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A key to extinct Eocene species of Atomaria 1. Body size smaller than 1.5 mm.......................................................................... 2 - Body size larger than 1.5 mm........................................................................... 4 2. Body size 1.0– 1.2 mm. Antennomere 3 short, subquadrate, as long as wide, much shorter than antennomere 2. Pronotal disc convex.............................................................................................. 3 - Body size 1.35 mm. Antennomere 3 elongated, 1.5 times as long as wide, slightly shorter than antennomere 2. Antennomere 5 conical, elongated, 1.5× as long as wide. Pronotal disc flat. Lateral side borders of pronotum visible from above only in basal half of its length. Bitterfeld amber......................................... A. saxonica Lyubarsky et Perkovsky 3. Lateral side borders of pronotum visible from above along entire its length. Pronotum not narrowed toward base. Pronotum maximum width before the middle of its length. Antennomere 5 conical, elongated, 1.5× as long as wide. Body size 1.0 mm. Baltic amber............................................................ A. groehni Perkovsky et Lyubarsky - Lateral side borders of pronotum visible from above only in basal half of its length. Pronotum narrowed toward base. Pronotum maximum width in the middle of its length. Antennomere 5 conical, nearly as long as wide, 1.1× as long as wide. Body size 1.19 mm. Rovno amber............................................................... A. perkovskyi sp. nov. 4. Pronotal disc convex.Antennomere 3 about 2× as long as wide.Antennomeres 9 and 10 strongly transverse. Pronotum maximum width nearly in the middle of its length. Base of pronotum with shallow depression. Body size 1.54 mm. Baltic amber......................................................................................... A. alekseevi sp. nov - Pronotal disc flat. Antennomere 3 about 1.5× as long as wide. Antennomere 10 subquadrate, nearly as long as wide. Pronotum maximum width much behind the middle of its length. Base of pronotum without depression. Body size 1.65 mm. Baltic amber.............................................................. A. gedanicola Lyubarsky et Perkovsky, Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on page 288, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693

Atomaria (Anchicera) perkovskyi sp. nov. (Figs 3–4) Type material. Holotype: collection number No 6832 [MAIG] (ex coll. Jonas Damzen JDC-10149 R); adult, sex unkown. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 38× 52 mm and a maximum thickness of 15 mm; preserved without supplementary fixation. Ventral side of specimen completely obscured by milky opacity. Syninclusions: four small Nematocera (Diptera) specimens. Type stratum. Rovno amber, Late Eocene (Perkovsky et al. 2007). Type locality. Rivne Oblast (region), Ukraine. Description. Measurements: body length (from anterior margin of head to elytral apex along midline) about 1.19 mm, body maximum width across both elytron 0.50 mm; head length 0.16 mm, head maximum width across eyes 0.27 mm; pronotum length 0.30 mm, pronotum maximum width 0.38 mm; elytra length 0.73 mm, elytra maximum width 0.50 mm. Body broadly elongate, moderately arched, slightly convex; integument uniformly darkbrown (as preserved); sparsely covered with semierect pubescence. Head transverse, about 1.7× as wide as long; coarsely punctate, distance between punctures equal to 0.5-1.0× diameter of one puncture, punctation denser laterally. Compound eyes hemispherical, convex, rather large, with large facets. Antennae inserted close to eyes, under lateral margin of forehead; antennal insertions widely separated; distance between antennal insertions larger than distance between antennae and аnterior margin of eye. Antennae 11-segmented with 3-segmented loose club, slender, short, reaching about to posterior one-third of pronotal length; antennomere 1 subcylindrical, elongate, 1.6× as long as wide; antennomere 2 conical, elongate, 1.5× as long as wide, slightly shorter than antennomere 1; antennomere 3 conical, short, nearly as long as wide, narrower and distinctly shorter than antennomere 2; antennomere 4 nearly as long as wide; antennomere 5 conical, nearly as long as wide, 1.1× as long as wide; antennomeres 6–8 trapezoidal, as long as wide; antennomeres 9–10 trapezoidal, strongly transverse, 1.5× as wide as long; antennomere 11 widely oval, with rounded apex, nearly as long as wide; relative length ratios of antennomeres 1–11 equal to 16:12:7:7:8:5:5:6:8:10:16. Pronotum transverse, 1.2× as wide as long, slightly narrowed posteriad and anteriad, with maximum width in middle of its length; with slightly convex disc; with finely margined posterior and lateral sides; lateral side borders visible from above only in basal half. Pronotal punctation moderately coarse and dense, distance between punctures equal to 1.5-2.0× diameter of one puncture; puncture diameter slightly smaller than eye facet diameter. Posterior and anterior angles obtuse. Anterior edge weakly sinuate, without excision; posterior edge strongly bisinuate, lobed, with shallow depression and basal pits; lateral edges widely rounded, without callosity and teeth. Scutellar shield small, oval, strongly transverse, 2.0× as wide as long, covered with fine punctation. Elytra short-oval, moderately convex, with maximum width about in anterior one-third of its length, 1.5× as long as wide and 2.4 times as long as pronotum length; completely covering the abdomen; lateral sides widely rounded; humeri rounded, slightly prominent. Elytral punctation irregular, moderately coarse and dense; (nearly as pronotal punctation), distance between punctures equal to 1.5-2.0× diameter of one puncture. Epipleura incomplete. Macropterous, metathoracic wings fully developed. Legs slender, long, relatively similar in shape, finely and densely punctate. Femora elongate-oval, flattened. Pretarsal claws simple, thin. Etymology. The species is named in honor of our dear colleague Dr. Evgeny E. Perkovsky (Kiev, Ukraine), leading researcher of inclusions in Rovno amber. Differential diagnosis. Atomaria perkovskyi sp. nov. can be distinguished from related species by a combination of characters: antennomere 3 short, nearly as long as wide, distinctly shorter than antennomere 2; antennomeres 9 and 10 strongly transverse; antennomere 5 conical, nearly as long as wide, 1.1× as long as wide; pronotum narrowed toward base; lateral side borders of pronotum visible from above only in basal half of its length; and pronotum maximum width in middle of its length. A new species can distinguished from all other extinct Eocene Atomaria representatives using the identification key provided below., Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on pages 286-288, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693, {"references":["Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Taraschuk, M. V. (2007) A comparative analysis of the Baltic and Rovno amber arthropod faunas: representative samples. African Invertebrates, 48, 229 - 245. [https: // hdl. handle. net / 10520 / EJC 84578]"]}

Atomaria (Anchicera) alekseevi sp. nov. (Figs 1–2) Type material. Holotype: collection number No 6831 [MAIG] (ex coll. Jonas Damzen JDC-10145); adult, sex unkown. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 17× 25 mm and a maximum thickness of 5 mm; preserved without supplementary fixation. Syninclusions: one Carabidae (Coleoptera) specimen, one small Nematocera (Diptera) specimen, and numerous stellate trichomes of Fagaceae. Type stratum. Baltic amber; Middle–Late Eocene (Sadowski et al. 2017; Seyfullah et al. 2018; Bukejs et al. 2019; Kasiński et al. 2020; Iakovleva et al. 2021). Type locality. Yantarny village (formerly Palmnicken), the Kaliningrad Region, Russia. Description. Measurements: body length (from anterior margin of head to elytral apex along midline) about 1.54 mm, body maximum width across both elytron 0.69 mm; head length 0.23 mm, head maximum width across eyes 0.38 mm; pronotum length 0.35 mm, pronotum maximum width 0.54 mm; elytra length 0.96 mm, elytra maximum width 0.69 mm. Body broadly elongate, moderately arched, slightly convex; integument uniformly lightbrown (as preserved); sparsely covered with semierect pubescence. Head transverse, about 1.7× as wide as long; coarsely punctate, distance between punctures nearly equal to diameter of one puncture. Compound eyes hemispherical, convex, rather large, with large facets. Antennae inserted close to eyes, under lateral margin of forehead; antennal insertions widely separated; distance between antennal insertions larger than distance between antennae and аnterior margin of eye. Antennae 11-segmented with 3- segmented loosed club, slender, rather long, reaching slightly beyond pronotal base; antennomere 1 subcylindrical, slightly elongate, 1.3× as long as wide; antennomere 2 conical, elongate, 1.6× as long as wide; antennomere 3 conical, elongate, about 2× as long as wide, distinctly narrower and slightly shorter than antennomere 2; antennomere 4 slightly elongate, 1.2× as long as wide; antennomere 5 conical, elongate, 1.5× as long as wide; antennomeres 6–8 trapezoidal, nearly as long as wide; antennomeres 9–10 trapezoidal, strongly transverse, 1.3–1.4× as wide as long; antennomere 11 widely oval, with rounded apex, 1.2× as long as wide; relative length ratios of antennomeres 1–11 equal to 10:10:7:5:6:5:5:5:6:7:13. Pronotum transverse, 1.5× as wide as long, narrowed posteriad and anteriad, with maximum width slightly beyond middle; with convex disc; with finely margined posterior and lateral sides; lateral side borders visible from above only in basal half. Pronotal punctation coarse and dense, distance between punctures equal to 0.5-1.5× diameter of one puncture, punctation distinctly sparser in posterior half; puncture diameter subequal to eye facet diameter; pronotal punctation larger than punctation of head. Posterior angles obtuse; anterior angles obtuse, slightly projecting anteriad. Anterior edge weakly sinuate, without excision; posterior edge strongly bisinuate, lobed, with shallow depression and basal pits; lateral edges widely rounded, without callosity and teeth, with distinct crenulation. Prohypomera slightly impressed; with sparse, fine punctation. Prosternum with disc convex; sparsely covered with coarse punctation, distance between punctures distinctly larger than diameter of one puncture. Prostenal process convex, rather narrow, with straight posterior margin, slightly dilated apically. Procoxal cavities apparently closed externally. Scutellar shield small, suboval, strongly transverse, 2.2× as wide as long, covered with fine punctation. Elytra short-oval, moderately convex, with maximum width in anterior one-third of its length, 1.4× as long as wide and 2.7 times as long as pronotum length; completely covering abdomen; lateral sides widely rounded; humeri rounded, slightly prominent. Elytral punctation irregular, coarse and dense; punctures in basal part slightly larger than punctures on pronotal disc, and distance between punctures equal to 1.5-2.0× diameter of one puncture. Epipleura narrowed posteriorly, incomplete, extending about to abdominal ventrite 2. Metaventrite with sparse, coarse punctation; disc convex. Macropterous, metathoracic wings fully developed. Legs slender, long, relatively similar in shape, finely and densely punctate. Procoxae large, widely suboval, transverse; mesocoxae round; metacoxae suboval, strongly transverse. Femora elongate-oval, flattened. Tibiae thin, almost straight, nearly as long as femora. Tarsi long and thin; tarsomeres simple, not lobed; tarsal formula 5-5-5. Abdomen with five visible, freely articulated ventrites; ventrite 1 sparsely covered with rather coarse punctation, ventrites 2–5 with small and denser punctures. Relative length ratios of ventrites 1–5 equal to 31:10:10:9:11 (measured medially). Etymology. The species is named in honor of our dear colleague Dr. Vitalii I. Alekseev (Kaliningrad, Russia), leading researcher of Coleoptera inclusions in Baltic amber. Differential diagnosis. Atomaria alekseevi sp. nov. can be distinguished from related species by a combination of characters: antennomere 3 elongated, about 2× as long as wide, slightly shorter than antennomere 2; antennomeres 9 and 10 strongly transverse; pronotal disc convex; pronotum narrowed toward base; lateral side borders of pronotum visible from above only in basal half of its length; pronotum maximum width nearly in middle of its length. A new species can distinguished from all other extinct Eocene Atomaria representatives using the identification key provided below., Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on pages 284-286, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693, {"references":["Sadowski, E. - M., Seyfullah, L. J., Schmidt, A. R. & Kunzmann, L. (2017) Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia, 106, 1 - 73.","Seyfullah, L. J., Beimforde, C., Dal Corso, J., Perrichot, V., Rikkinen, J. & Schmidt, A. R. (2018) Production and preservation of resins - past and present. Biological Reviews, 93, 1684 - 1714. https: // doi. org / 10.1111 / brv. 12414","Bukejs, A., Alekseev, V. I. & Pollock, D. A. (2019) Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa, 4664 (2), 261 - 273. https: // doi. org / 10.11646 / zootaxa. 4664.2.8","Kasinski, J. R., Kramarska, R., Slodkowska, B., Sivkov, V. & Piwocki, M. (2020) Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 bore hole, Kaliningrad region, Russia). Geological Quarterly, 64, 29 - 53. https: // doi. org / 10.7306 / gq. 1513","Iakovleva, A. I., Mychko, E. V. & Aleksandrova, G. N. (2021) New dinocyst data from amber quarry \" Primorsky \" (Kaliningrad region): clarification of the deposits age as base for the revision of paleogeographic reconstructions of Danish-Polish strait in late Paleogene. In: Golubev, V. K. & Nazarov, V. N. (Eds), Paleostrat- 2021. Annual meeting (scientific conference) of the Paleontological group of the Moscow Society of Naturalists and the Moscow branch of the Russian Paleontological Society, Russian Academy of Sciences (Moscow, 25 - 26 January 2021). Abstracts. Borissiak Paleontological Institute Press, Moscow, pp. 77 - 78. [in Russian]"]}

