Search

Asceua Thorell, 1887 Type species. Asceua elegans Thorell, 1887, by monotypy. Diagnosis. For genus description and diagnosis, see Jocqué (1991)., Published as part of Sankaran, Pradeep M., 2023, Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae), pp. 381-405 in Zootaxa 5296 (3) on page 382, DOI: 10.11646/zootaxa.5296.3.4, http://zenodo.org/record/7984039, {"references":["Jocque, R. (1991) A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History, 201, 1 - 160."]}

Asceua cingulata (Simon, 1905) Figs 1–6, 17 Suffucia cingulata Simon, 1905: 174 (♀).— Gravely, 1921: 409, fig. 2d (♁ ♀). Type material. Lectotype (here designated) ♀ and paralectotypes 2 ♀♀ from INDIA: Puducherry (formerly Pondicherry), 11°56’N 79°48’E, 5 m a.s.l., date unknown, leg. M.M. Maindron, repository MNHN, Paris (AR9978), examined based on photographs. Other material examined. INDIA: Andaman Islands: Ross Island, 11°40’N 92°45’E, 41 m a.s.l., 29 March 1911, leg. Paiva: 2 ♀♀ (NZC-ZSI 2718 /H2). West Bengal: Hooghly, Serampore, 22°45’N 88°20’E, 9 m a.s.l., July 1919, leg. Drake: 1 ♀ (NZC-ZSI 2719 /H2). Tamil Nadu: Chennai (formerly Madras), 13°04’N 80°16’E, 4 m a.s.l., 12 March, leg. Ramuni Menon: 1 ♀ (only prosoma) (NZC-ZSI 2720 /H2). Odisha (formerly Orissa): Ganjam, Chilika Lake (= Chilka Lake), Barkuda Island (formerly part of Madras Presidency), 19°33’N 85°08’E, 13 m a.s.l., 3-19 August 1919, leg. F.H. Gravely: 2 ♁♁, 4 ♀♀ (NZC-ZSI 2721 /H2). Kerala: Malappuram, Nilambur, Canolly’s Plot, 11°16’06.17’’N 76°12’22.21’’E, 20 m a.s.l., 16 May 2013, on the ground, by hand, leg. M.S. Pradeep: 3 ♀♀, 1 subadult ♁, 1 juvenile (ADSH192); Ernakulam, Karimpani, Bhoothathankettu Forest Reserve, 10°08’N 76°40’E, 24 m a.s.l., 8 September 2013, on the ground, by hand, leg. M.S. Pradeep: 1 ♀ (ADSH193); Ernakulam, Illithodu, 10°11’55.94’’N 76°33’00.57’’E, 24 m a.s.l., 25 February 2014, on the ground, by hand, leg. M.S. Pradeep: 1 ♁, 2 ♀♀, 1 juvenile (ADSH194). Diagnosis. Males of A. cingulata are closely related to those of A. thrippalurense sp. nov. by the shape of retrolateral, ventral and prolateral tibial apophyses, cymbium with basomedial conical process extending between ventral and retrolateral apophyses and embolus with similar course of running, but can be distinguished by the smaller cymbial retrolateral fold with less modified basoretrolateral extremity (vs. highly modified in A. thrippalurense sp. nov.), anterolateral margin of cymbial fold less prominent (vs. prominent in A. thrippalurense sp. nov.), retrolateral process of conductor directed at 3-o’ clock ventrally and basally acute (vs. 2-o’ clock and basally rounded in A. thrippalurense sp. nov.), median apophysis with long proximal part (vs. short in A. thrippalurense sp. nov.), and embolus with distal loop (vs. absent in A. thrippalurense sp. nov.) (compare Figs 4A–C, 5A–D with Figs 9A–C, 10A–D). Females are closely related to the females of Asceua similis Song & Kim, 1997 in the general shape and orientation of copulatory ducts, but can be separated from the latter by epigyne with anterior plate (vs. absent in A. similis), and spherical spermathecae (vs. irregular in A. similis) (compare Figs 4D–F, 5E–F with Song & Kim 1997: figs 14–15). Redescription. Male in alcohol (ADSH194) (Figs 2A–J). Body length 1.57. Carapace 0.87 long, 0.70 wide. Opisthosoma 0.74 long, 0.62 wide. Carapace, eye region, clypeus, chilum, chelicerae, endites, labium, sternum yellowish brown; leg segments creamy-white to yellowish brown; palp segments yellowish brown to brown; dorsal scutum brownish, opisthosoma shiny, greyish black with two pairs of large dull yellow patches (Fig. 2D), lateral and rear end black with creamy-white patches, streaks and dots (Figs 2C–D), venter creamy-white with yellow anterior part (Fig. 2B); spinnerets creamy-white (Fig. 2H); carapace, clypeus, chelicerae, margins of sternum, leg segments with black mottling. Carapace glossy; cephalic part without long black setae. Fovea short, longitudinal, straight, dark brown. Clypeus high (Fig. 2E). Chilum inverted triangular, unipartite (Fig. 2E). Cheliceral promargin with two tiny teeth, retromargin without tooth visible (Fig. 2F). Sternum glossy, shield-shaped with posterior end truncated, rebordered, provided with scattered black setae, with coxal and intercoxal extensions (Fig. 2G). Opisthosoma oval in dorsal view, dorsal scutum complete, without long black setae (Fig. 2D); rear and lateral opisthosoma with corrugations. Colulus and anal tubercle prominent (Figs 2C, H). Femur I without retrolateral striae; metatarsi II–III with distal preening brush (Fig. 2I); all metatarsi and tarsi without scopula (Fig. 2J); all tarsi with reduced claw tuft (Figs 2I–J). Eye diameters and interdistances: ALE 0.06, AME 0.08, PLE 0.07, PME 0.05; AME–ALE 0.03, AME– AME 0.05, AME–PME 0.05, ALE–PLE 0.04, PME–PLE 0.08, PME–PME 0.07. Clypeus height at AMEs 0.29, at ALEs 0.24. Chelicerae 0.25 long. Sternum 0.45 long, 0.44 wide. Measurements of palp and legs: palp 1.02 [0.35, 0.15, 0.10, 0.42], I 2.54 [0.66, 0.23, 0.61, 0.62, 0.42], II 2.10 [0.59, 0.23, 0.45, 0.50, 0.33], III 2.03 [0.58, 0.20, 0.41, 0.54, 0.30], IV 2.79 [0.72, 0.23, 0.62, 0.83, 0.39]. Leg formula: 4123. Spination of palp: femur do 2, patella spineless, tibia spineless, tarsus/cymbium spineless; legs: femora I–IV do 1; patellae I–IV spineless; tibiae I–IV spineless; metatarsi I–III spineless, IV plv 1 rlv 1; tarsi I–IV spineless. Palp (Figs 4A–C, 5A–D). Tibia with short retrolateral, broad ventral and narrow, finger-like prolateral apophyses (Figs 4A, C, 5A, C–D); RTA having triangular distal part (Fig. 5C); RTA and VTA enclose large concavity in which fits basomedial projection of cymbium (Figs 4C, 5C). Cymbium modified, with prominent cymbial retrolateral fold having less prominent anterolateral margin and less modified posteroretrolateral extremity (Figs 4B–C, 5B–D), with basomedial conical process extending between RTA and VTA (Fig. 5C). Tegulum broad, with posteromedian extension (Figs 4B, 5B). Median apophysis with broad proximal and long, slender, prolaterally oriented distal parts (Figs 4A–C, 5A–C), lying adjacent to retrolateral process of conductor (Figs 4B, 5B). Conductor broad, sclerotized, with retrolateral and mediolateral, blunt processes (Figs 4B, 5B). Embolus long, filiform, basal to tegulum, with short base, at first running anteriad along prolateral side of cymbium, then turning retrolaterad, then to posterolaterad forming distal loop, with blunt tip directed at 1-o’ clock ventrally (Figs 4A–B, 5A–B). Female in alcohol (ADSH194) (Figs 3A–I). Body length 2.00. Carapace 0.96 long, 0.68 wide. Opisthosoma 1.04 long, 0.88 wide. Habitus and details as in male except for the following: opisthosoma without scutum and corrugations (Fig. 3A); dorsum with three pairs of large patches, numerous tiny spots, and a single median patch at the rear end in front of anal tubercle, all creamy-white (Fig. 3A); laterals without stripes or dots (Fig. 3C); venter uniformly creamy-white (Fig. 3B). Palp segments yellowish brown to brown. Eye diameters and interdistances: ALE 0.06. AME 0.07. PLE 0.04. PME 0.05; AME–ALE 0.04. AME–AME 0.03. AME–PME 0.07. ALE–PLE 0.04. PME–PLE 0.08. PME–PME 0.07. Clypeus height at AMEs 0.25, at ALEs 0.21. Chelicerae 0.33 long. Sternum 0.45 long, 0.43 wide. Measurements of palp and legs: palp 0.85 [0.27, 0.16, 0.16, 0.26], I 2.28 [0.64, 0.21, 0.54, 0.53, 0.36], II 1.94 [0.55, 0.22, 0.40, 0.48, 0.29], III 2.00 [0.54, 0.22, 0.40, 0.53, 0.31], IV 2.70 [0.69, 0.22, 0.61, 0.79, 0.39]. Leg formula: 4132. Spination of palp: tibia pld 1; tarsus pl 2 pld 1 v 2; legs: metatarsi II–IV plv 1 rlv 1. Genitalia (Figs 4D–F, 5E–F). Epigyne with convex posterior margin, with an anteromedial, sclerotized, transversely oval plate (Figs 4D–E, 5E). Copulatory openings situated anteriorly behind epigynal plate, lying close to each other (Figs 4E, 5E). Copulatory ducts long, slender, with multiple windings (Figs 4F, 5F). Spermathecae small, globular, lying adjacent to each other, situated posteriorly (Figs 4F, 5F). Fertilization ducts long, narrow, diverging (Figs 4F, 5F). Variation. For checking variation in measurements, only the freshly collected specimens were considered. Female (n=6): body length 1.80–2.08. One of the female specimens collected from Malappuram presents a variation in the abdominal pattern as shown in Fig. 3J, and its epigyne also shows variation in the shape of the epigynal plate as shown in Fig. 4G. Remarks. All the specimens deposited in the NZC-ZSI, Kolkata are severely dried off due to bad preservation (Figs 6A–D). Distribution. India (Andaman Islands (Rose Island), Kerala (new record), Odisha, Puducherry, Tamil Nadu, West Bengal) (Gravely 1921; present data) (Fig. 17)., Published as part of Sankaran, Pradeep M., 2023, Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae), pp. 381-405 in Zootaxa 5296 (3) on pages 382-389, DOI: 10.11646/zootaxa.5296.3.4, http://zenodo.org/record/7984039, {"references":["Simon, E. (1905) Voyage de M. Maurice Maindron dans l'Inde meridionale (mai a novembre 1901). 7 me Memoire. Arachnides (1 re partie). Annales de la Societe Entomologique de France, 74 (1 - 2), 160 - 180.","Gravely, F. H. (1921) The spiders and scorpions of Barkuda Island. Records of the Indian Museum, Calcutta, 22, 399 - 421, pls. 17 - 19. https: // doi. org / 10.5962 / bhl. part. 1478","Song, D. X. & Kim, J. P. (1997) On seven new species of the family Zodariidae (Araneae) from China. Korean Arachnology, 13 (1), 7 - 17."]}

