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Hairy galinsoga (Galinsoga quadriradiata Cav.) is a troublesome weed in lettuce (Lactuca sativa L.) and other low-growing crops. Many troublesome weed species are more responsive to fertilizer than major crops, and hence potentially more competitive under fertile conditions. This study examined the responses of G. quadriradiata and lettuce, in monoculture or competition with each other, to nitrogen and phosphorus. A greenhouse experiment assessed the biomass and flower production of G. quadriradiata grown in 16 nutrient treatments. The nutrient treatments were arranged in a phosphorus series, in which phosphorus varied, while nitrogen was either low or high, and a nitrogen series, in which nitrogen varied, while phosphorus was either low or high. A field experiment examined G. quadriradiata biomass, flower production, and competition with lettuce using the same nutrient treatments as the greenhouse experiment. Galinsoga quadriradiata dry biomass increased with phosphorus addition, especially when nitrogen was abundant. In the phosphorus series at high nitrogen, G. quadriradiata produced more biomass when competing with lettuce than in monoculture. Nutrient addition generally increased flower production per unit mass. Lettuce fresh biomass increased with phosphorus, but lettuce dry biomass was largely unaffected by fertility. Lettuce fresh and dry biomass were sometimes decreased by competition from G. quadriradiata, relative to intraspecific competition. We conclude that high phosphorus rates increase the productivity of G. quadriradiata and may increase its competitiveness. Therefore, high phosphorus rates should be avoided unless good weed control can be assured.

PREMISE Understanding how drought and biomass allocation patterns influence competitive ability can help identify traits related to invasiveness and guide management. Vincetoxicum nigrum and V. rossicum are increasingly problematic herbaceous perennial vines in the northeastern United States and southeastern Canada. METHODS Using a greenhouse experiment, we investigated how biomass allocation and competition intensity of Vincetoxicum spp. responded to four competitive regimes at two levels of soil water availability in the presence of conspecific or congeneric neighbors. RESULTS Soil moisture was the most important influence on growth and biomass allocation. Vincetoxicum nigrum had a greater capacity for growth and reproduction than V. rossicum, especially under drought. Drought reduced the probability of reproduction for V. rossicum. Vincetoxicum rossicum had a higher root-to-shoot ratio than V. nigrum under adequate soil moisture. This difference more than doubled under drought. Under interspecific competition, V. nigrum maximized its biomass, while V. rossicum limited aboveground growth and reproduction. Root-only competition increased shoot and root biomass relative to shoot-only competition. The effects of root and shoot competition were additive under interspecific competition, but interacted under intraspecific competition (negative interaction under drought and positive interaction under sufficient soil moisture). CONCLUSIONS Management strategies targeting mixed populations of V. rossicum and V. nigrum are most important under ample water availability. Under drought conditions, strategies focused on V. nigrum should effectively limit Vincetoxicum growth and seed reproduction. Phenotypic plasticity and the positive competition intensity associated with drought in monocultures may contribute to drought resistance in these invasive species.

Macronutrient inputs to annual cropping systems can benefit weeds as well as crops, sometimes decreasing or eliminating the benefits of fertilization. This interaction between fertility management and integrated weed management is becoming increasingly important as these fields increase their focus on efficiency and prevention, respectively. The risk of increased weed competition reflects the fact that weed biomass and height may be highly responsive to nitrogen, phosphorus, and/or potassium. This generalization is supported by monoculture studies of species such as redroot pigweed (Amaranthus retroflexus L.), common lambsquarters (Chenopodium album L.), and barnyardgrass [Echinochloa crus-galli (L.) P. Beauv.] and by ecological theory. However, field studies indicate variation in the effects of macronutrients on weed–crop competition and crop yield, even within species groups. To address challenges in interpreting, comparing, and extrapolating from these diverse reports, we propose a conceptual framework that summarizes the mechanisms underlying observed variation within and between studies. This framework highlights functional traits and trends that help predict yield outcomes in binary weed–crop interactions. Important factors include timing of emergence, maximum heights of the weed and crop, and relative responsiveness to the added nutrient. We also survey recent work on the effects of nutrient source (e.g., the composition of organic amendments) on weed–crop competition. Because different sources vary in their nutrient release dynamics and supplied nutrient ratios, they may have dramatically different effects on weed–crop competition and crop yield. Finally, we offer a guide to best practices for studies of fertility effects on weed–crop competition. Although this review highlights several topics requiring further research, including fertility effects on multispecies interactions and interactions with other environmental factors, emerging methods offer considerable promise. Ultimately, an improved understanding of nutrient effects on weed–crop competition will contribute to the efficient and effective management of diverse cropping systems.

Weed management is a major challenge in organic crop production, and organic farms generally harbor larger weed populations and more diverse communities compared with conventional farms. However, little research has been conducted on the effects of different organic management practices on weed communities and crop yields. In 2014 and 2015, we measured weed community structure and soybean [Glycine max (L.) Merr.] yield in a long-term experiment that compared four organic cropping systems that differed in nutrient inputs, tillage, and weed management intensity: (1) high fertility (HF), (2) low fertility (LF), (3) enhanced weed management (EWM), and (4) reduced tillage (RT). In addition, we created weed-free subplots within each system to assess the impact of weeds on soybean yield. Weed density was greater in the LF and RT systems compared with the EWM system, but weed biomass did not differ among systems. Weed species richness was greater in the RT system compared with the EWM system, and weed community composition differed between RT and other systems. Our results show that differences in weed community structure were primarily related to differences in tillage intensity, rather than nutrient inputs. Soybean yield was lower in the EWM system compared with the HF and RT systems. When averaged across all four cropping systems and both years, soybean yield in weed-free subplots was 10% greater than soybean yield in the ambient weed subplots that received standard management practices for the systems in which they were located. Although weed competition limited soybean yield across all systems, the EWM system, which had the lowest weed density, also had the lowest soybean yield. Future research should aim to overcome such trade-offs between weed control and yield potential, while conserving weed species richness and the ecosystem services associated with increased weed diversity.

Incorporation of cover crop residue into the soil has been suggested as a means for reducing weed seedbanks. To explore this hypothesis, we buried mesh bags of seeds mixed with sand at 15-cm depth in late fall in plots that had been planted with rye (Secale cerealeL.) or hairy vetch (Vicia villosaRoth.) or left unplanted. Separate bags contained either velvetleaf (Abutilon theophrastiMedik.), giant foxtail (Setaria faberiHerrm.), Powell amaranth (Amaranthus powelliiS. Watson), or common lambsquarters (Chenopodium albumL.). The experiment used a randomized complete block design with five replications, and enough bags were buried to allow a final recovery in each of the following three springs. Each spring, bags were exhumed, and seeds were either counted and tested for viability or mixed with chopped cover crop material or simply stirred for control bags, and the material was reburied. The experiment was completed twice with initial burials in fall of 2011 and 2013. Rye had no consistent effect on persistence of seeds of any of the species. For two observation intervals, rye increased persistence of a species; for another two intervals, it decreased persistence relative to the control; but mostly rye did not affect persistence. Hairy vetch decreased persistence ofC. albumandA. powelliiin both runs of the experiment but had no effect on persistence ofA. theophrastiandS. faberi. Germination of the first two species is promoted by nitrate, whereasA. theophrastigermination is not sensitive to nitrate, andS. faberiis only rarely nitrate sensitive. We suggest that nitrate released during decomposition of hairy vetch may have promoted fatal germination ofC. albumandA. powellii. Incorporation of legume cover crops like hairy vetch may provide a means for decreasing the seedbanks of the many weed species whose germination is promoted by nitrate. The lack of any reduction ofA. theophrastiandS. faberiseed persistence in response to hairy vetch and the inconsistent and mostly negligible effect of rye indicate that a general increase in readily decomposable organic matter through incorporation of cover crops may be ineffective at reducing weed seedbanks.