Subgenus Anchicera Thomson, 1863 Note. The fossil specimens under consideration are placed in Cryptophagidae based on antennae 11-segmented with 3-segmented loose club; tarsal formula 5-5-5; incomplete epipleura; elytra completely covering abdomen, with irregular punctation; procoxal cavities closed externally; tarsomeres simple, not lobed; and abdominal ventrite 1 longest. A new extinct species are assigned to the subgenus Anchicera within the genus Atomaria based on the combination of the following characters: distance between antennal insertions larger than distance between antennae and аnterior margin of eye; and lateral sides of elytra widely rounded, not parallel., Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on page 284, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693

Two new species of Atomaria Stephens, 1829 from China and Taiwan are described and illustrated: Atomaria (Atomaria) cooterisp. nov. and Atomaria (Atomaria) johnsoni sp. nov. New records for China and Taiwan are added: Atomaria (Atomaria) klapperichi Johnson, 1971 and Atomaria (Atomaria) plecta Lyubarsky, 1995. New subgeneric combinations are proposed for Atomaria (Atomaria) klapperichi, Atomaria (Atomaria) plecta and Atomaria (Atomaria) torrida Johnson, 1971 SI

Bestimmungsschlüssel der Männchen der Atomaria ·Arten mit dunkler Flügeldeckenzeichnung Männchen der Arten mit Makeln, Verschattungen oder Querbänder auf den Flügeldecken, die aus der Türkei bekannt sind oder dort vorkommen könnten. Die Weibchen sind mit denselben Merkmalen in aller Regel auch zu trennen (mit Ausnahme der fehlen- den Vordertarsenmodifikationen), es sollten jedoch nur Tiere gleichen Geschlechts mit- einander verglichen werden. 1 Vordertarsen der Männchen deutlich erweitert, erstes Glied etwa doppelt so breit wie das letzte Glied. Die Unterscheidung der beiden Arten kann im Einzelfall sehr schwer sein, konstante Unterschiede im Bau des Aedoeagus sind bislang nicht feststellbar...........2 - Vordertarsen der Männchen nicht oder nur sehr geringfügig erweitert, erstes Glied nicht doppelt so breit wie das letzte Glied...........................................................................3 2 Durchschnittlich grösser, um 2 mm (meist grösser als 1,8 mm), grösste Halsschildbreite in der Mitte, Halsschild stärker gewölbt, Flügeldecken gewölbter, aber seitlich in flacherer Rundung verengt, subparallel (Abb. 7). Verbreitung: Südöstliches Europa, Türkei, Grossbritannien........................................................ A. unifasciata ERICHSON, 18461 - Durchschnittlich kleiner, um 1,7 mm (meist kleiner als 1,8 mm), grösste Halsschildbreite vor der Mitte, Halsschild flacher gewölbt, Flügeldecken flacher, aber seitlich weniger flach gerundet (Abb. 8). Verbreitung: Teile Südeuropas, Kaukasus, Asien (vereinzelte Funde, nicht revidiert), Nordafrika.............. A. fasciata KOLENATI, 1846 3 Halsschild mit nur schmalen Seitenrändern und stark gewölbt, feiner, dicht oder weitläufiger punktiert. Seitenrandkante einfach. Halsschildbasis schmal niedergedrückt. Paramerenplatte ohne charakteristische Bildungen an der Spitze (Abb. 10). Dunkle Zeichnung (nahezu schwarz) der Flügeldecken ist oft sehr ausgedehnt und nur das Spitzenfünftel und die Basis, oft nur Schultern aussparend. Fühler lang und schlank, Keulenglieder nicht quer und Keule nur schwach von der Geissel abgesetzt.......................4 - Halsschild, besonders kurz vor den Hinterecken, mit deutlicher Seitenrandkehle, schwächer gewölbt, immer dicht und grob punktiert. Seitenrandkante ganz oder teilweise doppelt. Halsschildbasis kräftig niedergedrückt. Paramerenplatte mit charakteristischen Bildungen an der Spitze (Abb. 11 & 12). Dunkle Zeichnung (braun) der Flügeldecken aus zwei zur Naht verschmälerten Flecken bestehend, Naht schmal hell gefärbt. Fühler kräftig, Keulenglieder quer und Keule von der Geissel gut abgesetzt..............................................................................................................................5 4 Halsschildpunktur in der Mitte weitläufiger, Punktabstand wenigstens so gross wie der Punktdurchmesser (Abb. 5). Verbreitung: Zypern, Israel, Jordanien.................................................................................................................................... A. semnusta JOHNSON, 19692 - Halsschildpunktur in der Mitte dichter, Punktabstand geringer als ihr Durchmesser (Abb. 6). Verbreitung: Weite Teile Europas, Nordafrikas, Türkei und im Nahen Osten (evtl. auf vorherige Art zu beziehen)............................ A. scutellaris MOTSCHULSKY, 18493 5 Oberseite fein und mässig dicht punktiert, glänzend (Abb. 4). Halsschild von der grössten Breite etwas vor der Mitte gerade oder fast gerade nach hinten und vorn verengt, stumpf gewinkelt. Vordertarsen erkennbar breiter als Mittel- und Hintertarsen. Aedoeagus: Paramerenplatte tief eingeschnitten, seitlich hochgebogen. Verbreitung: Besonders Nord- und Mitteleuropa, bis Ost- und Südosteuropa........ A. ornata HEER, 18414 [ähnlich ist A. suffusa JOHNSON, 1971 (Abb. 9) aus Marokko, der Aedoeagus jenem der A. ornata sehr ähnelnd] - Oberseite gröber und mässig dicht punktiert, glänzend (Abb. 2 und 3)................................6 ¹ A. unifasciata liegt dem Verfasser aus der Türkei vor aus den Umgegenden von Konya, Niǧde, Nevşhir und Tokat. ² A. semnusta fehlt bislang in der Türkei (Kleinasien), ist aber von Zypern (ohne genau Lokalität) beschrieben und kommt auch in Nahen Osten vor. Der Verfasser kennt die Art bislang nur aus dem griechischen Teil Zyperns, aber auch von der Insel Samos, die unmittelbar an das türkische Festland grenzt. ³ A. scutellaris liegt dem Verfasser aus der Türkei (Umgebung Antalya und Mersin) vor, weiterhin von den griechischen Inseln Kos und Kreta. 4 A. ornata ist derzeit nicht aus der Türkei bekannt. Da sie aber aus Griechenland gemeldet wird, ist ein Vorkommen zumindest im europäischen Teil der Türkei denkbar. 6 Halsschild von der Mitte zur Basis gerade, kaum merklich verengt (Abb. 2). Vordertarsen nicht deutlich breiter als Mittel- und Hintertarsen. Aedoeagus wie in Abb. 11: ̎Öhrchen̎ an der Spitze der Paramerenplatte kleiner und stärker aus dem Flucht der Paramerenplatte heraustretend. Verbreitung: Südtürkei.................. A. rudolfina nov.sp. - Halsschild von der Mitte zur Basis gerundet verengt, flacher als von der Mitte nach vorn (Abb. 3). Vordertarsen nicht deutlich breiter als Mittel- und Hintertarsen. Aedoeagus wie in Abb. 12: ̎Öhrchen̎ an der Spitze Paramerenplatte grösser und weniger aus der Flucht der Paramerenplatte hervortretend. Verbreitung: Südtürkei......................................................................................................................... A. anneliesae nov.sp., Published as part of Esser, Jens, 2019, Neue Arten der Gattung Atomaria STEPHENS, 1830 aus der Türkei und dem Irak (Coleoptera, Cryptophagidae), pp. 55-62 in Linzer biologische Beiträge 51 (1) on pages 58-60, DOI: 10.5281/zenodo.3763644

