108 results on '"Basílio, Daniel"'
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2. Catálogo Taxonômico da Fauna do Brasil: setting the baseline knowledge on the animal diversity in Brazil
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Boeger, Walter A, Valim, Michel P, Zaher, Hussam, Rafael, José A, Forzza, Rafaela C, Percequillo, Alexandre R, Serejo, Cristiana S, Garraffoni, André RS, Santos, Adalberto J, Slipinski, Adam, Linzmeier, Adelita M, Calor, Adolfo R, Garda, Adrian A, Kury, Adriano B, Fernandes, Agatha CS, Agudo-Padrón, Aisur I, Akama, Alberto, Silva Neto, Alberto M da, Burbano, Alejandro L, Menezes, Aleksandra, Pereira-Colavite, Alessandre, Anichtchenko, Alexander, Lees, Alexander C, Bezerra, Alexandra MR, Domahovski, Alexandre C, Pimenta, Alexandre D, Aleixo, Alexandre LP, Marceniuk, Alexandre P, Paula, Alexandre S de, Somavilla, Alexandre, Specht, Alexandre, Camargo, Alexssandro, Newton, Alfred F, Silva, Aline AS da, Santos, Aline B dos, Tassi, Aline D, Aragão, Allan C, Santos, Allan PM, Migotto, Alvaro E, Mendes, Amanda C, Cunha, Amanda, Chagas Júnior, Amazonas, Sousa, Ana AT de, Pavan, Ana C, Almeida, Ana CS, Peronti, Ana LBG, Henriques-Oliveira, Ana L, Prudente, Ana L, Tourinho, Ana L, Pes, Ana MO, Carmignotto, Ana P, Wengrat, Ana PG da Silva, Dornellas, Ana PS, Molin, Anamaria Dal, Puker, Anderson, Morandini, André C, Ferreira, André da S, Martins, André L, Esteves, André M, Fernandes, André S, Roza, André S, Köhler, Andreas, Paladini, Andressa, Andrade, Andrey J de, Pinto, Ângelo P, Salles, Anna C de A, Gondim, Anne I, Amaral, Antonia CZ, Rondón, Antonio AA, Brescovit, Antonio, Lofego, Antônio C, Marques, Antonio C, Macedo, Antonio, Andriolo, Artur, Henriques, Augusto L, Ferreira Júnior, Augusto L, Lima, Aurino F de, Barros, Ávyla R de A, Brito, Ayrton do R, Romera, Bárbara LV, Vasconcelos, Beatriz MC de, Frable, Benjamin W, Santos, Bernardo F, Ferraz, Bernardo R, Rosa, Brunno B, Sampaio, Brunno HL, Bellini, Bruno C, Clarkson, Bruno, Oliveira, Bruno G de, Corrêa, Caio CD, Martins, Caleb C, Castro-Guedes, Camila F de, Souto, Camilla, Bicho, Carla de L, Cunha, Carlo M, Barboza, Carlos A de M, Lucena, Carlos AS de, Barreto, Carlos, Santana, Carlos DCM de, Agne, Carlos EQ, Mielke, Carlos GC, Caetano, Carlos HS, Flechtmann, Carlos HW, Lamas, Carlos JE, Rocha, Carlos, Mascarenhas, Carolina S, Margaría, Cecilia B, Waichert, Cecilia, Digiani, Celina, Haddad, Célio FB, Azevedo, Celso O, Benetti, Cesar J, Santos, Charles MD dos, Bartlett, Charles R, Bonvicino, Cibele, Ribeiro-Costa, Cibele S, Santos, Cinthya SG, Justino, Cíntia EL, Canedo, Clarissa, Bonecker, Claudia C, Santos, Cláudia P, Carvalho, Claudio JB de, Gonçalves, Clayton C, Galvão, Cleber, Costa, Cleide, Oliveira, Cléo DC de, Schwertner, Cristiano F, Andrade, Cristiano L, Pereira, Cristiano M, Sampaio, Cristiano, Dias, Cristina de O, Lucena, Daercio A de A, Manfio, Daiara, Amorim, Dalton de S, Queiroz, Dalva L de, Colpani, Daniara, Abbate, Daniel, Aquino, Daniel A, Burckhardt, Daniel, Cavallari, Daniel C, Prado, Daniel de C Schelesky, Praciano, Daniel L, Basílio, Daniel S, Bená, Daniela de C, Toledo, Daniela GP de, Takiya, Daniela M, Fernandes, Daniell RR, Ament, Danilo C, Cordeiro, Danilo P, Silva, Darliane E, Pollock, Darren A, Muniz, David B, Gibson, David I, Nogueira, David S, Marques, Dayse WA, Lucatelli, Débora, Garcia, Deivys MA, Baêta, Délio, Ferreira, Denise NM, Rueda-Ramírez, Diana, Fachin, Diego A, Souza, Diego de S, Rodrigues, Diego F, Pádua, Diego G de, Barbosa, Diego N, Dolibaina, Diego R, Amaral, Diogo C, Chandler, Donald S, Maccagnan, Douglas HB, Caron, Edilson, Carvalho, Edrielly, Adriano, Edson A, Abreu Júnior, Edson F de, Pereira, Edson HL, Viegas, Eduarda FG, Carneiro, Eduardo, Colley, Eduardo, Eizirik, Eduardo, Santos, Eduardo F dos, Shimbori, Eduardo M, Suárez-Morales, Eduardo, Arruda, Eliane P de, Chiquito, Elisandra A, Lima, Élison FB, Castro, Elizeu B de, Orlandin, Elton, Nascimento, Elynton A do, Razzolini, Emanuel, Gama, Emanuel RR, Araujo, Enilma M de, Nishiyama, Eric Y, Spiessberger, Erich L, Santos, Érika CL dos, Contreras, Eugenia F, Galati, Eunice AB, Oliveira Junior, Evaldo C de, Gallardo, Fabiana, Hernandes, Fabio A, Lansac-Tôha, Fábio A, Pitombo, Fabio B, Dario, Fabio Di, Santos, Fábio L dos, Mauro, Fabio, Nascimento, Fabio O do, Olmos, Fabio, Amaral, Fabio R, Schunck, Fabio, Godoi, Fábio SP de, Machado, Fabrizio M, Barbo, Fausto E, Agrain, Federico A, Ribeiro, Felipe B, Moreira, Felipe FF, Barbosa, Felipe F, Silva, Fenanda S, Cavalcanti, Fernanda F, Straube, Fernando C, Carbayo, Fernando, Carvalho Filho, Fernando, Zanella, Fernando CV, Jacinavicius, Fernando de C, Farache, Fernando HA, Leivas, Fernando, Dias, Fernando MS, Mantellato, Fernando, Vaz-de-Mello, Fernando Z, Gudin, Filipe M, Albuquerque, Flávio, Molina, Flavio B, Passos, Flávio D, Shockley, Floyd W, Pinheiro, Francielly F, Mello, Francisco de AG de, Nascimento, Francisco E de L, Franco, Francisco L, Oliveira, Francisco L de, Melo, Francisco T de V, Quijano, Freddy RB, Salles, Frederico F, Biffi, Gabriel, Queiroz, Gabriel C, Bizarro, Gabriel L, Hrycyna, Gabriela, Leviski, Gabriela, Powell, Gareth S, Santos, Geane B dos, Morse, Geoffrey E, Brown, George, Mattox, George MT, Zimbrão, Geraldo, Carvalho, Gervásio S, Miranda, Gil FG, Moraes, Gilberto J de, Lourido, Gilcélia M, Neves, Gilmar P, Moreira, Gilson RP, Montingelli, Giovanna G, Maurício, Giovanni N, Marconato, Gláucia, Lopez, Guilherme EL, Silva, Guilherme L da, Muricy, Guilherme, Brito, Guilherme RR, Garbino, Guilherme ST, Flores, Gustavo E, Graciolli, Gustavo, Libardi, Gustavo S, Proctor, Heather C, Gil-Santana, Helcio R, Varella, Henrique R, Escalona, Hermes E, Schmitz, Hermes J, Rodrigues, Higor DD, Galvão Filho, Hilton de C, Quintino, Hingrid YS, Pinto, Hudson A, Rainho, Hugo L, Miyahira, Igor C, Gonçalves, Igor de S, Martins, Inês X, Cardoso, Irene A, Oliveira, Ismael B de, Franz, Ismael, Fernandes, Itanna O, Golfetti, Ivan F, S. Campos-Filho, Ivanklin, Oliveira, Ivo de S, Delabie, Jacques HC, Oliveira, Jader de, Prando, Jadila S, Patton, James L, Bitencourt, Jamille de A, Silva, Janaina M, Santos, Jandir C, Arruda, Janine O, Valderrama, Jefferson S, Dalapicolla, Jeronymo, Oliveira, Jéssica P, Hájek, Jiri, Morselli, João P, Narita, João P, Martin, João PI, Grazia, Jocélia, McHugh, Joe, Cherem, Jorge J, Farias Júnior, José AS, Fernandes, Jose AM, Pacheco, José F, Birindelli, José LO, Rezende, José M, Avendaño, Jose M, Duarte, José M Barbanti, Ribeiro, José R Inácio, Mermudes, José RM, Pujol-Luz, José R, Santos, Josenilson R dos, Câmara, Josenir T, Teixeira, Joyce A, Prado, Joyce R do, Botero, Juan P, Almeida, Julia C, Kohler, Julia, Gonçalves, Julia P, Beneti, Julia S, Donahue, Julian P, Alvim, Juliana, Almeida, Juliana C, Segadilha, Juliana L, Wingert, Juliana M, Barbosa, Julianna F, Ferrer, Juliano, Santos, Juliano F dos, Kuabara, Kamila MD, Nascimento, Karine B, Schoeninger, Karine, Campião, Karla M, Soares, Karla, Zilch, Kássia, Barão, Kim R, Teixeira, Larissa, Sousa, Laura D do NM de, Dumas, Leandro L, Vieira, Leandro M, Azevedo, Leonardo HG, Carvalho, Leonardo S, Souza, Leonardo S de, Rocha, Leonardo SG, Bernardi, Leopoldo FO, Vieira, Letícia M, Johann, Liana, Salvatierra, Lidianne, Oliveira, Livia de M, Loureiro, Lourdes MA El-moor, Barreto, Luana B, Barros, Luana M, Lecci, Lucas, Camargos, Lucas M de, Lima, Lucas RC, Almeida, Lucia M, Martins, Luciana R, Marinoni, Luciane, Moura, Luciano de A, Lima, Luciano, Naka, Luciano N, Miranda, Lucília S, Salik, Lucy M, Bezerra, Luis EA, Silveira, Luis F, Campos, Luiz A, Castro, Luiz AS de, Pinho, Luiz C, Silveira, Luiz FL, Iniesta, Luiz FM, Tencatt, Luiz FC, Simone, Luiz RL, Malabarba, Luiz R, Cruz, Luiza S da, Sekerka, Lukas, Barros, Lurdiana D, Santos, Luziany Q, Skoracki, Maciej, Correia, Maira A, Uchoa, Manoel A, Andrade, Manuella FG, Hermes, Marcel G, Miranda, Marcel S, Araújo, Marcel S de, Monné, Marcela L, Labruna, Marcelo B, Santis, Marcelo D de, Duarte, Marcelo, Knoff, Marcelo, Nogueira, Marcelo, Britto, Marcelo R de, Melo, Marcelo RS de, Carvalho, Marcelo R de, Tavares, Marcelo T, Kitahara, Marcelo V, Justo, Marcia CN, Botelho, Marcia JC, Couri, Márcia S, Borges-Martins, Márcio, Felix, Márcio, Oliveira, Marcio L de, Bologna, Marco A, Gottschalk, Marco S, Tavares, Marcos DS, Lhano, Marcos G, Bevilaqua, Marcus, Santos, Marcus TT, Domingues, Marcus V, Sallum, Maria AM, Digiani, María C, Santarém, Maria CA, Nascimento, Maria C do, Becerril, María de los AM, Santos, Maria EA dos, Passos, Maria I da S dos, Felippe-Bauer, Maria L, Cherman, Mariana A, Terossi, Mariana, Bartz, Marie LC, Barbosa, Marina F de C, Loeb, Marina V, Cohn-Haft, Mario, Cupello, Mario, Martins, Marlúcia B, Christofersen, Martin L, Bento, Matheus, Rocha, Matheus dos S, Martins, Maurício L, Segura, Melissa O, Cardenas, Melissa Q, Duarte, Mércia E, Ivie, Michael A, Mincarone, Michael M, Borges, Michela, Monné, Miguel A, Casagrande, Mirna M, Fernandez, Monica A, Piovesan, Mônica, Menezes, Naércio A, Benaim, Natalia P, Reategui, Natália S, Pedro, Natan C, Pecly, Nathalia H, Ferreira Júnior, Nelson, Silva Júnior, Nelson J da, Perioto, Nelson W, Hamada, Neusa, Degallier, Nicolas, Chao, Ning L, Ferla, Noeli J, Mielke, Olaf HH, Evangelista, Olivia, Shibatta, Oscar A, Oliveira, Otto MP, Albornoz, Pablo CL, Dellapé, Pablo M, Gonçalves, Pablo R, Shimabukuro, Paloma HF, Grossi, Paschoal, Rodrigues, Patrícia E da S, Lima, Patricia OV, Velazco, Paul, Santos, Paula B dos, Araújo, Paula B, Silva, Paula KR, Riccardi, Paula R, Garcia, Paulo C de A, Passos, Paulo GH, Corgosinho, Paulo HC, Lucinda, Paulo, Costa, Paulo MS, Alves, Paulo P, Roth, Paulo R de O, Coelho, Paulo RS, Duarte, Paulo RM, Carvalho, Pedro F de, Gnaspini, Pedro, Souza-Dias, Pedro GB, Linardi, Pedro M, Bartholomay, Pedro R, Demite, Peterson R, Bulirsch, Petr, Boll, Piter K, Pereira, Rachel MM, Silva, Rafael APF, Moura, Rafael B de, Boldrini, Rafael, Silva, Rafaela A da, Falaschi, Rafaela L, Cordeiro, Ralf TS, Mello, Ramon JCL, Singer, Randal A, Querino, Ranyse B, Heleodoro, Raphael A, Castilho, Raphael de C, Constantino, Reginaldo, Guedes, Reinaldo C, Carrenho, Renan, Gomes, Renata S, Gregorin, Renato, Machado, Renato JP, Bérnils, Renato S, Capellari, Renato S, Silva, Ricardo B, Kawada, Ricardo, Dias, Ricardo M, Siewert, Ricardo, Brugnera, Ricaro, Leschen, Richard AB, Constantin, Robert, Robbins, Robert, Pinto, Roberta R, Reis, Roberto E dos, Ramos, Robson T da C, Cavichioli, Rodney R, Barros, Rodolfo C de, Caires, Rodrigo A, Salvador, Rodrigo B, Marques, Rodrigo C, Araújo, Rodrigo C, Araujo, Rodrigo de O, Dios, Rodrigo de VP, Johnsson, Rodrigo, Feitosa, Rodrigo M, Hutchings, Roger W, Lara, Rogéria IR, Rossi, Rogério V, Gerstmeier, Roland, Ochoa, Ronald, Hutchings, Rosa SG, Ale-Rocha, Rosaly, Rocha, Rosana M da, Tidon, Rosana, Brito, Rosangela, Pellens, Roseli, Santos, Sabrina R dos, Santos, Sandra D dos, Paiva, Sandra V, Santos, Sandro, Oliveira, Sarah S de, Costa, Sávio C, Gardner, Scott L, Leal, Sebastián A Muñoz, Aloquio, Sergio, Bonecker, Sergio LC, Bueno, Sergio L de S, Almeida, Sérgio M de, Stampar, Sérgio N, Andena, Sérgio R, Posso, Sergio R, Lima, Sheila P, Gadelha, Sian de S, Thiengo, Silvana C, Cohen, Simone C, Brandão, Simone N, Rosa, Simone P, Ribeiro, Síria LB, Letana, Sócrates D, Santos, Sonia B dos, Andrade, Sonia CS, Dávila, Stephane, Vaz, Stéphanie, Peck, Stewart B, Christo, Susete W, Cunha, Suzan BZ, Gomes, Suzete R, Duarte, Tácio, Madeira-Ott, Taís, Marques, Taísa, Roell, Talita, Lima, Tarcilla C de, Sepulveda, Tatiana A, Maria, Tatiana F, Ruschel, Tatiana P, Rodrigues, Thaiana, Marinho, Thais A, Almeida, Thaís M de, Miranda, Thaís P, Freitas, Thales RO, Pereira, Thalles PL, Zacca, Thamara, Pacheco, Thaynara L, Martins, Thiago F, Alvarenga, Thiago M, Carvalho, Thiago R de, Polizei, Thiago TS, McElrath, Thomas C, Henry, Thomas, Pikart, Tiago G, Porto, Tiago J, Krolow, Tiago K, Carvalho, Tiago P, Lotufo, Tito M da C, Caramaschi, Ulisses, Pinheiro, Ulisses dos S, Pardiñas, Ulyses FJ, Maia, Valéria C, Tavares, Valeria, Costa, Valmir A, Amaral, Vanessa S do, Silva, Vera C, Wolff, Vera R dos S, Slobodian, Verônica, Silva, Vinícius B da, Espíndola, Vinicius C, Costa-Silva, Vinicius da, Bertaco, Vinicius de A, Padula, Vinícius, Ferreira, Vinicius S, Silva, Vitor CP da, Piacentini, Vítor de Q, Sandoval-Gómez, Vivian E, Trevine, Vivian, Sousa, Viviane R, Sant’Anna, Vivianne B de, Mathis, Wayne N, Souza, Wesley de O, Colombo, Wesley D, Tomaszewska, Wioletta, Wosiacki, Wolmar B, Ovando, Ximena MC, Leite, Yuri LR, Boeger, Walter A, Valim, Michel P, Zaher, Hussam, Rafael, José A, Forzza, Rafaela C, Percequillo, Alexandre R, Serejo, Cristiana S, Garraffoni, André RS, Santos, Adalberto J, Slipinski, Adam, Linzmeier, Adelita M, Calor, Adolfo R, Garda, Adrian A, Kury, Adriano B, Fernandes, Agatha CS, Agudo-Padrón, Aisur I, Akama, Alberto, Silva Neto, Alberto M da, Burbano, Alejandro L, Menezes, Aleksandra, Pereira-Colavite, Alessandre, Anichtchenko, Alexander, Lees, Alexander C, Bezerra, Alexandra MR, Domahovski, Alexandre C, Pimenta, Alexandre D, Aleixo, Alexandre LP, Marceniuk, Alexandre P, Paula, Alexandre S de, Somavilla, Alexandre, Specht, Alexandre, Camargo, Alexssandro, Newton, Alfred F, Silva, Aline AS da, Santos, Aline B dos, Tassi, Aline D, Aragão, Allan C, Santos, Allan PM, Migotto, Alvaro E, Mendes, Amanda C, Cunha, Amanda, Chagas Júnior, Amazonas, Sousa, Ana AT de, Pavan, Ana C, Almeida, Ana CS, Peronti, Ana LBG, Henriques-Oliveira, Ana L, Prudente, Ana L, Tourinho, Ana L, Pes, Ana MO, Carmignotto, Ana P, Wengrat, Ana PG da Silva, Dornellas, Ana PS, Molin, Anamaria Dal, Puker, Anderson, Morandini, André C, Ferreira, André da S, Martins, André L, Esteves, André M, Fernandes, André S, Roza, André S, Köhler, Andreas, Paladini, Andressa, Andrade, Andrey J de, Pinto, Ângelo P, Salles, Anna C de A, Gondim, Anne I, Amaral, Antonia CZ, Rondón, Antonio AA, Brescovit, Antonio, Lofego, Antônio C, Marques, Antonio C, Macedo, Antonio, Andriolo, Artur, Henriques, Augusto L, Ferreira Júnior, Augusto L, Lima, Aurino F de, Barros, Ávyla R de A, Brito, Ayrton do R, Romera, Bárbara LV, Vasconcelos, Beatriz MC de, Frable, Benjamin W, Santos, Bernardo F, Ferraz, Bernardo R, Rosa, Brunno B, Sampaio, Brunno HL, Bellini, Bruno C, Clarkson, Bruno, Oliveira, Bruno G de, Corrêa, Caio CD, Martins, Caleb C, Castro-Guedes, Camila F de, Souto, Camilla, Bicho, Carla de L, Cunha, Carlo M, Barboza, Carlos A de M, Lucena, Carlos AS de, Barreto, Carlos, Santana, Carlos DCM de, Agne, Carlos EQ, Mielke, Carlos GC, Caetano, Carlos HS, Flechtmann, Carlos HW, Lamas, Carlos JE, Rocha, Carlos, Mascarenhas, Carolina S, Margaría, Cecilia B, Waichert, Cecilia, Digiani, Celina, Haddad, Célio FB, Azevedo, Celso O, Benetti, Cesar J, Santos, Charles MD dos, Bartlett, Charles R, Bonvicino, Cibele, Ribeiro-Costa, Cibele S, Santos, Cinthya SG, Justino, Cíntia EL, Canedo, Clarissa, Bonecker, Claudia C, Santos, Cláudia P, Carvalho, Claudio JB de, Gonçalves, Clayton C, Galvão, Cleber, Costa, Cleide, Oliveira, Cléo DC de, Schwertner, Cristiano F, Andrade, Cristiano L, Pereira, Cristiano M, Sampaio, Cristiano, Dias, Cristina de O, Lucena, Daercio A de A, Manfio, Daiara, Amorim, Dalton de S, Queiroz, Dalva L de, Colpani, Daniara, Abbate, Daniel, Aquino, Daniel A, Burckhardt, Daniel, Cavallari, Daniel C, Prado, Daniel de C Schelesky, Praciano, Daniel L, Basílio, Daniel S, Bená, Daniela de C, Toledo, Daniela GP de, Takiya, Daniela M, Fernandes, Daniell RR, Ament, Danilo C, Cordeiro, Danilo P, Silva, Darliane E, Pollock, Darren A, Muniz, David B, Gibson, David I, Nogueira, David S, Marques, Dayse WA, Lucatelli, Débora, Garcia, Deivys MA, Baêta, Délio, Ferreira, Denise NM, Rueda-Ramírez, Diana, Fachin, Diego A, Souza, Diego de S, Rodrigues, Diego F, Pádua, Diego G de, Barbosa, Diego N, Dolibaina, Diego R, Amaral, Diogo C, Chandler, Donald S, Maccagnan, Douglas HB, Caron, Edilson, Carvalho, Edrielly, Adriano, Edson A, Abreu Júnior, Edson F de, Pereira, Edson HL, Viegas, Eduarda FG, Carneiro, Eduardo, Colley, Eduardo, Eizirik, Eduardo, Santos, Eduardo F dos, Shimbori, Eduardo M, Suárez-Morales, Eduardo, Arruda, Eliane P de, Chiquito, Elisandra A, Lima, Élison FB, Castro, Elizeu B de, Orlandin, Elton, Nascimento, Elynton A do, Razzolini, Emanuel, Gama, Emanuel RR, Araujo, Enilma M de, Nishiyama, Eric Y, Spiessberger, Erich L, Santos, Érika CL dos, Contreras, Eugenia F, Galati, Eunice AB, Oliveira Junior, Evaldo C de, Gallardo, Fabiana, Hernandes, Fabio A, Lansac-Tôha, Fábio A, Pitombo, Fabio B, Dario, Fabio Di, Santos, Fábio L dos, Mauro, Fabio, Nascimento, Fabio O do, Olmos, Fabio, Amaral, Fabio R, Schunck, Fabio, Godoi, Fábio SP de, Machado, Fabrizio M, Barbo, Fausto E, Agrain, Federico A, Ribeiro, Felipe B, Moreira, Felipe FF, Barbosa, Felipe F, Silva, Fenanda S, Cavalcanti, Fernanda F, Straube, Fernando C, Carbayo, Fernando, Carvalho Filho, Fernando, Zanella, Fernando CV, Jacinavicius, Fernando de C, Farache, Fernando HA, Leivas, Fernando, Dias, Fernando MS, Mantellato, Fernando, Vaz-de-Mello, Fernando Z, Gudin, Filipe M, Albuquerque, Flávio, Molina, Flavio B, Passos, Flávio D, Shockley, Floyd W, Pinheiro, Francielly F, Mello, Francisco de AG de, Nascimento, Francisco E de L, Franco, Francisco L, Oliveira, Francisco L de, Melo, Francisco T de V, Quijano, Freddy RB, Salles, Frederico F, Biffi, Gabriel, Queiroz, Gabriel C, Bizarro, Gabriel L, Hrycyna, Gabriela, Leviski, Gabriela, Powell, Gareth S, Santos, Geane B dos, Morse, Geoffrey E, Brown, George, Mattox, George MT, Zimbrão, Geraldo, Carvalho, Gervásio S, Miranda, Gil FG, Moraes, Gilberto J de, Lourido, Gilcélia M, Neves, Gilmar P, Moreira, Gilson RP, Montingelli, Giovanna G, Maurício, Giovanni N, Marconato, Gláucia, Lopez, Guilherme EL, Silva, Guilherme L da, Muricy, Guilherme, Brito, Guilherme RR, Garbino, Guilherme ST, Flores, Gustavo E, Graciolli, Gustavo, Libardi, Gustavo S, Proctor, Heather C, Gil-Santana, Helcio R, Varella, Henrique R, Escalona, Hermes E, Schmitz, Hermes J, Rodrigues, Higor DD, Galvão Filho, Hilton de C, Quintino, Hingrid YS, Pinto, Hudson A, Rainho, Hugo L, Miyahira, Igor C, Gonçalves, Igor de S, Martins, Inês X, Cardoso, Irene A, Oliveira, Ismael B de, Franz, Ismael, Fernandes, Itanna O, Golfetti, Ivan F, S. Campos-Filho, Ivanklin, Oliveira, Ivo de S, Delabie, Jacques HC, Oliveira, Jader de, Prando, Jadila S, Patton, James L, Bitencourt, Jamille de A, Silva, Janaina M, Santos, Jandir C, Arruda, Janine O, Valderrama, Jefferson S, Dalapicolla, Jeronymo, Oliveira, Jéssica P, Hájek, Jiri, Morselli, João P, Narita, João P, Martin, João PI, Grazia, Jocélia, McHugh, Joe, Cherem, Jorge J, Farias Júnior, José AS, Fernandes, Jose AM, Pacheco, José F, Birindelli, José LO, Rezende, José M, Avendaño, Jose M, Duarte, José M Barbanti, Ribeiro, José R Inácio, Mermudes, José RM, Pujol-Luz, José R, Santos, Josenilson R dos, Câmara, Josenir T, Teixeira, Joyce A, Prado, Joyce R do, Botero, Juan P, Almeida, Julia C, Kohler, Julia, Gonçalves, Julia P, Beneti, Julia S, Donahue, Julian P, Alvim, Juliana, Almeida, Juliana C, Segadilha, Juliana L, Wingert, Juliana M, Barbosa, Julianna F, Ferrer, Juliano, Santos, Juliano F dos, Kuabara, Kamila MD, Nascimento, Karine B, Schoeninger, Karine, Campião, Karla M, Soares, Karla, Zilch, Kássia, Barão, Kim R, Teixeira, Larissa, Sousa, Laura D do NM de, Dumas, Leandro L, Vieira, Leandro M, Azevedo, Leonardo HG, Carvalho, Leonardo S, Souza, Leonardo S de, Rocha, Leonardo SG, Bernardi, Leopoldo FO, Vieira, Letícia M, Johann, Liana, Salvatierra, Lidianne, Oliveira, Livia de M, Loureiro, Lourdes MA El-moor, Barreto, Luana B, Barros, Luana M, Lecci, Lucas, Camargos, Lucas M de, Lima, Lucas RC, Almeida, Lucia M, Martins, Luciana R, Marinoni, Luciane, Moura, Luciano de A, Lima, Luciano, Naka, Luciano N, Miranda, Lucília S, Salik, Lucy M, Bezerra, Luis EA, Silveira, Luis F, Campos, Luiz A, Castro, Luiz AS de, Pinho, Luiz C, Silveira, Luiz FL, Iniesta, Luiz FM, Tencatt, Luiz FC, Simone, Luiz RL, Malabarba, Luiz R, Cruz, Luiza S da, Sekerka, Lukas, Barros, Lurdiana D, Santos, Luziany Q, Skoracki, Maciej, Correia, Maira A, Uchoa, Manoel A, Andrade, Manuella FG, Hermes, Marcel G, Miranda, Marcel S, Araújo, Marcel S de, Monné, Marcela L, Labruna, Marcelo B, Santis, Marcelo D de, Duarte, Marcelo, Knoff, Marcelo, Nogueira, Marcelo, Britto, Marcelo R de, Melo, Marcelo RS de, Carvalho, Marcelo R de, Tavares, Marcelo T, Kitahara, Marcelo V, Justo, Marcia CN, Botelho, Marcia JC, Couri, Márcia S, Borges-Martins, Márcio, Felix, Márcio, Oliveira, Marcio L de, Bologna, Marco A, Gottschalk, Marco S, Tavares, Marcos DS, Lhano, Marcos G, Bevilaqua, Marcus, Santos, Marcus TT, Domingues, Marcus V, Sallum, Maria AM, Digiani, María C, Santarém, Maria CA, Nascimento, Maria C do, Becerril, María de los AM, Santos, Maria EA dos, Passos, Maria I da S dos, Felippe-Bauer, Maria L, Cherman, Mariana A, Terossi, Mariana, Bartz, Marie LC, Barbosa, Marina F de C, Loeb, Marina V, Cohn-Haft, Mario, Cupello, Mario, Martins, Marlúcia B, Christofersen, Martin L, Bento, Matheus, Rocha, Matheus dos S, Martins, Maurício L, Segura, Melissa O, Cardenas, Melissa Q, Duarte, Mércia E, Ivie, Michael A, Mincarone, Michael M, Borges, Michela, Monné, Miguel A, Casagrande, Mirna M, Fernandez, Monica A, Piovesan, Mônica, Menezes, Naércio A, Benaim, Natalia P, Reategui, Natália S, Pedro, Natan C, Pecly, Nathalia H, Ferreira Júnior, Nelson, Silva Júnior, Nelson J da, Perioto, Nelson W, Hamada, Neusa, Degallier, Nicolas, Chao, Ning L, Ferla, Noeli J, Mielke, Olaf HH, Evangelista, Olivia, Shibatta, Oscar A, Oliveira, Otto MP, Albornoz, Pablo CL, Dellapé, Pablo M, Gonçalves, Pablo R, Shimabukuro, Paloma HF, Grossi, Paschoal, Rodrigues, Patrícia E da S, Lima, Patricia OV, Velazco, Paul, Santos, Paula B dos, Araújo, Paula B, Silva, Paula KR, Riccardi, Paula R, Garcia, Paulo C de A, Passos, Paulo GH, Corgosinho, Paulo HC, Lucinda, Paulo, Costa, Paulo MS, Alves, Paulo P, Roth, Paulo R de O, Coelho, Paulo RS, Duarte, Paulo RM, Carvalho, Pedro F de, Gnaspini, Pedro, Souza-Dias, Pedro GB, Linardi, Pedro M, Bartholomay, Pedro R, Demite, Peterson R, Bulirsch, Petr, Boll, Piter K, Pereira, Rachel MM, Silva, Rafael APF, Moura, Rafael B de, Boldrini, Rafael, Silva, Rafaela A da, Falaschi, Rafaela L, Cordeiro, Ralf TS, Mello, Ramon JCL, Singer, Randal A, Querino, Ranyse B, Heleodoro, Raphael A, Castilho, Raphael de C, Constantino, Reginaldo, Guedes, Reinaldo C, Carrenho, Renan, Gomes, Renata S, Gregorin, Renato, Machado, Renato JP, Bérnils, Renato S, Capellari, Renato S, Silva, Ricardo B, Kawada, Ricardo, Dias, Ricardo M, Siewert, Ricardo, Brugnera, Ricaro, Leschen, Richard AB, Constantin, Robert, Robbins, Robert, Pinto, Roberta R, Reis, Roberto E dos, Ramos, Robson T da C, Cavichioli, Rodney R, Barros, Rodolfo C de, Caires, Rodrigo A, Salvador, Rodrigo B, Marques, Rodrigo C, Araújo, Rodrigo C, Araujo, Rodrigo de O, Dios, Rodrigo de VP, Johnsson, Rodrigo, Feitosa, Rodrigo M, Hutchings, Roger W, Lara, Rogéria IR, Rossi, Rogério V, Gerstmeier, Roland, Ochoa, Ronald, Hutchings, Rosa SG, Ale-Rocha, Rosaly, Rocha, Rosana M da, Tidon, Rosana, Brito, Rosangela, Pellens, Roseli, Santos, Sabrina R dos, Santos, Sandra D dos, Paiva, Sandra V, Santos, Sandro, Oliveira, Sarah S de, Costa, Sávio C, Gardner, Scott L, Leal, Sebastián A Muñoz, Aloquio, Sergio, Bonecker, Sergio LC, Bueno, Sergio L de S, Almeida, Sérgio M de, Stampar, Sérgio N, Andena, Sérgio R, Posso, Sergio R, Lima, Sheila P, Gadelha, Sian de S, Thiengo, Silvana C, Cohen, Simone C, Brandão, Simone N, Rosa, Simone P, Ribeiro, Síria LB, Letana, Sócrates D, Santos, Sonia B dos, Andrade, Sonia CS, Dávila, Stephane, Vaz, Stéphanie, Peck, Stewart B, Christo, Susete W, Cunha, Suzan BZ, Gomes, Suzete R, Duarte, Tácio, Madeira-Ott, Taís, Marques, Taísa, Roell, Talita, Lima, Tarcilla C de, Sepulveda, Tatiana A, Maria, Tatiana F, Ruschel, Tatiana P, Rodrigues, Thaiana, Marinho, Thais A, Almeida, Thaís M de, Miranda, Thaís P, Freitas, Thales RO, Pereira, Thalles PL, Zacca, Thamara, Pacheco, Thaynara L, Martins, Thiago F, Alvarenga, Thiago M, Carvalho, Thiago R de, Polizei, Thiago TS, McElrath, Thomas C, Henry, Thomas, Pikart, Tiago G, Porto, Tiago J, Krolow, Tiago K, Carvalho, Tiago P, Lotufo, Tito M da C, Caramaschi, Ulisses, Pinheiro, Ulisses dos S, Pardiñas, Ulyses FJ, Maia, Valéria C, Tavares, Valeria, Costa, Valmir A, Amaral, Vanessa S do, Silva, Vera C, Wolff, Vera R dos S, Slobodian, Verônica, Silva, Vinícius B da, Espíndola, Vinicius C, Costa-Silva, Vinicius da, Bertaco, Vinicius de A, Padula, Vinícius, Ferreira, Vinicius S, Silva, Vitor CP da, Piacentini, Vítor de Q, Sandoval-Gómez, Vivian E, Trevine, Vivian, Sousa, Viviane R, Sant’Anna, Vivianne B de, Mathis, Wayne N, Souza, Wesley de O, Colombo, Wesley D, Tomaszewska, Wioletta, Wosiacki, Wolmar B, Ovando, Ximena MC, and Leite, Yuri LR more...