Pale swallowwort and black swallowwort are European viny milkweeds that have become invasive in many habitats in the northeastern United States and southeastern Canada. A multiyear seedbank study was initiated in fall 2011 to assess annual emergence of seedlings and longevity of seeds of pale swallowwort and black swallowwort at four different burial depths (0, 1, 5, and 10 cm) over 4 yr. One hundred swallowwort seeds were sown in seed pans buried in individual pots, and emerged seedlings were counted and removed from May through September each year. A subset of seed pans was retrieved annually in October, and recovered seeds were counted and tested for viability. The majority of seedling emergence occurred during the first year (92% in 2012), and no new seedlings emerged in the third (2014) or fourth (2015) years. Pale swallowwort had relatively poor emergence at sowing depths of 0 cm (11%), 5 cm (6%), and 10 cm (0.05%—only one seedling), while 37% of pale swallowwort seeds emerged at 1 cm. The larger-seeded black swallowwort was more successful, with two-thirds of all sown seeds emerging at depths of 1 cm (71%) and 5 cm (66%), and 26% emerging at 10 cm. Only 16% of the surface-sown black swallowwort emerged. A large portion of the seeds that germinated at 10 cm, as well as at 5 cm for pale swallowwort, died before reaching the soil surface. Of filled seeds that were recovered in 2012 (black swallowwort at the 0-cm depth), 66% were viable. No viable seeds were recovered after the second growing season. Seeds recovered following the third year had become too deteriorated to accurately assess. Swallowwort seeds do not appear to survive more than 2 yr in the soil, at least in our experiment, suggesting that the elimination of seed production over 3 yr will exhaust the local seedbank. Seeds would need to be buried at least 10 cm for pale swallowwort but more than 10 cm for black swallowwort to prevent seedling emergence. Burial of swallowwort seeds as a management strategy may, however, only be practical in natural areas where high swallowwort densities occur.Nomenclature: Black swallowwort, Vincetoxicum nigrum (L.) Moench [Cynanchum louiseae Kartesz & Gandhi]; pale (or European) swallowwort, V. rossicum (Kleopow) Barbar. [Cynanchum rossicum (Kleopow) Borhidi].

To assess the effectiveness of interrow cultivation, counts were taken before and after cultivation of corn and soybean during the first two crop rotations in a corn–soybean–spelt organic grain cropping systems experiment. Overall control per cultivation event in soybean was 73%, about equal to the 67% of the interrow area actually covered by cultivator tools. Weed control per cultivation event in corn was higher, and exceeded 91% at later cultivations. The greater weed control in corn relative to soybean, particularly at later cultivations, was probably due to more soil being thrown into the corn row, burying a greater proportion of the weeds. Perennial weeds emerging from roots and rhizomes were less controlled by cultivation events than weeds emerging from seeds. Relatively poor control of perennials was due both to rapid resprouting during the few days between cultivation and assessment and to a lower probability of death in the zone indirectly disturbed by cultivator tools. Seedlings of perennial species suffered greater mortality from cultivation than annual weeds, probably because the low relative growth rate of perennials resulted in small seedlings that were susceptible to cultivation. Common ragweed was less controlled by cultivation than other annual weeds, probably because its heavier seeds produced larger seedlings at the time of cultivation. These larger seedlings were less likely to be buried during hilling-up operations at later cultivations. Counts of weeds before and after individual cultivation events provide insight into the processes affecting weed mortality during mechanical management.

Recovery of common agricultural weeds after burial by soil was studied in four greenhouse and three field experiments. Species studied included velvetleaf, Powell amaranth, common lambsquarters, barnyardgrass, and giant foxtail. Seedlings were bent over before burial to simulate the effect of the impact of soil thrown by a cultivator. Altogether, more than 35,000 seedlings were marked and observed for recovery. No seedlings recovered from 4 cm of burial. Recovery from complete burial under 2 cm of soil ranged from 0 to 24% depending on the experiment, species, and watering treatment, but recovery greater than 5% was rare. Large-seeded species tended to recover from complete burial under 2 cm of soil better than small-seeded species. The study did not reveal a difference in recovery of grasses relative to broadleaf weeds. Overall, seedlings tended to recover best when water was applied daily after burial, worst when water was applied once on the day of burial, and to an intermediate extent when no water was applied. However, difference in recovery between the no-water and watering-once treatments were usually small. Also, many experiment by species combinations showed no significant differences among watering treatments. When even a small portion of the seedling was left exposed, recovery generally exceeded 50%. Organic weed management systems commonly use burial of weed seedlings with tine weeders and soil thrown by sweeps and hilling disks to control weeds in crop rows. Recovery from burial could pose a substantial weed management problem in some circumstances, particularly for large-seeded weed species. Maximizing burial depth is important for limiting recovery. Recovery from burial can be minimized by withholding irrigation for several days after hilling-up operations.

The distinction between count nouns and mass nouns affects thinking and writing about various types of crops and produce. Count nouns are words that indicate discrete, countable objects (e.g., forks, viewpoints), whereas mass nouns are words that indicate some relatively undifferentiated substance (e.g., water, energy). We explain the grammar of these two forms and point out some writing pitfalls to avoid. The word seed is one of the few English nouns that is both a count noun and a mass noun. An argument is presented for using seeds as the plural when several individuals are counted and for using seed as the singular when referring to seeds in the aggregate.

Sufficient fertility is important for crop growth and yield but supplying a balanced amount of N, P, and K with compost and manure is a challenge and nutrient imbalances can benefit weeds more than crops. The goal of this study was to compare the aboveground growth responses of common northeastern U.S. crops and weeds to addition of composted poultry manure (CPM). A secondary goal was to test whether the observed growth responses to CPM could be attributed to the three macronutrients—N, P, and K—supplied in the CPM. Two field experiments were conducted over 2 yr. Species grown were corn, lettuce, kale, Powell amaranth, common lambsquarters, giant foxtail, and velvetleaf. Plants were grown in soil amended with CPM or single-nutrient organic N, P, and K fertility amendments. Single-nutrient P treatments with bone char did not adequately mimic P supply from CPM. In both years, biomass of all weeds studied increased with CPM amendment rate. Powell amaranth was the most responsive to CPM addition, increasing 1,775 and 159% from the control to the highest CPM rate in 2010 and 2011, respectively. Corn biomass increased by 22% with CPM rate in 2010 but did not increase with CPM rate in 2011. Lettuce biomass increased with CPM amendment rate (175% in 2010 and 109% in 2011), but due to the increased weed biomass at high CPM amendment rates, good weed control will be necessary to maintain this yield benefit. The increase in growth of Powell amaranth, common lambsquarters, and giant foxtail with CPM amendment was not due to N or K, and may have been influenced by P or another factor in the CPM. Velvetleaf was the only species that exhibited increased biomass with N addition (as blood meal), similarly to the increase with added CPM, suggesting the velvetleaf growth response to CPM was due to N in the CPM. The results show that nutrient ratios should be considered when applying organic amendments, because application rates of organic amendments that meet the crop's needs for one nutrient may oversupply other nutrients. Overfertilization will not benefit crop yield, but the results of this study show that high organic fertility application rates are likely to increase weed growth.