Atomaria anneliesae nov.sp (Abb. 3, 12) T y p e n m a t e r i a l: Holotypus ♂♂: ̎TR Mersin (42), road Slifke - Gülnar, 1000 m / 36°20'37N, 33°35'17E (42), leg. 6.5.200 4, Brachat & Meybohm ̎ [cES]. Paratypen: 2♀♀ mit denselben Daten wie der Holotypus [cES]; 1 ♀♀ ̎TR Karahmanmaras (18) Pass N Tekir S Göksun, 1400 - 1550 m / 37°56'56N, 36°34'E (18), leg. 26.4.200 4, Brachat & Meybohm ̎ [cES]; 1♀ ̎ N37°57'20, E36°06'22, Türkei Adana (35/36), Eyüplü 1560 m, s Saimbeyli (n Kozan) / Tannen, Polsterpfl., leg. 27.4.200 5 (35/36) Brachat & Meybohm ̎ [cES]; 2♀♀ ̎ Turkey, Gölhisar, 120 km W Antalya, N36°57'40'', E29°57'53'', 7 km SW Altinyayla, P-trap, Hollow Quercus, 2009- 06-28, Nicklas Jansson / Mustafa Avci ̎ [cJA]; 1♂, 1♀ ̎ Turkey, Gölhisar, 120 km W Antalya, N36°57'40'', E29°57'53'', 7 km SW Altinyayla, P-trap, Hollow Quercus, 2009-10-04, Nicklas Jansson / Mustafa Avci ̎ [cJA]; 1♂ ̎TR-Antalya, W Kemer, s Hisar, No. 24, Quercus Carpinus, 36°44'02N, 30°26'23E, 2.IV.2002, V. Assing ̎ [cPÜ]; 1♂, 1♀ ̎TR-Muǧla, No. 19, SE Fethiye, Baba Daǧ, above Ovacik, 1450 m, 36°31'59N, 29°11'26E, 30.III.2002, V. Assing ̎ [cES, cPÜ], 1♀ ̎TR-Kahramanmaraş [34], Ahar Daǧı, 815 m, 10 km WNW Kahramanmaraş, 37°38'46N, 36°49'59E, 11.IV.2004, leg. V. Assing ̎ [cPÜ]; 2♂♂ ̎ Turquie, Isparta, Eǧridin - Çandin, 950 m, 6. V.75, Besuchet, Löbl ̎ [MHNG]. E t y m o l o g i e: Zum Dank an meine Mutter, Anneliese Esser, die mich besonders in jungen Jahren stets bei meinen entomologischen Entwicklungen unterstützte und mir den Rücken frei hielt. B e s c h r e i b u n g: Männchen, 1,6mm,rotbraun,Flügeldeckenmiteinem gemeinschaftlichen und schlecht begrenzten dunkelbraunen Fleck, der die Naht, die Basis und das apikale Drittel ausspart, Hinterflügel voll entwickelt. Mit mässig langer, aber recht dichter heller Behaarung, die einzelnen Haare erreichen knapp die Basis des folgenden Haares (vorderes Drittel der Flügeldecken). Beine von gewöhnlicher Länge, nicht auffallend kräftig. Fühler kräftig, Geisselglieder sehr unterschiedlich in der Länge: Glied 1 kurz und dick, wenig länger als breit, Glied 2 schmaler als das vorhergehende und länger als breit, Glied 3 ebenso, aber noch etwas schmaler, Glied 4 kurz, Glied 5 ähnlich Glied 3, Glied 6 kurz, Glied 7 kurz aber breiter als Glied 6, Glied 8 kugelig und wieder schmaler als Glied 7. Die Glieder der Keule (9 bis 11) sind schwächer quer (9 stärker quer als 10), Glied 11 gedrungen spindelförmig und asymmetrisch. Die Punktur des Kopfes ist fein und weitläufig (Abstand fast doppelt so gross wie der Punktdurchmesser), die des Halsschildes gröber und dichter (Abstand kleiner als der Durchmesser), die der Flügeldecken ähnlich grob und weitläufig wie die des Halsschildes (im ersten Flügeldeckendrittel: Abstand entspricht etwa dem Durchmesser). Die Flügeldecken sind zwischen der Punktur verloschen und kaum wahrnehmbar chagriniert. Der Kopf ist ohne Auffälligkeiten, die Augen von gewöhnlicher Grösse. Der Halsschild ist 1,55 mal so breit wie lang, seine Seitenränder schmal verflacht, von oben gleichzeitig sichtbar. Die Seiten des Halsschildes sind fast gleichmässig gerundet, von der Mitte nach vorn etwas stärker als nach hinten. Vorderwinkel stumpf. Seitenrandkante vollständig verdoppelt. Die Halsschildbasis ist auf ganzer Breite verflacht, der Hinterrand ist nach hinten flachbogig vorgezogen. Die Behaarung ist auf die Mitte des Halsschildes gerichtet. Die Flügeldecken sind oval und ohne Auffälligkeiten. Aedoeagus s. Abb. 12. B e m e r k u n g e n: Die Art ähnelt sehr der vorhergehenden und steht mit ihr sicher in engstem verwandtschaftlichen Verhältnis. Da beide Arten auch nebeneinander vorkommen, bestand längere Zeit Zweifel an der Eigenständigkeit der Taxa. Da aber zumindest der Bau des Halsschildes und der Paramerenplatte konstant unterschiedlich zu sein scheint, erscheint die Trennung gerechtfertigt. Unterscheidung von anderen, ähnlich gefärbten Arten, insbesondere A. ornata HEER, 1841, siehe Bestimmungsschlüssel., Published as part of Esser, Jens, 2019, Neue Arten der Gattung Atomaria STEPHENS, 1830 aus der Türkei und dem Irak (Coleoptera, Cryptophagidae), pp. 55-62 in Linzer biologische Beiträge 51 (1) on pages 57-58, DOI: 10.5281/zenodo.3763644

Atomaria rudolfina nov.sp. (Abb. 2, 11) T y p e n m a t e r i a l: Holotypus ♂♂: ̎TR Gaziantep (12), N Birecik, 360 m, Ostufer des Euphrat / 37°3'30N, 37°57'43E (12), leg. 24.4.200 4, Brachat & Meybohm ̎ [cES]. Paratypen: 2♀♀ ̎ Turkey, Gölhisar, 120 km W Antalya, N36°57'40'', E29°57'53'', 7 km SW Altinyayla, P-trap, Hollow Quercus, 2009-06-28, Nicklas Jansson / Mustafa Avci ̎ [cES, cJA]; 1♀ ̎ Turkey, Gölhisar, 120 km W Antalya, N36°57'40'', E29°57'53'', 7 km SW Altinyayla, P-trap, Hollow Quercus, 2009- 05-22, Nicklas Jansson / Mustafa Avci ̎ [cJA]; 1♀ ̎ Turkey, Mersin, 40 km N Gülnar, N36°30'22.5, E33°07'43.3, Kösecobanli / Tasdüstü, W-trap, Holow Quercus, 2006-06-24, Nicklas Jansson / Mustafa Avci ̎ [cJA]; 1♂ ̎ Turkey, Adiyaman, Komur, N 37°55'900, E38°28'352, 2011- 04-20, Pinaryayla & Karaman mezrasi, 30 km N, Hollow Quercus, trap, T. Kayis / M. Coskun / N. Jansson ̎ [cES]; 1♀ ̎N-Iraq, S Rawandoz, Akoian valley, ~ 1400 m, ~ 36°30'N, 44°36'E, 17.- 25.IV.2017, pitfall trap, leg. Reuter ̎ [cRE]; 1♀ ̎TR-Antalya, W Kemer, s Hisar, No. 24, Quercus Carpinus, 36°44'02N, 30°26'23E, 2.IV.2002, V. Assing ̎ [cPÜ]. E t y m o l o g i e: Zum Dank an meinen Vater, Rudolf Esser, der mich stets unterstützte und besonders in meiner Kindheit und Jugend mit mir unzählige Male auf Pirsch ging. B e s c h r e i b u n g: Männchen, 1,6mm,rotbraun,Flügeldeckenmiteinem gemeinschaftlichen und undeutlich begrenzten dunkelbraunen Fleck, der die Naht, die Basis und das apikale Drittel ausspart, Hinterflügel voll entwickelt. Mit kurzer, aber recht dichter heller Behaarung, die einzelnen Haare erreichen nicht die Basis des folgenden Haares (vorderes Drittel der Flügeldecken). Beine von gewöhnlicher länge, nicht auffallend kräftig. Fühler kräftig, Geisselglieder sehr unterschiedlich in der Länge: Glied 1 kurz und dick, wenig länger als breit, Glied 2 schmaler als das vorhergehende und länger als breit, Glied 3 ebenso, aber noch etwas schmaler, Glied 4 fast kugelig, Glied 5 ähnlich Glied 3, Glied 6 und 7 kugelig, wobei Glied 7 breiter ist, Glied 8 sehr klein und quer. Die Glieder der Keule (9 bis 11) sind schwächer quer (9) bis quer (10), Glied 11 gedrungen spindelförmig und asymmetrisch. Die Punktur des Kopfes ist recht fein und weitläufig (Abstand grösser als der Punktdurchmesser), die des Halsschildes gröber und dichter (Abstand kleiner als der Durchmesser), die der Flügeldecken wieder feiner und weitläufiger (im ersten Flügeldeckendrittel: Abstand grösser als der Durchmesser). Die Flügeldecken sind zwischen der Punktur fein chagriniert. Der Kopf ist ohne Auffälligkeiten, die Augen von gewöhnlicher Grösse. Der Halsschild ist 1,35 mal so breit wie lang, seine Seitenränder schmal, nach hinten breiter werdend verflacht, von oben gleichzeitig sichtbar. Die Seiten des Halsschildes sind etwas vor der Mitte flachbogig gewinkelt, davor fast gerade und deutlich nach vorn verengt, dahinter gerade und fast parallel nach hinten verengt. Vorderwinkel wenig stumpf. Seitenrandkante vorn verdoppelt. Die Halsschildbasis ist auf ganzer Breite verflacht, der Hinterrand ist nach hinten flachbogig vorgezogen. Die Behaarung ist auf die Mitte des Halsschildes gerichtet. Die Flügeldecken sind oval und ohne Auffälligkeiten. Aedoeagus s. Abb. 11. B e m e r k u n g e n: Die Art ähnelt sehr der folgenden und steht mit ihr sicher in engstem verwandtschaftlichen Verhältnis. Da beide Arten auch nebeneinander vorkommen, bestand längere Zeit Zweifel an der Eigenständigkeit der Taxa. Da aber zumindest der Bau des Halsschildes und der Paramerenplatte konstant unterschiedlich zu sein scheinen, erscheint die Trennung gerechtfertigt. Unterscheidung von anderen, ähnlich gefärbten Arten, insbesondere A. ornata HEER, 1841, siehe Bestimmungsschlüssel., Published as part of Esser, Jens, 2019, Neue Arten der Gattung Atomaria STEPHENS, 1830 aus der Türkei und dem Irak (Coleoptera, Cryptophagidae), pp. 55-62 in Linzer biologische Beiträge 51 (1) on pages 56-57, DOI: 10.5281/zenodo.3763644

