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The Homalocnemidae of Chile is reviewed, including the descriptions of the male and female terminalia of the two known species: H. nigripennis Philippi and H. praesumpta Collin. All references pertaining to Chilean Homalocnemis Philippi and the two known species are provided. Geographical distributions of the species were compiled and mapped from bibliographic data and collections. An illustrated key to the two species is also presented.

The Niphta Theischinger fauna of South America is revised to include 11 species, nine of which are described as new to science (N. acus Pivar, sp. nov., N. bifurcata Pivar & Moulton, sp. nov., N. bispinosa Pivar & Sinclair, sp. nov., N. brunnea Pivar, sp. nov., N. courtneyi Pivar, sp. nov., N. daniellae Pivar, sp. nov., N. downesi Pivar, sp. nov., N. eurydactyla Pivar, sp. nov., N. mapuche Pivar, sp. nov.). The genus Niphta is redefined, both previously described Chilean species are redescribed, N. halteralis (Edwards) and N. nudipennis (Edwards), and females are described or redescribed where possible. The first descriptions of the immature stages of South American Niphta are provided, which represent a new larval morphotype in Thaumaleidae, as larvae and pupae possess ventral adhesive structures. Furthermore, these larvae were collected from vegetation rather than rocky substrates. Illustrations and micrographs are provided for all species, and scanning electron microscopy images are included for select immatures. A key to species, distribution maps, and discussions regarding phylogenetic affinities and habitat are also included.

Louse flies (Diptera, Hippoboscidae) are ectoparasites often found on birds. As they spend most of their life on their host, they are not often collected or studied. Hence, little is known about their species richness and prevalence on raptors in Canada. In this study, louse flies were collected from 184 out of 1467 raptors examined during the 2020 fall migration in southern Ontario, Canada, giving an overall prevalence of 12.5%. In total, 256 louse fly specimens were collected (mean intensity = 1.41) representing four species (identified morphologically, with support of DNA barcoding): Icosta americana (91.9%), Ornithomya anchineuria (0.3%), O. avicularia (7.3%) and O. bequaerti (0.3%). Mite clusters were found on 42% of O. avicularia. This study also presents the first North American record for O. avicularia and the presence of O. bequaerti in Europe was confirmed for the first time. Based on the different parameters recorded during banding, it appears that the host species and the month play a part in the presence of louse flies on the host. Further study of louse flies is needed to better understand their prevalence across different bird groups and geographic distribution., Graphical abstract Image 1, Highlights • The overall prevalence of louse flies on raptors was 12.5%. • Host species and month influence the presence of louse flies. • Ornithomya avicularia is recorded from North America for the first time. • Ornithomya bequaerti is confirmed for the first time from Europe. • Icosta americana was the most abundant louse fly (91.9%).

Pseudoscelolabes Collin, 1933, an endemic New Zealand genus, is revised, including the description of a new species, P. lesagei Sinclair & Barros sp. nov. The male and female terminalia of the genus are described and illustrated for the first time. An identification key to the two included species and a map with distribution records are provided. The relationships of the genus within Ocydromiinae are discussed.

The identity, richness, and abundance of true flies (Diptera) from the nests of three cavity-nesting raptors (Aves) were investigated in northern Nova Scotia, Canada. After fledging, flies were extracted from the nest material using Berlese funnels within an emergence chamber. Thirty-one species/morphospecies from 14 families were collected, including eight new records for Nova Scotia and two new records for eastern North America.

A new fossil genus of the family Hybotidae is described, based on male and female specimens. The new genus is monotypic: Syneproctus caridadi gen. et sp. nov. It belongs to the subfamily Hybotinae and shares some characters with the extant genera Syneches Walker, 1852, Stenoproctus Loew, 1858 and Chillcottomyia Saigusa, 1986; however, the differences, principally in the wing venation (shortened cua cell, horizontal m-cu crossvein) and sclerotized mouthparts, support the description of a new genus. This is the first description of a new taxon of the subfamily Hybotinae from specimens preserved in Dominican amber.