Most previous research on changes in weed abundance and community composition in cropping systems has focused on field crops. The study presented here examined changes in the weed seedbank and aboveground biomass in four organic vegetable cropping systems over a 10-yr period. The systems included an Intensive system with six crops per 4-yr rotation, an Intermediate system with one cash crop per year, a Bio-extensive system with alternating cash crop and tilled fallow years plus prevention of seed rain, and a Ridge-tillage system with one cash crop per year. Systems also differed in the types and number of cover crops between cash crops. During the course of the experiment, the weed community shifted from one dominated by summer annual broadleaf species that reproduce at the end of their lives to a community dominated by summer and winter annuals that mature rapidly. This shift in community composition can be attributed to the change in land use from conventionally managed corn (Zea maysL.) and alfalfa (Medicago sativaL.) to organic vegetable production. In particular, crop rotations with diverse preplantings and postharvest tillage dates interrupted the life cycle of common lambsquarters (Chenopodium albumL.) and pigweed species (Amaranthusspp.: mostly Powell amaranth [Amaranthus powelliiS. Watson], with small numbers of redroot pigweed [Amaranthus retroflexusL.] and smooth pigweed [Amaranthus hybridusL.]), while favoring a diverse assemblage of quickly maturing species. The study thus demonstrates that an appropriate crop rotation can control the seedbank of weeds likeC. albumthat potentially persist well in the soil. The Ridge-tillage system greatly reduced the frequency and depth of tillage relative to other systems while effectively suppressing perennial weeds. The early-reproducing annuals, however, became more abundant in the Ridge-tillage system than in the other systems, primarily due to escapes along the edge of the scraped ridges. The tilled fallow periods coupled with prevention of seed rain in the Bio-extensive system substantially reduced weed abundance through time and relative to the other systems.

Elucidating relationships between the soil food web, soil processes, and agroecosystem function is a critical step toward a more sustainable agriculture. Soil and crop management practices can alter these relationships, and their effects can persist even after imposing new management practices. In 2005, the Cornell Organic Grain Cropping Systems Experiment was established in central New York. Four cropping systems that varied in fertilizer inputs, tillage practices, and weed control were compared: High Fertility, Low Fertility, Enhanced Weed Management, Reduced Tillage. Two crop rotation entry points were included in the experiment. In June 2017, the entire experimental site (plots and alleyways) was plowed and seeded with sorghum sudangrass [Sorghum bicolor (L.) Moench x S. sudanense (Piper) Stapf] as part of a uniformity trial to assess legacy effects of past management practices. Prior to initiating the uniformity trial, soil samples were collected and analyzed for soil health indicators. Soil samples were also collected to assess soil invertebrate abundance and community structure 34 and 70 days after planting. Sorghum sudangrass and weed biomass were sampled at the end of the uniformity trial in September 2017. Legacy effects of past management that were observed during the uniformity trial were associated with differences in nutrient inputs and soil disturbance, as well as the preceding crop. The High Fertility system had greater soil phosphorus than the Low Fertility system, and in one of the two crop rotation entry points, soil aggregate stability and soil respiration were greater in the Reduced Tillage system compared to the Enhanced Weed Management system. The Enhanced Weed Management cropping system also had a soil invertebrate community characterized by more disturbance tolerant taxa. Weed biomass varied by crop rotation entry point, but not cropping system. Sorghum sudangrass biomass was greater in the Reduced Tillage system than the Low Fertility system, and the entry point that had greater weed biomass also had greater sorghum sudangrass biomass. Piecewise structural equation modelling (SEM) was used to test relationships between response variables and showed that soil phosphorus, soil aggregate stability, and soil respiration explained variation in abundance of some invertebrates, and that aggregate stability, soil respiration, soil moisture, weed biomass, and a select group of invertebrates affected sorghum sudangrass biomass production. Overall our findings show that soil invertebrates can mediate the relationship between soil health indicators and crop productivity, and provide support for including direct measurements of soil invertebrates in soil health assessments.

Indirect effects of insect control strategies on weed populations are important to consider when developing robust integrated pest management strategies. Weed seed predation rates were investigated in corn managed under three contrasting treatments based on control practices for corn rootworm: (1) the transgenic crop Cry3Bb Bt corn, (2) the broad-spectrum insecticide tefluthrin, and (3) no insecticide control. This 2-yr field study conducted near Ithaca, NY, involved quantifying seed loss from velvetleaf, common lambsquarters, and giant foxtail in arenas with and without vertebrate exclosures. Velvetleaf and giant foxtail were unaffected by the insecticide treatment; however, average seed predation of common lambsquarters was lower in both the Bt corn (11.9%) and insecticide-treated plots (11.8%) compared with control plots (17.5%) that did not receive any insecticide. Seed predation of common lambsquarters was not affected by the vertebrate exclosure. Lower seed predation in the transgenic Bt corn and insecticide treatments was likely due to nontarget effects on carabids (Coleoptera: Carabidae). Although the reduction in seed predation was modest and limited to only one of the three weed species tested, our results highlight the need for greater risk assessment that includes the ecosystem service of weed seed predation when considering insect pest management options.

Weed management is a major concern for organic farmers. Strategies and techniques used by organic farmers vary according to crops, soils, climate and other factors. Organic farmers in upstate New York experienced in weed management were interviewed about what weed–crop complexes they face, how they manage weeds, and what priorities they have for research. The interviews were complemented with an all-day focus group conducted with the farmers who were interviewed. All of the farmers interviewed relied heavily on cultural practices, such as crop rotation, fertility management, high-density planting, transplants and the use of cover crops. Mechanical practices were also important for the farmers interviewed. They emphasized selection of the appropriate tools for the situation and techniques for using tools effectively. Most of the farmers interviewed relied on intensive tillage to control perennial weeds. Differences in practices were notable among the farmers interviewed. Several farmers shared innovations both in practices and equipment modification. Farmers identified soil and nutrient management; tillage and cultivation tools and practices; natural herbicides; flame weeding and no-till systems without herbicides as research priorities. Some of the topics suggested for research have already received substantial research attention; identification of these topics by the farmers indicated a need to disseminate research findings through extension materials and demonstration of viable practices.

