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En este trabajo se presenta el listado florístico, incluyendo las formas de vida de las angiospermas presentes en las sabanas de San Miguel Allende (Campeche) y de Chacho Lugo (Yucatán). Se registraron 142 especies pertenecientes a 96 géneros y 36 familias, siendo las plantas herbáceas las que tienen la forma de vida dominante. El valor Ij obtenido fue de 0.169, lo que demuestra las notables diferencias en la composición florística de cada una de las sabanas, las cuales comparten sólo 24 de las 142 especies registradas. Las características geomorfológicas de cada lugar indican variaciones en la dinámica hidrológica, lo que se refleja en notables variaciones en la composición florística. Así, San Miguel Allende se clasifica como una sabana estacional, caracterizada por la presencia de géneros como Trachypogon y por la escasa presencia de representantes de la familia Cyperaceae; mientras que Chacho Lugo se clasifica como una sabana hiperestacional, donde abundan especies típicas de ambientes más húmedos (Cyperaceae, Eriocaulaceae y Lentibulariaceae). La presencia de endemismos como Echeandia campechiana, E. petenensis y Hampea trilobata, así como la baja presencia de plantas introducidas, muestran la importancia florística y ecológica de estos enclaves tan pobremente estudiados y afectados por las actividades antropogénicas., SEP (Secretaría de Educación Pública) del Gobierno de México, Promep (Programa para el Desarrollo Profesional Docente) del Gobierno de México, Secretaría de Relaciones Exteriores del Gobierno de México, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

This work has been funded by FONDECYT (Fondo Nacional de Desarrollo Científico y Tecnológico) from Chile by means of a postdoctoral fellowship of the first author (project Nº3170270), Two new Andean species belonging to the tribe Senecioneae (Compositae) are described herein, i.e., Dendrophorbium azoguesense from Ecuador and D. varicosum from Bolivia. The species Pentacalia todziae is recorded for the first time in Ecuador and the names Dendrophorbium onae and D. onae var. leonis synonymized to D. scytophyllum. Taxonomic discussions and diagnostic characters to differentiate each species are provided, as well as a distribution map and pictures of living plants when available., Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Coccoloba tunii is here described as a new species from Mexico. Morphological characters of the leaf, inflorescence, and fruit show discontinuities among populations of C. tunii and its relative, C. liebmannii. In addition, both species exhibit different geographical patterns, C. tunii is exclusive of the Central Depression and Plateau of Chiapas (Mexico) at mid elevations, while C. liebmannii ranges from Oaxaca to Colima on lowlands along the Pacific watershed., SEP (Secretaría de Educación Pública) del Gobierno de México, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Centaurea stuessyi is described as new to science from the eastern Iberian Peninsula, Spain. The range of this new species coincides with the zone of overlap of the ranges of two related species, C. antennata (subsp. antennata) and C. linifolia. The chromosome number found in the new species (2n = 44) reinforces its differences from C. linifolia (2n =22) and C. antennata (2n = 22). The new species is compared with related species, and a description is given of its ecological preferences., Ministerio de Educación y Ciencia, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

This work was partially financed by SEP-PROMEP “Fortalecimiento del Cuerpo Académico Diversidad de los Recursos Florísticos de Mesoamérica UADY-CA94”. One of the authors, I. Arnelas Seco, was the beneficiary of a postdoctoral grant (ARNITZ81031712) financed by S.R.E. of Mexican Government, A new species Neomillspaughia hondurensis (Polygonaceae) from Honduras is described and illustrated. This species is morphologically and ecollogicaly different from the other two currently known Neomillspaughia species, being more related to N. emarginata than to N. paniculata. A comprehensive comparison and a key for the three species are provided., Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