Atomaria schuhi nov.sp. Type material: Holotype ♂: " Indonesia: W Java, Telaga Patengan, 1400 m, 2 km SE Rancabali (40 km SSW Bandung) lg. Schuh 6.8.1994 " [cES]. Paratypes: 1♀ with the same data as the holotype [cES]; 1♀ " Indonesia, W Java, Tangkuban Prahu, 10 km N Bandung, 1500-1700 m, lg. Schuh, 5.8.1994 " [cES]; 1♀ " Java: W Java, Cibodas, 50 km E Bogor, 1400 m, 3.-6.XI.1989, Agosti. Löbl, Burckhardt #2a" [MHNG]. Etymology: To the honour of Rudolf Schuh, who was the collector of the type material. Description: Male, 1,4 mm, reddish-brown, shiny, hind wings developed. Covered with pale decumbent hairs. Legs slender and pale. Antennae also pale and slender, club well separated but with segments 9, 10 and 11 relatively small, not broader than long. Pronotum broad, convex and with margin slightly angled in middle, broadest also in the middle. Base of pronotum depressed, puncture on surface strong but not dense. Elytra (basal third) with puncture as on pronotum. Elytra rounded, broadest in the basal third, much broader than pronotum. Comment: Atomaria schuhi nov.sp. (fig. 2) is similar to a few Palaearctic species, especially Atomaria horridula REITTER, 1877 (fig. 4) and A. incertula JOHNSON, 1971 (fig. 5) and the cosmopolitan Atomaria lewisi REITTER, 1877 (fig. 3). They all are separated by their well developed and broad antennal clubs. Distribution: Known only from West Java from mountain ranges north and south of Bandung. Further record of Atomaria STEPHENS, 1829 from the Oriental Region, Published as part of Esser, Jens, 2018, Two new Atomaria S, 1829 (Coleoptera: Cryptophagidae) from the Oriental Region with remarks on further Atomaria species, pp. 229-233 in Linzer biologische Beiträge 50 (1) on pages 230-233, DOI: 10.5281/zenodo.3985414

Atomaria bisulcata nov.sp. T y p e m a t e r i a l: Holotype ♂: " Iran, Azarbayian-e Sharqi, Osku: Amghan, 2100 m (Kuh-e Sahand), N37��49���38������ E46��16���15������, 08.08.2005, leg. Frisch & Serri [MFNB]. Paratypes: 2 ex. with the same data as the holotype [MFNB, cES]. E t y m o l o g y: The Latin word bisulcata means split due to the parameres which has a deep split. D e s c r i p t i o n: Male, 1,4 mm, hindwings absent, uniformly dark reddish-brown. Body stout and convex, with coarse and dense puncture, the distance of puncture on pronotum (disc) and elytra (anterior third) like the diameter. Covered with short yellowish pubescence (fig. 1). Anterior part of margin of the pronotum not visible from above, medium part of the basis moderately depressed. Sides moderately angled before the middle. Posterior angles nearly rectangular. Elytra oviform, puncture in the posterior third less coarse. Antennae slender, club 3-segmented but not well developed. Segments 9, 10 and 11 elongated, segments 3 to 8 decreasing in length but increasing in width. Segment longer and wider than 3, segment 1 longer and wider than 2 and slightly curved. Legs slender, all tarsi 5-segmented, anterior tarsi in male inconspicuous widened. Aedoeagus with a paramere with a deep split (fig. 2). C o m m e n t: More or less similar to a few species like A. fuscata (SCH��NHERR, 1808) or A. gibbula ERICHSON, 1846 but separated by the parameres (fig. 2). D i s t r i b u t i o n: Currently known from the type locality in Iran., Published as part of Esser, Jens, 2018, Atomaria bisulcata nov. sp. (Cryptophagidae) from Iran, pp. 235-237 in Linzer biologische Beitr��ge 50 (1) on pages 235-236, DOI: 10.5281/zenodo.3985418

Atomaria natmataunga nov.sp. Type material: Holotype ♂: "Mya, Chin state, barkspray, Natmataung NP, 2960 m, wnw Kanpetlet, 6.6.201 0, leg. D. Zimmermann " [NHMW]. Paratypes: 6 ex. with the same data as the holotype [NHMW, cES]. Etymology: Named after Nat Ma Taung, the local name for Mount Victoria, the highest mountain into the Chin hills range in Myanmar (Chin state). Description: Male, 1,4 mm, reddish-brow, elytra with a dark band, hind wings developed. Covered with pale decumbent hairs, pubescence coarse, beetle less shiny. Legs slender and pale. Antennae also pale and slender, club well separated but with segments 9, 10 and 11 relatively small. Segments 6 to 8 are globose, but segment 7 is larger than 6 and 8. Segment 1 slightly elongated and curved, longer and broader than segment 2. Segment 2 also elongated but smaller and shorter than segment 1. Segment 9 and 10 not transverse. Pronotum somewhat broader than long, broadest at the base. Base fully depressed, margins slightly rounded, nearly straight. Puncture moderately strong and dense. Elytra also with puncture moderately strong and dense, somewhat more as on pronotum. Elytra in the middle with a dark band, more ore less distinct, widened to the margins. Comment: Atomaria natmataunga nov.sp. (fig. 1) is well separated from all known species by the combination of general shape, antennae and coloration. The aeodeagus is typically for the genus. The tip of the medianlobus is very acute. Distribution: Known only from the Nat Ma Taung National Park (Chin hills range in Myanmar)., Published as part of Esser, Jens, 2018, Two new Atomaria S, 1829 (Coleoptera: Cryptophagidae) from the Oriental Region with remarks on further Atomaria species, pp. 229-233 in Linzer biologische Beiträge 50 (1) on page 230, DOI: 10.5281/zenodo.3985414

Atomaria linearis Stephens, 1830 3 Exemplare aus Conthey (m.5.1988, e.5.1988, e.7.1988). Die Art ist bei Löbl & Smetana (2007) für die Schweiz nicht aufgeführt. Auch in der Verbreitungskarte in der Fauna Europaea (2013) wird sie für die Schweiz nicht aufgeführt. A. linearis wurde aber schon im vorletzten Jahrhundert für die Schweiz nachgewiesen und zum Beispiel von Stierlin (1898) für Genf, Zürich und Schaffhausen gemeldet. Auch Merz (2012) meldet die Art für den Kanton Genf. In der Sammlung des Natur-Museums Luzern befinden sich zahlreiche Belege aus 19 verschiedenen Fundplätzen in der Schweiz aus den Kantonen BE, LU, OW, SH, SZ, VS und ZG. A. linearis ist wohl häufig und weit verbreitet in der Schweiz., Published as part of Herger, Peter & Germann, Christoph, 2017, Käfer aus Lichtfallenfängen in Conthey (VS) und in Seseglio (TI) (Coleoptera), pp. 87-92 in Entomo Helvetica 10 on page 89, DOI: 10.5169/seals-986074, http://zenodo.org/record/7958079, {"references":["Lobl I. & Smetana A. (eds.) 2007. Catalogue of Palaearctic Coleoptera. Volume 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, Danemark, 935 pp.","Fauna Europaea 2013. Version 2.6 (29. April 2013). URL: http // www. fauna-eu. org eingesehen im November 2016.","Stierlin G. 1898. Fauna Coleopterorum Helvetica. Teil II. Bolli und Boecherer, Schaffhausen, 662 pp.","Merz B. (ed.) 2012. Liste annotee des insectes (Insecta) du canton de Geneve. Instrumenta Biodiversitatis VIII, Geneve, 532 pp."]}

Atomaria lewisi Reitter, 1877 Erstnachweis für TI. 4 Ex. 21.– 30.06.1995, 2 Ex. 1.– 10.07.1995, 1 Ex. 11.– 20.07.1995, 1 Ex. 1.– 10.08.1995, 6 Ex 1.– 10.06.1996, 2 Ex. 21.– 30.06.1996, 2 Ex. 11.– 20.07.1996, 2 Ex. 21.– 31.07.1996. 1 Ex. 1.– 10.08.1996, 2 Ex. 21.– 31.05.1997, 1 Ex. 1.– 10.06.1997, 2 Ex. 21.– 31.08.1997, 1 Ex. 1.– 10.10.1997, alle det. Esser. Adventivart aus Ostasien, Erstnachweis in der Schweiz 1958 im Kanton SG (Linder 1968)., Published as part of Herger, Peter, Germann, Christoph, Uhlig, Manfred, Vogel, Jürgen, Geiser, Michael & Kirejtshuk, Alexander, 2015, Käfer aus Lichtfängen am Monte San Giorgio bei Serpiano, Kanton Tessin (Coleoptera), pp. 89-96 in Entomo Helvetica 8 on page 91, DOI: 10.5281/zenodo.7706118, {"references":["Linder A. 1968. 4. Beitrag zur Coleopteren-Fauna der Schweiz. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 41 (1 - 4): 211 - 232."]}