The Wiedemannia zetterstedti species group is revised after examination of all available type specimens and includes one new species (W. ulrichi Ivković & Sinclair sp. nov.) and four redescribed species (W. czernyi (Bezzi), W. longipennis (Mik) stat. rev., W. rufipes (Oldenberg) stat. rev. and W. zetterstedti (Fallén)). The following new synonyms are proposed: W. (Roederella) ouedorum Vaillant, 1952 = W. czernyi (Bezzi, 1905); Paramesia riparia Robert, 1836 = W. zetterstedti (Fallén, 1826). Lectotypes are designated for the following species/subspecies: Atalanta hirtiloba Speiser, Brachystoma escheri Zetterstedt, Clinocera czernyi Bezzi, Clinocera longipennis Mik, Paramesia riparia Robert, and Roederia czernyi rufipes Oldenberg. In addition to morphological evidence, molecular species concepts were investigated using a molecular phylogenetic divergence-based species delimitation (bPTP) and results confirmed the morphological conclusions. A key to species is presented and geographic distributions are mapped.

A new empidoid genus, Saigusamyia new genus and three new species (S. denningi new species, S. harkrideri new species, S. uvasensis new species) (Diptera: Empidoidea: Brachystomatidae) are described and illustrated. This new genus is related to a group of genera in the Trichopezinae, with fused epandrium and hypandrium and highly modified sperm pump. Adults of Saigusamyia are present primarily during the colder months in California (USA). The phylogenetic relationships of the Trichopeza genus-group are analyzed based on morphological data.

Iteaphila Zetterstedt is redefined to include species with both branched and unbranched radial vein (R4+5) on the basis of a morphological cladistic analysis and parsimony analysis of COI mitochondrial DNA barcode sequences. As a result, Anthepiscopus Becker is hypothesized as a junior synonym of Iteaphila and species of the Iteaphila setosa group are transferred to the new genus, Paraiteaphila gen. nov. The following new combinations are proposed: P. arundela (Shamshev & Sinclair, 2009) comb. nov., P. caucasica (Shamshev & Sinclair, 2009) comb. nov., P. italica (Loew, 1873) comb. nov., P. kubaniensis (Shamshev & Sinclair, 2009) comb. nov., P. merzi (Shamshev & Sinclair, 2009) comb. nov. and P. setosa (Bezzi, 1924) comb. nov. These two genera are assigned to the family Iteaphilidae stat. rev. Thirty-five species of Iteaphila with unbranched radial vein are revised, including 26 new species: bulbosa species group (I. beringiensis sp. nov., I. bifida sp. nov., I. recta sp. nov., I. tribulosa sp. nov.), macquarti species group (I. bartaki sp. nov., I. falki sp. nov., I. kyrgyzstanensis sp. nov., I. ribesii (Becker, 1891) comb. nov.), nitidula species group (I. longiphallus sp. nov.), nupta species group (I. arnaudi sp. nov., I. bayarea sp. nov., I. brooksi sp. nov., I. dichoptica sp. nov., I. flavipilosa (Coquillett, 1900) comb. nov., I. glabricula sp. nov., I. gracilis sp. nov., I. grandis sp. nov., I. lolo sp. nov., I. longipalpis (Melander, 1928) comb. nov., I. luteitibia sp. nov., I. nupta (Melander, 1928) comb. nov., I. sierrensis sp. nov., I. spinosa sp. nov., I. subnupta sp. nov.), oedalina species group (I. aktruensis sp. nov., I. incus sp. nov., I. miranda sp. nov., I. oedalina (Zetterstedt, 1838) comb. nov., I. polygyna (Melander, 1928) comb. nov., I. recurvata sp. nov., I. sakhalinensis sp. nov., I. zontaki (Nowicki, 1871) comb. nov.), stentor species group (I. parastentor sp. nov., I. stentor (Melander, 1902) comb. nov.) and unplaced to species group (I. caelebs (Becker, 1891) comb. nov.). The following new synonyms are proposed: I. flavicoxa (Melander, 1928) is a junior synonym of I. polygyna (Melander, 1928); I. hirsutus (Melander, 1928) is a junior synonym of I. oedalina (Zettersedt, 1838). Lectotypes are designated for the following species: I. flavipilosa, I. fraternella Zetterstedt, I. nigra Zetterstedt, I. nupta, I. oedalina, I. polygyna and I. ribesii. All species of Iteaphila with unbranched R4+5 are described, key to species presented, male terminalia illustrated, distributions plotted and flowers visited by these species are listed. COI mitochondrial DNA barcode sequences were obtained for 18 identified Nearctic species of Iteaphila with both branched and unbranched R4+5.