- Abstract
The limited temporal completeness and taxonomic accuracy of species lists, made available in a traditional manner in scientific publications, has always represented a problem. These lists are invariably limited to a few taxonomic groups and do not represent up-to-date knowledge of all species and classifications. In this context, the Brazilian megadiverse fauna is no exception, and the Catálogo Taxonômico da Fauna do Brasil (CTFB) (http://fauna.jbrj.gov.br/), made public in 2015, represents a database on biodiversity anchored on a list of valid and expertly recognized scientific names of animals in Brazil. The CTFB is updated in near real time by a team of more than 800 specialists. By January 1, 2024, the CTFB compiled 133,691 nominal species, with 125,138 that were considered valid. Most of the valid species were arthropods (82.3%, with more than 102,000 species) and chordates (7.69%, with over 11,000 species). These taxa were followed by a cluster composed of Mollusca (3,567 species), Platyhelminthes (2,292 species), Annelida (1,833 species), and Nematoda (1,447 species). All remaining groups had less than 1,000 species reported in Brazil, with Cnidaria (831 species), Porifera (628 species), Rotifera (606 species), and Bryozoa (520 species) representing those with more than 500 species. Analysis of the CTFB database can facilitate and direct efforts towards the discovery of new species in Brazil, but it is also fundamental in providing the best available list of valid nominal species to users, including those in science, health, conservation efforts, and any initiative involving animals. The importance of the CTFB is evidenced by the elevated number of citations in the scientific literature in diverse areas of biology, law, anthropology, education, forensic science, and veterinary science, among others. more...
- Published
- 2024
Catalog
3. New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data.
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Cherman, Mariana A, Basílio, Daniel S, Clarkson, Bruno, Agostinis, André O, Smith, Andrew B T, Vaz-De-Mello, Fernando Z, and Almeida, Lúcia M
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SCARABAEIDAE , *SISTERS , *SPECIES , *PARSIMONIOUS models , *GENES - Abstract
The relationships among the Neotropical Diplotaxini and homology of revised and new taxa are revisited with parsimony analyses based on morphological characters, and probabilistic analyses based on concatenated ITS2 and COI genes. Neotropical Diplotaxini is now composed of five monophyletic genera: Pacuvia, Pachrodema, Homalochilus, Liogenys , and Careocallus gen. nov.. Pseudodiplotaxis is here transferred to incertae sedis of Melolonthinae. Careocallus is the sister genus of Liogenys and is composed of Careocallus tehuelche sp. nov. and Careocallus densicollis sp. nov. Sister phylogenetic relationship between Homalochilus and the rest of genera except Pacuvia is revealed. Homalochilus sensu nov. is comprised by Homalochilus punctatostriatus and Homalochilus nigripennis sp. nov. This genus recovered its monophyly by transferring Homalochilus niger to Liogenys , as Liogenys niger comb. nov. and this name is also synonymized with L. morio. The type species of Homalochilus is emended to be H. punctatostriatus. A neotype (Liogenys morio) and lectotypes (H. punctatostriatus, H. niger) are designated. Dichotomious key to Neotropical Diplotaxini genera and to species of Homalochilus and Careocallus are presented. Sequences and phylogenetic signals for Neotropical melolonthines are reported for the first time. [ABSTRACT FROM AUTHOR] more...
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- 2024
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4. A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species
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Basílio, Daniel Silva, primary, Vaz-de-Mello, Fernando Zagury, additional, Cherman, Mariana Alejandra, additional, and Almeida, Lúcia Massutti de, additional
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- 2023
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5. PERCEPÇÃO DOS IMPACTOS SOCIOAMBIENTAIS NO OLHAR DA COMUNIDADE LOCAL NA IMPLEMENTAÇÃO DO PARQUE EÓLICO NO MUNICÍPIO DE RIACHUELO-RN
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Basílio, Daniel, primary, Torres, Antônio, additional, and Dantas, Fabiola, additional
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- 2023
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6. New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data
- Author
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Cherman, Mariana A, primary, Basílio, Daniel S, additional, Clarkson, Bruno, additional, Agostinis, André O, additional, Smith, Andrew B T, additional, Vaz-De-Mello, Fernando Z, additional, and Almeida, Lúcia M, additional more...
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- 2023
- Full Text
- View/download PDF
7. Intra- and inter-rater reliability in ultrasonographic measurements of coracohumeral distance
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Guerra-Rodríguez, Diego, Guerrero-Henriquez, Juan, Basilio, Daniel, and Mendez-Rebolledo, Guillermo
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- 2024
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8. Crossing the Bridge: Foreign Language Students' Reciprocal Images in (Inter)Cultural Mediation between Portugal and Turkey
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Basílio, Daniel, Araújo e Sá, Maria Helena, and Simões, Ana Raquel
- Abstract
This study intends to highlight the role that Foreign Language Education (FLE), particularly in the Higher Education context, can play so as to contribute to the rapprochement of two distant and still mutually unknown countries such as Portugal and Turkey. In this sense, it ultimately aims at supporting the training of intercultural speakers, capable of promoting an effective Intercultural Dialogue between the two countries. A diagnosis is presented of the reciprocal images of Portuguese and Turkish students learning each other's language and culture. Conclusions are drawn on how their self- and hetero-images may pertain on the construction of their identities, on their awareness about and attitudes towards each other, on their motivation to learn each other's language, and on intercultural communication itself. Allying the study of Images of Languages and Cultures and the concept of Intercultural Competence in the study of students' representations within the FLE research tradition, a content analysis was carried out of the responses given to an inquiry by questionnaire. Conative implications of the students' images as revealed in the results were pointed out. Some important distinctions are highlighted in the reciprocal images of both groups. On this basis, recommendations for FLE are made focusing on the positive reconstruction of students' reciprocal images. more...
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- 2016
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9. A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species
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Basílio, Daniel Silva, Vaz-de-Mello, Fernando Zagury, Cherman, Mariana Alejandra, Almeida, Lúcia Massutti de, Basílio, Daniel Silva, Vaz-de-Mello, Fernando Zagury, Cherman, Mariana Alejandra, and Almeida, Lúcia Massutti de more...
- Abstract
Coilodes Westwood, 1846 is a Neotropical genus of Hybosorinae. Despite being a morphologically homogeneous genus, it presents a great deal of intraspecific variation, especially with reference to the colour of integuments. This makes identification of species even more difficult, since the majority of original descriptions present the colour as the main diagnostic character. To solve this problem, the first taxonomic revision of Coilodes is presented. The genus now comprises 13 species. Redescriptions of C. castaneus Westwood, 1846, C. fumipennis Arrow, 1909, C. humeralis (Mannerheim, 1829), C. niger (Mannerheim, 1829), C. ovalis Robinson, 1948, C. parvulus Westwood, 1846, and C. punctipennis Arrow, 1909 are presented. Coilodes niger (Mannerheim, 1829) has its status revalidated and three new synonyms are proposed: C. gibbus (Perty, 1830) and C. chilensis Westwood, 1846 with C. humeralis, and C. nigripennis Arrow, 1903 with C. castaneus. Biological and geographical distribution data are expanded. Lectotypes are designated for C. humeralis and C. niger. Six new species are described: C. bezerrai Basílio & Vaz-de-Mello sp. nov., C. edeiltae Basílio & Vaz-de-Mello sp. nov., C. lunae Basílio & Vaz-de-Mello sp. nov., C. mayae Basílio & Vaz-de-Mello sp. nov., C. ravii Basílio & Vaz-de-Mello sp. nov., and C. skelleyi Basílio & Vaz-de-Mello sp. nov. An identification key for the males of the species is presented. more...
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- 2023
10. Phylogenetic relationships in Hybosoridae (Coleoptera: Scarabaeoidea)
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Basílio, Daniel Silva, primary, Cherman, Mariana Alejandra, additional, Vaz-De-Mello, Fernando Zagury, additional, and Almeida, Lúcia Massutti, additional
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- 2023
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11. A morphological reappraisal of the immature stages and life history of Elachista synethes Meyrick (Lepidoptera, Elachistidae), an Australian leaf miner alien to Chile
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Vargas, Héctor Andrés, Brito, Rosângela, Basilio, Daniel Silva, and Moreira, Gilson Rudinei Pires
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- 2015
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12. Description and life history of a new cecidogenous species of Palaeomystella Fletcher (Lepidoptera, Momphidae) from Brazil
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Basilio, Daniel Silva, Casagrande, Mirna Martins, Bordignon, Sérgio Augusto Loreto, and Moreira, Gilson Rudinei Pires
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- 2015
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13. Education and Communication in Health as Strategies for Accomplishment of the Universal Access to Health in Uberaba
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Martins, Rosane A.S., Julião, Claudia H., Farinelli, Marta R., Bessa, Aline F., Maríngolo, Aline C.P., Nolasco, Brendda C.A., Ribeiro, Camila B., Basilio, Daniel S., Gomes, Nairana A.N., Severino, Renata R., and Costa, Tainara P. more...
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- 2015
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14. Energy landscape of the reactions governing the Na + deeply occluded state of the Na + /K + -ATPase in the giant axon of the Humboldt squid
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Castillo, Juan P., De Giorgis, Daniela, Basilio, Daniel, Gadsby, David C., Rosenthal, Joshua J. C., Latorre, Ramon, Holmgren, Miguel, and Bezanilla, Francisco
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- 2011
15. A new genus and species of Neotropical Hybosorinae Erichson, 1847 (Coleoptera: Hybosoridae)
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BASÍLIO, DANIEL S., primary, VAZ-DE-MELLO, FERNANDO Z., additional, and ALMEIDA, LÚCIA M., additional
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- 2022
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16. Liogenys schneiderae Cherman & Basílio & Mise & Frisch & Smith & Almeida 2021, new species
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys schneiderae ,Taxonomy - Abstract
Liogenys schneiderae Cherman, new species Figs. 8, 9. Type material. Holotype male, labeled: [green, handwritten] “4-dens Fbr / 4-dentata / kl. / Pg.[?] / Col. [Colombia]”, [white, typeset and handwritten] “MLU Halle / WB Zoologie / S. –Nr. 8/3/11”, [red typeset and handwritten] “ LIOGENYS / SCHNEIDERAE / HOLOTYPE / Cherman M. A.” (MLUH). Paratypes (81), all bearing the label [yellow, typeset and handwritten] “ LIOGENYS / SCHNEIDERAE / PARATYPE / Cherman M. A.”. One female with the labels: [white handwritten] “ Venezuela ”, [white typeset and handwritten] “ Liogenys quadri- / dens / F det. G. Frey, 1968” (ZMHB). One female with the label: [green typeset] “ Venezuela / F. Kummerow S.” (ZMHB). Two females and one male with the following label: [white handwritten] “ Guyana ” (ZMHB). Two females and two males with the label: [green, outlined, typeset] “Hist. Coll. (Coleoptera) / Nr. 24131 / Liogenys quadridens Burm. / Columb., Moritz. / Zool. Mus. Berlin” (ZMHB). One male with the labels: [white typeset] “24131”, [dark green handwritten] “ Amphicrania / 4 dentata Nr / Columb. Moritz”, [white handwritten] “quadridens / Burm / Melol. 4 dens Fab.” (ZMHB). Two males bearing the labels: [white handwritten] “ Colombia / Mus. Ber. / 4-dentata / Kl.”, [white typeset] “Mus. Western” (ZMUC). One male bearing: [white typeset] “ Coll. Kraatz ”, [white typeset] “ Moser det”, [white typeset] “Dtsch. Ent. Inst. / Eberswalde”, [white outlined green, handwritten] “quadridens. / Guyana /F.”, [white handwritten] “ Liogenys. /Guerin.”, [white handwritten] “ Liogenys / quadridens Fab ”, [white, typeset and handwritten] “ Liogenys quadri- / dens / F det. G. Frey, 1968”, [white handwritten] “ Liogenys /quadridens / F.”, [light green typeset] “DEI Münchenberg / Col – 03566” (SDEI). One male with the following data: [white typeset] “CARIACO / (Edo. SUCRE)”, [white typeset] “ Bordon leg.” / [handwritten] “10-IV-59” [white handwritten] “4”, [white, outlined, typeset] “H. & A. HOWDEN/COLLECTION / ex. A. Martinez coll.” (CMNC). One female with the data: [white typeset] “QUIAMARE[Libertad] / (BARCELONA)”, [white typeset] “5-4-58/BORDON LEG.”, [white, outlined, typeset] “H. & A. HOWDEN / COLLECTION / ex. A. Martinez coll.” (CMNC). Two females with the data: [white, typeset] “ Aragua / Venezuela / V.1978 ” (USNM). One male with the data: [white, typeset] “Chaguarramas / (Edo. Guarico)”, [white typeset] “13-3-60 / BORDON LEG.”, [white, handwritten and typeset] “ Liogenys quadridens ♂ / (Fab.) / A MARTINEZ-DET. 1974”, [white with black border, typeset] “H. & A. HOWDEN / COLLECTION / ex. A. Martinez coll.”, genitalia mounted (CMNC). Two males and four females with the data: [white, typeset] “West Indies: / Tobago: / Scarborough. / 16. vi. 1914. / W.E. Broadway. ” and “Brit. Mus. / 1920–487.” (1 male and 2 females at BMNH; 1 male and 2 females at CMNC). One male with the data: [white, typeset] “West Indies: / Tobago: / Scarborough. / 17. vi. 1914. / W.E. Broadway. ” and “Brit. Mus. / 1920–487.” (BMNH). One female with the data: [white, typeset] “West Indies: / Tobago: / Scarborough. / 10. vi. 1914. / W.E. Broadway. ” and “Brit. Mus. / 1920–487.” (BMNH). One female with the data: [white, typeset] “West Indies: / Tobago: / Scarborough. / 12. vi. 1914. / W.E. Broadway. ” and “Brit. Mus. / 1920–487.” (BMNH). Two females with the data: [white, typeset] “West Indies: / Tobago: / Scarborough. / 15. vi. 1914. / W.E. Broadway. ” and “Brit. Mus. / 1920–487.” (BMNH). Two females with the data: [white, typeset] “West Indies: / Tobago: / 9. vi. 1914. / W.E. Broadway. ” and “Brit. Mus. / 1920–487.” (BMNH). One male with the data: [white, handwritten] “quadridens / Fabr. / 4-dentata, / Klug. / Columbia”, [white, typeset] “So named / in Reiches / Collection. / C.W.”, [green, handwritten] “ ♂ ”, [white, handwritten] “nov. sp.”, [light blue, handwritten] “2258.”, [white, typeset] “67.45” (BMNH). Sixteen males and 15 females with the data: [white, typeset] “ VENEZUELA: Guarico, Hato / Masaguaral, 45kmS Calabozo / 8.57N, 67.58W, GalryForest / #28, 75m, 3-5 June 1988 / uv light, M. Epstein ” (eight males and eight females at USNM, four males and three females at CMNC, two males and two females at DZUP, two males and two females at CEMT). Paratypes of DZUP with the sequence of voucher numbers: [white typeset] “ DZUP / 402753 ” to “ DZUP / 402756 ”. One male and eight females with the data: [white, typeset] “ VENEZUELA: Guarico, Hato / Flores Morades, 45 km S / Calabozo, 8.57N, 67.58W / Galry Forest #2, 75 m / 24-26June 1989, uv light / M. Epstein & M. Rodiguez ” (USNM). One male and two females with the data: [white, typeset and handwritten] “ex? / intercepted / at Miami Fla / 1 May 1964 ” (USNM). One male and two females with the data: [white, typeset and handwritten] “ VENEZUELA: BOLIVAR / Guri, 16.VI.1996 / H.&A. Howden Lt / Wet forest ” (CMNC). One female with the data: [white, typeset and handwritten] “ VENEZUELA: GUARICO / 12km W Valle de la / Pascua, 21-22.VI.96 Lt / H.&A. Howden ” (CMNC). One female with the data: [white, typeset] “VEN: Bolivar, 20km / SW Ciudad Bolivar / 19. VI. 1987, S&JPeck / woodland on sand, UV” (CMNC). One female with the data: [white, typeset] “VENEZ: Anzoategui / 18 km E Pariaguan / 24JUN 1987, 240m / M.A. Ivie, beating” (CMNC). Holotype deposited at MLUH. Eleven paratypes at BMNH. Nineteen paratypes deposited at CMNC. Four paratypes at DZUP, four paratypes at CEMT. One paratype deposited at SDEI. Thirty paratypes deposited at USNM. Ten paratypes deposited at ZMHB. Two paratypes deposited at ZMUC. Additional material. One male (BMNH) missing the head and pronotum with the data: [white, typeset and handwritten] “Scarborough, / Tobago. / 15 May 1914. / W.E. Broadway. / 1914-403”. Diagnosis. Body brown, elongate, sides almost parallel in males (Fig. 8A), slightly wider on posterior third in females; elytra brown; pronotum reddish to purplish brown; clypeus quadridentate, clypeal emargination deep, rounded, and narrow; distance between lateral and anterior clypeal teeth slightly longer than basal width of one anterior tooth; right angle between lateral and anterior clypeal teeth; pronotal corners rounded (Fig. 8B), appearing obsolete especially in females; mesotibia quadrate in cross section (Fig. 8C); pygidium weakly convex; pygidial disc glabrous, with scattered bristles on apex (Fig. 8D); ventrites II, III, and sometimes IV undulate on midline, especially ventrite II (Fig. 8E); in males protarsomere II up to twice the length of protarsomere I in dorsal view; metatibia subbasally produced towards apex; basal region of parameres strongly narrowed, parameres more than five times the length of their apex; apex harpoon-shaped, lateral spine short, projected almost perpendicular to parameres (Fig. 8F). Holotype. Male. Length: 9.4 mm, width: 4.4 mm. Brown. Head: distance between eyes nearly twice the width of one eye; frons longer than clypeus; clypeus rugose, clypeal emargination deep, rounded, and narrow; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous tooth-like projection; distance between lateral and anterior teeth slightly longer than basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye longer than one eye length, angle between anterior and lateral teeth approximately 90º; canthus not exceeding the outer margin of the eye; distal maxillary palpomere maximum width slightly wider than at apex; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and as long as the funicle. Thorax: pronotal anterior margin slightly produced medially, undulate and flanged (Fig. 8B); maximum length of pronotum exceeding the length of tarsomeres I–III combined; disc glabrous, punctures very sparse and coarse, denser towards anterior margin; pronotal posterior corners rounded, appearing obsolete; hypomere with long bristles and thin scales; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the length of the metacoxa; scutellum triangular, few, scattered, fine punctures at the sides. Elytra: shiny, uniform light brown, lighter in color than the pronotum; elytra more than three times longer than the pronotum; elytral suture darker than the elytron and slightly elevated; all four elytral ridges weakly defined. Legs: procoxa scaly, thin scales on outer surface; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; protibial spur present; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia subquadrate in cross section, surface finely sculptured, mesotibia with two transverse carinae, the apical carina incomplete; metacoxa with bristles throughout, basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia carinate and abruptly subbasally produced towards apex, apical inner surface glabrous, metatibial surface coarsely sculptured on outer margin; metatibial transverse carina present posteriorly; protarsomere II very elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres equal in width to the mesotarsomeres, slightly wider than the metatarsomeres; metatarsomere I shorter than metatarsomere II; claw bifid, outer tooth of a claw longer and narrower than the inner tooth; distance between teeth shorter than the inner tooth. Abdomen: disc of ventrites with sparse bristles; ventrites II–III and sometimes IV with tubercles on disc (Fig. 8E); propygidium with bristles, pygidium slightly convex, subtrapezoidal, wide; pygidial width exceeding distance between spiracles of propygidium; pygidial disc glabrous, few bristles on apex, pygidial disc finely punctate; pygidial apex quadrate. Parameres: basal region dorsally narrowed, narrower than both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; parameres more than five times the length of their apex; inner margins opened, convergent; apex harpoon-shaped, lateral spine thin, short, projected almost perpendicular to parameres (approximately 80º angle). Parameres concave in lateral view (Fig. 8G). Variation. Female length: 9.2–10.2 mm, width 4.5–5.3 mm. females as the holotype except the clypeal anterior teeth wider; canthus exceeding the outer margin of the eye; pronotal corners more marked; metatibia coarsely sculptured, metatibial apical carina complete; ventrite tubercles more noticeable and pygidium wider, more trapezoidal. Male length: 9.0– 9.5 mm, width 4.4–4.8 mm. Males as the holotype except the head and pronotal punctures coarser; pronotal corners more marked, and pygidium flatter and bent downwards. Etymology. The species is dedicated to Dr. Karla Schneider, biologist and curator of the Zoological Collection of the Martin-Luther-Universität, Halle-Wittenberg, Halle, Germany. Karla made the discovery of this new species possible by loaning the holotype from the MLUH to the ZHMB. This allowed its thorough examination during the visit of M.A.C. to these collections. This name should be treated as a noun in the genitive case. Type locality: Colombia. Geographical distribution. TRINIDAD AND TOBAGO (Scarborough), COLOMBIA, GUYANA, VENEZUELA (Anzoátegui, Bolívar, Guaricó, Sucre). Remarks. Liogenys schneiderae Cherman, new species resembles L. quadridentata mainly in the body shape and size, color of the body and elytra, and in the quadridentate clypeus. However, L. schneiderae differs (L. quadridentata in parenthesis) in the clypeus being more rugose, with the impressions next to the frons more marked (not rugose); antenna with club shorter; anterior margin of pronotum slightly depressed throughout (not as so); elytral suture medially darker than the elytron (medially unicolored with elytron); metatibia flatter with inner margin abruptly subbasally produced towards apex (produced straight towards apex); ventrites I–IV with tubercles (without tubercles); and pygidium never inflated, with apex quadrate (pygidium convex in both sexes, inflated in females, pygidial apex rounded)., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 221-224, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886 more...
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17. Liogenys genieri Smith & Cherman 2021, new species
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
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Coleoptera ,Liogenys genieri ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys genieri Smith & Cherman, new species Figs. 3, 9. Type material. Holotype male, labeled: [white, typeset] “ VENEZUELA: Guarico, Hato / Flores Morades, 45 km S / Calabozo, 8.57N, 67.58W / Galry Forest #2, 75 m / 24–26June 1989, uv light / M. Epstein & M. Rodiguez ”, [red, typeset and handwritten] “ LIOGENYS / GENIERI / SMITH & CHERMAN / HOLOTYPE ”, genitalia mounted (USNM). Paratypes (8), all bearing the label [yellow, typeset and handwritten] “ LIOGENYS / GENIERI / SMITH & CHERMAN / PARATYPE ”: One male and six females with the same data as the holotype (one male and one female at CMNC, two females at DZUP, three females at USNM). One male with the data: [white, typeset] “ VENEZUELA: Guarico / Hato Masaguaral / (44km S Calabozo) / May 3–10 1985 / Menke & Carpenter” (USNM). Each female paratype of DZUP with the voucher: [white typeset] “ DZUP / 402748 ” and “ DZUP / 402749 ”. Holotype and four paratypes deposited at USNM. Two paratypes at CMNC. Two paratypes at DZUP. Diagnosis. Body light brown, elongate, sides almost parallel; elytra light brown; pronotum reddish brown (Fig. 3A); clypeus quadridentate; clypeal emargination deep, angulate, and narrow; distance between lateral and anterior clypeal teeth approximately equal to basal width of one anterior tooth; angle between lateral and anterior clypeal teeth approximately 90º; pronotal corners rounded (Fig. 3B); mesotibia subquadrate in cross section; pygidium slightly convex; pygidial disc glabrous, with scattered bristles on apex (Fig. 3D); in males protarsomere II more than twice the length of protarsomere I in dorsal view (Fig. 3C); parameres more than six times the length of their apex; lateral expansion along the parameres only seen subapically; inner margins convergent; apex expanded with backward-facing spine weakly projected, spine length less than one-third length of apex (Fig. 3E). Holotype. Male. Length: 9.6 mm, width: 4.6 mm. Light brown. Head: distance between eyes nearly twice the width of one eye; frons equal in length to clypeus; clypeal emargination deep, angulate, and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous tooth-like projection; distance between lateral and anterior teeth approximately equal to basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye approximately equal to one eye length, angle between anterior and lateral teeth approximately 90º; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width twice the width at apex; antenna with 10 antennomeres, club lighter in color and longer than funicle. Thorax: pronotal anterior margin slightly produced medially, undulate, and flanged; disc glabrous, punctures moderately dense and coarse; anterolateral margin reflexed; pronotal posterior corners rounded; hypomere with long bristles; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the length of the metacoxa; scutellum triangular with sparse, coarse punctures. Elytra: shiny, uniform light brown, lighter in color than the pronotum; elytra near three times the length of pronotum; elytral suture darker than the elytron and weakly elevated; all four elytral ridges weakly defined. Legs: procoxa with abundant, thin and long scales, short scales on outer surface; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; protibial spur present; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia subquadrate in cross section, surface coarsely sculptured on outer margin, mesotibia with two transverse carinae, the apical carina incomplete; metacoxa with bristles throughout, basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia straight, carinate, and produced on apex, apical inner surface with bristles, metatibial surface coarsely sculptured; metatibial transverse carina weakly defined, only present posteriorly; protarsomere II elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres slightly wider than the mesotarsomeres and two times wider than the metatarsomeres; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, outer tooth of a claw longer and as wide as the inner tooth; distance between teeth shorter than the inner tooth. Abdomen: disc of ventrites with bristles; propygidium with bristles, pygidium weakly convex, subquadrate, wide; pygidial width approximately equal to distance between spiracles of propygidium; pygidial disc glabrous except for a few bristles on apex, pygidial disc coarsely punctate; pygidial apex oval. Parameres: basal region dorsally narrowed, equal to both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; inner margins slightly convergent; parameres more than six times the length of their apex; apex harpoon shaped; laterally expanded with backward-facing spine weakly projected straight downward, spine length less than one-third length of apex. Parameres in lateral view straight, not coplanar (Fig. 3F). Variation. Female length: 10.2–11.2 mm, width 5.0– 5.6 mm. Females as the holotype except in the size; pronotum wider, with anterolateral margin distinctly reflexed; maximum width of distal maxillary palpomere slightly wider than at apex. Male length: 9.4–9.6 mm, width 4.0– 4.8 mm. Males as the holotype except in the punctures of pronotum slightly less dense; protibial teeth sharper and longer (worn in holotype). Etymology. This species is named after François Génier, scarab taxonomist from the CMNC, in appreciation for his help in photographing specimens for this paper. Type locality. Hato Masaguaral, Guárico, Venezuela (8.57ºN, 67.58ºW). Geographical distribution. VENEZUELA (Guárico). Remarks. Liogenys genieri Smith & Cherman, new species resembles L. clipeosetosa in the body color and size, shape of the quadridentate clypeus, head and pronotum coarsely punctate, anterior teeth of clypeus long, anterior and lateral teeth forming approximately 90º angle, and posterior corners of pronotum rounded. Males in both species have parameres very long, more than five times the apical length, and apex laterally weakly projected. Liogenys genieri differs (L. clipeosetosa in parenthesis) in the frontoclypeal surface without bristles (with bristles); clypeal emargination angulate, narrower (rounded, wider); pronotum with anterolateral margin reflexed (not as so); apex of pygidium oval (quadrate); males with parameres slightly shorter, lateral expansion along the parameres not visible dorsally and straight in lateral view (lateral expansion entirely visible dorsally)., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 207-209, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886 more...