Published in Agron. J. 106:871–880 (2014) doi:10.2134/agronj13.0286 Copyright © 2014 by the American Society of Agronomy, 5585 Guilford Road, Madison, WI 53711. All rights reserved. No part of this periodical may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopying, recording, or any information storage and retrieval system, without permission in writing from the publisher. ABSTRACT

The alien invasive vines black and pale swallowwort are currently spreading across eastern North America, invading parklands, old fields, restored forest sites, and other natural areas. These plants spread by wind-borne seed and can form dense stands where they become established. Although their current geographic ranges overlap, there is little known overlap locally. Preliminary observations and anecdotal information have associated black swallowwort with low-pH inceptisols and pale swallowwort with high-pH alfisols. We conducted a common garden field experiment repeated over two years and a growth chamber germination experiment to assess whether seedling emergence and performance of these two swallowworts are affected by soil type and/or soil pH. Soil pH was artificially modified. In the common garden experiment, plants of both species grown on the Onondaga County soil type (an alfisol) produced a smaller root mass than on the Orange County soil (an inceptisol). Also, in one of two years more seedlings emerged and plants produced more follicles on the Onondaga County soil. Soil pH did not affect seedling emergence, although plants grown on low pH soils had a smaller root dry mass compared with plants grown on higher pH soils. Soil pH effects on stem length, stem dry mass, and follicle production were inconsistent among years. Species differences were also evident, with more pale swallowwort seedlings emerging than black swallowwort seedlings, whereas black swallowwort plants mostly had greater biomass and fecundity than pale swallowwort plants. In the growth chamber experiment, final percentage seed germination was greater on the Onondaga County soil than on the Orange County soil. The germination speed index as well as the probability to reach 50% germination for black swallowwort was higher on the Onondaga County soil than the Orange County soil, but only at lower pH levels. The germination speed index of pale swallowwort on the Orange County soil was higher than black swallowwort at low, but not high, pH levels. In contrast, black swallowwort had a higher probability of reaching 50% germination than pale swallowwort on the Orange County soil at higher pH levels. Contrary to our expectations, interactions between the two swallowwort species with their associated soil type or with their presumably preferred soil pH were weak, contradictory, or non-existent in both experiments. This suggests that these two species can colonize and grow well in a relatively wide range of soil pH conditions. From a management perspective, our results suggest that the current range and local overlap of these two species will continue to increase and that early detection rapid response (EDRR) programs should be established in susceptible regions not yet colonized by these two invasive vines.

EA GESTAD INRA; International audience; Weed management is a major constraint in organic cropping systems. In 2004, the Cornell Organic Vegetable Cropping Systems Experiment was established in central New York state using a split-plot randomized complete block design with two crop rotation entry points (split-plot factor). Four organic vegetable cropping systems that varied in cropping intensity and tillage (main plot factor) were compared: (1) intensive, (2) intermediate, (3) bio-extensive, and (4) ridge tillage. The basic crop rotation was cabbage, lettuce, potato, and winter squash, with additional short-season crops in the intensive system and with cover crops and fallow substituted for cabbage and potato in the bio-extensive system. In 2014, two uniformity trials were conducted in which oat and then a mixture of sorghum-sudangrass plus Japanese millet were grown uniformly over the entire experiment. Prior to sowing oat, soil samples were collected from each plot and an emergence bioassay was conducted to assess the soil weed seedbank. Crop biomass, weed density, and weed biomass were sampled in the uniformity crops. Soil weed seedbank density was three to four times greater in the intensive, intermediate, and ridge-tillage systems than in the bio-extensive system. The bio-extensive system also had lower weed density and weed biomass in the oat uniformity trial compared with the other three systems. Oat biomass did not differ between the cropping systems. Weed density and biomass in oat were also affected by the crop rotation entry point. Cropping system legacy effects on weed abundance and community composition were greater in the oat than in the sorghum-sudangrass plus Japanese millet uniformity trial. Our results illustrate the effects of different organic vegetable production practices on weed community structure and highlight the value of tilled fallow periods, cover crops, and prevention of weed seed rain for reducing weed populations.

Mohler CL, Dykeman C, Nelson EB & Ditommaso A (2012). Reduction in weed seedling emergence by pathogens following the incorporation of green crop residue. Weed Research.52, 467–477. Summary Because tillage promotes the germination of many weed species and freshly killed plant material favours the growth of microbial pathogens, we hypothesised that the incorporation of green crop residue should temporarily reduce weed seedling emergence relative to unamended soil. Soil with field-incorporated green crop residue was compared with non-amended soil in glasshouse experiments by sowing several weed species at different times after incorporation. Species included Abutilon theophrasti, Chenopodium album, Amaranthus powellii, Setaria faberi, Echinochloa crus-galli and, in one year, lettuce and red clover. Soils with green crop residue reduced seedling emergence for 0–4 days after incorporation by an average of 30%. Comparison of emergence in non-sterilised soil with that in sterilised soil, with and without fresh crop residue, indicated that a biological agent caused the depressed emergence. In the third year of the study, the fungi Fusarium oxysporum and F. chlamydosporum were isolated from seeds exposed to soil amended with green crop residues, and their pathogenicity to seeds and seedlings was confirmed in bioassays. This study indicated that incorporation of fresh crop residue reduces the first flush of weed seedlings following tillage and that this depression in emergence is probably caused by pathogen attack on seeds and seedlings before emergence.

Black and pale swallowwort (BSW and PSW, respectively) are perennial, herbaceous vines in the Apocynaceae that are native to Europe. The species are becoming increasingly abundant in the northeastern United States and southeastern Canada and are difficult to manage. However, we know little about the demographic parameters of these species. We determined the survival, annual rate of vegetative growth, and fecundity of mature clumps of these swallowwort species. We selected four PSW sites (three of which comprised both old-field and forest habitats) in central New York and three BSW old fields in southeastern New York. BSW is largely restricted to higher light habitats in its introduced range. In each habitat, we followed the growth of 30 to 32 randomly selected clumps of similar size (2 to 5 stems clump−1 in the initial year) for 3 to 4 yr. Yearly survival was 99.6 ± 0.3% [mean ± standard error] for PSW and 100 ± 0% for BSW. In old fields, vegetative expansion varied from −0.01 ± 0.1 to 4.6 ± 0.4 stems clump−1 yr−1 for BSW and −0.02 ± 0.2 to 2.1 ± 0.5 stems clump−1 yr−1 for PSW. In forests, PSW growth was lower with vegetative expansion ranging from −0.01 ± 0.1 to 0.8 ± 0.2 stems clump−1 yr−1. Fecundity of PSW in 2007 and 2008 (130 ± 10 viable seeds stem−1 yr−1) was similar to BSW (100 ± 10 viable seeds stem−1 yr−1). Fecundity of PSW in forests was generally lower than PSW in old fields, but it varied greatly among sites (0 to 170 viable seeds stem−1 yr−1). We found that stem growth and fecundity did not vary with clump size (stems per clump). Since vegetative expansion and fecundity rates were high in old-field habitats, but were generally low or nonexistent in forest habitats, we suggest that management of these two invasive vines be focused in higher light environments to reduce overall seed production and its subsequent spread to surrounding areas.