The western Nearctic fauna of Androprosopa Mik is revised to include twenty-five species, three of which are new to science. Descriptions of the adult males of A. apache Pivar & Moulton sp.n., A. arnaudi Pivar sp.n. and A. rainierensis Pivar & Sinclair sp.n. are presented. Redescriptions of all remaining western Nearctic species are provided, as well as genitalic illustrations and updated distribution maps for each species. A diagnostic key to males of western Nearctic species of Androprosopa is offered. Species groups based on morphology are discussed.

A new species, Contarinia brassicola Sinclair (Diptera: Cecidomyiidae), which induces flower galls on canola (Brassica napus Linnaeus and Brassica rapa Linnaeus (Brassicaceae)), is described from Saskatchewan and Alberta, Canada. Larvae develop in the flowers of canola, which causes swelling and prevents opening, pod formation, and seed set. Mature larvae exit the galls, fall to the soil, and form cocoons. Depending on conditions, larvae will either pupate and eclose in the same calendar year or enter facultative diapause and emerge the following year. At least two generations of C. brassicola occur each year. Adults emerge from overwintering cocoons in the spring and lay eggs on developing canola flower buds. The galls produced by C. brassicola were previously attributed to the swede midge, Contarinia nasturtii (Kieffer) in Saskatchewan; here, we compare and list several characters to differentiate the two species.

The Canadian Diptera fauna is updated. Numbers of species currently known from Canada, total Barcode Index Numbers (BINs), and estimated numbers of undescribed or unrecorded species are provided for each family. An overview of recent changes in the systematics and Canadian faunistics of major groups is provided as well as some general information on biology and life history. A total of 116 families and 9620 described species of Canadian Diptera are reported, representing more than a 36% increase in species numbers since the last comparable assessment by JF McAlpine et al. (1979). Almost 30,000 BINs have so far been obtained from flies in Canada. Estimates of additional number of species remaining to be documented in the country range from 5200 to 20,400.

The species of the Nearctic genus Tritoxa Loew are revised. Seven species of Tritoxa are recognized, including two new species: T. californica sp. nov. (type locality: near Spring Garden, California), T. cuneata Loew, T. decipiens sp. nov. (type locality: near Smithers, British Columbia), T. flexa (Wiedemann), T. incurva Loew, T. pollinosa Cole and T. ra Harriot. One species from California and Nevada based on female specimens remains undescribed. A key to all species is provided, species are illustrated and their distributions mapped. Wing patterns may be used to differentiate among all species, although confident identification of certain sympatric species requires confirmation by examination of male genitalia.

We present a summary and analysis of the Diptera-related information published in Zootaxa from 2001 to 2020, with a focus on taxonomic papers. Altogether, 2,527 papers on Diptera were published, including 2,032 taxonomic papers and 1,931 papers containing new nomenclatural acts, equivalent to 22% of all publications with new nomenclatural acts for Diptera. The new nomenclatural acts include 7,431 new species, 277 new genera, 2,003 new synonymies, and 1,617 new combinations. A breakdown by family of new taxa and new replacement names proposed in the journal during the last two decades is provided, together with a comparison of Zootaxa’s output to that of all other taxonomic publications on Diptera. Our results show that the journal has contributed to 20% of all biodiversity discovery in this megadiverse insect order over the last 20 years, and to about 31% in the last decade.

The Nearctic species of Gimnomera Rondani are revised and includes eight species, of which three are new to science (G. aquilonia sp. nov., G. cerea (Coquillett), G. cuneiventris (Zetterstedt), G. incisurata Malloch, G. subvittata (Malloch), G. terrywheeleri sp. nov., G. tibialis (Malloch), G. vockerothi sp. nov.). Gimnomera cuneiventris is newly recognized as Holarctic in distribution. Gimnomera fasciventris Malloch is removed and transferred to the genus Acerocnema, comb. nov. An additional Nearctic species is recognized but not described due to lack of male specimens. Species descriptions, diagnoses and distribution maps are presented, including images of the male terminalia as well as additional diagnostic characters. A key to the Nearctic species is also provided. A morphological cladistic analysis of 12 species reveals that the Nearctic and Palearctic species do not form separate monophyletic groups. COI mitochondrial DNA barcode sequences were obtained for eight Nearctic species of Gimnomera.

