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The study of "Crioceras" barremense Kilian was undertaken as a part of the revision of the Hemihoplitidae. This species was considered "classic" and has been used as the index of an Upper Barremian subzone; this usage raises a number of problems. The type specimen from Tyrol was a fragment described and illustrated by Uhlig as Crioceras sp. ind. aff. roemeri. This specimen could not be retrieved, and a topotype could not be collected. Our study revealed that there is both a biostratigraphic hiatus and important differences between conceptions of this species: (1) that ascribed Uhlig's type specimen (Upper Barremian, Tyrol), (2) Kilian's concept of the specimen he found and named "Crioceras" barremense (probably a Camereiceras from the uppermost levels of the Vandenheckei Subzone or from the basal Sartousiana subzone of the Nauvin site, southeastern France) and (3) current interpretations of authors, who often synonymize the type specimen with Gassendiceras alpinum (d'Orbigny), which occurs in the middle of the Vandenheckei Subzone. So there is a real confusion concerning the synonymy of "Crioceras" barremense. The age of Uhlig's type specimen is too imprecise and its preservation too fragmentary to be reliably identifiable, because the same morphology and ornamentation exist in several species of other genera. Therefore, we recommend the use of the species "Crioceras" barremense Kilian be avoided, in particular as an index, along with that of the genus Barrancyloceras Vermeulen for which "C." barremense is used as reference. Some species formerly assigned to this genus have been referred to the genus Gassendiceras Bert et alii. Consequently, we also recommend the Barremense auctorum Subzone be renamed the Alpinum Subzone (new) [index-species: Gassendiceras alpinum (d'Orbigny)], without changing its limits as currently defined. The lower limit of this subzone is indicated by the first occurrence of Gassendiceras alpinum (a new biohorizon, introduced here), a common, easily identifiable species with a well-defined stratigraphic range.

A new ammonite biozonal scheme is proposed for the Upper Barremian of southeastern France. It takes into account faunal changes, sequence stratigraphy and historical data. Four new biohorizons are proposed. At present, three (formerly six) are accepted for the Upper Barremian: the Vandenheckei, Sartousiana and Giraudi biozones. The first contains three subzones, the Uhligi (current status discussed), the Sayni and the Barremense subzones, and two new biohorizons, the Marchandi and Breistrofferi biohorizons (top of the Barremense Subzone). The Sartousiana Biozone now includes the Limentinus, Provincialis and Feraudianus subzones. This last one lost its status of biozone because of paleontologic and stratigraphic arguments. Six biohorizons are recognized in the Sartousiana Biozone: the Limentinus, Provincialis, Casanovai, Feraudianus, Magnini and Autrani biohorizons. The Giraudi Biozone includes the Giraudi, Sarasini and Waagenoides subzones and the Giraudi, Emerici and Puzosianum biohorizons.

Phylogenetic reconstruction of the Upper Barremian ammonite genus Gassendiceras (Gassendiceratinae) was performed using a cladistic analysis incorporating continuous data. Some morphological features were found to vary identically among all the analysed species and therefore carry no phylogenetic information (= symplesiomorphic). The single obtained cladogram allows interpreting the evolution of the Gassendiceras as an anagenetic succession of eight species, in stratigraphic order of appearance, Gassendiceras multicostatum, G. alpinum, G. hoheneggeri, G. rebouleti, G. bosellii, G. quelquejeui, G. coulletae and G. enayi. The clade Pseudoshasticrioceras/ Imerites is derived from G. enayi, so the genus Gassendiceras appears to be paraphyletic. But here, we accept this fact as the best evolutive classification. The evolution over time of Gassendiceras is modulated by some processes, which could have constrained the inferred phylogenetic pattern with the drift of the global variability towards the most gracile forms over time. It is tempting to interpret this evolution as a constant selection over time of the Gassendiceras modulated by environmental control due to eustatic variation across a transgressive sequence. Thus, the most peramorphic (gracile) individuals seemed favoured at the expense of those most robust (paedomorphic). [ABSTRACT FROM AUTHOR]