Vincetoxicum rossicum (pale swallow-wort) is a non-native, perennial, herbaceous vine in the Apocynaceae. The species’ abundance is steadily increasing in the northeastern United States and southeastern Canada. Little is known about Vincetoxicum species recruitment and growth. Therefore, we conducted a field experiment in New York State to address this knowledge gap. We determined the establishment, survival, and growth of V. rossicum during the first 2 years after sowing in two old fields subjected to four disturbance regimens. We hypothesized that establishment and survival would be higher in treatments with greater disturbance. At the better-drained location, overall establishment was 15 ± 1% [mean ± standard error] and did not differ among treatments. At the poorly drained location, establishment varied by treatment; mowed and control plots had greater establishment [10 ± 2%] than herbicide + tillage and herbicide-only plots [1.6 ± 0.5%]. Of those seedlings that emerged, overall survival was high at both locations (70–84%). Similarly, total (above + belowground) biomass was greater in herbicide + tillage and herbicide-only plots than in mowed and control plots at both locations. Thus, V. rossicum was successful in establishing and surviving across a range of disturbance regimens particularly relative to other old field species, but growth was greater in more disturbed treatments. The relatively high-establishment rates in old field habitats help explain the invasiveness of this Vincetoxicum species in the northeastern U.S. and southeastern Canada.

Determining which traits may allow some introduced plant species to become invasive in their new environment continues to be a key question in invasion biology. Vincetoxicum rossicum is an invasive, perennial vine colonizing natural and seminatural habitats primarily in the northeastern United States and southeastern Canada. More than half its seeds exhibit polyembryony, a relatively uncommon condition in which a single seed produces multiple seedlings. For evaluating the potential consequences of polyembryony on invasiveness, V. rossicum plants derived from seeds of three embryonic classes-singlets, doublets, and triplets (one, two, and three seedlings per seed, respectively)-were paired in all combinations intraspecifically and with the co-occurring native herbs Solidago canadensis and Asclepias syriaca in a greenhouse study. Vincetoxicum rossicum biomass was 25-55% greater and follicle production 55-100% greater under intraspecific competition compared with interspecific competition. However, within a competitive environment, follicle production varied little. Regardless of competitive environment, V. rossicum originating from seeds with a greater number of embryos typically performed no better than plants arising from seed with fewer embryos (singlets = doublets = triplets)-except intraspecifically where doublets outperformed singlets, and with S. canadensis where triplets outperformed singlets. Our findings suggest that overall performance and fitness of V. rossicum is higher in monocultures than in mixed stands and that its ability to invade new habitats may not be attributable to the production of polyembryonic seeds.

A standard method for evaluating weed seed predation is needed to facilitate generalizations across studies. Identification of general trends could allow practical recommendations for enhancing weed seed predation in agricultural systems. The objective of this study was to compare the commonly used sandpaper and soil substrate methods for offering weed seeds when assessing seed predation rates. Invertebrate seed predators and associated weed seed predation levels were measured in June to July, August, and September of 2005 and 2006 within a conventionally managed corn system. Seed predation levels of three common weed species, velvetleaf, giant foxtail, and common lambsquarters, were estimated using feeding trials in which 40 seeds of each species were offered over a 48 h period using the two substrates. Exclosures were used to distinguish total predation from predation by invertebrates alone. In addition, we investigated the use of geospatial analysis to estimate spatial autocorrelation of inver...

The objectives of this research were to characterize the extent of intraspecifc variation in seed characteristics of Powell amaranth and to evaluate whether such variation was correlated with crop rotation history of the collection sites. We compared characteristics of seeds originating from dairy farms with a corn–alfalfa crop rotation history with seeds originating from farms with a history of intensive vegetable production. We hypothesized that (1) multiple years of perennial alfalfa would select for greater seed dormancy and longevity in seeds of the summer annual Powell amaranth, (2) earlier spring planting dates of corn and alfalfa compared with most vegetable crops would select for earlier emergence, and (3) greater competition and lower soil moisture in the nonirrigated corn–alfalfa rotation would select for greater seed size. Seeds from 10 to 20 plants from each of 10 farms from each habitat were collected in the fall of 2002 and 2003 in central New York. To control for maternal effects ...

The primary objectives of this research were (1) to characterize intraspecific variation in Powell amaranth seed germination and emergence response to nitrogen fertilization, and (2) to evaluate whether germination and emergence characteristics varied between seeds from populations originating on organic vs. conventional vegetable farms. We hypothesized that nonherbicide–based weed management and use of slower-releasing forms of N on organic farms may have selected for seeds with lower dormancy and lower germination sensitivity to N fertilization than seeds from conventional farms. Seeds were collected from five conventional and five organic vegetable farms in central New York State. A second generation of seeds, produced under common greenhouse conditions and stored for at least 3 mo at 5 C was tested for both germination in petri dishes and emergence in the field under multiple rates of ammonium nitrate (NH4NO3). Both seed germination and emergence were greater for seeds originating from organi...

Vertical position of weed seeds in the soil column is one of the critical factors governing the density of emerged seedlings, but data on movement of subsurface seeds and seed surrogates by tillage are limited. In this experiment ceramic beads were seeded at 0, 4, 8, 12, 16 and 20 cm using specially constructed equipment. Plots were then tilled at right angles to the bead slots with either a moldboard plow, a chisel plow with curved blades, a chisel plow with straight blades, heavy tandem disks, a rotary tiller, or left untilled. Soil was sampled to determine bead positions. The probability matrix describing movement of beads from each soil layer to any other was estimated for each tillage regimen by maximum likelihood. Movement patterns for moldboard plow, the two chisel plow treatments, and disk and rotary tillage differed significantly. The difference between disking and rotary tillage was only marginally significant (0.05 80% probability of remaining in at their original depth. Tillage by moldboard plow showed the expected soil inversion, but burial of surface beads was greater than return of buried beads to the surface. For example, the probability of a surface bead ending up below 10 cm was over 77% whereas the probability of a bead originally in the 14–18 cm layer moving to the top 10 cm was only 44%. Sampling of the no tillage and moldboard plow treatments the following spring showed an upward movement of beads by natural causes for beads below 14 cm. To allow flexibility in vertically structured population models, the movement probability matrices were fitted with a continuous model based on the beta distribution. The model showed that for a hypothetical uniformly distributed seed bank, 97% of seeds in the top 4 cm following moldboard plowing had arrived there from greater depths. In contrast, only about one third of seeds in the top 4 cm of soil after tillage by chisel, disk or rotary tiller arrived from deeper layers.

Summary Variations in climate are widely recognized as central factors governing the competitive balance in mixed-species plant communities. In agricultural systems, highly variable patterns of crop yield reduction as a function of weed density have been documented across sites and among years at the same site for several crop–weed combinations. This variation is typically attributed to contrasting environmental conditions. Despite broad acknowledgement of their importance, experimental and temporal limitations have constrained the investigation and systematic understanding of environmental controls on the dynamics of competition. For several well-studied crop–weed associations, aggregating historical data from similar competition experiments provides an opportunity to explore interference relationships over an array of conditions. In this study, 19 site-years of maize –Abutilon theophrasti (velvetleaf) data were compiled and the weather characterized (i.e. average ambient temperature and moisture regime) for discrete portions of each growing season. These features were then related to patterns of maize yield loss from A. theophrasti interference at high weed densities. Results of this analysis suggest that temperatures following establishment, together with the presence or absence of water stress during the maize crop's exponential growth phase, account for over 60% of the observed variation in relative yield loss.