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18. Liogenys quadridens
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
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Coleoptera ,Liogenys quadridens ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys quadridens (Fabricius, 1798) Figs. 6, 9. Melolontha quadridens Fabricius, 1798: 131 (original description). Liogenys quadridens: Burmeister 1855: 14 (redescription); Harold 1869: 1140 (checklist); Bates 1887:155 (checklist); Dalla Torre 1913: 318 (checklist); Blackwelder 1944: 228 (checklist); Frey 1969: 50 (key); Evans 2003: 213 (checklist); Evans & Smith 2009: 181–182 (checklist); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 11, 16; 2019: 62 (taxonomy). Type material examined. Melolontha quadridens female lectotype, present designation (ZMUC): [white handwritten] “4 Dens.”, [white typeset] “zmuc / 00022369”, [white, outlined in red, typeset and handwritten] “ LIOGENYS / QUADRIDENS / (Fabricius, 1798) / LECTOTYPE / Des. Cherman M. A.”. Additional material examined. PANAMA. Coclé: Río Hato, 22.V.1960, light trap, V.J. Tipton, 1 male, 1 female (CNCI). Panamá: Cerro Campana, 820 m, 8º40`N, 79º56`W, 22.V.1976, at blacklight, B.C. Ratcliffe, 2 males, 4 females (UNSM); Cerro Campana, 2600 feet, 10–20.V.1981, J.E. Wappes, 1 male, 1 female (UNSM). Panamá Oeste: Nueva Gorgona, 25.V.1962, R.&F. Zweifel, 1 male, 1 female (AMNH); Sajalices, 17.V.1982, E. Giesbert, 1 male, 3 females (UNSM). Veraguas: Santiago, 18.V.1977, H. Howden, 1 male (CMNC). VENEZUELA. Guárico: San Juan de Los Morros, 28.IV.1938, C.H. Ballou, 1 female (USNM); Monagas: El Tejero, 200 m, 2.V.1964, C. Bordon, 1 male (CMNC). Distrito Capital: Caracas, 1938, G. Vivas-R., 2 males (USNM). Without locality and date, E. Kummerow S., 1 female (ZMHB). Diagnosis. Body light brown, elongate, males with sides parallel (Fig. 6A), females with sides slightly wider on posterior third (Fig. 6B); elytra light brown to yellowish brown, pronotum reddish to purplish brown, head even darker; clypeus quadridentate; clypeal emargination deep, rounded, and wide (Fig. 1A), slightly narrower in females than in males (Fig. 1B); distance between lateral and anterior clypeal teeth longer than basal width of one anterior tooth; angle between lateral and anterior clypeal teeth approximately 90º; mesotibia subquadrate in cross section; pygidium flat or slightly convex, pygidial disc glabrous, with scattered bristles on apex (Fig. 6D); males with protarsomere II less than twice the length of protarsomere I in dorsal view; parameres elongate, up to three times the length of their apex (Fig. 6H), inner margins straight, lateral expansion on outer margins along the parameres partially hidden in dorsal view (Fig. 1E), apex harpoon-shaped, lateral spine projected backwards, slightly convergent to the outer margin of the parameres; spine length slightly more than twice the length of the apex, subapical and proximal part of the apex elevated medially (Fig. 6), abruptly flattened distally at apex and spine (Fig. 6G). Redescription. Length: 9.2–10.8 mm, width: 4.6–5.0 mm. Light brown to brown. Head: dark, distance between eyes nearly twice the width of one eye; frons equal in length to clypeus; clypeal emargination deep, rounded, and narrow; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous, tooth-like projection; distance between lateral and anterior teeth longer than basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye longer than one eye length, angle between anterior and lateral teeth approximately 90º; canthus exceeding or not the outer margin of the eye; distal maxillary palpomere maximum width twice the width at apex; fovea deep, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color, as long as the funicle in females, longer than the funicle in males. Thorax: pronotal anterior margin slightly produced medially, undulate; disc glabrous, punctures sparse and fine in females, coarser in males; pronotal posterior corners rounded in males, apparently obsolete in females; hypomere with long bristles; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the length of the metacoxa; scutellum triangular, punctures mainly at the sides. Elytra: shiny, uniform yellowish brown to light brown, lighter in color than pronotum; elytra near three times longer than the pronotum; elytral suture slightly darker than elytron and not elevated; all four elytral ridges weakly defined. Legs: procoxa with long bristles on infra-carinal surface and fine scales and short bristles on outer surface; three protibial teeth, middle and apical equal in size, distance between basal and middle teeth longer than between middle and apical; protibial spur present; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia quadrate in cross section, surface coarsely sculptured in females, finely sculptured in males, mesotibia with two transverse carinae, the apical carina complete in females, incomplete in males; metacoxa with bristles throughout and forming a row of scales parallel to the anterior margin (Fig. 6F); basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of male metatibia carinate and produced on apex, apical inner surface with scattered bristles, metatibial disc coarsely sculptured; metatibial transverse carina and discontinuous longitudinal carina present posteriorly; metatarsomere I and metatarsomere II equal in length; males with protarsomere II as wide as it is long; protarsomeres and mesotarsomeres I–IV equally enlarged, wider than metatarsi; claw bifid, symmetrical, outer tooth of a claw longer and narrower than inner tooth; distance between teeth shorter than the inner tooth. Abdomen: disc of ventrites with sparse bristles; propygidium with bristles, pygidium flat, subtrapezoidal, wide, pygidial width exceeding distance between spiracles of propygidium; pygidial disc glabrous, few bristles on apex, coarsely punctate, roughly in males; slightly depressed medially in males; pygidial apex quadrate. Parameres: basal region slightly wider than both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; parameres almost three times the length of their apex; inner margins straight; lateral expansion of the outer margins partially hidden, visible from the midline up to the subapical portion; apex harpoon-shaped, lateral spine straight backwards, slightly bent, spine length more than twice the length of the apex, subapical and proximal part of the apex elevated medially, flattened distally at apex and spine (Figs. 1E, 6G). Parameres concave in lateral view (Fig. 6E). Type locality. “ India Orientali” [Burmeister 1855: 14, misinterpreted locality of West Indies]. Species from Panama and Venezuela. Geographic distribution. PANAMA (Coclé, Panamá, Panamá Oeste, Veraguas), VENEZUELA (Distrito Capital, Monagas). Remarks. Liogenys quadridens was redescribed for the first time by Burmeister (1855), who transferred it from Melolontha Fabricius, 1775 to Liogenys. Fabricius (1798) mentioned in the original description that the species is from “ India Orientali” [East Indies], we interpreted that he meant West Indies instead. According to Burmeister (1855) and Harold (1869), L. quadridens occurs in Colombia and Guyana. Bates (1887) stated that this species occurs in “Amazons” but this record should be disregarded as it was surely based on the misidentification of other species of Liogenys. Bates (1887) also cited Panama, “Volcán de Chiriqui 2500 to 4000 feet (Champion)” but these specimens are here identified as L. quadridentata (based on our examination of these specimens in the BMNH). Frey (1969) cited the distributional limits of this species, from Florida (United States of America) (presumably based on port interception specimens) to Pernambuco (Brazil). In his key, Frey (1969) mentioned an intraspecific variation in the pronotal punctures and in the shape of the parameres, which he drew. At NHMB, we found the specimen from Pernambuco and it is here identified as L. piauiensis Cherman, 2017. Therefore, we refute the occurrence of L. quadridens in Brazil. Among the 41 specimens studied from seven collections bearing L. quadridens identification, we found one male from Nueva Gorgona (Panama) matching with the parameres drawn by Frey (1969), and a female of the same collection event which matches with L. quadridens lectotype. Liogenys quadridens has been cited by the United States Department of Agriculture (1969, 1971, 1974) in its list of intercepted plant pests, carried by aircraft, in unknown hosts. The first time cited (1967–1968) it was being carried from Panama, and the following (1971–1972) probably from Venezuela (United States Department of Agriculture 1969, 1974). This species was also cited as a tomato plant host from Maracaibo, Venezuela (Morales- Valles et al. 2003), and there exists reports of this species in imported ripe tomato fruit throughout the European and Mediterranean Plant Protection Organization (EPPO) region. Nonetheless, as it is not likely to be transported in tomatoes it is classified non dangerous (Grousset et al. 2015). In a study on scarab beetles abundance of a tropical dry forest in Colombia, García-Atencia et al. (2015) found that L. quadridens was the most abundant species (1,667 individuals), characterized as an important pest of semestral crops in the Colombian Caribbean. After studying several specimens of the Caribbean region we cannot confirm that all those specimens mentioned on the previous works are L. quadridens. Although we do believe that L. quadridens occurs in Colombia (based on the known distribution in Panama and Venezuela), we did not examine any specimens to confirm this country record., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 214-215, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886, {"references":["Fabricius, J. C. (1798) Supplementum entomologiae systematicae. Proft et Storch, Copenhagen, 572 pp.","Burmeister, H. (1855) Handbuch der Entomologie (Coleoptera Lamellicornia Phyllophaga chaenochela). Vierter Band. Zweite Abtheilung. Theod. Chr. Fr. Enslin, Berlin, 570 pp.","Harold, E. F. von (1869) Scarabaeidae. In: Gemminger, M. & von Harold, E. F. (Eds.), Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Vol. 4. E. H. Gummi, Munich, pp. 976 - 1346.","Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. & Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160.","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, S. (Ed.), Coleopterorum catalogus. Vol. XX. Junk, W., Berlin, pp. 291 - 450.","Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies and South America. Part 2. Smithsonian Institution, Washington, D. C., pp. 189 - 265. https: // doi. org / 10.5479 / si. 03629236.185.2","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64.","Evans, A. V. (2003) A checklist of the NewWorld chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211, 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) version 3 2009. Papers in Entomology, 1, 1 - 344.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","United States Department of Agriculture [USDA] (1969) List of intercepted plant pests, 1968 (Pest recorded from July 1, 1967, through June 30, 1968). Hayattsville, Maryland. U. S. G. P. O., Washington, D. C., pp. 46 - 58.","United States Department of Agriculture [USDA] (1971) Insects not known to occur in the United States: A Scarab (Liogenys macropelma Bates). Cooperative Economic Insect Report, 21, 335 - 336.","United States Department of Agriculture [USDA] (1974) List of intercepted plant pests, 1972 (Pest recorded from July 1, 1971, through June 30, 1972). Hayattsville, Maryland. U. S. G. P. O., Washington, D. C., pp. 55 - 68.","Grousset, F., Suffert, M. & Petter, F. (2015) EPPO Study on pest risks associated with the import of tomato fruit. EPPO Bulletin, 45, 153 - 156. https: // doi. org / 10.1111 / epp. 12180","Garcia-Atencia, S. P., Martinez-Hernandez, N. & Pardo-Locarno, L. C. (2015) Escarabajos fitofagos (Coleoptera: Scarabaeidae) en un fragmento de bosque seco tropical del departamento del Atlantico, Colombia. Revista Mexicana de Biodiversidad, 86, 754 - 763. https: // doi. org / 10.1016 / j. rmb. 2015.07.009"]} more...
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19. Liogenys macropelma Bates. The 1887
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys macropelma ,Taxonomy - Abstract
Liogenys macropelma Bates, 1887 Figs. 5, 9. Liogenys macropelma Bates, 1887: 155 (original description); Nonfried 1892: 268 (checklist); Dalla Torre 1913: 318 (checklist); Blackwelder 1944: 227 (checklist); Frey 1969: 42 (key); United States Department of Agriculture 1971: 335; 1974: 28 (natural history); Evans 2003: 210 (checklist); Evans & Smith 2009: 179 (checklist); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 14, 16, 69, 86 (taxonomy); 2019: 18, 64 (taxonomy), 53–55 (redescription). Liogenys gebieni Moser, 1921: 140 (original description); Blackwelder 1944: 227 (checklist); Frey 1969: 41 (key); Evans 2003: 209 (checklist); Evans & Smith 2009: 178 (checklist); Krajčík 2012: 145 (checklist). New junior subjective synonym. Type material examined. Liogenys macropelma male lectotype (MNHN): [white typeset] “V. de Chiriqui, / 25–4000 ft. / Champion.”, [white handwritten] “ Liogenys / macropelma / Bates ♂ ”, [red typeset] “ SYNTYPE ”, [white handwritten] “micropygus”, [white, outlined in red, typeset] “ LECTOTYPE / Liogenys macropelma / Bates, 1887 / Cherman M. A. 2015”. Genitalia mounted. Six males with the same labels as the lectotype, plus the label: [white, outlined in red, typeset] “ PARALECTOTYPE / Liogenys macropelma / Bates, 1887 / Cherman M. A. 2015”. One female with one different label: [white handwritten] “ Liogenys / macropelma / Bates ♀ ”. Six male and one female paralectotypes (BMNH): [white, typeset] “V. de Chiriqui, / 25–4000 ft. / Champion.”, [white, typeset and handwritten] “B.C.A.Col.II. (2) / Liogenys / macropelma”. One male paralectotype also has the labels: [round, white with red border] “Type”, [white, typeset] “Sp. figured.”. Liogenys gebieni male lectotype, present designation (ZMHB): [white typeset] “Columbien / LaGarita [Los Patios, Norte de Santander] (Gebirge [mountains]) / W. Fritsche / leg. 1912” [genitalia mounted on the label], [white handwritten] “Isonychus [misspelled Liogenys] / gebieni / Typen Mos.”, [red typeset] “TYPUS”, [white typeset] “ Liogenys / gebieni / Mos. ”,. [white, outlined in red, typeset and handwritten] “ LIOGENYS / GEBIENI / Moser, 1921 / LECTOTYPE / Des. Cherman M. A.”. One female paralectotype with the same labels as the lectotype, plus the label: [white, outlined in red, typeset] “ PARALECTOTYPE / Des. Cherman M. A. [handwritten] LIOGENYS / GEBIENI Mos.”. Additional material examined. PANAMA. Colón: Coco Solo, 21.V.1976, B.C. Ratcliffe, 10 males, 20 females (UNSM); Coco Solo Hospital, V.1981, 2 males (UNSM); Fort Gulich, 17.V.1976, at lights, B.C. Ratcliffe, 1 male, 2 females; IV.1981, at lights, 5 males (UNSM); Fort Sherman, 26.IV.1980, D. Engleman, 1 female (UNSM). Panamá: Balboa, 16–17.V.1977, light, H.&A. Howden, 3 males, 1 female (CMNC); 19.VI.1977, H.&A. Howden, 1 male (CMNC); Corazal, 27.IV.1911, A. Busck, 2 males (USNM); Curundú, 8.V.1972, R.&E. Froeschner, 1 male (USNM); Diablo Heights, 3.V.1971, W.E. Bivin, 1 male, 1 female (USNM); Fort Clayton, 15.V.1961, C.E. Yunker, 8 males, 2 females (CNCI); Fort Clayton, V.1961, C.E. Yunker, 1 male (CMNC); La Sabanas, Panama City, IV.1927, at light, J. Zetek, 7 males, 19 females (USNM); Paraiso, 24.IV.1911, A. Busck, 2 males, 1 female (USNM); Punta Paitilla, Panama City, 8º59’N, 79º31`W, 17.V.1970, H. Hespenheide, 1 male (CMNC); Trivoli Hotel, Panama City, 15.IV.1967, H. Hespenheide, 1 male (CMNC). Geographical distribution. PANAMA (Chiriquí, Colón, Panamá), COLOMBIA (Norte de Santander). Remarks. After studying the type series of L. gebieni (ZMHB) (Fig. 5A–C), and comparing its external features and male genitalia (Fig. 5A–F) with that of L. macropelma type series and non-type material (Fig. 5G), we concluded that these specimens belong to the same species. The only difference is the color, which is darker in the L. gebieni types. Nonetheless, the difference in body color is a common intraspecific variation within Liogenys. Herein, we propose L. gebieni Moser as a junior subjective synonym of L. macropelma Bates. The parameres of each L. macropelma and L. gebieni drawn by Frey (1969) seem to be different. However, there were no specimens labeled as L. gebieni among Frey’s material housed at NHMB or USNM, thus, the material used by Frey (1969) to illustrate L. gebieni parameres remains unknown. According to the United States Department of Agriculture (1971, 1974), L. macropelma has been intercepted in Florida, Texas, and Puerto Rico (United States of America) coming by aircraft from El Salvador, Panama, and South America (probably Brazil) several times in a period from 1948–1971, always between April and May (identification verified based on the illustration in the report). Also, it was recorded being a host of okra plant Hibiscus esculentus L. (Malvaceae) coming from Guatemala, Panama, and Venezuela, which were intercepted in Florida, California, and Puerto Rico (United States of America). Several specimens in the USNM collection had been caught in port interceptions from Houston, Texas, originated from Panama, 1966 with no other explanation.Another specimen in the USNM bears the following data: 27 April 1962 port interception from Miami, Florida originating from “South America”. Based on the poor state of taxonomy of the genus Liogenys when these port interceptions took place, all identifications should be considered questionable unless they can be verified using voucher specimens., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 212-214, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886, {"references":["Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. & Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160.","Nonfried, A. F. (1892) Verzeichniss der seit 1871 neu beschriebenen Glaphyriden, Melolonthiden und Euchiriden. Berliner Entomologische Zeitschrift, 37, 249 - 290. https: // doi. org / 10.1002 / mmnd. 18920370302","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, S. (Ed.), Coleopterorum catalogus. Vol. XX. Junk, W., Berlin, pp. 291 - 450.","Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies and South America. Part 2. Smithsonian Institution, Washington, D. C., pp. 189 - 265. https: // doi. org / 10.5479 / si. 03629236.185.2","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64.","United States Department of Agriculture [USDA] (1971) Insects not known to occur in the United States: A Scarab (Liogenys macropelma Bates). Cooperative Economic Insect Report, 21, 335 - 336.","United States Department of Agriculture [USDA] (1974) List of intercepted plant pests, 1972 (Pest recorded from July 1, 1971, through June 30, 1972). Hayattsville, Maryland. U. S. G. P. O., Washington, D. C., pp. 55 - 68.","Evans, A. V. (2003) A checklist of the NewWorld chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211, 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) version 3 2009. Papers in Entomology, 1, 1 - 344.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312"]} more...
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20. Liogenys quadridentata Blanchard 1851
- Author
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys quadridentata ,Taxonomy - Abstract
Liogenys quadridentata Blanchard, 1851 Figs. 7, 9. Liogenys quadridentatus Blanchard, 1851: 168 (original description); Lacordaire 1855: 269 (checklist); Harold 1869:1140 (checklist); Dalla Torre 1913: 318 (checklist); Krajčík 2012: 145 (checklist). Liogenys quadridentata: Blackwelder 1944: 228 (checklist); Evans 2003: 213 (checklist); Evans & Smith 2009: 182 (checklist). Amphicrania 4-dentata [quadridentata] Klug in litt. Burmeister 1855: 14 (synonym of Liogenys quadridens, nomen nudum). Amphicrania quadridentatus Klug in litt.: Harold 1869: 1140 (synonym of L. quadridens). Type material examined. Liogenys quadridentatus male lectotype, present designation (MNHN): [white handwritten] “292 / 39”, [green handwritten] “ Amphicrania / 4 Dentata.”, [light green typeset] “MUSEUM PARIS”, [green handwritten] “ L. quadridentatus / Cat Mus”, [red typeset] “ HOLOTYPE ”, [white, outlined in red, typeset and handwritten] “ LIOGENYS / QUADRIDENTATA / Blanchard, 1851 / LECTOTYPE / Des. Cherman M. A.”, genitalia mounted. Additional material examined. PANAMA. Chiriquí: Volcán de Chiriquí, 2500–4000 feet, G.C. Champion, 4 males (BMNH). Panamá: La Chorrera, 2 females (BMNH). COLOMBIA. Magdalena: 12 miles W Santa Marta, 28.IV.1973, J.M. Campbell & H.F. Howden, 1 female (CMNC); 12 miles W Santa Marta, 29.IV.1973, J.M. Campbell & H.F. Howden, 1 male, 3 females (CMNC); 12 miles W Santa Marta, 12.V.1973, J.M. Campbell & H.F. Howden, 2 males (CMNC); 18.V.1973, 3 males, 5 females (CMNC); Parque Tayrona, 21 miles E Santa Marta, 15.V.1973, H.F. Howden & J.M. Campbell, 1 female (CMNC); Aracataca, 25.III.28, Darlington, 1 female (AMNH); Santa Marta, 3.V.1928, Darlington, 1 male (AMNH); Bonda, VI, 12 females (CMNH). VENEZUELA. Delta Amacuro: 15 km E Los Casillos, 45 km NE Ciudad Guyana, 26.VI.1985, ultraviolet light, M.A. Ivie, 3 females (CMNC). Guárico: 12 km W Valle de la Pascua, 21–22.VI.1996, light, H.&A. Howden, 1 female (CMNC); Estación Biológica Los Llanos, 12 km S Calabozo, 6–12.II.1969, ultraviolet light, P.&P. Spangler, 16 males, 9 females (USNM). Sucre: Cariaco, 20.V.1959, C. Bordon, 1 female (CMNC). Zulia: Maracaibo, without date, E. v. Jess., 1 male (ZMHB). Without locality and date, Moequerys, 2 females (SDEI); with erroneous locality (Brazil), without date, Mniszech, 1 female (MNHN). Diagnosis. Body light brown, elongate, sides parallel in males (Fig. 7A), slightly wider on posterior third in females; elytra light brown to yellowish brown, pronotum reddish to purplish brown; clypeus quadridentate; clypeal emargination deep, rounded, and narrow (Fig. 1C); anterior margin noticeable bent upwards, especially in females (Fig. 1D); distance between lateral and anterior clypeal teeth as long as basal width of one anterior tooth; acute angle between lateral and anterior clypeal teeth; mesotibia cylindrical in cross section (Fig. 7D); pygidium convex in both sexes, females with pygidium strongly convex medially (Fig. 7E); pygidial disc glabrous, with scattered bristles on apex; males with protarsomere II more than twice the length of protarsomere I in dorsal view; parameres up to three times the length of their apex (Fig. 7F); inner margins opened, slightly convergent; apex harpoon-shaped, lateral spine projected backwards, almost parallel to the outer margin of the parameres; spine length slightly more than twice the length of apex, subapical and part of the apex elevated medially, abruptly flattened at apex and spine. Redescription. Length: 9.4–11.2 mm, width: 4.7–5.2 mm. Brown. Head: distance between eyes nearly twice the width of one eye; frons somewhat swollen; clypeal emargination deep, rounded, and narrow; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous tooth-like projection; in females anterior margin noticeable bent upwards; distance between lateral and anterior teeth equal to basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye longer than one eye length, acute angle between anterior and lateral teeth; distal maxillary palpomere maximum width wider than at apex; fovea deep, extending up to the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than funicle. Thorax: pronotal anterior margin slightly produced medially; punctures very sparse, denser and coarser than the disc towards anterior margin; pronotal posterior corners rounded; hypomere with long bristles; mesepisternum with sparse, thin scales; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice as long as metacoxal width; scutellum triangular, sparse punctures at the base and sides. Elytra: shiny, uniform light brown or yellowish brown, lighter in color than the pronotum; elytral suture unicolored with elytron and slightly elevated or flat; all four elytral ridges weakly defined. Legs: procoxa with bristles, fine, thin scales on outer surface; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; protibial spur present; mesotibia cylindrical in cross section, surface finely sculptured, mesotibia with two transverse carinae, the apical carina incomplete in males, complete in females; metacoxa with bristles throughout; inner margin of male metatibia carinate and produced straight towards apex, apical inner surface glabrous, metatibial disc coarsely sculptured along the outer margin; metatibial transverse carina present posteriorly; protarsomere II elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres slightly wider than the mesotarsomeres, more than twice the width of metatarsomeres; metatarsomere I slightly shorter and wider than metatarsomere II; claw bifid, outer tooth of a claw equal in length and narrower than the inner tooth; distance between teeth shorter than the inner tooth. Abdomen: disc of ventrites with sparse bristles; propygidium with bristles, pygidium convex, convexity with intraspecific variation, females with pygidium inflated medially; pygidium subtrapezoidal, apex rounded; pygidial maximum width exceeding the distance between spiracles of propygidium; pygidial disc glabrous, few bristles on apex. Parameres: basal region narrowed at the midline; parameral split at 2/3 the length of the basal region; length of parameres nearly three times the length of their apex; inner margins convergent; apex harpoon-shaped, lateral spine projected backwards, slightly divergent from the outer margin of the parameres, spine length slightly more than twice as long as apex, subapical and proximal part of the apex elevated medially, abruptly flattened distally at apex towards the spine. Parameres concave, not coplanar in lateral view (Fig. 7G). Type locality. Not established. Geographic distribution. COLOMBIA (Magdalena), PANAMA (Chiriquí), VENEZUELA (Delta Amacuro, Guárico, Sucre, Zulia). Remarks. The lectotype of Liogenys quadridentata bears a green label “ Amphicrania 4-dentata ”, a name credited to “Klug MSC” by Burmeister (1855) but not formally validated until Blanchard (1851) described the species. Burmeister (1855) listed the name as a junior synonym of L. quadridens. Soon after, Harold (1869) mentioned “ Amphicrania 4-dentata ” as a synonym of L. quadridens, followed by “in litt.”, which means: “new species mentioned in litteris (in correspondence), that will be described later in a separate paper”. Harold (1869) probably did not see the L. quadridentata type series, so he did not associate the name “ Amphicrania 4-dentata ” to the latter. After the mentioned synonymy of “ Amphicrania 4-dentata ” (Burmeister 1855; Harold 1869), L. quadridentata was suggested to be a junior synonym of L. quadridens by later authors (Bates 1887; Dalla Torre 1913). We here confirm that L. quadridentata is a different species from L. quadridens. Liogenys quadridentata differs from L. quadridens (in parenthesis) mainly in the anterior margin of clypeus strongly bent upwards (not strongly bent); anterior and lateral tooth closer and acute angle between them (anterior and lateral tooth less close and right angle between them); mesotibia cylindrical in cross section (mesotibia subquadrate); pygidium noticeably convex, inflated medially in females (pygidium flat in both sexes); males with the protarsomere II longer (as wide as it is long), and the parameres are different in shape. The lectotype of L. quadridentata does not bear any locality data, and Blanchard (1851) did not mention the type locality in the original description. This fact agrees with Lacordaire (1855), who described the locality of L. quadridentata as “Patrie inconnue”. However, Burmeister (1855) and then Harold (1869) mentioned “Columbia” [current Colombia and Panama] as a locality record of “ Amphicrania 4-dentata ”, which is likely the actual distribution of this species. Dalla Torre (1913) cited L. quadridentata from Montevideo (Uruguay). This is here interpreted as an error, which was reproduced by Blackwelder (1944) and subsequent authors. Due to the layout of Blanchard (1851) descriptions when they are contiguous, it is easy to attribute the type locality of one species to the next one. It is most likely that Dalla Torre (1913) attributed the type locality of Liogenys parva Blanchard, 1851 to L. quadridentata., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 217-218, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886, {"references":["Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp.","Lacordaire, M. T. (1855) Genera des Coleopteres. Famille XXX. Pectinicornes. In: Histoire Naturelle des Insectes. Librairie Encyclopedique de Roret, Paris, pp. 1 - 594.","Harold, E. F. von (1869) Scarabaeidae. In: Gemminger, M. & von Harold, E. F. (Eds.), Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Vol. 4. E. H. Gummi, Munich, pp. 976 - 1346.","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, S. (Ed.), Coleopterorum catalogus. Vol. XX. Junk, W., Berlin, pp. 291 - 450.","Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies and South America. Part 2. Smithsonian Institution, Washington, D. C., pp. 189 - 265. https: // doi. org / 10.5479 / si. 03629236.185.2","Evans, A. V. (2003) A checklist of the NewWorld chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211, 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) version 3 2009. Papers in Entomology, 1, 1 - 344.","Burmeister, H. (1855) Handbuch der Entomologie (Coleoptera Lamellicornia Phyllophaga chaenochela). Vierter Band. Zweite Abtheilung. Theod. Chr. Fr. Enslin, Berlin, 570 pp.","Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. & Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160."]} more...
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21. Liogenys undetermined
- Author
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
- Subjects
Coleoptera ,Liogenys undetermined ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Key to Liogenys species from northern South America and Central America 1. Clypeus bidentate (Fig. 5A–B), only anterior margin of clypeus indented; distal maxillary palpomere evenly convex, without a fovea…................................................................... Liogenys macropelma Bates, 1887 - Clypeus quadridentate (Fig. 1A–D), anterior and lateral margin of clypeus indented; distal maxillary palpomere with an elongate, concave fovea................................................................................ 2 2. Angle between lateral and anterior teeth on clypeus slightly or strongly acute (Fig. 1C); males with lateral expansion along the parameres length entirely visible in dorsal view (Fig. 1F)...................................................... 3 - Angle between lateral and anterior teeth on clypeus right or obtuse (Fig. 1A); males with lateral expansion along the parameres at least partially hidden in dorsal view or absent (Fig. 1E)..................................................... 4 3. Frons and clypeus glabrous (Fig. 1C–D); angle between lateral and anterior teeth on clypeus strongly acute; pronotum moderate coarsely punctate; males with pygidium convex medially (Fig. 7D) and females with pygidium strongly inflated (Fig. 7E)...................................................................... Liogenys quadridentata Blanchard, 1851 - Frons and clypeus with bristles (Fig. 2C); angle between lateral and anterior teeth on clypeus slightly acute; pronotum strongly coarsely punctate; pygidium almost flat, sometimes slightly depressed medially (Fig. 2F)............................................................................................ Liogenys clipeosetosa Cherman, new species 4. Ventrites II, III, and sometimes IV with tubercles medially, tubercles variable in strength (Fig. 8E)...................................................................................... Liogenys schneiderae Cherman, new species - All ventrites smooth, without tubercles.................................................................... 5 5. Clypeal emargination clearly wider than the dorsal diameter of one eye (Fig. 1B)..... Liogenys quadridens (Fabricius, 1798) - Clypeal emargination as wide as or narrower than the dorsal diameter of one eye (Fig. 4B)........................... 6 6. Body and pronotum purplish brown, strongly darker than elytra; clypeal emargination strongly angulate; pronotum with anterolateral margin reflexed (Fig. 3C); pygidium without sexual dimorphism, weakly convex in both sexes.......................................................................... Liogenys genieri Smith & Cherman, new species - Body and pronotum reddish brown, slightly darker than elytra; clypeal emargination rounded; pronotum with anterolateral margin not reflexed, pygidium weakly convex in males and strongly convex to inflated in females....................................................................................... Liogenys granadina Cherman, new species, Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on page 203, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886, {"references":["Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. & Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160.","Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp.","Fabricius, J. C. (1798) Supplementum entomologiae systematicae. Proft et Storch, Copenhagen, 572 pp."]} more...