Asceua thrippalurense sp. nov. Figs 7–10, 17 Type material. Holotype ♁ (ADSH196) from INDIA: Kerala: Palakkad, Thrippalur, Pullodu, 10°38’16.58’’N 76°33’52.87’’E, 70 m a.s.l., 24 January 2016, on the ground, by hand, leg. M.S. Pradeep. Paratype: 2 ♀♀ (ADSH197), same data as for the holotype. Etymology. The specific epithet is an adjective and referring to Thrippalur, where the type locality of the new species is located. Diagnosis. Males are similar to A. cingulata, but can be distinguished from it as stated above (compare Figs 9A–C, 10A–D with Figs 4A–C, 5A–D). Females are closely related to the females of Asceua wallacei Bosmans & Hillyard, 1990 as both share spherical spermathecae and copulatory ducts with prominent windings near copulatory openings, but can be separated from the latter species by epigyne with anteromedian pore (vs. absent in A. wallacei), and without anteromedian process (vs. present in A. wallacei) (compare Figs 9D–F, 10E–F with Bosmans & Hillyard 1990: figs 54–55). Females of A. thrippalurense sp. nov. can easily be distinguished from the females of A. cingulata by the presence of longitudinal striae retrolaterally on femur I (Fig. 8H) which is absent in A. cingulata. Description. Male in alcohol (holotype, ADSH196) (Figs 7A–J). Body length 2.06. Carapace 1.09 long, 0.88 wide. Opisthosoma 0.97 long, 0.73 wide. Carapace, eye region, clypeus dark brown; chilum, chelicerae, endites, labium, sternum brown; leg and palp segments yellowish brown to creamy-white; dorsal and lateral opisthosoma black with creamy-white streaks on laterals, dorsal scutum shiny, brownish black (Fig. 7A), venter creamy-white with yellowish brown and greyish patches; spinnerets creamy-white with black shades (Fig. 7G). Carapace finely rugose; cephalic part provided with scattered few long black setae. Fovea short, longitudinal, straight, dark. Clypeus high (Fig. 7D). Chilum inverted triangular, unipartite (Fig. 7D). Cheliceral promargin with two tiny teeth, retromargin without tooth visible (Fig. 7E). Sternum rebordered, rugose, shield-shaped with posterior end truncated, provided with scattered greyish black setae, with coxal and intercoxal extensions (Fig. 7F). Opisthosoma oval, scutum covering 2/3 rd of dorsum area (Fig. 7A), covered with long black setae; rear and lateral opisthosoma with corrugations. Colulus and anal tubercle prominent (Figs 7A, G). Femur I retrolaterally provided with longitudinal striae almost along its entire length (Fig. 7H); metatarsi II–III with distal preening brush (Fig. 7I); all metatarsi and tarsi without scopula (Fig. 7J); all tarsi with reduced claw tuft (Figs 7I–J). Eye diameters and interdistances: ALE 0.08, AME 0.10, PLE 0.07, PME 0.05; AME–ALE 0.02, AME–AME 0.03, AME–PME 0.07, ALE–PLE 0.03, PME–PLE 0.08, PME–PME 0.09. Clypeus height at AMEs 0.32, at ALEs 0.30. Chelicerae 0.28 long. Sternum 0.54 long, 0.52 wide. Measurements of palp and legs: palp 1.36 [0.42, 0.17, 0.13, 0.64], I 3.22 [0.87, 0.27, 0.79, 0.82, 0.47], II 2.55 [0.73, 0.28, 0.54, 0.64, 0.36], III 2.46 [0.68, 0.26, 0.52, 0.67, 0.33], IV 3.54 [0.93, 0.28, 0.81, 1.08, 0.44]. Leg formula: 4123. Spination of palp: femur rld 1, patella do 1, tibia spineless, tarsus/cymbium spineless; legs: femora I–IV do 1; patellae I–IV spineless; tibiae I–IV spineless; metatarsus I spineless, II–IV plv 1 rlv 1; tarsi I–IV spineless. Palp (Figs 9A–C, 10A–D). Tibia with short retrolateral, broad ventral and narrow, finger-like prolateral apophyses (Figs 9A, C, 10A–D); RTA with prolaterally oriented distal part (Figs 9C, 10C); RTA and VTA enclose large concavity in which fits basomedial projection of cymbium (Figs 9C, 10C). Cymbium with large retrolateral fold having prominent anterolateral margin and highly modified basoretrolateral extremity (Figs 9B–C, 10B–C), with basomedial conical process extending between RTA and VTA (Figs 9C, 10C). Tegulum broad, with basomedian extension (Figs 9B, 10B). Median apophysis with short proximal and long, slender, prolaterally oriented distal parts (Figs 9B–C, 10A– C), lying adjacent to retrolateral process of conductor (Figs 9B, 10B). Conductor broad, sclerotized, with retrolateral and mediolateral, blunt processes (Figs 9B, 10B); retrolateral process anteriorly oriented (Figs 9B, 10B). Embolus long, filiform, with short embolar base originated basally to tegulum, at first running anteriad along prolateral side of cymbium, then turning retrolaterad, then to posterolaterad, then to anteroretrolaterad without forming distal loop, with blunt tip directed at 1-o’ clock ventrally (Figs 9A–B, 10A–B). Female in alcohol (paratype, ADSH197) (Figs 8A–J). Body length 2.46. Carapace 1.04 long, 0.88 wide. Opisthosoma 1.42 long, 1.00 wide. Habitus and details like male except for the following: leg and palp segments yellowish brown to light brown. Opisthosoma without scutum and corrugations (Fig. 8A); dorsum with three pairs of creamy-white patches, anterior C-shaped, median V-shaped and posterior dot-like (Fig. 8A); laterals black and creamy-white without streaks; venter uniformly creamy-white with black at the rear end. Eye diameters and interdistances: ALE 0.07. AME 0.08. PLE 0.04. PME 0.05; AME–ALE 0.02. AME–AME 0.04. AME–PME 0.09. ALE–PLE 0.05. PME–PLE 0.10. PME–PME 0.09. Clypeus height at AMEs 0.33, at ALEs 0.28. Chelicerae 0.42 long. Sternum 0.58 long, 0.55 wide. Measurements of palp and legs: palp 1.04 [0.34, 0.20, 0.21, 0.29], I 2.97 [0.82, 0.30, 0.67, 0.72, 0.46], II 2.44 [0.66, 0.28, 0.52, 0.62, 0.36], III 2.51 [0.69, 0.28, 0.52, 0.67, 0.35], IV 3.42 [0.91, 0.30, 0.78, 1.01, 0.42]. Leg formula: 4132. Spination of palp: tibia pld 1; tarsus pl 1 pld 1 v 2; legs: metatarsi I–III spineless. Genitalia (Figs 9D–F, 10E–F). Epigyne with W-shaped posterior margin, with a central pore leading to copulatory openings (Figs 9D–E, 10E). Copulatory openings contiguous, situated inside the central pore. Copulatory ducts long, slender, with multiple windings (Figs 9F, 10F). Spermathecae large, globular, lying adjacent to each other, situated posteriorly (Figs 9F, 10F). Fertilization ducts long, narrow, diverging (Figs 9F, 10F). Distribution. Known only from the type locality (Fig. 17)., Published as part of Sankaran, Pradeep M., 2023, Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae), pp. 381-405 in Zootaxa 5296 (3) on pages 389-394, DOI: 10.11646/zootaxa.5296.3.4, http://zenodo.org/record/7984039, {"references":["Bosmans, R. & Hillyard, P. (1990) Spiders of the family Zodariidae from Sulawesi, Indonesia (Arachnida: Araneae: Zodariidae). Bulletin of the British Arachnological Society, 8, 147 - 160."]}