Invasive species have received considerable attention in recent years, but research has primarily focused on invasive species of natural habitats. Furthermore, cropland weeds have often been viewed as possessing a “general-purpose genotype” and therefore exhibiting relatively static genetics. However, a more current view is that weeds are capable of rapid genetic change, thereby making analysis of their evolutionary ecology a potentially valuable component for the development of sustainable weed management systems. In particular, further analysis of ongoing evolutionary change in cropland weeds is important because (1) most cropland weed species exhibit considerable adaptability, (2) cropland agriculture is continuously changing, and (3) further research on weed adaptability is needed to design cropping systems to address evolutionary change. In this review, we examine the potential of cropland weeds to evolve so as to affect their invasiveness. There is abundant evidence of genetic variation within and among weed populations in traits relevant to invasiveness, including seed germination patterns, life history traits, physiological adaptability and adaptation to disturbance and resource fluctuations. Approximately half of cropland weed species are primarily selfing, and species with a high degree of selfing tend to exhibit homogeneity within populations but divergence among populations. We identified four critical areas for future research: conflicting selection pressures on weeds in agroecosystems, feed-back driven dynamics of human-weed co-evolution, co-evolutionary mechanisms of weed adaptation in conjunction with other weed species or organisms, and the role of weed evolution in the restoration of agroecosystems.

We extend the resource concentration hypothesis (herbivorous insects are more likely to find and stay in more dense and less diverse patches of their host plants) to plant communities. Specifically, whenever superior plant competitors spread to form dense stands, they will be found and attacked by their specialist insect enemies. This will decrease host plant abundance, causing a reduction in standing crop biomass, which will indirectly increase subordinate competitors and plant species richness. In this study, we found that a native, specialist chrysomelid beetle (Trirhabda virgata) in an old-field community decreased total standing crop biomass, leading to an increase in plant species richness. This reduction in biomass was due solely to a reduction in the biomass of the beetle's host plant, meadow goldenrod (Solidago altissima), which was the dominant plant species in this community. Our results demonstrate that when a superior competitor increases in density, the per-stem impact of herbivores increase...

The unimbibed crush test, in which seed viability is evaluated by crushing and visual inspection of dry seeds, was compared with tetrazolium staining, an established method of testing seed viability. The unimbibed crush test potentially provides an immediate and rapid method for determining seed viability. Six sets of seed lots, involving the four weed species, velvetleaf, common lambsquarters, redroot pigweed, and smooth pigweed, were tested by each method. For four of the six sets, results from the crush test were statistically indistinguishable from those of the tetrazolium test. For the other two sets of seed lots, the crush test indicated higher viability than did the tetrazolium test. The crush test may be most useful for seed bank surveys in which many samples are typically processed, and most of the variation in density of viable seeds is associated with number of seeds present rather than percentage viability. Its use in more exacting circumstances like seed survival studies requires ca...

Mulches on the soil surface are known to suppress weed emergence, but the quantitative relationships between emergence and mulch properties have not been clearly defined. A theoretical framework for describing the relationships among mulch mass, area index, height, cover, light extinction, and weed emergence is introduced. This theory is applied to data from experiments on emergence of four annual weed species through mulches of selected materials applied at six rates. Mulch materials, in order from lowest to highest surface-area-to-mass ratio, were bark chips, Zea mays stalks, Secale cereale, Trifolium incarnatum, Vicia villosa, Quercus leaves, and landscape fabric strips. The order of weed species' sensitivity to mulches was Amaranthus retroflexus > Chenopodium album > Setaria faberi > Abutilon theophrasti, regardless of mulch material. The success of emergence through mulches was related to the capacity of seedlings to grow around obstructing mulch elements under limiting light conditions. Mul...

Soil chemical and biological dynamics in mixed use landscapes are dependent on the distribution and pattern of soil moisture and water transport. In this paper we examine the effect of different grid sizes on soil water content for a spatially explicit, variable-source-area hydrology model applied to a watershed in central New York. Data on topography, soil type, and land use were input at grid sizes from 10 to 600 m. Output data consisted of runoff and spatial pattern of soil moisture. To characterize the spatial variability at different grid sizes, information theory was used to calculate the information content of the input and output variables. Simulation results showed higher average soil water contents and higher evaporation rates for large grid sizes. During a wet year, runoff was not affected by grid size, whereas during a dry year runoff was greatest for the smallest grid size. While the information content (i.e., spatial variability) of soil type and land use maps was not affected by the different grid sizes, increasing grid sizes caused the information content of the slope gradient to decrease slightly and the Laplacian (or curvature of the landscape) to decrease greatly. In other words, increasing grid cell size misrepresented the curvature of the landscape. During wet periods the decrease in information content of the soil moisture data was the same as for the Laplacian as grid size increased. During dry periods, when local fluxes such as evaporation and runoff determine the moisture content, this relation did not exist. The Laplacian can be used to provide a priori estimates of the moisture content deviations by aggregation. These deviations will be much smaller for the slowly undulating landscapes than the landscape with steep valleys simulated in this study.

Summary Tillage causes both vertical redistribution of weed seeds and changes in soil physical properties. These two factors are rarely distinguished in studies of the impact of tillage on seedling emergence or seed survival. In this study, seeds of Chenopodium album L., Amaranthus retroflexus L., and Abutilon theophrasti Medik, were planted at particular depths in pots of undisturbed or stirred soil to separate these effects. Emergence and survival data were analysed by non-linear regression to determine the nature of significant differences between treatments. Emergence increased with slight burial and then decreased exponentially at greater depths. Average emergence over all depths was generally greater in tilled sou than in unfilled soil, particularly for C. album and A. retroflexus. Seed survival approached a maximum with increasing depth. Average survival of seeds that did not produce emerged seedlings was greater in tilled soil than in untilled soil for C. album and A. retroflexus. Thus, tillage affects emergence and seed survival of weeds through changes in soil conditions independently of effects resulting from redistribution of seeds in the soil profile.

Methods were developed to evaluate the performance of a decision-tree model used to predict landscape-level patterns of potential forest vegetation in central New York State. The model integrated environmental databases and knowledge on distribution of vegetation. Soil and terrain decision-tree variables were derived by processing state-wide soil geographic databases and digital terrain data. Variables used as model inputs were soil parent material, soil drainage, soil acidity, slope position, slope gradient, and slope azimuth. Landscapescale maps of potential vegetation were derived through sequential map overlay operations using a geographic information system (GIS). A verification sample of 276 field plots was analyzed to determine: (1) agreement between GIS-derived estimates of decision-tree variables and direct field measurements, (2) agreement between vegetation distributions predicted using GIS-derived estimates and using field observations, (3) effect of misclassification costs on prediction agreement, (4) influence of particular environmental variables on model predictions, and (5) misclassification rates of the decision-tree model. Results indicate that the prediction model was most sensitive to drainage and slope gradient, and that the imprecision of the input data led to a high frequency of incorrect predictions of vegetation. However, in many cases of misclassification the predicted vegetation was similar to that of the field plots so that the cost of errors was less than expected from the misclassification rate alone. Moreover, since common vegetation types were more accurately predicted than rare types, the model appears to be reasonably good at predicting vegetation for a randomly selected plot in the landscape. The error assessment methodology developed for this study provides a useful approach for determining the accuracy and sensitivity of landscape-scale environmental models, and indicates the need to develop appropriate field sampling procedures for verifying the predictions of such models.

