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In New Zealand, the genus Pselaphaulax Reitter is represented by thirteen species. Six new species are described: Pselaphaulax hornabrooki Owens and Carlton, sp. nov.; Pselaphaulax intermedius Owens and Carlton, sp. nov., Pselaphaulax nunni Owens and Carlton, sp. nov.; Pselaphaulax pelorus Owens and Carlton, sp. nov.; Pselaphaulax ramsayi Owens and Carlton, sp. nov.; and Pselaphaulax tararua Owens and Carlton, sp. nov. Pselaphus trifoveatus Broun, 1914 is synonymized under Pselaphus pilifrons Broun, 1914 syn. nov. and is transferred to Pselaphaulax comb. nov. Two species, Pselaphus pilistriatus Broun, 1880 and Pselaphus fuscopilus Broun, 1886 are synonymized under Pselaphus pauper Sharp syn. nov., which is also transferred to Pselaphaulax comb. nov. Pselaphus meliusculus Broun, 1893 is also transferred to Pselaphaulax comb. nov. Lectotypes and paralectotypes are designated for the Broun and Sharp species. Distributional maps and line drawings of diagnostic characters for each species are provided. A regional checklist and key are provided to species of the genus that occur in New Zealand.

Pselaphaulax hornabrooki Owens & Carlton, sp. nov. (Fig. 6a–c) Type Material. Holotype: (male) Wellington 10-28 1946 R.Hornabrook (NZAC). Paratypes: (3 females) holotype locality (NZAC). Etymology. This species is named after the collector of the type series, R. Hornabrook. Diagnosis. Pselaphaulax hornabrooki can be distinguished from other species in the genus by the combination of a median lobe curved to the right and an associated left lateral process that is narrow and elongate, extending beyond the median lobe. Description. MALE holotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 19 facets. Frontal margin of rostrum in dorsal view slightly rounded with sparse suberect setae. Gular mound narrowed apically and posteriorly, swelling distinctly longer than wide, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, apical swelling roughened, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus faint, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae and single sutural fovea; single discal stria, sutural striae present; apical fringe of setae absent. Abdomen: Tergites of usual form for tribe. Ventrite II with median area slightly flattened at base, apically margined by short, sparse setae. Genitalia: aedeagus asymmetrical. Median lobe distally curved to the right. Ventral process downward curved, distally, shorter than median lobe. Left lateral process elongate narrow, extending beyond median lobe. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median ovate depression and thickened area at apex. Distribution. Pselaphaulax hornabrooki is known only from four collecting events in Wellington (WN). Material examined. Wellington Region: (1 male, 3 females) New Zealand: WN: Otaki Forks, Hector Trig Moss, 6 Sep 1965 JI Townsend, 65/480 (LSAM); (8 males, 6 females) New Zealand: WN: Hutt Valley, Taita Litter, 1958-1961 K Lee, 61/16 (LSAM); (1 male, 1 female) Masterton 1410 10/2/1952 R. Hornabrook // Entomology Division D.S.I.R. New Zealand (NZAC)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, Revision of the New Zealand species of the Genus Pselaphaulax Reitter (Staphylinidae: Pselaphinae: Pselaphitae: Pselaphini), pp. 221-244 in Zootaxa 5155 (2) on pages 226-227, DOI: 10.11646/zootaxa.5155.2.3, http://zenodo.org/record/6673412

Pselaphaulax pilifrons (Broun), 1914, comb. nov. Pselaphus pilifrons (Broun 1914a: 93) Pselaphus trifoveatus (Broun 1914b: 172), synon. nov. (Fig. 9a–c) Type Material Examined. After examining the type series of Pselaphus pilifrons and P. trifoveatus in the NHM and reviewing the original species descriptions of Broun, we determined the two species to be identical; we hereby synonymize them and transfer them to the genus Pselaphaulax. In the case of P. trifoveatus, lectotype designation is necessary to fix the taxon to a name bearing type. Broun did not specifically identify the holotype specimen, and the syntype series includes multiple species Pselaphus trifoveatus: The following specimen is here designated the lectotype: (male) 3538.// New Zealand Broun Coll. Brit. Mus. 1922–482//Mc Clennans 25-3-1912 (lectotype). The following four specimens are included in the syntype series in the NHM and have been determined to represent additional specimens of P. trifoveatus, and are here designated as paralectotypes: (1 male) TYPE (red/orange label)// 3538. ♂ // New Zealand Broun Coll. Brit. Mus. 1922–482// Mc Clennans 25-3-1912; (2 females) 3538. ♀ // New Zealand Broun Coll. Brit. Mus. 1922–482// Rakaia 9-8-1912; (1 female) New Zealand Broun Coll. Brit. Mus. 1922–482// Mc Clennans 25-3-1912. These specimens match the manuscript number, “3583”, locality “McClennans Bush and Raikaia Gorge”, and collecting date, 1912, mentioned in Broun’s original description of the syntype series (1914b). Pselaphus pilifrons: (1 female) 3402// Tisbury 9/10/10// Pselaphus pilifrons // TYPE (red/orange label) (Holotype). This specimen matches the manuscript number, “3402”, as well as the locality, “Tisbury, Invercargill” of the one specimen mentioned by Broun in the original description (1914b), so this specimen was determined to represent the holotype of P. pilifrons. No other specimens are included in the type series in the NHM. Diagnosis. Pselaphaulax pilifrons may primarily be separated from the other species in group 1 by the reduction of the ventral process with a short, blunt apex. The well-developed fleshy process associated with the median lobe is also unique among species in the group. Redescription. MALE lectotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 19 facets. Frontal margin of rostrum in dorsal view weakly emarginate with sparse suberect setae. Gular mound modified, slightly longer than wide, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, distal swollen area smooth, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus faint, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, single sutural fovea; discal stria and sutural striae present; sparse apical fringe of setae present. Abdomen: Tergites of usual form for tribe. Ventrite II with median area slightly flattened throughout length of ventrite, apically margined by short, sparse setae. Genitalia: aedeagus asymmetrical. Median lobe distally curved downwards, protruding from fleshy dorsal extension. Ventral process blunt, shortened, barely extending beyond basal bulb. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median ovate depression and thickened area at apex. Distribution. Pselaphaulax pilifrons is known from a number of collecting events in Mid Canterbury (MC) and the lectotype collection locality in Southland (SL). Material examined. Middle Canterbury region: (3 males, 1 female) New Zealand: MC: Foot of Mt. Somers Moss, 28 Jul 1966 AD Lowe, 66/220 (LSAM, one m slide-mounted); (1 female) New Zealand: MC: Foot of Mt. Somers Moss, 28 Jul 1966 AD Lowe, 66/219 (LSAM); (1 male) New Zealand: MC: McClennan's Bush Moss, 28 Jul 1966 AD Lowe, 66/221 (LSAM); (1 female) New Zealand: MC: Mt. Hutt, McClennan's Bush Moss, 11 Dec 1973 G Kuschel, 73/142 (LSAM); (2 males, 1 female) NEW ZEALAND MC Mt Alford 1067m NW of Methven 2 Feb 1976 G.W. Ramsay // Litter 76/16 (NZAC); (1 female) NEW ZEALAND MC Peel Forest Park Blandswood 7 Nov 1979 J.C.Watt // Litter 79/148 (NZAC); (1 female) NEW ZEALAND MC Sharplin Falls Bowyers stream 610m Mt Somers 2 Feb 1976 // G.W. Ramsay Litter and moss 76/18 (NZAC); (2 males) Pudding Hill, nr. Methven, Canterbury // Coll. T. Hall 13.4.1915 // 3538// T.Broun (NZAC); (5 females) Pudding Hill, nr. Methven Canterbury //3538 Pselaphus trifoveatus Broun (NZAC); (6 specimens) 21.i.52, moss, Riccarton Bush, Christchurch, S.I. (NZAC); (3 specimens) 13.x.51 moss Riccarton Bush, Christchurch, S.I. (NZAC); (2 specimens) 9.x.51 moss, 7 miles Christchurch (Bell-Bush) S.I. (NZAC); (2 specimens) 31.i.52 moss K. Valley, Cant., S.I. (NZAC); (3 specimens) 17.i.52, moss n. Ashley Gorge Cant. S.I. (NZAC); (1 male) Ex Lucerne Lincoln College, N.Z. ex: (turf) 10 pm 3 10. 1967 Coll. R. Macfarlane (NZAC); (1 male) Oakden 23-9-13// 3538// T.Broun Collector (NZAC); (2 males, 1 female) Mt. Algidus 9.10.1913 // T.Broun Collection (NZAC); (1 female) Algidus 25-9-13// 3538// T.Broun Collection (NZAC).