The empidoid flies of the Russian Arctic islands and Svalbard Archipelago are revised including families Empididae (4 genera, 32 species), Hybotidae (2 genera, 3 species) and Brachystomatidae (1 genus, 2 species). The following ten new species are described: Empididae—Empis septentrionalis Shamshev & Sinclair sp. nov. (Russia: Wrangel Island); Rhamphomyia (Dasyrhamphomyia) gorodkovi Shamshev, Sinclair & Saigusa sp. nov. (Russia: Chukotka mainland, Wrangel Island, Taymyr, Yakutia; USA: Alaska); R. (D.) mallochi Shamshev, Sinclair & Saigusa sp. nov. (Canada: Yukon; Russia: Chukotka mainland, Kolyuchin and Wrangel Islands; USA: Alaska); R. (Pararhamphomyia) subfilicauda Shamshev & Sinclair sp. nov. (Russia: Wrangel Island); R. (P.) submacrura Shamshev & Sinclair sp. nov. (Russia: Wrangel Island); R. (P.) subtenuiterfilata Shamshev & Sinclair sp. nov. (Russia: Wrangel Island, Yakutia); R. (P.) wrangeli Shamshev & Sinclair sp. nov. (Russia: Wrangel Island); R. sublongiseta Shamshev, Sinclair & Saigusa sp. nov. (Canada: Yukon; Russia: Wrangel Island); one species of Rhamphomyia was retained as unnamed due to insufficient material; Hybotidae—Platypalpus septentrionalis Shamshev & Sinclair sp. nov. (Russia: Wrangel Island), P. subtectifrons Shamshev & Sinclair sp. nov. (Canada: Northwest Territories, Yukon; Russia: Wrangel Island; USA: Alaska). For previously described species a short diagnosis and a habitus photo are provided to enable easier recognition. In addition, a key to the species of Rhamphomyia (Empididae) occurring on the Russia Arctic islands and Svalbard Archipelago is compiled. A lectotype is designated for Rhamphomyia armipes Sack, 1923. The male and the female of R. taimyrensis Frey and R. kaninensis Frey are described for the first time, respectively. Tachypeza sericeipalpis Frey is newly recorded from North America (Canada: Northwest Territories, Nunavut) and first illustration of the male terminalia is provided. Trichoclinocera lapponica (Ringdahl) is recorded for the first time from the territory of Russia.

Neothaumalea atlanticanew genus, new species (Diptera: Thaumaleidae), is described from the state of Santa Catarina in southern Brazil. This represents the first thaumaleid collected east of the Andes mountain range. The egg, larva, pupa, and both adults are described and illustrated, distribution map presented, and phylogenetic affinities discussed. A key to the genera of South America is also provided.

We report first record of the occurrence of the aquatic empidid genus Trichoclinocera Collin in Missouri based upon aquatic macroinvertebrate samples collected during March 2016 from two riffles in East Fork of the Black River in Reynolds County in the Ozark Highlands. Substrate characteristics and notes on habitat from where larval specimens of Trichoclinocera were collected are also provided.