This work has been supported by a Spanish Government grant project CGL005-25122-E to J. A. Devesa, a co-funded Spanish Aragón Government and European Social Fund grant to the University of Zaragoza BIOFLORA research group, and a BBVA Foundation post-doctorate fellowship to E. Pérez-Collazos., Valerianella (cornsalad) is a taxonomically complex genus formed by 50–65 annual Holarctic species classified into at least four main sections. Carpological traits (sizes and shapes of achenes and calyx teeth) have been used to characterize its sections and species. However, the potential systematic value of these traits at different taxonomic ranks (from sections to species (and infraspecific taxa)) has not been tested phylogenetically yet. Here, we have assessed the evolutionary systematic value of Valerianella diagnostic carpological traits at different hierarchical ranks and have demonstrated their ability to separate taxa at the sectional level but not at species level for species of several species pairs. A total of 426 individuals (17 species, 4 sections) of Valerianella were analyzed using AFLP and plastid data. Genetic clusters, phylogenetic trees, and haplotype networks support the taxonomic classification of Valerianella at the four studied sectional levels (V. sects. Valerianella, Cornigerae, Coronatae, Platycoelae) but show admixture for ten taxa from five species pairs (V. locusta—V. carinata, V. coronata—V. pumila, V. multidentata—V. discoidea, V. dentata—V. rimosa, V. eriocarpa—V. microcarpa), which are not reciprocally monophyletic. Dating analyses indicate that the Valerianella sections are relatively old (mid-Miocene), while most species diverged in the Pliocene–Pleistocene. A new section Valerianella sect. Stipitae is described to accommodate the highly divergent and taxonomically distinct V. fusiformis type species. Taxonomic treatments that recognize the sectional ranks and that subsume the separate species of each species pair into single species represent a natural classification for Valerianella., Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Oritrophium yacuriense se describe como una nueva especie de los Andes del sur de Ecuador. Se utilizan caracteres morfológicos para diferenciarla de O. repens, especie similar que difiere principalmente en la forma de la hoja. La nueva especie crece en páramos herbáceos y húmedos del complejo lagunar del Parque Nacional Yacuri., Ministerio de Medio Ambiente (Ecuador), Universidad Técnica Particular de Loja, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

En este estudio se llevó a cabo una sinopsis del género Paspalum L. en la Península de Yucatán, México. Se reconocen 23 especies en el área de estudio, se proporcionan claves de identificación, descripciones morfológicas, mapas de distribución, así como información acerca de su fenología, hábitat y distribución. Se reconocen dos especies endémicas de la Península de Yucatán, P. mayanumy P. sparsum, localizadas en áreas abiertas de la selva baja caducifolia., Programa de Mejoramiento del Profesorado, Secretaría de Educación Pública, Gobierno de México (PROMEP), Secretaria de Relaciones Exteriores del Gobierno de México, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Este trabajo ha sido parcialmente financiado por la Universidad Técnica Particular de Loja (PY-1897). El Ministerio del Ambiente de Ecuador (MAE) otorgó los permisos de investigación para desarrollar el presente trabajo (MAE-DNB-CM-2015-0016). JC ha sido financiado por FONDECYT a través de una beca postdoctoral ―proyecto n.º 3170270., Oritrophium (Kunth) Cuatrec. is a neotropical genus with a disjunct distribution in the Andes (Bolivia, Colombia, Ecuador, Peru, and Venezuela), the center of Mexico, and the mountains of the Guiana Shield in Venezuela, with a remarkable diversity in Ecuador. We present a taxonomic study for Ecuador that represents the first modern and exhaustive revision of this genus in this country. We recognize eight species and four subespecies. We provide a dichotomous key, descriptions, synonyms, morphological and nomenclatural notes, distribution maps, and photographs of living plants. Three names are lectotypified and two names are neotypified., Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Coccoloba floresii is here described as a new species from Mexico. Morphological characters of the leaf, inflorescence, and fruit show discontinuities among populations of C. floresii and its relatives, C. barbadensis and C. cozumelensis. In addition, C. floresii is exclusive of the tropical dry forests of Central Depression and Plateau of Chiapas (Mexico) at high elevations., SEP (Secretaría de Educación Pública) del Gobierno de México, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