The spider genus Asceua Thorell, 1887 contains 34 species, almost entirely limited to Indochina, India, Sri Lanka and China, with a regional distribution. Eleven species of Asceua are currently only known from China, five of them are described only from one sex. Two new spider species of the genus Asceua are reported from China, A. haocongi sp. n. (♂♀, Hainan) and A. zijin sp. n. (♂♀, Jiangsu). Photos and a morphological description of the new species are provided.

Asceua digitata sp. nov. Figures 4, 5, 9 Type material. Holotype ♂: CHINA, Yunnan Province: Longyang County, Bawan Township, Nankang forest station, 24.82260ºN, 98.78201ºE, 2060 m, 23 May 2005, Heng-mei Yan and Ke-ji Guo leg. (GKJ018) Etymology. The specific epithet is derived from the Latin “ digitata ” (finger-shaped), referring to the fingershaped ventral ramus of RTA, adjective. Diagnosis. The male of this new species resembles males of A. menglun Song & Kim, 1997 (see Song & Kim, 1997, figs 11, 12) and A. similis Song & Kim, 1997 (see Song & Kim, 1997, figs 16, 17) in having a bifurcate VTA and a V-shaped RTA, but can be distinguished from them by: (1) dorsal ramus of RTA much shorter than ventral ramus in retrolateral view in A. digitata sp. nov. (Fig 5B) (vs. dorsal ramus longer than ventral ramus); (2) median apophysis spoon-shaped, distal end located in the inner side of cymbium in ventral view in A. digitata sp. nov. (Fig 5C) (vs. hook-shaped, distal end beyond the lateral margin of cymbium). Description. Male (holotype) (Fig. 4). Total length 3.47; carapace 1.60 long, 1.12 wide; abdomen 1.66 long, 1.26 wide. Carapace brown, with a V-shaped dark patch medianly; fovea and radial grooves distinct, cervical grooves indistinct. Clypeus 0.37 high. Eye sizes and interdistances:AME 0.07, ALE 0.09, PME 0.08, PLE 0.10; AME–AME 0.04, AME–ALE 0.06, PME–PME 0.10, PME–PLE 0.11, MOA 0.28 long, anterior width 0.18, posterior width 0.29. Chelicerae yellowish brown, with two promarginal teeth and with one retromarginal tooth. Endites yellow. Labium yellowish brown, triangular. Sternum yellowish brown, edge slightly darker. Legs yellow. Measurements of legs: I, 5.30 (1.49, 1.79, 1.28, 0.74); II, 4.34 (1.22, 1.44, 1.08, 0.60); III, 4.22 (1.21, 1.33, 1.11, 0.57); IV, 5.94 (1.55, 1.83, 1.75, 0.81). Leg formula: 4123. Abdomen oval, dorsum dark brown, median portion with two pairs of light patches medianly, posterior ones much smaller than anterior ones, posterior portion with three transversal light bands medianly; venter yellow, with dense field of short setae anteriorly, a dark chevron posteriorly and one pair of longitudinal dark patches laterally. Spinnerets pale yellow. Palp (Fig. 5). Tibia with two apophyses and a dentation medially; VTA bifurcate, all rami thumb-shaped; RTA bifurcate, ventral ramus longer and finger-shaped, dorsal ramus shorter and dentiform. Median apophysis with two lateral processes. Conductor membranous, with expanded tip and a tooth basally. Embolus slender and whip-shaped, distal portion cross over the conductor in retrolateral view extending clockwise. Female unknown. Distribution. Known only from the type locality (Fig. 9)., Published as part of Li, Song-Lin, Liu, Ping & Peng, Xian-Jin, 2022, Three new species of the genus Asceua Thorell, 1887 (Araneae: Zodariidae) from Yunnan Province, China, pp. 56-68 in Zootaxa 5190 (1) on pages 61-63, DOI: 10.11646/zootaxa.5190.1.2, http://zenodo.org/record/7119793, {"references":["Song, D. X. & Kim, J. P. (1997) On seven new species of the family Zodariidae (Araneae) from China. Korean Arachnology, 13 (1), 7 - 17."]}

Genus Asceua Thorell, 1887 Type species: Asceua elegans Thorell, 1887, Published as part of Li, Song-Lin, Liu, Ping & Peng, Xian-Jin, 2022, Three new species of the genus Asceua Thorell, 1887 (Araneae: Zodariidae) from Yunnan Province, China, pp. 56-68 in Zootaxa 5190 (1) on page 57, DOI: 10.11646/zootaxa.5190.1.2, http://zenodo.org/record/7119793