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22. Liogenys granadina Cherman 2021, new species
- Author
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
- Subjects
Coleoptera ,Liogenys granadina ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys granadina Cherman, new species Figs. 4, 9. Type material. Holotype male, labeled: [white handwritten] “N. Grenada / Cartagena / coll[?] Chev.”, [red typeset and handwritten] “ LIOGENYS / GRANADINA / HOLOTYPE / Cherman M. A.”, genitalia mounted (ZMHB). Paratypes (45), all bearing the label [yellow, typeset and handwritten] “ LIOGENYS / GRANADINA / PARATYPE / Cherman M. A.”: One male bearing: [white handwritten] “ Venezuela / Orinoco river [?]”, [white typeset] “ Liogenys / [handwritten] quadri- / dens F / [typeset] det. G. Frey, 1968”, genitalia mounted (ZMHB). One female with the labels: [white handwritten] “nov. grenada”, [white typeset] “ Liogenys / [handwritten] quadridens / Mos / typeset] det. G. Frey, 1968” (ZMHB). One male with the data: [white, typeset] “ TRINIDAD: St. George Co. / Monos Island. 10º42’Nx / 61º41’W. 31 MAY 1987 / ROM#872012. M.L. Pickles ” (ROME). One male with the data: [pink, handwritten] “Columbia”, [white, typeset] “Pascoe / Coll. / 93—60”, [white, handwritten] “Telea / 4dentata / [illegible]” (BMNH). One male with the data: [green, handwritten] “Columbia / Lebas.”, [light blue, handwritten] “4-dentatus / Blanch. Cat. / p. 168”, [white, typeset] “So named / in Reiches / Collection. / C.W.”, [light blue, handwritten] “2259”, [green, handwritten] “ ♂ ”, [white, typeset] “67.45”, [white, handwritten] “4-dentatus. / Blanch. Dej. / brevicollis, / Reich. / Carthagena” (BMNH). Seven males and three females with the data [white, typeset] “ VENEZUELA, Guar. / 12kmS Carabozo / 6 II 1969 ”, [white, typeset] “P&PSpangler / Est. Bio. Los / Llanos uv lite” (two males and one female at CMNC, two males and one female at DZUP, three males and one female at USNM). Paratypes of DZUP with the sequence of voucher numbers: [white typeset] “ DZUP / 402750 ” to “ DZUP / 402752 ”. Ten males and seven females with the data [white, typeset] “ VENEZUELA / Guar. 12 Km S. / Carabozo / II-6-12-1969 / P.&P. Spangler ”, [white, typeset] “Est. Biologica / Los Llanos / Black Light” (three males and two females at CMNC, two males and two females at CEMT, five males and three females at USNM). Five males and two females with the data [white, typeset] “ VENEZUELA, Guar. / 12kmS Carabozo”, [white, typeset] “ 6-12 II 1969 / P&PSpangler / BiolSta uv lite” (USNM). One male with the data [white, typeset] “ VENEZUELA, Guar. / 12kmS Carabozo”, [white, typeset] “ 6-12 II 1969 / P&PSpangler” (USNM). One female with the data [white, typeset and handwritten] “Cd Bolivar / 23-6 Venez”, [white, typeset] “EA Klages / Collector” (USNM). One female with the data [white, typeset and handwritten] “Cd Bolivar / 20-6 Venez”, [white, typeset] “EA Klages / Collector”, [white, typeset and handwritten] “ Liogenys / quadri- / dens / det. G.Frey,1968” (USNM). One female with the data [white, typeset and handwritten] “Cd Bolivar / 20-6 Venez”, [white, typeset] “EA Klages / Collector” (USNM). One female with the data [white, typeset] “ VENEZUELA: Guarico / Hato Masaguaral / (44km S Calabozo) / May 3–10 1985 / Menke & Carpenter” (USNM). Two females with the data [white, typeset and handwritten] “ VENEZUELA / Maracay / 450 m. / 15.VI.1949. / H.E. Box ” (USNM). Holotype and two paratypes deposited at ZMHB. Two paratypes at BMNH. Eight paratypes at CMNC. Three paratypes at DZUP. Four paratypes at CEMT. One paratype at ROME. Twenty-five paratypes at USNM. Diagnosis. Body light brown, elongate, sides almost parallel; elytra light brown; pronotum reddish brown (Fig. 4A); clypeus quadridentate; clypeal emargination deep, rounded, and narrow; distance between lateral and anterior clypeal teeth longer than basal width of one anterior tooth; angle between lateral and anterior clypeal teeth 90º or slightly acute; pronotal corners rounded (Fig. 4B); mesotibia quadrate in cross section; pygidium convex; pygidial disc glabrous, with scattered bristles on apex (Fig. 4D); in males protarsomere II more than twice the length of protarsomere I in dorsal view (Fig. 4C); parameres four times the length of their apex; lateral expansion along the parameres only seen subapically; inner margins convergent; apex harpoon shaped, lateral spine projected at approximately a 45º angle, spine length as long as the apex (Fig. 4E). Holotype. Male. Length: 9.2 mm, width: 4.6 mm. Light brown. Head: distance between eyes nearly twice the width of one eye; frons equal in length to clypeus; clypeal emargination deep, rounded, and narrow; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous tooth-like projection; distance between lateral and anterior teeth longer than basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye longer than one eye length, angle between anterior and lateral teeth approximately 90º; canthus not exceeding the outer margin of the eye; distal maxillary palpomere not seen; antenna with 10 antennomeres, club lighter in color and longer than funicle. Thorax: pronotal anterior margin moderately produced medially, undulate and flanged; maximum length of pronotum exceeding the length of tarsomeres I–III combined; disc glabrous, punctures very sparse and coarse; pronotal posterior corners rounded; hypomere with long bristles; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the length of the metacoxa; scutellum triangular with sparse, coarse punctures, mainly at the sides. Elytra: shiny, uniform light brown, slightly lighter in color than the pronotum; elytra more than three times longer than the pronotum; elytral suture darker than the elytron and weakly elevated; all four elytral ridges weakly defined. Legs: procoxa with abundant, thin and long scales, outer surface with short scales; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; protibial spur present; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia subquadrate in cross section, surface coarsely sculptured on outer margin, mesotibia with two transverse carinae, the apical carina incomplete; metacoxa with bristles throughout, basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia carinate and produced on apex, apical inner surface with sparse bristles, metatibial surface coarsely sculptured; metatibial transverse carina present posteriorly; protarsomere II elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres wider than mesotarsomeres and two times wider than metatarsomeres; metatarsomere I slightly shorter than metatarsomere II; claw bifid, outer tooth of a claw longer and as wide as the inner tooth; distance between teeth shorter than the inner tooth. Abdomen: disc of ventrites with bristles; propygidium with bristles, pygidium convex, subquadrate, wide; pygidial width exceeding distance between spiracles of propygidium; pygidial disc glabrous and apex with few bristles, pygidial disc coarsely punctate; pygidial apex subquadrate. Parameres: basal region dorsally narrowed, equal to both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; parameres four times the length of their apex; inner margins convergent; lateral expansion along the parameres only seen subapically; apex harpoon-shaped, lateral spine projected straight, at approximately a 45º angle, spine length as long as the apex. Parameres concave in lateral view (Fig. 4F). Variation. Female length: 9.5–11.4 mm, width 4.8–5.4 mm. Females as the holotype except in the size, head, and pronotum darker than elytra; pronotum wider, with pronotal corners obsolete; maximum width of distal maxillary palpomere slightly wider than at apex; fovea deep, extending up to the transverse midline of the palpomere; scutellum ogival. Male length: 8.4–10.2 mm, width 4.2–5.0 mm. Males as the holotype except in the scutellum ogival; punctures of pronotum and tibiae less marked; elytral suture unicolored with the elytron; ventral expansion of the outer margins of the parameres more apparent. Etymology. This species is named after Nueva Grenada, a defunct nation corresponding to present-day Colombia and parts of many other countries in South America (Brookes 1819). This name should be considered an adjective in the nominative singular. Type locality. Nueva Grenada (Republic of New Grenada, today a region that comprises Panama, many South American countries, and the total territory of Colombia). Geographical distribution. TRINIDAD AND TOBAGO (Saint George), COLOMBIA (Cartagena), VENEZUELA (Guaricó, Maracay). Remarks. Liogenys granadina Cherman, new species resembles L. quadridens in the shape of the quadridentate clypeus and of the pygidium. However, L. granadina is smaller than L. quadridens; the distance between anterior and lateral clypeal teeth is shorter; and in the pronotal corners are defined (obsolete in L. quadridens). Frey (1969) considerated specimens of L. granadina as being L. quadridens, judging by his description in the key, one of the two parameres illustration (the other one corresponds to L. quadridens), and because two paratypes (USNM and ZMHB) bear his determination label. Moreover, Frey (1969) refers to a specimen from Pernambuco, Brazil, which was found at the NHMB. This Brazilian specimen was identified as Liogenys piauiensis Cherman, 2017 by Cherman et al. (2019). There are three specimens in the USNM that were intercepted at the port of Miami, Florida on 1 May 1964 with no other information on the host or origin. This indicates that some previous port interceptions or pest reports for L. quadridens possibly included or even all of them are specimens of L. granadina., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 209-212, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886, {"references":["Brookes, R. (1819) General Gazeteer ... Illustrated with maps ... The fifteenth edition ... with considerable additions and improvements. 15 th Edition. Plummer and Brewis, Printers, Love-Lane, Little-Eastcheap, London, 796 pp.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Cherman, M. A., Basilio, D. S., Mise, K. M. & Almeida, L. M. (2019) Unraveling the puzzle of Liogenys biodiversity: fifteen new species, nomenclatural acts and new geographical records (Coleoptera: Scarabaeidae). Insect Systematics & Evolution, 51, 1 - 69. https: // doi. org / 10.1163 / 1876312 X- 00001040"]} more...
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23. Liogenys clipeosetosa Cherman 2021, new species
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Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T., and Almeida, Lúcia M.
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Coleoptera ,Insecta ,Arthropoda ,Liogenys clipeosetosa ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys clipeosetosa Cherman, new species Figs. 2, 9. Type material. Holotype male, labeled: [green typeset] “Venezuela / F. Kummerow S.”, [red typeset and handwritten] “ LIOGENYS / CLIPEOSETOSA / HOLOTYPE / Cherman M. A.”, genitalia mounted (ZMHB). Paratypes (10) all bearing the label [yellow, typeset and handwritten] “ LIOGENYS / CLIPEOSETOSA / PARATYPE / Cherman M. A.”. Six female paratypes bearing the same data of the holotype (ZMHB). Three female paratypes with the label: [green, outlined, typeset] “Hist. Coll. (Coleoptera) / Nr. 24131 / Liogenys quadridens Burm. / Columb., Moritz. / Zool. Mus. Berlin” (ZMHB). One female paratype with the data: [white, typeset] “ VENEZUELA: Lara / 20 km. E. Carora / 24-VI-1976 / Holotype and nine paratypes deposited at ZMHB. One paratypes at USNM. Additional material. Three females (FSCA) were port interceptions from Gulfport, Mississippi, United States of America in Musa (Musaceae) (bananas, plantains, etc.) originating from Colombia. We examined photographs of these specimens provided by John Leavengood (United States Department of Agriculture, Tampa, Florida, United States of America). No specific locality was available within Colombia and it is likely that these specimens were attracted to lights during the packing or loading of this commodity. Diagnosis. Body light brown; elongate (Fig. 2A–B), elytra light brown to yellowish brown, pronotum reddish brown, head darker; clypeus quadridentate; clypeal emargination deep, rounded, and wide; anterior teeth sometimes twice the length of the lateral teeth (Fig. 2C); distance between lateral and anterior clypeal teeth longer than basal width of one anterior tooth; angle between lateral and anterior clypeal teeth 90º or acute; clypeus and part of the frons with bristles; pronotum roughly punctate; pronotal corners rounded or subangulate; mesotibia quadrate in cross section (Fig. 2D); pygidium flat or slightly convex, depressed medially (Fig. 2E–F), pygidial disc with short bristles throughout, bristles longer on apex; males with protarsomere II up to twice the length of protarsomere I in dorsal view; parameres elongate, inner margins straight; lateral expansion along the parameres length entirely visible in dorsal view; apex harpoon-shaped, lateral spine projected backwards, slightly convergent compared to the outer margin of the parameres; spine length slightly longer than the apex, subapical and part of the apex not elevated on midline (Figs. 1E, 2G). Holotype. Male. Length: 12.0 mm, width: 5.8 mm. Light brown. Head: darker than pronotum, distance between eyes less than twice the width of one eye; frons equal in length to clypeus; frontoclypeal surface with bristles, clypeal emargination deep, rounded, and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous, tooth-like projection; distance between lateral and anterior teeth longer than basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye equal to one eye length, angle between anterior and lateral teeth approximately 90º; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width more than twice the width at apex; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than funicle. Thorax: pronotal anterior margin slightly produced medially, undulate and flanged; anteriorly and on posterolateral surface depressed, sulcate medially; pronotal disc glabrous, punctures sparse medially and rugose on anterior margin; pronotal posterior corners rounded; hypomere with few long bristles; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the width of the metacoxa; scutellum ogival, sparse punctures. Elytra: shiny, uniform yellowish brown, lighter in color than pronotum; elytra near three times the length of pronotum; elytral suture slightly darker than elytron and not elevated; all four elytral ridges weakly defined. Legs: procoxa with long bristles on infra-carinal surface, fine scales and short bristles on outer surface; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia quadrate in cross section, surface coarsely sculptured, mesotibia with two transverse carinae, the apical carina incomplete; metacoxa with bristles throughout; basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia carinate and produced on apex, apical inner surface with scattered bristles, metatibial surface coarsely sculptured; metatibial transverse carina and discontinuous longitudinal carina present posteriorly; protarsomere II elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres slightly wider than mesotarsomeres, more than twice the width of metatarsomeres; metatarsomere I and metatarsomere II equal in length; claw bifid, symmetrical, outer tooth of a claw longer and as wide as the inner tooth; distance between teeth shorter than the inner tooth. Abdomen: disc of ventrites with sparse bristles; propygidium with bristles, pygidium flat, subtrapezoidal, wide, pygidial width exceeding distance between spiracles of propygidium; pygidial disc glabrous except for a few bristles on apex, pygidial disc coarsely punctate; depressed medially; pygidial apex quadrate. Parameres: basal region slightly wider than both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; inner margins straight; lateral expansion along the parameres length entirely visible in dorsal view; apex harpoon-shaped, laterally expanded and blunt projected. Parameres concave in lateral view (Fig. 2H). Variation. Female paratypes. Length: 11.2–12.6 mm; width 5.4–6.3 mm. As the holotype except pygidial apex rounder, wider and somewhat convex, not depressed medially. Some females have the clypeal teeth noticeable longer. One female from Venezuela has a subtrapezoidal pygidium. Etymology. Adjective in the nominative singular. From the Latin clipeus (clypeus) + Latin setosus, from seta (“bristle”). Type locality. Venezuela. Geographical distribution. COLOMBIA, VENEZUELA (Lara). Remarks. Liogenys clipeosetosa Cherman, new species resembles L. quadridens in the body color and size, shape of the quadridentate clypeus, pygidium, and somewhat in the shape of the male parameres. Liogenys clipeosetosa differs (L. quadridens in parenthesis) in the clypeus with bristles (clypeus glabrous), fovea of the distal maxillary palpomere shallow, surface almost flat (fovea deep); pronotal anterior margin flanged (flange less conspicuous or absent); pronotum depressed anteriorly and on posterolateral surface, sulcate medially (not as so); pronotal disc roughly punctate mainly on anterior margin (pronotal disc less coarse and uniformly punctate); males with the protarsomere II longer than wide (as wide as it is long); apex of parameres entirely flat and lateral expansion along the parameres length entirely visible in dorsal view (subapical and proximal part of the apex elevated medially and lateral expansion of parameres partially hidden, visible from the midline up to the subapical portion)., Published as part of Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2021, Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species, pp. 201-226 in Zootaxa 4990 (2) on pages 203-206, DOI: 10.11646/zootaxa.4990.2.1, http://zenodo.org/record/5025886, {"references":["Fabricius, J. C. (1798) Supplementum entomologiae systematicae. Proft et Storch, Copenhagen, 572 pp.","Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp."]} more...
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24. Dinâmicas de (investig)ação em contextos de promoção da língua portuguesa: a presença do CIDTFF na rede Camões, I.P
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Araújo e Sá, M. H. (coord.), Ambrósio, Susana (coord.), Alexandre, Andrea Maria do Nascimento Martins, Almeida, Bárbara Mota de, Almenar, Ana, Basílio, Daniel, Bastos, Mónica, Batoréo, Hanna, Cardoso, Inês, Carinhas, Raquel, Carlos, Timóteo José, Neves, Alexandra Fidalgo das, Doval, Jimena, Faustino, Inês, Ferrari, Lucía, Freitas, Helena, Gomes, Maria Agostinha, Gonçalves, Maria de Lurdes, Gonçalves, Maria Jacinta, Graça, Luciana, Marialva, Marta, Melo, Sónia Rita, Melo-Pfeifer, Sílvia, Mendes, Pedro, Monteiro, Ana Catarina, Morais, Sónia, Moreira, Elisabete Vaz, Naturra, Barroso Paulo Roque Afonso, Nhacarize, Olga, Oliveira, Álvaro, Pereira, Cristina Baptista, Rodrigues, Brigite, Silva, Teresa, and Ussene, Abacar more...
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Ensino do Português no Estrangeiro - EPE ,Língua portuguesa ,Português língua de herança ,Investigação linguística ,Aquisição e aprendizagem de competências linguísticas ,Língua portuguesa - Educação intercultural ,Linguística portuguesa ,Ensino investigação linguística ,Português língua não-materna ,Instituto Camões ,Língua portuguesa - Ensino ,Investigação educativa - Abstract
Obra coordenadado por: Maria Helena Araújo e Sá e Susana Ambrósio Através da rede de Ensino de Português no Estrangeiro (EPE) ao nível do ensino básico, secundário e superior sob a tutela do Camões – Instituto da Cooperação e da Língua (Camões, I.P.) – pretende assegurar-se o objectivo da promoção da língua portuguesa enquanto desígnio inscrito nas grandes opções estratégicas da política externa portuguesa. A rede interage com comunidades portuguesas e luso-descendentes em diversos países na Europa, na América e em África, proporcionando o acesso à língua e à cultura portuguesa por parte de crianças e jovens e dinamizando, desse modo, comunidades escolares. O livro pretende reflectir sobre as vertentes de investigação e acção nas áreas abrangidas, reunindo 21 testemunhos provenientes de diversas partes do mundo liusófono. Este trabalho é financiado por Fundos Nacionais através da FCT – Fundação para a Ciência e a Tecnologia, I.P., no âmbito do projeto UIDB/00194/2020 info:eu-repo/semantics/publishedVersion more...
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25. Liogenys pallens Blanchard 1851
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Liogenys pallens ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys pallens Blanchard, 1851 Figs. 17; 25. Liogenys pallens Blanchard, 1851: 167; Lacordaire 1855: 269 (revision, systematics); Harold 1869: 1140 (checklist); Bruch 1911: 200 (checklist); Dalla Torre 1913: 318 (checklist); Blackwelder 1944: 227 (checklist); Frey 1969: 49, 61 (key, redescription); Krajčík 2012: 145 (checklist); Evans 2003: 212 (checklist); Evans & Smith 2009: 180 (checklist). Liogenys fulvescens Blanchard, 1851: 167; Lacordaire 1855: 269 (revision, systematics); Harold 1869: 1140 (checklist); Dalla Torre 1913: 318 (checklist); Blackwelder 1944: 227 (checklist); Frey 1969: 50, 61 (key, redescription); Evans 2003: 209 (checklist); Evans & Smith 2009: 178 (checklist); Krajčík 2012: 145 (checklist) New synonym. Liogenys flavicollis Frey, 1964: 693; Frey 1969: 40 (key); Evans 2003: 209 (checklist); Evans & Smith 2009: 177 (checklist); Krajčík 2012: 144 (checklist). New synonym. Type material. Liogenys pallens male lectotype here designated (MNHN): [white handwritten] “ Montevideo. / 7 bre 8 bre / 1820.”, [light green, typeset and handwritten] “MUSÉUM PARIS / Montevideo ”, [green handwritten] “ L. pallens / Cat. Mus. / Montevideo / M. A. St Hilaire. ”, [red typeset] “ SYNTYPE ”, [white, outlined red, typeset] “ LECTOTYPE / Liogenys pallens / Blanchard, 1851 / Cherman M. A. 2015”, genitalia mounted. Liogenys fulvescens female lectotype here designated (MNHN): [white handwritten] “ Montevideo. / 7 bre 8 bre / 1820.”, [light green, typeset and handwritten] “MUSÉUM PARIS / Montevideo ”, [green handwritten] “ L. fulvescens / Cat. Mus. / Montevideo / M. A. St Hilaire.”, [red typeset] “ SYNTYPE ”, [white, outlined red, typeset] “ LECTOTYPE / Liogenys fulvescens / Blanchard, 1851 / Cherman M. A. 2015”. Paralectotype (1): One female (MNHN) with the labels: [white handwritten] “Del’emb. / del’uruguay / jusqu’aux / missions”, [light green typeset] “MUSÉUM PARIS”, [red typeset] “ SYNTYPE ”, [white, outlined red, typeset] “ PARALECTOTYPE / Liogenys fulvescens / Blanchard, 1851 / Cherman M. A. 2015”. Liogenys flavicollis female holotype (NHMB): [white typeset] “ ♀ ”, [white handwritten] “ Formosa / Gran Guardia / XI.52”, [red typeset] “TYPE”, [white handwritten] “ Typus ♀ / Liogenys flavicollis / n. sp. / [typeset] det. G. Frey 1964 ”. Paratypes (2): One female with the same label as the holotype (NHMB). One female with the labels: [white handwritten] “ Formosa / Gran Guardia / XI.52”, [red typeset] “TYPE”, [white handwritten] “Typus F / Liogenys flavicollis / n. sp. / [typeset] det. G. Frey 1964 ”, [white typeset] “H. & A. HOWDEN / COLLECTION / ex. A. Martínez coll.”, [white typeset] “Canadian Museum of / Musée Canadian de la / NATURE / CMNEN 00018922” (CMNC). Non-type material (79). Without country locality, date, and collector, 2 females (ZMHB). PARAGUAY: Asunción, IX.1952, M. Alvarenga, 3 females (DZUP). ARGENTINA: without locality: Stempelm, 3 females (MZSP); Kraatz, 1 male and 5 females (SDEI); Formosa: Clorinda, 26.IX.1937, without collector, 2 males (MLPA); Chaco: Antequera, Departamento Primero de Mayo, 27°26.500’S 58°51.459’W, 52 m, 22.XII.2014, M. Ibarra Polesel, 1 specimen; Estación San Francisco, Departamento San Fernando, 27°30’35.90’’S 59° 4’47.60’’W, 54 m, 16– 17.X.2013, M. Ibarra Polesel, 1 specimen (CARTROUNNE); Santiago del Estero: without locality, E.R. Wagner, 1 female (MLPA); 7.XI.1963, Cartwright, 1 female (NHMB); Girardet, without date and collector, 1 male (MLPA); Choya, 27.XI.1962, without collector, 1 male (NHMB); Catamarca: without locality, date, and collector, 1 male (MLPA); La Rioja: 20 km N La Rioja, 25.IX.1968, without collector, 5 males and 1 female (AMNH), Córdoba: without locality, date, and collector, 3 males and 6 females (ZMHB); without locality, date, and collector, 2 females (MLPA); Córdoba, 10.XI.1903, C. Bruch, 1 male (ZMHB); San Luis: Potrero de los Funes, 20.XII.2012, M.A. Cherman, 1 female (DZUP); El Milagro, Ayacucho, XI.1966, A. Martínez, 1 female (CMNC); Mendoza: without locality, date, and collector, 5 males and 11 females. (ZMHB), without date, Jensen-Haarup, 1 female (ZMUC); Pedregal Railway Station, El Pedregal, Maipú, without date, Jensen-Haarup, 6 males and 8 females (ZMUC); Santa Rosa Railway Station, Santa Rosa, Jensen-Haarup, 1 female (ZMUC); Santa Rosa, Ñancuñan, 33º50’81.5’’S 67º53’71.7’’W, 5.I. 2005, 540 m, E. Ruiz, 1 male (IADIZA); Santa Rosa, Ñancuñan, 34°2’57.840’’S 67° 58’0.480’’W, 11.II.1982, S. Claver & S. Roig, 1 male (IADIZA); La Pampa: Santa Rosa, XI.1961, Daguerre, 1 female (NHMB); (1) Buenos Aires: La Plata, without date and collector, 1 female (ZMHB). Diagnosis. Length: 12.0– 15.5 mm; width 5.7–7.3 mm. Body elongate, dull yellow, more uniform in color in males (Fig. 17A) than in females (Fig. 17 B–C) semi-opaque to shiny, head and base of elytra darker; distance between eyes twice as wide as one eye in males, wider in females; clypeus in males long, clypeal emargination shallow, rounded, almost straight, wider than the frons; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; clypeal lateral margin convex, projection sharp or blunt; distance between clypeal lateral projection and anterior margin of eye as long as one eye; distance between clypeal lateral projection and the adjacent anterior tooth equal to the basal width of the anterior tooth; obtuse angle between outer side of anterior teeth and clypeal lateral projection; in females, clypeal emargination deep, rounded, wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; clypeal lateral margin convex, projection sharp; distance between clypeal lateral projection and anterior margin of eye longer than one eye; distance between clypeal lateral projection and the adjacent anterior tooth equal to or longer than the basal width of the anterior tooth; right angle between outer side of anterior teeth and clypeal lateral projection; antenna with 10 antennomeres; pronotal posterior corners rounded; scutellum ogival, finely punctate throughout; three dark protibial teeth, middle and apical equally large; distance between basal and middle teeth longer than between middle and apical; mesotibia cylindrical in cross section, with two transversal carinae, the apical carina incomplete in males, complete in females; inner margin of male metatibia not carinate; two metatibial transverse carina present posteriorly; metatarsomere I shorter than metatarsomere II; in males protarsomeres and mesotarsomeres I to IV strongly enlarged; claw bifid, symmetrical, inferior tooth strongly reduced; pygidium convex, pygidial disc glabrous or slightly with bristles on apex, finely punctate, coarser in females; pygidial apex rounded (Fig. 17H); parameres, basal region equal in width to the parameres at the midline; parameral split at the apical third; inner margins of parameres straight; parameres narrower subapically; apex strongly bent downwards, ventral surface expanded laterally forming an acute angle (Fig. 17I); parameres in lateral view straight, not coplanar (Fig. 17J). Type locality. Liogenys pallens and L. fulvescens: URUGUAY, Montevideo; L. flavicollis: ARGENTINA, Formosa, Gran Guardia. Geographical distribution. PARAGUAY (Asunción); ARGENTINA (Formosa, Chaco, Santiago del Estero, Catamarca, La Rioja, Córdoba, San Luis, Mendoza, La Pampa, Buenos Aires); URUGUAY (Montevideo). Remarks. Frey (1969) designated a neotype for L. pallens, which is housed at the NHMB. As we found the primary type of Blanchard (1851) at the MNHN (Fig. 17A, D), the former must be invalidated (Article 75.8, International Commission on Zoological Nomenclature 1999). Despite Blanchard (1851) mentioned a range of length in L. pallens description, we found a single type specimen, which is here designated as the lectotype. This species has a strong sexual dimorphism and many characters vary within its distribution. A large series with males and females from the same collection event from Mendoza (Argentina) allowed the association of males and females. After comparing the male lectotype of L. pallens, the female lectotype of L. fulvescens (Fig. 17B, E) and the female holotype of L. flavicollis (Fig. 17C, F), we propose that both latter names are synonyms of L. pallens., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 39-41, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240.","Lacordaire, J. T. (1855) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes. Librairie Encyclopedique de Roret, Paris, 594 pp. https: // doi. org / 10.5962 / bhl. title. 67686","Harold, E. F. von (1869) Scarabaeidae. In: Gemminger, M. & von Harold, E. F. (Eds.), Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus, Vol. 4. E. H. Gummi, Munich, pp. 976 - 1346.","Bruch, C. (1911) Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de la Plata, 17, 181 - 225. https: // doi. org / 10.5962 / bhl. title. 8799","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, W. (Ed.), Coleopterorum catalogus, Volumen XX. W. Junk, Berlin, Germany, pp. 291 - 450.","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Frey, G. (1964) Neue Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 15, 691 - 701.","International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp."]} more...