The morphological variability of Centaurea langei Nyman (Centaurea sect. Centaurea), an endemic to the north-central and north-west Iberian Peninsula, was studied. Seven subspecies with reasonably well-defined areas are recognized, of which C. langei subsp. dominguezii E. López, Devesa & Arnelas is described as new, and for the others the following combinations are proposed: C. langei subsp. exilis (Arènes)E. López, Devesa & Arnelas, C. langei subsp. coutinhoi (Franco)E. López, Devesa & Arnelas, C. langei subsp. geresensis (Arènes)E. López, Devesa & Arnelas, C. langei subsp. kheilii (Pau)E. López, Devesa & Arnelas, and C. langei subsp. rothmalerana (Arènes)E. López, Devesa & Arnelas. Nomenclatural and taxonomic synonyms are provided for each taxon, most of which are typified. For C. langei subps. dominguezii, the chromosome number was also studied (2n = 18)., Ministerio de Educación y Ciencia, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

The taxonomic boundaries of Centaurea L. species and infraspecific taxa are often blurred by hybridizations. However, counterbalancing genetic isolation contributes to maintaining taxonomic limits and fostering rapid speciation processes. Radiant (R) and non-radiant (NR) capitula are two major floral morphs present in Centaurea. However, it is unclear how gene flow affects the distribution of floral morphotypes across populations and taxa. We have investigated the taxonomic differentiation and potential effect of genetic isolation in R and NR populations of Iberian Centaurea sects. Jacea (Mill.) Pers. ex Dumort. and Lepteranthus (Neck. ex DC.) Dumort. A total of 510 individuals from 58 populations (12 taxa) were analyzed using 165 amplified fragment length polymorphism (AFLP) markers. Genetic diversity and structure parameters were estimated at the taxon, population, and floral morphotypic levels. We tested whether there was correlation between population pairwise fixation index (Fst) genetic distances and the inbreeding coefficient (Fis), taken as a surrogate of reproductive isolation between the R and NR morphotypes of each group, and also taking into account geographic distances, using distance-based redundancy analysis (dbRDA). We also performed isolation by distance (IBD) tests between different floral morphotypes, aiming to infer the impact of genetic isolation on evolutionary and taxonomic divergence of the individuals. The taxa of Centaurea sects. Jacea and Lepteranthus constitute independent evolutionary lineages. Most of the detected genetic clusters match the taxonomic circumscription proposed in our most recent treatments. Genetic diversity was highest in C. debeauxii Godr. & Gren. subsp. debeauxii and subsp. grandiflora (Gaudin ex Schübl. & G. Martens) Devesa & Arnelas and in C. jacea L. subsp. angustifolia (DC.) Gremli (R) and lowest in C. nevadensis Boiss. & Reut. (NR) within the Centaurea sect. Jacea clade, whereas it was highest in C. linifolia L. w, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Centaurea is one of the richest but less investigated genera of tribe Cardueae (Asteraceae). We reconstructed an updated phylogeny of Iberian species of Centaurea sects. Centaurea, Jacea, Lepteranthus and Phalolepis by sampling representatives from 48 taxa (~60% of the taxonomic variation). Phylogenetic trees and haplotypic networks were based on ITS and trnLF sequences, and Bayesian dating analysis on ITS data. We found two main lineages within this monophyletic group: (1) a W Mediterranean Centaurea and Phalolepis plus C. diffusa (CP) clade; and (2) an E Mediterranean – Eurosiberian Jacea and Lepteranthus (JL) clade. Our analyses provide new reconstructions for species of C. sects. Jacea and Lepteranthus that support new taxonomic circumscriptions. The inferred divergence times indicated succesive splits of the CP and JL lineages during the Miocene (e.g. CP/JL, Serravalian; Jacea/Lepteranthus, Tortonian; C. hyssopifolia, C. gr. janeri, C. gr. exarata, Messinian) and the Pliocene (C. linifolia/C. antennata). Interspecific hybridizations and polyploidizations likely took place during the Pleistocene (1.5–0.7 Ma) interglacials, when several hybrid lineages apparently originated (e.g. C. x decipiens). Allopatric speciation probably played an important role in the diversification of the Iberian Centaurea taxa, since the origins of several endemic species occurred during mid-late Pleistocene times (2.2–1.3 Ma) in several disjunct Iberian ranges., Ministerio de Ciencia e Innovación (MCIN), Gobierno de Aragón, Unión Europea. Fondo Europeo de Desarrollo Regional (European Regional Development Fund), Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