Pselaphogenius cornus­­­Owens­­­&­­­Carlton,­­­sp.­­­nov. (Fig. 9a–c) Type­­­Material.­­­ Holotype: (1­­­male)­­­ New Zealand:AK: Mt. Dome, litter 15 Mar 1978 SB Peck (NZAC). Paratypes: (1­­­male,­­­1­­­female) holotype locality (NZAC). Etymology.­­­ This specific epithet of this species refers to the horn-shaped, upturned median lobe of the aedeagus in males of this species. Diagnosis.­­­ Externally, this species is difficult to distinguish from many other species of Pselaphogenius in New Zealand. The form of the aedeagus, particularly the wing-shaped lateral processes, is unique. Description.­­­ MALE holotype. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes small, composed of five facets. Frontal margin of rostrum acute, with sparse suberect setae in dorsal view. Gular mound modified, longer than wide, broad anteriorly and narrowed posteriorly; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV longer than head, narrow, widened at base and distal 1/5, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae and median antebasal fovea reduced to minute pits. Mesoventrite with median shield unmodified. Metaventrite afoveate, medially flattened, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae and sutural fovea present; single discal stria, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, shallow oval median depression reaching apex. Genitalia: aedeagus symmetrical. Median lobe compressed laterally, apical end upturned. Lateral processes almost as long as median lobe, bearing row of feather-like ventral extensions giving wing-like appearance. Ventral process narrow, straight, slight ventral curve, distally, about as long as median lobe. Parameres present, as long as median lobe. Females.­­­ Integument,­­­ Head,­­­Thorax,­­­Abdomen:­­­ similar to male except ventrite II lacking median ovate depression. Distribution.­­­ This species is known from several localities in Auckland AK. Material­­­ Examined. AK: (1­­­ female) New Zealand: AK: Auckland, Lynfield Ground plants 6 Dec 1975, G Kuschel (LSAM); (1­­­female) New Zealand: AK: Auckland, Lynfield Ground plants 13 Dec 1975, G Kuschel (LSAM); (2­­­males,­­­1­­­slide-mounted) New Zealand:AK: Auckland, Lynfield Litter, 17 Oct 1975 G Kuschel, 75/155 (LSAM); (3­­­females) New Zealand: AK: Mt. Dome, litter 15 Mar 1978 SB Peck (LSAM); (6­­­males,­­­1­­­female) New Zealand: AK: Ponga Bush, Drury, nr. Pukekohe Litter, 4 Jan 1973 BM May, 73/1 (LSAM); (1male) ­­­ New Zealand AK (illegible) Gully 5 Nov 1985 R.C. Craw // Sifted litter (NZAC). Pselaphogenius delicatus ­­­(Broun),­­­1886 Pselaphus delicatus Broun (1886: 943), transferred (Jeannel 1951). (Fig. 10a–c) Type­­­Material­­­Examined.­­­ After examining the type series of Pselaphus delicatus in the NHM and specimens from the Broun collection in the NZAC, and reviewing Broun’s original description, the following specimen from the NZAC is here designated the lectotype (1­­­male)­­­ Near Howick, Auckland.// 1697.// Pselaphus delicatus, Broun, 1886 (NZAC) (lectotype). In Broun’s original description (1886) he mentioned “near Howick” as the locality, but did not indicate number of specimens in the type series. The manuscript number and locality of this specimen in the NZAC most closely matches that found in Broun’s original description. Diagnosis.­­­ Externally, P. delicatus is difficult to distinguish from other Pselaphogenius in New Zealand. The form of the aedeagus, with the two pairs of short, curved lateral processes and the spatula-like ventral process, is unique. Redescription.­­­ MALE. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes small, composed of five facets. Frontal margin of rostrum acute, with sparse suberect setae in dorsal view. Gular mound modified, longer than wide, narrowly rounded posteriorly; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV longer than head, narrow, widened at base and distal 1/5, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae present, median antebasal fovea reduced, pit like. Mesoventrite with median shield unmodified. Metaventrite afoveate, medially depressed, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, sutural fovea present; discal stria and sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, large shallow oval median depression not reaching apex. Genitalia: aedeagus asymmetrical. Median lobe distally displaced towards left, curved ventrad and slightly towards midline. Two pairs of lateral processes, more dorsal pair horn-like and curved inward, more ventral pair slightly expanded distally, also curved inwards, slightly. Ventral process straight and flattened, longer than median lobe, constricted distally to small, spatula-shaped extension. Parameres present, slightly longer than median lobe. Females.­­­ Integument,­­­ Head,­­­Thorax,­­­Abdomen:­­­ similar to male except ventrite II lacking median ovate depression. Distribution.­­­ This species is known from several localities in AK and ND. Material­­­ examined.­­­AK:­­­ (1­­­ male,­­­ 1­­­ female)­­­ New Zealand AK Kirks Bush Papakura 26.5.92// JT Nunn Collection (JN); (2­­­males) 1697.// ♂ // Hunua Maketu // New Zealand Broun Coll. Brit. Mus. 1922–482; (1­­­female) TYPE (red/orange label)// Paparoa // New Zealand Broun Coll. Brit. Mus. 1922–482; (1­­­ female) 1697.// ♀ // Hunua // New Zealand Broun Coll. Brit. Mus. 1922–482; (2­­­females)­­­ 1697.// ♀ // Hunua Maketu // // New Zealand Broun Coll. Brit. Mus. 1922–482; (1­­­male) 1697.// ♂ // Hunua Clevedon // // New Zealand Broun Coll. Brit. Mus. 1922–482; (1­­­male)­­­ NEW ZEALAND ND Waitangi SF 2 Nov 1981 G. Kuschel // Litter and decayed wood 81/117 (NZAC); (1­­­male) ­­­ Titirangi Wairakere Rg. Auckland // 1694// coll. A.E. Brookes 26.2-1915// Pselaphus delicatus Broun (NZAC); (2­­­males,­­­2­­­females) ­­­ Hunua Ra. Clevedon Auckland // Ex. Broun Collection // 1697// Pselaphus delicatus Broun (NZAC); ­­­(1­­­male)­­­ Lynfield, AK Tropicana Dr. 9.6.74 G. Kuschel // Pselaphus delicatus Broun, 1886 (NZAC); (5­­­males) ­­­ New Zealand AK Lynfield WB 2 May 1981 G. Kuschel // Pselaphus delicatus Broun, 1886 (NZAC); (1­­­male) ­­­ New Zealand AK Lynfield 4 Sep 1976 // ground plants// Pselaphus delicatus Broun, 1886 (NZAC); (1­­­male) ­­­ New Zealand AK Lynfield Wattle Bay 29 Jun 1980 G. Kuschel // Litter at hollow Metrosideros escelsa// Pselaphus delicatus Broun, 1886 (NZAC); (1­­­male) ­­­ New Zealand AK Omeru SR 15-22 Dec 1998 # 162 P. Paquin & N. Duperre Pit traps (NZAC); (2­­­males) ­­­ Titirangi, Waitakere Rg Auckland // 1697// Coll. A.E. Brookes 26.2.1915 // Pselaphus delicatus Broun (NZAC); (1­­­male,­­­1­­­female) ­­­ Ex. Broun Collection // Hunua Rg. Cleveden, Auckland // 1697.// Pselaphus delicatus Broun, 1886 // A.E. Brookes Collection (NZAC). ­­­, Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on pages 198-200, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382, {"references":["Broun, T. (1886) s. n. In: Manual of the New Zealand Coleoptera. Parts III & IV. Colonial Museum and Geological Survey Department, Wellington, pp. 745 - 973.","Jeannel, R. (1951) Sur la systematique des genres de la tribu Pselaphini Raffray (Coleoptera Pselaphidae). Revue Francaise d'Entomologie, 18, 5 - 11."]}

Pselaphaulax pelorus Owens & Carlton, sp. nov. (Fig. 8a–c) Type Material. Holotype: (male) New Zealand: MB: Pelorus Reserve Litter, 15 Sep 1964 JI Townsend, 64/93 (NZAC). Paratypes: (1 male, 2 females) holotype locality (NZAC). Etymology. This species is named after the type locality in Pelorus Reserve. Diagnosis. Of the species in group 1, only P. pelorus possesses a ventral process with serrations on the dorsal surface. Additionally, the elongate, curved processes associated with the elongate median lobe (4/5 length of basal bulb) are unique. Description. MALE holotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 19 facets. Frontal margin of rostrum in dorsal view straight with sparse suberect setae. Gular mound modified, ovate, narrowed posteriorly and anteriorly, longer than wide; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, apical swelling roughened, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus distinct, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, single sutural fovea; single discal stria, sutural striae present; sparse apical fringe of setae present. Abdomen: tergites of usual form for tribe. Ventrite II with median area slightly flattened throughout length of ventrite, apically with row of small tubercles. Genitalia: aedeagus asymmetrical. Median lobe broad, about 4/5 length of basal bulb; associated processes narrow, elongate, curved. Ventral process serrate on dorsal surface, about as long as median lobe. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median ovate depression and thickened area at apex. Distribution. This species is known from the Pelorus and Wakamarina Valleys in Marlborough (MB) and Cobb Reservoir and Dun Mt. in Nelson (NN). Material examined. Marlborough region: (2 males) New Zealand: MB: Pelorus Bridge Litter, 15 Sep 1964 JI Townsend, 64/95 (LSAM); (1 male) New Zealand: MB: Pelorus Valley, 214 m Moss, 13 Jul 1965 JI Townsend, 65/388 (LSAM); (1 male) New Zealand: MB: Wakamarina Valley Litter, 12 Aug 1966 AK Walker, 66/237 (LSAM); (1 male) New Zealand: MB Ronga Saddle Moss, 13 Jul 1965 LPMarchant, 65/391(LSAM). Nelson region: (1 female) New Zealand: NN: Cobb Reservoir, 1037 m Moss, 18 Sep 1964 TG Wood, 64/100 (LSAM); (1 male) New Zealand: NN: Dun Mt., Third House 560 m, moss, 66/121 29 Mar 1966, JI Townsend (LSAM); (1 male) NEW ZEALAND: So. Island, 70 km NE Nelson, Tennyson Inlet 480m, 27-v-1982, FMHD #82-604, Beech forest litter, S. Peck (FMNH, EtOH)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, Revision of the New Zealand species of the Genus Pselaphaulax Reitter (Staphylinidae: Pselaphinae: Pselaphitae: Pselaphini), pp. 221-244 in Zootaxa 5155 (2) on page 234, DOI: 10.11646/zootaxa.5155.2.3, http://zenodo.org/record/6673412

