercles; labral corners rounded, without projecting hook. Ommatidia absent. Antennae short, robust. Mandibular stipes (Fig. 2, m) with sawtooth distal edge, acute apical projection. Color of live specimen white to pale yellow (Fig. 1). Collum (Fig. 2, col) broadly expanded, with 10 subequal crests limited to posterior two-thirds; anterior third with pebbled sculpture. Anterior rings (Fig. 3) with C2 and C6 exaggerated, the latter expanded into prominent paranota with deep posterior notch; C1 reduced to a few knobs, absent on more posterior rings, C2 also progressively lower on midbody ring, exaggerated again on most posterior 6–8 rings (Fig. 4). Metazonital setae not observed, probably absent. Limbus serrate. Telson without crests, lobes deeply incised (Fig. 4). First legpair crassate (Fig. 2, L 1), larger than legpairs 2 and 3 (Fig. 2, L2, L3), postfemora and tibia set with long, stiff setae, tarsus with comb. Legpair 2 relatively small, similar to postgonopodal legs. Legpair 3 with prominent, broad, slightly sigmoidal coxal flasks bearing curled setae on anterior surface (Fig. 2, cf); prefemora expanded, flattened. Legpairs 4–6 strongly crassate. Legpair 7 similar to pairs 4–6, but with large, flattened coxal plates covering bases of gonopods. Gonopods in mesal view (Figs 5, 50) with anterior angiocoxites erect, narrow, hooked and expanded at tip; posterior edge with continuous fimbriate lamina, lamina ends in long distal projection; tuft of strong, thin cuticular projections subapically (aac, Figs 5, 49, 50). Posterior angiocoxites as broad, rounded plate (pac, Figs 5, 50). Flagellocoxites absent. Colpocoxite (cc, Figs 5, 49, 50) developed as compact mass of finely divided, sinuous fimbriae with small posterior portion unmodified, sclerotized. Ninth legpair (Figs 6, 51) strongly reduced, sternum (s9), coxae (cx) and telopodites (t9) fused; coxal part with blunt triangular process (cp), telopodite part with long, finger-like basal process bearing acute setae. Tenth leg with enlarged, glandular coxae (Fig 6, cx10). Female paratype similar to male in nonsexual characters, but exaggerated crests significantly lower. A living female specimen is depicted in Fig. 1. Distribution: As for the genus. The Grays Harbor County record (WASHINGTON: Grays Harbor Co.: Canyon River, 47.303517°N, - 121.508167°W, 1500’ asl, 12 December 2004, W. P. Leonard) is based on a female that strongly resembles the females from other localities., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on pages 504-506, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Lamparia bentonensis Shear & Marek, n. sp. Figs 20–25, 63–65 Types: Male holotype and male and female paratypes from Clemens Park, Seely Creek Road, 0.3 mi from SR34, north fork of Alsea River, Benton Co., Oregon, 400’ asl, 44.4092°N, - 123.5678°W, collected 4 December 2005 by W. Leonard and C. Richart. Male and female paratypes from localities along Wells Creek Road, Benton Co., Oregon as follows: at SR34, 44.4707°N, - 123.4797°W, collected 29 November 2004 by W. Leonard; 0.6 mi W of SR34, 44.4722°N, - 123.4794°W, collected 25 February 2005 by W. Leonard, C. Richart; 0.7 mi from SR34, 6.3 mi from SR20, 44.4686°N, - 123.4941°W, 1100’ asl, collected 9 December 2005 by W. Leonard, C. Richart. Etymology: The species epithet refers to Benton County, Oregon, which includes all known collection localities for this species. Diagnosis: Very similar to L. curryensis, n. sp., but distinct in details of the gonopods (compare Figs 63, 64 with Figs 61, 62) and in the shorter coxosternal processes of the ninth legs (Fig. 64, cp). The sixth crests of this species (Fig. 21) are only about half the height of those of L. curryensis n. sp.. Description: Male paratype from Clemens Park. Length, 5.1 mm, width 0.6 mm. Two ommatidia on each side of head. Sixth crests well extended as small paranota (Figs 20, 21). Telson lobes nearly completely suppressed (Fig 22, tel). Color white, suffused light purplish brown. Second coxae with common opening for vas deferens (Fig. 23). Flasks of third coxae long, when extended posteriorly reaching to anterior margin of seventh coxae (Fig. 25, cf). Other characters as described for genus. Gonopods (Figs 24, 63, 64) moderately large. Coxae with 5–6 setae, elongate.Anterior angiocoxites (Figs 24, 63, 63, aac) narrow, erect, with flattened, bulbous tip, posterior surface finely fimbriate from base to just below bulbous tip. Posterior angiocoxite complex (Figs 63, 64, pac) with three branches, one of which sheaths flagellocoxite. Flagellocoxite (possibly 2 of these) narrow, slightly curved, comparatively short (Fig. 63, fc). Colpocoxite (Fig. 64, cc) small, densely set with triangular, fine cuticular teeth. Ninth legs (Fig. 65) with coxosternum. Coxosternal processes (Figs 24, 65, cp) flattened, drawn out laterally to points.Telopodites (Figs 24, 65, t 9) free, apically flattened, mesally curved anteriorly, with sawtooth edges, interlocking with gonopods, laterally fitting into deep indentations in ventrolateral edges of seventh ring (Fig. 24). Tenth coxae not much swollen, gland openings anteriodorsal. Female paratypes similar to males in nonsexual characters. Distribution: Presently limited to Benton Co., Oregon. Note: Initially we thought this species and the preceding were the same, but careful examination of the gonopods at 400X under the compound microscope and a study of SEM photographs of the ninth legs showed that they are clearly distinct., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on pages 513-514, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605

Key to species of Plaramia, based on males 1a. Gonopod anterior angiocoxite slightly curved; 9th leg telopodite not excavate......................... P. arcata, n. sp. 1b. Gonopod anterior angiocoxite strongly sigmoidially curved, tip as a definite hook, 9 th leg telopodite strongly excavate with a deep notch mesally..................................................................... P. johnsonae, n. sp., Published as part of Shear, William A. & Marek, Paul E., 2022, The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), pp. 501-531 in Zootaxa 5205 (6) on page 520, DOI: 10.11646/zootaxa.5205.6.1, http://zenodo.org/record/7318605