Rhamphomyia of the Canadian Arctic Archipelago, Greenland and Iceland, comprising 23 species, including five new species, are revised: R. (Ctenempis) albopilosa Coquillett, R. (Dasyrhamphomyia) erinacioides Malloch, R. (Dasyrhamphomyia) hovgaardii Holmgren, R. (Dasyrhamphomyia) leptidiformis Frey, R. (Dasyrhamphomyia) nigrita Zetterstedt, R. (Eorhamphomyia) shewelli Sinclair, Vajda, Saigusa & Shamshev sp. nov., R. (Pararhamphomyia) diversipennis Becker, R. (Pararhamphomyia) filicauda Henriksen & Lundbeck, R. (Pararhamphomyia) frigida Sinclair, Vajda, Saigusa & Shamshev sp. nov., R. (Pararhamphomyia) helleni Frey, R. (Pararhamphomyia) hilariformis Frey, R. (Pararhamphomyia) hoeli Frey, R. (Pararhamphomyia) kjellmanii Holmgren, R. (Pararhamphomyia) lymaniana Sinclair, Vajda, Saigusa & Shamshev sp. nov., R. (Pararhamphomyia) omissinervis Becker, R. (Pararhamphomyia) petervajdai Sinclair, Vajda, Saigusa & Shamshev sp. nov., R. (Pararhamphomyia) septentrionalis Sinclair, Vajda, Saigusa & Shamshev sp. nov., R. (Pararhamphomyia) simplex Zetterstedt, R. (Pararhamphomyia) ursinella Melander, R. herschelli Malloch, R. hirtula Zetterstedt, R. laevigata Loew, R. setosa Coquillett. The following six new synonyms are proposed: R. calvimontis Cockerell, 1916 and R. wuorentausi Frey, 1922 = R. albopilosa Coquillett, 1900; R. fridolini Frey, 1950 = R. laevigata Loew, 1861; R. hirticula Collin, 1937 = R. setosa Coquillett, 1895; R. uralensis Becker, 1915 = R. kjellmanii Holmgren, 1880; R. zaitzevi Becker, 1915 = R. hovgaardii Holmgren, 1880. Lectotypes are designated for the following species: R. diversipennis Becker, R. filicauda Henriksen & Lundbeck, R. helleni Frey, R. herschelli Malloch, R. hirticula Collin, R. hoeli Frey, R. leptidiformis Frey, R. omissinervis Becker, R. setosa Coquillett, R. uralensis Becker, R. wuorentausi Frey, R. zaitzevi Becker. A neotype is designated for R. laevigata Loew. Keys to male and female species of Rhamphomyia and distribution maps of this region are provided. DNA barcode data are presented for 16 species of arctic Rhamphomyia.

Paraleucopidae Wheeler fam. nov. is proposed for the previously unplaced New World genera Paraleucopis Malloch, Mallochianamyia Santos-Neto and Schizostomyia Malloch and undescribed Australian species. A key to genera of Paraleucopidae is provided. Paraleucopis is revised and includes nine species: P. auripes Wheeler & Sinclair sp. nov. (type locality: Andalgala, Argentina); P. bispinosa Wheeler & Sinclair sp. nov. (type locality: Socos, Coquimbo, Chile); P. boharti Wheeler & Sinclair sp. nov. (type locality: Andalgala, Argentina); P. boydensis Steyskal (type locality: nr. Palm Desert, California, USA); P. corvina Malloch (type species of genus; type locality: New Mexico, USA); P. mexicana Steyskal (type locality: Kino Bay, Mexico); P. nigra Wheeler & Sinclair sp. nov. (type locality: Portal, Arizona, USA); P. paraboydensis Wheeler & Sinclair sp. nov. (type locality: Willis Palms Oasis, California, USA); P. saguaro Wheeler & Sinclair sp. nov. (type locality: Usery Mtn Park, Arizona, USA). A key to the species of Paraleucopis is provided. The distribution of Paraleucopis is disjunct, with six species in the western United States and northwestern Mexico and three species in northern Chile and northern Argentina. The sister group and superfamilial assignment of the Paraleucopidae cannot be established based on current knowledge although the family has affinities to some families of the Asteioinea sensu J.F. McAlpine. A well-supported hypothesis on the relationships of the families of the Acalyptratae will be required before the sister group relationships of Paraleucopidae can be determined.

The Holarctic genus Gloma Meigen is revised and includes five species (G. fuscipennis Meigen, G. fuscipes Melander, G. luctuosa Melander, G. pectinipes Melander, G. pyricornis Saigusa & Sinclair sp. nov.) and two undescribed species from China, presently known only from females. The monotypic Baltic amber genus Palaeoparamesia Meunier is discussed as possibly congeneric with Gloma. The Baltic amber species Gloma hirta Loew is considered a nomen dubium. A lectotype is designated for Gloma luctuosa. All species are described or diagnosed, and their geographic distributions mapped. A World key to the species of Gloma is provided. Phylogenetic placement of the genus, including a possible new relationship with Oreogeton Schiner is considered, and the relationships and zoogeographic history of the species are discussed based on a morphological cladistic analysis.

A large diversity of saproxylic Empidoidea from a temperate deciduous forest in southern Quebec is documented. Adults of 43 empidoid species representing 19 genera in 12 subfamilies and three families were collected from in situ sealed emergence traps placed over decayed logs of American beech (Fagus grandifolia Ehrhart) and sugar maple (Acer saccharum Marshall). The results of this technique give a clear indication of the saproxylic larval and pupal habitat of these species. The importance of Empidoidea as larval predators in decayed wood niches is noted.