Pselaphogenius dismukesi ­­­Owens­­­&­­­Carlton,­­­sp.­­­nov. (Fig. 21a–c) Type­­­Material.­­­ Holotype:­­­(1­­­male)­­­ New Zealand: ND: Waipoua SF Te Matua Ngahere, litter, 4 Feb 1975 JC Watt, 75/94 (NZAC). Paratypes:­­­(1­­­male,­­­3­­­females)­­­ holotype locality (NZAC). Etymology.­­­ This species is named in honor of Dr. David Dismukes, for his support of this project as member of the first author’s advisory committee. Diagnosis.­­­ Externally, P. dismukesi is similar to other species in the genus. The form of the aedeagus, with a median lobe enlarged, ventral process simple and two lateral processes simple and curved dorsally is unique. Redescription.­­­ MALE. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 15 facets. Frontal margin of rostrum straight in dorsal view. Gular mound modified, as wide as long, broad anteriorly and rounded posteriorly; area of head posterior to gular mound flattened, slightly concave, nude, maxillary palpomere IV as long as head, narrow, widened at base and distal ½ to 1/3, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae present, median antebasal fovea reduced. Mesoventrite with median shield unmodified. Metaventrite afoveate, medially flattened, bearing short, fine setae, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae and single sutural fovea present; discal stria absent, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, oval median depression shallow, reaching apex. Genitalia: aedeagus symmetrical. Median lobe enlarged, produced in thickened extension from phallobase. Lateral processes curved dorsally, narrowed and simple. Ventral process ventrally curved, distally, longer than median lobe. Parameres present, as long ventral process. Females.­­­Integument,­­­Head,­­­Thorax,­­­Abdomen:­­­ Females are similar to males except eyes slightly smaller and ventrite II lacking median ovate depression. Distribution.­­­ This species is only known from collection events in Northland (ND) and Westland (WD). Material­­­examined.­­­(18­­­males,­­­25­­­females)­­­ New Zealand: ND: Waipoua Stm, 100m Litter 19 May 1978, SB Peck (LSAM); (3­­­males,­­­5­­­females) New Zealand: ND: Omahutu S.F. Litter, 18 Mar 1978 SB Peck (LSAM); (5­­males,­­­7­­­females) ­­­ New Zealand: ND: Waipoua S.F. Litter, 4 Feb 1975 SE Nichols, 75/84; (1­­­male,­­­1­­­female)­­­ New Zealand: ND: Waipoua S.F., Te Matua Ngahere, litter, 75/19 4 Feb 1975, JC Watt (LSAM); (10­­­males,­­­5­­­females) New Zealand: WD: Hokitika Gorge 19 Jan 1978 SB Peck (LSAM); (6­­­males,­­­6­­­females) New Zealand: WD: Okuku Scenic reserve 46.7 km W Otira, 75 m 42’43S 171’14E, #050// zelandiae leaf litter Berlese, 12 Jan 1998 C.Carlton, R.Leschen (LSAM)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on pages 215-216, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382

Pselaphogenius ventralis ­­­(Broun),­­­1895 Pselaphus ventralis Broun (1895: 72), transferred (Jeannel 1951). (Figs. 5c, 19a–c) Type ­­­Material­­­Examined.­­­ After examining the type series of Pselaphus ventralis in the NHM and reviewing Broun’s original description, the following specimen from the NHM is here designated the lectotype: (1­­­male) TYPE (red/orange label)// 2742.// Mount Pirongia // New Zealand Broun coll. Brit. Mus. 1922-482 (lectotype). The additional specimen in the syntype series is also a member of P. ventralis: (1­­­male) 2742.// ♂ // Mount Pirongia // New Zealand Broun coll. Brit. Mus. 1922-482 and is here designated a paralectotype. Diagnosis.­­­ Externally, this species is most similar to P. calantica, which also bears a thickened region at the apex of ventrite II. The greatly reduced form of the aedeagus, with the median lobe truncate and the absence of lateral and ventral processes, is unique. Redescription.­­­ MALE. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes small, composed of five facets. Frontal margin of rostrum emarginate in dorsal view. Gular mound swollen, ovate, longer than wide, narrowed anteriorly and posteriorly, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV longer than head, narrow, widened at base and distal 1/4, apical area lightly tuberculate, v-shaped sensory patches reduced. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae and median antebasal fovea reduced, pit like. Mesoventrite with median shield unmodified. Metaventrite afoveate, with median depression, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, sutural fovea present; discal stria absent, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying width of ventrite, sulcus densely setose, large deep oval median depression not reaching ventrite apex, apical portion of depression thickened into small, lateral ridge. Genitalia: aedeagus symmetrical. Median lobe shortened, truncate. Lateral processes absent. Ventral process absent. Parameres present, slightly exceeding length of median lobe. Females.­­­ Integument,­­­ Head,­­­Thorax,­­­Abdomen:­­­ similar to male except ventrite II lacking median ovate depression. Distribution.­­­ This species is known from localities in: Auckland (AK), Hunua Range, Waipipi Scenic Reserve, and the Waitakere Range; Bay of Plenty (BP), Kaimai-Mamaku Range; Nelson (NN), Kahurangi N.P.; Waikato (WO), Pirongia Forest Park and Te Mata. Material­­­examined.­­­(2­­­males)­­­ NEW ZEALAND: AK: Hunua Ranges, Vining Reserve, Mangatangi Track, 240m, 37 ∘ 0.7.747/S, 175 ∘ 13.13.024/E, 17.xi.2005, Nothofagus-Agathis-Phyllocladus trichomanoides, etc.; FMHD#2005-008, berl. Leaf & log litter, A. Newton, A. Solodovnikov, et al.; ANMT site 1141 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (3­­­males,­­­2­­­females)­­­ NEW ZEALAND: AK: Waipipi Scen. Res., 2km. Wsw Waipipi 40m 37.14S 174.40E 22.ii.1987 native hardwood forest in gully A.Newton & M.Thayer 835 FMHD #87-269 berl., leaf & log litter, forest floor (FMNH, EtOH); (4­­­males,­­­3­­­females)­­­ NEW ZEALAND: AK: Waitakere Ra., Nohoanga Scen. Res., 260m 36.57S 174.35E 23.ii.1987 hardwood-podocarp forest A.Newton & M.Thayer 836 FMHD #87-271 berl., frond litter at base of tree ferns. (FMNH, EtOH); (4­­­males,­­­2­­­females)­­­ NEW ZEALAND: BP: Kaimai-Mamaku Forest Park, Mt. Te Aroha, upper end Tul Mine Track, nr. Summit road, 775m, 37 ∘ 31.658'S, 175 ∘ 44.684'E, 19.xi.2005, low Nothofagus menziesii forest w/ Astelia ground layer; FMHD#2005-021, berl., forest litter, A. Newton & M. Thayer; ANMT site 1145 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (3­­males,­­­4­­­females)­­­ NEW ZEALAND: NN: Kahurangi N.P., Cobb Dam Rd., Asbestos Track 450m, 41 ∘ 06.333'S, 172 ∘ 43.174'E, 18.xii.2005, mixed broadleaf (incl. Nothofagus fusca)- podocarp forest; FMHD#2005-111, berl., leaf & log litter. A. Solodovnikov, D. Clarke, et al.; ANMT site 1160 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (1­­­male,­­­1­­­female)­­­ NEW ZEALAND: NN: Kahurangi N.P., Arthur Range, above Flora Saddle, 1000m, 41 ∘ 11.351'S, 172 ∘ 44.456'E, 28.xi-19.xii.2005, Nothofagus -dominant forest; FMHD#2005-044, flight intercept trap, A. Newton & M. Thayer, ANMT site 1156 propylene glycol to 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (2­­­males,­­­2­­­females)­­­ NEW ZEALAND: WO: Pirongia Forest Park, Tirohanga Track (above end Corcoran Rd.), 500m, 37 ∘ 58.409'S, 175 ∘ 07.173'E, 20.xi.2005, broadleaf forest; FMHD#2005-015, berl., forest litter, D. Clarke & A. Solodovnikov; ANMT site 1146 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (3­­­males,­­­5­­­females)­­­ NEW ZEALAND: WO: Pirongia Forest Park, Mahaukura Track (above end Grey Rd.), 270m, 37 ∘ 58.218'S, 175 ∘ 06.523'E, 18.xi.2005, mixed broadleaf forest; FMHD#2005-011, berl., leaf & log litter, M. Thayer & A. Newton; ANMT site 1142 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (1­­­male,­­­1­­­female)­­­ NEW ZEALAND WO Te Mata Bridal Veil Falls 19 Sep 1981 C.F.Butcher// Litter 81/88 (NZAC)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on pages 210-212, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382, {"references":["Broun, T. (1895) Descriptions of new Coleoptera from New Zealand. Annals and Magazine of Natural History, Series 6, 15, 1 - 419. https: // doi. org / 10.1080 / 00222939508677906","Jeannel, R. (1951) Sur la systematique des genres de la tribu Pselaphini Raffray (Coleoptera Pselaphidae). Revue Francaise d'Entomologie, 18, 5 - 11."]}