Asceua calciformis sp. nov. Figures 1–3, 9 Type material. Holotype ♂: CHINA, Yunnan Province, Tengchong County: Jietou Township, Datang Village, 25.73947ºN, 98.69630ºE, 2010 m, 16 May 2006, D. H. Kavanaugh and R. L. Brett leg. (DHK-2006-025C-01). Paratypes: 1♂ 2♀, same data as holotype (DHK-2006-025C-02); 1♂ 2♀, Jietou Township, Datang Village, Longtang River, papaya orchard, 25.75720°N, 98.69459°E, 2078 m, 16 May 2006, X. J. Peng, X. P. Wang and P. Hu leg. (Peng060516); 1♂ 1♀, Longjiang Township, Xiaoheishan Nature Reserve, 24.82888°N, 98.76001°E, 2020 m, 26 May 2005, C. Griswold and D. H. Kavanaugh leg. (CGY127). Etymology. The specific epithet is the combination of the Latin “ calceamen ” (shoe) and “ formis ” (form), referring to the shoe-shaped median apophysis, adjective. Diagnosis. The males of this new species resembles that of A. adunca Zhang & Zhang, 2018 (see Zhang & Zhang, 2018a, figs 1–3) in having similar embolic course, but can be distinguished by: (1) RTA longer than VTA, distal end bifurcated in retrolateral view in A. calciformis sp. nov. (Fig. 2B) (vs. RTA as long as VTA, distal end not bifurcated); (2) median apophysis shoe-shaped in ventral view in A. calciformis sp. nov. (Fig. 2C) (vs. hookshaped); (3) conductor terminal expanded in retrolateral view in A. calciformis sp. nov. (Fig. 2B) (vs. not expanded). The females of this new species resembles that of A. maculosa Logunov, 2010 (see Logunov, 2010: figs 4, 5), A. menglun Song & Kim, 1997 (see Song & Kim, 1997, figs 9, 10) and A. similis Song & Kim, 1997 (see Song & Kim, 1997, figs 14, 15) in having similar course of copulatory ducts, but can be easily distinguished from them by epigyne without atrium in A. calciformis sp. nov. (Fig. 3A) (vs. with atrium). Description. Male (holotype) (Fig. 1A, B). Total length 3.06; carapace 1.61 long, 1.32 wide; abdomen 1.43 long, 1.10 wide. Carapace brown, with a V-shaped dark patch medianly; fovea and radial grooves distinct, cervical grooves indistinct. Clypeus 0.46 high. Eye sizes and interdistances: AME 0.06, ALE 0.06, PME 0.08, PLE 0.11; AME–AME 0.03, AME–ALE 0.05, PME–PME 0.09, PME–PLE 0.14, MOA 0.33 long, anterior width 0.15, posterior width 0.24. Chelicerae brown, with two promarginal teeth and with one retromarginal tooth. Endites yellow. Labium yellowish brown, triangular. Sternum yellowish brown, edge slightly darker. Legs pale yellow. Measurements of legs: I, 5.14 (1.39, 1.69, 1.30, 0.76); II, 4.26 (1.24, 1.32, 1.06, 0.64); III, 4.22 (1.18, 1.33, 1.11, 0.60); IV, 5.97 (1.66, 1.79, 1.77, 0.75). Leg formula: 4123. Abdomen oval, dorsum black, anterior portion with two pairs of light patches medianly, posterior portion with three transversal light bands medianly; venter yellow, with a dark chevron posteriorly and one pair of dark patches laterally. Spinnerets pale yellow. Palp (Fig. 2). Tibia with two apophyses; VTA trifurcate, all rami with blunt tip; RTA much longer than wide, distal end with two protuberances, forcipate, dorsal side with two small teeth. Median apophysis shoe-shaped, with acerose upper ramus and wide lower ramus. Conductor membranous and overlapped with distal portion of embolus, with expanded tip and a tooth basally. Embolus slender and whip-shaped, extending clockwise. Female (one paratype of DHK-2006-025C-02) (Fig. 1C, D). Total length 4.13; carapace 1.79 long, 1.21 wide; abdomen 2.34 long, 1.70 wide. Clypeus 0.46 high. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.09, PLE 0.11; AME–AME 0.05, AME–ALE 0.05, PME–PME 0.11, PME–PLE 0.14. MOA 0.35 long, anterior width 0.18, posterior width 0.29. Measurements of legs: I, 4.71 (1.33, 1.55, 1.13, 0.70); II, 3.84 (1.23, 1.24, 0.86, 0.51); III, 4.17 (1.16, 1.32, 1.16, 0.53); IV, 5.64 (1.47, 1.79, 1.67, 0.71). Leg formula: 4132. Abdomen lighter than male. Other color and pattern same as in male. Epigyne (Fig. 3). Epigynal plate about two times wider than long, without pocket. Copulatory openings located on the median portion of epigyne. Copulatory ducts intertwined. Spermathecae tubular, almost as wide as copulatory ducts. Distribution. Known only from the type locality (Fig. 9)., Published as part of Li, Song-Lin, Liu, Ping & Peng, Xian-Jin, 2022, Three new species of the genus Asceua Thorell, 1887 (Araneae: Zodariidae) from Yunnan Province, China, pp. 56-68 in Zootaxa 5190 (1) on page 57, DOI: 10.11646/zootaxa.5190.1.2, http://zenodo.org/record/7119793, {"references":["Zhang, B. S. & Zhang, F. (2018 a) Two new species of the spider family Zodariidae from Laos. Zootaxa, 4459 (2), 285 - 300. https: // doi. org / 10.11646 / zootaxa. 4459.2.4","Logunov, D. V. (2010) New species of Zodariidae (Arachnida: Aranei) from Vietnam. Arthropoda Selecta, 19 (4), 249 - 260. https: // doi. org / 10.15298 / arthsel. 19.4.05","Song, D. X. & Kim, J. P. (1997) On seven new species of the family Zodariidae (Araneae) from China. Korean Arachnology, 13 (1), 7 - 17."]}

Asceua forcipiformis sp. nov. Figures 6–9 Type material. Holotype ♂: CHINA, Yunnan Province: Tengchong County, Jietou Township, Longtang River, 25.74622ºN, 98.69612ºE, 2030 m, 15 May 2006, D. H. Kavanaugh and D. Z. Dong leg. (DHK-2006-024-01). Paratypes: 16♂ 5♀, same data as holotype (DHK-2006-024-02); 12♂ 4♀, Longling County, Longjiang Township, Xiaoheishan Nature Reserve, 24.82888°N, 98.76001°E, 2020 m, 26 May 2005, Charles Griswold and D. H. Kavanaugh leg. (CGY127); 6♂ 1♀, Lonyang County, Bawan Township, Nankang Col, 24.83178°N, 98.76472°E, 2180 m, 22 May 2005, Charles Griswold and D. H. Kavanaugh leg. (CGY114). Etymology. The specific epithet is the combination of the Latin “ forceps ” (forceps) and “ formis ” (form), referring to the forcipate distal end of RTA, adjective. Diagnosis. The males of this new species resembles that of A. maculosa Logunov, 2010 (see Logunov, 2010, figs 1–3, 6) in having a strong median apophysis and a bifurcate VTA, but can be distinguished by: (1) tibia with DTA in A. forcipiformis sp. nov. (Fig. 7B) (vs. without DTA); (2) RTA with two rami in A. forcipiformis sp. nov. (Fig. 7F) ( vs. uniramous); (3) median apophysis without the spur in A. forcipiformis sp. nov. (Fig. 7C) (vs. with spur); (4) distal portion of embolus far away from the margin of genital bulb in ventral view in A. forcipiformis sp. nov. (Fig. 7C) (vs. covers the genital bulb). The females of this new species resembles that of A. maculosa Logunov, 2010 (see Logunov, 2010, figs 4, 5, 7) in having similar course of copulatory ducts and two pockets, but can be easily distinguished by: (1) the top of copulatory ducts located medially in A. forcipiformis sp. nov. (Fig. 8B) (vs. located laterally); (2) epigyne with two atria in A. forcipiformis sp. nov. (Fig. 8A) (vs. with one atrium). Description. Male (holotype) (Fig. 6A, B). Total length 3.85; carapace 1.96 long, 1.52 wide; abdomen 2.90 long, 1.32 wide. Carapace brown, with a V-shaped dark patch medianly; fovea and radial grooves distinct, cervical grooves indistinct. Clypeus 0.46 high. Eye sizes and interdistances: AME 0.06, ALE 0.07, PME 0.09, PLE 0.10; AME–AME 0.06, AME–ALE 0.06, PME–PME 0.13, PME–PLE 0.15, MOA 0.34 long, anterior width 0.18, posterior width 0.31. Chelicerae yellowish brown, with two promarginal teeth and without retromarginal tooth. Endites yellow. Labium brown, triangular. Sternum yellowish brown, edge slightly darker. Legs yellow. Measurements of legs: I, 5.23 (1.44, 1.75, 1.30, 0.74); II, 4.48 (1.33, 1.52, 1.03, 0.60); III, 4.47 (1.31, 1.41, 1.23, 0.52); IV, 6.28 (1.73, 1.90, 1.85, 0.80). Leg formula: 4123. Abdomen oval, dorsum black, anterior portion with three pairs of light patches medianly, posterior portion with three transversal light bands medianly; venter yellow, with dense field of short setae anteriorly, with a dark chevron posteriorly and one pair of dark patches laterally. Spinnerets pale yellow. Palp (Fig. 7). Tibia with three apophyses and a membranous lamella medially; VTA bifurcate, ventral ramus thicker with truncated tip, lateral ramus thinner with blunt tip in ventral view; RTA stronger and forcipate; DTA taper. Median apophysis leaf-shaped, distal end slightly curved. Conductor membranous, far away from distal portion of embolus in ventral view, with expanded tip and a tubercule basally. Embolus slender and whip-shaped, extending clockwise. Female (one paratype of DHK-2006-024-02) (Fig. 6C, D). Total length 4.12; carapace 1.77 long, 1.41 wide; abdomen 2.21 long, 1.71 wide. Clypeus 0.07 high. Eye sizes and interdistances: AME 0.09, ALE 0.09, PME 0.11, PLE 0.12; AME–AME 0.04, AME–ALE 0.06, PME–PME 0.11, PME–PLE 0.12. MOA 0.40 long, anterior width 0.20, posterior width 0.30. Measurements of legs: I, 4.66 (1.31, 1.66, 1.09, 0.60); II, 3.89 (1.10, 1.39, 0.96, 0.44); III, 4.06 (1.06, 1.35, 1.14, 0.51); IV, 5.82 (1.51, 1.84, 1.77, 0.70). Leg formula: 4132. Other color and pattern same as in male. Epigyne (Fig. 8). Epigynal plate about two times wider than long, with two triangular pockets anteriorly. Atria subcircular, located anteriorly. Copulatory openings located on the inner sides of atria. Copulatory ducts intertwined, original portion thicker. Spermathecae invisible, covered by copulatory ducts. Distribution. Known only from the type locality (Fig. 9)., Published as part of Li, Song-Lin, Liu, Ping & Peng, Xian-Jin, 2022, Three new species of the genus Asceua Thorell, 1887 (Araneae: Zodariidae) from Yunnan Province, China, pp. 56-68 in Zootaxa 5190 (1) on page 63, DOI: 10.11646/zootaxa.5190.1.2, http://zenodo.org/record/7119793, {"references":["Logunov, D. V. (2010) New species of Zodariidae (Arachnida: Aranei) from Vietnam. Arthropoda Selecta, 19 (4), 249 - 260. https: // doi. org / 10.15298 / arthsel. 19.4.05"]}