This work was possible thanks to funding from the Living Earth Collaborative (LEC) at Washington University in St. Louis, for the working group ‘A synthesis of patterns and mechanisms of diversity and forest change in the Andes: A global biodiversity hotspot’ (organized by JST, JAM, and SB), and by the German Centre for Integrative Biodiversity Research (iDiv), for the working group ‘sAndes: Tree diversity, composition and carbon storage in Andean tropical montane forests’ (organized by LC and MJM). The authors are grateful to Jens Kattge for making available the TRY data presented in this paper. AA-Q thanks the Herbario Nacional de Bolivia, the German Academic Exchange Service (DAAD) and DFG for funding (HE3041/20-1); to Ricardo Sonco and Marcelo Reguerin, Arely Palabral, Heike Heklau, and the Chulumani community. SB thanks the financial support of the Alexander von Humboldt Foundation through a Georg Foster Fellowship; SB, MB and HV acknowledge VLIR-UOS grants COFOREC (EC2018SIN223A103) and COFOREC II (EC2020SIN279A103). GBD was funded through a PhD grant by the Spanish Ministry of Education (MINEDU; FPU14/05303) and is indebted to Alex Nina, Jorge Armijos, Gonzalo Bañares, José Sánchez, Ángel Delso, Anselmo Vergaray, “Rosho” Tamayo, Reynerio Ishuiza, and national parks rangers for assistance in the field. CB thanks CONICET, Argentina, for a Doctoral Scholarship. LC and MJM were partly funded through projects CGL2013-45634-P, CGL2015-72431-EXP, CGL2016-75414-P, PID2019-105064GB-I00, and S2018/EMT-4 338. JGA and CBS acknowledge Luis Torres, Mara Paneghel, Iñigo Gómez, Maaike Pyck, Manuel Marca, Daniel Irygoin, Piher Maceda and Pontificia Universidad Católica de Perú (PUCP). JH thanks DFG (projects HO3296/2, HO3296/4, HO3296/6) and the Erasmus Go International Plus Program for funding and Katherine Angulo-Schipper, Michel Edelmann, Julius Joosten, Julia Falk, Roman Link, Johanna Lindemann, Stephanie Lorenz, Phillip Obst, Jaime Peña, Bernhard Schuldt, Julia Siegel an, We introduce the FunAndes database, a compilation of functional trait data for the Andean flora spanning six countries. FunAndes contains data on 24 traits across 2,694 taxa, for a total of 105,466 entries. The database features plant-morphological attributes including growth form, and leaf, stem, and wood traits measured at the species or individual level, together with geographic metadata (i.e., coordinates and elevation). FunAndes follows the field names, trait descriptions and units of measurement of the TRY database. It is currently available in open access in the FIGSHARE data repository, and will be part of TRY’s next release. Open access trait data from Andean plants will contribute to ecological research in the region, the most species rich terrestrial biodiversity hotspot., Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