Pselaphaulax tararua Owens & Carlton, new species (Fig. 11a–c) Type Material. Holotype: (male) NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 990m 13 Feb 1985 // C.F. Butcher Litter 85/16 (NZAC). Paratypes: (2 males, 3 females) holotype locality (NZAC). Etymology. This species is named after the type locality in the Tararua Range. Diagnosis. This species can be readily be distinguished from all others in the genus by the curvature of the median lobe to the right and the posterior direction of the terminal end. Description. MALE holotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 19 facets. Frontal margin of rostrum in dorsal view straight, weakly emarginate with sparse suberect setae. Gular mound swollen, ovate, longer than wide, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, apical swelling roughened, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus faint, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate, raised into median tumosity; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, single sutural fovea present; single discal stria and sutural striae present; apical fringe of setae absent. Abdomen: tergites of usual form for tribe. Ventrite II with narrow, flattened median depression, extending length of ventrite. Genitalia: aedeagus asymmetrical. Median lobe distally curved far to the right, elongate, terminal end directed posteriorly. Right lateral process well-developed, narrowed distally. Ventral process downward curved, distally, shorter than median lobe. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median ovate depression and thickened area at apex. Distribution. This species is known from collecting events in Nelson (NN), Taranaki (TK), Taupo (TO), and Wellington (WN). Its distribution is split across the North and South islands of NZ, but it has not been collected farther south than the extreme northern tip of the South Island. Material examined. (2 males, 1 female) New Zealand: NN: Stanley Brook, Sunday Creek Litter, 11 Oct 1963 JI Townsend, 63/27 (LSAM); (2 males) NEW ZEALAND: So. Island, 20 km NE Takaka, Tasman Nat. Pk., 21-v-1982, FMHD #82-591, Mixed forest litter, S. Peck (FMNH); (1male) New Zealand: TK: Mt. Egmont Nat. Park 29 Nov 1970 AW Don (LSAM); (1 male) NEW ZEALAND TO Chakune Mtn Rd Waitonga Falls 4 Nov 2004 A C Eyles ground moss (NZAC); (3 males, 3 females) NEW ZEALAND: WN: Tararua Forest Park, above Akatarawa Saddle, 455m, 40 ∘ 56.936'S, 175 ∘ 06.529'E, 26.ix.2005, broadleaf-podocarp forest on slope; FMHD#2005-033, berl., leaf & log litter, A. Newton & M. Thayer, ANMT site 1151 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH); (3 males, 2 females) NEW ZEALAND: WN: Tararua Forest Park, Judd Ridge, Field's Track, Field Hut vic., 855m, 40 ∘ 54.474'S, 175 ∘ 15.371'E, 24.xi-26.xi.2005, broadleaf (Noth. Menziesii-Weinmannia racemosa) -podocarp forest; FMHD#2005-039, berl., forest leaf litter (mostly Nothofagus), A. Solodovnikov & D. Clarke; ANMT site 1153 95% EtOH FIELD MUSEUM NAT. HIST. (FMNH); (1 female) NEW ZEALAND, WN, Kaitoke Regional Park, Pakuratahi Forks, 15 Apr 2005, R. Leschen, C. McGuiness,// leaf litter, RL975, 41.03'S, 175.11'E (NZAC); (1 male) NEW ZEALAND WN Tararua Ra Logan East Basin 12 Feb 1985 C.F.Butcher // Mixed swards 85/27 (FMNH). NEW ZEALAND, WN Tararua Forest Park off Waiotauru Rd 25 Aug 1998 E. Spur // found on bait (NZAC); (1 male, 1 female) NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 950m 3 Dec 1984 // R.C.Craw Sifted litter 84/88 (NZAC); (1 male) NEW ZEALAND WN Tararua Ra Ridge to Dundas Hut 1275m 6 Dec 1984 // R.C.Craw Moss and plants 84/98 (NZAC); (1 male, 2 females) NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 800m 13 Feb 1985 // G.W.Ramsay Litter 85/18 (NZAC); (1 female) NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 990m 3 Dec 1984 // B. G. Bennett & T. K. Crosby Sifted litter 84/83 (NZAC); (1 male) NEW ZEALAND WN Tararua Ra Dundas Hut 1250m 6 Dec 1984 J.S. Dugdale // Moss,plants 84/93 (NZAC); (1 male) NEW ZEALAND WN Tararua Ra Dundas Hut Ridge, River Forks 900m 11 Feb 1985 // G.W.Ramsay Litter 85/12 (NZAC); (1 male, 3 females) NEW ZEALAND WN Wainuiomata Whakanui Tk 10 Jan 1993 J R Grehan // Beech /kamahai (NZAC)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, Revision of the New Zealand species of the Genus Pselaphaulax Reitter (Staphylinidae: Pselaphinae: Pselaphitae: Pselaphini), pp. 221-244 in Zootaxa 5155 (2) on page 238, DOI: 10.11646/zootaxa.5155.2.3, http://zenodo.org/record/6673412

Pselaphaulax Reitter 1909 Pselaphaulax Reitter (1909: 218) Type species: Pselaphus dresdensis Herbst, 1792 (monot.) Redescription (based on Owens et al. 2019). Males. Head: eyes rounded, composed of 15–20 facets. Vertex convex; shallow sulcus beginning at base of vertex, extending to level of vertexal foveae; vertexal foveae large, set into margins of vertexal depression parallel to middle of eyes; vertexal depression widest between eyes, confluent with sulcus extending to apex of rostrum. Rostral sulcus bounded by lateral, flattened ridges extending to antennal acetabula. Frontal margin of rostrum in dorsal view typically straight to slightly emarginate, with sparse, suberect setae. Antennal segment one as long as antennomeres 2-4, antennae otherwise unmodified. Gular mound modified, hemispherical, about as long a wide or narrowed and longer than wide, glabrous; area of head posterior to gular mound flattened, slightly concave, sometimes bearing flattened modified scales; gular foveae present, paired. Maxillary palpomere I elongate, 1/2 as long as palpomere II; palpomere II narrow, enlarged distally, shorter than head; maxillary palpomere III quadrate; maxillary palpomere IV as long as head, widened in distal 1/3-1/2, apical area smooth to roughened, v-shaped sensory patch present, margined by carina. Thorax: Pronotum widest at 1/2 length, narrowed anteriorly and posteriorly; basal sulcus present, delimited by lateral antebasal foveae, median antebasal fovea reduced to depression, margined by two short carinae. Prosternum in front of coxae slightly convex, anteriorly with modified flattened setae; lateral procoxal foveae present. Mesoventrite with median shield; lateral mesoventral foveae present in large cavities filled with dense setae, meeting internally; lateral mesocoxal foveae absent; median mesoventral foveae present, paired, meeting internally; mesocoxae separated by extension of meso- and metaventrite. Metaventrite afoveate, variously modified; metacoxae separated by extension of ventrite I. Elytra with three basal foveae, single sutural fovea; single discal stria, sutural striae present; apical fringe of setae present. Abdomen: Tergites of usual form for tribe, unmodified; tergite 2 without obvious basal foveae, lateral basal sulcus extending along width of ventrite, obscured by dense band of setae along posterior edge. Ventrite I covered in dense, shining setae; ventrite II variously modified with median depressions, sulci, or projections; ventrites otherwise unmodified. Females: similar to males except eyes may be smaller, composed of fewer facets; secondary sexual modifications of the legs and second ventrite absent. Comments. NZ species in this genus are primarily united by the presence of an antebasal sulcus on the pronotum margined by lateral antebasal foveae, the median antebasal foveae present and sometimes delimited by two small carinae (may be reduced to a median pit), maxillary palpi IV widened in the distal 1/3 to1/2 with a v-shaped sensory patch margined by a carina, and the gular swelling prominent, about as long as wide or narrower and slightly longer than wide. While several groups in NZ likely representing undescribed genera may superficially resemble Pselaphaulax, no other groups possess a combination of these character states. Pselaphaulax is most similar to Pselaphotumulus and Pselaphogenius, but can quickly be distinguished, most noticeably, by the rounded to slightly emarginate apical margin of the rostrum (pointed in Pselaphotumulus and some Pselaphogenius), the gular process rounded or ovate-rounded (greatly expanded to cover the entire gular area of the head in Pselaphotumulus), and the presence of an antebasal sulcus on the pronotum (absent in Pselaphogenius). Superficially, NZ specimens in the genus can be divided into two “species groups” which can consistently be distinguished by several characters, including: shape of gular mound (ovate, narrowed apically and posteriorly in group 1, rounded, widened apically and rounded or narrowed posteriorly in group 2); modifications of the area behind gular mound (nude in group 1, covered in flattened modified scales in group 2); the degree of reduction of the median antebasal fovea (present and margined by carinae in group 1, reduced and pit-like in group 2); and modifications of the mesoventral shield (bearing a single, setose fovea in group 1, lacking setose fovea in group 2). All species described in the previous Chatham Islands manuscript (Owens et al. 2019) are members of group 2. Pselaphaulax intermedius possesses a mixture of characters (gular mound rounded posteriorly as in group 2, area behind gular mound nude as in group 1, median antebasal fovea present as in group 1, and mesoventral shield bearing a setose fovea as in group 2). Further investigation and comparison with Asian fauna may eventually warrant the designation of subgeneric status to these species groups within NZ Pselaphaulax, but under the current system they should be considered informal designations encompassing morphological variation within the genus., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, Revision of the New Zealand species of the Genus Pselaphaulax Reitter (Staphylinidae: Pselaphinae: Pselaphitae: Pselaphini), pp. 221-244 in Zootaxa 5155 (2) on page 223, DOI: 10.11646/zootaxa.5155.2.3, http://zenodo.org/record/6673412, {"references":["Owens, B. E., Leschen, R. A. B. & Carlton, C. E. (2019) The Pselaphini (Coleoptera: Staphylinidae: Pselaphinae) of the Subantarctic and Chatham Islands of New Zealand; description of five species and one genus. Insect Systematics and Diversity, 3, 1 - 14. https: // doi. org / 10.1093 / isd / ixz 011"]}