The environmental impact quotient (EIQ) developed by Kovach et al. (1. Kovach, C. Petzoldt, J. Degni & J. Tette. 1992. A method to measure the environmental effect of pesticides. N.Y. Food Life Sci. Bull. No. 139.) is an effort to fill an important gap: the need to provide growers and others with easy-to-use information about the adverse effects of pesticides. However, flaws in both the formula and its conceptual underpinnings serve to make the information provided misleading. Although Kovach et al. provides a great deal of information and many interesting ideas, we recommend that EIQ presented there not be used as a tool to evaluate field applications of pesticides. Further, current understanding of pesticides and their effects is not sufficient to allow the environmental effects of a pesticide to be captured by a single number. We discuss alternate ways to provide growers and policymakers with usable information about pesticides.

The nonnative vine Vincetoxicum rossicum threatens several ecosystems in the Lower Great Lakes Basin of North America. One feature that may contribute to its invasiveness is the production of some seeds with multiple embryos (polyembryony), which may be beneficial as a bet-hedging strategy in variable environments. However, lower seed reserves per embryo in polyembryonic seeds may entail costs in low-light environments. The effect of seed from three embryonic classes (1, 2, or 3 embryos/seed) on V. rossicum survival and growth was studied under two forest understory light environments: full canopy (shade) or canopy gaps (light) in New York state. Two seedling cohorts were planted, in May 2004 and in May 2005. The survival and growth of seedlings was monitored biweekly for two (2005 cohort) or three (2004 cohort) seasons. For both cohorts, plants grown in canopy shade had reduced survival and growth compared with those grown in gaps. Contrary to expectations, seed embryo number had no effect on the final height, survival, or dry mass of plants in either habitat. Our results suggest that any fitness advantage provided by polyembryony may be habitat (light) dependent and not a general trait that affords V. rossicum a benefit in all habitats colonized.

Summary: Resume: Zusammenfassung Vicia villosa Roth (hairy vetch) or Secale cereale L. (rye) residue at 2 to 4 times the natural field rate reduced seedling emergence of a wide variety of weed species. However, the natural rate of residue was insufficient for control of most weed species. The form of the response of weed emergence to increasing rates of residue varied among species: emergence of some species showed a monotonic decline in response to increasing rates of residue whereas emergence of others increased at low rates before declining at higher rates. In addition to reducing number of seedlings, high rates of residue delayed the date of emergence. Seed size was generally not a good indicator of a species' ability to emerge through residue, although one very large seeded weed, Abutilon theophrasti. was especially insensitive to residue. V. villosa and 5. cereale did not differ greatly in their effect on weeds early in the growing season, but V. villosa decomposed more rapidly and allowed greater emergence later in the season. Effet de residus de Vicia villosa Roth et de Secale cereale L. sur la levee des mauvaises herbes Des residus de Vicia villosa Roth (Vesce velue) et de Secale cereale L. (seigle) a 2–4 fois la quan-tite normale reduisaient la levee d'un large spectre de mauvaises herbes. Cependant la quantite normale de residus etait insuffisante pour lutter contre la plupart d'entre elles. La reponse de la levee des mauvaises herbes a des quantites crois-santes de rVsidus differait selon les especes. Pour certaines, la levee subissait un declin monotone; pour d'autres, elle augmentait en presence de faible quantites de reesidus puis decroissait pour des quantites plus importantes. Ces dernieres reduisaient la levee et la retardaient egalement. La taille des graines n'etait generalement pas un bon indicateur de la capacite des especes a lever au travers des residus, bien qu'une treAs grosse grains, Abutilon theophrasti Medic., etait particu-lierement insensible aux residus. V. villosa et 5. cereale ne differaient pas beaucoup dans leur effet sur les mauvaises herbes en debut de saison, mais V. villosa se decomposait plus rapidement et permettait des levees plus importantes en fin de saison. Wirkung von Emteruckstanden von Vicia villosa Roth und Secale cereale L. aufdie Keimung von Unkrautern Durch 2- bis 4 fache Ernterackstande der Be-haarten Wicke (Vicia villosa) und des Roggens (Secale cereale) wurde die Keimung einer grosen Zahl von Unkrautarten herabgesetzt. Normale Ernteruckstandsmengen behinderten jedoch die Keimung nur unzureichen. Die Art und Weise der Reaktion der Keimung auf hohere Riick-stande war von Art zu Art verschieden: Die Keimung einiger Arten nahm mit zunehmender Menge an Ernteruckstanden gleichmasig ab, wahrend sie bei anderen bei niedrigen Mengen anstieg, urn dann bei hoheren Mengen abzunehmen. Zusatzlich zur Verringerung der Keimpflanzendichte wurde bei hohen Ruck-standsmengen die Keimung verzogert. Die Samengrose war im allgemeinen kein guter Indikator fur die Fahigkeit einer Art, durch das Ruckstandsmaterial zu keimen, obwohl eine Art mit grosen Samen, Abutilon theophrasti, gegenuber diesen Ruckstanden besonders unempfindlich war. Die Ruckstande der beiden Kulturpflanzen zeigten zu Beginn der Vegetationsperiode keine grosen Wirkungsunterschiede, aber die Ruckstande von Vicia villosa zerfielen schneller und liesen spater eine starkere Keimung zu.

Cover crop residue on the surface of soils in no-tillage systems can suppress weed emergence and growth. Although allelopathy often is invoked to explain weed suppression by residue, physical alterations of the seed environment could be important as well. This experiment was conducted to determine the light, temperature, and moisture conditions under cover crop residue. Hairy vetch (Vicia villosa Roth) and rye (Secale cereale L.) were desiccated with a contact herbicide and residue rates ranging from one-fourth to four times the natural residue biomass were established in experiments at Beltsville, MD and Ithaca, N.Y. Photosynthetic photon flux density (PPFD) was determined above and below residue at approximately monthly intervals after initiation of the experiment. Transmittance of PPFD through residue declined according to an exponential decay function of residue biomass. Transmittance was similar through hairy vetch and rye residue initially, but as the experiment progressed, transmittance through hairy vetch residue was greater than that through rye because of faster decomposition of hairy vetch residue. Spectral analysis from 400 to 1100 nm showed a slight increase in transmittance as wavelength increased resulting in a slight lowering of the red (660 nm) to far-red (730 nm) ratio relative to that of unobstructed sunlight. Soil maximum temperature and daily soil temperature amplitude were reduced by cover crop residue. Residue prevented the decline of soil water content during droughty periods. Results indicated that reductions in light transmittance and daily soil temperature amplitude by cover crop residue were sufficient to reduce emergence of weeds but that maintenance of soil moisture could increase weed emergence.