While databasing type specimens of Hydropeza Sinclair in the Australian Museum (Sydney), different spellings of a species name in Sinclair (2016) were identified. I herein resolve this nomenclatural issue.

Study of all flies (Diptera) collected for one year from a four-hectare (150 x 266 meter) patch of cloud forest at 1,600 meters above sea level at Zurquí de Moravia, San José Province, Costa Rica (hereafter referred to as Zurquí), revealed an astounding 4,332 species. This amounts to more than half the number of named species of flies for all of Central America. Specimens were collected with two Malaise traps running continuously and with a wide array of supplementary collecting methods for three days of each month. All morphospecies from all 73 families recorded were fully curated by technicians before submission to an international team of 59 taxonomic experts for identification. Overall, a Malaise trap on the forest edge captured 1,988 species or 51% of all collected dipteran taxa (other than of Phoridae, subsampled only from this and one other Malaise trap). A Malaise trap in the forest sampled 906 species. Of other sampling methods, the combination of four other Malaise traps and an intercept trap, aerial/hand collecting, 10 emergence traps, and four CDC light traps added the greatest number of species to our inventory. This complement of sampling methods was an effective combination for retrieving substantial numbers of species of Diptera. Comparison of select sampling methods (considering 3,487 species of non-phorid Diptera) provided further details regarding how many species were sampled by various methods. Comparison of species numbers from each of two permanent Malaise traps from Zurquí with those of single Malaise traps at each of Tapantí and Las Alturas, 40 and 180 km distant from Zurquí respectively, suggested significant species turnover. Comparison of the greater number of species collected in all traps from Zurquí did not markedly change the degree of similarity between the three sites, although the actual number of species shared did increase. Comparisons of the total number of named and unnamed species of Diptera from four hectares at Zurquí is equivalent to 51% of all flies named from Central America, greater than all the named fly fauna of Colombia, equivalent to 14% of named Neotropical species and equal to about 2.7% of all named Diptera worldwide. Clearly the number of species of Diptera in tropical regions has been severely underestimated and the actual number may surpass the number of species of Coleoptera. Various published extrapolations from limited data to estimate total numbers of species of larger taxonomic categories (e.g., Hexapoda, Arthropoda, Eukaryota, etc.) are highly questionable, and certainly will remain uncertain until we have more exhaustive surveys of all and diverse taxa (like Diptera) from multiple tropical sites. Morphological characterization of species in inventories provides identifications placed in the context of taxonomy, phylogeny, form, and ecology. DNA barcoding species is a valuable tool to estimate species numbers but used alone fails to provide a broader context for the species identified.