Three new species of the genusAsceuaThorell, 1887, from the natural forests of Malaysia, are described asAsceuabifurcasp. n.(♂♀),A.curvasp. n.(♂), andA.trimaculatasp. n.(♀). The genusAsceuais reported from Malaysia for the first time.

Genus Asceua Thorell, 1887 Type species: Asceua elegans Thorell, 1887, Published as part of Zhang, Bao-Shi & Zhang, Feng, 2018, Two new species of the spider Family Zodariidae from Laos, pp. 285-300 in Zootaxa 4459 (2) on page 286, DOI: 10.11646/zootaxa.4459.2.4, http://zenodo.org/record/1458618

Asceua torquata (Simon, 1909) (Figs 7–8) Storena torquata Simon, 1909: 79. Asceua torquata: Ono, 2004: 68, f. 6-7; Zhang, Zhang & Jia, 2012: 66, f. 12-18. Material examined. LAOS, Champasak Province: 3 males (Z-Laos-11-31 - Z-Laos-11-33) and 1 female (Z-Laos- 11-34), Tad Etu (N 15°11', E 106°06'), Pakse City, 18 Nov. 2011, Z.Z. Gao leg. Diagnosis. Asceua torquata can be distinguished from other Asceua species by the wide retrolateral prong of tibial apophysis, the dorsal pattern on the opisthosoma and the details of genitalia (Figs 7A–D, 8A–E). Description. See Ono (2004) and Zhang et al. (2012) Distribution. Laos (Champasak), China, Vietnam., Published as part of Zhang, Bao-Shi & Zhang, Feng, 2018, Two new species of the spider Family Zodariidae from Laos, pp. 285-300 in Zootaxa 4459 (2) on page 294, DOI: 10.11646/zootaxa.4459.2.4, http://zenodo.org/record/1458618

Asceua adunca sp. nov. (Figs 1���3) Type material. Holotype male (Z-Laos-11-21), LAOS, Champasak Province: Pakse City, Tad Etu (N 15��11', E 106��06'), 17 Nov. 2011, Z.Z. Gao leg. Paratypes: 3 males (Z-Laos-11-22 - Z-Laos-11-24) and 6 females (Z-Laos- 11-25 - Z-Laos-11-30), same data as holotype; 2 males (Z-Laos-11-60,Z-Laos-11-61) and 2 females (Z-Laos-11- 62, Z-Laos-11-63), Pakse City, Tad Etu (N 15��11', E 106��06'), 21 Nov. 2011, Z.Z. Gao leg. Diagnosis. The males of Asceua adunca sp. nov. can be distinguished from other Asceua species by the combination of the following characters: membranous conductor long and parallel to the terminal part of embolus, the median apophysis with a very long and thin hook-like extension; the shape of tegulum, the RTA bifurcate (Figs 2A���C, 3A���C). Asceua adunca sp. nov. resembles to Asceua quadrimaculata Zhang, Zhang & Jia, 2012 by the shape of tegulum and the RTA bifurcate, two small epigynal copulatory openings and a small central epigynal pocket, but it differs from the latter by the hook-like extension of the median apophysis, the two distant copulatory openings and the smaller spermathecae (Figs 2D, E, 3D, E). Etymology. The specific name is from Latin word ��� adunca ���, in reference to the hook-like extension of the median apophysis; adjective. Description. Male (holotype): Total length 2.03: prosoma 1.07 long, 0.73 wide; opisthosoma 0.82 long, 0.70 wide. Carapace (Fig. 1A) longer than wide, brown, median part with faint black patch in front of black fovea, margins black, thorax elevated, tegument smooth and shiny. Clypeus 0.24 high, brown. Anterior eye row slightly procurved, posterior eye row strongly procurved in dorsal view. Each eye ringed with black. Eye diameters: AME 0.08, ALE 0.08, PME 0.07, PLE 0.09. Eye interdistances: AME���AME 0.02, AME���ALE 0.08, ALE���ALE 0.30, AME���PME 0.07, PME���PME 0.08, PME���PLE 0.08, PLE���PLE 0.41, ALE���PLE 0.07. MOQ 0.22 long, anterior width 0.18, posterior width 0.21. Chilum poorly defined. Mouthpart (Fig. 1B): chelicerae brown, with two anterior and one posterior teeth on margins of fang furrows, terminal part with row of black scopulae, fangs short; endites brown, apices paler and provided with dense black scopula; labium triangular, 0.16 long, 0.15 wide, brown, median part with semi-orbicular dark band. Sternum (Fig. 1B) 0.51 long, 0.50 wide, brown, lateral margin dark, provided with sparse black setae, its lateral margin with inter- and intra-coxal triangles. Legs (Figs 1A, B) yellow-brown, each femur, patella, tibia, metatarsus with some black patches at extremities. Leg measurements: leg?3.10 (0.86 + 0.53 + 0.50 + 0.76 + 0.45),? 2.42 (0.67 + 0.29 + 0.44 + 0.65 + 0.37),? 1.63 (0.63 + 0.17 + 0.21 + 0.38 + 0.24),? 3.19 (0.89 + 0.18 + 0.57 + 1.05 + 0.50). Leg formula: 4123. Opisthosoma (Figs 1A, B) oval, longer than wide, covered with black short hair. Dorsum of opisthosoma slightly sclerotized, dark brown, with three pairs of lighter brown patches, posterior one connected by narrow transversal median light band; anterior part of venter yellow. Spinnerets (Fig. 1B) yellow pale, laterally with two blackish patches. Male palp (Figs 2A���C, 3A���C): Median apophysis wide and strong, retrolaterally with long and thin hook-like extension; palpal tibia with strong ventral and retrolateral blunt process, retrolateral tibial apophysis wide, triangular and dorsal tibial apophysis finger-shaped. Cymbium with terminal spine and with pro- and retrolateral folds not reaching tip. Female. One of the specimens (Z-Laos-11-25) measured: Total length 2.74: prosoma 1.14 long, 0.85 wide; opisthosoma 1.61 long, 1.26 wide. Carapace brown. Clypeal height 0.20. Eye diameters: AME 0.07, ALE 0.06, PME 0.08, PLE 0.08. Eye interdistances: AME���AME 0.03, AME���ALE 0.04, ALE���ALE 0.35, AME���PME 0.16, PME���PME 0.10, PME���PLE 0.09, PLE���PLE 0.47, ALE���PLE 0.14. MOQ 0.31 long, front width 0.19, back width 0.23. Mouthpart (Fig. 1D): labium 0.19 long, 0.25 wide. Sternum 0.55 long, 0.57 wide. Leg measurements: I 2.67 (0.63 + 0.23+ 0.65 + 0.68 + 0.48), II 2.22 (0.55 + 0.25 + 0.54 + 0.57 + 0.31), III 2.60 (0.60 + 0.30 + 0.58 + 0.71 + 0.41), IV 3.70 (0.78 + 0.29 + 0.88 + 1.16 + 0.59). Leg formula: 4132. Dorsum of opisthosoma black, without dorsal scutum, with two pairs of white anterior patches, followed posteriorly by four transversal median bands; venter yellowish-white. Other color as in male (Figs 1C, D). Female genitalia (Figs 2D, E, 3D, E) with central epigynal pocket, copulatory openings situated in posterior part of epigyne; long and winding copulatory ducts visible through integument; spermathecae small and oval, situated posteriorly and well separated. Distribution. Laos (Champasak)., Published as part of Zhang, Bao-Shi & Zhang, Feng, 2018, Two new species of the spider Family Zodariidae from Laos, pp. 285-300 in Zootaxa 4459 (2) on pages 286-290, DOI: 10.11646/zootaxa.4459.2.4, http://zenodo.org/record/1458618