A simple model is developed in which the density of weed seedlings emerging in a field is related to (1) the ability of seedlings to emerge from various depths in the soil, (2) the survival of seeds at different depths, and (3) the depth of seed burial in no tillage, rotary tillage, and plow tillage. Other tillage regimes are considered by analogy. Literature is reviewed to determine biologically reasonable functions describing seedling emergence, seed survival, and distribution of seeds with depth, and parameters of these equations are estimated from data in the literature. Problems related to the mathematical description of these phenomena are discussed, and it is noted that some commonly held beliefs regarding survival of seeds in the soil are mutually incompatible. Although many studies have investigated the persistence of seeds as a function of depth in the soil, few have distinguished death from the production of seedlings. The model indicates that in the first year following input of seeds to the soil, no tillage will have more seedlings than tillage, but in later years no tillage will likely have fewer seedlings unless innate or induced dormancy is high or seed survival near the soil surface is unusually good. If seed return is allowed, no tillage or minimum tillage will have more seedlings perennially. Recovery of good weed control following a year with substantial seed input may be easiest if the soil is plowed deeply to bury the seeds, and then shallow or no tillage is used in subsequent years to avoid returning seeds to the surface. Much of the literature on the effects of tillage on weed density is difficult to interpret because little indication is given of the vertical distribution of seeds in the soil at the beginning of the experiment.

Effects of several stale seedbed procedures on weed density and biomass were evaluated on a silt loam soil in central New York. After an initial rotary tillage, weeds were allowed to emerge and either single or multiple applications of glyphosate, propane flame, spring tine weeder, springtooth harrow, or rotary tiller were used to kill the weeds over a 4-week period. The last (or only) application occurred immediately prior to simulated seeding of a crop performed by passing an empty seeder through the plots. These stale seedbed treatments were compared with a control consisting of a single rotary tillage just before simulated planting. Flaming or glyphosate stale seedbed techniques significantly reduced density and biomass of the principal broadleaf species, common purslane (Portulaca oleracea L.) and common chickweed (Stellaria media (L.) Cyrillo), in most cases. A single delayed flame or glyphosate stale seedbed treatment was usually as effective as multiple treatments. None of the stale seedbed techniques was effective against yellow nutsedge (Cyperus esculentus L.). A flexible tine weeder was not effective as a stale seedbed weed-killing treatment in this study because of poor penetration of crusted soil. Penetra- tion was better with a springtooth harrow, but this failed to reduce weed density. None of the stale seedbed treatments fully controlled weeds. However, glyphosate or flaming a stale seedbed could be incorporated into integrated weed management programs to improve control and reduce the need for herbicides. Broadleaf weed density within 3.8 cm of the center of the seeder wheel track was greater than elsewhere in the plot. Chemical name used: N-(phosphonomethyl)glycine (glyphosate). The premise behind the stale seedbed prac- tice is that by delaying seeding after crop seed- bed preparation, flushes of weeds can be in- duced to sprout, and then be killed. If the weeds are killed with minimal soil disturbance, the weed seedbank in the upper few centimeters of soil will be depleted, resulting in less weed pressure against subsequent crops. Most investigations of the stale seedbed technique have involved herbicide-based soy- bean (Glycine max (L.) Merrill) systems (Heatherly et al., 1993; Lanie et al., 1993, 1994; Oliver et al., 1993), despite wide use of the

Sweet corn was grown with a living mulch of white clover, a dead mulch of rye, and without mulch, in both till and no-till conditions. Unplanted controls were also included in the experimental design. Corn yields were highest in clover treatments early in the experiment but lowest in later years. The declining yields in the clover living mulch were related to the strip application of glyphosate which allowed establishment of perennial and biennial weeds, notably dandelion and horseweed. These overwintering weeds apparently prevented effective control of summer annuals, especially redroot pigweed, common lambsquarters and large crabgrass, by atrazine and metolachlor. Presence of a rye mulch decreased weed biomass and had no detrimental effect on corn yield. In general, corn yield was not affected by tillage, although the number of marketable ears was reduced in the no-till treatments during the drought year of 1988. The much greater weed biomass in the unplanted control treatments showed the importance of crop competition for weed control in sweet corn cropping systems.

Weeds from an ecological perspective Weeds share certain ecological characteristics that distinguish them from other plants. Specifically, weeds are plants that are especially successful at colonizing disturbed, but potentially productive, sites and at maintaining their abundance under conditions of repeated disturbance . That is, weeds are the plants that thrive where soil and climate are favorable to plant growth, but disturbance frequently reduces competition among plants to low levels. Unlike previous conceptions of weediness (Baker, 1965; Harlan & de Wet, 1965; Buchholtz et al. , 1967), this ecologically based definition lacks reference to humans and human disturbance. The species people refer to as weeds mostly existed prior to human disturbance, and the repertoire of behaviors that makes them invasive and persistent in human-dominated habitats largely evolved independently of human society. Nevertheless, as discussed in Chapter 10, human activities selectively modify weed characteristics such that weeds are becoming better adapted to human disturbance regimes. The subcategory of weeds dealt with in this book consists of the weeds of agriculture – specifically, the plants that colonize and increase in the disturbances created by farming. These are sometimes termed agrestal weeds, as distinguished from the ruderal weeds of roadsides, waste piles, and other non-agricultural disturbances (Baker, 1965). Agricultural weeds share certain life-history characteristics that adapt them for life on farms (Table 2.1). The thesis of this chapter is that understanding life-history characteristics provides insights into how weed management practices work and how they can be improved.

Concerns over environmental and human health impacts of conventional weed management practices, herbicide resistance in weeds, and rising costs of crop production and protection have led agricultural producers and scientists in many countries to seek strategies that take greater advantage of ecological processes and thereby allow a reduction in herbicide use. This book provides principles and practices for ecologically based weed management in a wide range of temperate and tropical farming systems. After examining weed life histories and processes determining the assembly of weed communities, the authors describe how tillage and cultivation practices, manipulations of soil conditions, competitive cultivars, crop diversification, grazing livestock, arthropod and microbial biocontrol agents, and other factors can be used to reduce weed germination, growth, competitive ability, reproduction and dispersal. Special attention is given to the evolutionary challenges that weeds pose and the roles that farmers can play in the development of new weed-management strategies.

Effects of tillage (till, no-till), mulch (none, dead Secale cereale), and crop competition (none, sweet corn) on weed seed production and seed banks are reported. Data are presented for Amaranthus retroflexus, Chenopodium album, Portulaca oleracea and Digitaria sanguinalis. These constituted the majority of the weeds present during the cropping season. Seed bank data for a suite of winter annuals active during the fallow season are also presented. Weed seed production was estimated by dimension analysis. Most seeds were produced by the first cohort of plants to emerge. Amaranthus and Digitaria showed significantly greater seed production in no-till than in till treatments. Seed production by Amaranthus, Chenopodium and Portulaca was significantly greater in the absence of a corn crop. The only effect of Secale cereale mulch was to decrease seed production by Portulaca in 1 year. Differences among treatments in the number of seeds produced per unit area were primarily due to effects on individual plant size and seed production. Effects on seedling density and rate of survival to maturity were of secondary importance. Seed banks of Digitaria were greater in no-till than in till treatments, whereas the seed banks of winter annual forbs were greater in till treatments. Presence of a corn crop did not affect seed banks until the final year of the study, at which time seed banks of Amaranthus, Chenopodium and Portulaca were greater in the absence of a crop. An uncropped treatment which did not receive herbicides had exceptionally large seed production and seed banks of Amaranthus and Chenopodium, but not Portulaca and Digitaria, which apparently suffered from competition with the two larger species.