This study was funded by the Spanish Aragon Government grant project LMP82 _21, the Spanish Aragon Government and European Social Fund Bioflora research group grant A01-20R, and the European and Spanish Government (SEPIE) Erasmus + KA107 mobility grant 2019-1-ES01-KA107-062605. MM-A was supported by a University of Zaragoza Bioflora research contract., The repeatome is composed of diverse families of repetitive DNA that keep signatures on the historical events that shaped the evolution of their hosting species. The cold seasonal Loliinae subtribe includes worldwide distributed taxa, some of which are the most important forage and lawn species (fescues and ray-grasses). The Loliinae are prone to hybridization and polyploidization. It has been observed a striking two-fold difference in genome size between the broad-leaved (BL) and fine-leaved (FL) Loliinae diploids and a general trend of genome reduction of some high polyploids. We have used genome skimming data to uncover the composition, abundance, and potential phylogenetic signal of repetitive elements across 47 representatives of the main Loliinae lineages. Independent and comparative analyses of repetitive sequences and of 5S rDNA loci were performed for all taxa under study and for four evolutionary Loliinae groups [Loliinae, Broad-leaved (BL), Fine-leaved (FL), and Schedonorus lineages]. Our data showed that the proportion of the genome covered by the repeatome in the Loliinae species was relatively high (average ∼ 51.8%), ranging from high percentages in some diploids (68.7%) to low percentages in some high-polyploids (30.7%), and that changes in their genome sizes were likely caused by gains or losses in their repeat elements. Ty3-gypsy Retand and Ty1-copia Angela retrotransposons were the most frequent repeat families in the Loliinae although the relatively more conservative Angela repeats presented the highest correlation of repeat content with genome size variation and the highest phylogenetic signal of the whole repeatome. By contrast, Athila retrotransposons presented evidence of recent proliferations almost exclusively in the Lolium clade. The repeatome evolutionary networks showed an overall topological congruence with the nuclear 35S rDNA phylogeny and a geographic-based structure for some lineages. The evolution of the Loliinae repeatome suggests, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

This study was funded by the Spanish Aragon Government LMP82-21 and the Spanish Aragon Government and European Social Fund Bioflora A01-17R research grants. The taxonomic study was funded by an Erasmus+ mobility grant 2019-1-ES01-KA107-062605 between the Universities of Zaragoza (Spain) and UTPL (Ecuador). M.F.M.-A. was supported by a University of Zaragoza—Santander Ph.D. fellowship., Allopolyploidy is considered a driver of diversity in subtribe Loliinae. We investigate the evolution and systematics of the poorly studied Mesoamerican and South American polyploid broad-leaved Festuca L. species of uncertain origin and unclear taxonomy. A taxonomic study of seven diagnostic morphological traits was conducted on a representation of 22 species. Phylogenomic analyses were performed on a representation of these supraspecific taxa and all other Loliinae lineages using separate data from the entire plastome, nuclear rDNA 45S and 5S genes, and repetitive DNA elements. F. subgen. Mallopetalon falls within the fine-leaved (FL) Loliinae clade, whereas the remaining taxa are nested within the broad-leaved (BL) Loliinae clade forming two separate Mexico–Central–South American (MCSAI, MCSAII) lineages. MCSAI includes representatives of F. sect. Glabricarpae and F. subgen. Asperifolia plus F. superba, and MCSAII of F. subgen. Erosiflorae and F. sect. Ruprechtia plus F. argentina. MCSAII likely had a BL Leucopoa paternal ancestor, MCSAI and MCSAII a BL Meso-South American maternal ancestor, and Mallopetalon FL, American I–II ancestors. Plastome vs. nuclear topological discordances corroborated the hybrid allopolyploid origins of these taxa, some of which probably originated from Northern Hemisphere ancestors. The observed data indicate rapid reticulate radiations in the Central–South American subcontinent. Our systematic study supports the reclassification of some studied taxa in different supraspecific Festuca ranks., Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Many studies have tried to assess the role of both deterministic and stochastic processes in community assembly, yet a lack of consensus exists on which processes are more prevalent and at which spatial scales they operate. To shed light on this issue, we tested two nonmutually exclusive, scale-dependent hypotheses: (1) that competitive exclusion dominates at small spatial scales; and (2) that environmental filtering does so at larger ones. To accomplish this, we studied the functional patterns of tropical montane forest communities along two altitudinal gradients, in Ecuador and Peru, using floristic and functional data from 60 plots of 0.1 ha. We found no evidence of either functional overdispersion or clustering at small spatial scales, but we did find functional clustering at larger ones. The observed pattern of clustering, consistent with an environmental filtering process, was more evident when maximizing the environmental differences among any pair of plots. To strengthen the link between the observed community functional pattern and the underlying process of environmental filtering, we explored differences in the climatic preferences of the most abundant species found at lower and higher elevations and examined whether their abundances shifted along the elevation gradient. We found (1) that greater community functional differences (observed between lower and upper tropical montane forest assemblies) were mostly the result of strong climatic preferences, maintained across the Neotropics; and (2) that the abundances of such species shifted along the elevational gradient. Our findings support the conclusion that, at large spatial scales, environmental filtering is the overriding mechanism for community assembly, because the pattern of functional clustering was linked to species’ similarities in their climatic preferences, which ultimately resulted in shifts in species abundances along the gradient. However, there was no evidence of competitive exclusion at more, Ministerio de Educación y Ciencia, Ministerio de Economía y Competitividad (MINECO), Universidad Rey Juan Carlos, Depto. de Biodiversidad, Ecología y Evolución, Fac. de Ciencias Biológicas, TRUE, pub