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26. Liogenys denticeps Blanchard 1851
- Author
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys denticeps ,Liogenys ,Taxonomy - Abstract
Liogenys denticeps Blanchard, 1851 Figs. 5; 25. Liogenys denticeps Blanchard, 1851: 167; Lacordaire 1855: 269 (systematics); Harold 1869: 1140 (checklist); Dalla Torre 1913: 318 (checklist); Blackwelder 1944: 227 (checklist); Evans 2003: 208 (checklist); Evans & Smith 2009: 176 (checklist); Krajčík 2012: 144 (checklist); Cherman et al. 2017: 4, 20 (generic history, natural history). Liogenys denticulatus Moser, 1918: 98; Frey 1969: 51 (key); Krajčík 2012: 144; Cherman et al. 2017: 4 (generic history). New synonym. Liogenys denticulata: Blackwelder 1944: 227 (checklist); Evans 2003: 208 (checklist); Evans & Smith 2009: 176 (checklist). Type material. Liogenys denticeps female lectotype (MNHN): [white handwritten] “1496”, [white handwritten] “649934”, [light green, typeset and handwritten] “MUSÉUM PÁRIS / Bolivie / d’Orbigny”, [green handwritten] “ L. denticeps / Cat Mus / Santa-Cruz (Bolivie). / M. d’Orbigny. ”, [red typeset] “ HOLOTYPE ”. Liogenys denticulatus male lectotype here designated (ZMHB): [white typeset] “ Argentinien / Santiago del Estero ”, [white handwritten] “ Liogenys / denticulatus / Mos / Typen ♂”, [light red typeset] “Typus”, [white, outlined red, typeset and handwritten] “ LIOGENYS / DENTICULATA / Moser, 1918 / LECTOTYPE / Des. M. A. Cherman”, genitalia mounted. Paralectotypes (1) (ZMHB): [white typeset] “ Argentinien / Santiago del Estero ”, [white handwritten] “ Liogenys / denticulatus / Mos / Typen ♂”, [light red typeset] “Typus”, [white, outlined red, typeset and handwritten] “ LIOGENYS / DENTICULATA / Moser, 1918 / PARALECTOTYPE / Des. M. A. Cherman”, genitalia mounted. Non-type material (38). BOLIVIA. Santa Cruz: San Antonio de Parapetí, 1–2.IX.1964, B. Malkin, 1 male (MZSP). ARGENTINA. Jujuy: Los Perales, I.1951, F. Justus Jor., 1 female (DZUP); Salta: Los Andes, without date and collector, 1 female (CMNC); “ Chaco austral” [Argentinian area from the Bermejo river southward to latitude 30°S, where the Pampas begin]: without locality and date, Auld [?], 1 female (MLPA); Santiago del Estero: Chuna pampa, XII–III.1909, E.R. Wagner, 4 females (MNHN); Icaño, Mistol Paso, 1909 [without month], E.R. Wagner, 2 males and 3 females (MNHN); Icaño, Mistol Paso, XII–I.1909, E.R. Wagner, 2 females (MNHN); Icaño, Mistol Paso, XII–III.1909, E.R. Wagner, 1 male and 8 females (MNHN); Icaño, Mistol Paso, XII.1909, E.R. Wagner, 1 male and 1 female (MNHN); La palisa del bracho (25 km NW Icaño), 1909 [without month], E.R. Wagner, 3 females; XII.1909, 1 female; XII–III.1909, 2 males (MNHN); without locality, date, and collector, 1 female (NHMB); El Pinto [overwritten], XI.1956, 1 male (CNCI); Colonia Dora, X.1956, without collector, 1 female (CNCI); XI.1945, F. Justus Jor., 2 females (DZUP). Diagnosis. Body elongate, brown, shiny; clypeal emargination deep, rounded and wide; lateral margin convex and strongly produced, forming a tooth-like projection; acute angle between the outer side of anterior teeth and clypeal lateral projection; pronotal posterior corners rounded (Fig. 5 A–C); basal apophysis of metacoxa not produced (see Cherman et al. 2016); metatibia weakly or not carinate along the inner margin; inferior tooth of a tarsal claw perpendicular to the axis of the superior tooth; ventrites with scattered bristles (Fig. 5J); pygidium glabrous (Fig. 5 G–I); in males protarsomere II more than twice as long as it is wide; parameres straight, apex curved downwards forming a reniform structure, longitudinally carinate, narrowed on inferior margin, sculptured (Fig. 5K). Redescription. Length 9.8–12.7 mm; width: 4.5–6.9 mm. Brownish. Head: distance between eyes twice as wide as one eye, wider in females; clypeal emargination deep, rounded, and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth as long as the eye or slightly shorter; lateral margin convex and strongly produced, forming a tooth-like projection; distance between clypeal lateral projection and anterior margin of the eye as long as one eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth equal to the basal width of the anterior tooth; acute angle between outer side of anterior teeth and clypeal lateral projection; distal maxillary palpomere, maximum width slightly wider than the apex; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than the funicle. Thorax: anterior margin of pronotum straight; glabrous, punctures coarse and dense, even denser near the anterior margin; pronotal posterior corners rounded; hypomere with short and long bristles; mesepisternum scaly; sides of metaventrite with sparse erect bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, coarsely punctate throughout. Elytra: shiny, glabrous, from brown to purplish brown; sometimes apparently burnt; elytral suture and elytron unicolored, weakly elevated; pair of inner ridges more defined than the three outer pairs. Legs: three protibial teeth, the middle and apical equal in size; distance between basal and middle teeth longer than between middle and apical, especially in males; mesofemural surface setose, thick erect bristles on posterior margin; mesotibia subquadrate or cylindrical in cross section, surface coarsely sculptured; two transverse carinae, the apical carina in general incomplete in males and complete in females; metacoxa with sparse bristles; basal apophysis of metacoxa not produced; metatibia weakly or not carinate along the inner margin, apical inner surface setose; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV enlarged; slightly wider than the metatarsomeres; protarsomere II more than twice longer than it is wide; metatarsomere I shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer than the inferior and equal in width, distance between teeth more than twice the length of the inferior tooth; inferior tooth perpendicular to the axis of the superior tooth. Abdomen: ventrites with scattered bristles on disc; propygidium slightly visible, glabrous; pygidium slightly convex in males (Fig. 5G), flat in females (Fig. 5 H–I); subquadrate, as wide as it is long in males, elongate in females; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex rounded. Parameres: basal region equal in width to both sections of the parameres at its maximum width, parameral split at the third portion, inner margins straight and close together; curved downwards at the apex forming in a frontal view a reniform structure, sculptured, longitudinally carinate, narrowed on inferior margin; parameres in lateral view straight, coplanar (Fig. 5L). Type locality. Liogenys denticeps: BOLIVIA, Santa Cruz [Santa Cruz de la Sierra Department]; L. denticulata: ARGENTINA, Santiago del Estero. Geographical distribution. BOLIVIA (Santa Cruz), ARGENTINA (Jujuy, Salta, Santiago del Estero). Remarks. Liogenys denticeps female lectotype (Fig. 5C, F) was compared with L. denticulata female paralectotype (Fig. 5B, E), so we here propose that L. denticulata is the junior synonym of L. denticeps. Liogenys denticeps resembles L. cartwrighti (Fig. 2), mainly in the size, color, shape of the clypeus, and pronotal punctuation. It differs from L. cartwrighti (in parenthesis) in the pronotal posterior corners rounded (obtuse angle); in the punctuation of elytra; the pygidium is wide in males—although the maximum width does not exceed the distance between spiracles of propygidium, long in females (pygidium as wide as it is long in both sexes), with disc glabrous (with sparse bristles throughout); in males the protarsomeres and mesotarsomeres are slightly enlarged (strongly enlarged), and in the shape of the parameres. The parameres of L. denticeps are quite similar to L. pallens Blanchard, 1851 (Fig. 17), and the females are similar, but the external morphology of the males of each species are distinguishable (see diagnosis of L. pallens). The redescription and parameres illustration of L. denticeps in Frey (1969) actually belong to L. latitarsis., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 13-14, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240.","Lacordaire, J. T. (1855) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes. Librairie Encyclopedique de Roret, Paris, 594 pp. https: // doi. org / 10.5962 / bhl. title. 67686","Harold, E. F. von (1869) Scarabaeidae. In: Gemminger, M. & von Harold, E. F. (Eds.), Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus, Vol. 4. E. H. Gummi, Munich, pp. 976 - 1346.","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, W. (Ed.), Coleopterorum catalogus, Volumen XX. W. Junk, Berlin, Germany, pp. 291 - 450.","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Cherman, M. A., Moron, M. A. & Almeida, L. M. (2016) Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology, 41, 744 - 770. https: // doi. org / 10.1111 / syen. 12188"]} more...
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27. Liogenys Guerin-Meneville 1831
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Identification key to Liogenys species from the Chacoan province and its boundaries 1. Antenna with 10 antennomeres; clypeal emargination wide or narrow; outer sides of anterior clypeal teeth parallel (Figs. 2B, 3B) or following the lateral margin of clypeus (Figs. 9B, 21B).................................................. 2 - Antenna with 9 antennomeres; clypeal emargination narrow and semicircular (Fig. 12E); outer sides of anterior clypeal teeth following the lateral margin of clypeus.......................................... Liogenys mendozana Moser, 1918 2. Clypeal teeth very short (Fig. 4B), outer sides of teeth not forming an angle with the lateral margin of clypeus; lateral margin convex or straight, not produced......................................................................... 3 - Clypeal teeth long or short (Figs. 2B, 8B), outer sides of teeth forming an angle with the lateral margin of clypeus, lateral margin convex and sometimes strongly produced (Fig. 6B)....................................................... 8 3. Pronotal disc densely punctate (Fig. 3B)................................................................... 4 - Pronotal disc sparsely punctate (Figs. 2B, 10C).............................................................. 6 4. Pronotum and elytra glabrous; punctures of pronotum very fine and dense (Fig. 4B); pygidium with dense scales (Fig. 4D), scales sparse only at the apex...................................................... Liogenys densata Frey, 1969 - Pronotum and elytra setose; punctures of pronotum coarse and dense (Fig. 3B); pygidium with sparse bristles and scales... 5 5. Clypeal emargination semicircular, narrow, lateral margin of clypeus never produced; in males inner margin of metatibia concave subapically (Fig. 16F)................................................ Liogenys opacicollis Fairmaire, 1892 - Clypeal emargination rounded, wide, lateral margin of clypeus concave, produced in most of the cases, forming a blunt projection (Fig. 3B); in males inner margin of metatibia convex subapically (Fig. 3F).......... Liogenys cribricollis Moser, 1921 6. Lateral margin of clypeus concave, in males tarsomeres almost equal in width in all legs, cylindrical (Fig. 9E)............................................................................... Liogenys isotarsis Cherman, new species - Lateral margin of clypeus straight (Fig. 21B), in males protarsomeres and mesotarsomeres enlarged.................... 7 7. Elytra setose, shiny; body length less than 8 mm............................. Liogenys truncata Cherman, new species - Elytra glabrous, semiopaque; body length 9–10 mm.................................. Liogenys rectangula Frey, 1969 8. Pronotum roughly punctate; lateral sides of abdomen, propygidium, and pygidium covered by wide scales (Fig. 20C)................................................................................. Liogenys rugosicollis Frey, 1969 - Pronotum somewhat coarsely punctate; sides of abdomen, propygidium, and pygidium with bristles (Fig. 10D)........... 9 9. Width of clypeal teeth combined three or more times longer than the length of one tooth (Figs. 14B, 18B).............. 10 - Width of clypeal teeth combined less than three times longer than the length of one tooth (Figs. 6B, 8B)............... 17 10. Epipleuron of elytra uniformly clothed by fringed-like, long, erect bristles (Fig. 18A, E); legs with tarsomere II twice as long as tarsomere I, or almost so, especially in male....................................... Liogenys parallela Frey, 1965 - Epipleuron of elytra with sparse, short bristles; legs with tarsomere II less than twice as long as tarsomere I............. 11 11. Pygidium subquadrate (Fig. 14D), apex angulate, punctures very coarse; clypeal emargination deeper than the outer margin of anterior teeth (Fig. 14B)...................................................... Liogenys nigrofusca Moser, 1918 - Pygidium trapezoidal (Figs. 2 D–E; 5G–H; 15G), apex subangulate or rounded; clypeal emargination shallower than the outer margin of anterior teeth (Fig. 10C)....................................................................... 12 12. Lateral margin of clypeus weakly produced, outer sides of anterior teeth obtuse or subparallel, without angle among anterior teeth and lateral margin of clypeus (Fig. 15E).............................................................. 13 - Lateral margin of clypeus strongly produced, outer sides of anterior teeth almost parallel, right or acute angle between anterior and lateral teeth (Fig. 10C)............................................................................. 14 13. Propygidium scaly; pygidium with erect and thick setae............................ Liogenys obscura Blanchard, 1851 - Propygidium with sparse bristles; pygidium with decumbent bristles.................... Liogenys cartwrighti Frey, 1969 14. Pygidium glabrous, or with few bristles on apex; distance between teeth of each tarsal claw as long as the length of superior tooth.............................................................................................. 15 - Pygidium setose, distance between teeth of each tarsal claw shorter than the length of inferior tooth................... 16 15. Body shiny, head and pronotum unicolored brown or dark brown in both sexes; clypeal emargination narrow, without clear sexual dimorphism in the clypeal margin; males with protarsomere II longer than wide (Fig. 5J)............................................................................................... Liogenys denticeps Blanchard, 1851 - Body opaque, head and pronotum dull yellow unicolored in males, not unicolored in females, head dark brown or black; clypeal emargination very wide, in females emargination as wide as one half the distance between eyes, in males wider; males with protarsomere II as long as wide or wider than long (Fig. 10D)........................ Liogenys pallens Blanchard, 1851 16. Metafemur glabrous, row of bristles along its margins; males with parameres angulate at the midline, without subapical constriction (Fig. 11)............................................................. Liogenys latitarsis Moser, 1918 - Metafemur setose throughout; males with parameres globose at the midline, with subapical constriction (Fig. 22E)....................................................................................... Liogenys vicina Frey, 1969 17. Clypeal emargination angulate (Fig. 8B); fovea of distal maxillary palp occupying the entire dorsal surface, almost reaching the apex (Fig. 8C)......................................................... Liogenys foveata Cherman new species - Clypeal emargination rounded; fovea of distal maxillary palp extending past the midline but not reaching the apex....... 18 18. Base of pygidium less than twice as wide as the apex (Fig. 7E); in males, the three pairs of tarsi equally enlarged...................................................................................... Liogenys tarsalis Moser, 1921 - Base of pygidium twice as wide as the apex (Figs. 7F, 13D); in males, protarsi and mesotarsi wider than the metatarsi.... 19 19. Base of metacoxae and trochanter setose (Fig. 13E); in males, parameres glabrous (Fig. 13F).......................................................................................... Liogenys neoforcipata Cherman, new species - Base of metacoxae and trochanter almost glabrous (Fig. 6D); in males, parameres setose (Fig. 6F)... Liogenys forcipata Frey, Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 3-4, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Fairmaire, M. L. (1892) Descriptions de quelques coleopteres Argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253.","Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312","Frey, G. (1965) Neue s ʾ damerikanische Melolonthidae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 16, 143 - 151.","Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240."]} more...
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28. Liogenys obscura Blanchard 1851
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Liogenys obscura ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys obscura Blanchard, 1851 Figs. 15; 24. Liogenys obscurus Blanchard, 1851: 167; Lacordaire 1855: 269 (systematics); Harold 1869: 1140 (checklist); Steinheil 1874: 557 (checklist); Kolbe 1907: 5 (checklist); Bruch 1911: 200 (checklist); Dalla Torre 1913: 318 (checklist); Frey 1969: 43 (key); Krajčík 2012: 145 (checklist). Liogenys obscura: Blackwelder 1944: 227 (checklist); Evans 2003: 211 (checklist); Evans & Smith 2009: 180 (checklist). Liogenys bruchi Moser, 1924: 122; Blackwelder 1944: 227 (checklist); Frey 1969: 58 (synonymy). Type material. Liogenys obscurus female lectotype here designated (MNHN): [green handwritten] “ L. obscurus / Cat. Mus. / Rio negro (Patagonie) / M. D’ Orbigny”; [white handwritten] “6049 / 34”, [white handwritten] “1050”, [light green typeset and handwritten] “MUSÉUM PARIS / Chiquitos d’Orbigny”, [white, outlined black, typeset] “Southern Neotropical Scarabs / database # AS 2615205 / Liogenys obscura / Blanchard, 1851 ♀ / DET: A.B.T.SMITH 2008”, [red, outlined black, typeset and handwritten] “ LIOGENYS / OBSCURA / BLANCHARD, 1851 / LECTOTYPE / Des. Cherman M. A. Paralectotypes (2): one female (MNHN): [white handwritten] “6049 / 34”, [white typeset] “MUSEUM PARIS / PATAGONIE / (PATAGONES) / D’ORBIGNY 1834”, [white, outlined black, handwritten and typeset] “ Liogenys / obscurus / DET. / L. W. SAYLOR”, [white, outlined black, typeset] “Southern Neotropical Scarabs / database # AS 2615206 / Liogenys obscura / Blanchard, 1851 ♀ / DET: A.B.T.SMITH 2008”; one female (MNHN): [white handwritten] “6049 / 34”, [white typeset and handwritten] “MUSÉUM PARIS / Chiquilos / d’Orbigny”, [white, outlined black, typeset] “Southern Neotropical Scarabs / database # AS 2615207 / Liogenys obscura / Blanchard, 1851 ♀ / DET: A.B.T.SMITH 2008”. Liogenys bruchi Moser male lectotype here designated: [white handwritten] “ Liogenys / bruchi / Moser / Typen”, [white, typeset and handwritten] “Alta Gracia La Granja / Sierras de Córdoba / IX.922 [typeset] C. Bruch Leg”, [white, typeset and handwritten] “ Liogenys / obscurus Bl / det. G. Frey, 1968”, [red typeset] “Typus” (ZMHB), [white, outlined red, typeset] “ LECTOTYPE / Liogenys bruchi / Moser, 1924 / des. M. A. Cherman” (ZMHB). Paralectotypes (2): one male (MACN): [white, typeset and handwritten] “Alta Gracia La Granja / Sierras de Córdoba / XII.921 / C. Bruch Leg.”, [red handwritten] “ Liogenys / bruchi / Moser 1924 / Syntypus?”, [green typeset] “Typus”, [white handwritten] “FICHADO”, [white, outlined black, typeset] “MACN–En / 952”, [white, outlined red, typeset] “ PARALECTOTYPE / Liogenys bruchi / Moser, 1924 / Des. M. A. Cherman”. One male (MACN): [white, typeset and handwritten] “Alta Gracia La Granja / Sierras de Córdoba I.922 / C. Bruch Leg”, [red handwritten] “ Liogenys / bruchi / Moser 1924 / Syntypus?”, [green typeset] “Cotypus”, [white handwritten] “FICHADO”, [white, outlined black, typeset] “MACN –En / 953”, [white, outlined red, typeset] “ PARALECTOTYPE / Liogenys bruchi / Moser, 1924 / Des. M. A. Cherman”. Non-type material (65). ARGENTINA. Without locality and collector, 1944, 1 female (MZSP); Catamarca: Anacasti, 18.XII. 2009, 573 m, 28º52’57’’S 65º25’04’’W, Ocampo, Dominguez, & Campon, 1 male and 2 females (IADIZA); Pomán (Camping), 7.XII.2013, Arriagada & Castillo, 1 female (CMVD); Santiago del Estero: Río Salado, without date, E.R. Wagner, 1 male (MZUC); Córdoba: Cruz del Eje (4 km NE), 20.II.1982, A. Martínez, 3 males and 20 females (CMNC); Capilla del Monte. Punilla, 21.XI.1965, A. Martínez, 1 male and 1 female (CMNC); 3.XII.1997, G. Arriagada, 2 males and 2 females (CMVD); Diquecito, II.1977, A. Martínez, 1 male (CMNC); Diquecito. El Sauce, XII.1964, A. Martínez, 3 males (MZSP); Río Cuarto (45 mi N), 11.VII.1951, Ross & Michelbacher, 1 male (MZSP); without date, Bremer, 2 females (ZMHB); San Vicente, without date, Frenzel, 1 female (ZMHB); San Luis: La Toma, km 73, Toll Autopista El Saladillo, 33,0715633ºS 65,654522ºW, 23.XII.2012, M.A. Cherman, 2 females (DZUP); Desaguadero, 15.I.2001, J. Jensen, 2 females (CMVD); Quines, 24.XI.1995, Roig & Flores, 2 males and 1 female (IADIZA); El Molle. Departamento Belgrano, 33º02’22.8’’S 66º30’47.6’’W, 7.II. 2017, 622 m, G. Arriagada, 1 male and 7 females (CEMT); Mendoza: Telteca. Lavalle, 32º22’59.58’’S 68º03’14.16’’W, 5.II. 2008, 548 m, Ocampo, Ruiz, & San Blas, 1 female (IADIZA); Buenos Aires: Solá, II.1961, A. Martínez, 1 female (MZSP); Base Naval Puerto Belgrano, Punta Alta, [38.883333ºS, 62.083333ºW], 2.II.1938, Biraben & Scott, 3 males and 3 females (MLPA); “ Patagonia ”: without date, Guerin, 1 female (ZMUC). URUGUAY. Colonia: Carmelo, without date, M. Palmer, 1 female (CMNC); Montevideo: Montevideo, without date and collector, 1 female (CMNC). Diagnosis. Length: 10.9–11.6 mm; width: 5.5–6.2 mm. Dark brown, shiny, pronotum and elytra unicolored; head and pronotum coarsely and densely punctate (Fig. 15B); distance between eyes more than three times the width of one eye; frons swollen and somewhat scaly; clypeal emargination deep, wide and rounded in males, more angulate in females; outer sides of anterior teeth parallel, subparallel in females; outer margin of anterior teeth shorter than the eye; clypeal lateral margin convex, blunt projection, in males sharper (Fig. 15E); distance between clypeal lateral projection and anterior margin of eye longer than one eye; distance between clypeal lateral projection and the adjacent anterior tooth equal to the basal width of the anterior tooth; in females obtuse angle between outer side of anterior teeth and clypeal lateral projection (Fig. 15A); in males angle obtuse or straight; antenna with 10 antennomeres; pronotal posterior corners rounded; scaly at the insertion with the pterothorax; scutellum rounded to ogival; mesepisternum, sides of metaventrite and metacoxae scaly abundantly (Fig. 15F); three protibial teeth, distance between basal and middle teeth slightly longer than between middle and apical; mesotibia quadrate to subquadrate in cross section, two transverse carinae, the apical one incomplete in males, complete in females; metatibial transverse carina present posteriorly; metatarsomere I one half the length of metatarsomere II and equal in width; in males pro and mesotarsomeres I to IV equally enlarged, protarsomere II as wide as it is long; claw bifid, symmetrical, superior tooth of a claw longer than the inferior; band of abundant scales beneath the outer margin of elytra; disc of ventrites scaly and with scattered bristles; pygidium convex, flatter in females; wide, subtrapezoidal; pygidial disc setose throughout, with thick erect bristles (Fig. 15G); parameres widened and swollen at the level of their split and narrowed towards the apex, parameral split at 2/3; inner margins straight (Fig. 15 H–I); parameres in lateral view straight, not coplanar (Fig. 15J). Type locality. Liogenys obscura: ARGENTINA, Río Negro (Patagonie); L. bruchi: ARGENTINA, “Alta Gracia La Granja, Sierras de Córdoba ”. Geographical distribution. URUGUAY (Montevideo, Colonia); ARGENTINA (Catamarca, Santiago del Estero, Córdoba, San Luis, Mendoza, Buenos Aires, Río Negro). Remarks. The type series specimens of L. bruchi were found at the MACN and ZMHB, although in their labels the date varies: XII.1921, I.1922 (MACN), and IX.1922 (ZMHB). Moreover, none of them match exactly with the month and/or year stated by Moser (1924) in the original description, which is XI—XII.1922. To maintain the stability of the name we designate the specimen of ZMHB as the lectotype, as this is the only one bearing the original label of type specimen written by Moser (Fig. 15B, D)., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 33-36, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240.","Lacordaire, J. T. (1855) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes. Librairie Encyclopedique de Roret, Paris, 594 pp. https: // doi. org / 10.5962 / bhl. title. 67686","Harold, E. F. von (1869) Scarabaeidae. In: Gemminger, M. & von Harold, E. F. (Eds.), Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus, Vol. 4. E. H. Gummi, Munich, pp. 976 - 1346.","Steinheil, E. (1874) Symbolae ad historian coleopterorum Argentiniae meridionalis, ossia enumeriazione dei coleotteri raccolti dal prof. P. Strobel nell'Argentinia meridionale, e descrizione dell species nuove, di Edoardo Stenheil. (Traduzione dal manoscritto tedesco inedito). Atti della Societa Italiana di Scienze Naturali, 15, 554 - 578.","Kolbe, H. (1907) Coleopteren. Ergebnisse der Hamburger Magalhaensische Sammelreise, 8, 1 - 125.","Bruch, C. (1911) Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de la Plata, 17, 181 - 225. https: // doi. org / 10.5962 / bhl. title. 8799","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, W. (Ed.), Coleopterorum catalogus, Volumen XX. W. Junk, Berlin, Germany, pp. 291 - 450.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Moser, J. (1924) Beitrag zur Kenntnis der Melolonthiden (Col.). Stettiner Entomologische Zeitung, 84, 114 - 136.","Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312"]} more...
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29. Liogenys neoforcipata Cherman 2021, new species
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Liogenys neoforcipata ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys neoforcipata Cherman, new species Figs. 13; 27. http://zoobank.org/ D8037B33-8328-4A65-883D-C1306A12B0EB Type material: Holotype labeled (CNCI): [white, typeset and handwritten] “ ARGENTINA / Santi. del Estero / Colonia Dora / X.1956 ”, [red, typeset and handwritten] “ LIOGENYS / NEOFORCIPATA / HOLOTYPE / Cherman M. A.”. Paratype (1) male (CNCI): [white typeset] “ ARGENTINA / Stgo. Del Estero / El Pinto [strikethrough], XI- 1956 ”, [yellow, typeset and handwritten] “ LIOGENYS / NEOFORCIPATA / PARATYPE Cherman M. A.”. Holotype and paratype deposited at CNCI. Diagnosis. Body elongate; brown, pronotum darker than elytra (Fig. 13A), head dark at vertex; clypeal emargination deep, rounded and wide; outer sides of anterior teeth parallel; clypeal lateral margin with subangulate projection (Fig. 13B); basal apophysis of metacoxa not produced, scaly as well as the margin of trochanter (Fig. 13E); in males tarsi shiny, protarsi and mesotarsi slightly enlarged; pygidium flat or slightly convex; trapezoidal; pygidial disc glabrous (Fig. 13D); parameres glabrous, slightly convergent; narrowed apically and apex rounded, curved downwards (Fig. 13 F–G). Holotype. Male. Length: 9.4 mm; width: 5.1 mm. Brown. Head: distance between eyes almost twice the width of one eye; clypeal emargination deep, rounded and wide; outer sides of anterior teeth parallel; inner margin of anterior teeth divergent, outer margin of anterior teeth shorter than the eye; clypeal lateral margin convex and strongly produced, forming a subangulate tooth-like projection; distance between clypeal lateral projection and anterior margin of the eye as long as one eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth equal to the basal width of the anterior tooth; angle between outer side of anterior teeth and clypeal lateral projection approximately 90º; distal maxillary palpomere, maximum width more than twice the apical width; sensorial region smooth or forming a shallow fovea, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and slightly longer of the funicle. Thorax: anterior margin of pronotum slightly produced medially; pronotal disc glabrous, punctures coarse and sparse; posterior corners rounded; hypomere with short bristles, mesepisternum scaly; sides of metaventrite with sparse bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, sparsely and finely punctate. Elytra: shiny, glabrous, brownish, lighter in color than pronotum; elytral suture darker than elytron and elevated, pair of inner ridges more defined than the three outer pairs. Legs: three protibial teeth, middle and apical equal in size, slightly longer than the basal tooth; distance between basal and middle teeth slightly shorter than between middle and apical; mesofemural surface setose, thick erect bristles on posterior margin; mesotibia subquadrate in cross section, mesotibial surface coarsely sculptured; two transverse carinae, the apical carina complete and strong; metacoxa scaly beneath the leg; basal apophysis of metacoxa not produced beyond the outer margin of trochanter, outer margin of the trochanter and metacoxa scaly; metafemur abundantly setose; inner margin of metatibia carinate towards the apex, inner surface setose; two strong metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males tarsi shiny; protarsomeres and mesotarsomeres I to III slightly enlarged; protarsomere II elongate; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and equal in width to the inferior, distance between teeth more than twice the length of the inferior tooth; teeth parallel. Abdomen: ventrites scarcely with bristles on disc, ventrite II more with bristles; propygidium slightly visible, glabrous; pygidium flat or slightly convex, subtrapezoidal, wide; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex subquadrate. Parameres: basal region as wide as both sections of the parameres at its maximum width, parameral split at 2/3; inner margins slightly convergent; apex rounded and curved downwards, inner margins slightly overlapped; parameres entirely glabrous; parameres in lateral view flattened and curved downwards from midline up to the apex; straight coplanar to the basal portion. Variation. Male paratype. Length: 10.0 mm; width: 5.4 mm. The paratype matches the holotype in all other characters described above. Etymology. Adjective in the nominative singular. Prefix from the Ancient Greek νέος (néos, “new, young”) + Latin forcipâtum (shaped like pincers or tongs). Liogenys neoforcipata is named in reference to the similarity with L. forcipata in external morphology, male genitalia, and geographical distribution. Type locality. ARGENTINA, Santiago del Estero, Colonia Dora [28°36’03.0’’S 62°57’00.3’’W]. Geographical distribution. ARGENTINA (Santiago del Estero). Remarks. Liogenys neoforcipata strongly resembles L. forcipata (in parenthesis) (Fig. 6), differing only in a few amount of features: the clypeal lateral projection weakly angulate (rounded); the antennal club shorter; the three protibial teeth almost equal in size, distance between basal and middle teeth slightly shorter than between middle and apical (middle and apical teeth equal in size, the three teeth equally spaced); the trochanter abundantly setose along its outer margin (glabrous, sparsely setose on its outer margin) (Fig. 6D); the mesotibia and metatibia subquadrate in cross section (cylindrical in cross section), the posterior carinae on metatibia stronger; the protarsi and mesotarsi shiny and slightly enlarged (opaque, almost twice the width of the metatarsomeres), the metatarsi are not enlarged (metatarsi enlarged); the space between the teeth of the tarsal claws is narrower than the width of the inferior tooth (more than twice the length of the inferior tooth); and in the shape of the parameres, with apex more rounded, not bilobed; the parameres are subapically glabrous, in lateral view flattened upon the apex and more curved downwards (setose, upon the apex uniform in height and straight). Females remain unknown., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 29-31, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258 more...
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30. Liogenys cartwrighti Frey 1969
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys cartwrighti ,Taxonomy - Abstract
Liogenys cartwrighti Frey, 1969 Figs. 2; 26. Liogenys cartwrighti Frey, 1969: 57; Evans 2003: 207 (checklist); Evans & Smith 2009: 176 (checklist); Krajčík 2012: 144 (checklist); Cherman et al. 2017: 5, 20 (generic history, natural history). Type material. Liogenys cartwrighti male holotype (USNM): [white handwritten] “Sgo. d. Estero / Argentinien ”, [white, typeset and handwritten] “ Liogenys / cartwrighti / n.sp. / Type / det. G Frey, 1968”, [red typeset] “TYPUS”, [pink typeset] “ Cartwright / Smithson.”, [red, typeset and handwritten] “ Type No / 74011 / USNM”, [white typeset] “ Loan from / USNMNH / 2065060”. Paratypes (2): one female paratype (NHMB) [white handwritten] “ Salta / Dep. Rivadavia / XII.950 Daguerre ”, [orange empty], [white typeset] “ ♀ ”, [pink typeset] “Property / USNM”, [red typeset] “ PARATYPE ”, [white typeset and handwritten] “ Liogenys / cartwrighti / n.sp. / det. G. Frey, 1968”, [white typeset] “Museum Frey / M̹nchen”. One female paratype (NHMB) [white handwritten] “ Salta / Dep. Rivadavia”, [white typeset] “ ♀ ”, [pink typeset] “Cartwrighti / Smithson.”, [red typeset] “ PARATYPE ”, [white, typeset and handwritten] “ Liogenys / cartwrighti / n.sp. / det. G. Frey, 1968”. Non-type material (12). ARGENTINA: Jujuy: without locality and date, R. Richter 1 male and one female (MLPA); Los Perales, I.1951, F. Justus Jor., 1 female (DZUP); Parque Nacional Calilegua, 29.XI.1986, J. Muzón, 1 male (IADIZA); near Parque Nacional Calilegua, 10.XII.2008, F.C. Ocampo, G. San Blas, & F. Campón, 1 male (IADIZA); Dique la Ciénaga, III.1953, A. Martínez, 1 male (MZSP); Salta: Orán, 11–12.X.1968, L. Peña, 2 males (NHMB); Dique Itiyuro- 5 km Salvador Mazza [22.0509ºS, 63.7101ºW], 15.XI.1995, S. Flores & S. Roig, 1 male (IADIZA); Formosa: Ruta Provincial 39, 23 km S Ingeniero Juárez, 24º05’27’’S 61º56’49’’W, 13–17.XII.2008, F. Ocampo, G. San Blas, & F. Campón, mercury vapor & ultraviolet light trap, 1 male (IADIZA); Tucumán: Lules, XII.1957, A. Martínez, 1 male (MZSP); Santiago del Estero: Ojo de Agua, III.1969, A. Martínez, 1 male (CMNC). Diagnosis. Body elongate, brown to dark brown, shiny (Fig. 2A); clypeal emargination deep but not reaching the base of the teeth, rounded and wide; lateral margin strongly convex and produced, forming a tooth-like projection; pronotal posterior corners in obtuse angle (Fig. 2B); basal apophysis of metacoxa produced; metatibia weakly or not carinate along the inner margin; superior tooth of a tarsal claw slightly longer than the inferior, distance between teeth shorter than the inferior tooth; pygidium with sparse bristles throughout (Fig. 2 D–E); ventrites with abundant bristles (Fig. 2C); in males protarsomere II as long as it is wide; parameres lyre shaped; ventral surface visible in dorsal view, concave (Fig. 2F). Redescription. Length 9.4–12.3 mm; width: 4.4–5.9 mm. Dark brown. Head: distance between eyes twice as wide as one eye; clypeal emargination deep, rounded and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; lateral margin strongly convex and produced, forming a tooth-like projection; distance between clypeal lateral projection and anterior margin of the eye as long as one eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth longer than the basal width of the anterior tooth; acute angle between outer side of anterior teeth and clypeal lateral projection; distal maxillary palpomere, maximum width twice the apical width; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than the funicle in males, as long as in females. Thorax: anterior margin of pronotum straight; glabrous, punctures fine and very sparse, denser and coarser near the anterior margin; pronotal posterior corners in obtuse angle; hypomere with long bristles, scales near the inner margin; mesepisternum scaly; sides of metaventrite with brisltes and scales; distance between mesocoxae and metacoxae slightly longer than the length of the metacoxa; scutellum ogival, weakly punctate at the sides. Elytra: shiny, glabrous, brown or dark brown; elytral suture and elytron unicolored, weakly elevated; all elytral ridges weakly defined. Legs: three protibial teeth, the middle and apical equal in size, basal one slightly shorter; distance between basal and middle teeth slightly longer than between middle and apical; mesofemural surface setose; mesotibia subquadrate or quadrate in cross-section, surface coarsely sculptured; two transverse carinae, the apical one incomplete in males and complete in females; metacoxa scaly, scales long; basal apophysis of metacoxa produced; metatibia not carinate along the inner margin, apical inner surface setose; metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger longer than the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV strongly enlarged; metatarsomeres slightly enlarged; protarsomere II as long as it is wide; metatarsomere I two times shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw slightly longer than the inferior and equal in width, distance between teeth shorter than the inferior tooth. Abdomen: ventrites with abundant bristles on disc; propygidium slightly visible, with bristles; pygidium more or less convex; subquadrate, as wide as it is long; pygidial width equal in distance or slightly wider than the distance between spiracles of propygidium; pygidial disc with sparse bristles throughout; pygidial apex quadrate. Parameres: basal region narrower than both sections of the parameres at its maximum width, parameral split at 2/3, lyre shaped; ventral surface seen from dorsal view, concave; parameres convex in lateral view (Fig. 2G). Type locality. ARGENTINA, Santiago del Estero. Geographical distribution. ARGENTINA (Jujuy, Salta, Formosa, Tucumán, Santiago del Estero). Remarks. Liogenys cartwrighti is similar to L. denticeps Blanchard, 1851 (Fig. 5) in the body size and color and in the clypeus and pygidium in shape. Liogenys cartwrighti differs from L. denticeps (in parenthesis) by the obtuse (rouded) pronotal posterior corners; pronotum and elytra finely punctate (coarsely); pygidium with bristles throughout (glabrous); males with protarsomeres and mesotarsomeres strongly enlarged (slightly wider than the metatarsomeres); and in the different shape of the parameres (Figs. 2F, G; 5K, L). After examining the specimen at NHMB, from which Frey (1969) had drawn the genitalia of L. denticulata Moser, 1918 (sensu Frey 1969), we realized that it is a misidentified specimen of L. cartwrighti. Frey (1969) mentioned “ Salta, Rivadavia” [Argentina] as the type locality of the complete series, and that the type [holotype] is deposited at the USNM. The paratypes examined at NHMB match with this locality, but the holotype at the USNM bears “ Santiago del Estero ” [Argentina]. As this type specimen bears all the original labels of Frey, we confirm that Santiago del Estero is the type locality of L. cartwrighti, and not Salta. Besides the Chacoan region, L. cartwrighti can be also found at the Yungas Province, in sympatry with L. denticeps and their similar species L. latitarsis Moser, 1918., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 6-8, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240.","Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167."]} more...