Asceua piperata Ono, 2004 (Figs 4���6) Asceua piperata Ono, 2004: 68, f. 1-5. Material examined. LAOS, Champasak Province: 1 male (Z-Laos-11-40) and 2 females (Z-Laos-11-41, Z-Laos- 11-42), Pakse City, Tad Etu (N 15��11', E 106��06'), 20 Nov. 2011, Z.Z. Gao leg. Diagnosis. Asceua piperata Ono, 2004 resembles to A. wallacei Bosmans & Hillyard, 1990 with the roughly triangular base of the embolus, the very long embolus following dorso-prolateral curvature of cymbium, the spherical and adjoining spermathecae and anteromedian epigynal pocket. However, it can be distinguished from the latter by the thin and bifurcate retrolateral tibial apophysis, the tip of cymbium without pointed lateral process, the tip of conductor not bifurcate and the course of the internal copulatory ducts (Figs 4E, 5A���E, 6A���E). Description. Male (Z-Laos-11-40): Total length 2.92: prosoma 1.55 long, 1.23 wide; opisthosoma 1.29 long, 1.03 wide. Carapace (Fig 4A) longer than wide, brown, thorax elevated, tegument smooth and shiny, median part with V-shaped black patch in front of black fovea. Clypeus 0.50 high, brown. Anterior eye row slightly procurved and posterior eye row strongly procurved in dorsal view. Each eye ringed with black. Eye diameters: AME 0.12, ALE 0.04, PME 0.11, PLE 0.10. Eye interdistances: AME���AME 0.07, AME���ALE 0.04, ALE���ALE 0.50, AME��� PME 0.06, PME���PME 0.11, PME���PLE 0.13, PLE���PLE 0.65, ALE���PLE 0.06. MOQ 0.29 long, anterior width 0.29, posterior width 0.31. Chilum single sclerite, triangular. Mouthpart (Fig 4B): chelicerae brown, with two anterior and one posterior teeth on margins of fang furrows, terminal part armed with row of black scopula; fangs short; endites brown, apices pale, furnished with dense black scopula; labium triangular, 0.25 long, 0.31 wide, brown. Sternum (Fig 4B) 0.69 long, 0.74 wide, brown, with sparse black setae, lateral margin with intracoxal triangles. Legs yellow, coxae whitish, each femur, patella, metatarsus with black patches at extremities, each tibia with black patches at base and extremities. Leg measurements:? 5.54 (1.40 + 0.35 + 1.31 + 1.79 + 0.69),? 4.30 (1.10 + 0.38 + 1.00 + 1.24 + 0.58),? 4.45 (1.11 + 0.48 + 1.04 + 1.26 + 0.56),? 6.09 (1.31 + 0.38 + 1.76 + 1.77 + 0.87). Leg formula: 4132. Opisthosoma (Figs 4A, B) oval, longer than wide, covered with grey short hair, with small rectangular dorsal scutum. Dorsum of opisthosoma black, with two pairs of anterior white patches, followed by two transversal median bands; anterior part of venter yellow, posterior part white and with some black patches. Spinnerets (Fig 4B) yellow brown, laterally with blackish patches. Male palp (Figs 4E, 5A���C, 6A���C): tibia with wide dorsal apophysis and bifurcate retrolateral apophysis, with large concavity between them, fitting by posterior, globose cymbial projection; tegulum with thin sharp posterior hook; conductor wide and semi-circular; embolus long, thread-like, at first following dorso-prolateral curvature of cymbium on mesal side of palp, then turning to lateral side of palp and following dorsal margin of conductor; cymbium tareperd apically, without terminal spine, with deep, sinuous pro-and lateral folds not reaching tip. Female. One of the specimens (Z-Laos-11-41) measured: Total length 3.06: prosoma 1.44 long, 1.04 wide; opisthosoma 1.38 long, 1.16 wide. Clypeal height 0.49. Eye diameters: AME 0.07, ALE 0.04, PME 0.11, PLE 0.13. Eye interdistances: AME���AME 0.07, AME���ALE 0.05, ALE���ALE 0.42, AME���PME 0.10, PME���PME 0.13, PME���PLE 0.12, PLE���PLE 0.61, ALE���PLE 0.03. MOQ 0.28 long, front width 0.25, back width 0.30. Labium 0.19 long, 0.28 wide. Sternum 0.61 long, 0.66 wide. Leg measurements: I 3.42 (0.91 + 0.37 + 0.72 + 0.92 + 0.50), II 3.10 (0.76 + 0.31 + 0.71 + 0.87 + 0.45), III 3.18 (0.85 + 0.31+ 0.75 + 0.81 + 0.46), IV 4.45 (1.13 + 0.37 + 1.03 + 1.36 + 0.56). Leg formula: 4132. Dorsum of opisthosoma without dorsal scutum and with two pairs of white anterior patches, followed by three transversal median bands (Fig 4C); venter white. Spinnerets (Fig 4D) yellow brown, ringed with blackish patches. Other color as in male (Figs 4C���D). Female genitalia (Figs 5D���E, 6D���E): copulatory openings situated in anterior part of epigyne, median part with short central epigynal pocket; long and winding copulatory ducts visible through integument; adjoining spermathecae big and spherical, situated posteriorly. Distribution. Laos (Champasak), Vietnam., Published as part of Zhang, Bao-Shi & Zhang, Feng, 2018, Two new species of the spider Family Zodariidae from Laos, pp. 285-300 in Zootaxa 4459 (2) on pages 290-294, DOI: 10.11646/zootaxa.4459.2.4, http://zenodo.org/record/1458618