Pselaphaulax meliusculus (Broun), 1893, comb. nov. Pselaphus meliusculus (Broun 1893: 1044) (Figs. 1 –2, 3a–b, 4a–b, 7a–c) Type Material Examined. The following specimen from the NHM is here designated the lectotype of Pselaphus meliusculus: (male) TYPE (red/orange label)// 1868.// Dunedin// New Zealand Broun coll. Brit. Mus. 1922-482 (lectotype). Broun’s original description does not mention a single type specimen, and the manuscript number (1868.) matches the number included in Broun’s description (Broun, 1893 a). Likewise, the locality of the specimen, Dunedin, is near the locality, Mount Maungatua, Otago, mentioned in Broun’s original description. Diagnosis. Pselaphaulax meliusculus can be distinguished from other species in the genus by the curvature of the median lobe to the right and the projection of a dorsal process over the median lobe. Redescription. MALE Lectotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 22 facets. Frontal margin of rostrum in dorsal view straight with tuft of sparse suberect setae. Gular mound modified, broadened anteriorly, as long as wide, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/2 to 1/3, distal swollen area smooth, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus faint, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate, lateral margins of depression with short erect setae; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with single basal fovea present, single sutural fovea; single discal stria and sutural striae present; apical fringe of setae present. Abdomen: Tergites of usual form for tribe. Ventrite II flattened in narrow band along length of ventrite, flattened area margined by fine, short setae in some specimens. Genitalia: aedeagus asymmetrical. Dorsal process extended over median lobe, projected ventrally. Median lobe distally curved to the right. Ventral process curved ventrad distally, about as long as median lobe. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median ovate depression and thickened area at apex. Distribution. Pselaphaulax meliusculus is known from Central Otago (CO), Dunedin (DN), Fiordland (FD), and Southland (SL) regions on South Island. Material examined. Central Otago region: (1 male) New Zealand CO Leaning Lodge Rock & Pillar Rng 25-Jan-02 (JN) ; (1 male, 1 female) New Zealand CO Ailsa Craig Lammermoor Rng 7-Dec-08// Sifted moss and tussock litter// J T Nunn collection (JN); (1 male, 1 female) New Zealand CO Carrick Ra Watts Rock 8 ii 2015 // Sifted moss and tussock litter J. Nunn (JN). Dunedin region: (1 male) New Zealand DN Flagstaff Hill 8-Oct-03// in tussock litter (JN); (1 male) NEW ZEALAND DN Flagstaff Hill Dunedin 23-IV-2011 // in tussock litter (JN); (2 males, 1 female) NEW ZEALAND: DN: Mt. Cargill Scen. Res., Organ Pipes Track, nr. Mt. Cargill summit, 600m, 45 ∘ 48.505'S, 170 ∘ 33.999'E, 5.xii.2005, podocarp-broadleaf forest; FMHD#2005-076, berl., leaf litter (wet), A. Solodovnikov & D. Clarke; ANMT site 1165 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (1 male) Dunedin 1408/15/1952 R.Hornabrook (NZAC). Fiordland region: (1 male) New Zealand FD Borland Saddle 24-Jan-08 (JN); New Zealand FD Thicket Burn L Hauroko 04-Jan-96// Nothofagus litter (JN); (1 male) New Zealand FD Mount Burns 18-Feb-07// Tussock litter (JN); (1 male) New Zealand FD First Bay L Hauroko 25 Sep 09// Washed soil sample, beech forest// J T Nunn collection (JN); (1 male) NEW ZEALAND FD Hump Ridge 914m 8 Feb 1968 J. I. Townsend Litter 68/9// New Zealand Arthropod Collection, NZAC Private Bat 92170 AUCKALND New Zealand (NZAC). Southland region: (3 males, 1 female) New Zealand SL Manuka Ridge Rd Blue mountains 9-Aug-03// in ground litter and litter from tussock zone// J T Nunn collection (JN); (1 male) New Zealand SL MW relat stn Blue Mountains 19-Mar-06// J T Nunn collection (JN); (1 male) New Zealand SL Round Hill Orepuki 6- Jul-02// In forest floor litter// J T Nunn collection (JN); (1 male) New Zealand SL Spence Basin Takitimu Rng 16- Jan-00// c. 1,150 m // J T Nunn (JN); (1 male) New Zealand SL Mores SR Riverton 6-Jul-02// In ground litter from kamahi forest// J T Nunn (JN); (1 male, 1 female) New Zealand SL Kaka Point SR 5-Jun-99// In kamahi forest floor litter (JN); (1 male) NEW ZEALAND SL Longwood Range 8 Feb 1976 G.W. Ramsay litter 76/36// Entomology Division D.S.I.R. New Zealand (NZAC).

Pselaphaulax nunni Owens & Carlton, new species (Fig. 12a–c) Type Material. Holotype: (male) New Zealand: MB Black Birch Ra summit 7 Feb-07// JTN collection (NZAC). Paratype: (1 female) holotype locality (NZAC). Etymology. This species is named in honor of John T. Nunn, an avid supporter of Pselaphinae research in New Zealand, who collected the type series. Diagnosis. This species can be distinguished from all other in the genus by the form of the deep ovate depression on the male ventrite II. Additionally, the absence of a setose pit on the mesoventral shield places this species in “species group 2”, although the absence of setae on the gular area behind the hemispherical swelling sets it apart from others in this division. The aedeagus, with a reduced median lobe and asymmetrical production of the right lateral process and absence of a left lateral process is also unique within the genus. Description. MALE. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 19 facets. Frontal margin of rostrum in dorsal view weakly emarginate with sparse suberect setae. Gular mound modified, hemispherical, as long as wide, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, distal swollen area smooth, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus faint, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with mesoventral shield unmodified. Metaventrite afoveate, flattened and bearing vestiture of sparse short setae; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, single sutural fovea; discal stria and sutural striae present; sparse apical fringe of setae present. Abdomen: tergites of usual form for tribe. Ventrite II with deep ovate depression occupying length of ventrite. Genitalia: aedeagus asymmetrical. Median lobe reduced. Right lateral process developed, compressed laterally. Ventral process absent. Parameres present, well-developed (longer than length of basal bulb). Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median ovate depression and thickened area at apex. Distribution. Pselaphaulax nunni is only known from the type locality., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, Revision of the New Zealand species of the Genus Pselaphaulax Reitter (Staphylinidae: Pselaphinae: Pselaphitae: Pselaphini), pp. 221-244 in Zootaxa 5155 (2) on pages 240-241, DOI: 10.11646/zootaxa.5155.2.3, http://zenodo.org/record/6673412