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31. Liogenys rugosicollis Frey 1969
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy ,Liogenys rugosicollis - Abstract
Liogenys rugosicollis Frey, 1969 Figs. 20; 24. Liogenys rugosicollis Frey, 1969: 51, 63; Evans 2003: 214 (checklist); Evans & Smith 2009: 182 (checklist); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 5 (generic history). Type material. Liogenys rugosicollis male holotype labeled (USNM): [white handwritten] “ Formosa / P. P. Icomajo [Puerto Pilcomayo] / Argentinien ”, [red typeset] “TYPE”, [white, typeset and handwritten] “ Liogenys / rugosi- / collis n sp / Type / det. G Frey, 1968”, [pink handwritten] “Cartwright / Smithson”, [red, typeset and handwritten] “Type Nº / 74009 / USNM”, genitalia mounted. Paratypes (1) (NHMB): [white handwritten] “ Formosa / P.P. Icomayo [Puerto Pilcomayo] / XII.950—Daguerre”, [orange, round, empty label], [pink typeset] “Property / USNM”, [red typeset] “ PARATYPE ”, [white, typeset and handwritten] “ Liogenys / rugosi / collis / n sp. / det. G. Frey, 1968”, genitalia mounted. Non-type material (2). ARGENTINA. Salta: Chaco Occidental, 100 km NE Los Colorados [20 km NE Cafayate], I.1993, M. McLaqughlin & B. Bestelmeyer, 1 female (UNSM); Santiago del Estero: Ciudad de Santiago del Estero, XII.1939, J. Bosq, 1 female (MZUC). Diagnosis. Length: 11.0–12.0 mm; width: 5.5–6.1 mm. Body and elytra purplish brown, elongate; pronotum sometimes darker than elytra, disc roughly punctate, punctures dense (Fig. 20A); frons somewhat swollen and depressed medially, especially in females; clypeal emargination deep, rounded and narrow, slightly wider in males; outer sides of anterior teeth parallel; clypeal lateral margin with a sharp tooth-like projection, stronger in females, forming an acute angle between this projection and the anterior teeth, angle narrower in females; antennae with 10 antennomeres; pronotal posterior corners sharp, in obtuse angle (Fig. 20B); hypomere, mesepisternum, sides of metaventrite, metacoxae and ventrites beneath the elytra scaly abundantly (Fig. 20C); mesotibia cylindrical in cross section; in males mesotibial and metatibial inner margin strongly concave; in females only on mesotibial inner margin; two transverse carinae present posteriorly, the apical one complete; pygidium flat; subtrapezoidal, apex subrounded; propygidium and pygidial disc densely scaly throughout, except on apex, with long, erect bristles (Fig. 20D); in males, metatibial inner margin strongly concave, carinate on apical portion; protarsomere II elongate, mesotarsi weakly enlarged; parameres strongly concave on inner margin and convergent apically; parameral split at 2/3; apex harpoon-shaped, parameres five times longer than the apex (Fig. 20E); parameres in lateral view convex, not coplanar (Fig. 20F). Type locality. ARGENTINA, Formosa, Puerto Pilcomayo. Geographical distribution. ARGENTINA (Salta, Formosa, Santiago del Estero). Remarks. Liogenys rugosicollis resembles L. densata (Fig. 4) in the body shape, size, and color and in the white-scaled vestiture covering the propygidium and most of the pygidium; and in males, the metatibial inner margin concave, although stronger in L. rugosicollis. It differs from L. densata (in parenthesis) in the lateral margin of clypeus with a sharp tooth-like projection, and acute angle between this projection and the anterior teeth (blunt projection, obtuse angle or angle absent); pronotal disc rugose, very coarsely punctate (finely punctate); mesotibiae and metatibiae shorter; pygidium narrower; in males the metatibial inner margin strongly concave (slightly concave); the inner margin of metatibia carinate only apically (weakly carinate along the entire margin); and mesotarsi slightly more enlarged. The parameres look quite alike except in their curvature in lateral view; the basal region slightly longer and the apex slightly shorter. Frey (1969) mentioned that there is only one primary type of Liogenys rugosicollis, reportedly deposited at the USNM, but we found this paratype at the NHMB., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 45-47, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031"]} more...
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32. Liogenys forcipata Frey 1970
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys forcipata ,Liogenys ,Taxonomy - Abstract
Liogenys forcipata Frey, 1970 Figs. 6; 7B, D, F; 23. Liogenys forcipatus Frey 1970: 282; Krajčík 2012: 144. Liogenys forcipata: Evans 2003: 209 (checklist); Evans & Smith 2009: 177 (checklist); Cherman et al. 2016: 764, 766 (systematics); Cherman et al. 2017: 10, 16, 77–78 (generic redescription). Type material. Liogenys forcipatus male holotype (NHMB): [white typeset] “ ARGENTINA / Pronunciamento / Entre Rios / XII.1962, leg. Pena”, [white handwritten] “Type / Liogenys / forcipatus / n sp / [typeset] det. G. Frey, 1970 ”, [red typeset] “TYPE”. Paratypes (2): one male (CMNC) with the same locality label as the holotype, plus: [white handwritten] “13 / ♂ / Type / Liogenys / forcipatus / n sp / [typeset] det. G. Frey, 1970 ”, [red typeset] “PARATYPE”, [white, outlined black, typeset] “Canadian Museum of / Musée Canadien de la / NATURE / CMNEN 00018923”, [white, outlined black, typeset] “H. & A. HOWDEN / COLLECTION / ex. A. Martinez coll.”. One male (NHMB): [white typeset] “ ARGENTINA / Pronunciamento / Entre Rios / XI.1963, leg. Pena”, [white handwritten] “ Liogenys / forcipatus / n sp / [typeset] det. G. Frey, 1970 ”, [red typeset] “ PARATYPE ”. Non-type material (4). ARGENTINA. Entre Ríos: without locality, I.1942, without collector, 1 male (MZUC); Liebig, XI.1987, without collector, 1 male (UNSM); Ubajay, 48 m, 31º57.326ʹS 58º17.814ʹW, I.2008, J. Corbelli, 1 female (IADIZA); Buenos Aires: Rio de la Plata, [without month].1906, without collector, 1 male (MNHN). Diagnosis. Body elongate, wider on the posterior third; elytra non-uniform brown in color that varies from light to dark brown, basal portion darker, as are the head and pronotum (Figs. 6A, 7B); clypeal emargination deep, rounded and wide; outer sides of anterior teeth subparallel; clypeal lateral margin rounded and produced (Figs. 6B, 7D); basal apophysis of metacoxa not produced (Fig. 6D); in males tarsi semi-opaque and enlarged, mainly the protarsomeres and mesotarsomeres; pygidium flat or slightly convex; trapezoidal; pygidial disc glabrous (Figs. 6E, 7F); parameres setose at the subapical portion; inner margins strongly convergent, narrowed apically and apex bilobed, curved inwards (Fig. 6F). Redescription. Length 11.4–12.0 mm; width: 6.0– 6.6 mm. Brown. Head: distance between eyes twice the width of one eye; clypeal emargination deep, rounded, and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth slightly shorter than the eye; lateral margin convex and produced, sometimes forming a rounded projection; distance between clypeal lateral projection and anterior margin of eye as long as one eye or shorter; distance between clypeal lateral projection and apex of the adjacent anterior tooth equal to the basal width of the anterior tooth; angle between outer side of anterior teeth and clypeal lateral projection approximately 90º; distal maxillary palpomere, maximum width slightly wider than the apex; sensorial region smooth or forming a shallow fovea, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and 1.5 times the length of the funicle in males, slightly longer in females. Thorax: anterior margin of pronotum slightly produced medially; pronotal disc sometimes sulcate medially, glabrous, punctures coarse and sparse, denser near the anterior margin; pronotal posterior corners rounded; hypomere with short bristles and scattered scales, mesepisternum scaly; sides of metaventrite with sparse erect bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival to rounded, sparsely and coarsely punctate. Elytra: shiny, glabrous, brown to dark brown, sometimes apparently burnt; elytral suture darker than elytron and elevated, pair of inner ridges more defined than the three outer pairs. Legs: three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface setose, thick erect bristles on posterior margin; mesotibia cylindrical in cross section, surface coarsely sculptured; two transverse carinae, the apical one complete; metacoxa with sparse bristles near the base of the leg; basal apophysis of metacoxa not produced (Fig. 6D); metatibia carinate along the inner margin, apex not produced, apical inner surface setose; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males tarsi opaque, with tarsomeres I to IV enlarged in every leg; protarsomeres and mesotarsomeres almost twice the width of the metatarsomeres; protarsomere II wide; metatarsomere I shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer than the inferior and equal in width, distance between teeth more than twice the length of the inferior tooth; inferior tooth oblique to the axis of the superior tooth. Abdomen: ventrites with scattered bristles on disc; propygidium slightly visible, glabrous; pygidium flat or slightly convex, subtrapezoidal, wide; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex rounded. Parameres: basal region narrower than both sections of the parameres at its maximum width, parameral split at the third portion, inner margins convergent; apex bilobed and curved inwards, inner margins overlapped; parameres abundantly setose on subapical portion; parameres in lateral view not flattened subapically; straight, coplanar (Fig. 6G). Type locality. ARGENTINA, Entre Ríos, Pronunciamento. Geographical distribution. ARGENTINA (Entre Ríos, Buenos Aires). Remarks. Liogenys forcipata resembles L. tarsalis Moser, 1921 (Fig. 75, Cherman et al. 2017) in the nonuniform color of the elytra and somewhat in the shape of the clypeus, which is strongly emarginate. It differs from L. tarsalis (in parenthesis) in the body size slightly wider on posterior third (uniform width (Fig. 6A)); the clypeal emargination slightly wider, with teeth shorter; the clypeal lateral margin more convex and produced (Figs. 6C, 7D); the pronotum globose (flatter); the metacoxae glabrous (scaly throughout); the pygidium wider (Figs. 6E, 7F); and in males the tarsi strongly enlarged but less than in L. tarsalis, and the metatarsi less enlarged than the protarsi and mesotarsi (tarsi strongly enlarged in all legs). Liogenys forcipata does not occur in the Chaco Province, but in the Espinal boundaring ecorregion (Figs. 23, 24). Nonetheless, we opted to include this species due to its similarity with L. tarsalis, especially the females (Fig. 7B, D, F), and with L. neoforcipata Cherman, new species (Fig. 13), both species present in the Chaco. Liogenys forcipata can be found in sympatry with L. latitarsis, L. obscura Blanchard, 1851, L. pallens Blanchard, 1851, and sometimes with L. cribricollis and L. opacicollis Fairmaire, 1892 in relicts of xerophytic vegetation., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 16-19, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Cherman, M. A., Moron, M. A. & Almeida, L. M. (2016) Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology, 41, 744 - 770. https: // doi. org / 10.1111 / syen. 12188","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312","Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240.","Fairmaire, M. L. (1892) Descriptions de quelques coleopteres Argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253."]} more...
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33. Liogenys foveata Cherman 2021, new species
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys foveata ,Liogenys ,Taxonomy - Abstract
Liogenys foveata Cherman, new species Figs. 8, 27. http://zoobank.org/ DB19CD93-CCCD-4A3F-A80E-74114513BDAC Type material. Holotype labeled (IADIZA): [white typeset] “ ARGENTINA: Formosa / RP 39, 23 km S Ing. Juárez, / 24º05ʹ27ʺS 61º56ʹ49ʺW. / 13-17-XII-08. F. Ocampo G. San Blas, F. Campón. ”, [red, typeset and handwritten] “ LIOGENYS FOVEATA / HOLOTYPE / Cherman M. A.”. Holotype deposited at IADIZA. Diagnosis. Body elongate; brown, elytra brown or light brown, pronotum purplish brown, head dark at vertex (Fig. 8A); clypeal emargination deep but not reaching the base of the teeth, angulate and wide; outer sides of anterior teeth parallel; clypeal lateral margin projection subangulate; outer side of anterior teeth and clypeal lateral projection forming an angle (Fig. 8B); fovea of the distal maxillary palpomere shallow, occupying almost the entire surface of the palpomere (Fig. 8C); pronotal posterior corners subangulate (Fig. 8B); mesofemural surface setose abundantly, long thick erect bristles, as well as the outer margin of the metafemur (Fig. 8D, E); pygidial width exceeding distance between spiracles of propygidium; pygidial disc with long bristles at the apex (Fig. 8F); parameres almost uniform along its length; outer margins slightly convex at the midline; apex spatulate (Fig. 8G). Holotype. Male. Length: 10.5 mm; width: 5.6 mm. Brown. Head: distance between eyes almost twice the width of one eye; clypeal emargination deep but not reaching the base of the teeth, angulate and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth almost equal to the eye length; lateral margin slightly convex, forming a subangulate projection; distance between clypeal lateral projection and anterior margin of eye shorter than the eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth longer than the basal width of the anterior tooth; outer side of anterior teeth and clypeal lateral projection not forming an angle; distal maxillary palpomere, maximum width more than twice the apical width; fovea shallow, extending past the transverse midline, occupying almost the entire surface of the palpomere; antenna with 10 antennomeres, club lighter in color and slightly longer of the funicle. Thorax: anterior margin of pronotum straight; pronotal disc glabrous, punctures coarse and sparse; posterior corners subangulate; hypomere with short bristles, mesepisternum scaly; sides of metaventrite with bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, sparsely and finely punctate throughout. Elytra: shiny, glabrous, brown, lighter in color than pronotum; elytral suture darker than elytron and weakly elevated; all four elytral ridges weakly defined. Legs: three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface setose abundantly, as well as the outer margin of the metafemur, bristles long, thick, erect; mesotibia cylindrical in cross section, surface finely sculptured; two transverse carinae, the apical one present posteriorly; metacoxa with sparse, long scales; basal apophysis of metacoxa not produced; metatibia carinate along the inner margin towards the apex, apical inner surface setose; a metatibial transverse carina present posteriorly and posterior discontinuous longitudinal carina (Fig. 8E); metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV enlarged; protarsomere II slightly longer than it is wide; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and slightly narrower than the inferior, distance between teeth equal to the length of the inferior tooth. Abdomen: ventrites with sparse, long bristles, abundantly on ventrite V; propygidium visible, with bristles; pygidium slightly convex, subquadrate, as wide as it is long; pygidial width exceeding distance between spiracles of propygidium; pygidial disc with long bristles at the apex; pygidial apex rounded. Parameres: basal region as wide as both sections of the parameres at its maximum width, parameral split at 2/3; inner margins straight, opened; outer margins slightly convex at the midline; apex spatulate; parameres slightly concave in lateral view (Fig. 8H). Etymology. Adjective in the nominative singular. From the Latin foveatus (fovea “ditch, pit”): having a slight depression or pit in a bone or organ. In this species the fovea of the maxillary distal palpomere is shallow but quite broadly extended, almost reaching the apex. Type locality. ARGENTINA, Formosa, Ruta Provincial 39, 23 km S Ingeniero Juárez, 24º05’27’’S 61º56’49’’W. Geographical distribution. ARGENTINA (Formosa). Remarks. Liogenys foveata resembles L. tarsalis, mainly in the size, body color, and clypeus deeply emarginate. Liogenys foveata differs from L. tarsalis (in parenthesis) mainly in the clypeus with emargination angulate and lateral margin with a weak subangulate projection (emargination subangulate, lateral margin with a distinct, rounded projection); in the large fovea on the maxillary distal palpomere, occupying almost the entire surface of the palpomere; in the pronotal posterior corners subangulate (rounded); and in the shape of the pygidium, tarsi and parameres. Females remain unknown., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 19-21, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258 more...
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34. Liogenys vicina Frey 1969
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys vicina ,Taxonomy - Abstract
Liogenys vicina Frey, 1969 Figs. 22; 26. Liogenys vicinus Frey, 1969: 62, 51 (key); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 5 (generic history). Liogenys vicina: Evans 2003: 215 (checklist); Evans & Smith 2009: 183 (checklist); Cherman et al. 2017: 20 (natural history). Type material. Liogenys vicinus male holotype (USNM): [white handwritten] “Formosa / Clorinda / 8.XI.950. Daguerre”, [red, typeset and handwritten] “Type No / 74010 / USNM”, [red typeset] “TYPUS”, [white, typeset and handwritten] “ Liogenys / vicinus / n.sp. / Type / det. G Frey, 1968”, [pink typeset] “Property / USNM”, [white typeset] “Loan from / USNMNH / 2065060”, genitalia mounted. Paratypes (2): One female (NHMB) [white handwritten] “ Formosa / P. Pilcomayo ”, [white typeset] “ ♀ ”, [red typeset] “ PARATYPE ”, [white, typeset and handwritten] “ Liogenys / vicinus / n.sp. / det. G. Frey, 1968”. One male (NHMB) [white typeset] “ Argentina / Formosa / Cindad XII.1949 / A. Martinez leg.”, [red typeset] “ PARATYPE ”, [white, typeset and handwritten] “ Liogenys / vicinus / n.sp. / det. G. Frey, 1968”, genitalia mounted. Non-type material (9). ARGENTINA. Formosa: Laguna Yema: Centro de Validación-(CEDEVA), barrio CEDEVA, 24º16’03’’S 61º14’54’’W, 163 m, 14.XII.2008, F.C. Ocampo, G. San Blas, & F. Campon, 1 male (IADIZA); Reserva de Biósfera Teuquito, Ruta Provincial 37, 1 km S “Canal”, 24º21’20,95’’S 61º18’55,23’’W, 11.XII.2008, 1 male and 1 female (IADIZA); Chaco: Fuerte Esperanza, XI.1978, A. Martínez & Fritz, 2 males and 2 females (CMNC); Santiago del Estero: without locality, II.1960, without collector, 1 male and 1 female (IADIZA). Diagnosis. Length 10.8–11.9 mm; width: 4.8–5.9 mm. Body elongate, brown to dark brown, shiny; coarsely punctate (Fig. 22A); clypeal emargination deep but not reaching the base of the teeth, rounded and narrowed; clypeal lateral margin strongly convex and produced, forming a tooth-like projection; pronotal posterior corners subangulate to rounded (Fig. 22B); antennae with 10 antennomeres; first two pair of elytral ridges equally defined and more noticeable than the other two; basal apophysis of metacoxa produced; metafemur with bristles, sometimes abundantly; metatibia not carinate along the inner margin; pygidium convex, wider than it is long; pygidial width wider than the distance between spiracles of propygidium, disc with bristles throughout, erect bristles (Fig. 22D); in males protarsomeres and mesotarsomeres strongly enlarged, metatarsomeres slightly enlarged; parameres strongly expanded at the midline and abruptly narrowed at the apex (Fig. 22E, F); parameres strongly convex in lateral view (Fig. 22G). Geographical distribution. ARGENTINA (Formosa, Chaco, Santiago del Estero). Remarks. Liogenys vicina resembles the sympatric L. latitarsis (Fig. 10), and these similarities difficult their identification. Liogenys vicina differs from L. latitarsis (in parenthesis) in the punctures on pronotum and elytra coarser; clypeal emargination slightly narrower, lateral margins of pronotum slightly rounder, elytral inner ridges evenly elevated (sometimes the second ridge less elevated), metafemur more with bristles; and in the shape of parameres inflated basally and abruptly narrowed at the apex (not inflated basally and apical constriction absent)., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on page 50, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)"]} more...
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- 2021
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35. Liogenys opacicollis Fairmaire 1892
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys opacicollis ,Liogenys ,Taxonomy - Abstract
Liogenys opacicollis Fairmaire, 1892 Figs. 16; 23. Liogenys opacicollis Fairmaire, 1892: 243; Dalla Torre 1913: 318 (checklist); Moser 1921: 140 (L. densicollis original description); Blackwelder 1944: 227 (checklist); Frey 1969: 39, 53 (key, systematics); Krajčík 2012: 145 (checklist); Evans 2003: 212 (checklist); Evans & Smith 2009: 180 (checklist). Liogenys densicollis Moser, 1921: 140; Blackwelder 1944: 227 (checklist); Frey 1969: 38–39 (key); Krajčík 2012: 144 (checklist); Evans 2003: 208 (checklist); Evans & smith 2009: 176 (checklist). New synonym. Type material. Liogenys opacicollis female lectotype here designated (MNHN): [white typeset] “MUSÉUM PARIS / 1906 / Coll. Léon FAIRMAIRE”, [white handwritten] “ Liogenys / opacicollis / Fairm / Rioja / Planta ”, [red, outlined black, handwritten] “ LIOGENYS / OPACICOLLIS / FAIRMAIRE ♀ / [typeset] LECTOTYPE / des. M. A. Cherman ”. Liogenys densicollis male lectotype here desigated (ZMHB): [white, outlined black, typeset] “ Argentinien / Santiago del Estero ”, [white handwritten] “ Liogenys / densicollis / Mos / Typen ♂”, [light red typeset] “Typus”, [white typeset] “ Liogenys / densicollis / Mos.”, [red typeset] “ SYNTYPUS / Liogenys / denticulata Moser, 1918 [misspelled] / labelled by MNHUB 2012”, [white, outlined red, typeset] “ LECTOTYPE / Liogenys densicollis / Moser, 1921 / Cherman M. A. 2013”, genitalia mounted. Paralectotypes (10) (ZMHB): one male with the labels [white typeset] “ Argentinien / Santiago del Estero ”, [white handwritten] “ Liogenys / densicollis / Mos / Typen ♂”, [light red typeset] “Typus”, [white typeset] “ Liogenys / densicollis / Mos.”, [red typeset] “ SYNTYPUS / Liogenys / densicollis Moser, 1921 / labelled by MNHUB 2012”, [white, outlined red, typeset] “ PARALECTOTYPE / Liogenys densicollis / Moser, 1921 / Cherman M. A.” Nine female specimens with the same labels, but instead of: [white handwritten] “ Liogenys / densicollis / Mos / Typen ♂”, the label is: [white handwritten] “ Liogenys / densicollis / Mos / Typen ♀ ”. Non-type material (81). BOLIVIA. Santa Cruz: Caballero, ~ 9 km SW Comarapa, XII.2008, 1694 m, 17º58.926’S 64º34.365’W, W.D. Edmonds & T. Vidaurre, thorn forest, human feces, 2 males (CEMT); Samaipata. Bolivia Andina, 25.XI.1959, Zischka, 1 female (NHMB). ARGENTINA. Salta: Las Tablillas, without date, G.L. Harrington, 1 female (CASC); Viñaco, 15 km S El Carril, 12.II.1982, H. & A. Howden, 2 females (CMNC); 1 km N Cafayate, 26º03’52’’S 65º56’19’’W; 24.XI.2006, F. Ocampo, E. Ruiz, G. San Blas, & G. Salazar, ultraviolet & mercury vapor light trap, 8 males and 2 females (IADIZA); El Naranjo, Rosario de la Frontera, I.1944, Duret-Martínez, 1 male (CMNC); Departamento Anta. Las Lajitas, XII.1982, A. Martínez, 1 male (CMNC); Olleros [Chañar Muyo], II.1958, Daguerre, 2 males and 2 females (USNM); Rivadavia, XII.1950, Daguerre, 3 females (USNM); [without month].1905, Steinbach, 1 male (ZMHB); Formosa: without locality and date, Mallo, 1 female (MLPA); Pilcomayo, XII.1950, Daguerre, 2 males and 2 females (USNM); Chaco: Gancedo, 8.XII.1939, Biraben & Bezzi, 1 female (MLPA); Santiago del Estero: Río Salado, without date, E.R. Wagner, 2 females (CMNC); La Palisa del Bracho, 25 km NNW d’Icaño, Laguna Mamaita, [without month].1904, E.R. Wagner, 1 male (CASC); banks of Río Salado, D’Icaño environs, Mistol Paso, [without month].1909, E.R. Wagner, 3 females (CASC); El Pinto, XI.1960, without collector, 1 male and 2 females (NHMB), 1 female (CNCI); Catamarca: without locality, C. Bruch, 1.II.1907, 1 male (MLPA); without locality, C. Bruch, 4.II.1910, 1 female (MLPA); without locality, date, and collector, 1 male and 1 female (MLPA); Andalgala km 1508 [5 km W, along Ruta Nacional 9, 1000 m, in Larrea cuneifolia bushes (Zigophyllaceae) (Schultz 1981)], 12.XI.1973, J.C. Schultz, 2 males (CASC); La Rioja: Sañogasta (Camping), 7.XII.2013, Arriagada & Castillo, light trap, 2 males and 1 female (CMVD); 6 km E Chepes, without date and collector, 1 female (CMNC); Santa Fé: Carcarañá, without date and collector, 1 male (USNM); Córdoba: without locality and date, Davis, 1 male; without locality, date, and collector, 1 female (AMNH); Mediolanza, 24.XII.2004, E. Ruiz, 1 male (IADIZA); San Vicente, without date, Frenzel, 2 males and 1 female (ZMHB); Potrero de Garay, II.1948, A. Martínez, 2 males (CMNC); Diquecito, without date, Frenzel, 1 female (NHMB); San Luis: Quines, 24.XI.1995, S. Roig & G. Flores, 1 male (IADIZA); El Molle, 7.II.2017, 33º02’22.8’’S 66º30’47.6’’W, 622 m, light, G. Arriagada, 1 male and 1 female (CEMT); Sa Jerónimo, I.1979, Williner, 2 females (CMNC); Mendoza: without locality and date, C. Bruch, 1 female (MLPA); Buenos Aires: Felipe Solá. Puán, 17.II.1944, A. Martínez, 2 males (CMNC); Neuquén: San Martín de los Andes, II.1956, A. Martínez, 1 male (MZSP); Río Negro: Gral. Conesa, I.1931, without collector, 1 male (CMNC); Coronel Gómez, XI.1945, J. Grosso, 1 male (CMNC); Departament Gral Roca, Catriel, 4.I.2005, F.C. Ocampo, 2 females (CMNC); San Antonio Oeste, I.1976, M. Zunino, 1 male (CMNC); Valcheta, XII.1992, at lights, without collector, 4 males, 1 female (CMNC). Diagnosis. Dark purplish brown. Head and pronotum densely punctate, cribrose-like surface; elytra uniformly with bristles, vertex scaly; clypeal emargination forming a semicircle, clypeal lateral margins convex; pronotal posterior corners in obtuse angle (Fig. 16 A–B); in males inner margin of male metatibia carinate towards the apex, subapically slightly concave (Fig. 16F); apical inner margin with tooth-like projection contiguous to the smaller spur (Fig. 16H); metatibial transverse carina complete; mesotarsomere I ventrally produced, proximal margin sometimes prickled (Fig. 16 G–H); abdominal ventrites, propygidium, and pygidium setose, with bristles and scales throughout; pygidial apex subquadrate to rounded (Fig. 16J); parameres with apex spatulate shaped in a single plane, rounded, or spiked laterally, apical length from 1/3 to 1/4 the length of the parameres after the basal split (Fig. 16K). Redescription. Length: 9.8–12.1 mm; width: 4.4–6.0 mm. Dark purplish brown. Head. Densely punctate, cribrose-like surface; vertex scaly; distance between eyes twice the width of one eye in males, three times in females; vertex scaly, sometimes with bristles on frontoclypeal surface and inner margins of eye; clypeal emargination deep, wide and rounded, forming a semicircle; outer sides of anterior teeth follow the lateral margin of clypeus, clypeal lateral margin convex; distal maxillary palpomere, maximum width more than twice the apical width; fovea deep, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than funicle. Thorax: anterior margin of pronotum slightly produced medially; sides of pronotum with sparse, long bristles, pronotal disc coarsely punctate, punctures dense, disposed tightly; pronotal posterior corners in obtuse angle; hypomere with long bristles and scaly; mesepisternum, sides of metaventrite and metacoxae scaly abundantly, scales dense; distance between mesocoxae and metacoxae less than twice longer than the metacoxa; scutellum rounded, coarsely punctate throughout. Elytra: shiny, erect bristles disposed uniformly; elytral suture and elytron unicolored, weakly elevated; all four elytral ridges weakly defined. Legs: procoxa scaly and punctate; three protibial teeth, middle and apical equal in size, distance between basal and middle teeth equal to or slightly longer than between middle and apical; mesofemural surface setose; mesotibia subquadrate to cylindrical in cross section, mesotibial surface coarsely sculptured; apical mesotibial transverse carina strong and complete, the basal one weakly defined or absent; metafemur setose throughout; inner margin of male metatibia carinate towards the apex, slightly concave subapically (Fig. 16F); apex with tooth-like projection below the smaller spur (Fig. 16I); metatibial transverse carina complete in males and females; surface coarsely sculptured; metatibial apical spurs of different lengths, the larger as long as the diameter of the tibial apex; metatarsomere I shorter than metatarsomere II and wider; in males protarsomeres I to IV enlarged; protarsomere II elongate, piriform; mesotarsomeres not enlarged; mesotarsomere I in males ventrally produced, proximal margin sometimes prickled (Fig. 16 G–H); claw bifid, symmetrical, superior tooth of a claw longer and slightly narrower than the inferior; distance between teeth equal to the length of the inferior tooth. Abdomen: band of abundant thick scales disposed tightly beneath the outer margin of elytra; disc of ventrites setose, with bristles and scales throughout in males, sides scaly and with bristles at the midline in females, as well as the propygidium; pygidium flat or slightly convex; subquadrate; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc setose, erect bristles throughout and scales mainly from the base up to the midline; pygidial apex subquadrate to rounded. Parameres: basal region equal in width to both sections of the parameres at its maximum width, parameral split at 2/3; inner margins straight or sometimes convex; apex spatulate-like in a single plane, rounded or angulate laterally, apical length from 1/3 to 1/4 times the length of the parameres after the basal split; parameres in lateral view straight, not coplanar (Fig. 16L). Type locality. Liogenys opacicollis: ARGENTINA, La Rioja; L. densicollis: ARGENTINA, Santiago del Estero. Geographical distribution. BOLIVIA (Santa Cruz); ARGENTINA (Salta, Formosa, Santiago del Estero, Chaco, Catamarca, La Rioja, Santa Fé, Córdoba, San Luis, Mendoza, Buenos Aires, Neuquén, Río Negro). Remarks. Frey (1969) mentioned that L. opacicollis and L. densicollis could be the same species; but did not propose the synonymy because he could not find the type of L. opacicollis. After examining the type material of both species we determined that the male lectotype of L. densicollis (Fig. 16B, D) matches with the female lectotype of L. opacicollis (Fig. 16A, C), and this is identical to the female paralectotype of L. densicollis. Here, we propose the synonymy of both names. There is a population from Catamarca (Argentina) that has some noteable differences from the type specimens, as the clypeal teeth more rounded, pronotal punctures finer and in males the apex of parameres shorter and more angulate. We conclude that these are intraspecific variations, although we recommend further studies on our concept of this species. The specimens from Catamarca were collected on “Jarilla”, Larrea cuneifolia Cavanilles, 1800 (Zygophyllaceae), an evergreen xerophytic shrub common in western South America (Mercado et al. 2018). The confirmed association of L. opacicollis with xerophytic vegetation, typical from Arid Chaco and Monte ecoregions, is shared with L. cribricollis, with the latter more associated with Schinus shrubs., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 36-39, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Fairmaire, M. L. (1892) Descriptions de quelques coleopteres Argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253.","Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: Schenkling, W. (Ed.), Coleopterorum catalogus, Volumen XX. W. Junk, Berlin, Germany, pp. 291 - 450.","Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167.","Schultz, J. C. (1981) Adaptive changes in antipredator behavior of a grasshopper during development. Evolution, 35, 175 - 179. https: // doi. org / 10.2307 / 2407951","Mercado, M. I., Moreno, M. A., Ruiz, A. I., Rodriguez, I. F., Zampini, I. C., Isla, M. I., Ponessa, G. I., Mercado, M. I., Moreno, M. A., Ruiz, A. I., Rodriguez, I. F., Zampini, I. C., Isla, M. I. & Ponessa, G. I. (2018) Morphoanatomical and histochemical characterization of Larrea species from northwestern of Argentina. Revista Brasileira de Farmacognosia, 28, 393 - 401. https: // doi. org / 10.1016 / j. bjp. 2018.05.012"]} more...