Made available in DSpace on 2019-10-06T15:56:52Z (GMT). No. of bitstreams: 0 Previous issue date: 2018-12-01 Estimations of tropical insect diversity generally suffer from lack of known groups or faunas against which extrapolations can be made, and have seriously underestimated the diversity of some taxa. Here we report the intensive inventory of a four-hectare tropical cloud forest in Costa Rica for one year, which yielded 4332 species of Diptera, providing the first verifiable basis for diversity of a major group of insects at a single site in the tropics. In total 73 families were present, all of which were studied to the species level, providing potentially complete coverage of all families of the order likely to be present at the site. Even so, extrapolations based on our data indicate that with further sampling, the actual total for the site could be closer to 8000 species. Efforts to completely sample a site, although resource-intensive and time-consuming, are needed to better ground estimations of world biodiversity based on limited sampling. Entomology Section Natural History Museum of Los Angeles County, 900 Exposition Boulevard Royal British Columbia Museum and the American Museum of Natural History, 691-8th Ave. SE Department of Plant and Environmental Sciences Poole Agricultural Center Clemson University, 130 McGinty Court, E-143 Depto. de Biologia FFCLRP Universidade de São Paulo, Av. Bandeirantes 3900 Great Lakes Forestry Centre Canadian Forest Service Natural Resources Canada, 1219 Queen St. E. Australian Museum, 1 William Street Department of Natural Resource Sciences McGill University Macdonald Campus Canadian National Collection of Insects Invertebrate Biodiversity Agriculture and Agri-Food Canada, K.W. Neatby Building, 960 Carling Avenue Department of Biological Sciences Spaulding Hall University of New Hampshire Department of Biological Sciences Wright State University, 3640 Colonel Glenn Hwy Instituto Federal do Triângulo Mineiro – Campus Uberaba. Rua João Batista Ribeiro 4000 Distrito Industrial II Departamento de Zoologia Universidade Federal do Paraná Jardim das Américas Mississippi Entomological Museum Mississippi State University, 100 Old Highway 12, P.O. Drawer 9775 Department of Biology Western Kentucky University Integrative Research Center Field Museum of Natural History Independent Investigator California State Collection of Arthropods, 2683 Tam O’ Shanter Dr., El Dorado Hills California Department of Food and Agriculture California State Collection of Arthropods, 3294 Meadowview Rd. The Academy of Natural Sciences of Drexel University, 1900 Ben Franklin Parkway American Museum of Natural History, Central Park West at 79th St. Department of Entomology University of California Riverside, 900 University Ave. Zoological Museum Biodiversity Unit FI-20014 University of Turku Instituto de Ecología A.C. (INECOL) Red Ambiente y Sustentabilidad, Carretera Antigua a Coatepec 351, Col El Haya, Xalapa Station Linné, Ölands Skogsby 161 I. I. Schmalhausen Institute of Zoology of the National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15 Universidad de Panama Department of Natural History University Museum of Bergen University of Bergen, P.O. Box 7800 Agriculture and Agri-Food Canada, 960 Carling Avenue School of Environmental Sciences University of Guelph Department of Entomology Smithsonian Institution, PO Box 37012, MRC 169 Natural History Museum of Denmark, Universitetsparken 15 Institute of Evolutionary Biology and Environmental Studies University of Zurich, Winterthurerstrasse 190 Systematic Entomology Laboratory USDA ARS c/o National Museum of Natural History, MRC-168, P.O. Box 37012 Departamento de Sistemática e Ecologia CCEN Universidade Federal da Paraíba, Castelo Branco, s/n Research Institute for Nature and Forest (INBO), Kliniekstraat 25 Research Group Terrestrial Ecology (TEREC) Ghent University, K.L.Ledeganckstraat 35 Entomology Unit Royal Belgian Institute for Natural Sciences (RBINS), Vautierstraat 29 USDA Forest Service Rocky Mountain Research Station Forestry Sciences Laboratory, 1648 S. 7th Avenue Department of Biological Sciences Bishop’s University, 2600 College Street UNESP - Univ Estadual Paulista Faculdade de Ciências Agrárias e Veterinárias Departamento de Morfologia e Fisiologia Animal; Via de Acesso Prof. Paulo Donato Castellane s/n Canadian National Collection of Insects & Canadian Food Inspection Agency OPL-Entomology, K.W. Neatby Bldg., C.E.F., 960 Carling Ave. Department of Biological Sciences University of Calgary, 2500 University Drive NW Finnish Museum of Natural History Zoology Unit University of Helsinki Washington State University, 2533 Inter Avenue Bohart Museum of Entomology University of California, One Shields Avenue Instituto Nacional de Biodiversidad (INBio) UNESP - Univ Estadual Paulista Faculdade de Ciências Agrárias e Veterinárias Departamento de Morfologia e Fisiologia Animal; Via de Acesso Prof. Paulo Donato Castellane s/n

The subfamily Clinocerinae (Empididae, Diptera) are aquatic dance flies, most commonly encountered in fast flowing rocky streams and creeks. The adults often rest on emergent rocks and are rather common in temperate regions.

One new species of Afrothaumalea is described from South Africa (A. stuckenbergi sp. n.). The pupa of Afrothaumalea is described for the first time and a key to all three species of Afrothaumalea is provided.