Pselaphogenius citimus ­­­(Broun),­­­1893 Pselaphus citimus Broun (1893: 1043), transferred (Jeannel 1951: 9) (Figs. 5d, 20a–c) Type­­­Material­­­Examined.­­­ After examining the type series of Pselaphus citimus in the NHM and reviewing Broun’s original description, the following specimen was determined to be the holotype: (1­­­male) TYPE (red/orange label)// 1867.// Howick// New Zealand Broun coll. Brit. Mus. 1922-482 (Holotype). This matches the manuscript number, physical description, and locality information provided in the original description (Broun 1893). Diagnosis.­­­ Externally, males of this species can be distinguished from other NZ species by the modification of ventrite II bearing a flattened median area instead of an ovate depression. The aedeagus is also distinct, with a large, shortened median lobe bearing distal complex of processes, a short straight ventral process and the absence of lateral processes. Redescription.­­­ MALE. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 15 facets. Frontal margin of rostrum slightly rounded in dorsal view. Gular mound modified, wider than long, broad anteriorly and rounded posteriorly; area of head posterior to gular mound flattened, slightly concave, bearing flattened scale-like setae. Maxillary palpomere IV as long as head, narrow, widened at base and distal ½ to 1/3, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae reduced to pits, median antebasal fovea absent. Mesoventrite with median shield unmodified. Metaventrite afoveate, raised into tumosity bearing small patches of short setae, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with one basal fovea and single sutural fovea present; discal stria present, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, median flattened area along length of ventrite. Genitalia: aedeagus symmetrical. Median lobe protruding from fleshy extension of phallobase, produced into distal complexities. Ventral process simple, straight, shorter than median lobe. Lateral processes absent. Parameres broad and extending beyond median lobe. Females.­­­Integument,­­­Head,­­­Thorax,­­­Abdomen:­­­ Females are similar to males except eyes slightly smaller and ventrite II lacking median flattened area. Distribution.­­­ Pselaphogenius citimus is known from a large number of localities in the north island, including sites in Auckland (AK), Bay of Plenty (BP), Coromandel (CL) Gisborne (GB), Hawkes Bay (HB), Northland (ND), Taranaki (TK), Taupo (TO), and Waikato (WO). This species is also known from Nelson (NN) on the South Island. Material­­­examined.­­­(1­­­male)­­­ New Zealand TO Waituhi Sdl 1.9.93// In leaf litter// JT Nunn Collection (JN); ­­­ (1­­­male,­­­4­­­females)­­­ New Zealand: AK: Hunua Ra., 150 m Litter 4 Jan 1973 BM May, 73/2 (LSAM); (­­­2­­­males,­­­ 1­­­female)­­­ New Zealand: AK: Hunua Ra., Kohukohunui, 550 m Litter, 30 Mar 1974 G Kuschel, 74/19 (LSAM); (4­­­males,­­­2­­­females)­­­ New Zealand: AK: Hunua Ra., Kohukohunui, 600 m Litter, 30 Mar 1974 G Kuschel, 74/20 (LSAM); (3­­­ males,­­­ 1­­­ female)­­­ New Zealand: AK: Mt. Dome, litter 15 Mar 1978 SB Peck (LSAM); (3­­­ males,­­­ 1­­­ female)­­­ New Zealand: AK: Ponga Bush, Drury, nr. Pukekohe Litter, 4 Jan 1973 BM May, 73/1 (LSAM); (1­­female)­­­ New Zealand: AK: Ponga Bush, Drury, nr. Pukekohe Ground plants 13 Dec 1975, G Kuschel (LSAM); (1­­­female)­­­ New Zealand: GB: Urewera, Orangihikoia Stm., litter, 6 Feb 1970 AW Don, 68/37 (LSAM); (1­­­male)­­­ New Zealand: GB: Urewera N.P., near Aniwaniwa Visitor Center, Sifting toetoe litter, 21 Mar 2000 C Carlton, A Weir, #074 (LSAM); (3­­­males,­­­4­­­females)­­­ New Zealand: GB: Urewera N.P., Lake Waikareiti 38 ∘ 47S 176 ∘ 10E / Dracophyllum / Nothofagus litter berlese 19 Mar 2000 C Carlton, A Weir, #058 (LSAM); (1­­­male,­­­1­­­female)­­­ New Zealand: HB: Kaweka Ra., Makahu Hut 976 m, litter, 24 Feb 1971 JI Townsend, AC Eyles 71/21 (LSAM); (4­­females)­­­ New Zealand: HB: Kaweka Ra., Makahu Spur 1036m, moss, 24 Feb 1971 JI Townsend, 71/23 (LSAM); (4­­­males,­­­1­­­female)­­­ New Zealand: ND: Waipoua Stm., 100 m Litter 19 May 1978, SB Peck (LSAM); (1­­­male)­­­ New Zealand: TK: Awakino Gorge, Kuriwera Stm, litter, 23 Jan 1972 GW Ramsay, 72/65 (LSAM); (1­­­female)­­­ New Zealand: TK: Dawson Falls, 975 m Mt. Egmont, litter, 65/387 16 Jun 1965, JI Townsend (LSAM); (3­­­females)­­­ New Zealand: TK: Mt Egmont, Dawson Falls 946 m, litter, 62/1 14 Jan 1962, G Kuschel (LSAM); (­­­1­­­male,­­­1­­­female)­­­ New Zealand: TO: Whirinaki Forest Park End of Okahu Rd. FIT, 23 Mar 2000 C Carlton, A Weir,#083 (LSAM); (3­­males,­­­3­­­females)­­­ NEW ZEALAND: AK: Hunua Ranges, Vining Reserve, Mangatangi Track, 240m, 37 ∘ 0.7.747/S, 175 ∘ 13.13.024/E, 17.xi.2005, Nothofagus-Agathis-Phyllocladus trichomanoides, etc.; FMHD#2005-008, berl. Leaf & log litter, A. Newton, A. Solodovnikov, et al.; ANMT site 1141 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (1­­­male)­­­ NEW ZEALAND: AK: Hunua Ranges Reg. Park, Kohukohunui Track nr. Mine Rd., 475m, 37 ∘ 02.396'S, 175 ∘ 11.251'E, 17.xi.2005, broadleaf-podocarp forest, numerous tree ferns; FMHD#2005-004, berl., leaf & log litter, A. Newton, A. Solodovnikov, et al.; ANMT site 1140 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (1­­­ male)­­­ NEW ZEALAND: AK: Waipipi Scen. Res., 2km. Wsw Waipipi 40m 37.14S 174.40E 22.ii.1987 native hardwood forest in gully A.Newton & M.Thayer 835 FMHD #87-269 berl., leaf & log litter, forest floor (FMNH, EtOH); (5­­­males,­­­3­­­females)­­­ NEW ZEALAND: AK: Waitakere Ra., Nohoanga Scen. Res., 260m 36.57S 174.35E 23.ii.1987 hardwood-podocarp forest A.Newton & M.Thayer 836 FMHD #87-271 berl., frond litter at base of tree ferns. (FMNH EtOH); (1­­­male)­­­ NEW ZEALAND: BP: Kaimai-Mamaku Forest Park, Mt. Te Aroha summit road, 450m, 37 ∘ 31.429'S, 175 ∘ 44.005'E, 19.xi.2005, mixed broadleaf forest w/many tree ferns, nikau palms; FMHD#2005-018, berl., leaf & log litter, A. Newton & M. Thayer; ANMT site 1144 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (5­­­males)­­­ NEW ZEALAND: BP: Kaimai-Mamaku Forest Park, Mt. Te Aroha, upper end Tul Mine Track, nr. Summit road, 775m, 37 ∘ 31.658'S, 175 ∘ 44.684'E, 19.xi.2005, low Nothofagus menziesii forest w/ Astelia ground layer; FMHD#2005-021, berl., forest litter, A. Newton & M. Thayer; ANMT site 1145 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (21­­­males,­­­7­­­females)­­­ NEW ZEALAND: GB: Urewera N.P., Waikaremoana Rd., S end Matanunui Ridge, 720m, 38 ∘ 44.404'S, 177 ∘ 05.806'E, 22.xii.2005, mixed broadleaf (incl. Nothofagus fusca)- podocarp; FMHD#2005-118, berl., leaf & log litter, A. Newton, M. Thayer, & A. Solodovnikov; ANMT site 1149 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (2­­­males,­­­3­­­females)­­­ NEW ZEALAND: GB: Urewera N.P., Maungapohatu Rd., 3.2 km E Taupeupe Saddle, 1110m, 38 ∘ 36.975'N, 177 ∘ 02.753'E, 22.xi.2005, mossy Nothofagus menzeisii forest; FMHD#2005-027, berl., leaf & log litter, A. Newton, M. Thayer, et al.; ANMT site 1148 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (3­­­males,­­­1­­­female)­­­ NEW ZEALAND: NN: Kahurangi N.P., Arthur Range, above Flora Saddle, 1000m, 41 ∘ 11.351'S, 172 ∘ 44.456'E, 28.xi-19.xii.2005, Nothofagus -dominant forest; FMHD#2005-044, flight intercept trap, A. Newton & M. Thayer, ANMT site 1156 propylene glycol to 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (12­­­males,­­­13­­­females)­­­ NEW ZEALAND: WO: Pirongia Forest Park, Tirohanga Track (above end Corcoran Rd.), 500m, 37 ∘ 58.409'S, 175 ∘ 07.173'E, 20.xi.2005, broadleaf forest; FMHD#2005-015, berl., forest litter, D. Clarke & A. Solodovnikov; ANMT site 1146 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (2­­­males)­­­ NEW ZEALAND: WO: Pirongia Forest Park, Mahaukura Track (above end Grey Rd.), 270m, 37 ∘ 58.218'S, 175 ∘ 06.523'E, 18.xi.2005, mixed broadleaf forest; FMHD#2005-011, berl., leaf & log litter, M. Thayer & A. Newton; ANMT site 1142 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH); (1­­­male)­­­ NEW ZEALAND AK Birkenhead Kauri Park 4 Oct 1978 B.M.May (NZAC); (1­­­male)­­­ Waitakere Drive 15.10.42// A.E.Brookes Collection (NZAC); (6­­­ specimens)­­­ Waimarino, Main Trunk Line. N.Is.// 1867// Coll. T. Broun (NZAC); (1­­­ male,­­­ 1­­­ female)­­­ NEW ZEALAND BP Tikitiki Stream Horohoro SF Mamaku Plateau 24 Jul 1076 // J.S. Dugdale litter 76/57 (NZAC); (1­­male)­­­ NEW ZEALAND BP Horohoro SF Mamaku Plat 550m 24 Jul 1976 J.S.Dugdale// Litter 76/44 (NZAC); (2­­males,­­­1­­­female)­­­ NEW ZEALAND BP Kaimai Ra 427m Leyland Tramline 22 Sep 1981 B.M.May// litter 81/92 (NZAC); (1­­­female)­­­ Kaimai Ra. Matamara, Waikare M.Is.// 1600 feet // A.E.Brookes Coll. (NZAC); (1­­­female)­­­ Kaimai 1520 1-1-31. AEB// A.E.Brookes Collection (NZAC); (1­­­female)­­­ NEW ZEALAND, BP 9km from Opotiki 20 Aug 1976 A.R.Ferguson// Litter 76/63 (NZAC); (2­­­males)­­­ NEW ZEALAND: BP Orete Forest Te Puia Hut Bush 15 Sep 1992 G. Hall &// R.C. Henderson Sifted litter 92/50 (NZAC); (2­­­males,­­­2­­­females)­­­ NEW ZEALAND, BP Waiaroho 21 Oct 1992 J.S. Dugdale// litter & mosses 92/70 (NZAC); (1­­­male)­­­ Leafmould Fitzgeralds Rotorua // R.M. Bull Collection (NZAC); (2­­­males,­­­1­­­female)­­­ NEW ZEALAND, BP Te Koau, Hovelis Watching Dog 240m 24 Oct 1992 J.S. Dugdale// Litter 92/78 (NZAC); (2­­­males,­­­3­­­females)­­­ NEW ZEALAND, BP Te Koau, 243m 15 Mar 1993 J.S. Dugdale Litter 93/43 (NZAC); (1­­­male,­­­1­­­female)­­­ NEW ZEALAND, BP Waiaroho 10 Mar 1993 J.S. Dugdale Litter 93/36 (NZAC); (1­­­male)­­­ NEW ZEALAND CL Gt Barrier I Mt Hobson Upper Kauri Dam 11 Apr 1982 // moss and litter (NZAC); (3­­­males,­­­1­­­female)­­­ NEW ZEALAND ND Kauri Reserve Omahutu SF 10 Oct 1974 J.C. Watt// Litter 74/81 (NZAC); (1­­­male)­­­ NEW ZEALAND, ND Kohuronaki, 270m 24 Nov 1982 G. Kuschel// Litter and decayed wood 82/123 (NZAC); (4­­­males)­­­ NEW ZEALAND ND Parahaki Park 5 Nov 1981 G. Kuschel// litter and decayed wood 81/127 (NZAC); (3­­­males,­­­1­­­female)­­­ NEW ZEALAND ND Parahaki Park 5 Nov 1981 G. Kuschel// litter and decayed wood 81/126 (NZAC); (2­­­males,­­­1­­­female)­­­ NEW ZEALAND ND Waipoua SF Toatoa Grove 25 Nov 1980 G.Kuschel// Sifted litter 80/121 (NZAC); (2­­­males,­­­1­­­female)­­­ NEW ZEALAND TK Mt Egmont NP Stratford Mtn House, 846m 23 May 1986 // C.L. Lyal Leaf litter CL 474 (NZAC); (1­­­male)­­­ NEW ZEALAND, WO Mt Pirongia SF Rangitukia Stream 13 June 2001 R. Leschen// berlesate of tree holes leaf litter RL664 (NZAC); (1­­­male)­­­ NEW ZEALAND WO Mt Pirongia 962m 16 Jan 1977 A.K. Walker litter 77/2 (NZAC); (4­­­males,­­­3­­­females)­­­ NEW ZEALAND WO Mt Pirongia 962m 16 Jan 1977 A.K. Walker litter 77/2 (NZAC)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on pages 212-213, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382, {"references":["Broun, T. (1893) s. n. In: Manual of the New Zealand Coleoptera. Parts V, VI & VIII. Colonial Museum and Geological Survey Department, Wellington, pp. 975 - 1504.","Jeannel, R. (1951) Sur la systematique des genres de la tribu Pselaphini Raffray (Coleoptera Pselaphidae). Revue Francaise d'Entomologie, 18, 5 - 11."]}