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36. Liogenys mendozana Moser 1918
- Author
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Liogenys mendozana ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys mendozana Moser, 1918 Figs. 12; 26. Liogenys mendozanus Moser, 1918: 102; Frey 1969: 38, 41 (key); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 4 (generic history). Liogenys mendozana: Blackwelder 1944: 227 (checklist). Liogenys mendozanus mendozanus: Frey 1969: 52. Liogenys mendozana mendozana: Evans 2003: 210 (checklist); Evans & Smith 2009: 179 (checklist). Liogenys mendozanus incisus Frey, 1969: 52; Krajčík 2012: 145 (checklist). New synonym Liogenys mendozana incisa: Evans 2003: 210 (checklist); Evans & Smith 2009: 179 (checklist). Type material. Liogenys mendozanus male lectotype here designated (ZMHB): [white, outlined black, typeset and handwritten] “ Argentinien / Mendoza ”, [white handwritten] “ Liogenys / mendozanus / Mos / Typen ♂”, [light red typeset] “Typus”, [white, outlined red, typeset and handwritten] “ LIOGENYS / MENDOZANA / Moser, 1918 / LECTOTYPE / Des. Cherman M. A.”, genitalia mounted. Paralectotypes (1) (ZMHB) with the same data of the lectotype, plus the label: [white, outlined red, typeset and handwritten] “ LIOGENYS / MENDOZANA / Moser, 1918 / PARALECTOTYPE / Des. Cherman M. A.”. Liogenys mendozanus incisus female neotype here designated (MZSP): [white typeset] “ Argentina / Salta / XII.1959 / Aguaray / A. Martinez leg.” [red, typeset and handwritten] “ LIOGENYS / MENDOZANUS / INCISUS / NEOTYPE / [Des.] Cherman M. A.”. Non-type material (14). BOLIVIA. Santa Cruz: Río Pailon, 320 m, 14.X.2001, P. Schmith, 1 female (CMVD). ARGENTINA. Salta: Urundel, 10.X.1968, L. Peña, 1 female (NHMB); 1 km N Cafayate, 24.XI.2006, 26º03’52’’S 65º56’19’’W, F. Ocampo, E. Ruiz, G. SanBlas, & G. Zalazar, 2 females (IADIZA); Tucumán: without locality, date and collector, 2 males and 3 females (MLPA); Santiago del Estero: without locality and date, Wagner, 1 male (MLPA); Córdoba: San Vicente, Frenzel, 1 female (ZMHB); Mendoza: without locality, 1957, H. Kulzer, 1 male (ZMHB); Estación Pedregal, without date, Jensen-Haarup, 1 female (ZMUC); Santa Rosa, Reserva de Ñancuñan, 11–13.XII.2002; V.M. Diéguez, 1 female (CMVD). Diagnosis. Body elongate, dark brown, shiny (Fig. 12 A–B); frons swollen; clypeal emargination shallow, rounded and narrow; outer sides of anterior teeth follow the lateral margin of clypeus (Fig. 12E); antenna with 9 antennomeres; elytra coarsely punctate, elytral suture unicolored with the elytron, slightly or not elevated; three protibial teeth equal in size; distance between basal and middle teeth longer than between middle and apical; inferior tooth of a claw very short and perpendicular to the axis of the superior tooth; male metatibia strongly expanded subapically towards the apex (Fig. 12F), protarsomeres and mesotarsomeres I to IV slightly enlarged; parameres equal in width along their total length; apex strongly truncate (Fig. 12H). Redescription. Length 10.7–12.4 mm; width: 5.8–6.5 mm. Dark brown. Head: distance between eyes more than twice the width of one eye in males, three times in females; frons swollen; clypeal emargination shallow, rounded and narrow; outer sides of anterior teeth follow the lateral margin of clypeus; lateral margin straight or slightly convex, not produced; distal maxillary palpomere, maximum width slightly wider than the apex; fovea deep shallow, elongate, extending past the transverse midline of the palpomere; antenna with 9 antennomeres, club lighter in color and equal to the funicle. Thorax: anterior margin of pronotum straight; glabrous, punctures coarse and sparse; pronotal posterior corners rounded to subangulate; hypomere with short bristles; mesepisternum and sides of metaventrite with bristles; distance between mesocoxae and metacoxae twice the length of the metacoxa; scutellum ogival, sparsely and randomly punctate. Elytra: shiny, glabrous, dark brown; elytral suture slightly unicolored with the elytron, not elevated; all four elytral ridges weakly defined, sometimes inner pair of ridges more defined than the others. Legs: three protibial teeth equal in size; distance between basal and middle teeth longer than between middle and apical; mesotibia cylindrical in cross section, surface finely sculptured; two transverse carinae, the apical one complete; metacoxa sparsely with bristles, thick bristles; basal apophysis of metacoxa not produced; two metatibial transverse carinae present in females, absent or weakly marked in males; male metatibia carinate along the inner margin; abruptly produced and strongly expanded subapically towards apex, apical inner surface setose abundantly; metatibial apical spurs of different lengths; the larger longer than the diameter of the tibial apex; protarsomeres and mesotarsomeres I to IV slightly enlarged, protarsomere long; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and equal in width to the inferior, distance between teeth more than twice the length of the inferior tooth; inferior tooth perpendicular to the axis of the superior tooth. Abdomen: ventrites scarcely with bristles on disc; propygidium glabrous; pygidium flat; subtrapezoidal, as wide as it is long (Fig. 12G), narrower in females (Fig. 12H); pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex quadrate. Parameres: basal region equal in width to both sections of the parameres at its maximum width, parameral split at third portion, inner margins straight and close together; apically slightly curved downwards; apex strongly truncate, forming in a frontal view rounded structure, sculptured, spiked at the inferior margin; parameres slightly convex in lateral view (Fig. 12J). Type locality. Liogenys mendozana: ARGENTINA, Mendoza; L. mendozana incisa: ARGENTINA, Salta. Geographical distribution. BOLIVIA (Santa Cruz); ARGENTINA (Salta, Tucumán, Santiago del Estero, Córdoba, Mendoza). Remarks. Among the specimens of L. mendozana examined, we found a range of variation in the coarseness of the punctures on pronotum and elytral disc, which has been previously mentioned by Frey (1969). Frey (1969) used this variation to distinguish two subspecies, while all the other morphological features are identical.After examining the material we understand that this is only an intraspecific variation, so here we propose L. mendozana incisa as the junior synonym of L. mendozana. The type series Frey (1969) used to describe L. mendozana incisa, which comprises five specimens with males and females, was not found at USNM or NHMB, even though both depositories were stated by the author in the original description. We found one specimen from Salta (MZSP) that was identified and labeled by Frey as L. mendozana incisa, and matches with the description and type locality mentioned by the author [Article 76.1, International Code of Zoological Nomenclature (1999)] (Frey 1969). Due to the purpose of clarifying the taxonomic status by fixing this name to a specimen, according to the Article 75.3 of the International Code of Zoological Nomenclature (1999), this specimen was selected as the neotype (Fig. 12B, D). Liogenys mendozana is distributed along the eastern Andes, from Bolivia (new country record presented here) through Mendoza (Argentina), at the Monte Province where xerophytic vegetation predominates. It can be found in sympatry with L. cribricollis, L. densata, L. obscura, L. opacicollis, and L. pallens., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 27-29, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp."]} more...
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37. Liogenys cribricollis Moser 1921
- Author
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys cribricollis ,Taxonomy - Abstract
Liogenys cribricollis Moser, 1921 Figs. 3; 23. Liogenys cribricollis Moser, 1921:140; Blackwelder 1944: 227 (checklist); Biezanko et al. 1949: 192 (natural history); United States Department of Agriculture 1971: 335 (natural history); Evans 2003: 208 (synonym of L. densicollis). Liogenys densicollis var. cribricollis: Frey, 1969: 38–39, (key, systematics). Type material. Liogenys cribricollis male lectotype here designated (ZMHB): [white typeset] “Republ. Argentina / CHACO DE SANTIAGO / DEL ESTERO, RIO SALADO”, [white handwritten] “ Liogenys / cribricollis / Mos / Typen ♂”, [light red typeset] “Typus”, [white typeset] “ Liogenys / cribricollis / Mos.”, [white, outlined red, typeset and handwritten] “ Liogenys / cribricollis / Moser, 1921 / LECTOTYPE / Des. Cherman M. A.”, genitalia mounted. Paralectotypes (1): one female with the same data of the lectotype, plus the label: [white, outlined red, typeset and handwritten] “ Liogenys / cribricollis / Moser, 1921 / PARALECTOTYPE / Des. Cherman M. A.” (ZMHB). Non-type material (24). ARGENTINA. Salta: without locality, I.1956, Vaqueros, 1 male and 2 females (CNCI); Santiago del Estero: without locality, date and collector, 1 male (ZMHB); without locality and date, Wagner, 1 male (MLPA); Río Salado, without date, E. Wagner, 1 male and 1 female (MLPA); 1 female (CASC); Chaco: Gancedo, 8.XII.1939, Biraben & Bezzi, 3 males (MLPA); Santa Fé: Carcarañá, without date and collector, 1 female (USNM); Córdoba: Marull, 22.I.1940, Biraben, 1 female (MLPA), Diquecito, XI.1958, A. Martínez, 1 male (MZSP); Alta Gracia, “Las Rosas”, I.1941, Monrós, 1 female (CMNC); without locality, XI.1968, Arguello, 2 females (CNCI); San Luis: 18 km S Arizona, 18–23.I. 1982, 250 m, H. & A. Howden, 1 female (CMNC), Río Negro: without locality, date and collector, 1 male and 2 females; Conesa, I.1931, J. Bosq, 1 female (MLPA); Villa Regina, XII.1961, A. Maller, 1 male and 1 female (DZUP). Diagnosis. Dark purplish brown. Head and pronotum densely punctate, cribrose-like surface; elytra uniformly with bristles, clypeal emargination rounded; anterior teeth rounded; lateral margin convex and produced roundly; angle between the outer side of anterior teeth and clypeal lateral projection approximately 90º; posterior corners rounded to subangulate (Fig. 3 A–B); inner margin of male metatibia carinate towards the apex, subapically convex, sometimes produced (Fig. 3F); apical inner margin with or without tooth-like projection contiguous to the smaller spur (Fig. 3J); two metatibial transverse carinae present only posteriorly in males, apical carina complete in females; in males protarsomere II elongate, rectangular and mesotarsomere I forming a hook-like projection towards the base (Fig. 3G); abdominal ventrites at the midline with abundant bristles and sometimes with scales in males (Fig. 3E), with sparse bristles in females; apex of parameres spatulate-like shaped, expanded laterally at the lower plane and more or less angulate (Fig. 3K). Redescription. Length: 9.7–13.0 mm; width: 5.0– 6.5 mm. Dark purplish brown. Head. Densely punctate, cribrose-like surface; distance between eyes from two times to three times as wide as one eye; vertex scaly, sometimes also the inner margins of the eye, clypeal margin sometimes with bristles; clypeal emargination deep, wide and rounded, anterior teeth rounded; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; lateral margin convex and more (Fig. 3B) or less (Fig. 3C) produced roundly; distance between clypeal lateral projection and the adjacent anterior tooth longer than the basal width of the anterior tooth, distance between clypeal lateral projection and anterior margin of eye longer than one eye, angle between outer side of anterior teeth and clypeal lateral projection approximately 90º; distal maxillary palpomere, maximum width less than twice the apical width; fovea deep, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than the funicle. Thorax: anterior margin of pronotum slightly produced medially; sides of pronotum with sparse, long bristles; pronotal disc coarsely punctate, punctures disposed tightly; posterior corners rounded to subangulate; hypomere with long bristles and scaly; mesepisternum, sides of metaventrite and metacoxae scaly abundantly, scales disposed tightly; distance between mesocoxae and metacoxae two times longer than the metacoxa; scutellum rounded, coarsely punctate. Elytra: shiny, with erect bristles disposed uniformly; elytral suture and elytron unicolored, weakly elevated; all four elytral ridges weakly defined. Legs: procoxa scaly and punctate; three protibial teeth, middle and apical equal in size, distance between basal and middle teeth equal to or longer than between middle and apical; mesofemural surface setose; mesotibia subquadrate to cylindrical in cross section, mesotibial surface coarsely sculptured; two transverse carinae, the apical carina complete; metafemur setose throughout; inner margin of male metatibia carinate towards the apex, subapically convex, sometimes produced; apical inner margin with or without tooth-like projection contiguous to the smaller spur; surface coarsely sculptured; two metatibial transverse carinae present only posteriorly in males, apical carina complete in females; metatibial apical spurs of different lengths, the larger shorter than the diameter of the tibial apex; metatarsomere I shorter than metatarsomere II and wider; in males protarsomeres I to IV enlarged, protarsomere II elongate, rectangular; mesotarsomeres not enlarged; mesotarsomere I in males ventrally produced, proximal margin spinose forming a hook-like projection towards the base; claw bifid, symmetrical, superior tooth of a claw slightly longer and narrower than the inferior; distance between teeth shorter than the inferior tooth. Abdomen: band of abundant thick scales disposed tightly beneath the outer margin of elytra; disc of ventrites scaly at the sides and with bristles at the midline, with abundant bristles and sometimes with scales at the midline in males, with sparse bristles in females; propygidium scaly tightly; pygidium slightly convex or flat; quadrate or subquadrate; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc setose throughout, with scales and erect bristles; pygidial apex quadrate in males (Fig. 3H), rounded in females (Fig. 3I). Parameres: basal region slightly wider than both sections of the parameres at its maximum width, parameral split at 2/3; inner margins convex; apex spatulate, expanded laterally at the lower plane and more-or-less angulate; paramere length after the split twice as long as the basal portion or even longer; in the first case the apex is short and more angulate laterally; parameres in lateral view straight, not coplanar (Fig. 3L). Type locality. ARGENTINA, “ Chaco de Santiago del Estero ” [Santiago del Estero Province]. Geographical distribution. BRAZIL (Rio Grande do Sul) (Biezanko et al. 1949), ARGENTINA (Salta, Santiago del Estero, Chaco, Santa Fé, Córdoba, Río Negro). Remarks. Liogenys cribricollis resembles L. opacicollis Fairmaire, 1892 (Fig. 16) mainly in the body color; in the head and pronotum densely punctate, forming a cribrose-like surface; the elytra uniformly bristled; the metatarsomere I short and wide; and in males the mesotarsomere I has a ventral hook-like projection (Fig. 3G). This species was considered by Frey (1969) as a variety of L. densicollis. Evans (2003) proposed this name as a synonym of L. densicollis. We found many differences between the primary types of both L. densicollis (in parenthesis) and L. cribricollis: in L. cribricollis the clypeal anterior teeth are well noticeable, the clypeal lateral margin convex and roundly produced, forming an angle between the outer side of anterior teeth and the clypeal lateral projection approximately 90º (outer margins of teeth obsolete, undistinguished from the lateral margin of clypeus, which is not produced); pronotal posterior corners rounder (obtuse angle); mesotibia with basal transverse carina more conspicuous (less distinct); in females the abdominal disc with sparse bristles (bristles more abundant at the midline of the ventral disc, sides scaly); and in males, the protarsomere II is rectangular (piriform); the mesotarsomere I forms a strong hook-like projection towards its base, almost reaching it (projection sharp but less wide and long); on metatibia the inner margin is subapically convex, and the transverse carina is present only posteriorly (metatibia uniform in width, transverse carina complete); and the apex of the parameres expanded laterally in a lower plane (in a single plane). Due to all these differences, herein we revalidate the status of Liogenys cribricollis as a valid species. As it happens among populations of L. opacicollis, populations of L. cribricollis, mainly those from Córdoba and Rio Negro (Argentina), show differences in the shape of the clypeal margin, with emargination wider and teeth weaker; and with the apex of the parameres shorter and sharper laterally. We interpret those specimens to belong to L. cribricollis because their morphological variation is not discrete enough to consider them as a different species. Adults of L. cribricollis were reported damaging corn in Pelotas, Rio Grande do Sul, Brazil (Biezanko et al. 1949). Here we want to state that in Biezanko et al. (1949) it was written “molho”, and Cherman et al. (2017) misspelled it with “milho”, which is corn in Portuguese. Biezanko et al. (1949) were referring to Schinus dependens Ortega “molho, assobiadeira” (Anacardiaceae), currently a junior synonym of the Peppertree, Schinus polygamus (Cav.) Cabrera. The United States Department of Agriculture (1971) reported L. cribricollis attacking “Molle”, Schinus latifolius (Gillies ex Lindl.) (Anacardiaceae) in southern Brazil. These Schinus species are shrubs to tall trees native to temperate South America, although the latter is more commonly spread in Argentina and Chile (Barroso et al. 1991). Despite Biezanko et al. (1949) and United States Department of Agriculture (1971) reports, we did not find specimens from Brazil among the studied material in any collection, so we recommend further studies reviewing the Brazilian geographical distribution for this species., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 8-11, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Biezanko, C. M., Bertholdi, R. E. & Baucke, O. (1949) Relac \" o dos principais insetos prejudiciais observados nos arredores de Pelotas nas plantas cultivadas e selvagens. Agros, 2, 156 - 213.","United States Department of Agriculture (1971) Insects not known to occur in the United States: a scarab (Liogenys macropelma Bates). Cooperative Economic Insect Report, 21, 335 - 336.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Fairmaire, M. L. (1892) Descriptions de quelques coleopteres Argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Barroso, J. G., Pedro, L. G., Pais, M. S. S. & Scheffer, J. J. C. (1991) Composition of the essential oil of Schinus latifolius Engl. Flavour and Fragrance Journal, 6, 237 - 239. https: // doi. org / 10.1002 / ffj. 2730060315"]} more...
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38. Liogenys latitarsis Moser 1918
- Author
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Liogenys latitarsis ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys latitarsis Moser, 1918 Figs. 10; 11; 26. Liogenys latitarsis Moser, 1918: 99; Blackwelder 1944: 227 (checklist); Frey 1969: 50 (key); Frey 1969: 51, 63 (key, redescription [named as L. denticeps]); Evans 2003: 208 (checklist); Krajčík 2012: 145 (checklist); Evans & Smith 2009: 176 (checklist); Cherman et al. 2017: 4 (generic history). Type material. Liogenys latitarsis male lectotype here designated (ZMHB): [white, outlined black, typeset] “ Argentinien / Santiago del Estero ”, [white handwritten] “ Liogenys / latitarsis / Mos / Typen ♂”, [light red typeset] “Typus”, [white handwritten] “latitarsis / Mos.”, [white, outlined red, typeset and handwritten] “ LIOGENYS / LATITARSIS / Moser, 1918 / LECTOTYPE / Des. Cherman M. A.”, genitalia mounted. Paralectotypes (1): one female (ZMHB): with the same data of the lectotype, plus the label: [white, outlined red, typeset and handwritten] “ LIOGENYS / LATITARSIS / Moser, 1918 / PARALECTOTYPE / Des. Cherman M. A.”. Non-type material (36). BOLIVIA. Santa Cruz: Caballero, ~ 9 km SW Comarapa, 17º58.926’S 64º34.365’W, XII.2008, 1694 m, W.D.E. Edmonds & T. Vidaurre, [collected in] human feces, 6 males (CEMT); Cochabamba: Cochabamba, Bolivia Andina, Zischka, 1.XII.1969, 1 male; 10.I.1960, 1 female (NHMB). PARAGUAY. Boquerón: Chaco, Teniente Enciso, 4.XI.2001, C. Aguilar J., 2 males (CMVD); Parque Nacional Teniente Inciso, administración, 21º12’S 61º39’W, 253 m, 18.IX.2003, B. Garcete, 3 males and 1 female (DZUP). ARGENTINA. Jujuy: Estación Yuto, without date and collector, 1 male (NHMB); Salta: Chaco Occidental, 100 km NE Los Colorados [20 km NE Cafayate], I.1993, M. McLaqughlin & B. Bestelmeyer, 1 male (UNSM); Formosa: Ruta Provincial 39, 23 km S Ingeniero Juárez, 24º05’27’’S 61º56’49’’W, 13–17.XII.2008, F. Ocampo, G. San Blas, & F. Campon, mercury vapor & ultraviolet light trap, 1 male and 1 female (IADIZA); Tucumán: Gobernador Garmendia, 22.I.1981, Roig, 1 male (IADIZA); without locality, date, and collector, 1 male (MLPA); Catamarca: Quiros (2 km S) on Ruta Nacional 157, 28º48’43.3’’S 65º06’22.6’’W, 9.XII. 2008, 305 m, F. Ocampo, G. San Blas, & F. Campon, at light, 1 female (IADIZA); El Peñón, [26.478056°S, 67.265°W], II.1947, 4000 m, A. Martínez, 1 female (CMNC); Ciudad de Catamarca, XI.1945, J. Bosq, 1 female (MZUC); Santiago del Estero: Tintina, without collector, X.1946, 1 female (MZUC); Santiago del Estero: Tintina, without collector, XI.1946, 1 male and 2 females (MZUC); Fernandez, XII.1935, J. Bosq, 2 males (MLPA); El Pinto, without date and collector, 1 female (NHMB); Mendoza: Santa Rosa, Reserva Biológica Ñancuñan, 10–15.III.2014, G. Arriagada, at light, 1 female (CMVD); 11–13.XII.2002, V.M. Dieguez, 1 male (CMVD); Reserva Biológica Ñancuñan, Médanos, 34º00’12’’S 67º55’08’’W, 4.II. 2006, 569 m, F.C. Ocampo, E. Ruiz, & G. Zalazar, at light, 2 males and 2 females (DZUP). URUGUAY. Montevideo: SAPar Labo Nº288 [34°53’01’’S 056°10’55’’W?], without collector, 1 male (USNM). Diagnosis. Body elongate, brownish to dark brown, elytra shiny to semiopaque (Fig. 10A); clypeal emargination deep but not reaching the base of the teeth, rounded and wide; clypeal lateral margin strongly convex and produced, forming a tooth-like projection; pronotal posterior corners rounded to subangulate (Fig. 10C); the first pair of elytral ridges more defined than the other three, sometimes the second pair also noticeable; basal apophysis of metacoxa produced; metafemur with scattered bristles; metatibia not carinate along the inner margin (Fig. 10D); pygidium convex, wider than it is long; pygidial width wider than the distance between spiracles of propygidium, disc with bristles throughout, erect bristles (Fig. 10E); in males protarsomeres and mesotarsomeres strongly enlarged, metatarsomeres slightly enlarged; parameres expanded after the midline towards the apex (Fig. 10F); outer margin dentate, undulate or straight (Fig. 11 A–E); parameres strongly convex in lateral view (Fig. 10G). Redescription. Length 8.0– 11.9 mm; width: 4.1–6.2 mm. Light brown to brown. Head: distance between eyes twice the width of one eye; clypeal emargination deep, rounded and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; lateral margin convex and produced, forming a tooth-like projection; distance between clypeal lateral projection and anterior margin of eye slightly longer than one eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth equal to the basal width of anterior tooth; angle acute or approximately 90º between outer side of anterior teeth and clypeal lateral projection; distal maxillary palpomere, maximum width almost twice the apical width; fovea deep, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and equal to the funicle. Thorax: anterior margin of pronotum slightly produced medially; glabrous, punctures coarse and sparse, denser and coarser towards the anterior margin; pronotal posterior corners rounded to subangulate; hypomere with long bristles near the outer margin and scales near the inner margin; mesepisternum scaly; sides of metaventrite scaly and with bristles; distance between mesocoxae and metacoxae longer than the length of the metacoxa; scutellum ogival, weakly punctate at the base or sides. Elytra: shiny to semiopaque, glabrous, light brown to dark brown; elytral suture slightly darker than elytron, weakly elevated; first pair of ridges more defined than the other three, sometimes the second pair also noticeable. Legs: three protibial teeth, the middle and apical equal in size, basal one slightly shorter; distance between basal and middle teeth slightly longer than between middle and apical; mesofemural surface setose; mesotibia subquadrate in cross section, surface coarsely sculptured; two transverse carinae, the apical carina incomplete or complete in males, always complete in females; metacoxa scaly; basal apophysis of metacoxa produced; metafemur with scattered bristles; metatibia not carinate along the inner margin, apical inner surface setose; strong metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger longer than the diameter of the tibial apex; in males tarsomeres I to IV enlarged in all legs; protarsomeres and mesotarsomeres strongly enlarged, metatarsomeres slightly enlarged; protarsomere II as long as it is wide; metatarsomere I twice shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw less than twice as long and as wide as the inferior tooth, distance between teeth equal to the length of the inferior tooth. Abdomen: ventrites abundantly with bristles on disc, with scattered bristles in females; propygidium slightly visible, with bristles; pygidium convex; trapezoidal, wide; pygidial width wider than the distance between spiracles of propygidium; pygidial disc with bristles throughout, erect bristles; pygidial apex quadrate to rounded. Parameres: basal region narrower than both sections of the parameres at its maximum width, parameral split at third portion, expanded after the midline towards the apex; outer margin indented (Fig. 11A, D, C, F, H, I), undulate (Fig. 11E, J) or almost straight (Fig. 11B, G); parameres strongly convex in lateral view. Type locality. ARGENTINA, Santiago del Estero. Geographical distribution. BOLIVIA (Santa Cruz, Cochabamba); PARAGUAY (Boquerón); ARGENTINA (Jujuy, Salta, Formosa, Tucumán, Catamarca, Santiago del Estero, Mendoza). Remarks. Liogenys latitarsis is the species with the broadest geographical distribution among the Chacoan Liogenys, here expanded to Bolivia and Paraguay, and intraspecific variations were found among the specimens examined. The varying features are mainly the presence and the amount of bristles at the clypeus; somewhat in the sharpness of the anterior clypeal teeth; the posterior corners of pronotum from rounded to subangle; the more or less elevated inner pair of elytral ridges; elytra shiny or semiopaque (less frequent); and in the shape of the outer margins of the parameres. Liogenys latitarsis strongly resembles externally L. vicina Frey, 1969 (Fig. 22) (for more details see L. vicina diagnosis)., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 23-27, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167.","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031"]} more...