Oritrophium (Kunth) Cuatrec. is a neotropical genus with a disjunct distribution in the Andes (Bolivia, Colombia, Ecuador, Peru, and Venezuela), the center of Mexico, and the mountains of the Guiana Shield in Venezuela, with a remarkable diversity in Ecuador. We present a taxonomic study for Ecuador that represents the first modern and exhaustive revision of this genus in this country. We recognize eight species and four subespecies. We provide a dichotomous key, descriptions, synonyms, morphological and nomenclatural notes, distribution maps, and photographs of living plants. Three names are lectotypified and two names are neotypified., Oritrophium (Kunth) Cuatrec. es un género neotropical con distribución disyunta a lo largo de los Andes (Bolivia, Colombia, Ecuador, Perú y Venezuela), el centro de México y las montañas del Escudo Guayanés en Venezuela, con una notable diversidad en Ecuador. Presentamos un estudio taxonómico del género para Ecuador, el cual representa la primera revisión moderna y exhaustiva del género en este país. Se reconocen ocho especies y cuatro subespecies. Se incluye una clave taxonómica, descripciones, sinonimia, observaciones morfológicas y nomenclaturales, mapas de distribución y fotografías de las plantas en su hábitat. Tres nombres se han lectotipificado y otros dos se han neotipificado.

In this study, a synopsis of Paspalum L. was performed in the Yucatan Peninsula, Mexico. Twenty-three taxa are recognized in the study area, identification keys are provided, as well as morphological descriptions, distribution maps, habitat, phenology and distribution data. Two endemic species are recognized for the Yucatan Peninsula, P. mayanum and P. sparsum, which are found in open areas of tropical deciduous forest., En este estudio se llevó a cabo una sinopsis del género Paspalum L. en la Península de Yucatán, México. Se reconocen 23 especies en el área de estudio, se proporcionan claves de identificación, descripciones morfológicas, mapas de distribución, así como información acerca de su fenología, hábitat y distribución. Se reconocen dos especies endémicas de la Península de Yucatán, P. mayanumy P. sparsum, localizadas en áreas abiertas de la selva baja caducifolia.

Taxonomic review of Centaurea sect. Lepteranthus (Neck.) Dumort. (Asteraceae) in the Iberian Peninsula. A taxonomic review of Centaurea sect. Lepteranthus (Neck.) Dumort. in the Iberian Peninsula was done. Twelve taxa are recognized and a new combination is proposed, C. gallaecica (Laínz) Arnelas & Devesa, a taxon which is included in a different section, Jacea (Mill.) Pers. A key for the identification of taxa is made, also their morphological description, information on geographical distribution, their ecology, their phenology and their chromosome numbers are given., Revisión taxonómica de Centaurea sect. Lepteranthus (Neck.) Dumort. (Asteraceae) en la Península Ibérica. Se ha efectuado la revisión taxonómica de Centaurea sect. Lepteranthus (Neck.) Dumort. en la Península Ibérica. Se reconocen 12 táxones y se hace una nueva combinación: C. gallaecica (Laínz) Arnelas & Devesa, proponiéndose la inclusión de esta especie en la sect. Jacea (Mill.) Pers. Se aporta una clave para la dentificación de los táxones, así como su descripción morfológica e información sobre la distribución geográfica, apetencias ecológicas, fenología y número cromosómico.