Pselaphaulax intermedius Owens & Carlton, sp. nov. (Fig. 5a–c) Type Material. Holotype: (male) New Zealand: CO: Rock and Pillar Ra., 1158 m Swards, 13 Nov 1969 J McBurney, 69/208 (NZAC). Paratypes: (1 male, 1 female) New Zealand: CO: Old Man Ra., 1432 m, Litter and debris, 24 Feb 1974 JC Watt, 74/11 (NZAC). Etymology. The specific epithet refers to the species’ intermediate status between species groups 1 and 2. Diagnosis. Externally, Pselaphaulax intermedius can be distinguished from all others in the genus by possessing a combination of characters of species groups 1 and 2: gular mound rounded posteriorly (group 2); pronotum with median antebasal foveae well-developed, mesoventral shield with setose foveae (group 1). The form of the genitalia is also unique, with the median lobe large and complex, lateral processes absent, and ventral process simple and short. Description. MALE holotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 16 facets. Frontal margin of rostrum in dorsal view straight with sparse suberect setae. Gular mound narrowed apically and rounded posteriorly, swelling as long as wide, glabrous; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus faint, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with one basal fovea and single sutural fovea; discal stria absent, sutural striae present; apical fringe of setae present, sparse. Abdomen: Tergites of usual form for tribe. Ventrite II with median area slightly flattened at base, apically margined by short, sparse setae. Genitalia: aedeagus asymmetrical. Median lobe distally large and complex. Ventral process short, downward curved, distally, much shorter than median lobe. Lateral processes absent. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller. Distribution. Pselaphaulax intermedius is known only from the type localities in Central Otago (CO)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, Revision of the New Zealand species of the Genus Pselaphaulax Reitter (Staphylinidae: Pselaphinae: Pselaphitae: Pselaphini), pp. 221-244 in Zootaxa 5155 (2) on page 226, DOI: 10.11646/zootaxa.5155.2.3, http://zenodo.org/record/6673412

Pselaphogenius chandleri Owens ­­­&­­­Carlton,­­­sp.­­­nov. (Fig. 8a–c) Type­­­Material.­­­ Holotype: (1­­­male) New Zealand, BP Orete Forest Te Puia Hut 26 Apr 1993 G. Hall // Sifted litter 93/96 (NZAC). Paratypes:­­­(2­­­males,­­­3­­­females)­­­ holotype locality (NZAC). Etymology.­­­ This species is named after Donald. S. Chandler, fellow pselaphine worker, whose years of support to both authors has been much appreciated. Diagnosis.­­­ Externally, P. chandleri is difficult to distinguish from other species of Pselaphogenius in New Zealand. The form of the aedeagus, with the simple, straight, median lobe and the absence of lateral and ventral processes, is unique. Description.­­­ MALE holotype. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes small, composed of three facets. Frontal margin of rostrum narrowly rounded, with sparse suberect setae in dorsal view. Gular mound modified, longer than wide, broad anteriorly and narrowed posteriorly; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV longer than head, narrow, widened at base and distal 1/5 to 1/6, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae and median antebasal fovea absent. Mesoventrite with median shield unmodified. Metaventrite afoveate, posterior margin with elongate setae extending to base of ventrite II, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, single sutural fovea present; discal stria absent, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, shallow oval median depression small, restricted to basal portion of ventrite. Genitalia: aedeagus asymmetrical. Median lobe straight, simple, slightly down-curved. Lateral processes absent. Ventral process absent. Parameres present, as long as median lobe. Females.­­­ Integument,­­­ Head,­­­Thorax,­­­Abdomen:­­­ similar to male except ventrite II lacking median ovate depression. Distribution.­­­ This species is known from several localities in BP. Material­­­examined.­­­BP:­­­(1­­­female)­­­ NEW ZEALAND, BP Hicks Bay 25 Oct 1992 J.S. Dugdale Litter 92/79 (NZAC); (1­­­female) ­­­ NEW ZEALAND BP Lake Okataina 9 Oct 1995 M-C Lariviere & A Larochelle // Litter 95/12 (NZAC); (2­­­females) ­­­ NEW ZEALAND, BP Orete Forest Te Puia Hut Bush 19 Oct 1992 J.S. Dugdale (NZAC); (1­­­male,­­­2­­­females) ­­­ NEW ZEALAND, BP Orete Forest Te Puia Hut 13 Apr 1992 G. Hall // Liter 92/34 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Papatea 5 Nov 1993 R.C. Henderson Litter 93/115 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Rereauira 26 Apr 1993 J.S. Dugdale Litter 93/97 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Te Koau, Hovells Watching Dog 240m 24 Oct 1992 J.S. Dugdale // Litter 92/78 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Te Koau, 243m 15 Mar 1993 J.S. Dugdale Litter 93/43 (NZAC); (2­­­females) ­­­ NEW ZEALAND, BP Te Koau, Hovells Watching Dog Track 360m 31 Jan 1993 (NZAC); (1­­­male,­­­2­­­females) ­­­ NEW ZEALAND, BP Te Koau, 120m 29 Apr 1993 J.S. Dugdale Litter 93/101 (NZAC); (3­­­males,­­­1­­­female) ­­­ NEW ZEALAND, BP Te Koau, 125m 2 May 1993 R.C. Henderson Litter 93/107 (NZAC); (1­­­male) ­­­ NEW ZEALAND, BP Te Koau 140m Twin Puriri's 31 Jan 1993 J.S. Dugdale // Litter 93/10 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Te Koau Main Ridge, 220m 23 Sep 1992 J.S. Dugdale // Litter 92/83 (NZAC); (1­­­male) ­­­ NEW ZEALAND, BP Lottin Pt Rd Waenga Bush 15 Mar 1984 G. Hall &// R.C. Henderson Litter 54/5 (NZAC); (1­­­male,­­­3­­­females) ­­­ NEW ZEALAND, BP Waiaroho 10 Mar 1993 J.S. Dugdale Litter 93/36 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Waiaroho 2 Nov 1995 G. Hall Litter 95/13 (NZAC); (1­­­female) ­­­ NEW ZEALAND, BP Waiotuma Stm 85m 28 Jan 1993 J.S. Dugdale Litter 93/9 (NZAC)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on pages 196-198, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382