Atomaria fuscata Sch��nherr, 1808 LABRADOR: Happy Valley, near Churchill River, July 29, 2008, G.R. Pohl & D.W. Langor, at dusk, 15 m (1, MUN). NEW BRUNSWICK: Albert Co.: Mary���s Pt., August 23, 2003, August 12, 2004, C.G. Majka, ocean beach, in beach drift (4, CGMC); Charlotte Co.: Kent Island, Grand Manan archipelago, June 25, 2008, M. Steck, west beach (1, KIC); Kent Co.: Bouctouche, May 25, 2007, J.-P. Michaud, on decaying pig (1, UMNB); Westmorland Co.: Pointe-du-Chene, July 6, 1972, June-July, 1972, D.H. Murray (6, MMUE); Sackville, August 27, 1982, P. Maltais (1, CGMC). NEW- FOUNDLAND: St. Barbe, July 26, 1949, E. Palmen (1, MZHF); Piccadilly, July 7, 1949, E. Palmen (1, MZHF); Glide Lake, August 30, 1994, pitfall (1, MUN); Goulds, May 22, 1953, May 29, 1953, June 3, 1953, June 4, 1953, June 5, 1953, June 6, 1953, June 9, 1953, June 10, 1953, June 11, 1953, June 19, 1953 (52, AAFC); Goulds, June 9, 1953, R.F. Morris (1, AAFC); Kilbride, St. John���s, June 6, 1982, R.F. Morris (1, AAFC); Mount Pearl, November 10, 1995, Dave McKenzie (1, AAFC); Portugal Cove: Indian Meal Line, June 13, 1978, D. Larson (1, MUN); Portugal Cove: Indian Meal Line, May 16, 1979, May 21, 1979, July 1, 1979, July 31, 1982 (19, MUN); Portugal Cove: Indian Meal Line, July 1, 1979 (4, CFS); St John���s, July 12, 1974, July 16, 1974, R.F. Morris (2, AAFC); St John���s, October 18, 1977, D. Larson (1, MUN); St John���s, September, 1988, Biology 4150 (6, MUN); St John���s, Long Pond, May 29, 1981 (1, MUN); Bishop���s Falls, August 17, 1979, E. John (1, AAFC); South Pond near South Brook, June 27, 1980, Brennan & Larson, drift (1, MUN); Millville, Grandaddys Brook, June 16, 1979, Larson & Swales (1, MUN); 1 mi. east of Traytown, July 27, 1970, D.E. Bright (4, MUN); Burnt Cape, July 10���24, 2003, August 7���21, 2003, August 21-September 4, 2003, A.M. Hynes, shoreline, pitfall (8, MUN); Burnt Cape, July 10���24, 2003, July 24-August 7, 2003, A.M. Hynes, coastal meadow, pitfall (8, MUN); Burnt Cape, July 24-August 7, 2003, A.M. Hynes, cow parsnip patch, pitfall (7, MUN); Brunette Island, June 26���30, 1995, A.G. Raske, dense vegetation, pitfall (2, MUN). NOVA SCOTIA: Cape Breton Co.: Scatarie Island: E. Harbour, July 15, 2006, A. MacDonald, freshwater pond (2, CBU); Scatarie Island: NW Cove, August 8, 2005, A. MacDonald, beach, under wrack (1, CBU); Colchester Co.: 4 km N of Brookfield, June 23, 1995, J. Cook, mixed forest, uv light trap (1, JCC); Bible Hill, May 31, 2005, June 15���24, 2005, K.R. Aikens, pasture (3, CBU); Bible Hill, May 31, 2005, June 30, 2005, July 21, 2005, S.M. Townsend, pasture, sweeping (12, CBU); Bible Hill, May 27, 2004, D.B. McCorquodale, pasture, sweeping (1, CBU); Bible Hill, May 31, 2007, C.W. D���Orsay, pasture, sweeping (2, CBU); Masstown, June 7, 1990, T.D. Smith (1, NSNR); Cumberland Co.: Chignecto Game Sanctuary, July 30, 1988, E. Georgeson, uv light trap (1, NSMC); Oxford, August 16, 1986, August 12, 1988, E. Georgeson, uv light trap (4, NSMC); Wentworth Park, July 12, 1993, J. & T. Cook, car net (5, JCC); Westchester-Londonderry, July 20, 1992, S. & J. Peck, forest road, car net (12, JCC); Guysborough Co.: Malay Lake, 2���15 June, 1997, D.J. Bishop, red spruce, FIT (1, NSMC); Trafalgar, Liscomb Sanctuary, July 19, 1992, S. & J. Peck, car net (25, JCC); Halifax Co.: Middle Musquodoboit, September 3, 1988, E. Georgeson, uv light trap (1, NSMC); Halifax, July, 1988, P.M. Hammond, 5 specimens brachypterous (8, BMNH); Pt. Pleasant Park, July 5, 1990, S. Robertson, coniferous forest, stovepipe trap (1, NSMC); West Dover, September 7, 2003, C.G. Majka, salt marsh (3, CGMC); Hants Co.: Upper Rawdon, June 25, 2008, J. Renkema, highbush blueberry field, pitfall trap (4, CGMC); Kings Co.: Kentville, August 19, 2007, D.H. Webster, compost, moldy corncobs (1, DHWC); Lunenburg Co.: Chester, June 22, 1960, B. Wright (1, NSMC); Pictou Co.: McLellan���s Brook, August 5, 1988, E. Georgeson, uv light trap (1, NSMC); Seafoam, August 11, 2008, C.G. Majka, seashore, beach drift (1, CGMC); Queens Co.: Medway River, July 13, 1993, J. & T. Cook, car net (5, JCC); Ponhook Lake nr. Greenfield, July 13, 1993, J. Cook, uv light trap (1, JCC); Kejimkujik National Park, September 26���19 October, 1994, B. Wright, hemlock stand (1, NSMC); Shelburne Co.: Clyde River Rd, July 16, 1992, S. & J. Peck, forest, car net (3, JCC); Sebim Beach, July 19, 1993, J. & T. Cook (1, JCC); Yarmouth Co.: Carleton: Perry Rd., August 22, 1992, J. & F. Cook, mixed forest, car net (2, JCC); Carleton: Perry Rd., July 18, 1993, J. & T. Cook, car net (3, JCC). PRINCE EDWARD ISLAND: Queens Co.: Harrington, June 14, 2004, C. Noronha, barley field, pitfall trap (1, ACPE); Harrington, June 1, 1987, L.S. Thompson, potato field (3, ACPE); Harrington, September 8, 2005, M.E.M. Smith, barley & hedgerows, sweep (1, CGMC). We have examined the holotype of Atomaria saginata Casey 1900 deposited in the Smithsonian Institution and have found it to be identical in all respects to A. fuscata Sch��nherr. We therefore designate A. saginata (syn. n.) as a junior synonym of A. fuscata. Atomaria fuscata is newly recorded in Labrador, New Brunswick, insular Newfoundland, Nova Scotia, and Prince Edward Island (Figs 13, 17). Johnson et al. (2007) designated Atomaria ochracea Zimmerman, 1869 as a junior synonym of A. fuscata, and Atomaria lacustris Casey, 1900 and Atomaria pennsylvanica Casey, 1900 had previously been designated as junior synonyms of A. ochracea (by Leng 1920). Majka and Shaffer���s (2008) discussion of A. ochracea is attributable to A. fuscata. This species has been recorded in British Columbia, the District of Columbia, Idaho, Indiana, Maine, Manitoba, Maryland, Michigan, New Hampshire, New York, Ontario, Oregon, Pennsylvania, Rhode Island, Qu��bec, Virginia, Washington, and Wisconsin (LeConte 1869; Blatchley 1910; Leng 1920; Procter 1946; Hatch 1961; Oatman et al. 1964; Bousquet 1991; Downie and Arnett 1996; Chandler 2001; Sikes 2004). In the Palaearctic region it is known throughout Europe, across Siberia to the Russian Far East, south to Tibet and northern China, and in the Middle East in Turkey and Israel (Johnson et al. 2007). Hatch (1961) reported it as very common on herbiage and in grass clippings, compost, leaf litter, and humus. In Atlantic Canada it has been collected in mixed and coniferous forests, pastures, blueberry and potato fields, hedgerows, in coastal dunes and on ocean beaches under drift, by freshwater ponds, and in compost. Johnson (1993) reported that it, ���occurs in a range of habitats ��� grassland, wetland, and woodland ��� especially in damp situations, and has been collected in litter, moss, flood refuse, cut vegetation and compost heaps.��� Description: Dorsum and venter usually dark red-brown castaneous, pronotum occasionally darker than elytra. Legs reddish-testaceous. Pronotal punctures moderately fine and dense apically, becoming progressively coarser and denser basally; pronotum widest at middle, strongly narrowing basally and apically; base with a distinct transverse impression, the middle third of the basal margin with a discernable, fine marginal bead. Elytral punctures very fine and sparse; interspaces between punctures 3.0+ times the diameter of punctures. Setae the length of the interspaces, appressed (Fig. 5). Antennae: antennomere 1 swollen, longer and wider than 2; 3 as long as or longer than 2 but more slender; 4���8 small and somewhat bead-like; antennal club distinct; antennomeres 9 and 10 quadrate, not discernibly transverse (Fig. 1.4). Body: width/length ratio, 0.47; length, 1.4���1.7 mm., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on pages 48-50, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Casey TL (1900) Revision of the North American Cryptophagidae. Journal of the New York Entomological Society 8: 75 - 128.","Johnson C, Otero JC, Leschen RAB (2007) Cryptophagidae Kirby, 1837. In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Volume 4: Elateroidea, Derontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, Denmark, 513 - 531.","Leng CW (1920) Catalogue of the Coleoptera of America north of Mexico. John D. Sherman, Jr., Mount Vernon, New York, 470 pp.","Majka CG, Shaffer F (2008) Beetles (Coleoptera) in the diet of Piping Plovers in the Iles de la Madeleine, Quebec, Canada. Wader Study Group Bulletin, 115 (2): 77 - 83.","LeConte JL (1869) Synonymical notes on Coleoptera of the United States, with descriptions of new species, from the mss. of the late Dr. C. Zimmerman. Transactions of the American Entomological Society 2: 243 - 259.","Blatchley WS (1910) An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana. The Nature Publishing Company, Indianapolis, Indiana, 1386 pp.","Procter W (1946) Biological Survey of the Mount Desert Region, Part VII: The Insect Fauna. Wistar Institute of Anatomy and Biology, Philadelphia, Pennsylvania, 566 pp. http: // www. biodiversitylibrary. org / item / 29157 [accessed 11. XI. 2009]","Hatch MH (1961) The beetles of the Pacific Northwest, Part III: Pselaphidae and Diversicornia I. University of Washington Press, Seattle, Washington, 503 pp.","Oatman ER, Legner EF, Brooks RF (1964) An ecological study of arthropod populations on apple in northeastern Wisconsin: insect species present. Journal of Economic Entomology 57 (6): 978 - 983.","Bousquet Y (1991) Family Cryptophagidae: silken fungus beetles. In: Bousquet Y (Ed) Checklist of Beetles of Canada and Alaska. Agriculture Canada: Research Branch Publication 1861 / E, 221 - 223. http: // www. canacoll. org / Coleo / Checklist / PDF % 20 files / CRYPTO- PHAGIDAE. pdf [accessed 11. XI. 2009]","Downie NM, Arnett RH, Jr (1996) The Beetles of Northeastern North America. Sandhill Crane Press. Gainsville, Florida, 1721 pp.","Chandler DS (2001) University of New Hampshire Insect and Arachnid Collections. http: // insectcoll. unh. edu / [accessed 11. XI. 2009]","Sikes DS (2004) The beetle fauna of Rhode Island: an annotated checklist. Rhode Island Natural History Survey, Volume 3. Kingston, Rhode Island, 296 pp.","Johnson C (1993) Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Biological Records Centre, Natural Environmental Research Council, Institute of Terrestrial Ecology, Huntingdon, Cambridgeshire, Great Britain. 91 pp. http: // nora. nerc. ac. uk / 7280 / 1 / Cryptophagidae-Atomarinae. pdf [accessed 11. XI. 2009]"]}