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39. Liogenys parallela Frey 1965
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Liogenys parallela ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys parallela Frey, 1965 Figs. 18; 25. Liogenys parallelus Frey, 1965: 144; Frey 1969: 51 (key); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 4 (generic history). Liogenys parallela: Evans 2003: 212 (checklist); Evans & Smith 2009: 180–181 (checklist). Type material. Liogenys parallelus male holotype (NHMB): [white, outlined violet, typeset and handwritten] “ Bolivia andina- / Samaipata- 1500 m / 25. XI.59 Zischka ”, [red typeset] “TYPE”, [white, typeset and handwritten] “ Type ♂ / Liogenys / parallelus / n sp. / det. G. Frey, 1964 ”. Paratypes (1) (NHMB) with the same data of the holotype. Non-type material (17). BOLIVIA. Santa Cruz: 10 km E Comarapa, 17º57’8.47’’S 64º27’7.45’’W, 26.XI.2008, 1825 m, Edmonds & Vidaurre, 3 males and 3 females (CEMT); 9 km SW Comarapa, 17º58.926’S 64º34.365’W, XII.2008, 1694 m, Edmonds & Vidaurre, human feces, 2 males (CEMT); Cochabamba: Cochabamba, 12.X.1949, 3100 m, R. Zichka, 5 males and 3 females (MZUC); without date, R. Zischka & A. Martínez, 1 male (CMNC). Diagnosis. Length: 8.1–10.7 mm; width: 3.7–5.6 mm. Body and elytra reddish brown, elongate, sides parallel; pronotum darker than elytra; elytra glabrous, reddish brown to testaceous, shiny, or semiopaque (Fig. 18A); clypeal emargination deep, rounded, and narrow to very narrow; outer sides of anterior teeth parallel; clypeal lateral margin in females straight but undulate, in males convex with a sharp, tooth-like projection, angle between this projection and the anterior teeth approximately 90º; antennae with 10 antennomeres; pronotal posterior corners rounded (Fig. 18B); mesepisternum, sides of metaventrite, metacoxae and ventrites with bristles abundantly; epipleura covered by long bristles (Fig. 18A, C); pygidium flat; subquadrate; with bristles throughout, bristles erect, long (Fig. 18 D–E); in males, inner margin of metatibia strongly carinate straight towards apex; protarsomere II elongate, almost three times longer than protarsomere I in dorsal view; parameres uniformly wide along its length; parameral split slightly beyond the midline (Fig. 18G); apex slightly expanded, beaked on lateral margin (Fig. 18F); parameres in lateral view convex, not coplanar (Fig. 18H). Type locality. BOLIVIA, Samaipata [18°10’37.7’’S 63°52’58.4’’W], 1500 m. Geographical distribution. BOLIVIA (Santa Cruz, Cochabamba). Remarks. Frey (1965) described L. parallela from a type series comprised of a holotype and 16 specimens collected from Samaipata (Santa Cruz) and Cochabamba [Bolivia]. At the NHMB, where the series was deposited, we found only two specimens (the holotype and one male paratype), both from Samaipata. There is a female paratype specimen from Cochabamba at ZHMB, but it was identified by us as L. latitarsis specimen. Frey (1965) compared L. parallela with L. nigrofusca, as mentioned earlier (see remarks in L. nigrofusca), and also with L. obscura. According to Frey (1965), the latter is darker in color and the clypeal margin is less undulate than L. parallela. Liogenys cartwrighti (Fig. 2) and L. vicina Frey, 1969 (Fig. 22) were described a few years later and both look very much like L. parallela. Liogenys parallela differs from all these other species in the clypeal emargination even narrower, the bristles on epipleura long; the pygidial bristles more erect and longer; in males, the protarsomere II more than twice as long as the protarsomere I in dorsal view, and the inner margin of metatibia strongly carinate towards the apex. This latter feature is also shared with L. nigrofusca. Specimens of L. parallela from Comarapa (Santa Cruz) match exactly with the holotype. However, specimens from Cochabamba differ in the size slightly smaller, the elytra lighter, and in the parameres dorsally smooth at the midline (without longitudinal wrinkles). These differences might be due to populational variation, taking into account that they were collected in a transitional zone among the Chaco, the Yungas, and the Rondonia Amazonian Province (Morrone 2014) (Fig. 25)., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 41-43, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Frey, G. (1965) Neue s ʾ damerikanische Melolonthidae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 16, 143 - 151.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Frey, G. (1964) Neue Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 15, 691 - 701.","Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1"]} more...
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40. Liogenys bidenticeps Moser 1919
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys bidenticeps ,Taxonomy - Abstract
Liogenys bidenticeps Moser, 1919 Fig. 1. Liogenys bidenticeps Moser, 1919: 13; Blackwelder 1944: 227; (checklist), Frey 1969: 43 (key); Evans 2003: 207 (checklist); Evans & Smith 2009: 175 (checklist); Cherman et al. 2011: 620 (immatures key); Krajčík 2012: 144 (checklist); Cherman et al. 2013: 273, 277 (natural history); Cherman et al. 2014a: 285 (natural history); Rodrigues et al. 2014: 235–238 (natural history); Cherman et al. 2016: 759 (systematics); Cherman et al. 2017: 26 (redescription); Rodrigues et al. 2017: 339–348 (immatures description, adult natural history). Liogenys bicuspis Moser, 1919: 14; Blackwelder 1944: 227; (checklist), Frey 1969: 43 (key); Evans 2003: 207 (checklist); Evans & Smith 2009: 175 (checklist); Krajčík 2012: 144 (checklist); Cherman et al. 2017: 26 (synonymy). Liogenys ophtalmicus Frey, 1973: 256; Krajčík 2012: 145 (checklist); Cherman et al. 2017: 5 (generic history). New synonym. Liogenys opthalmica [sic]: Evans 2003: 212 (checklist); Evans & Smith 2009: 180 (checklist). Type material. Types of Liogenys bidenticeps (male lectotype and three paralectotypes; ZMHB) and of Liogenys bicuspis (male lectotype and one paralectotype; ZMHB) were discussed in Cherman et al. (2017). Liogenys ophtalmicus male holotype (CASC) bearing the labels: [white typeset] “Ipane River / Chaco, Par / A. Schulze”, [white typeset] “L.W. Saylor / Collection”, [white, typeset and handwritten] “ California Academy / of Sciences / Type / No. 11816”, [red, typeset and handwritten] “Type / G. Frey 1973 / Liogenys / ophtalmicus”, genitalia mounted. Paratypes (1) (NHMB) with the labels: [white typeset] “Ipane River / Chaco, Par / A. Schulze”, [white typeset] “L.W. Saylor / Collection”, [white, red typeset] “Leech”, [red, typeset and handwritten] “ Paratype / G. Frey 1973 / Liogenys / ophtalmicus / male”, “♂”, genitalia mounted. Diagnosis and redescription. See Cherman et al. (2017). Type locality. Liogenys bidenticeps: BRAZIL, S„o Paulo; L. bicuspis: BRAZIL, Cuiabá, Mato Grosso; L. ophtalmicus: PARAGUAY, Chaco, Río Ypané [natural border between Concepción and San Pedro departments]. Geographical distribution. BRAZIL (Mato Grosso, Mato Grosso do Sul, Bahia, S„o Paulo, Paraná, Rio Grande do Sul); PARAGUAY (Concepción, San Pedro, Itapúa); ARGENTINA (Formosa, Catamarca, Santiago del Estero). Remarks. The holotype and paratype of L. ophtalmica, at CASC and NHMB, respectively, were studied. Frey (1973) stated and drew the “extraordinary shape of the head” and the “eyes exceedingly large, with distance between them less than one half the diameter of the eye”, and we verified these features on both type specimens. Surprisingly, the author did not recognize that the two type specimens, from the same collection event, bear a teratologic head (Fig. 1 A–B). In the case of the holotype, the canthus is absent, so the insertion of the antennae is exposed (Fig. 1A). In the paratype, a palp-like flagellum grew from the scape of the right antennae (Fig. 1G, J). Besides the abnormal head, most of external body features are not teratological, although the body is dorsoventrally flattened and the legs more slender than B. bidenticeps. We conclude that L. bidenticeps and L. ophtalmica are synonyms principally based on the parameres (Fig. 1L, M), which are an exact match., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 4-6, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1919) Beitrag zur Kenntnis der Melolonthiden (Col.). Stettiner Entomologische Zeitung, 80, 3 - 64.","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Cherman, M. A., Guedes, J. V. C., Moron, M. A., Dal Pra, E., Perini, C. R. & Jung, A. H. (2011) First record of species of Liogenys (Coleoptera, Melolonthidae) associated with winter crops in Rio Grande do Sul (Brazil). Revista Brasileira de Entomologia, 55, 618 - 620. https: // doi. org / 10.1590 / S 0085 - 56262011005000052","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Guedes, J. V. C., Moron, M. A., Dal Pra, E. & Bigolin, M. (2013) White grubs (Coleoptera, Melolonthidae) in the \" Planalto Region \", Rio Grande do Sul state, Brazil: key for identification, species richness and distribution. Revista Brasileira de Entomologia, 57, 271 - 278. https: // doi. org / 10.1590 / S 0085 - 56262013000300005","Cherman, M. A., Moron, M. A., Dal Pra, E., Valmorbida, I. & Guedes, J. V. C. (2014 a) Ecological characterization of white grubs (Coleoptera: Melolonthidae) community in cultivated and noncultivated fields. Neotropical Entomology, 43, 282 - 288. https: // doi. org / 10.1007 / s 13744 - 014 - 0214 - 0","Rodrigues, S. R., Nogueira, G. A. de L. & Gomes, E. S. (2014) Biological aspects of Liogenys bidenticeps Moser, 1919 (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 68, 235 - 238. https: // doi. org / 10.1649 / 0010 - 065 X- 68.2.235","Cherman, M. A., Moron, M. A. & Almeida, L. M. (2016) Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology, 41, 744 - 770. https: // doi. org / 10.1111 / syen. 12188","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Rodrigues, S. R., Fuhrmann, J., Gomes, E. S. & Amaro, R. A. (2017) Description of immatures and mating behavior of Liogenys bidenticeps Moser, 1919 (Coleoptera: Melolonthidae: Melolonthinae). Revista Brasileira de Entomologia, 61, 339 - 348. https: // doi. org / 10.1016 / j. rbe. 2017.07.006","Frey, G. (1973) Neue Macrodactylini (Col., Scarab., Melolonthinae). 20. Beitrag zur Kenntnis der s ʾ damerikanischen Melolonthiden. Entomologische Arbeiten aus dem Museum G. Frey, 24, 255 - 279."]} more...
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41. Liogenys nigrofusca Moser 1918
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
- Subjects
Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Liogenys nigrofusca ,Animalia ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys nigrofusca Moser, 1918 Figs. 14; 24. Liogenys nigrofuscus Moser, 1918: 103; Frey 1969: 50 (key); Krajčík 2012: 145 (checklist); Cherman et al. 2017: 4 (generic history). Liogenys nigrofusca: Blackwelder 1944: 227 (checklist); Evans 2003: 211 (checklist); Evans & Smith 2009: 179 (checklist). Liogenys longicrus Moser, 1918: 100; Frey 1965: 145 (synonymy). Liogenys longicra: Blackwelder 1944: 227 (checklist). Type material. Liogenys nigrofuscus male holotype (ZMHB): [white, typeset and handwritten] “ Argentinien / Gran Chaco ”, [white handwritten] “ Liogenys / nigrofuscus / Typen Mos ”, [red typeset] “ Typus ”, [white typeset] “ Liogenys / nigrofuscus / Mos.” Liogenys longicrus male holotype (ZMHB): [white typeset] “ Argentinien / Santiago del Estero ”, [white handwritten] “ Liogenys / longicrus / Typen Mos ”, [red typeset] “ Typus ”, [white handwritten] “ Liogenys / nigrofuscus / Mos. [typeset] det. G. Frey 1964 ” Non-type material (10). PARAGUAY. Boquerón: Colonia Menonitas, VII.1945, Podtiaguin 2 males (MZUC); Guachalla, A. Pilcomayo, 23.VIII.1945, P. Willim, 1 male (MZUC). ARGENTINA. Salta: Viñaco (15 km El Carril), 12.II.1982, H. Howden, 1 female (CMNC); Coronel Moldes, XII.1958, A. Martínez, 1 female (MZSP); Chaco: Pampa del Infierno [26º31’S, 61º10’W], IX.1982, A. Martínez, 1 female (CMNC); Estación San Francisco, Departamento San Fernando, 16–17.X.2013, 54 m, 27°30’35.90’’S, 59°4’47.60’’W, M. Ibarra Polesel, 1 specimen; Antequera, Departamento Primero de Mayo, 22.XII.2014, 52 m, 27°26.500’S, 58°51.459’W, M. Ibarra Polesel, 1 specimen (CARTROUNNE); Santiago del Estero: El Pinto, without date and collector, 2 females (NHMB). Diagnosis. Length: 9.1–10.0 mm; width: 4.1–4.8 mm. Body and elytra dark brown, elongate, shiny, sides parallel; pronotum and elytra unicolored, head darker; elytra glabrous (Fig. 14A); pronotum roughly punctate; frons somewhat swollen and depressed medially in females; clypeal emargination deep, rounded and narrow; outer sides of anterior teeth parallel; teeth sharp; clypeal lateral margin bearing a sharp tooth-like projection, angle between this projection and the anterior teeth approximately 90º; antennae with 10 antennomeres; pronotal posterior corners rounded (Fig. 14B); hypomere, mesepisternum, sides of metaventrite, metacoxae and sides of ventrites with sparse long bristles (Fig. 14C); protibial teeth, distance between medial and apical shorter than basal and medial; mesotibia cylindrical in cross section; two strong transverse carinae posteriorly, the basal sloped, the apical complete; pygidium flat; quadrate; pygidial disc very coarsely and densely punctate, sparser in females; with bristles throughout, bristles erect, long (Fig. 14D); in males, inner margin of metatibia strongly carinate straight towards the apex, somewhat curved outwards (Fig. 14C); protarsomere II elongate; parameres narrowed medially; parameral split at the midline; apex club shaped and strongly truncate on upper margin (Fig. 14E); parameres in lateral view convex, not coplanar (Fig. 14F). Type locality. Liogenys nigrofusca: ARGENTINA, “Gran Chaco ”; L. longicra: ARGENTINA, Santiago del Estero. Geographical distribution. PARAGUAY (Boquerón); ARGENTINA (Salta, Chaco, Santiago del Estero). Remarks. Frey (1965) synonymized L. nigrofusca with L. longicra, and compared this species with the preceding one in order of description, L. parallela Frey, 1965 (Fig. 18). According to Frey (1965), both are almost undistinguishable but the parameres. Although L. nigrofusca resembles L. parallela in the body size and in the shape of the clypeus, especially in females, they have external differences. This species differs from L. parallela (in parenthesis) mainly in the color darker, basal transverse carina on the posterior margin of mesotibia and metatibia strong and sloped (basal transverse carina less marked and straight on mesotibia, absent on metatibia); and in the pygidium quadrate, with punctures on disc coarser and uniformly disposed., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 31-33, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Frey, G. (1965) Neue s ʾ damerikanische Melolonthidae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 16, 143 - 151.","Frey, G. (1964) Neue Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 15, 691 - 701."]} more...
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42. Liogenys isotarsis Cherman 2021, new species
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Liogenys isotarsis ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys isotarsis Cherman, new species Figs. 9; 27. http://zoobank.org/ 0D51DD77-5048-4285-B0F6-A0785885A6F0 Type material: Holotype labeled (USNM): [white typeset] “ Trinidad, Bolivia / 1922-23 / M. S. Pennington ”, [red, typeset and handwritten] “ LIOGENYS ISOTARSIS / HOLOTYPE / Cherman M. A.”, genitalia mounted. Holotype deposited at USNM. Diagnosis. Body elongate; purplish brown, pronotum and elytra coarsely punctate (Fig. 9A); clypeal emargination angulate, teeth very short; outer sides of clypeal teeth subparallel; pronotal posterior corners rounded (Fig. 9B); in males metatibia not carinate along the inner margin, apical inner surface glabrous; tarsomeres almost equal in width, cylindrical (Fig. 9 D–E); pygidium flat, glabrous (Fig. 9C); parameres narrowed at the midline; widened towards the apex, apex equal in width to the basal portion (Fig. 9F); parameres convex in lateral view (Fig. 9G). Holotype. Male. Length: 9.1 mm; width: 5.1 mm. Purplish brown. Head: distance between eyes more than three times the width of one eye; clypeal emargination deep, angulate and wide; teeth short, outer sides of anterior teeth subparallel; outer margin of anterior teeth noticeably shorter than the eye length; lateral margin slightly convex, not produced; distal maxillary palpomere, maximum width more than twice the apical width; fovea shallow, extending past the transverse midline; antenna with 10 antennomeres, club lighter in color and longer than the funicle. Thorax: anterior margin of pronotum slightly produced medially; pronotal disc glabrous, punctures strongly coarse and very sparse; posterior corners rounded; hypomere with short bristles, mesepisternum and sides of metaventrite with sparse thick bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, with two longitudinal rows of punctures strongly coarse. Elytra: shiny, glabrous, unicolored with the pronotum, coarsely punctate; elytral suture unicolored with the elytron and weakly elevated; ridges I, II, and IV more defined than ridge III. Legs: three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface almost glabrous, setae on margins; mesotibia cylindrical in cross section, surface coarsely sculptured; two transverse carinae, the apical carina complete; metacoxa with sparse bristles, more abundant near the insertion of the metafemur; basal apophysis of metacoxa produced; metatibia not carinate along the inner margin, apical inner surface glabrous; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; protarsomeres and mesotarsomeres I to IV almost equal in width to the metatarsomeres I to IV (Fig. 9E); metatarsomere I equal in size to metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and equal in width to the inferior, distance between teeth slightly longer than the inferior tooth. Abdomen: ventrites with scattered short bristles; propygidium slightly visible, glabrous; pygidium flat, subtrapezoidal, wide; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex rounded. Parameres: narrowed at the midline; basal region wider than the parameres combined at the transverse midline, parameral split at 2/3; inner margins slightly convergent, after the midline widened towards the apex, apex equal in width to the basal portion; parameres convex in lateral view. Etymology. Adjective in the nominative singular. From Ancient Greek ἴΣΟΣ (ísos, “equal”) + tarsus. In Liogenys isotarsis all the three pairs of tarsi have almost the same width, even being a male. Type locality. BOLIVIA, Trinidad [14°49’55.6’’S 64°54’09.6’’W]. Geographical distribution. BOLIVIA (Beni). Remarks. Liogenys isotarsis resembles L. unicolor Evans, 2003 (Cherman et al. 2017, 2019a) in the size and body color. Liogenys isotarsis differs from L. unicolor mainly in the outer sides of clypeal teeth subparallel (follow the lateral margin of clypeus); the pronotum wider; the punctures of scutellum and elytra coarser; the elytral ridges more defined; and in the pygidium flat (convex). In males, the protarsomeres are even more cylindrical and less enlarged than in L. unicolor. The parameres resemble those of L. angustitarsis Cherman, 2019 (Cherman et al. 2019a), but they differ mainly in the lateral view, which are concave in the new species. Females remain unknown. Liogenys isotarsis is possibly associated only to Bolivian Chiquitano Forests, and might occur in sympatry with L. parallela Frey, 1965 and L. mendozana Moser, 1918 in their northernmost distribution., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 21-23, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Cherman, M. A., Basilio, D. S., Mise, K. M. & Almeida, L. M. (2019 a) Unraveling the puzzle of Liogenys biodiversity: fifteen new species, nomenclatural acts and new geographical records (Coleoptera: Scarabaeidae). Insect Systematics & Evolution, 51, 1 - 69. https: // doi. org / 10.1163 / 1876312 X- 00001040","Frey, G. (1965) Neue s ʾ damerikanische Melolonthidae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 16, 143 - 151.","Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167."]} more...
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43. Liogenys tarsalis Moser 1921
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Liogenys tarsalis ,Biodiversity ,Liogenys ,Taxonomy - Abstract
Liogenys tarsalis Moser, 1921 Figs. 7A, C, E; 23. Liogenys tarsalis Moser, 1921: 54; Blackwelder 1944: 228 (checklist); Cherman et al. 2016: 23 (new combination); Cherman et al. 2017: 75 (redescription). Homoliogenys tarsalis: Gutiérrez 1952: 216 (genus description); Frey 1969: 44 (key); Evans 2003: 206 (checklist); Evans & Smith 2009: 175 (checklist); Krajčík 2012: 129 (checklist). Type material. Liogenys tarsalis male lectotype and two female paralectotypes at ZMHB, two male paralectotypes at NHRS; data in Cherman et al. (2017). Non-type material (5). ARGENTINA. Santa Fé: without locality and date, R. Richter, 2 females (MLPA). Santiago del Estero: without locality, date, and collector, 1 female (NHMB); Corrientes: Mercedes, Reserva Itatí, 12.XII.2013, 60 m, 28°45’52.30’’S, 58°7’0.50’’W, at light, grassland, M. Ibarra Polesel, 1 male (CARTROUNNE); San Luis: Villa Mercedes, Facultad de Ingeniería y Ciencias Agropecuarias. Universidad Nacional de San Luis, 22.X.2019, 33.641641ºS 65,446229ºW, at light, M.A. Cherman, S.L. Bonivardo, & A.N. Martínez, 1 male (DZUP). Geographical distribution. ARGENTINA (Chaco, Santiago del Estero, Corrientes Santa Fé, San Luis). Diagnosis. Male. Length 10.0– 10.5 mm; width: 5.2–5.4 mm. Brown, elongate, sides almost parallel; elytra with non-uniform brown, base darker, as are the head and pronotum; clypeal emargination deep, rounded or subangulate and wide; in males teeth as long as the eye or slightly longer; outer sides of anterior teeth parallel; clypeal lateral margin convex, not produced; club of antenna longer than the funicle; metacoxae scaly throughout, pronotal posterior corners rounded; metacoxal basal apophysis not produced; apical inner surface glabrous; metatibial spurs with equal lengths; tarsi opaque and enlarge in all legs, mainly the tarsomere II; pygidium convex, as wide as it is long; parameres glabrous with inner margins strongly convergent, narrowed subapically; apex bilobed, curved inwards, inner margins overlapped (Cherman et al. 2017). Female. Length: 11.0– 11.4 mm; width 5.8–6.3 mm. Burned brown (Fig. 7A), clypeus wider than in males; emargination shallower, teeth slightly shorter than the eye, parallel or subparallel; clypeal lateral margins more convex (Fig. 7C); mesotibia and metatibia with transverse carina more-or-less strong; pygidial apex more quadrate and narrower (Fig. 7E). Remarks. Cherman et al. (2017) suggested that Rio de Janeiro is a doubtful type locality of L. tarsalis, because almost all the material collected by E.R. Wagner was from Río Salado (Santiago del Estero, Argentina), therefore, it might be written “Rio Sal” instead. One of the females housed at the NHMB was collected in Río Salado. Moreover, no L. tarsalis specimens were found from Rio de Janeiro [Brazil], among all the 30 collections studied. Thus, it is more likely that the type locality of L. tarsalis is Río Salado [Argentina] instead of Rio de Janeiro [Brazil]. The Brazilian distribution of L. tarsalis is disregarded, and confirmed for Chaco, Santiago del Estero, Corrientes, Santa Fe, and San Luis Argentinian provinces. There are Frey type labels for “ Liogenys nigricans ” attached to a female specimen from Santiago del Estero (NHMB) and in other collections where Frey deposited or exchange specimens as well (e.g., USNM). Since the name L. nigricans was only used on labels, it is a nomem nudum and is unavailable (Article 13.1.1, International Commission on Zoological Nomenclature 1999)., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 47-48, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312","Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2","Cherman, M. A., Moron, M. A. & Almeida, L. M. (2016) Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology, 41, 744 - 770. https: // doi. org / 10.1111 / syen. 12188","Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031","Gutierrez, R. (1952) Notas sobre Scarabaeidae Neotropicos. Revista Chilena de Entomologia, 2, 207 - 227.","Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64.","Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1","Evans, A. V. & Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)","Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp.","International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp."]} more...
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44. Liogenys truncata Cherman 2021, new species
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Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes, and Almeida, Lúcia Massutti De
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Coleoptera ,Insecta ,Arthropoda ,Melolonthidae ,Animalia ,Biodiversity ,Liogenys ,Liogenys truncata ,Taxonomy - Abstract
Liogenys truncata Cherman, new species Figs. 21; 27. http://zoobank.org/ D09F18D9-F0DA-429B-A7DF-C70B253EAC08 Type material. Holotype labeled (CMNC): [white typeset] “ ARGENTINA, Prov. Salta / 3 km W Cabra Corral Dam / 30.I.1982 / H & A Howden”, [red, typeset and handwritten] “ LIOGENYS TRUNCATA / HOLOTYPE / Cherman M. A.”, genitalia mounted. Paratypes (3), one male and two females (CNCI): “ ARGENTINA / Stgo. del Estero / Choya [strikethrough] XII-1958 ”, [yellow, typeset and handwritten] “ LIOGENYS TRUNCATA / PARATYPE / Cherman M. A.”. Holotype deposited at CMNC. Paratypes deposited at CNCI. Diagnosis. Body elongate; yellowish brown, pronotum and elytra unicolored, elytra uniformly with bristles (Fig. 21A); clypeal emargination very wide and shallow, almost truncate (Fig. 21B); procoxal and mesotibial surface smooth, metatibial surface coarsely punctate; apical inner margin of metatibia with sparse, long bristles (Fig. 21C); pygidium with bristles and as wide as it is long (Fig. 21D); in males protarsomeres I to IV weakly enlarge; parameres narrowed subapically; apex flap shaped, outer margin angulate inferiorly, angle in a plane below the apex (Fig. 21E). Holotype. Male. Length: 7.7 mm; width: 3.5 mm. Yellowish brown, head and margins of pronotum darker. Head: distance between eyes almost three times the width of one eye; frons with bristles; anterior margin of clypeus apparently truncate; clypeal emargination very shallow, almost straight, as wide as the distance between eyes; lateral margin of clypeus straight, slightly convex near the canthus; distal maxillary palpomere, maximum width less than twice the apical width; fovea shallow, extending up to the transverse midline; antenna with 10 antennomeres, club equal in color and longer than the funicle. Thorax: anterior margin of pronotum slightly produced medially; pronotum anteriorly narrowed forming a depressed ring; pronotal disc glabrous, punctures coarse and very sparse; posterior corners subangulate, angle obtuse; hypomere with long bristles, mesepisternum and sides of metaventrite with thick and fine bristles; distance between mesocoxae and metacoxae slightly longer than the metacoxa; scutellum ogival, finely punctate throughout. Elytra: shiny, uniformly with bristles, bristles longer at the margins of the elytra, unicolored with the pronotum; elytral suture unicolored with the elytron and not elevated; all elytral ridges weakly defined. Legs: procoxa with bristles, smooth; three protibial teeth, the apical the longest; the three teeth equally spaced; mesofemural surface almost glabrous, setae on margins; mesotibia cylindrical in cross section, surface smooth; two transverse carinae, the apical carina incomplete; metacoxa with sparse, thick bristles; basal apophysis of metacoxa produced; metatibia carinate along the inner margin towards the apex, apical inner surface sparsely setose, bristles long; metatibial surface coarsely punctate; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; protarsomeres I to IV enlarged; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and equal in width to the inferior, distance between teeth shorter than the inferior tooth. Abdomen: ventrites and propygidium with bristles; pygidium slightly convex, subquadrate, as wide as it is long; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc with bristles throughout; pygidial apex quadrate. Parameres: basal region slightly wider than both sections of the parameres at the transverse midline, parameral split at 2/3; inner margins convergent at the apex, narrowed subapically; apex flap shaped, outer margin angulate inferiorly, angle in a plane below the apex; parameres in lateral view slightly convex, coplanar (Fig. 21F). Variation. Male paratype: length: 8.2 mm; width: 4.2 mm. Equal to the holotype except in the pronotal punctures finer and sparser and pygidial apex rounder. Female paratypes: length: 7.8–8.0 mm; width: 3.6–3.9 mm. Equal to the holotype except in the abdomen with bristles less abundant and more sparse. Etymology. Adjective in the nominative singular. From Latin truncâtus, perfect passive participle of truncô (“maim, reduce to a trunk”), meaning that a structure is ending abruptly as if cut across at the base or tip. This term is referring to the clypeus truncate in Liogenys truncata. Type locality. ARGENTINA, Salta, 3 km W Cabra Corral [25°16’37.6’’S 65°23’20.4’’W]. Geographical distribution. ARGENTINA (Salta, Santiago del Estero). Remarks. Liogenys truncata resembles L. rufoflava Moser, 1918 in the body color and the bristle pattern of pronotum, elytra, and pygidium. Liogenys truncata differs from L. rufoflava (in parenthesis) mainly in the body size smaller, the clypeus almost truncate, not forming noticeable anterior teeth (two distinct teeth and a wide emargination between them), lateral margin of clypeus straight (convex, with a rounded projection); the pronotal punctures denser at the disc; the elytra entirely shiny (pruinose at the margins); the pygidium as long as it is wide (longer than it is wide); and in males the protarsi and mesotarsi slightly enlarged (distinctly enlarged); and the parameres longer, with apex spatulate (lanceolate)., Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 48-50, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258, {"references":["Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167."]} more...
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45. Figures 40-43 from: Del G. da R. Celli N, Almeida LM, Basílio DS, Castro CF (2021) The way to maturity: taxonomic study on immatures of Southern Brazilian Coccinellini (Coleoptera: Coccinellidae) species important in biological control. Zoologia 38: 1-18. https://doi.org/10.3897/zoologia.38.e64154
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Del G. da R. Celli, Nathália, primary, Almeida, Lúcia M., additional, Basílio, Daniel S., additional, and Castro, Camila F., additional
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46. Figures 57-65 from: Del G. da R. Celli N, Almeida LM, Basílio DS, Castro CF (2021) The way to maturity: taxonomic study on immatures of Southern Brazilian Coccinellini (Coleoptera: Coccinellidae) species important in biological control. Zoologia 38: 1-18. https://doi.org/10.3897/zoologia.38.e64154
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Del G. da R. Celli, Nathália, primary, Almeida, Lúcia M., additional, Basílio, Daniel S., additional, and Castro, Camila F., additional
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47. Figures 66-70 from: Del G. da R. Celli N, Almeida LM, Basílio DS, Castro CF (2021) The way to maturity: taxonomic study on immatures of Southern Brazilian Coccinellini (Coleoptera: Coccinellidae) species important in biological control. Zoologia 38: 1-18. https://doi.org/10.3897/zoologia.38.e64154
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Del G. da R. Celli, Nathália, primary, Almeida, Lúcia M., additional, Basílio, Daniel S., additional, and Castro, Camila F., additional
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48. Figures 1-11 from: Del G. da R. Celli N, Almeida LM, Basílio DS, Castro CF (2021) The way to maturity: taxonomic study on immatures of Southern Brazilian Coccinellini (Coleoptera: Coccinellidae) species important in biological control. Zoologia 38: 1-18. https://doi.org/10.3897/zoologia.38.e64154
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Del G. da R. Celli, Nathália, primary, Almeida, Lúcia M., additional, Basílio, Daniel S., additional, and Castro, Camila F., additional
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49. Figures 71-79 from: Del G. da R. Celli N, Almeida LM, Basílio DS, Castro CF (2021) The way to maturity: taxonomic study on immatures of Southern Brazilian Coccinellini (Coleoptera: Coccinellidae) species important in biological control. Zoologia 38: 1-18. https://doi.org/10.3897/zoologia.38.e64154
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Del G. da R. Celli, Nathália, primary, Almeida, Lúcia M., additional, Basílio, Daniel S., additional, and Castro, Camila F., additional
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50. Figures 18-26 from: Del G. da R. Celli N, Almeida LM, Basílio DS, Castro CF (2021) The way to maturity: taxonomic study on immatures of Southern Brazilian Coccinellini (Coleoptera: Coccinellidae) species important in biological control. Zoologia 38: 1-18. https://doi.org/10.3897/zoologia.38.e64154
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Del G. da R. Celli, Nathália, primary, Almeida, Lúcia M., additional, Basílio, Daniel S., additional, and Castro, Camila F., additional
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