51. Asceua longji Barrion-Dupo, Barrion & Heong, new species [Figure 51 A-E] FEMALE. Holotype female, total length 2.11 mm. Cephalothorax length 0.97 mm. width 0.77 mm. Abdomen length 1.16 mm, width 0.76 mm. Carapace. Reddish brown with black eye margins and a brown cross band in the foveal area, 1.23 x longer than wide. Cephalic elevated and higher than thoracic area. Chelicerae reddish brown, vertical and with a small fang. Promargin bears a small tooth and none in the retromargin. Pedipals yellowish brown except femur with gray tinge. Maxillae yellowish gray, longer than wide, slantly triangular with narrow apex converging, and broad basally. Sternum broadly heart-shaped, yellow except brownish rebordered margins. Anterior margins of sternum widely truncated and posterior tip narrowly blunt to truncate. Eyes. In two procurved rows of four each. AER slightly shorter than PER. Eye diameter (mm): AME (0.08)> PLE (0.07)> PME (0.06)> ALE (0.04). Eye separation (mm): PME-PME=PME-PLE (0.06)> AME-AME=AME-ALE (0.03). Clypeus height wide, 2.63 x AME diameter. Legs. Yellow brown with apical and basal black annulations in femora, tibiae and tips of metatarsi II and III.Metatarsi I-IV each with a long trichobothrium on apical one-fifth. Femora I-IV each with a subbasodorsal spine. Tibiae II and III equally long. Metatarsus I as long as III. Leg formula 4132. Abdomen. Black with an anterior pair of white longitudinal band (=dorsum sepia), white band present midlaterally extended to subposterior lateral end of abdomen, a pair of circular dots submedially and a triangular yellowish white band on posterior tip. Anal tubercle and spinnerets yellow. Venter yellow except black lateral margins of spinnerets. Epigynum forms a simple plate with a lip-like band in the center in uncleared state. Cleared epigynum has a U-shaped indention towards posterior epigynal margin, copulatory tube coiled posteriorly, and anteriorly forming copulatory openings on the anterolateral side. MALE. Unknown. Material Examined. Holotype female (coll. no. La 1), CHINA, Hainan Island, Chengmai, Laocheng, Longji hills, 24 April 2012, ATBarrion, JLACatindig and SCVillareal; paratype one female (coll. no. So 99), Panja town, Shuiku vill., Songtao Reservoir, 2 April 2011, ATBarrion, JLACatindig and SCVillareal. Diagnostic Features. Closely resembles in general morphology, Asceua elegans Thorell, 1887 but differs from the latter in the following (1) smaller body size, (2) abdominal markings where dorsum sepia has two parallel longitudinal bands, (3) epigynal structures, and (4) leg measurements. Etymology. Named after the type locality., Published as part of Alberto T. Barrion, Aimee Lynn A. Barrion-Dupo, Josie Lynn A. Catindig, Sylvia C. Villareal, Ducheng Cai, Qianhua Yuan & Kong Luen Heong, 2016, NEW SPECIES OF SPIDERS (ARANEAE) FROM HAINAN ISLAND, CHINA, pp. 1-103 in UPLB Museum Publications in Natural History 3 on pages 44-45, DOI: 10.5281/zenodo.269136

Asceua anding sp. nov. (Figs 8–11) Type material. Holotype Ƥ, CHINA, Hainan Province: Anding County (19 ° 31 ʹN, 110 ° 18 ʹE), 21 Aug. 2007, F. Zhang leg. Diagnosis. The female can be distinguished from other Asceua species by the dorsal pattern on the opisthosoma, the details of genitalia and the very long copulatory ducts (Figs 7–9). Etymology. The specific name is a noun in apposition taken from the type locality. Description. Female (holotype). Total length 2.70: prosoma 1.31 long, 0.90 wide; opisthosoma 1.35 long, 0.95 wide. Color. Carapace (Fig. 8) brown; median furrow black; radial furrow dark brown. Clypeus brown. Each eye ringed with black. Chelicerae, endites and labium yellow brown, apices of endites and labium yellowish. Sternum brown, lateral margin slightly dark. Legs brown. Dorsum of opisthosoma with a pair of anterior patches, followed by three transverse median bands and pre-anal spot; venter yellowish. Spinnerets yellowish. Prosoma (Fig. 8). Carapace longer than wide, thorax swell, tegument smooth and shiny. Both the eye rows procurved in dorsal view. Diameters of eyes: AME 0.05, ALE 0.05, PME 0.07, PLE 0.08. Interdistances of eyes: AME–AME 0.02, AME–ALE 0.03, ALE–ALE 0.36, PME–PME 0.07, PME–PLE 0.11, PLE–PLE 0.48, ALE–PLE 0.09. MOA 0.27 long, front width 0.18, back width 0.21. Clypeal height 0.32. Fangs of chelicerae short. Labium triangular, 0.27 long, 0.23 wide. Sternum 0.59 long, 0.50 wide and furnished with sparse black setae. Legs. Each femur with a dorsal spine. Measurements of legs: I: 3.34 (0.90, 0.27, 0.77, 0.86, 0.54), II: 2.44 (0.68, 0.27, 0.45, 0.59, 0.45), III: 2.75 (0.72, 0.27, 0.50, 0.81, 0.45), IV: 3.84 (0.95, 0.32, 0.81, 1.22, 0.54). Leg formula: 4132. Opisthosoma (Fig. 8). Oval, longer than wide, covered with short, black hairs, without dorsal scutum. Female genitalia (Figs 9–11). Copulatory openings situated in median part of epigynum; long and winding copulatory ducts visible through integument; spermathecae situated posteriorly, small, long oval and partly hidden in copulatory ducts. Distribution. China (Hainan).

Asceua torquata (Simon, 1909) (Figs 12���18) Storena torquata Simon, 1909: 79. Asceua torquata Ono, 2004: 68, f. 6���7 Material examined. CHINA, Hainan Province: 13, Jianfengling Mountain, 9 Nov. 2008, G.X. Han leg.; Guangxi Province: 1 ��, Wuming County (23 �� 10 ��N, 108 �� 12 ��E), Nanning City, 1 May 2006, Z.S. Guan leg.; Hunan Province: 3 ��, Dao County (25 �� 32 ��N, 110 �� 36 ��E), 26 May 2002, Z.S. Zhang leg. Diagnosis. Males can be distinguished from other Asceua species by the wide retrolateral prong of tibial apophysis. It differs from A. japonica and A. expugnatrix Jocqu��, 1995 by the shape of retrolateral prong of tibial apophysis and the dorsal pattern on the opisthosoma (Figs 12, 15). Females can be distinguished from other Asceua species by the dorsal pattern on the opisthosoma and the details of genitalia (Figs 16���18). Description. Male (from Jianfengling Mountain). Total length 3.38: prosoma 1.58 long, 1.13 wide; opisthosoma 1.62 long, 1.13 wide. Color. Carapace (Fig. 12) brown, elevated; median furrow black; radial furrow dark brown. Clypeus brown. Each eye ringed with black. Chelicerae, endites and labium brown, apices of endites and labium yellow. Sternum brown, lateral margin slightly dark. Legs brown. Dorsum of opisthosoma black, median with a brown band, lateral part with a pair of white patches followed by two white median patches, the first white median patch with a small black median patch; venter yellowish. Spinnerets yellowish. Prosoma (Fig. 12). Carapace longer than wide, thorax swell, tegument smooth and shiny. Both the eye rows procurved in dorsal view. Diameters of eyes: AME 0.10, ALE 0.10, PME 0.08, PLE 0.13. Interdistances of eyes: AME���AME 0.04, AME���ALE 0.03, ALE���ALE 0.51, PME���PME 0.09, PME���PLE 0.12, PLE���PLE 0.61, ALE��� PLE 0.03. MOA 0.35 long, front width 0.27, back width 0.30. Clypeal height 0.41. Fangs of chelicerae short. Labium triangular, 0.27 long, 0.23 wide. Sternum 0.77 long, 0.68 wide and furnished with sparse black setae. Legs. Each femur with a dorsal spine. Measurements of legs: I: 4.11 (1.04, 0.41, 0.90, 1.08, 0.68), II: 3.79 (0.99, 0.45, 0.77, 0.95, 0.63), III: 3.82 (0.99, 0.45, 0.81, 0.95, 0.62), IV: 4.56 (1.17, 0.50, 1.04, 1.26, 0.59). Leg formula: 4132. Opisthosoma (Fig. 12). Oval, longer than wide, covered with short, black hairs. Male palp (Figs 11, 13��� 15). Retrolateral prong of tibial apophysis wide, ventrolateral prong thin; cymbium narrowed, with typical huge lateral fold, and with a projection near the apex of cymbium; bulb with a sharply process pointing retrolateraliy; membranous conductor large; embolus threadlike. Female (from Wuming County). Total length 3.47: prosoma 1.53 long, 1.22 wide; opisthosoma 1.89 long, 1.26 wide. Prosoma. Both the eye rows slightly procurved in dorsal view. Diameters of eyes: AME 0.10, ALE 0.12, PME 0.10, PLE 0.12. Interdistances of eyes: AME���AME 0.03, AME���ALE 0.04, ALE���ALE 0.53, PME���PME 0.10, PME���PLE 0.13, PLE���PLE 0.66, ALE���PLE 0.04. MOA 0.36 long, front width 0.23, back width 0.30. Clypeal height 0.32. Labium 0.36 long, 0.36 wide. Sternum 0.81 long, 0.72 wide. Legs. Each femur with one dorsal spine. Measurements of legs: I: 4.10 (0.99, 0.45, 0.95, 0.99, 0.72), II: 3.65 (0.95, 0.45, 0.81, 0.90, 0.54), III. 3.61 (0.95, 0.45, 0.72, 0.99, 0.50), IV: 4.23 (1.08, 0.45, 1.08, 0.99, 0.63). Leg formula: 4123. Female genitalia (Figs 16���18). Copulatory openings situated in anterior part of epigynum; long and winding copulatory ducts visible through integument; spermathecae almost spherical, situated posteriorly and near each other. Distribution. China (Hainan, Guangxi, Hunan); Vietnam. Remark. While examining the Asceua specimens collected from southern China, we found some Asceua females captured from Guangxi and Hunan provinces, recognized them as A. torquata which described originally from Vietnam. In 2008, we also collected one Asceua male from Hainan Province, and with the same habitus and dorsal pattern on the opisthosoma as the females of A. torquata. In addition, Hunan, Guangxi and Hainan are near in geographical distribution, We therefore identified it as the male of A. torquata and describe it here., Published as part of Zhang, Bao-Shi, Zhang, Feng & Jia, Xiao-Mei, 2012, Two new species of the ant spider genus Asceua Thorell, 1887 (Araneae: Zodariidae) from China, pp. 62-68 in Zootaxa 3307 on page 66, DOI: 10.5281/zenodo.212786, {"references":["Ono, H. (2004) Spiders of the family Zodariidae (Araneae) from Dambri, Lam Dong Province, southern Vietnam. Bulletin of the National Science Museum, Tokyo (A), 30, 67 - 75.","Jocque, R. (1995) Notes on Australian Zodariidae (Araneae), I. New taxa and key to the genera. Records of the Australian Museum, 47, 117 - 140."]}