Atomaria testacea Stephens, 1830 NEW BRUNSWICK: Albert Co.: Mary���s Pt., August 12, 2004, C.G. Majka, coastal field, in compost (4, CGMC). NOVA SCOTIA: Cape Breton Co.: Scatarie Island: Savage Cove, August 10, 2005, K.R. Aikens, beach, under wrack (1, CBU); Colchester Co.: Bible Hill, May 31, 2005, June 14, 2005, K.R. Aikens, pasture, sweeping (4, CBU); Bible Hill, May 31, 2005, July 21, 2006, S.M. Townsend, sweeping (2, CBU); Bible Hill, June 3, 2007, June 15, 2007, August 1, 2007, C.W. D���Orsay, pasture, sweeping (5, CBU); Debert, June 19, 1993, J. Ogden (1, NSNR); Guysborough Co.: Dayspring Lake, 2���15 June, 1997, D.J. Bishop, red spruce, FIT (1, NSMC); Malay Lake, 14 May-2 June, 1997, 1���16 July, 1997, D.J. Bishop, red spruce (mature), FIT (2, NSMC); Seloam Lake, 14 May-2 June, 1997, 2���15 June, 1997, D.J. Bishop, red spruce, FIT (2, NSMC); Halifax Co.: Abraham���s Lake, 1���16 July, 1997, D.J. Bishop, red spruce (old), FIT (1, NSMC); Anti Dam Lake, 1���16 July, 1997, D.J. Bishop, black spruce (90 years), FIT (1, NSMC); Antrim, June 9, 2005, J. Gordon, FIT (1, NSNR); Burnside, May 19, 2004, C. Cormier, field, on dead pig (1, SMU); Campbell Hill, 15��� 30 June, 1997, D.J. Bishop, red spruce (mature), FIT (1, NSMC); Long Lake, May 25, 2002, C.G. Majka, along stream (1, CGMC); Pockwock Lake, 14 May-2 June, 1997, D.J. Bishop, red spruce (mature), FIT (1, NSMC); Pogwa Lake, 15���30 June, 1997, D.J. Bishop, red spruce, FIT (1, NSMC); Pt. Pleasant Park, July 2, 2001, July 7, 2001, August 18, 2001, June 9, 2002, June 18, 2002, July 23, 2002, C.G. Majka, red spruce forest, boggy area (6, CGMC); Sandy Lake, 16���29 July, 1997, D.J. Bishop, red spruce (120+ years), FIT (1, NSMC); south-end Halifax, May 26, 2001, May 20, 2002, June 21, 2002, C.G. Majka, open area (3, CGMC); south-end Halifax, August 22, 2002, C.G. Majka, compost (1, CGMC); Kings Co.: Kentville, August 10, 2005, July 9, 2006, August 19, 2007, D.H. Webster, compost (11, DHWC); Kentville, August 12, 2007, D.H. Webster, at light (1, DHWC). Atomaria testacea is newly recorded in New Brunswick, Nova Scotia, and Canada (Fig. 15). In North America it has previously been recorded from southern California (Leng 1920) under the name Atomaria ruficornis Marsham, 1802, synonymized by Johnson et al. (2007). LeConte (1869, 258) reported a species from South Carolina under the name of Atomaria testacea Zimmerman; however, as Bousquet (1989) makes clear, this refers to Cryptophagus ferrugineus Sahlberg. In the Palaearctic region it is found throughout Europe, in Algeria, Morocco, and Tunisia in North Africa, and in Asia in Turkey, Kazhakhstan, Mongolia, and western Siberia (Johnson et al. 2007). In Atlantic Canada it has been found in red spruce forests, boggy areas and marshes, in pastures, fields, meadows, and open areas, on beaches under wrack, in a ravine, and in compost. Johnson (1993) reported that it was, ���mainly a grassland species which has been recorded around farms, gardens and parks, but it often occurs in other habitats. It has been collected in man-made heaps of refuse, especially cut vegetation, grass, hay, haystack bottoms and compost, more rarely in dung or dung heaps.��� Description: Head and prontotum piceous. Elytra piceous basally, gradually becoming paler in the apical half. Venter: pro-, meso-, and meta-sterna piceous; abdominal sterna and legs dark testaceous. Head finely, moderately sparsely, punctate. Pronotum and elytra moderately finely and densely punctate, interspaces 1.0���1.5 times the diameter of punctures, elytral punctures becoming finer apically; pronotum widest at midpoint, constricted towards base and apex; base with only a very shallow transverse impression (Fig. 9). Antennae: antennomere 1 swollen, slightly curved and expanded at apex; 2 smaller than 1 but also apically expanded; 3 almost as long as 2 and only slightly more slender; 4���8 short and somewhat bead-like; antennal club distinct; antennomeres 9 and 10 transverse (Fig. 1.8). Body: width/length ratio, 0.46; length, 1.3���1.6 mm. Qu��bec Newfoundland Gaspe Prince Edward Cape Breton Island Island New Brunswick Haine Nova Atomaria apicalis Scotia Atomaria nigrirostris Figure ��6. Distribution of Atomaria apicalis and Atomaria nigrirostris in Atlantic Canada., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on pages 54-56, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Leng CW (1920) Catalogue of the Coleoptera of America north of Mexico. John D. Sherman, Jr., Mount Vernon, New York, 470 pp.","Johnson C, Otero JC, Leschen RAB (2007) Cryptophagidae Kirby, 1837. In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Volume 4: Elateroidea, Derontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, Denmark, 513 - 531.","LeConte JL (1869) Synonymical notes on Coleoptera of the United States, with descriptions of new species, from the mss. of the late Dr. C. Zimmerman. Transactions of the American Entomological Society 2: 243 - 259.","Bousquet Y (1989) A review of the North American genera of Cryptophaginae (Coleoptera: Cryptophagidae). The Coleopterists Bulletin 43 (1): 1 - 17.","Johnson C (1993) Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Biological Records Centre, Natural Environmental Research Council, Institute of Terrestrial Ecology, Huntingdon, Cambridgeshire, Great Britain. 91 pp. http: // nora. nerc. ac. uk / 7280 / 1 / Cryptophagidae-Atomarinae. pdf [accessed 11. XI. 2009]"]}

Atomaria apicalis Erichson, 1846 NEWFOUNDLAND: Portugal Cove: Indian Meal Line, June 2, 1979, July 1, 1979, June 22, 1981, September 1, 1981, July 21, 1982 (14, MUN); Gander, July-August, 1998, D. Russell (2, MUN); South Pond near South Brook, June 27, 1980, Brennan & Larson, drift (2, MUN). NEW BRUNSWICK: Albert Co.: Mary���s Pt., September 8, 2002, C.G. Majka, old field, in compost (7, CGMC); Westmorland Co.: Figures 8�������. 8 Dorsal habitus photograph of Atomaria pusilla (Paykull) 9 Dorsal habitus photograph of Atomaria testacea Stephens ��0 Dorsal habitus photograph of Atomaria nigrirostris Stephens ���� Dorsal habitus photograph of Atomaria wollastoni Sharp. Figure ��2. Adeagus illustrations of Atomaria species. ��2.�� Atomaria apicalis ��2.2 Atomaria ephippiata ��2.3 Atomaria fuscata ��2.4 Atomaria lederi ��2.5 Atomaria lewisi ��2.6 Atomaria pusilla ��2.7 Atomaria testacea ��2.8 Atomaria nigrirostris ��2.9 Atomaria wollastoni. Adapted from Sj��berg (1947), Lohse (1967), Johnson (1970, 1971) with additional material. Note: no illustration of the adeagus of A. distincta was available. Pointe-du-Chene, July 6, 1972, August 24, 1972, D.H. Murray (5, MMUE); NOVA SCOTIA: Halifax Co.: Halifax, September 7, 1972, D.H. Murray (2, MMUE); Halifax, July 1988, P.M. Hammond (1, BMNH); Lunenburg Co.: Bridgewater, June 30, 1965, B. Wright, red oak, window trap (1, NSMC); Queens Co.: Medway River, July 13, 1993, J. & T. Cook, car net (2, JCC). Atomaria apicalis is newly recorded in insular Newfoundland, New Brunswick, Nova Scotia, and Atlantic Canada (Fig. 16). We have examined the holotype of Atomaria ovalis Casey 1900 deposited in the Smithsonian Institution and have found it to be identical in all respects to A. apicalis Erichson. We therefore designate A. ovalis syn. n. as a junior synonym of A. apicalis. In North America this species has previously been reported from Connecticut, Indiana, Iowa, Maine, Manitoba, Massachusetts, New Hampshire, New York, Qu��bec (Blatchley 1910; Britton 1920; Leng 1920; Procter 1946; Bousquet 1991; Downie and Arnett 1996; Chandler 2001). It is widely distributed in the Palaearctic region being found throughout Europe and North Africa, east across Siberia to the Russian Far East, across the Middle East, to Central Asia, Mongolia, and Fujian in China (Johnson et al. 2007). Johnson (1993) reported that it was, ���a grassland species which has been recorded especially around farms, gardens and parks but often occurs in other habitats. It has been collected in man-made heaps of refuse, especially cut vegetation, grass, compost and dung heaps, flood debris, and more rarely on carrion and rotting fungi.��� Description: A variable species; head, pronotum, and elytra piceous to castaneous or (more rarely) black; elytra sometimes paler in apical 2/5. Legs, prosternum, and abdominal sterna reddish-testaceous. Pronotum weakly constricted from middle to base; from middle to apex even more strongly. Rarely, in males the basal half of the pronotum is parallel-sided, or very weakly constricted basally. Punctures of the pronotum and elytra very variable; on the base of the pronotum, interspaces between punctures �� 1.0 times the diameter of puctures. Punctation of elytra similar to that of pronotum, or finer and shallower, but always less dense. Distance between furrows of prosternal process approximately half the width of the prosternal process. Body strongly convex in cross-section (Fig. 2). Antennae: antennomere 1 swollen, slightly longer than 2; 3 as long as 2 but narrower; 4���8 short and bead-like; club distinct, antennomeres 9 and 10 slightly transverse (Fig. 1.1). Body: width/length ratio, 0.41; length, 1.3���1.6 mm., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on pages 43-45, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Sjoberg O (1947) Die Ergebnisse einer Genitaluntersuchung der nordischen Arten der Gattung Atomaria Steph. (Col., Cryptophagida). Entomologisk Tidskrift 69: 91 - 119.","Lohse GA (1967) Fam. Cryptophagidae. In: Freude H, Harde KW, Lohse GA. (Eds) Die Kafer Mitteleuropas, Band 7: Clavicornia. Goecke & Evers, Krefeld, Germany, 110 - 157.","Johnson C (1970) Three new species of Atomaria Stephens (Coleoptera: Cryptophagidae) from the eastern Palaearctic. Notulae Entomologicae 50: 112 - 116.","Johnson C (1971) Atomariinae (Col., Cryptophagidae) from the northern parts of the Indian sub-continent with descriptions of seven new species. Entomologists Monthly Magazine 107 [1970]: 224 - 232.","Casey TL (1900) Revision of the North American Cryptophagidae. Journal of the New York Entomological Society 8: 75 - 128.","Blatchley WS (1910) An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana. The Nature Publishing Company, Indianapolis, Indiana, 1386 pp.","Britton WE (1920) Check-list of the Insects of Connecticut. State Geological and Natural History Survey Bulletin of Connecticut 31. Hartford, Connecticut. 397 pp.","Leng CW (1920) Catalogue of the Coleoptera of America north of Mexico. John D. Sherman, Jr., Mount Vernon, New York, 470 pp.","Procter W (1946) Biological Survey of the Mount Desert Region, Part VII: The Insect Fauna. Wistar Institute of Anatomy and Biology, Philadelphia, Pennsylvania, 566 pp. http: // www. biodiversitylibrary. org / item / 29157 [accessed 11. XI. 2009]","Bousquet Y (1991) Family Cryptophagidae: silken fungus beetles. In: Bousquet Y (Ed) Checklist of Beetles of Canada and Alaska. Agriculture Canada: Research Branch Publication 1861 / E, 221 - 223. http: // www. canacoll. org / Coleo / Checklist / PDF % 20 files / CRYPTO- PHAGIDAE. pdf [accessed 11. XI. 2009]","Downie NM, Arnett RH, Jr (1996) The Beetles of Northeastern North America. Sandhill Crane Press. Gainsville, Florida, 1721 pp.","Chandler DS (2001) University of New Hampshire Insect and Arachnid Collections. http: // insectcoll. unh. edu / [accessed 11. XI. 2009]","Johnson C, Otero JC, Leschen RAB (2007) Cryptophagidae Kirby, 1837. In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Volume 4: Elateroidea, Derontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, Denmark, 513 - 531.","Johnson C (1993) Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Biological Records Centre, Natural Environmental Research Council, Institute of Terrestrial Ecology, Huntingdon, Cambridgeshire, Great Britain. 91 pp. http: // nora. nerc. ac. uk / 7280 / 1 / Cryptophagidae-Atomarinae. pdf [accessed 11. XI. 2009]"]}