Pselaphogenius lescheni ­­­Owens­­­&­­­Carlton,­­­sp.­­­nov. (Figs. 5f, 24a–c) Type­­­Material.­­­ Holotype:­­­(1­­­male)­­­ New Zealand: MB: Black Birch, 1800 m Litter, 16 Feb 1970 AC Eyles, 70/139 (NZAC). Paratypes:­­­(1­­­male,­­­1­­­female)­­­ holotype locality (NZAC); (2­­­males,­­­1­­­female)­­­ New Zealand: MB: Black Birch Ra., Mt. Altimarloch 1700 m, litter, 16 Feb 1970 G Kuschel, 70/119 (NZAC). Etymology.­­­ This species is named after Dr. Richard Leschen, a specialist of New Zealand Coleoptera, supporter of this project, and friend of the authors. Diagnosis.­­­ Externally, P. lescheni is most similar to P. kalleri in possessing a tubercle at the apex of the medial oval depression on ventrite II. However, the form of the aedeagus, with the deeply scoop-shaped parameres and the presence of only a right lateral process is unique. Redescription.­­­ MALE. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 15 facets. Frontal margin of rostrum straight in dorsal view. Gular mound modified, as wide as long, broad anteriorly and rounded posteriorly; area of head posterior to gular mound flattened, slightly concave, nude, maxillary palpomere IV as long as head, narrow, widened at base and distal ½ to 1/3, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae present, median antebasal fovea reduced. Mesoventrite with median shield unmodified. Metaventrite afoveate, median area broadly flattened, bearing short, fine setae, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae and single sutural fovea present; discal stria absent, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, deep oval median depression reaching apex, apex thickened into a tubercle bearing patch of fine, short setae. Genitalia: aedeagus asymmetrical. Median lobe short, simple. Lateral process only produced on right side of aedeagus, inner and outer process simple. Ventral process absent. Parameres large, deep and scoop shaped, almost completely encasing the terminal portion of the aedeagus in lateral view. Females.­­­Integument,­­­Head,­­­Thorax,­­­Abdomen:­­­ Females are similar to males except eyes slightly smaller and ventrite II lacking median ovate depression. Distribution.­­­ This species is only known from collection events in Marlborough (MB) near Black Birch Range and a single collection event from Buller (BR) near Mt. Robert. Material­­­examined.­­­ (4­­­males,­­­2­­­females)­­­ New Zealand: MB: Black Birch Ra., Mt. Altimarlock 1700 m, litter, 16 Feb 1970 G Kuschel, 70/121 (LSAM); (3­­­males,­­­7­­­females)­­­ New Zealand: MB: Black Birch Ra., Mt.Altimarlock 1700 m, litter, 16 Feb 1970 G Kuschel, 70/123; (2­­­males)­­­ New Zealand: MB: Black Birch Ra., Mt.Altimarlock 1700 m, mats, 16 Feb 1970 G Kuschel, 70/125 (LSAM); (7­­­males,­­­5­­­females)­­­ New Zealand: MB: Black Birch Ra., Mt. Altimarlock 5550 m, mats, 16 Feb 1970 G Kuschel, 70/126 (LSAM). (1­­­male)­­­ NEW ZEALAND: BR: Nelson Lakes N.P., Mt. Robert, Relax Shelter 1420m, 41 ∘ 50.2'S, 172 ∘ 48.6'E, 1.xii.2005, alpine tussock herbfields; FMHD#2005- 062, berl., debris at bases of tussocks, D. Clarke 95% ethanol FIELD MUSEUM NAT. HIST. (FMNH, EtOH)., Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on pages 219-220, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382

Pselaphaulax ramsayi Owens & Carlton, new species (Fig. 10a–c) Type Material. Holotype (male) New Zealand: NN: Farewell Spit Lighthouse Litter, 28 Jan 1967 AK Walker, 67/47 (NZAC). Etymology. This species is named in honor of GW Ramsay, collector of the holotype. Diagnosis. This species is the only species in the genus with a completely reduced (absent) ventral process of the aedeagus. The median lobe is broad, also uncharacteristic for the genus. Description. MALE holotype. Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head: wide, width between eyes greater than ½ length of head from base of vertex to apex of rostrum. Eyes rounded, composed of 19 facets. Frontal margin of rostrum in dorsal view straight with sparse suberect setae. Gular mound modified, ovate, narrowed posteriorly and anteriorly, longer than wide; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV as long as head, widened at base and distal 1/3, apical swelling roughened, v-shaped sensory patches present, margined by carinae. Thorax: pronotum with basal sulcus distinct, delimited by lateral antebasal foveae; median antebasal foveae margined by small longitudinal carinae, present in groove alongside basolateral pits. Mesoventrite with median shield bearing patch of spongeose setae in small concavity behind margin of procoxae. Metaventrite afoveate, raised into median tumosity; metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae, single sutural fovea; single discal stria, sutural striae present; sparse apical fringe of setae present. Abdomen: tergites of usual form for tribe. Ventrite II with median area slightly flattened throughout length of ventrite, apically margined by short, sparse setae. Genitalia: aedeagus asymmetrical. Median lobe distally curved downwards, broad. Ventral process reduced, apparently absent. Parameres present, as long as median lobe. Females. Integument, Head, Thorax, Abdomen: similar to male except eyes slightly smaller and ventrite II lacking median flattened area. Distribution. This species is only known from the holotype from Nelson.

Pselaphogenius elephantus Owens ­­­&­­­Carlton,­­­sp.­­­nov. (Fig. 11a–c) Type­­­Material.­­­ Holotype: (1­­­male)­­­ NEW ZEALAND ND Waipoua SF Wairau Smt, 390m 14 Apr 1980 J.C.Watt // Sifted litter 80/51 (NZAC). Etymology.­­­ The specific epithet refers to the elongate processes on the aedeagus, which resemble the trunk of an elephant. Diagnosis.­­­ Externally, this species is similar to other species of Pselaphogenius in New Zealand. The form of the aedeagus, with the greatly elongate ventral process, is unique. Description.­­­ MALE holotype. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes small, composed of five facets. Frontal margin of rostrum acute with sparse suberect setae in dorsal view. Gular mound modified, longer than wide, broad anteriorly and narrowed posteriorly; area of head posterior to gular mound flattened, slightly concave, nude, maxillary palpomere IV longer than head, narrow, widened at base and distal 1/5, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae absent, antebasal fovea reduced to minute pit. Mesoventrite with median shield unmodified. Metaventrite afoveate, flattened, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae and sutural fovea present; discal stria absent, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, shallow oval median depression not reaching apex, hind margin of depression with median tubercle. Genitalia: aedeagus asymmetrical. Median lobe straight, short. Lateral processes present, narrow, elongate, right process curved inwards towards left, left process gradually curving inwards towards right. Ventral greatly elongate, longer than phallobase, distally flattened and bifurcate. Parameres present, as long as ventral process. Females.­­­Integument,­­­Head,­­­Thorax,­­­Abdomen:­­­ females of this species are unknown. Distribution.­­­ This species is known from the holotype locality in ND, Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on page 202, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382

Pselaphogenius otteai Owens ­­­&­­­Carlton,­­­sp.­­­nov. (Fig. 16a–c) Type­­­ Material.­­­ Holotype: (1­­­ male)­­­ NEW ZEALAND: ND Waipoua State Forest, 0.8km nw Wairau Summit 350m, 27.xi.1984 hdwd.-podocarp forest A. Newton / M. Thayer 689// berl., leaf & log litter, forest floor (NZAC). Paratypes: (5­­­males,­­­2­­­females)­­­ holotype locality (FMNH). Etymology.­­­ This species is named in honor of Dr. James Ottea, for his support of this project as member of the first author’s advisory committee. Diagnosis.­­­ Externally, P. otteai is similar to other species in the genus. The form of the aedeagus, with the bifurcate lateral process on the right side is unique. Description.­­­ MALE holotype. ­­­Integument: head, pronotum, elytra, abdomen, and appendages light brown, palpi lighter in color. Dorsal surface with scattered, stiff, elongate setae. Head:­­­ width between eyes equal to ½ length of head from base of vertex to apex of rostrum. Eyes small, composed of three facets. Frontal margin of rostrum acute, with sparse suberect setae in dorsal view. Gular mound ovate, longer than wide, narrowed posteriorly; area of head posterior to gular mound flattened, slightly concave, nude. Maxillary palpomere IV longer than head, narrow, widened at base and distal 1/5, v-shaped sensory patches present. Thorax:­­­ pronotum basal sulcus absent, lateral antebasal foveae and median antebasal fovea absent. Mesoventrite with median shield unmodified. Metaventrite afoveate, medially depressed, metacoxae separated by extension of ventrite I. Legs unmodified. Elytra with two basal foveae and sutural fovea present; discal stria absent, sutural striae present. Abdomen:­­­ tergites of usual form for tribe. Ventrite I covered in dense, shining setae; ventrite II with basal sulcus occupying length of ventrite, sulcus densely setose, shallow ovate median depression occupying length of ventrite. Genitalia: aedeagus asymmetrical. Median lobe simple. Dorsal processes short, curved ventrad. Lateral process on right side bifurcate, as long as ventral process; left lateral process straight, simple, as long as ventral process. Ventral process ventrally curved, distally, as long as median lobe. Parameres present, as long as median lobe. Females.­­­Integument,­­­Head,­­­Thorax,­­­Abdomen:­­­ similar to male except eyes slightly smaller and ventrite II lacking median ovate depression. Distribution.­­­ This species is known from only the type locality in ND, Published as part of Owens, Brittany E. & Carlton, Christopher E., 2022, --- Revision- -- of- -- the- -- New- -- Zealand- -- species- -- of- -- the- -- Genus- -- Pselaphogenius Reitter- -- (Staphylinidae: --- Pselaphinae: --- Pselaphitae: --- Pselaphini) ---, pp. 187-220 in Zootaxa 5155 (2) on page 208, DOI: 10.11646/zootaxa.5155.2.2, http://zenodo.org/record/6673382