The Nearctic species of Calliphora Robineau-Desvoidy are revised and all species are redescribed and/or diagnosed. Diagnostic characters to permit reliable identification of both sexes of Calliphora aldrichia (Shannon) and C. montana (Shannon) and detailed distributional records for both species are provided for the first time. A lectotype is designated for Calliphora loewi Enderlein, 1903. A revised key to the 13 species of Nearctic Calliphora is also included. The key is based on examination of over 1,000 specimens from across North America and the structure of the terminalia of both sexes of each species. Complete illustrations of the terminalia of both sexes are provided for all species, including those of eight poorly known species: Calliphora alaskensis (Shannon), C. aldrichia (Shannon), C. coloradensis Hough, C. grahami Aldrich, C. latifrons Hough, C. livida Hall, C. montana (Shannon) and C. terraenovae Macquart. The female terminalia of C. alaskensis , C. aldrichia , C. coloradensis , C. livida and C. montana are illustrated for the first time. Barcode data for all 13 species of Nearctic Calliphora are provided, several for the first time. Results support current species concepts but barcodes failed to distinguish C. aldrichia and C. montana .

The North American Plant Protection Organization (NAPPO) is an organization comprising plant protection regulatory officials of the three signatory countries: the United States, Canada, and Mexico. NAPPO develops Regional Standards for Phytosanitary Measures (RSPMs) as well as discussion papers on important issues related to plant protection. The Fruit Panel of NAPPO organized a Technical Assistance Group (TAG) to develop a discussion paper on the status of pest species of Rhagoletis (Diptera: Tephritidae) within the NAPPO countries. As regulations are developed to prevent the spread of these pests both within and outside of the NAPPO countries, it was important to understand the pest status of these species. This article reviews the basic biology, host range, distribution, potential to spread, management, and regulatory status of pest species of Rhagoletis within the NAPPO countries.

A new species of gall midge, Stenodiplosis phragmicola Sinclair and Ahee (Diptera: Cecidomyiidae), is described. The host plant, Phragmites australis (Cavanilles) Trinius ex Steudel (Poaceae), is a tall, widely distributed emergent aquatic macrophyte. An introduced subspecies of the plant is considered invasive in North America (although a native subspecies also occurs). Insect specimens were collected during September 2010 and 2011 from the florets of common reed at 12 sites around Peterborough, Ontario, Canada. Preliminary data on ecological interactions between the plant and the insect are presented. Out of 2400 florets sampled, 9.5% were host to larvae of S. phragmicola. Three sites had much higher rates of infestation, with between 20% and 30% florets, and up to 100% of sampled shoots containing fly larvae. The largest stands in the sample all hosted fly populations and there was a positive association between the reproductive output of P. australis stands (measured as the average inflorescence mass per stand) and the proportion of shoots per stand containing larvae. The occurrence of S. phragmicola on the native subspecies of P. australis is also documented. Insects that consume reproductive structures of the common reed have not previously been reported from the plant's introduced range in North America.

The male terminalia character system in Diptera is reviewed. The phylogenetic implications of apomorphic changes are traced on published cladograms. New synapomorphies include: anteroventral parameral apodeme for the Tipulomorpha; parameral sheath encompassing desclerotized aedeagus for Neodiptera (exclusive of Axymyiidae); endoaedeagus for Xylophagomorpha + Tabanomorpha. Apystomyiidae are classified as the sister group to the Eremoneura based on four synapomorphies (lateral ejaculatory processes absent, subepandrial sclerite extending from base of hypoproct to phallus, bacilliform sclerites extending to tips of the epandrium and surstyli functionally developed, but not articulated) and lack of eremoneuran synapomorphies (i.e., loss of gonostyli, presence of postgonites and phallic plate). The Diptera sperm pump with a functional ejaculatory apodeme is a possible autapomorphy of Diptera, exclusive of Nymphomyiidae and Deuterophlebiidae. Internal details of the male terminalia ofSylvicolaandMycetobia(Anisopodidae),Hilarimorpha(Hilarimorphidae) andApystomyia(Apystomyiidae) are newly illustrated and homologies of the aedeagus, paramere and sperm pump of the Tipuloidea are clarified.

Three genera (Isodrapetis Collin, Chersodromia Walker, Elaphropeza Macquart) and six species of Tachydromiinae (Hybotidae) are recorded from the Galápagos Islands. One new species of Isodrapetis (I. meridionalis sp. n.) and three new species of Chersodromia (C. floreana sp. n., C. galapagensis sp. n., C. isabela sp. n.) are described. Elaphropeza zonalis (Curran) is redescribed and a fourth species of Chersodromia is identified based on a single female specimen. A key to all Galápagos species is provided. Isodrapetis is recorded for the first time outside of New Zealand. The affinities of the Galápagos Tachydromiinae to the New World fauna are discussed.