Atomaria lederi Johnson, 1970 NOVA SCOTIA: Colchester Co.: Five Islands Park, September 6, 1988, E. Georgeson, uv light trap (2, NSMC); Cumberland Co.: Oxford, August 11, 1988, E. Georgeson, uv light trap (1, NSMC); Guysborough Co.: Dayspring Lake, 2���15 June, 1997, 1���16 July, 1997, D.J. Bishop, red spruce, FIT (2, NSMC); George Lake, 14 May-2 June, 1997, D.J. Bishop, red spruce (young), FIT (1, NSMC); Malay Lake, 14 May-2 June, 1997, 15���30 June, 1997, 1���16 July, 1997, D.J. Bishop, red spruce, FIT (8, NSMC); Melopseketch Lake, 1���16 July, 1997, D.J. Bishop, red spruce (young), FIT (1, NSMC); Seloam Lake, 14 May-2 June, 1997, 2���15 June, 1997, 15���30 June, 1997, D.J. Bishop, red spruce, FIT (3, NSMC); Halifax Co.: Big St. Margarets Bay, 2���15 June, 1997, 29 July-13 August, 1997, D.J. Bishop, red spruce, FIT (2, NSMC); Grassy Lake, 15���30 June, 1997, 1���16 July, 1997, D.J. Bishop, red spruce, FIT (4, NSMC); Moser Lake, 15���30 June, 1997, 1���16 July, 1997, D.J. Bishop, red spruce, FIT (2, NSMC); Pockwock Lake, 29 July-13 August, 1997, D.J. Bishop, red spruce (mature), FIT (1, NSMC); Porters Lake, September 2, 1993, B. Wright (2, NSMC); Pt. Pleasant Park, June 7, 2001, June 27, 2001, July 7, 2001, May 9, 2002, C.G. Majka, boggy area, (5, CGMC); Sandy Lake, 2���15 June, 1997, 29 July-13 August, 1997, D.J. Bishop, red spruce (120+ years), FIT (2, NSMC); Ten Mile Lake, 15���30 June, 1997, D.J. Bishop, red spruce, FIT (1, NSMC); Hants Co.: Armstrong Lake, 14 May-2 June, 1997, D.J. Bishop, red spruce (75 years), FIT (2, NSMC); Leminister, 15���30 June, 1997, D.J. Bishop, red spruce/eastern hemlock, FIT (1, NSMC); Little Armstrong Lake, 14 May-2 June, 1997, D.J. Bishop, red spruce (75 years), FIT (1, NSMC); Lunenburg Co.: Bridgewater, May 27, 1965, B. Wright, red oak, window trap (1, NSMC); Card Lake, 15���30 June, 1997, D.J. Bishop, red spruce/eastern hemlock (old), FIT (1, NSMC); Pictou Co.: Lorne, 15���30 June, 1997, D.J. Bishop, red spruce/ eastern hemlock (mature), FIT (1, NSMC); McLellan���s Brook, July 29, 1988, E. Georgeson, uv light trap (1, NSMC); Shelburne Co.: Barrington, May 18, 1966, collector not noted, attic of old meeting house (1, NSMC). Atomaria lederi is newly recorded in North America (Fig. 14). The reference to North America in Johnson et al. (2007) refers to some of the above records. In the Palaearctic region this species has been recorded from central and northern Russia, east across Siberia to the Russian Far East, and south to Mongolia, North Korea, and northern China (Liaoning) (Johnson et al. 2007). In Nova Scotia it has primarily been found in mixed red spruce (Picea rubens Sarg.) and eastern hemlock (Tsuga canadensis (L.) Carr.) (Pinaceae) forests of various ages, occasionally in boggy areas in such forests, on Spiraea alba Duroi (Rosaceae), and balsam fir (Abies balsamea (L.) Mill) (Pinaceae). Description: Dorsum and venter reddish-testaceous throughout, including legs. Head, pronotum, and elytra moderately coarsely, moderately densely punctate; interspaces 1.0���1.5 times the diameter of punctures; elytral setae the length of the interspaces, appressed. Pronotum widest at middle, narrowing slightly to base and more strongly to apex (Fig. 6). Antennae: antennomere 1 quadrate, swollen; 2 the same length and almost the same width as 1; 3 slightly shorter and narrower; 4���8 similar in size and bead-like; club distinct; antennomeres 9 and 10 quadrate, not transverse (Fig. 1.5). Body: width/length ratio, 0.43; length, 1.2���1.4 mm., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on pages 50-52, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Johnson C (1970) Three new species of Atomaria Stephens (Coleoptera: Cryptophagidae) from the eastern Palaearctic. Notulae Entomologicae 50: 112 - 116.","Johnson C, Otero JC, Leschen RAB (2007) Cryptophagidae Kirby, 1837. In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Volume 4: Elateroidea, Derontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, Denmark, 513 - 531."]}

Atomaria lewisi Reitter, 1877 NEW BRUNSWICK: Albert Co.: Mary���s Pt., September 8, 2002, C.G. Majka, old field, in compost (3, CGMC); NOVA SCOTIA: Cumberland Co.: Oxford, June 16, 1986, E. Georgeson (1, NSNR); Wentworth Park, July 12, 1993, J. & T. Cook, car net (5, JCC); Halifax: Pt. Pleasant Park, August 10, 2001, August 18, 2001, C.G. Majka, coniferous forest, boggy area, Spiraea alba (3, CGMC); Kings Co.: Kentville, August 10, 2005, D.H. Webster, compost, moldy corncobs (5, DHWC); Queens Co.: Caledonia, July 25, 1992, J. & F. Cook, interior mixed forest, car net (4, JCC); Medway River, July 13, 1993, J. & T. Cook, car net (1, JCC); Ponhook Lake nr. Greenfield, July 13, 1993, J. Cook, uv light trap (1, JCC); Richmond Co.: Louisdale, November 12, 1991, M. Boudreau (4, NSMC); Shelburne Co.: Clyde River Rd, July 16, 1992, S. & J. Peck, forest, car net (1, JCC); Yarmouth Co.: Carleton, Perry Rd., August 22, 1992, J. & F. Cook, mixed forest, car net (7, JCC); Carleton, Perry Rd., July 18, 1993, J. & T. Cook, car net (3, JCC); Coldstream Rd. E of Quinan, July 19, 1993, J. & T. Cook, car net (1, JCC); Hebron, June 4, 1999, J. Cook, on Hawthorne (1, JCC); north of North Kemptville, August 23, 1992, J. & F. Cook, car net (6, JCC); Wellington, July 11���12, 1997, J. Cook, uv light trap (3, JCC); Yarmouth-Shelburne Co.: Oak Park Rd., August 27, 1992, J. & F. Cook, car net (9, JCC). We have examined the holotype of Atomaria curtula Casey 1900 deposited in the Smithsonian Institution and have found it to be identical in all respects to A. lewisi Reitter. We therefore designate A. curtula syn. n. as a junior synonym of A. lewisi. Atomaria lewisi is newly reported in New Brunswick, Nova Scotia, Canada, and North America (Fig. 14). Under the name A. curtula it was reported from Connecticut, Ontario (Casey 1900; Leng 1920; Bousquet 1991; Downie and Arnett 1996), and Maine (Procter 1946). In the Palaearctic region it is found throughout Europe, the Russian Far East, Central Asia, Mongolia, China, Japan, and North Korea, south to Bhutan and northern India (Johnson et al. 2007). Lohse (1967) called it an adventive species in Europe, and Johnson et al. (2007) referred to it as cosmopolitan. In Nova Scotia it has been found in coniferous and mixed forests, in boggy areas, on Craetegus sp. (Rosaceae), and on compost. Johnson (1993) reported that it was, ���a grassland species which has been recorded especially around farms, gardens and parks but it often also occurs in other habitats. It has been collected in man-made heaps of refuse, especially cut vegetation, hay, grass, and compost.��� Description: Dorsum and venter brownish-yellow or reddish-yellow throughout, including legs. Head moderately densely, moderately finely punctate. Pronotum moderately densely and moderately coarsely punctate; interspaces 0.5 times the diameter of punctures; base with a distinct flattened depression. Elytra somewhat more finely and sparsely punctate; interspaces 1.0���1.5 times the diameter of punctures; elytral margins strongly rounded; setae rather long and prominently erect giving a ���bristling��� appearance (Fig. 7). Antennae: antennomere 1 long, swollen, and curved, apex nearly 1.5���2.0 times the width of base, 1.5 times longer than 2; 3 the same length as 2 but more slender; 4���8 almost equal in length and bead-like; club distinct; antennomeres 9 and 10 slightly transverse (Fig. 1.6). Body: width/length ratio, 0.46; length, 1.3���1.6 mm., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on pages 52-53, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Casey TL (1900) Revision of the North American Cryptophagidae. Journal of the New York Entomological Society 8: 75 - 128.","Leng CW (1920) Catalogue of the Coleoptera of America north of Mexico. John D. Sherman, Jr., Mount Vernon, New York, 470 pp.","Bousquet Y (1991) Family Cryptophagidae: silken fungus beetles. In: Bousquet Y (Ed) Checklist of Beetles of Canada and Alaska. Agriculture Canada: Research Branch Publication 1861 / E, 221 - 223. http: // www. canacoll. org / Coleo / Checklist / PDF % 20 files / CRYPTO- PHAGIDAE. pdf [accessed 11. XI. 2009]","Downie NM, Arnett RH, Jr (1996) The Beetles of Northeastern North America. Sandhill Crane Press. Gainsville, Florida, 1721 pp.","Procter W (1946) Biological Survey of the Mount Desert Region, Part VII: The Insect Fauna. Wistar Institute of Anatomy and Biology, Philadelphia, Pennsylvania, 566 pp. http: // www. biodiversitylibrary. org / item / 29157 [accessed 11. XI. 2009]","Johnson C, Otero JC, Leschen RAB (2007) Cryptophagidae Kirby, 1837. In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Volume 4: Elateroidea, Derontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, Denmark, 513 - 531.","Lohse GA (1967) Fam. Cryptophagidae. In: Freude H, Harde KW, Lohse GA. (Eds) Die Kafer Mitteleuropas, Band 7: Clavicornia. Goecke & Evers, Krefeld, Germany, 110 - 157.","Johnson C (1993) Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Biological Records Centre, Natural Environmental Research Council, Institute of Terrestrial Ecology, Huntingdon, Cambridgeshire, Great Britain. 91 pp. http: // nora. nerc. ac. uk / 7280 / 1 / Cryptophagidae-Atomarinae. pdf [accessed 11. XI. 2009]"]}