Asceua quadrimaculata sp. nov. (Figs 1���7) Type material. Holotype 3, CHINA, Hainan Province: Datian National Nature Reserve (19 ��09��N, 108 �� 47 ��E), 5 Aug. 2007, F. Zhang leg. Paratypes: 1 ��, same data as holotype; 2 ��, Bawangling Mountain (19 ��07��N, 109 ��04��E), 3 Aug. 2007, F. Zhang leg.; 1 ��, Bawangling Mountain, 21 May 2009, C. Zhang leg.; 13, 3��, Jianfengling Mountain (18 �� 37 ��N, 108 �� 42 ��E), 9 Nov. 2008, G.X. Han leg.; 23, 2��, Jianfengling Mountain, 28 May 2009, C. Zhang leg. Diagnosis. The male of the new species can be distinguished from other Asceua species by the wide median apophysis and long membranous conductor. It differs from A. jianfeng, A. menglun, A. similis and A. maculosa Logunov, 2010 by the shapes of median apophysis and retrolateral tibial apophysis, and the dorsal pattern on the opisthosoma (Figs 1���4). The female can be distinguished from other Asceua species by the central epigynal pocket, the dorsal pattern on the opisthosoma and the details of genitalia; it differs from A. piperata Ono, 2004 by the spermathecae well separated, and from A. jianfeng by the central epigynal pocket and the course of internal copulatory ducts (Figs 5���6). Etymology. The specific name is a nominative neuter singular of the Latin adjective quadrimaculatus, meaning with four dots and refers to the pattern of dorsal opisthosoma. Description. Male (holotype). Total length 1.76: prosoma 0.86 long, 0.63 wide; opisthosoma 0.88 long, 0.54 wide. Color. Carapace (Fig. 1) brown; median furrow black. Clypeus brown, median part dark brown. Each eye ringed with black. Chelicerae, endites and labium brown, apices of endites and labium yellow, median part of labium with a black patch. Sternum brown, lateral margin slightly dark. Legs brown, apices and bases of femora and tibia ringed with black. Dorsum of opisthosoma black, with an ovoid scutum, lateral part with two pairs of white patches; venter yellowish. Spinnerets yellowish. Prosoma (Fig. 1). Carapace longer than wide, tegument smooth and shiny. Both the eye rows slightly procurved in dorsal view. Diameters of eyes: AME 0.09, ALE 0.08, PME 0.07, PLE 0.08. Interdistances of eyes: AME���AME 0.03, AME���ALE 0.01, ALE���ALE 0.31, PME���PME 0.07, PME���PLE 0.06, PLE���PLE 0.39, ALE��� PLE 0.07. MOA 0.23 long, front width 0.20, back width 0.18. Clypeal height 0.18. Labium triangular, 0.17 long, 0.16 wide. Sternum 0.50 long, 0.50 wide and sparsely covered with black setae. Legs. Each femur with a dorsal spine. Measurements of legs: I: 2.76 (0.68, 0.27, 0.68, 0.63, 0.50), II: 2.03 (0.54, 0.18, 0.45, 0.50, 0.36), III: 2.22 (0.59, 0.23, 0.50, 0.54, 0.36), IV: 3.12 (0.72, 0.23, 0.77, 0.90, 0.50). Leg formula: 4132. Opisthosoma (Fig. 1). Oval, longer than wide, covered with short, black hairs. Dorsal scutum distinct. Male palp (Figs 2���4). Retrolateral tibial apophysis thin, digitiform, palpal tibia with an elevated ridge on the ventral and retrolateral side respectively; cymbium with typical huge retrolateral fold; bulb prolaterally with a blunt tubercle; median apophysis wide and strong; membranous conductor wide and long; embolus threadlike. Female. Paratype (from Datian National Nature Reserve, Hainan). Total length 2.07: prosoma 0.90 long, 0.59 wide; opisthosoma 1.13 long, 0.68 wide. Prosoma. Both eye rows slightly procurved in dorsal view. Diameters of eyes: AME 0.09, ALE 0.07, PME 0.08, PLE 0.08. Interdistances of eyes: AME���AME 0.02, AME���ALE 0.03, ALE���ALE 0.30, PME���PME 0.08, PME���PLE 0.07, PLE���PLE 0.36, ALE���PLE 0.07. MOA 0.22 long, front width 0.17, back width 0.20. Clypeal height 0.23. Labium 0.18 long, 0.16 wide. Sternum 0.41 long, 0.41 wide. Legs. Each femur with a dorsal spine. Measurements of legs: I: 2.12 (0.54, 0.18, 0.45, 0.54, 0.41), II: 1.85 (0.45, 0.18, 0.45, 0.41, 0.36), III. 1.94 (0.50, 0.18, 0.45, 0.45, 0.36), IV: 2.58 (0.59, 0.23, 0.54, 0.77, 0.45). Leg formula: 4132. Opisthosoma without dorsal scutum. Female genitalia (Figs 5���7). Median part with a central epigynal pocket, copulatory openings situated in anterior part of epigynum; long and winding copulatory ducts visible through integument; spermathecae small and spherical, situated posteriorly and well separated. Distribution. China (Hainan)., Published as part of Zhang, Bao-Shi, Zhang, Feng & Jia, Xiao-Mei, 2012, Two new species of the ant spider genus Asceua Thorell, 1887 (Araneae: Zodariidae) from China, pp. 62-68 in Zootaxa 3307 on pages 62-64, DOI: 10.5281/zenodo.212786, {"references":["Logunov, D. V. (2010) New species of Zodariidae (Arachnida: Araneae) from Vietnam. Arthropoda Selecta, 19 (4), 249 - 260.","Ono, H. (2004) Spiders of the family Zodariidae (Araneae) from Dambri, Lam Dong Province, southern Vietnam. Bulletin of the National Science Museum, Tokyo (A), 30, 67 - 75."]}