Atomaria pusilla (Paykull, 1798) NOVA SCOTIA: Cumberland Co.: Wentworth Park, July 12, 1993, J. & T. Cook, car net (2, JCC); Yarmouth Co.: Carleton: Perry Rd., July 18, 1993, J. & T. Cook, car net (1, JCC); Coldstream Rd. E of Quinan, July 19, 1993, J. & T. Cook, car net (2, JCC). NEW BRUNSWICK: Albert Co.: Mary���s Pt., August 9, 2002, C.G. Majka, coniferous forest, decaying Russula virescens (1, CGMC); Westmorland Co.: Shediac, September 30, 1972 (1, MMUE). Atomaria pusilla is newly recorded in New Brunswick and Nova Scotia (Fig. 15). In North America it has been recorded in Alaska, British Columbia, Connecticut, Idaho, Indiana, New York, Ohio, Oregon, Qu��bec, and Washington (Blatchley 1910; Britton 1920; Notman 1920; Hatch 1961; Bousquet 1991; Williams et al. 1995; Downie and Arnett 1996). Blatchley (1910) referred to it as a ���European��� species and Bousquet (1991) listed it as ���probably introduced��� in North America. In the Palaearctic region it is found throughout Europe (including Turkey), across North Africa, and in central Asia in Iran, Afghanistan, and Kazhakhstan, northeast to Mongolia and the Russian Far East (Johnson et al. 2007). Blatchley (1910) reported it from the borders of a sphagnum marsh and in other damp vegetable debris. In New Brunswick it was found in a coniferous forest on a decaying gill fungus. Johnson (1993) reported that it was, ���a grassland species which is found especially around farms and gardens and only rarely elsewhere, It has been collected in man-made heaps of refuse, cut vegetation, hay, grass, and compost heaps. Description: Body (dorsally and ventrally) uniformly reddish-yellow. Antennae and legs uniformly yellow-colored. Head very finely and sparsely punctate. Pronotum widest at midpoint, constricted basally and apically. Pronotum moderately finely and closely punctate, interspaces 1.0 times the diameter of punctures; pronotal base with a complete fine marginal line running along the bottom of the basal impression. Elytral punctation similar to pronotum (Fig. 8). Antennae: antennomere 1 short, straight, not or slightly expanded at apex, slightly longer than wide, approximately as long as antennomere 2; 3 slightly shorter than 2; 4���8 short and bead-like; antennal club distinct; antennomeres 9 and 10 transverse (Fig. 1.8). The smallest species of the genus; body width/length ratio, 0.40; length, 1.0���1.2 mm., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on page 54, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Blatchley WS (1910) An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana. The Nature Publishing Company, Indianapolis, Indiana, 1386 pp.","Britton WE (1920) Check-list of the Insects of Connecticut. State Geological and Natural History Survey Bulletin of Connecticut 31. Hartford, Connecticut. 397 pp.","Notman H (1920) Coleoptera collected at Windsor, Broome Co., N. Y., 26 May to 5 June, 1918, with notes and descriptions. Journal of the New York Entomological Society 28 (2): 178 - 194.","Hatch MH (1961) The beetles of the Pacific Northwest, Part III: Pselaphidae and Diversicornia I. University of Washington Press, Seattle, Washington, 503 pp.","Bousquet Y (1991) Family Cryptophagidae: silken fungus beetles. In: Bousquet Y (Ed) Checklist of Beetles of Canada and Alaska. Agriculture Canada: Research Branch Publication 1861 / E, 221 - 223. http: // www. canacoll. org / Coleo / Checklist / PDF % 20 files / CRYPTO- PHAGIDAE. pdf [accessed 11. XI. 2009]","Williams RN, Ellis MS, Fickle DS (1995) Insects in the Killbuck Marsh Wildlife Area: 1993 Survey. Ohio Journal of Science 95 (3): 226 - 232.","Downie NM, Arnett RH, Jr (1996) The Beetles of Northeastern North America. Sandhill Crane Press. Gainsville, Florida, 1721 pp.","Johnson C, Otero JC, Leschen RAB (2007) Cryptophagidae Kirby, 1837. In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Volume 4: Elateroidea, Derontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, Denmark, 513 - 531.","Johnson C (1993) Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Biological Records Centre, Natural Environmental Research Council, Institute of Terrestrial Ecology, Huntingdon, Cambridgeshire, Great Britain. 91 pp. http: // nora. nerc. ac. uk / 7280 / 1 / Cryptophagidae-Atomarinae. pdf [accessed 11. XI. 2009]"]}

Atomaria distincta Casey, 1900 NEW BRUNSWICK: Westmorland Co.: Moncton, July 6, 1972, D.H. Murray (1, MMUE). NOVA SCOTIA: Halifax Co.: West Dover, September 7, 2003, C.G. Majka, salt marsh (4, CGMC). Atomaria distincta is newly recorded in New Brunswick, Nova Scotia, and Canada (Fig. 14). This Nearctic species has previously been reported from the District of Columbia, Massachusetts, New York, and Qu��bec (Leng 1920; Downie and Arnett 1996). Description: Head piceous, finely sparsely punctate. Pronotum piceous, finely sparsely punctate; pronotum widest at or slightly ahead of midpoint, not appreciably flattened across base; finely sparsely punctate; interspaces between punctures 1.5���2.0 times the diameter of punctures. Scutellum black. Basal 3/5 of elytra black, becoming abruptly yellow in apical 2/5, the yellow portion extending slightly anterad in the region of the suture. Punctation very fine and sparse; interspaces between punctures 2.0���3.0 times the diameter of punctures; elytral setae very fine and short, shorter than interspace intervals and appressed. Legs, prosternum, and abdominal sterna reddishtestacous; sterna darker basally and laterally (Fig. 3). Antennae: antenomere 1 short, almost quadrate, not longer than 2; 2 slightly shorter than 1 and slightly narrower; 3 as long as 2 and slightly more slender; 4, 5 and 7 slightly shorter than 3; 6 and 8 still shorter, each approximately 1/2 the length of 3, not bead-like; antennal club distinct, antennomeres 9 and 10 almost quadrate, not noticibly transverse (Fig. 1.2). Body: width/length ratio, 0.43; length, 1.4���1.5 mm., Published as part of Majka, Christopher, Johnson, Colin & Langor, David, 2010, Contributions towards an understanding of the Atomariinae (Coleoptera, Cryptophagidae) of Atlantic Canada, pp. 37-63 in ZooKeys 35 (35) on page 46, DOI: 10.3897/zookeys.35.318, http://zenodo.org/record/576616, {"references":["Casey TL (1900) Revision of the North American Cryptophagidae. Journal of the New York Entomological Society 8: 75 - 128.","Leng CW (1920) Catalogue of the Coleoptera of America north of Mexico. John D. Sherman, Jr., Mount Vernon, New York, 470 pp.","Downie NM, Arnett RH, Jr (1996) The Beetles of Northeastern North America. Sandhill Crane Press. Gainsville, Florida, 1721 pp."]}

The Atomariinae (Coleoptera: Cryptophagidae) fauna of Atlantic Canada are surveyed. Ten species in the genus Atomaria are reported in the region, six of which, including Atomaria distincta Casey, Atomaria fuscata Schonherr, Atomaria lederi Johnson, Atomaria lewisi Reitter, Atomaria testacea Stephens, and Atomaria nigrirostris Stephens, are newly recorded in Canada. Atomaria lederi and A. lewisi are newly recorded in North America. Twenty-three new provincial records are reported. Atomaria curtula Casey, 1900 (syn. n.) is designated as a synonym of Atomaria lewisi Reitter; Atomaria ovalis Casey, 1900 (syn. n.) is designated as a synonym of Atomaria apicalis Erichson, 1846; and Atomaria saginata Casey, 1900 (syn. n.) is designated as a synonym of Atomaria fuscata Schonherr. A key to species is provided, as are species descriptions. Records of all species are given and their distribution in the region is plotted. The global distribution of all Holarctic and adventive species is briefly summarized, and the jurisdictions in North America where they have been reported are compiled. Habitat and bionomic information is also provided for all species, both from within the region, and from observations in European studies. The fauna is briefly discussed from a zoogeographic perspective; two species are Nearctic in origin, five may be Holarctic in distribution, and three others may be adventive Palaearctic species, although further evidence is required to confirm such preliminary assessments. Similarly, because of the paucity of information on this genus, further fieldwork is required to fully discern the distribution of these spe- cies in Atlantic Canada. Three are apparently saproxylic species characteristic of forested habitats; three are primarily grassland species; and the remaining four are eurytopic species found in a wide variety of forested and open habitats.